identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03A587F0FF83FF94FEF2FE15A7FCFE64.text	03A587F0FF83FF94FEF2FE15A7FCFE64.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Varanus bushi	<div><p>Varanus bushi sp. nov.</p><p>Figures 1, 3–6</p><p>Holotype: Western Australian Museum R108999, adult male from Marandoo, Western Australia in 22 0 37’ S 118 0 08’ E. Collected on 20 June 1991 by Greg Harold. Specimen fixed in 10% formalin then stored in 70% ethanol. Liver sample stored in ­80 0 C ultrafreeze at WAM.</p><p>Paratypes: Three additional specimens from Marandoo: WAM R54230, an adult male collected on 25 October 1976; WAM R56834, an adult female collected in April 1977; and WAM R62171 a juvenile of undetermined sex collected on 28 February 1979.</p><p>Diagnosis: A small­bodied member of the subgenus Odatria distinguished from most others by the combination of a longitudinally striped and only moderately spinose tail, unkeeled head and body scales, non­overlapping ventral primary scales, and an absence of longitudinal streaks on throat. Distinguished from V. g i l l e n i by its slightly lesser average size, more elongate dorsal scales, more densely spotted venter and more irregularly spotted dorsum, less prominent linear patterning on the head and neck, and its more numerous presacral vertebrae, pedal subdigital lamellae and ventral scales. Males are further distinguished from V. gilleni by having hemipenes with an undivided inner hemibaculum. Distinguished from V. caudolineatus by its slightly greater average and maximum size, proportionally shorter fore­ and hind­limbs, more elongate snout, higher average midbody and ventral scale counts, higher average sub­digital lamellar counts on pes, more finely scaled and less rugose proximal portion of the tail, more numerous presacral vertebrae, less conspicuously spotted head, more orderly alignment of dorsal pattern into transverse rows, and presence of transverse bands on the basal one­third of the tail. Males are further distinguished from V. caudolineatus by having a shorter hemipenis with more numerous papillose distal frills.</p><p>Etymology: We take pleasure in naming this species after naturalist and educator Brian Bush who has contributed enormously to our knowledge of the herpetofauna of Western Australia and of the Pilbara region in particular.</p><p>Distribution and sympatry: Endemic to the Pilbara region of Western Australia, northwest to the vicinity of Cooya Poonya and Tambrey, southwest to Mt Brockman and Mt Tom Price in the Hamersley Range, southeast to Mt Whaleback and northeast to Marillana and Hope Downs (Fig. 9). Two instances of regional sympatry with V. caudolineatus are recorded—at West Angelas and at Hope Downs, both on the southern margin of the Pilbara Uplands. Details of the relevant specimens are provided in the Discussion.</p><p>Description of holotype: Adult male measuring SVL 145 mm, Tail 190 mm, Forelimb 32 mm, Hind limb 43 mm, Head and neck 50 mm. Both hemipenes are fully everted.</p><p>Vertebral column includes 30 presacral vertebrae. Phalangeal formula of manus 1.2.3.4.2; and of pes 3.4.3.2.1.</p><p>Head moderately depressed, depth at pineal organ 8.7 mm. Snout relatively elongate; lacking canthus rostralis. Nostril positioned 6.8 mm from tip of snout; 3.4 mm from anterior corner of eye. Ear aperture is vertically narrow and obliquely oriented.