taxonID	type	description	language	source
03A587F0FF83FF94FEF2FE15A7FCFE64.taxon	materials_examined	Holotype: Western Australian Museum R 108999, adult male from Marandoo, Western Australia in 22 0 37 ’ S 118 0 08 ’ E. Collected on 20 June 1991 by Greg Harold. Specimen fixed in 10 % formalin then stored in 70 % ethanol. Liver sample stored in ­ 80 0 C ultrafreeze at WAM. Paratypes: Three additional specimens from Marandoo: WAM R 54230, an adult male collected on 25 October 1976; WAM R 56834, an adult female collected in April 1977; and WAM R 62171 a juvenile of undetermined sex collected on 28 February 1979.	en	Aplin, Kenneth P., Fitch, Alison J., King, Dennis J. (2006): A new species of Varanus Merrem (Squamata: Varanidae) from the Pilbara region of Western Australia, with observations on sexual dimorphism in closely related species. Zootaxa 1313: 1-38, DOI: 10.5281/zenodo.173907
03A587F0FF83FF94FEF2FE15A7FCFE64.taxon	diagnosis	Diagnosis: A small­bodied member of the subgenus Odatria distinguished from most others by the combination of a longitudinally striped and only moderately spinose tail, unkeeled head and body scales, non­overlapping ventral primary scales, and an absence of longitudinal streaks on throat. Distinguished from V. g i l l e n i by its slightly lesser average size, more elongate dorsal scales, more densely spotted venter and more irregularly spotted dorsum, less prominent linear patterning on the head and neck, and its more numerous presacral vertebrae, pedal subdigital lamellae and ventral scales. Males are further distinguished from V. gilleni by having hemipenes with an undivided inner hemibaculum. Distinguished from V. caudolineatus by its slightly greater average and maximum size, proportionally shorter fore­ and hind­limbs, more elongate snout, higher average midbody and ventral scale counts, higher average sub­digital lamellar counts on pes, more finely scaled and less rugose proximal portion of the tail, more numerous presacral vertebrae, less conspicuously spotted head, more orderly alignment of dorsal pattern into transverse rows, and presence of transverse bands on the basal one­third of the tail. Males are further distinguished from V. caudolineatus by having a shorter hemipenis with more numerous papillose distal frills.	en	Aplin, Kenneth P., Fitch, Alison J., King, Dennis J. (2006): A new species of Varanus Merrem (Squamata: Varanidae) from the Pilbara region of Western Australia, with observations on sexual dimorphism in closely related species. Zootaxa 1313: 1-38, DOI: 10.5281/zenodo.173907
03A587F0FF83FF94FEF2FE15A7FCFE64.taxon	etymology	Etymology: We take pleasure in naming this species after naturalist and educator Brian Bush who has contributed enormously to our knowledge of the herpetofauna of Western Australia and of the Pilbara region in particular.	en	Aplin, Kenneth P., Fitch, Alison J., King, Dennis J. (2006): A new species of Varanus Merrem (Squamata: Varanidae) from the Pilbara region of Western Australia, with observations on sexual dimorphism in closely related species. Zootaxa 1313: 1-38, DOI: 10.5281/zenodo.173907
