identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03A2856AFF8FA3134B99FBF7E1CDFE08.text	03A2856AFF8FA3134B99FBF7E1CDFE08.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amaurodera projecta Assing & V 2015	<div><p>Amaurodera projecta nov.sp. (Figs 1-4, 19-23)</p><p>T y p e m a t e r i a l: Holotype 3: "N-Vietnam - 7 km NW Sa Pa, 22°22'48''N, 103°47'12''E, 1810 m, field margin, 31.VII.2013, V. Assing [4+2] / Holotypus 3 Amaurodera projecta sp. n., det. V. Assing 2015" (cAss). Paratype 3: "N-Vietnam - 7 km NW Sa Pa, 22°22'47''N, 103°47'10''E, 1850 m, decid. forest, 31.VII.2013, V. Assing [5+2]" (cAss).</p><p>E t y m o l o g y: The specific epithet (Latin, adjective: projecting) alludes to the conspicuous shape of the crista apicalis of the aedeagus.</p><p>D e s c r i p t i o n: Body length 5.7-6.2 mm; length of forebody 2.6 mm. Other measurements: head width: 0.69-0.71 mm; length of pronotum: 1.00- 1.05 mm; width of pronotum: 0.75-0.77 mm; elytral length at suture: 0.50-0.51 mm; elytra width: 0.90-0.98 mm. Habitus as in Fig. 19. Coloration: forebody dark-brown; abdomen: tergites III and IV reddish to reddish-brown with the posterior portions of the paratergites infuscate, tergite V dark-brown, with the anterior half slightly to distinctly paler, tergites VI and VII dark-brown, tergite VIII dark-brown with the posterior portion yellowish-brown, segments IX-X yellowish-brown; legs with brown tibiae and yellowish tarsi, profemora brown, meso- and metafemora dark-brown to blackish with broadly yellowish bases; head blackish; pronotum and blackish brown, with the humeral angles of the elytra dark- reddish and the elytra posterior margins narrowly pale-reddish; antennae with antennomeres I-II reddish, III-VII more or less distinctly infuscate, and VIII-XI pale reddish; maxillary palpi dark-brown with the terminal palpomere pale-yellowish.</p><p>Head (Fig. 1) weakly oblong, 1.03-1.05 times as long as broad, broadest across eyes; postero-lateral outline between eyes and posterior constriction very weakly convex in dorsal view; median dorsal portion impressed; median and anterior dorsal portions extensively without punctation and glossy, with nearly obsolete microsculpture; posterior and postero-lateral dorsal portions with shallow microsculpture and sparse setiferous punctures (Fig. 20). Eyes strongly convex, slightly more than half as long as distance from posterior margin of eye to posterior constriction. Antenna elongate, 3.3-3.4 mm long, and very slender (Fig. 2).</p><p>Pronotum (Fig. 1) approximately 1.35 times as long as broad and 1.06-1.09 times as broad as head; dorsal surface opaque due to very dense microgranules; midline with long and narrow furrow reaching neither anterior nor posterior margins; antero-lateral portions slightly elevated, each with four granules arranged in an oblique series and bearing stout black setae (Fig. 21); antero-lateral surface nearly vertical, without microsculpture and glossy.</p><p>Elytra (Fig. 1) 0.48-0.49 times as long as pronotum; punctation fine and moderately sparse; interstices without microsculpture, several times as broad as diameter of punctures. Hind wings present, but apparently of reduced length. Metatarsomere I as long as, or slightly longer than, the combined length of II and III.</p><p>Abdomen narrower than elytra; tergites III-IV with moderately deep, tergite V with very shallow anterior impressions; tergites III-VII with fine punctation only at and near posterior margins, otherwise impunctate; tergite VIII with granulose setiferous punctation in posterior third (male secondary sexual character?) (Fig. 3); microsculpture present and composed of short transverse meshes, but nearly obsolete and visible only at high magnification (at least 100 x); posterior margin of tergite VII with palisade fringe.