</p><p>Dorsal head scales unornamented; most are irregular polygons, sub­rounded in shape, and majority have a single scale organ. Supraocular scales are smaller than those on the rostrum, frontal and parietal regions. Granules are absent from head except in temporal region and on throat forward into genal groove. Dorsal primary scales on neck are raised but lack keels; all have scale organs. The dorsal primary scales on neck are rounded anteriorly but become more ovate posteriorly, merging smoothly with body scalation. Ventral primary scales on neck are small and almost round anteriorly, but become larger and progressively more elongate to rear; all ventral primary scales on neck are ringed laterally and posteriorly by small granules. Approximately one third of ventral neck scales bear scale organs, usually one per scale but not infrequently two or three. Gular fold is distinct, located 5 mm forward of anterior base of forelimb, and consisting of 4 rows of small scales, all lacking scale organs.</p><p>Body scalation is relatively unspecialised. Dorsal primary scales are ovate, approximately twice as long as wide and lack keels; the majority have a single scale organ. Transverse scale rows are well­organised on the dorsum but tend to subdivide on flanks. Ventral primary scales are approximately 50% longer than and twice as wide as dorsal primary scales. All are ringed laterally and posteriorly by small granules. Single scale organs are present on the majority of scales towards the flanks but only about 15% of scales located along the mid­ventral area. Inguinal fold indistinct but located 1.5 mm forward of the anterior base of hind­limb. Position is marked by a change in scale size and shape, those behind the fold and back to vent being smaller, rounded and not arranged in transverse rows, and by occurrence of incomplete transverse series that terminate before reaching flank.</p><p>Scales on the inner surface of the fore­ and hind­limbs are small and rounded while those on the outer surfaces are larger, more elongate and weakly keeled. Plantar surfaces of manus and pes with primary scales on small mounds of fine granules; all primary scales bear scale organs and some are pigmented. Subdigital surfaces with transverse lamellae made up of two or three rounded primary scales surrounded by granules. The apical lamella on each digit is single and more intensely pigmented. Longest digit (IV) of manus has 20 lamellae; longest digit (IV) of pes has 23. Claws on manus and pes are moderately large and darkly pigmented; all are laterally compressed and bear sharp, recurved tips.</p><p>Tail is slightly wider than deep at base, becoming more rounded in cross­section towards tip; it lacks a dorsal keel or other obvious specialisation. Dorsal primary scales near tail base are elongate but otherwise unspecialised. Moving distally, the primary scales first become more elongate, then develop a distinct midline keel, and finally develop smaller lateral keels. Ventral primary scales also change from elongate and lacking keels near the tail base, to even longer and with a strong midline keel distally. The overall effect is that the tail feels relatively smooth near the base but becomes more distinctly rasp­like distally. Circumferential scale counts decrease from 52 near tail base to 32 at one third of total length, then to 22 at two­thirds of total length. Longitudinal scale counts at the same positions are 8, 7 and 6 rows per cm.</p><p>The post­cloacal scale cluster is relatively poorly developed and consists of two rows of modified but unpigmented scales. The outer row consists of six nodular scales of which the lateral four scales are distinctly spinose. The inner row consists of four smaller, bluntly pointed scales.