03A587F0FF83FF94FEF2FE15A7FCFE64.taxon	distribution	Distribution and sympatry: Endemic to the Pilbara region of Western Australia, northwest to the vicinity of Cooya Poonya and Tambrey, southwest to Mt Brockman and Mt Tom Price in the Hamersley Range, southeast to Mt Whaleback and northeast to Marillana and Hope Downs (Fig. 9). Two instances of regional sympatry with V. caudolineatus are recorded — at West Angelas and at Hope Downs, both on the southern margin of the Pilbara Uplands. Details of the relevant specimens are provided in the Discussion.	en	Aplin, Kenneth P., Fitch, Alison J., King, Dennis J. (2006): A new species of Varanus Merrem (Squamata: Varanidae) from the Pilbara region of Western Australia, with observations on sexual dimorphism in closely related species. Zootaxa 1313: 1-38, DOI: 10.5281/zenodo.173907
03A587F0FF83FF94FEF2FE15A7FCFE64.taxon	description	Description of holotype: Adult male measuring SVL 145 mm, Tail 190 mm, Forelimb 32 mm, Hind limb 43 mm, Head and neck 50 mm. Both hemipenes are fully everted. Vertebral column includes 30 presacral vertebrae. Phalangeal formula of manus 1.2.3.4.2; and of pes 3.4.3.2.1. Head moderately depressed, depth at pineal organ 8.7 mm. Snout relatively elongate; lacking canthus rostralis. Nostril positioned 6.8 mm from tip of snout; 3.4 mm from anterior corner of eye. Ear aperture is vertically narrow and obliquely oriented. Dorsal head scales unornamented; most are irregular polygons, sub­rounded in shape, and majority have a single scale organ. Supraocular scales are smaller than those on the rostrum, frontal and parietal regions. Granules are absent from head except in temporal region and on throat forward into genal groove. Dorsal primary scales on neck are raised but lack keels; all have scale organs. The dorsal primary scales on neck are rounded anteriorly but become more ovate posteriorly, merging smoothly with body scalation. Ventral primary scales on neck are small and almost round anteriorly, but become larger and progressively more elongate to rear; all ventral primary scales on neck are ringed laterally and posteriorly by small granules. Approximately one third of ventral neck scales bear scale organs, usually one per scale but not infrequently two or three. Gular fold is distinct, located 5 mm forward of anterior base of forelimb, and consisting of 4 rows of small scales, all lacking scale organs. Body scalation is relatively unspecialised. Dorsal primary scales are ovate, approximately twice as long as wide and lack keels; the majority have a single scale organ. Transverse scale rows are well­organised on the dorsum but tend to subdivide on flanks. Ventral primary scales are approximately 50 % longer than and twice as wide as dorsal primary scales. All are ringed laterally and posteriorly by small granules. Single scale organs are present on the majority of scales towards the flanks but only about 15 % of scales located along the mid­ventral area. Inguinal fold indistinct but located 1.5 mm forward of the anterior base of hind­limb. Position is marked by a change in scale size and shape, those behind the fold and back to vent being smaller, rounded and not arranged in transverse rows, and by occurrence of incomplete transverse series that terminate before reaching flank. Scales on the inner surface of the fore­ and hind­limbs are small and rounded while those on the outer surfaces are larger, more elongate and weakly keeled. Plantar surfaces of manus and pes with primary scales on small mounds of fine granules; all primary scales bear scale