taxonID	type	description	language	source
03A5792B772F707AFF1FF89DFA10FC9D.taxon	materials_examined	Type: — PHILIPPINES, Leyte, 500 m, 5 February 1914. C. A. Wenzel 572 (holotype P 00370339! designated here, isotypes P 00370338!, P 00370340!, C 0364253 F (F!), US 01516038!).	en	Salgado, Arthur Edward (2021): Taxonomic and nomenclatural notes on Philippine ferns. IV. Asplenium pahli (Polypodiales, Aspleniineae, Aspleniaceae), a new species from Leyte, Philippines. Phytotaxa 524 (2): 71-84, DOI: 10.11646/phytotaxa.524.2.1
03A5792B772F707AFF1FF89DFA10FC9D.taxon	diagnosis	Diagnosis: — Asplenium pahli and A. anisodontum have a proliferous rachis but differ in the pinna shape and serration. A. pahli differs from A. caudatum, A. falcatum, A. gueinzianum, A. planicaule and A. polyodon in the proliferous rachis, the pinna margins being irregularly serrate, sinuses of different depth and width, fewer sori distant from the costa, not contiguous, and straight or slightly falcate. Terrestrial. Rhizome short, erect, 0.4 – 0.5 cm thick, paleae and palea bases few, scattered, clathrate, black, lanceate, margin entire, <1 mm long, upper part of the paleae often broken off, cells small, polygonal. Roots many, crowded at the base of fronds, brown, thin, fragile when dried. Fronds erect, pinnate, crowded at the apex of the rhizome, spirally arranged, 25 – 45 cm long. Stipes 10.5 (13.0 – 17.0) 21.0 cm long, adaxial side grooved, abaxial side rounded, greenish, glabrous or glabrescent, with sparse paleae, paleae bicolorous, base and margin light brown, central area black, lanceate, ≤ 1 mm long, adpressed against the surface, margin entire, irregular with an occasional acute projection formed by a clathrate cell wall, palea apex acute. Lamina oblong, 15 – 35 cm long, (5.5) 6.5 – 8 cm wide, base truncate, pinnate, chartaceous, 12 – 15 alternating pinna pairs, pinnae slightly reduced at near apex. Rachis proliferous, abaxial side rounded, adaxial side with a wide U-shaped groove with prominent ridges, greenish, glabrescent, with adpressed small paleae, ≤ 1 mm long, lanceate, black, margin entire, apex acute, paleae on both sides of the rachis. Pinnae 3.2 – 4.0 cm long, 1.0 – 1.3 cm wide in the middle, noncontiguous, alternating, generally trapezoid, stalked to sessile, stalk ≤ 1 mm long, naked or with few paleae, lamina adaxial side glabrous, abaxial side glabrous or with a few reduced paleae on the costa, costa prominent of both sides, adaxial side grooved with prominent ridges gradually disappearing toward the pinna apex, abaxial side rounded, basal pinna pair sub-opposite or alternating, generally oblong, of the same length and shape as median pinnae or slightly smaller; pinna base basiscopic side cuneate, acroscopic side cuneate to broadly cuneate, pinna apex acute, pinna margins irregularly serrate, sinuses of different depth and width, acroscopic margin with 5 – 6 oblique widely separated teeth, larger teeth with 2 – 3 apical teeth, teeth blunt or acute, basiscopic margin serration similar to acroscopic margin, 3 – 5 oblique widely separated teeth, lamina apical segment usually missing, when present deformed with a proliferous bud at the insertion with the rachis. Venation free, costa visible with prominent ridges, 5 – 7 pairs of veins oblique to the costa, abaxial side veins not visible, adaxial side veins hardly visible, once forking, ending near the margin; hydathodes absent. Sori in a single row distal from pinna base, on both sides of the costa, originating about 1 mm from costa, oblique, widely separated, 3 – 6 sori on the acroscopic side, 2 – 5 on the basiscopic side, 5 – 7 mm long, occupying middle portion of lamina, ending before reaching the margin. Indusium thick, brown, entire, rolls back to expose mature