</p><p>The hemipenis measures 6.3 mm without the projecting hemibacula and 6.8 including these structures. The nude basal portion measures 3.3 mm. Further details were provided in an earlier section.</p><p>Ground colour of dorsum in preservative is a uniform pale­brown from rostrum to base of tail, becoming greyer on the flanks. Patterning on the head includes an irregular mottling of dark brown on the dorsum and sides, tending towards longitudinal streaking on the occiput, and a subdued temporal stripe on each side, running from the posterior corner of the eye to above the ear. Irregular dark brown mottling is also present on the dorsum of the body, each ‘spot’ usually consisting of one or a few dark primary scales. The pattern is irregular on the anterior body but tends towards transverse alignment on the lower half of the back and above the hind limbs. The last three bars of this series are quite distinct and alternate with rows of small dark spots. The under surface of the throat, neck and body bears distinct pale grey spotting, each ‘spot’ usually comprised of two or three pigmented primary scales. These are most abundant on the throat and neck, and along the sides of the body, and less so in the mid­ventral region. The insides of the limbs are similarly patterned. The proximal 35 mm of the tail bears a series of eight more or less complete transverse bars. The next 38 mm bears a series of broken bars and irregular spots. The remainder of the tail supports a linear pattern consisting of five, more or less continuous dark stripes, each of which is one scale wide. The ventral surface of the tail is unpatterned throughout its length.</p><p>Variation among referred specimens: Mensural and meristic data for the complete sample of V. b u s h i sp. nov. is presented in Table 1; locality details for referred specimens are given in Appendix II. The holotype is the largest male specimen of V. b u s h i sp. nov.; SVL of the largest female (WAM R135340) is 140 mm.</p><p>Dorsal patterning in most individuals is similar to that of the holotype, with fine spotting on the anterior body, conspicuous banding only on the rear of the body and on the proximal segment of the tail, and longitudinal striping of the distal tail. In some individuals (e.g. WAM R125521 from the ‘northern Pilbara’) the entire head and body are more intensely marked with variable sized spots; these are randomly distributed on the head and neck but are aligned into more or less regular transverse rows posterior to the level of attachment of the forelimbs. All specimens of V. b u s h i sp. nov. have ventral spotting on the throat, neck and body. In comparison with the holotype, most individuals show more intense and evenly distributed spotting on the body. Some individuals also show a strong differentiation between abundant fine spotting on the throat and neck, with each spot consisting of a single pigmented primary scale, and less abundant but larger blotches on the body.</p><p>Dorsal ground colour in life is pale grey­brown, typically with a reddish tinge on the upper surface, from the crown of the head to between the hind limbs (Fig. 1). The ground colour of the tail is always paler than the body and generally has a cream wash on the distal one third of the upper surface.