organs and some are pigmented. Subdigital surfaces with transverse lamellae made up of two or three rounded primary scales surrounded by granules. The apical lamella on each digit is single and more intensely pigmented. Longest digit (IV) of manus has 20 lamellae; longest digit (IV) of pes has 23. Claws on manus and pes are moderately large and darkly pigmented; all are laterally compressed and bear sharp, recurved tips. Tail is slightly wider than deep at base, becoming more rounded in cross­section towards tip; it lacks a dorsal keel or other obvious specialisation. Dorsal primary scales near tail base are elongate but otherwise unspecialised. Moving distally, the primary scales first become more elongate, then develop a distinct midline keel, and finally develop smaller lateral keels. Ventral primary scales also change from elongate and lacking keels near the tail base, to even longer and with a strong midline keel distally. The overall effect is that the tail feels relatively smooth near the base but becomes more distinctly rasp­like distally. Circumferential scale counts decrease from 52 near tail base to 32 at one third of total length, then to 22 at two­thirds of total length. Longitudinal scale counts at the same positions are 8, 7 and 6 rows per cm. The post­cloacal scale cluster is relatively poorly developed and consists of two rows of modified but unpigmented scales. The outer row consists of six nodular scales of which the lateral four scales are distinctly spinose. The inner row consists of four smaller, bluntly pointed scales. The hemipenis measures 6.3 mm without the projecting hemibacula and 6.8 including these structures. The nude basal portion measures 3.3 mm. Further details were provided in an earlier section. Ground colour of dorsum in preservative is a uniform pale­brown from rostrum to base of tail, becoming greyer on the flanks. Patterning on the head includes an irregular mottling of dark brown on the dorsum and sides, tending towards longitudinal streaking on the occiput, and a subdued temporal stripe on each side, running from the posterior corner of the eye to above the ear. Irregular dark brown mottling is also present on the dorsum of the body, each ‘ spot’ usually consisting of one or a few dark primary scales. The pattern is irregular on the anterior body but tends towards transverse alignment on the lower half of the back and above the hind limbs. The last three bars of this series are quite distinct and alternate with rows of small dark spots. The under surface of the throat, neck and body bears distinct pale grey spotting, each ‘ spot’ usually comprised of two or three pigmented primary scales. These are most abundant on the throat and neck, and along the sides of the body, and less so in the mid­ventral region. The insides of the limbs are similarly patterned. The proximal 35 mm of the tail bears a series of eight more or less complete transverse bars. The next 38 mm bears a series of broken bars and irregular spots. The remainder of the tail supports a linear pattern consisting of five, more or less continuous dark stripes, each of which is one scale wide. The ventral surface of the tail is unpatterned throughout its length.	en	Aplin, Kenneth P., Fitch, Alison J., King, Dennis J. (2006): A new species of Varanus Merrem (Squamata: Varanidae) from the Pilbara region of Western Australia, with observations on sexual dimorphism in closely related species. Zootaxa 1313: 1-38, DOI: 10.5281/zenodo.173907