sporangia, margin entire or slightly jagged. Distribution and ecology: — Island of Leyte, Central-Eastern Philippines. A terrestrial species growing on the eastern slopes of the Leyte Cordillera, at 500 m elevation, in lowland tropical evergreen rainforest. Climate warm and humid; rainfall evenly distributed throughout the year (Pagasa 2020). Etymology: — The species epithet honors Brother George Pahl, FSC, Ph. D., an American biologist and university professor who taught, mentored, and inspired thousands of students in the United States and the Philippines during his teaching career spanning more than 50 years. Conservation status: — A. pahli was collected in an area near the towns of Dagami and Jaro that is presently denuded of its original primary growth forest (Lasco et al. 2001; Mongabay 2020). The only collection known is that of Wenzel in 1914. This is a possible indication that A. pahli is endangered or possibly extinct in its type locality. There might be more plants scattered in the remaining stands of old growth deep in the Leyte Cordillera. Asplenium caudatum, A. falcatum, A. anisodontum and A. pahli represent different species as shown by the characters presented in this key.	en	Salgado, Arthur Edward (2021): Taxonomic and nomenclatural notes on Philippine ferns. IV. Asplenium pahli (Polypodiales, Aspleniineae, Aspleniaceae), a new species from Leyte, Philippines. Phytotaxa 524 (2): 71-84, DOI: 10.11646/phytotaxa.524.2.1
03A5792B77277074FF1FFADDFDE7FE9E.taxon	materials_examined	Holotype (designated by C. Presl 1849): — PHILIPPINES. Luzon: Cuming 128 (PRC non vide); isotypes L 0050982!, L 0050983!, BM 001045207!, BM 001045208!, BM 001045209!, BM 001045210, B 20 0011335!, B 20 0011341!, E 00210809!, MICH 1003398 B!, MW 0000516!, US 00065552!, W 0052502!.	en	Salgado, Arthur Edward (2021): Taxonomic and nomenclatural notes on Philippine ferns. IV. Asplenium pahli (Polypodiales, Aspleniineae, Aspleniaceae), a new species from Leyte, Philippines. Phytotaxa 524 (2): 71-84, DOI: 10.11646/phytotaxa.524.2.1
03A5792B77277074FF1FFB81FA31FD5A.taxon	materials_examined	FRENCH POLYNESIA, Society islands, Tahiti: vers 1200 m, Nadeaud s. n. (P!); Herbier E. Drake, Savatier s. n. (two sheets, P!); Herbier E. Drake, Nadeaud s. n. (P!); Vallée de la rivière Paho, Toahotu, crête sud, 149 ° 16 ’ W, 17 ° 47 ’ S, 650 m, Florence & Jourdan 11075 (US 01517559!); Island of Moorea: Mt. Muaroa, elev. 429 m, 17.54735 ºS, 149.84485 ºW, 12 August 2010, J. Nitta 570 (P 00847390!). Marquesas Islands, Ua Pou Island, Poumaka, mid-elevation wet forest, 680 m, 9 ° 23 ′ 26 ″, 140 ° 4 ′ 58 ″, 30 October 2017, Schuettpelz, E. Wood, K. R. & Butaud, J. - F. 1857 (US 01936878!); Hivaoa, F. W. Christian s. n. (P!); Fatu Hiva, K. R. Wood 4491, 2 sheets (P!).	en	Salgado, Arthur Edward (2021): Taxonomic and nomenclatural notes on Philippine ferns. IV. Asplenium pahli (Polypodiales, Aspleniineae, Aspleniaceae), a new species from Leyte, Philippines. Phytotaxa 524 (2): 71-84, DOI: 10.11646/phytotaxa.524.2.1
03A5792B77277074FF1FF8C5FB1AFC3A.taxon	materials_examined	Lectotype (designated by Salgado & Fraser-Jenkins 2013): — SRI LANKA: 1737, J. Burman Thesaurus Zeylanicus t. 43. Epitype (designated by Salgado & Fraser-Jenkins 2013): — SRI LANKA. North Central Province: Ritigala, 14 January 1951, W. A. Sledge 873 (US 01514788!). SRI LANKA, 1717. P. Herrmann s. n., Historical Collections, Herb. Herrmann 3: 47, no. 385, (BM 000621974!). North Western Province: Kurunegala District, R. B. Faden & A. J. Faden 76 / 400 (US 01514785!). INDIA. Courtallam: East India Company; s. coll. (K 001109576!, K 00119577!); Assam: Darrauz District, Balipara forest; M. Gustav s. n. (US 01514780!).	en	Salgado, Arthur Edward (2021): Taxonomic and nomenclatural notes on Philippine ferns. IV. Asplenium pahli (Polypodiales, Aspleniineae, Aspleniaceae), a new species from Leyte, Philippines. Phytotaxa 524 (2): 71-84, DOI: 10.11646/phytotaxa.524.2.1