</p><p>Morphological comparisons: Varanus bushi sp. nov. is most similar overall to V. gilleni, a phenetic resemblance that is consistent with the phylogenetic conclusions of the molecular analyses. These species share a number of morphological attributes including a relatively elongate body with proportionally shorter limbs, high mid­body and ventral scale counts, high subdigital lamellar counts on the pes, a relatively non­spinose basal portion of the tail, and a tendency to include transverse bands in the dorsal patterning. As indicated in the diagnosis, V. bushi sp. nov. differs from V. gilleni in its lower average and maximum body size, its more elongate dorsal scales, its more densely spotted venter, its less conspicuous linear patterning on the head and neck, its more numerous presacral vertebrae, its simpler heimbacular morphology, and some minor meristic differences.</p><p>Var a nu s bushi sp. nov. is more similar to V. caudolineatus in body patterning and this probably explains the former confusion between these species. Both species share a spotted rather than banded pattern on the dorsum, and both are typically spotted on the throat and to some degree on the venter. However, Varanus bushi sp. nov. is usually less heavily pigmented than typical V. caudolineatus, with smaller and less intense spots. The lower back and basal portion of the tail are distinctly banded in V. b u s h i sp. nov. but more irregularly patterned in V. caudolineatus . Body proportions also distinguish the two species, V. b u s h i sp. nov. having a more elongate body form with more numerous presacral vertebrae, higher ventral scale counts, more elongate dorsal primary scales, and relatively shorter fore­ and hind­limbs. Although tail length is quite variable in both taxa, the majority of V. caudolineatus have proportionally shorter tails than V. b u s h i sp. nov. The head is also shorter and stockier in V. caudolineatus, produced mainly by a shortening of the rostrum. Midbody scale counts are typically higher in V. bushi sp. nov. and the hemipenes differ in several important details from those of V. caudolineatus .</p><p>The evolutionary polarity of most of the morphological characteristics that distinguish each of the three members of the V. caudolineatus species group is uncertain. Sprackland (1991) suggests that more heavily rugose tails and smaller body size are probably apomorphic within Odatria . On both counts, V. caudolineatus would rate as more derived than either of the other taxa. Varanus caudolineatus appears to be more primitive in hemipeneal morphology than either V. bushi sp. nov. or V. g i l l e n i; of the latter two species, V. gilleni possesses a more derived hemibacular morphology.</p><p>Taxonomic remarks: Although the type specimens of Varanus caudolineatus Boulenger, 1885 and Varanus gilleni Lucas &amp; Frost, 1895 were not examined as part of this study, the type localities of each of these taxa [Champion Bay, W.A. (= Geraldton) and Charlotte Waters, N.T.; see Fig. 9] is remote from the known geographic range of V. bushi sp. nov. and well within the ranges of these other taxa as currently understood. We are therefore confident that the various names are correctly associated with the biological entities as defined herein.</p><p>Ecological notes: Relatively little information is available on the ecology of Pilbara varanids and the new taxon is no exception. Most recent specimens of V. bushi sp. nov. have been taken from fallen or standing hollow trees in mulga or eucalypt woodland associations.