03A587F0FF83FF94FEF2FE15A7FCFE64.taxon	materials_examined	Variation among referred specimens: Mensural and meristic data for the complete sample of V. b u s h i sp. nov. is presented in Table 1; locality details for referred specimens are given in Appendix II. The holotype is the largest male specimen of V. b u s h i sp. nov.; SVL of the largest female (WAM R 135340) is 140 mm. Dorsal patterning in most individuals is similar to that of the holotype, with fine spotting on the anterior body, conspicuous banding only on the rear of the body and on the proximal segment of the tail, and longitudinal striping of the distal tail. In some individuals (e. g. WAM R 125521 from the ‘ northern Pilbara’) the entire head and body are more intensely marked with variable sized spots; these are randomly distributed on the head and neck but are aligned into more or less regular transverse rows posterior to the level of attachment of the forelimbs. All specimens of V. b u s h i sp. nov. have ventral spotting on the throat, neck and body. In comparison with the holotype, most individuals show more intense and evenly distributed spotting on the body. Some individuals also show a strong differentiation between abundant fine spotting on the throat and neck, with each spot consisting of a single pigmented primary scale, and less abundant but larger blotches on the body. Dorsal ground colour in life is pale grey­brown, typically with a reddish tinge on the upper surface, from the crown of the head to between the hind limbs (Fig. 1). The ground colour of the tail is always paler than the body and generally has a cream wash on the distal one third of the upper surface. Morphological comparisons: Varanus bushi sp. nov. is most similar overall to V. gilleni, a phenetic resemblance that is consistent with the phylogenetic conclusions of the molecular analyses. These species share a number of morphological attributes including a relatively elongate body with proportionally shorter limbs, high mid­body and ventral scale counts, high subdigital lamellar counts on the pes, a relatively non­spinose basal portion of the tail, and a tendency to include transverse bands in the dorsal patterning. As indicated in the diagnosis, V. bushi sp. nov. differs from V. gilleni in its lower average and maximum body size, its more elongate dorsal scales, its more densely spotted venter, its less conspicuous linear patterning on the head and neck, its more numerous presacral vertebrae, its simpler heimbacular morphology, and some minor meristic differences.	en	Aplin, Kenneth P., Fitch, Alison J., King, Dennis J. (2006): A new species of Varanus Merrem (Squamata: Varanidae) from the Pilbara region of Western Australia, with observations on sexual dimorphism in closely related species. Zootaxa 1313: 1-38, DOI: 10.5281/zenodo.173907