</p><p>3: posterior margin of tergite VIII strongly serrate, on either side of middle with 3-4 pronounced tooth-like projections, median pair of projections distinctly longer than lateral teeth (Fig. 4); sternite VIII with broadly and weakly convex posterior margin; median lobe of aedeagus 0.9 mm long; ventral process of distinctive shape; crista apicalis enormous, strongly projecting (Figs 22-23).</p><p>♀: unknown.</p><p>C o m p a r a t i v e n o t e s: From all its congeners, except A. darenae YAN &amp; LI, 2015, Amaurodera projecta differs by the pronounced crista apicalis of the aedeagus and by the presence of a series of four setae in the antero-lateral portions of the pronotum. As can be inferred from these shared derived characters, as well as from the similarly shaped posterior margin of tergite VIII, A. projecta is closely allied to A. darenae (China: Guangxi), most likely its sister species, from which it is distinguished by larger size (no overlap; see measurements in YAN &amp; LI 2015a), different coloration particularly of the legs (uniformly pale reddish-yellow in A. darenae), the different shape of the apex of the ventral process of the aedaegus both in lateral and in ventral view, the more oblique crista apicalis (nearly parallel to ventral process and of slightly different shape in A. darenae), as well as the more massive apical and more slender basal internal structures of the aedeagus. For illustrations of A. darenae see YAN &amp; LI (2015a), for those of other</p><p>geographically close congeners from China and Thailand see ASSING (2009), PACE (1986, 1992), and YAN &amp; LI (2015a).</p><p>D i s t r i b u t i o n a n d n a t u r a l h i s t o r y: The type specimens were collected in two close localities 7 km to the northwest of Sa Pa, North Vietnam, by sifting debris and leaf litter in a field margin and in a secondary deciduous forest at altitudes of 1810 and 1850 m.</p></div>	https://treatment.plazi.org/id/03A2856AFF8FA3134B99FBF7E1CDFE08	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Assing, Volker;V, North	Assing, Volker, V, North (2015): On the Lomechusini fauna of Vietnam (Coleoptera: Staphylinidae: Aleocharinae). Linzer biologische Beiträge 47 (2): 1257-1282, DOI: 10.5281/zenodo.5282217
03A2856AFF89A3134B99FE0DE3E2FC9D.text	03A2856AFF89A3134B99FE0DE3E2FC9D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drusilla zyrasoides DVORAK 1988	<div><p>Drusilla zyrasoides DVOŘÁK, 1988 (Figs 41-50)</p><p>Drusilla zyrasoides DVORAK, 1988: 4 f.</p><p>T y p e m a t e r i a l e x a m i n e d Holotype 3: " Vietnam, Tam Dao, 4.-11.VI.1985,</p><p>Vinh phu prov., Jan Víša lgt. / Holotypus / ex coll. M. Dvořák, National Museum Prague, Czech</p><p>Republic / Drusilla zyrasoides sp. n., det. M. Dvořák.86" (NMP). Paratypes: 333, 3♀♀: same</p><p>data as holotype (NMP).</p><p>M a t e r i a l e x a m i n e d: Vietnam: 13, 3♀♀ Vinh Phu province, Tam Dao, 20.-28.VI.1990, leg. Dvořák (NMP, cAss).</p><p>C o m m e n t: The original description is based on a male holotype, a female allotype, and seven paratypes (five males and two females) from " Vietnam, Tam Dao, Vinh phu Provinz" (DVOŘÁK 1988). This species is characterized by a pronounced sexual dimorphism of the pronotum and of the abdominal segment VIII, as well as by the primary sexual characters. The external and sexual characters are illustrated in Figs 41-50.</p></div>	https://treatment.plazi.org/id/03A2856AFF89A3134B99FE0DE3E2FC9D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Assing, Volker;V, North	Assing, Volker, V, North (2015): On the Lomechusini fauna of Vietnam (Coleoptera: Staphylinidae: Aleocharinae). Linzer biologische Beiträge 47 (2): 1257-1282, DOI: 10.5281/zenodo.5282217