</p><p>The sex ratio among the total sample of V. b u s h i sp. nov. is 34 males to 13 females. This situation is typical for opportunistically collected samples of varanids (King and Rhodes 1982; Greer 1989) and probably reflects different activity patterns between the sexes. The lack of immature specimens of V. b u s h i sp. nov. and the small numbers of very young individuals in the large samples of V. caudolineatus and V. g i l l e n i is consistent with previous comment that hatchling varanids are especially cryptic and poorly represented in wild caught samples (Horn and Visser 1991).</p><p>Identifiable stomach contents were observed in two of four V. b u s h i sp. nov. examined by D.J. King. WAM R73142 (male, SVL 96 mm) contained remains of a spider and a skink tail; WAM R54230 (male, SVL 135 mm) contained remains of a mole cricket ( Gryllotalpidae).</p></div>	https://treatment.plazi.org/id/03A587F0FF83FF94FEF2FE15A7FCFE64	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Aplin, Kenneth P.;Fitch, Alison J.;King, Dennis J.	Aplin, Kenneth P., Fitch, Alison J., King, Dennis J. (2006): A new species of Varanus Merrem (Squamata: Varanidae) from the Pilbara region of Western Australia, with observations on sexual dimorphism in closely related species. Zootaxa 1313: 1-38, DOI: 10.5281/zenodo.173907
03A587F0FFB6FFADFEF2FE37A4D6FCA9.text	03A587F0FFB6FFADFEF2FE37A4D6FCA9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Varanus caudolineatus	<div><p>Varanus caudolineatus</p><p>Males: R330 – 331, R10812 Quinns 27°04’S 118°38’E; R1191 Warriedar, via Yalgoo 29°08’S 117°11’E; R3415 – 3416 Laverton 28°38’S 122°24’E; R3818 Narloo, via Wurarga 28°19’S 116°11’E; R4732 Gullewa, via Yalgoo 28°39’S 116°19’E; R4933 Murchison Downs 26°48’S 118°39’E; R5048 Marilla 22°58’S 114°28’E; R7378 Belele, via Meekatharra 26°22’S 117°38’E; R8168 Warroora 23°29’S 113°48’E; R10611 Minilya 23°51’S 113°58’E; R13362, 26069 Jiggalong Mission 23°22’S 120°47’E; R14918 11 km S of Cue 27°32’S 117°54’E; R15785 Ejah Paddock, Mileura 26°22’S 117°20’E; R15789 Charlies Creek, Mileura 26°22’S 117°20’E; R19771 Kathleen Valley 27°24’S 120°39’E; R21100 32 km NE of Yelma HS 26°20’S 121°56’E; R21137 24 km SW of Wiluna 26°45’S 120°03’E; R21139 70 km SW of Wiluna 27°03’S 119°43’E; R21150 51 km SW of Sandstone 28°18’S 118°56’E; R22652 Fields Find 29°02’S 117°15’E; R22995 Ajana 27°57’S 114°38’E; R25149 2 Miles SE of Turee Creek 23°38’S 118°40’E; R26320 ‘Greenough’ (probably Eastern Goldfields) 30°45’S 121°30’E; R28028 The Well Spring 25°01’S 121°35’E; R28290 Albion Downs 27°17’S 120°23’E; R28394, 28955 Coordewandy 25°36’S 115°58’E; R30967 – 30968 Albion Downs (within 13 km of HS) 27°17’S 120°23’E; R31381 – 31385 35 km NE of Mingenew 28°58’S 115°42’E; R34563, 39765 Callagiddy, 32 km SE Of Carnarvon 25°03’S 114°02’E; R34685 64 km N of Beacon. 29°53’S 117°52’E; R41793 Kalli, near Cue 26°54’S 117°07’E; R47347 Byro 26°05’S 116°09’E; R48151 5 km NW of Kirkalocka HS 28°32’S 117°43’E; R48152 16 km S of Mt Magnet 28°14’S 117°51’E; R49943 Woodleigh 26°11’S 114°33’E; R49992 Wilroy Reserve, 19 km S of Mullewa 28°44’S 115°30’E; R51092 60 km S of Leonora 29°25’S 121°20’E; R51189 1 km N of Orabanda 30°22’S 121°04’E; R52893, 52894, 52896 Mt Augustus 24°18’S 116°55’E; R54594 15 km E of Hamelin Pool 26°22’S 114°20’E; R54595 Wooramel HS 25°44’S 114°17’E; R57378 Woodleigh 26°11’S 114°33’E; R59607 20 km ENE of Meadow Station HS 26°39’S 114°48’E; R59620 34 km SSE of Nerren Nerren 27°19’S 114°51’E; R60130 Youanmi 28°37’S 118°50’E; R60429 2 km S of Overlander 26°26’S 114°28’E; R60642 13 km W of Mungawolagudi Claypan 26°48’S 115°19’E; R62865 22 km SE of Mt Keith 27°19’S 