03A5792B77277074FF1FFE25FB80FBAE.taxon	materials_examined	Holotype (verified by R. Vianey): — SOUTH AFRICA: Port Natal: Gueinzius s. n. (B 20 0015752!).	en	Salgado, Arthur Edward (2021): Taxonomic and nomenclatural notes on Philippine ferns. IV. Asplenium pahli (Polypodiales, Aspleniineae, Aspleniaceae), a new species from Leyte, Philippines. Phytotaxa 524 (2): 71-84, DOI: 10.11646/phytotaxa.524.2.1
03A5792B77277074FF1FFE51FDCBFB3E.taxon	materials_examined	Holotype (designated here by A. E. Salgado): — PHILIPPINES. Leyte, C. A. Wenzel 572, P 00370339!; isotypes P 00370338!, P 00370340!, US 01516038!, UC 267862!, C 0364253 F! (F).	en	Salgado, Arthur Edward (2021): Taxonomic and nomenclatural notes on Philippine ferns. IV. Asplenium pahli (Polypodiales, Aspleniineae, Aspleniaceae), a new species from Leyte, Philippines. Phytotaxa 524 (2): 71-84, DOI: 10.11646/phytotaxa.524.2.1
03A5792B77277074FF1FFF21FB98F94A.taxon	materials_examined	Lectotype (designated by Morton 1973): — NEPAL. 1820, Numerical List (Catalogue) No. 189 - 1, Wallich s. n., isotypes BM 001045235!, K 001109500!, K 001044577!, NY 214847!, UC 267862!. INDIA. Srinugur [Srinagar], 1820, Numerical List (Catalogue) No. 189 - 2, W. Moorcroft s. n., isotypes K 001109501!, NY 214848!; Nilgiris, Flora of Madras, 6000 ft [1828 m], May 1886, Gamble 17334 (US 01515892!). CHINA. Yunnan, Yangbi Xian, W side of Diancang Shan mountain range. En route from Xueshanhe to Dapingzi, growing on crevices of a ledge, 25 º 43 ’ N, 100 º 02 ’ E, 2300 - 3000 m, 17 June 1984, Sino-American Botanical Expedition to Yunnan Province 242 (US 3043222!); Yunnan Red River forest across from Maubao Mongtze, W. Hancock 187 (K 001092464!). COUNTRY UNKNOWN. Himalayas, J. F. Royle s. n. (PH 00025140!);	en	Salgado, Arthur Edward (2021): Taxonomic and nomenclatural notes on Philippine ferns. IV. Asplenium pahli (Polypodiales, Aspleniineae, Aspleniaceae), a new species from Leyte, Philippines. Phytotaxa 524 (2): 71-84, DOI: 10.11646/phytotaxa.524.2.1
03A5792B77277074FF1FFF21FB98F94A.taxon	discussion	Note: Morton (1973) designated Wallich No. 189 (B) as the lectotype of A. planicaule. The present author was not able to locate this specimen. There are doubtful isotypes at BM, K, LE, PH and S: BM 001045200!, BM 001045235!, BM 000523633!, BM 000523634!, BM 000523635!, K 001044578!, LE 00008417!, PH 00004925!, S-P- 1376! (S), UC 267862!.	en	Salgado, Arthur Edward (2021): Taxonomic and nomenclatural notes on Philippine ferns. IV. Asplenium pahli (Polypodiales, Aspleniineae, Aspleniaceae), a new species from Leyte, Philippines. Phytotaxa 524 (2): 71-84, DOI: 10.11646/phytotaxa.524.2.1
03A5792B77277074FF1FFCF5FEBBF82A.taxon	materials_examined	Lectotype (Lectotype designated by Nicolson & Fosberg 2003: 123) G. Forster s. n. (UPS-THUNB 24832!): — AUSTRALIA. Queensland: South-east region, Queen Mary Falls, 8 miles SE of Killarney, 13 December 1956. R. Schodde 316 (BM!); North of Moreton Bay, from Glasshouse Mountain to Mount Flinders and Stradborke Island, 1850 - 1851, no coll. (BM!); West Springbrook, 2,200 ft [670 m], 9 October 1938. A. Goy & L. S. Smith 544 (BM!). NEW ZEALAND, North Island, Hawera, Mere Mere Rd., Hughie & Black’s Gully, 200 ft [61 m], 2 January 1956. A. M Lysaght 1 (BM!); Auckland District: Waitemata Co., Huia, 5 August 1950, K. Wood s. n. (BM!).	en	Salgado, Arthur Edward (2021): Taxonomic and nomenclatural notes on Philippine ferns. IV. Asplenium pahli (Polypodiales, Aspleniineae, Aspleniaceae), a new species from Leyte, Philippines. Phytotaxa 524 (2): 71-84, DOI: 10.11646/phytotaxa.524.2.1
