identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03A5792B772F707AFF1FF89DFA10FC9D.text	03A5792B772F707AFF1FF89DFA10FC9D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Asplenium pahli Salgado, Wenzel 2021	<div><p>Asplenium pahli Salgado, sp. nov. (Figure 2).</p><p>Type:— PHILIPPINES, Leyte, 500 m, 5 February 1914. C. A. Wenzel 572 (holotype P00370339! designated here, isotypes P00370338!, P00370340!, C0364253 F (F!), US 01516038!).</p><p>Diagnosis: — Asplenium pahli and A. anisodontum have a proliferous rachis but differ in the pinna shape and serration. A. pahli differs from A. caudatum, A. falcatum, A. gueinzianum, A. planicaule and A. polyodon in the proliferous rachis, the pinna margins being irregularly serrate, sinuses of different depth and width, fewer sori distant from the costa, not contiguous, and straight or slightly falcate.</p><p>Terrestrial. Rhizome short, erect, 0.4–0.5 cm thick, paleae and palea bases few, scattered, clathrate, black, lanceate, margin entire, &lt;1mm long, upper part of the paleae often broken off, cells small, polygonal. Roots many, crowded at the base of fronds, brown, thin, fragile when dried. Fronds erect, pinnate, crowded at the apex of the rhizome, spirally arranged, 25–45 cm long. Stipes 10.5(13.0–17.0)21.0 cm long, adaxial side grooved, abaxial side rounded, greenish, glabrous or glabrescent, with sparse paleae, paleae bicolorous, base and margin light brown, central area black, lanceate, ≤ 1 mm long, adpressed against the surface, margin entire, irregular with an occasional acute projection formed by a clathrate cell wall, palea apex acute. Lamina oblong, 15–35 cm long, (5.5) 6.5–8 cm wide, base truncate, pinnate, chartaceous, 12–15 alternating pinna pairs, pinnae slightly reduced at near apex. Rachis proliferous, abaxial side rounded, adaxial side with a wide U-shaped groove with prominent ridges, greenish, glabrescent, with adpressed small paleae, ≤ 1 mm long, lanceate, black, margin entire, apex acute, paleae on both sides of the rachis. Pinnae 3.2–4.0 cm long, 1.0– 1.3 cm wide in the middle, noncontiguous, alternating, generally trapezoid, stalked to sessile, stalk ≤ 1 mm long, naked or with few paleae, lamina adaxial side glabrous, abaxial side glabrous or with a few reduced paleae on the costa, costa prominent of both sides, adaxial side grooved with prominent ridges gradually disappearing toward the pinna apex, abaxial side rounded, basal pinna pair sub-opposite or alternating, generally oblong, of the same length and shape as median pinnae or slightly smaller; pinna base basiscopic side cuneate, acroscopic side cuneate to broadly cuneate, pinna apex acute, pinna margins irregularly serrate, sinuses of different depth and width, acroscopic margin with 5–6 oblique widely separated teeth, larger teeth with 2–3 apical teeth, teeth blunt or acute, basiscopic margin serration similar to acroscopic margin, 3–5 oblique widely separated teeth, lamina apical segment usually missing, when present deformed with a proliferous bud at the insertion with the rachis. Venation free, costa visible with prominent ridges, 5–7 pairs of veins oblique to the costa, abaxial side veins not visible, adaxial side veins hardly visible, once forking, ending near the margin; hydathodes absent. Sori in a single row distal from pinna base, on both sides of the costa, originating about 1 mm from costa, oblique, widely separated, 3–6 sori on the acroscopic side, 2–5 on the basiscopic side, 5–7 mm long, occupying middle portion of lamina, ending before reaching the margin. Indusium thick, brown, entire, rolls back to expose mature sporangia, margin entire or slightly jagged.</p><p>Distribution and ecology:—Island of Leyte, Central-Eastern Philippines. A terrestrial species growing on the eastern slopes of the Leyte Cordillera, at 500 m elevation, in lowland tropical evergreen rainforest. Climate warm and humid; rainfall evenly distributed throughout the year (Pagasa 2020).</p><p>Etymology:—The species epithet honors Brother George Pahl, FSC, Ph.D., an American biologist and university professor who taught, mentored, and inspired thousands of students in the United States and the Philippines during his teaching career spanning more than 50 years.