120°40’E; R64436 Cooloomia HS 26°57’S 114°18’E; R65820 12.0 km ENE of Comet Vale 29°54’25”S 121°14’05”E; R65883 8.3 km SSE of Mt Linden 29°23’20”S 122°28’05”E; R65886 8 km SSE of Mt Linden 29°23’00”S 122°27’30”E; R65961 2.5 km N of Mt Linden 29°18’20”S 122°25’00”E; R69118 18.5 km ENE of Yuinmery HS 28°31’55”S 119°11’35”E; R69315 12.5 km SSE of Banjawarn HS 27°47’55”S 121°40’55”E; R70831 24 km at 333°N from Mt Windarra 28°18’S 122°07’E; R70895 3.5 km at 5°N from Yowie Rockhole 30°26’30”S 122°21’00”E; R71062 Billabong Roadhouse 26°49’S 114°37’E; R72653 – 72654 Comet Vale 29°55’S 121°35’E; R72830 7.75 km SE of Mt Linden 29°22’55”S 122°27’05”E; R72892 7.75 km SSE of Mt Linden 29°22’55”S 122°28’05”E; R73223 2 km at 15°N from Yowie Rockhole 30°27’30”S 122°21’00”E; R73309 5.5 km at 137°N from Black Flag 30°35’30”S 121°16’30”E; R73433 9 km at 190°N from Mt Elvire HS 29°26’30”S 119°35’00”E; R73624 Ophthalmia Range 23°17’S 119°07’E; R74671 18.5 km ENE of Yuinmery 28°31’55”S 119°11’35”E; R78488 7 km NE of Yowie Rockhole 30°25’S 122°22’E; R78545 21 km SE of Mt Keith 27°19’S 120°40’E; R81917 Jibberding 29°51’30”S 116°58’00”E; R81922 White Well, 6 km W of Jibberding 29°53’S 116°55’E; R84443 14 km S of Dromedary Hill 29°11’S 118°24’E; R85239, 85241 Dead Horse Rocks 29°22’S 121°17’E; R87661 13 km SSW of Mt. Phillip HS 24°31’S 116°14’E; R87770 12 km SW of Yinnietharra 24°45’S 116°06’E; R92900 14 km SW of Hamelin HS 26°30’49”S 114°05’44”E; R94715 Agnew 28°01’S 120°31’E; R95287 7 km N of Goolthan Goolthan Hill 28°03’S 116°44’E; R97786 Yoothapinna 26°32’S 118°30’E; R 122622 6 km W of Meedo HS 25°42’38”S 114°35’58”E; R123783 14.5 km W of Pells Creek 25°07’09”S 115°25’36”E; R129980 Mt Joel 27°17’55”S 121°03’15”E; R129984 Mandilla Well 27°31’05”S 121°07’21”E; R131829 Lake Carey 29°10’S 122°21’E; R132181 Bandya Hill area 27°50’S 122°15’E; R132507 Jundee 26°35’59”S 120°47’57”E; R135601 Lake Carey 28°50’49”S 122°18’39”E; R138950 West Angelas 23°11’46”S 118°31’10”E; R 140705 Hope Downs 22°40’25”S 119°24’58”E.</p><p>Females: R3423 Laverton 28°38’S 122°24’E; R3903 Well 29.Canning Stock Route 22°33’S 123°53’E; R4934 Murchison Downs 26°48’S 118°59’E; R7379 Belele via Meekatharra 26°22’S 117°38’E; R8167 Warroora 23°29’S 113°48’E; R12407 Kathleen Valley near Lenora 27°24’S 120°39’E; R13711 Overlander Roadhouse 26°24’S 114°28’E; R13857 Cosmo Newbery 28°00’S 122°54’E; R14240 Kalgoorlie 30°44’S 121°28’E; R14917 27 km E of Marillana 22°38’S 119°40’E; R15786 – 15788 Ejah Paddock, Mileura 26°22’S 117°20’E; R17681 Mt Margaret 28°48’S 122°11’E; R19600 Cosmo Newbery Mission 28°00’S 122°54’E; R21138 70 km SW of Wiluna 27°03’S 119°43’E; R21149 35 km N of Sandstone 27°40’S 119°18’E; R21178, 21179, 21181 11 km SW of Youanmi 28°41’S 118°44’E; R22870 Yalgoo 28°21’S 116°41’E; R25883 Ajana 27°57’S 114°38’E; R26068, 26070 Jiggalong, vicinity of Mission 23°22’S 120°47’E; R27231 Kathleen Valley, Wanjarri 27°19’S 120°33’E; R28956 Coordewandy 25°36’S 115°58’E; R29111 32 km S of Mt Magnet 28°21’S 117°51’E; R30969 Albion Downs, within 13 km of HS 27°17’S 120°23’E; R31681 14 km S of Menzies 29°49’S 121°02’E; R34564 Callagiddy, 32 km SE of Carnarvon 25°03’S 114°02’E; R39043 Youanmi 28°37’S 118°50’E; R44529 Woodleigh 26°11’S 114°33’E; R47374 Sandstone 27°59’S 119°18’E; R47793 30 km SE of Bulloo Downs 24°13’S 119°47’E; R49993 Wilroy Reserve, 19 km S of Mullewa 28°44’S 115°30’E; R52895 Mt Augustus 24°19’S 116°55’E; R57377 Woodleigh 26°11’S 114°33’E; R62866 – 62867 21 km SE of Mt Keith 27°18’S 120°40’E; R63655 25 km NW of Winning