03A587F0FF83FF94FEF2FE15A7FCFE64.taxon	description	Var a nu s bushi sp. nov. is more similar to V. caudolineatus in body patterning and this probably explains the former confusion between these species. Both species share a spotted rather than banded pattern on the dorsum, and both are typically spotted on the throat and to some degree on the venter. However, Varanus bushi sp. nov. is usually less heavily pigmented than typical V. caudolineatus, with smaller and less intense spots. The lower back and basal portion of the tail are distinctly banded in V. b u s h i sp. nov. but more irregularly patterned in V. caudolineatus. Body proportions also distinguish the two species, V. b u s h i sp. nov. having a more elongate body form with more numerous presacral vertebrae, higher ventral scale counts, more elongate dorsal primary scales, and relatively shorter fore­ and hind­limbs. Although tail length is quite variable in both taxa, the majority of V. caudolineatus have proportionally shorter tails than V. b u s h i sp. nov. The head is also shorter and stockier in V. caudolineatus, produced mainly by a shortening of the rostrum. Midbody scale counts are typically higher in V. bushi sp. nov. and the hemipenes differ in several important details from those of V. caudolineatus. The evolutionary polarity of most of the morphological characteristics that distinguish each of the three members of the V. caudolineatus species group is uncertain. Sprackland (1991) suggests that more heavily rugose tails and smaller body size are probably apomorphic within Odatria. On both counts, V. caudolineatus would rate as more derived than either of the other taxa. Varanus caudolineatus appears to be more primitive in hemipeneal morphology than either V. bushi sp. nov. or V. g i l l e n i; of the latter two species, V. gilleni possesses a more derived hemibacular morphology. Taxonomic remarks: Although the type specimens of Varanus caudolineatus Boulenger, 1885 and Varanus gilleni Lucas & Frost, 1895 were not examined as part of this study, the type localities of each of these taxa [Champion Bay, W. A. (= Geraldton) and Charlotte Waters, N. T.; see Fig. 9] is remote from the known geographic range of V. bushi sp. nov. and well within the ranges of these other taxa as currently understood. We are therefore confident that the various names are correctly associated with the biological entities as defined herein. Ecological notes: Relatively little information is available on the ecology of Pilbara varanids and the new taxon is no exception. Most recent specimens of V. bushi sp. nov. have been taken from fallen or standing hollow trees in mulga or eucalypt woodland associations. The sex ratio among the total sample of V. b u s h i sp. nov. is 34 males to 13 females. This situation is typical for opportunistically collected samples of varanids (King and Rhodes 1982; Greer 1989) and probably reflects different activity patterns between the sexes. The lack of immature specimens of V. b u s h i sp. nov. and the small numbers of very young individuals in the large samples of V. caudolineatus and V. g i l l e n i is consistent with previous comment that hatchling varanids are especially cryptic and poorly represented in wild caught samples (Horn and Visser 1991). Identifiable stomach contents were observed in two of four V. b u s h i sp. nov. examined by D. J. King. WAM R 73142 (male, SVL 96 mm) contained remains of a spider and a skink tail; WAM R 54230 (male, SVL 135 mm) contained remains of a mole cricket (Gryllotalpidae).	en	Aplin, Kenneth P., Fitch, Alison J., King, Dennis J. (2006): A new species of Varanus Merrem (Squamata: Varanidae) from the Pilbara region of Western Australia, with observations on sexual dimorphism in closely related species. Zootaxa 1313: 1-38, DOI: 10.5281/zenodo.173907