03A2856AFF89A31A4B99FCF0E333FDB8.text	03A2856AFF89A31A4B99FCF0E333FDB8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Drusilla perforans Assing & V 2015	<div><p>Drusilla perforans nov.sp. (Figs 5-7, 24-26, 60)</p><p>T y p e m a t e r i a l: Holotype 3: "N-Vietnam - 10 km NW Sa Pa, 22°22'26''N, 103°45'27''E, 1850 m, stream valley, 8.VIII.2013, V. Assing [9+2] / Holotypus 3 Drusilla perforans sp. n., det. V. Assing 2015" (cAss). Paratypes: 233, 2♀♀: same data as holotype (cAss).</p><p>E t y m o l o g y: The specific epithet is the present participle of the Latin verb perforare (to pierce) and alludes to the claw-shaped internal structures of the aedeagus.</p><p>D e s c r i p t i o n: Body length 4.5-5.1 mm; length of forebody 2.1-2.2 mm. Coloration: head and pronotum reddish (Fig. 5); elytra brown to dark-brown (Fig. 5); abdomen (Fig. 7): paratergites III-VI yellowish in anterior half and blackish in posterior half; tergites III-IV reddish to yellowish-brown; tergite V of similar coloration as tergites III- IV or somewhat darker in posterior half; tergite VI dark-brown, except for the diffusely yellowish lateral margins; tergite VII dark-brown with broadly yellowish to reddish posterior margin; segments VIII-X dark-yellowish; legs pale-yellowish, with the apical third to two-fifths of the metafemora broadly infuscate; antennae (Fig. 6) dark-brown with antennomeres I-II and the base of III yellowish and the apical 1-3 antennomeres reddish; maxillary palpi yellowish.</p><p>Head (Fig. 5) weakly oblong, approximately 1.1 times as long as broad, broadest across eyes; postero-lateral outline between eyes and posterior constriction very weakly convex in dorsal view; dorsal surface with sparse and extremely fine punctation. Eyes large and strongly convex, slightly longer than distance from posterior margin of eye to posterior constriction. Antenna 2.1-2.3 mm long and shaped as in Fig. 6.</p><p>Pronotum (Fig. 5) approximately 1.02 times as broad as long and 1.01-1.04 times as broad as head, broadest anteriorly; lateral margins distinctly sinuate in dorsal view; posterior angles obtusely marked; median portion impressed; midline with distinct median furrow reaching neither anterior nor posterior margins; punctation dense, rather coarse, and sexually dimorphic.</p><p>Elytra (Fig. 5) approximately as long as pronotum; punctation very dense, with the interstices reduced to narrow ridges, and sometimes weakly defined or even partly confluent. Hind wings fully developed. Metatarsomere I of very variable length, relatively short and slightly longer than II (much shorter than the combined length of II and III) to long and approximately as long as the combined length of II and III.</p><p>Abdomen (Fig. 7): tergites III-V with deep, tergite VI with very shallow anterior impressions; tergites III-VI with a baso-lateral puncture on either side; tergites III-IV with a median pair of punctures; tergites III-VII otherwise impunctate, except for few setiferous punctures near posterior margins; posterior margin of tergite VII with palisade fringe.</p><p>3: pronotum with more pronounced median impression and with somewhat granulose punctation; posterior margin of tergite VIII truncate in the middle; sternite VIII somewhat longer than tergite VIII and with convex posterior margin; median lobe of aedeagus (Figs 24-25) 0.66-0.69 mm long and of distinctive shape; internal sac with a pair of clawshaped sclerotized structures (ventral view).</p><p>♀: pronotum with less pronounced median impression and with non-granulose punctation; posterior margin of tergite VIII indistinctly truncate in the middle; posterior margin of sternite VIII weakly convex; spermatheca of distinctive shape (Fig. 26).</p><p>C o m p a r a t i v e n o t e s: Drusilla perforans differs from all other Drusilla species by the morphology of the aedeagus and the shape of the spermatheca, as well as by the coloration. It is additionally distinguished from D. zyrasoides DVOŘÁK, 1988, the only other species of the genus previously recorded from Vietnam, by different coloration ( D. zyrasoides: head, pronotum, and abdomen blackish; elytra distinctly bicoloured, darkyellowish with the postero-lateral portions blackish), the absence of a sexual dimorphism of the head, the oblong head, and numerous other characters (see Figs 41-50). For illustrations of other geographically close Drusilla species recorded from China, Thailand, and Laos see ASSING (in press a), MARUYAMA (2004a, 2004b), PACE (1986, 1992, 1993, 1998, 2000, 2003, 2004a, 2012a, 2012b).