HS 22°56’S 114°27’E; R69293 9.5 km SSE of Banjawarn HS 27°46’55”S 121°39’45”E; R69295 12.5 km SSE of Banjawarn HS 27°47’55”S 121°40’55”E; R72749 12.25 km ENE of Comet Vale 29°55’25”S 121°14’35”E; R72893 7.75 km SSE of Mt Linden 29°22’55”S 122°28’05”E; R74689 8 km ENE of Yuinmery 28°31’15”S 119°05’30”E; R75858 Coomalbidgup, Lort River 33°43’S 121°22’E; R78546 21 km SE of Mt Keith 27°19’S 120°40’E; R78582 Mt Windarra area 28°29’S 122°14’E; R84028 37 km SE of Ashburton Downs HS 23°38’S 117°18’E; R85240 Dead Horse Rocks 29°22’S 121°17’E; R87637 12 km SW of Yinnetharra HS 24°45’S 116°06’E; R87754 4 km NNE of Mt. Phillip HS 24°23’S 116°19’E; R88098 11 km N of Nerren Nerren HS 27°02’S 114°38’E; R91684 15 km N of Karalunde 26°00’S 118°41’E; R95527 3 km S of Nannowtharra Hill 28°18’S 117°00’E; R 101298 2 km N of Kathleen Valley shearing shed, Wanjarri Nature Reserve 27°20’S 120°39’E; R116670 Black Range, 15 km WSW of Marangaroon 23°47’18”S 115°28’02”E; R117117 Upper Gascoyne 24°43’S 116°04’E; R 122854 5.6 km E of Mardathuna HS 24°26’35”S 114°30’42”E; R125540 Randall Well, 210 km NNE of Meekatharra” 25°21’S 119°24’E; R125889 31.2 km from Binthalya HS 24°31’23”S 114°57’56”E; R138077 10 km SE of Mt Augustus HS 24°29’S 116°54’E 140714 Hope Downs 22°40’25”S 119°24’58”E.</p><p>Unknown sex: R3881 Well 5, Canning Stock Route 25°22’S 121°00’E; R7279 Caron 29°35’S 116°19’E; R8257 Grants Patch 30°27’S 121°07’E; R12278 Mundiwindi 23°48’S 120°15’E; R19789 – 19790 Albion Downs 27°17’S 120°23’E; R21180 11 km SW of Youanmi 28°41’S 118°44’E; R23908 – 23910 Laverton 28°38’S 122°24’E; R37897 Callagiddy 25°03’S 114°02’E; R46618 Yarri Battery, NE of Kalgoorlie 29°47’S 122°22’E; R46621 Linden 29°18’S 122°25’E; R47630 Macquarie Mill, Mileura 26°22’S 117°20’E; R47794 23 km NW of Mt Bruce 22°30’S 117°58’E; R51157 5 km S of Warriedar HS 29°12’S 117°11’E; R66004 7.75 km SSE of Mt Linden 29°22’55”S 122°28’05” E 78594 22 km S of Mt Elvire HS 29°33’S 119°36’E; R96122 – 96125 Millrose area 26°10’S 120°43’E; R96679 17 km WNW of Wadina HS 27°56’S 115°28’E; R 100314 Mt Lawrence Wells 26°48’S 120°12’E; R 102016 5.4 km N of Joy Helen MS 23°14’15”S 115°46’23”E; R 102098 Wongida Well, Barlee Range” 22°58’S 115°51’E; R115204 Eurardy 27°34’S 114°40’E; R 122576 1.9 km W of Meedo HS 25°40’50”S 114°37’18”E; R125186 31.2 km from Binthalya HS 24°31’23”S 114°57’56”E; R127246 130 km NW of Mount Magnet 27°42’S 117°05’E.</p><p>Varanus gilleni</p><p>Males: R3970 Well 37, Canning Stock Route 22°09’S 125°27’E; R8715 Well 43, Canning Stock Route 21°12’S 125°59’E; R14653, 14654, 14656 Warburton Range 26°08’S 126°35’E; R14657 Newbore, 45 km NW of Warburton Mission 25°40’S 126°15’E; R15707 32 km SE of Warburton Range 26°20’S 126°48’E; R20608, 22007, 22021, 22022, 22211, 22212 Warburton Mission 26°08’S 126°35’E; R21000 – 21001 10 km SE of Warburton Range Mission 26°12’S 126°39’E; R28027 Injudinah Creek, La Grange 18°38’S 121°52’E; R28814 14 km NW of Mt Beadell 25°47’S 124°38’E; R28864 77 km SW of Mt Beadell. Sutherland Range 26°01’S 124°44’E; R40111 60 Miles E of No.24 Well, Canning Stock Route 23°22’S 124°08’E; R45768 – 45769 Between Emily Gap and Amoonguna, near Alice Springs 23°14’S 133°59’E; R46166 Anna Plains 19°15’S 121°29’E; R54073 Edgar Ranges Reserve 18°55’S 123°15’E; R57047 30 km NNE of Stretch Range 20°44’S 127°51’E; R57304 McLarty Hills 19°29’S 123°28’E; R63421 Twin Heads 20°15’S 126°32’E; R63960 Well No 30, Canning Stock Route 22°30’S 124°08’E; R75808 Nita Downs 19°05’S 121°41’E; R81443 Wildlife Well 22°53’S 125°10’E; R82606 approx. 