03A587F0FFB6FFADFEF2FE37A4D6FCA9.taxon	description	Females: R 3423 Laverton 28 ° 38 ’ S 122 ° 24 ’ E; R 3903 Well 29. Canning Stock Route 22 ° 33 ’ S 123 ° 53 ’ E; R 4934 Murchison Downs 26 ° 48 ’ S 118 ° 59 ’ E; R 7379 Belele via Meekatharra 26 ° 22 ’ S 117 ° 38 ’ E; R 8167 Warroora 23 ° 29 ’ S 113 ° 48 ’ E; R 12407 Kathleen Valley near Lenora 27 ° 24 ’ S 120 ° 39 ’ E; R 13711 Overlander Roadhouse 26 ° 24 ’ S 114 ° 28 ’ E; R 13857 Cosmo Newbery 28 ° 00 ’ S 122 ° 54 ’ E; R 14240 Kalgoorlie 30 ° 44 ’ S 121 ° 28 ’ E; R 14917 27 km E of Marillana 22 ° 38 ’ S 119 ° 40 ’ E; R 15786 – 15788 Ejah Paddock, Mileura 26 ° 22 ’ S 117 ° 20 ’ E; R 17681 Mt Margaret 28 ° 48 ’ S 122 ° 11 ’ E; R 19600 Cosmo Newbery Mission 28 ° 00 ’ S 122 ° 54 ’ E; R 21138 70 km SW of Wiluna 27 ° 03 ’ S 119 ° 43 ’ E; R 21149 35 km N of Sandstone 27 ° 40 ’ S 119 ° 18 ’ E; R 21178, 21179, 21181 11 km SW of Youanmi 28 ° 41 ’ S 118 ° 44 ’ E; R 22870 Yalgoo 28 ° 21 ’ S 116 ° 41 ’ E; R 25883 Ajana 27 ° 57 ’ S 114 ° 38 ’ E; R 26068, 26070 Jiggalong, vicinity of Mission 23 ° 22 ’ S 120 ° 47 ’ E; R 27231 Kathleen Valley, Wanjarri 27 ° 19 ’ S 120 ° 33 ’ E; R 28956 Coordewandy 25 ° 36 ’ S 115 ° 58 ’ E; R 29111 32 km S of Mt Magnet 28 ° 21 ’ S 117 ° 51 ’ E; R 30969 Albion Downs, within 13 km of HS 27 ° 17 ’ S 120 ° 23 ’ E; R 31681 14 km S of Menzies 29 ° 49 ’ S 121 ° 02 ’ E; R 34564 Callagiddy, 32 km SE of Carnarvon 25 ° 03 ’ S 114 ° 02 ’ E; R 39043 Youanmi 28 ° 37 ’ S 118 ° 50 ’ E; R 44529 Woodleigh 26 ° 11 ’ S 114 ° 33 ’ E; R 47374 Sandstone 27 ° 59 ’ S 119 ° 18 ’ E; R 47793 30 km SE of Bulloo Downs 24 ° 13 ’ S 119 ° 47 ’ E; R 49993 Wilroy Reserve, 19 km S of Mullewa 28 ° 44 ’ S 115 ° 30 ’ E; R 52895 Mt Augustus 24 ° 19 ’ S 116 ° 55 ’ E; R 57377 Woodleigh 26 ° 11 ’ S 114 ° 33 ’ E; R 62866 – 62867 21 km SE of Mt Keith 27 ° 18 ’ S 120 ° 40 ’ E; R 63655 25 km NW of Winning HS 22 ° 56 ’ S 114 ° 27 ’ E; R 69293 9.5 km SSE of Banjawarn HS 27 ° 46 ’ 55 ” S 121 ° 39 ’ 45 ” E; R 69295 12.5 km SSE of Banjawarn HS 27 ° 47 ’ 55 ” S 121 ° 40 ’ 55 ” E; R 72749 12.25 km ENE of Comet Vale 29 ° 55 ’ 25 ” S 121 ° 14 ’ 35 ” E; R 72893 7.75 km SSE of Mt Linden 29 ° 22 ’ 55 ” S 122 ° 28 ’ 05 ” E; R 74689 8 km ENE of Yuinmery 28 ° 31 ’ 15 ” S 119 ° 05 ’ 30 ” E; R 75858 Coomalbidgup, Lort River 33 ° 43 ’ S 121 ° 22 ’ E; R 78546 21 km SE of Mt Keith 27 ° 19 ’ S 120 ° 40 ’ E; R 78582 Mt Windarra area 28 ° 29 ’ S 122 ° 14 ’ E; R 84028 37 km SE of Ashburton Downs HS 23 ° 38 ’ S 117 ° 18 ’ E; R 85240 Dead Horse Rocks 29 ° 22 ’ S 121 ° 17 ’ E; R 87637 12 km SW of Yinnetharra HS 24 ° 45 ’ S 116 ° 06 ’ E; R 87754 4 km NNE of Mt. Phillip HS 24 ° 23 ’ S 116 ° 19 ’ E; R 88098 11 km N of Nerren Nerren HS 27 ° 02 ’ S 114 ° 38 ’ E; R 91684 15 km N of Karalunde 26 ° 00 ’ S 118 ° 41 ’ E; R 95527 3 km S of Nannowtharra Hill 28 ° 18 ’ S 117 ° 00 ’ E; R 101298 2 km N of Kathleen Valley shearing shed, Wanjarri Nature Reserve 27 ° 20 ’ S 120 ° 39 ’ E; R 116670 Black Range, 15 km WSW of Marangaroon 23 ° 47 ’ 18 ” S 115 ° 28 ’ 02 ” E; R 117117 Upper Gascoyne 24 ° 43 ’ S 116 ° 04 ’ E; R 122854 5.6 km E of Mardathuna HS 24 ° 26 ’ 35 ” S 114 ° 30 ’ 42 ” E; R 125540 Randall Well, 210 km NNE of Meekatharra ” 25 ° 21 ’ S 119 ° 24 ’ E; R 125889 31.2 km from Binthalya