</p><p>D i s t r i b u t i o n a n d n a t u r a l h i s t o r y: The type specimens were sifted and roots from moist leaf litter in a steep stream valley with deciduous trees at an altitude of 1850 m (Fig. 60), approximately 10 km to the northwest of Sa Pa, North Vietnam. An association with ants or termites was not observed.</p></div>	https://treatment.plazi.org/id/03A2856AFF89A31A4B99FCF0E333FDB8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Assing, Volker;V, North	Assing, Volker, V, North (2015): On the Lomechusini fauna of Vietnam (Coleoptera: Staphylinidae: Aleocharinae). Linzer biologische Beiträge 47 (2): 1257-1282, DOI: 10.5281/zenodo.5282217
03A2856AFF80A3194B99FDDDE157FC08.text	03A2856AFF80A3194B99FDDDE157FC08.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Orphnebius fansipanicus Assing & V 2015	<div><p>Orphnebius fansipanicus nov.sp. (Figs 8-9, 27-29, 39)</p><p>T y p e m a t e r i a l: Holotype 3: "N-Vietnam - pass 8 km NW <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.766945&amp;materialsCitation.latitude=22.353611" title="Search Plazi for locations around (long 103.766945/lat 22.353611)">Sa Pa</a>, 22°21'13''N, 103°46'01''E, 2030 m, forest margin, 10.VIII.2013, V.Assing [10a+2] / Holotypus 3 Orphnebius fansipanicus sp. n., det. V. Assing 2015" (cAss).</p><p>E t y m o l o g y: The specific epithet is an adjective derived from Fansipan, the highest peak of Vietnam, near which the type locality is situated.</p><p>D e s c r i p t i o n: Body length 3.6 mm; length of forebody 1.6 mm. Coloration: forebody black; abdomen dark-reddish; legs with dark-brown femora and with reddish tibiae and tarsi; antennae with antennomeres I-IV reddish, V brown, and VI-XI blackishbrown; maxillary palpi dark-brown with pale-yellowish terminal palpomere.</p><p>Head (Fig. 8) 1.4 times as broad as long; dorsal surface with few scattered punctures and without microsculpture. Eyes large and bulging, more than twice as long as distance from posterior margin of eye to posterior constriction. Antenna 1.1 mm long, nearly symmetric, shaped as in Fig. 9.</p><p>Pronotum (Fig. 8) approximately 1.2 times as broad as long and 1.7 times as broad as head; disc with very few scattered setiferous punctures, nearly impunctate; lateral margins each with four long black setae, one at anterior angle, one a short distance (about half the length of seta) behind anterior angle, one just behind middle, and one near posterior angle.</p><p>Elytra (Fig. 8) approximately 0.85 times as long as pronotum; punctation fine and sparse; pubescence long, pale, and depressed to suberect; microsculpture absent. Hind wings fully developed. Metatarsomere I nearly as long as the combined length of II and III.</p><p>Abdomen: tergites III-VI each with a lateral puncture and few punctures at posterior margins, otherwise impunctate; tergite VII with extensive oblong puncture-like impressions in anterior two-thirds, posteriorly with a transverse row of setiferous punctures; posterior margin of tergite VII with pronounced palisade fringe.</p><p>3: median lobe of aedeagus 0.75 mm long, with rather short and straight (lateral view) ventral process (Figs 27-28); paramere (Figs 29, 39) much shorter than median lobe, paramerite without conspicuous modifications, at apical margin of sclerotized portion with four setae, condylite stout, much shorter than paramerite, and with short velum.</p><p>♀: unknown.