</p><p>Conservation status:— A. pahli was collected in an area near the towns of Dagami and Jaro that is presently denuded of its original primary growth forest (Lasco et al. 2001; Mongabay 2020). The only collection known is that of Wenzel in 1914. This is a possible indication that A. pahli is endangered or possibly extinct in its type locality. There might be more plants scattered in the remaining stands of old growth deep in the Leyte Cordillera.</p><p>Asplenium caudatum, A. falcatum, A. anisodontum and A. pahli represent different species as shown by the characters presented in this key.</p><p>1. Fronds non proliferous .......................................................................................................................................................................2</p><p>1. Fronds proliferous ..............................................................................................................................................................................3</p><p>2. Basal pinnae not decreasing in size, lamina base truncate; acroscopic basal pinna lobe with a prominent sharp tooth, sori single row, oblique to costa.......................................................................................................................................................... A. falcatum</p><p>2. Basal pinnae gradually decreasing in size, lamina base attenuate; acroscopic basal pinna lobe without a prominent sharp tooth, sori in two rows, one parallel to costa and one oblique to costa ............................................................................................ A. caudatum</p><p>3. Fronds up to 120 cm long. Stipe and rachis dark brown; rachis hirsute, paleae projecting away from the stipe and rachis, oblong to lanceate or reduced to long twisted biserrate filaments, margin with prominent biserrate projections up to five cells long, palea base and margin concolorous with the rest of the palea, apex a long twisted biseriate filament; median pinnae falcate, 10.0– 15.6 cm long, 0.9–1.2 cm wide; sori close and parallel to costa then curving towards the margin, 7–11mm long, contiguous ................. ..................................................................................................................................................................................... A. anisodontum</p><p>3. Fronds up to 45 cm long. Stipe and rachis greenish; rachis with a few scattered paleae, paleae adpressed against the stipe and rachis surface, deltoid to lanceate in shape, margin entire and irregular, with an occasional short, acute projection caused by a cell wall, palea base and margin bicolorous with light brown margins and black central area, apex acute; median pinna straight, 3.5–4.0 cm long and 1.0– 1.3 cm wide; sori distant from the costa, straight or slightly falcate, oblique, 5–7 mm long, not contiguous............... ................................................................................................................................................................................................. A. pahli</p></div>	https://treatment.plazi.org/id/03A5792B772F707AFF1FF89DFA10FC9D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Salgado, Arthur Edward	Salgado, Arthur Edward (2021): Taxonomic and nomenclatural notes on Philippine ferns. IV. Asplenium pahli (Polypodiales, Aspleniineae, Aspleniaceae), a new species from Leyte, Philippines. Phytotaxa 524 (2): 71-84, DOI: 10.11646/phytotaxa.524.2.1
03A5792B77277074FF1FFADDFDE7FE9E.text	03A5792B77277074FF1FFADDFDE7FE9E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Asplenium anisodontum C. Presl, Cuming 1849	<div><p>Asplenium anisodontum C.Presl</p><p>Holotype (designated by C. Presl 1849):— PHILIPPINES. Luzon: Cuming 128 (PRC non vide); isotypes L0050982!, L0050983!, BM001045207!, BM001045208!, BM001045209!, BM001045210, B 20 0011335!, B 20 0011341!, E 00210809!, MICH1003398 B!, MW0000516!, US00065552!, W0052502!.</p></div>	https://treatment.plazi.org/id/03A5792B77277074FF1FFADDFDE7FE9E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Salgado, Arthur Edward	Salgado, Arthur Edward (2021): Taxonomic and nomenclatural notes on Philippine ferns. IV. Asplenium pahli (Polypodiales, Aspleniineae, Aspleniaceae), a new species from Leyte, Philippines. Phytotaxa 524 (2): 71-84, DOI: 10.11646/phytotaxa.524.2.1