HS 24 ° 31 ’ 23 ” S 114 ° 57 ’ 56 ” E; R 138077 10 km SE of Mt Augustus HS 24 ° 29 ’ S 116 ° 54 ’ E 140714 Hope Downs 22 ° 40 ’ 25 ” S 119 ° 24 ’ 58 ” E. Unknown sex: R 3881 Well 5, Canning Stock Route 25 ° 22 ’ S 121 ° 00 ’ E; R 7279 Caron 29 ° 35 ’ S 116 ° 19 ’ E; R 8257 Grants Patch 30 ° 27 ’ S 121 ° 07 ’ E; R 12278 Mundiwindi 23 ° 48 ’ S 120 ° 15 ’ E; R 19789 – 19790 Albion Downs 27 ° 17 ’ S 120 ° 23 ’ E; R 21180 11 km SW of Youanmi 28 ° 41 ’ S 118 ° 44 ’ E; R 23908 – 23910 Laverton 28 ° 38 ’ S 122 ° 24 ’ E; R 37897 Callagiddy 25 ° 03 ’ S 114 ° 02 ’ E; R 46618 Yarri Battery, NE of Kalgoorlie 29 ° 47 ’ S 122 ° 22 ’ E; R 46621 Linden 29 ° 18 ’ S 122 ° 25 ’ E; R 47630 Macquarie Mill, Mileura 26 ° 22 ’ S 117 ° 20 ’ E; R 47794 23 km NW of Mt Bruce 22 ° 30 ’ S 117 ° 58 ’ E; R 51157 5 km S of Warriedar HS 29 ° 12 ’ S 117 ° 11 ’ E; R 66004 7.75 km SSE of Mt Linden 29 ° 22 ’ 55 ” S 122 ° 28 ’ 05 ” E 78594 22 km S of Mt Elvire HS 29 ° 33 ’ S 119 ° 36 ’ E; R 96122 – 96125 Millrose area 26 ° 10 ’ S 120 ° 43 ’ E; R 96679 17 km WNW of Wadina HS 27 ° 56 ’ S 115 ° 28 ’ E; R 100314 Mt Lawrence Wells 26 ° 48 ’ S 120 ° 12 ’ E; R 102016 5.4 km N of Joy Helen MS 23 ° 14 ’ 15 ” S 115 ° 46 ’ 23 ” E; R 102098 Wongida Well, Barlee Range ” 22 ° 58 ’ S 115 ° 51 ’ E; R 115204 Eurardy 27 ° 34 ’ S 114 ° 40 ’ E; R 122576 1.9 km W of Meedo HS 25 ° 40 ’ 50 ” S 114 ° 37 ’ 18 ” E; R 125186 31.2 km from Binthalya HS 24 ° 31 ’ 23 ” S 114 ° 57 ’ 56 ” E; R 127246 130 km NW of Mount Magnet 27 ° 42 ’ S 117 ° 05 ’ E. Varanus gilleni Males: R 3970 Well 37, Canning Stock Route 22 ° 09 ’ S 125 ° 27 ’ E; R 8715 Well 43, Canning Stock Route 21 ° 12 ’ S 125 ° 59 ’ E; R 14653, 14654, 14656 Warburton Range 26 ° 08 ’ S 126 ° 35 ’ E; R 14657 Newbore, 45 km NW of Warburton Mission 25 ° 40 ’ S 126 ° 15 ’ E; R 15707 32 km SE of Warburton Range 26 ° 20 ’ S 126 ° 48 ’ E; R 20608, 22007, 22021, 22022, 22211, 22212 Warburton Mission 26 ° 08 ’ S 126 ° 35 ’ E; R 21000 – 21001 10 km SE of Warburton Range Mission 26 ° 12 ’ S 126 ° 39 ’ E; R 28027 Injudinah Creek, La Grange 18 ° 38 ’ S 121 ° 52 ’ E; R 28814 14 km NW of Mt Beadell 25 ° 47 ’ S 124 ° 38 ’ E; R 28864 77 km SW of Mt Beadell. Sutherland Range 26 ° 01 ’ S 124 ° 44 ’ E; R 40111 60 Miles E of No. 24 Well, Canning Stock Route 23 ° 22 ’ S 124 ° 08 ’ E; R 45768 – 45769 Between Emily Gap and Amoonguna, near Alice Springs 23 ° 14 ’ S 133 ° 59 ’ E; R 46166 Anna Plains 19 ° 15 ’ S 121 ° 29 ’ E; R 54073 Edgar Ranges Reserve 18 ° 55 ’ S 123 ° 15 ’ E; R 57047 30 km NNE of Stretch Range 20 ° 44 ’ S 127 ° 51 ’ E; R 57304 McLarty Hills 19 ° 29 ’ S 123 ° 28 ’ E; R 63421 Twin Heads 20 ° 15 ’ S 126 ° 32 ’ E; R 63960 Well No 30, Canning Stock Route 22 ° 30 ’ S 124 ° 08 ’ E; R 75808 Nita Downs 19 ° 05 ’ S 121 ° 41 ’ E; R 81443 Wildlife Well 22 ° 53 ’ S 125 ° 10 ’ E; R 82606 approx. 45 km NW of Mt Crofton 21 ° 25 ’ S 121 ° 45 ’ E; R 88546 55 km S of Anna Plains HS 19 ° 44 ’ S 121 ° 28 ’ E; R 100593 15 km SSW of Giles 25 ° 05 ’ S 128 ° 20 ’ E; R 101434 Plumridge Lakes Nature Reserve 29 ° 36 ’ S 124 ° 54 ’ E; R 102728 Little Sandy Desert 24 ° 35 ’ 33 ” S 120 ° 15 ’ 47 ” E; R 108565 7 – 8 km WSW of Point Salvation 28 ° 15 ’ S 123 ° 36 ’ E. Females: R 3995 Well 49, Canning Stock Route 20 ° 10 ’ S 126 ° 41 ’ E; R 14655 Warburton Range 26 ° 08 ’ S 126 ° 35 ’ E; R 15179 Elder Creek, 8 km NNW of Warburton 26 ° 