</p><p>C o m p a r a t i v e n o t e s: Based on the modifications of the abdominal segments IX and X (with long and dense dark setae) and on the morphology of the aedeagus (shapes of median lobe and of parameres), Orphnebius fansipanicus belongs to the O. hauseri group (see ASSING 2006a, 2006b), whose distribution ranges from the western Himalaya southeastwards to China and Taiwan (ASSING 2015a). Except for O. thai PACE, 2000 (Thailand), whose aedeagal shape suggests that it may belong to this group, too, all other species previously described from Thailand and Laos belong to other species groups. Among the species of the O. hauseri group, O. fansipanicus is most similar to O. bihamatus ASSING, 2015 (East Yunnan), from which it differs by the proportions of head and pronotum ( O. dishamatus: pronotum 1.35 times as broad as long and 1.25 times as broad as head), the coloration ( O. dishamatus: abdomen pale-reddish; legs with brown femora and reddish-brown tibiae), the slightly longer antennomere XI, the chaetotaxy of the pronotum ( O. dishamatus: the two antero-lateral long setae separated by a greater distance), the shape of the median lobe of the aedeagus ( O. dishamatus: ventral process broader in ventral view; crista apicalis of different orientation and slightly different shape), and by the shape of the paramere (O. bihamatus: apex of paramere of slightly different shape; condylite longer in relation to paramerite). For illustrations of the Orphnebius species recorded from Thailand and China see ASSING (2006a, 2006b, 2009, 2015a) and PACE (1986, 1992, 2000, 2004a), for figures of O. bihamatus and a key to the species known from China see Figs 30, 40 and ASSING (2015a), respectively.</p><p>D i s t r i b u t i o n a n d n a t u r a l h i s t o r y: The type locality is situated at a pass 8 km to the northwest of Sa Pa, North Vietnam, at an altitude of 2030 m. The holotype was sifted from litter and roots under bushes near the margin of a secondary deciduous forest, together with Tetrabothrus bicolor . For a habitat photo see figure 27 in ASSING (2015c).</p></div>	https://treatment.plazi.org/id/03A2856AFF80A3194B99FDDDE157FC08	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Assing, Volker;V, North	Assing, Volker, V, North (2015): On the Lomechusini fauna of Vietnam (Coleoptera: Staphylinidae: Aleocharinae). Linzer biologische Beiträge 47 (2): 1257-1282, DOI: 10.5281/zenodo.5282217
03A2856AFF83A3194B99FB08E230FA2D.text	03A2856AFF83A3194B99FB08E230FA2D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Orphnebius willersi PACE 2001	<div><p>Orphnebius willersi PACE, 2001 (Figs 51-52)</p><p>Orphnebius willersi PACE, 2001: 196 .</p><p>T y p e m a t e r i a l e x a m i n e d: Holotype 3: " Vietnam, N, 100 km SSW Hanoi, 40 km SW Than Hoa, Ben En National Park, 50 m, 23.-29.VII.1997, leg. A. Napolov / Holotypus Orphnebius willersi mihi, det. R. Pace 2001 / Orphnebius willersi mihi, det. R. Pace 2001 " (NME).</p><p>C o m m e n t: The original description is based on a unique holotype (PACE 2001). The specimen is in very poor condition (antennomeres III-XI of left antenna and IV-XI of right antenna, right protarsus, left mesotarsus, right mid-leg, and right hind-leg missing). It is illustrated in Figs 51-52.</p></div>	https://treatment.plazi.org/id/03A2856AFF83A3194B99FB08E230FA2D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Assing, Volker;V, North	Assing, Volker, V, North (2015): On the Lomechusini fauna of Vietnam (Coleoptera: Staphylinidae: Aleocharinae). Linzer biologische Beiträge 47 (2): 1257-1282, DOI: 10.5281/zenodo.5282217