03A5792B77277074FF1FFB81FA31FD5A.text	03A5792B77277074FF1FFB81FA31FD5A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Asplenium caudatum G. Forster 1786	<div><p>Asplenium caudatum G.Forst.</p><p>ISLANDS OF THE PACIFIC OCEAN. Isotypes, islands of the Pacific Ocean, G. Forster s.n. (BM001048371!); G. Forster s. n. (LIV!); islands of Pacific Sea, G. Forster s.n. (GOET012797!).</p><p>FRENCH POLYNESIA, Society islands, Tahiti: vers 1200 m, Nadeaud s. n. (P!); Herbier E. Drake, Savatier s. n. (two sheets, P!); Herbier E. Drake, Nadeaud s. n. (P!); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-149.84485&amp;materialsCitation.latitude=-17.54735" title="Search Plazi for locations around (long -149.84485/lat -17.54735)">Vallée de la rivière Paho, Toahotu</a>, crête sud, 149°16’W, 17°47’S, 650 m, Florence &amp; Jourdan 11075 (US01517559!) ; Island of Moorea: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-149.84485&amp;materialsCitation.latitude=-17.54735" title="Search Plazi for locations around (long -149.84485/lat -17.54735)">Mt. Muaroa</a>, elev. 429 m, 17.54735ºS, 149.84485ºW, 12 August 2010, J. Nitta 570 (P00847390!) . Marquesas Islands, Ua Pou Island, Poumaka, mid-elevation wet forest, 680 m, 9° 23′ 26″, 140° 4′ 58″, 30 October 2017, Schuettpelz, E. Wood, K.R. &amp; Butaud, J.-F. 1857 (US01936878!); Hivaoa, F.W. Christian s.n. (P!); Fatu Hiva, K.R. Wood 4491, 2 sheets (P!) .</p></div>	https://treatment.plazi.org/id/03A5792B77277074FF1FFB81FA31FD5A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Salgado, Arthur Edward	Salgado, Arthur Edward (2021): Taxonomic and nomenclatural notes on Philippine ferns. IV. Asplenium pahli (Polypodiales, Aspleniineae, Aspleniaceae), a new species from Leyte, Philippines. Phytotaxa 524 (2): 71-84, DOI: 10.11646/phytotaxa.524.2.1
03A5792B77277074FF1FF8C5FB1AFC3A.text	03A5792B77277074FF1FF8C5FB1AFC3A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Asplenium falcatum Lamarck 1786	<div><p>Asplenium falcatum Lam.</p><p>= Trichomanes adiantoides</p><p>Lectotype (designated by Salgado &amp; Fraser-Jenkins 2013):— SRI LANKA: 1737, J. Burman Thesaurus Zeylanicus t. 43. Epitype (designated by Salgado &amp; Fraser-Jenkins 2013):— SRI LANKA. North Central Province: Ritigala, 14 January 1951, W.A. Sledge 873 (US 01514788!).</p><p>SRI LANKA, 1717. P. Herrmann s.n., Historical Collections, Herb. Herrmann 3: 47, no. 385, (BM 000621974!). North Western Province: Kurunegala District, R. B. Faden &amp; A. J. Faden 76/400 (US 01514785!). INDIA. Courtallam: East India Company; s. coll. (K 001109576!, K 00119577!); Assam: Darrauz District, Balipara forest; M. Gustav s.n. (US01514780!).</p></div>	https://treatment.plazi.org/id/03A5792B77277074FF1FF8C5FB1AFC3A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Salgado, Arthur Edward	Salgado, Arthur Edward (2021): Taxonomic and nomenclatural notes on Philippine ferns. IV. Asplenium pahli (Polypodiales, Aspleniineae, Aspleniaceae), a new species from Leyte, Philippines. Phytotaxa 524 (2): 71-84, DOI: 10.11646/phytotaxa.524.2.1
03A5792B77277074FF1FFE25FB80FBAE.text	03A5792B77277074FF1FFE25FB80FBAE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Asplenium gueinzianum Mett. ex Kuhn 1868	<div><p>Asplenium gueinzianum Mett. ex Kuhn</p><p>Holotype (verified by R. Vianey):— SOUTH AFRICA: Port Natal: Gueinzius s.n. (B20 0015752!).</p></div>	https://treatment.plazi.org/id/03A5792B77277074FF1FFE25FB80FBAE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Salgado, Arthur Edward	Salgado, Arthur Edward (2021): Taxonomic and nomenclatural notes on Philippine ferns. IV. Asplenium pahli (Polypodiales, Aspleniineae, Aspleniaceae), a new species from Leyte, Philippines. Phytotaxa 524 (2): 71-84, DOI: 10.11646/phytotaxa.524.2.1