03 ’ S 126 ° 33 ’ E; R 15706, 15708 32 km SE of Warburton Range 26 ° 20 ’ S 126 ° 48 ’ E; R 24437 16 km S of Ewaninga HS, Northern Territory 24 ° 08 ’ S 133 ° 56 ’ E; R 26714 No locality but probably SW Kimberley 17 ° 30 ’ S 123 ° 30 ’ E; R 46165, 46167 Anna Plains 19 ° 15 ’ S 121 ° 29 ’ E; R 47674 Point Massie, Canning Stock Route 20 ° 43 ’ S 126 ° 30 ’ E; R 53787 – 53789 near Alice Springs 23 ° 42 ’ S 133 ° 52 ’ E; R 57302 – 57303 McLarty Hills 19 ° 29 ’ S 123 ° 28 ’ E; R 60135 Injudinah Creek, La Grange 18 ° 38 ’ S 121 ° 52 ’ E; R 63959 Well No. 30, Canning Stock Route 22 ° 30 ’ S 124 ° 08 ’ E; R 64202 2 km at 90 ° N from Murguga Well No 39, Canning Stock Route 21 ° 45 ’ S 125 ° 40 ’ E; R 67579, 94371 Balgo Mission 20 ° 09 ’ S 128 ° 57 ’ E; R 75791 24 km W of Joanna Spring 20 ° 05 ’ S 123 ° 55 ’ E; R 75792 Dragon Tree Soak 19 ° 39 ’ S 123 ° 23 ’ E; R 82607 approx. 45 km NW of Mt Crofton 21 ° 25 ’ S 121 ° 45 ’ E; R 100859 10 km N of Charlies Knob 23 ° 05 ’ S 125 ° 00 ’ E. Unknown sex: R 3998 between Wells 49 and 50, Canning Stock Route 20 ° 12 ’ S 126 ° 50 ’ E; R 4020 between Wells 39 and 51, Canning Stock Route 20 ° 50 ’ S 126 ° 10 ’ E; R 19598 – 19599 Warburton Range 26 ° 08 ’ S 126 ° 35 ’ E; R 40887 Mt Romilly, Canning Stock Route 20 ° 28 ’ S 126 ° 30 ’ E; R 45262 Well 40, Canning Stock Route 21 ° 40 ’ S 125 ° 47 ’ E; R 46123 32 km SW of Christmas Creek HS 19 ° 05 ’ S 125 ° 43 ’ E; R 54125 Edgar Ranges 18 ° 49 ’ S 123 ° 17 ’ E; R 67578 Balgo Mission 20 ° 09 ’ S 128 ° 57 ’ E; R 102674 Little Sandy Desert 24 ° 04 ’ 46 ” S 120 ° 20 ’ 15 ” E; R 112140 Warburton 26 ° 08 ’ S 126 ° 35 ’ E; R 136033 Little Sandy Desert 24 ° 55 ’ 50 ” S 120 ° 31 ’ 18 ” E; R 136034 Little Sandy Desert 24 ° 35 ’ 33 ” S 120 ° 15 ’ 47 ” E. Varanus bushi sp. nov. Males: R 4288 Tambrey 21 ° 38 ’ S 117 ° 36 ’ E; R 54230 Marandoo 22 ° 37 ’ S 118 ° 08 ’ E; R 73142 26.6 km at 225 ° N from Marillana HS 22 ° 48 ’ 17 ” S 119 ° 13 ’ 30 ” E; R 76453 10 km SSW of Cooya Pooya HS 21 ° 07 ’ S 117 ° 07 ’ E; R 108999 Marandoo 22 ° 37 ’ S 118 ° 08 ’ E; R 125105 Yandicoogina 22 ° 43 ’ 08 ” S 119 ° 03 ’ 07 ” E; R 125520 – 125521 northern Pilbara region 21 ° 30 ’ S 117 ° 30 ’ E; R 129632 120 km NW of Newman 22 ° 57 ’ S 119 ° 21 ’ E; R 129912, 138949 West Angelas MS, 100 km NE of Newman 23 ° 11 ’ 09 ” S 118 ° 45 ’ 16 ” E; R 131494 Mt Whaleback 23 ° 22 ’ 31 ” S 119 ° 38 ’ 25 ” E; R 135467 Mt Brockman area 22 ° 28 ’ S 117 ° 18 ’ E; R 145260 5 km S of Mt Tom Price MS 22 ° 49 ’ 02 ” S 117 ° 45 ’ 50 ” E. Females: R 20241 near Kangiangi HS 21 ° 41 ’ S 117 ° 17 ’ E; R 56834 Marandoo 22 ° 37 ’ S 118 ° 08 ’ E; R 73143 26.6 km at 225 ° N from Marillana HS 22 ° 48 ’ 17 ” S 119 ° 13 ’ 30 ” E; R 127789 5 km S of Mount Tom Price MS 22 ° 47 ’ 49 ” S 117 ° 47 ’ 20 ” E; R 131037 Unknown locality; R 135340 Hope Downs 22 ° 40 ’ S 119 ° 24 ’ E. Unknown sex: R 62171 Marandoo 22 ° 37 ’ S 118 ° 08 ’ E; R 75834 Weano Gorge 22 ° 22 ’ S 118 ° 15 ’ E; R 131751 Four Corners Bore, Hamersley 22 ° 26 ’ 43 ” S 117 ° 52 ’ 47 ” E; R 138169 Hamersley Range National Park 22 ° 35 ’ S 118 ° 12 ’ E.	en	Aplin, Kenneth P., Fitch, Alison J., King, Dennis J. (2006): A new species of Varanus Merrem (Squamata: Varanidae) from the Pilbara region of Western Australia, with observations on sexual dimorphism in closely related species. Zootaxa 1313: 1-38, DOI: 10.5281/zenodo.173907