03A2856AFF83A3074B99FA60E0D6FC78.text	03A2856AFF83A3074B99FA60E0D6FC78.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Peltodonia atripalpis Assing & V 2015	<div><p>Peltodonia atripalpis nov.sp. (Figs 10-11, 37-38, 61)</p><p>T y p e m a t e r i a l: Holotype 3: "N-Vietnam - 6 km NW <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.78861&amp;materialsCitation.latitude=22.360834" title="Search Plazi for locations around (long 103.78861/lat 22.360834)">Sa Pa</a>, 22°21'39''N, 103°47'19''E, 1810 m, pasture margin, 7.VIII.2013, Wunderle [8+2] / Holotypus 3 Peltodonia atripalpis sp.n., det. V. Assing 2015" (cAss).</p><p>E t y m o l o g y: The specific epithet (Latin, adjective: with black palpi) alludes to the dark coloration of the maxillary palpi, one of the characters distinguishing this species from the similar P. chinensis (PACE, 1998) .</p><p>D e s c r i p t i o n: Body length 3.1 mm; length of forebody 1.3 mm. Coloration: head pale-brown with yellowish labrum; pronotum pale-brown with yellowish margins; elytra yellowish-brown; abdomen pale-reddish, with the posterior portion of tergite VII and the anterior portion of tergite VIII darker; legs yellow; antennae pale-reddish; maxillary palpi blackish-brown.</p><p>Head (Fig. 10) strongly wedge-shaped and strongly transverse; posterior margin sharply angled in lateral view, its outline nearly straight in dorsal view. Eyes large and bulging, nearly four times as long as postocular region. Antenna 1.0 mm long and shaped as in Fig. 11.</p><p>Pronotum (Fig. 10) strongly transverse, 1.75 times as broad as long and 1.3 times as broad as head; posterior margin broadly convex; posterior angles not marked; disc without microsculpture and with sparse punctures bearing moderately long pale setae; lateral margins each with three very long and stout black setae, posterior margin laterally with a somewhat shorter black seta on either side.</p><p>Elytra (Fig. 10) approximately as long as pronotum; punctation and pubescence similar to those of pronotum; laterally with three conspicuously long (even longer than lateral setae of pronotum) and stout black setae. Hind wings fully developed. Metatarsomere I as long as the combined length of II and III.</p><p>Abdomen: sternites III-VII posteriorly with a row of very long and stout black setae; tergites III-VI postero-laterally with two long and stout black setae on either side; tergite VII with pronounced longitudinal sculpture (gland openings?) in posterior third; tergite VIII with dense and coarse longitudinal rugae in anterior half and with oval impressions posteriorly, in posterior fourth with two transverse series each composed of six punctures bearing fine black setae, posterior margin with pronounced obtuse projection in the middle; posterior margin of tergite VII with pronounced palisade fringe.</p><p>3: posterior margin of sternite VIII moderately convex; median lobe of aedeagus 0.38 mm long, slender, weakly sclerotized, and with a long and straight flagellum in internal sac (Figs 37-38).</p><p>♀: unknown.</p><p>C o m p a r a t i v e n o t e s: According to HLAVÁČ et al. (2011), Peltodonia was was represented by seven species, one from the West Palaearctic, four from Borneo, one from Peninsular Malaysia, and one from China. Eight additional species from Borneo were described as Chaetosogonocephus PACE, 1987 (a junior synonym of Peltodonia since 2009) by PACE (2014, 2015).</p><p>In external characters, the new species is similar to P. chinensis (PACE, 1998), whose original description is based on a unique female from the Chinese province Zhejiang (PACE 1998), from where the species was again recorded by ASSING (2009). Peltodonia atricalpis is distinguished from P. chinensis by slightly smaller body size, shorter antennae, a less transverse pronotum, dark maxillary palpi, and particularly by the much smaller and differently shaped median lobe of the aedeagus. For comparison, the male median lobe of the aedeagus of P. chinensis is illustrated in Figs 35-36.</p><p>D i s t r i b u t i o n a n d n a t u r a l h i s t o r y: The type locality is situated 6 km to the northwest of Sa Pa, North Vietnam, at an altitude of 1810 m. The holotype was sifted from litter and roots under bushes and small trees at the margin of a pasture (Fig. 61).</p></div>	https://treatment.plazi.org/id/03A2856AFF83A3074B99FA60E0D6FC78	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Assing, Volker;V, North	Assing, Volker, V, North (2015): On the Lomechusini fauna of Vietnam (Coleoptera: Staphylinidae: Aleocharinae). Linzer biologische Beiträge 47 (2): 1257-1282, DOI: 10.5281/zenodo.5282217