45 km NW of Mt Crofton 21°25’S 121°45’E; R88546 55 km S of Anna Plains HS 19°44’S 121°28’E; R 100593 15 km SSW of Giles 25°05’S 128°20’E; R 101434 Plumridge Lakes Nature Reserve 29°36’S 124°54’E; R 102728 Little Sandy Desert 24°35’33”S 120°15’47”E; R108565 7–8 km WSW of Point Salvation 28°15’S 123°36’E.</p><p>Females: R3995 Well 49, Canning Stock Route 20°10’S 126°41’E; R14655 Warburton Range 26°08’S 126°35’E; R15179 Elder Creek, 8 km NNW of Warburton 26°03’S 126°33’E; R15706, 15708 32 km SE of Warburton Range 26°20’S 126°48’E; R24437 16 km S of Ewaninga HS, Northern Territory 24°08’S 133°56’E; R26714 No locality but probably SW Kimberley 17°30’S 123°30’E; R46165, 46167 Anna Plains 19°15’S 121°29’E; R47674 Point Massie, Canning Stock Route 20°43’S 126°30’E; R53787 – 53789 near Alice Springs 23°42’S 133°52’E; R57302 – 57303 McLarty Hills 19°29’S 123°28’E; R60135 Injudinah Creek, La Grange 18°38’S 121°52’E; R63959 Well No. 30, Canning Stock Route 22°30’S 124°08’E; R64202 2 km at 90°N from Murguga Well No39, Canning Stock Route 21°45’S 125°40’E; R67579, 94371 Balgo Mission 20°09’S 128°57’E; R75791 24 km W of Joanna Spring 20°05’S 123°55’E; R75792 Dragon Tree Soak 19°39’S 123°23’E; R82607 approx. 45 km NW of Mt Crofton 21°25’S 121°45’E; R 100859 10 km N of Charlies Knob 23°05’S 125°00’E.</p><p>Unknown sex: R3998 between Wells 49 and 50, Canning Stock Route 20°12’S 126°50’E; R4020 between Wells 39 and 51, Canning Stock Route 20°50’S 126°10’E; R19598 – 19599 Warburton Range 26°08’S 126°35’E; R40887 Mt Romilly, Canning Stock Route 20°28’S 126°30’E; R45262 Well 40, Canning Stock Route 21°40’S 125°47’E; R46123 32 km SW of Christmas Creek HS 19°05’S 125°43’E; R54125 Edgar Ranges 18°49’S 123°17’E; R67578 Balgo Mission 20°09’S 128°57’E; R 102674 Little Sandy Desert 24°04’46”S 120°20’15”E; R 112140 Warburton 26°08’S 126°35’E; R136033 Little Sandy Desert 24°55’50”S 120°31’18”E; R136034 Little Sandy Desert 24°35’33”S 120°15’47”E.</p><p>Varanus bushi sp. nov.</p><p>Males: R4288 Tambrey 21°38’S 117°36’E; R54230 Marandoo 22°37’S 118°08’E; R73142 26.6 km at 225°N from Marillana HS 22°48’17”S 119°13’30”E; R76453 10 km SSW of Cooya Pooya HS 21°07’S 117°07’E; R108999 Marandoo 22°37’S 118°08’E; R125105 Yandicoogina 22°43’08”S 119°03’07”E; R125520 – 125521 northern Pilbara region 21°30’S 117°30’E; R129632 120 km NW of Newman 22°57’S 119°21’E; R129912, 138949 West Angelas MS, 100 km NE of Newman 23°11’09”S 118°45’16”E; R131494 Mt Whaleback 23°22’31”S 119°38’25”E; R135467 Mt Brockman area 22°28’S 117°18’E; R145260 5 km S of Mt Tom Price MS 22°49’02”S 117°45’50”E.</p><p>Females: R20241 near Kangiangi HS 21°41’S 117°17’E; R56834 Marandoo 22°37’S 118°08’E; R73143 26.6 km at 225°N from Marillana HS 22°48’17”S 119°13’30”E; R127789 5 km S of Mount Tom Price MS 22°47’49”S 117°47’20”E; R 131037 Unknown locality; R135340 Hope Downs 22°40’S 119°24’E.</p><p>Unknown sex: R62171 Marandoo 22°37’S 118°08’E; R75834 Weano Gorge 22°22’S 118°15’E; R131751 Four Corners Bore, Hamersley 22°26’43”S 117°52’47”E; R138169 Hamersley Range National Park 22°35’S 118°12’E.</p></div>	https://treatment.plazi.org/id/03A587F0FFB6FFADFEF2FE37A4D6FCA9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Aplin, Kenneth P.;Fitch, Alison J.;King, Dennis J.	Aplin, Kenneth P., Fitch, Alison J., King, Dennis J. (2006): A new species of Varanus Merrem (Squamata: Varanidae) from the Pilbara region of Western Australia, with observations on sexual dimorphism in closely related species. Zootaxa 1313: 1-38, DOI: 10.5281/zenodo.173907