03A5792B77277074FF1FFE51FDCBFB3E.text	03A5792B77277074FF1FFE51FDCBFB3E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Asplenium pahli Salgado, Wenzel 2021	<div><p>Asplenium pahli Salgado</p><p>Holotype (designated here by A. E. Salgado):— PHILIPPINES. Leyte, C. A. Wenzel 572, P00370339!; isotypes P00370338!, P00370340!, US01516038!, UC 267862!, C0364253 F! (F).</p></div>	https://treatment.plazi.org/id/03A5792B77277074FF1FFE51FDCBFB3E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Salgado, Arthur Edward	Salgado, Arthur Edward (2021): Taxonomic and nomenclatural notes on Philippine ferns. IV. Asplenium pahli (Polypodiales, Aspleniineae, Aspleniaceae), a new species from Leyte, Philippines. Phytotaxa 524 (2): 71-84, DOI: 10.11646/phytotaxa.524.2.1
03A5792B77277074FF1FFF21FB98F94A.text	03A5792B77277074FF1FFF21FB98F94A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Asplenium planicaule Mett.	<div><p>Asplenium planicaule Wall. ex Mett.</p><p>= Asplenium yoshinagae (Makino) subsp. indicum (Sledge) Fraser-Jenk.</p><p>Lectotype (designated by Morton 1973):— NEPAL. 1820, Numerical List (Catalogue) No. 189-1, Wallich s.n., isotypes BM001045235!, K001109500!, K001044577!, NY 214847!, UC 267862!. INDIA. Srinugur [Srinagar], 1820, Numerical List (Catalogue) No. 189- 2, W. Moorcroft s.n., isotypes K001109501!, NY 214848!; Nilgiris, Flora of Madras, 6000 ft [1828 m], May 1886, Gamble 17334 (US01515892!).</p><p>CHINA. Yunnan, Yangbi Xian, W side of Diancang Shan mountain range. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.03333&amp;materialsCitation.latitude=25.716667" title="Search Plazi for locations around (long 100.03333/lat 25.716667)">En</a> route from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.03333&amp;materialsCitation.latitude=25.716667" title="Search Plazi for locations around (long 100.03333/lat 25.716667)">Xueshanhe</a> to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.03333&amp;materialsCitation.latitude=25.716667" title="Search Plazi for locations around (long 100.03333/lat 25.716667)">Dapingzi</a>, growing on crevices of a ledge, 25º 43’N, 100º 02’E, 2300-3000 m, 17 June 1984, Sino-American Botanical Expedition to Yunnan Province 242 (US3043222!) ; Yunnan Red River forest across from Maubao Mongtze, W. Hancock 187 (K001092464!). COUNTRY UNKNOWN. Himalayas, J. F. Royle s.n . (PH 00025140!);</p><p>Note: Morton (1973) designated Wallich No. 189 (B) as the lectotype of A. planicaule . The present author was not able to locate this specimen. There are doubtful isotypes at BM, K, LE , PH and S: BM001045200!, BM001045235!, BM000523633!, BM000523634!, BM000523635!, K001044578!, LE00008417 !, PH 00004925!, S-P-1376! (S), UC 267862 !.</p></div>	https://treatment.plazi.org/id/03A5792B77277074FF1FFF21FB98F94A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Salgado, Arthur Edward	Salgado, Arthur Edward (2021): Taxonomic and nomenclatural notes on Philippine ferns. IV. Asplenium pahli (Polypodiales, Aspleniineae, Aspleniaceae), a new species from Leyte, Philippines. Phytotaxa 524 (2): 71-84, DOI: 10.11646/phytotaxa.524.2.1
03A5792B77277074FF1FFCF5FEBBF82A.text	03A5792B77277074FF1FFCF5FEBBF82A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Asplenium polyodon G. Forst.	<div><p>Asplenium polyodon G.Forst.</p><p>Lectotype (Lectotype designated by Nicolson &amp; Fosberg 2003: 123) G. Forster s.n. (UPS-THUNB 24832!): — AUSTRALIA. Queensland: South-east region, Queen Mary Falls, 8 miles SE of Killarney, 13 December 1956. R. Schodde 316 (BM!) ; North of Moreton Bay, from Glasshouse Mountain to Mount Flinders and Stradborke Island, 1850-1851, no coll. (BM!) ; West Springbrook, 2,200 ft [670 m], 9 October 1938. A. Goy &amp; L.S. Smith 544 (BM!) . NEW ZEALAND, North Island, Hawera, Mere Mere Rd., Hughie &amp; Black’s Gully, 200 ft [61 m], 2 January 1956. A.M Lysaght 1 (BM!) ; Auckland District: Waitemata Co., Huia, 5 August 1950, K. Wood s.n. (BM!).</p></div>	https://treatment.plazi.org/id/03A5792B77277074FF1FFCF5FEBBF82A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Salgado, Arthur Edward	Salgado, Arthur Edward (2021): Taxonomic and nomenclatural notes on Philippine ferns. IV. Asplenium pahli (Polypodiales, Aspleniineae, Aspleniaceae), a new species from Leyte, Philippines. Phytotaxa 524 (2): 71-84, DOI: 10.11646/phytotaxa.524.2.1