03A2856AFF9DA3074B99FB9DE3B9FA51.text	03A2856AFF9DA3074B99FB9DE3B9FA51.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tetrabothrus bicolor Cameron 1939	<div><p>Tetrabothrus bicolor CAMERON, 1939 (Figs 12-14, 31-32)</p><p>Tetrabothrus bicolor CAMERON, 1939: 375 .</p><p>T y p e m a t e r i a l e x a m i n e d: Lectotype 3, present designation: "64504 / Birmah, Ruby M.es / Doherty / Fry Coll., 1905.100. / Tetrabothrus bicolor Cam Type / Type / Lectotypus 3 Tetrabothrus bicolor Cameron, desig. V. Assing 2015 / Tetrabothrus bicolor Cameron, det. V. Assing 2015" (BMNH). Paralectotype ♀: "Birmah, Ruby M.es / Doherty / Fry Coll., 1905.100. / Tetrabothrus bicolor Cam Cotype / Syntype / M. Cameron Bequest., B.M. 1955-147 / Tetrabothrus bicolor Cameron, det. V. Assing 2015" (BMNH).</p><p>M a t e r i a l e x a m i n e d: Vietnam: 13, 5♀♀, pass 8 km NW <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.76667&amp;materialsCitation.latitude=22.35" title="Search Plazi for locations around (long 103.76667/lat 22.35)">Sa Pa</a>, 22°21'N, 103°46'E, 2030 m, margin of secondary deciduous forest with bushes, bamboo, and herbs, litter, roots, and moss sifted, 9.VIII.2013, leg. Assing (cAss) ; 13, 1♀, same data, but 10.VIII.2013 (cAss) .</p><p>C o m m e n t: The original description is based on an unspecified number of syntypes from " Burma: Ruby Mines" (CAMERON 1939). Two syntypes, a male and a female, were located in the collections of the BMNH. The male is designated as the lectotype. The additional material listed above represents the first records since the original description. The external and male sexual characters are illustrated in Figs 12-14, 31-32.</p></div>	https://treatment.plazi.org/id/03A2856AFF9DA3074B99FB9DE3B9FA51	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Assing, Volker;V, North	Assing, Volker, V, North (2015): On the Lomechusini fauna of Vietnam (Coleoptera: Staphylinidae: Aleocharinae). Linzer biologische Beiträge 47 (2): 1257-1282, DOI: 10.5281/zenodo.5282217
03A2856AFF9DA3064B99FA45E39AFD9F.text	03A2856AFF9DA3064B99FA45E39AFD9F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tetrabothrus japonicus Nakane 1991	<div><p>Tetrabothrus japonicus NAKANE, 1991</p><p>Tetrabothrus vietnamiculus PACE, 2013: 375; nov.syn.</p><p>T y p e m a t e r i a l e x a m i n e d: Holotype 3: " Viet Nam N ( Ha Nang) 160 km NNW Ha Noi, 50-200 m, NE env. of Na Hang, 1/14.VI-96, J. Roma &amp; A. Napalov / Ex R. Pace Collection / Tetrabothrus vietnamiculus n. sp., det. R. Pace 2012 / Holotypus Tetrabothrus vietnamiculus mihi, det. R. Pace 2012 / Tetrabothrus japonicus Nakane, det. V. Assing 2015" (NME).</p><p>A d d i t i o n a l m a t e r i a l e x a m i n e d: Vietnam: 13, Cao Bang province, 12 km NE Cao Bang, 22°46'N, 106°19'E, 600-700 m, 15.-16.V.2010, leg. Dembický (NHMB).</p><p>C o m m e n t: The original description of Tetrabothrus vietnamiculus is based on a unique male from Vietnam (PACE 2013). The external and male sexual characters of the holotype (see also the figures in the original description) are identical to those illustrated for T. japonicus by YAN &amp; LI (2015b) for T. japonicus . According to MARUYAMA (email, 24.IX.2015), the interpretation of T. japonicus in YAN &amp; LI (2015b) is correct. Consequently, T. vietnamiculus is placed in synonymy with T. japonicus .</p></div>	https://treatment.plazi.org/id/03A2856AFF9DA3064B99FA45E39AFD9F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Assing, Volker;V, North	Assing, Volker, V, North (2015): On the Lomechusini fauna of Vietnam (Coleoptera: Staphylinidae: Aleocharinae). Linzer biologische Beiträge 47 (2): 1257-1282, DOI: 10.5281/zenodo.5282217
03A2856AFF9CA3064B99FCEBE18AFB41.text	03A2856AFF9CA3064B99FCEBE18AFB41.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zyras (Glossacantha) laocaiensis (: HLAVAC, NEWTON & MARUYAMA 2008) PACE 2008	<div><p>Zyras (Glossacantha) laocaiensis PACE, 2008, nov.comb. (Fig. 54)</p><p>Zyras (Pella) laocaiensis PACE, 2008: 11 f.</p><p>Pella laocaiensis: HLAVÁČ, NEWTON &amp; MARUYAMA (2011) .</p><p>T y p e m a t e r i a l e x a m i n e d: Holotype 3 [in poor condition]: " Vietnam, Lao Cai, ca. 40 km SE Sapa, 1400 m, 15-16. II.05, G. de Rougemont leg. / Holotypus Zyras laocaiensis n. sp., det. R. Pace 2005 / Zyras laocaiensis n. sp., det. R. Pace 2005" (cRou).</p><p>C o m m e n t: PACE (2008) assigned Z. laocaiensis to Pella, which he regarded as a subgenus of Zyras, although Pella had been attributed generic status in a revision by MARUYAMA (2006). The species was listed as Pella laocaiensis by HLAVÁČ et al. (2011). An examination of the holotype revealed that it does not belong to Pella, but to Zyras . According to MARUYAMA (e-mail 7 October, 2015), it belongs to the subgenus Glossacantha GEMMINGER &amp; HAROLD, 1868 sensu lato . The forebody of the holotype is illustrated in Fig. 54.</p></div>	https://treatment.plazi.org/id/03A2856AFF9CA3064B99FCEBE18AFB41	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Assing, Volker;V, North	Assing, Volker, V, North (2015): On the Lomechusini fauna of Vietnam (Coleoptera: Staphylinidae: Aleocharinae). Linzer biologische Beiträge 47 (2): 1257-1282, DOI: 10.5281/zenodo.5282217
03A2856AFF9CA3064B99FDFFE31EFCAB.text	03A2856AFF9CA3064B99FDFFE31EFCAB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zyras (Glossacantha) pokarzhevskii DVORAK 1996	<div><p>Zyras (Glossacantha) pokarzhevskii DVOŘÁK, 1996 (Fig. 55)</p><p>Zyras (Glossacantha) pokarzhevskii DVOŘAK, 1996: 7 f.</p><p>T y p e m a t e r i a l e x a m i n e d: Holotype ♀: " Vietnam, 30.XII.1980, Taj Nguen, Bud Lioj, 65 km N Ankche, A. Pokarževskij lgt. / Holotypus / ex coll. M. Dvořák, National Museum Prague, Czech Republic / Zyras pokarzhevskii sp. n., det. M. Dvořák. 88" (NMP).</p><p>C o m m e n t: The original description is based on a unique female holotype. Its forebody is illustrated in Fig. 55. The subgeneric assignment was confirmed by MARUYAMA (e-mail 16 October, 2015) based on a photo of the holotype.</p></div>	https://treatment.plazi.org/id/03A2856AFF9CA3064B99FDFFE31EFCAB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Assing, Volker;V, North	Assing, Volker, V, North (2015): On the Lomechusini fauna of Vietnam (Coleoptera: Staphylinidae: Aleocharinae). Linzer biologische Beiträge 47 (2): 1257-1282, DOI: 10.5281/zenodo.5282217
03A2856AFF9CA3064B99FB55E320FA3E.text	03A2856AFF9CA3064B99FB55E320FA3E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zyras (Zyras) malaisei SCHEERPELTZ 1965	<div><p>Zyras (Zyras) malaisei SCHEERPELTZ, 1965</p><p>M a t e r i a l e x a m i n e d: Vietnam: 1♀, pass 8 km NW Sa Pa, 22°21'N, 103°46'E, 2030 m, margin of secondary deciduous forest with bushes, bamboo, and herbs, litter, roots, and moss sifted, 9.VIII.2013, leg. Assing (cAss).</p><p>C o m m e n t: This species was previously known only from the type locality in the northeast of Burma, close to the border with the Chinese province Yunnan. The above female represents the first record of Z. malaisei from Vietnam.</p></div>	https://treatment.plazi.org/id/03A2856AFF9CA3064B99FB55E320FA3E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Assing, Volker;V, North	Assing, Volker, V, North (2015): On the Lomechusini fauna of Vietnam (Coleoptera: Staphylinidae: Aleocharinae). Linzer biologische Beiträge 47 (2): 1257-1282, DOI: 10.5281/zenodo.5282217
03A2856AFF9CA3054B99FA50E2BAFE5C.text	03A2856AFF9CA3054B99FA50E2BAFE5C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zyras (Zyras) neoparageminus HLAVAC, NEWTON & MARUYAMA 2011	<div><p>Zyras (Zyras) neoparageminus HLAVÁČ, NEWTON &amp; MARUYAMA, 2011</p><p>Zyras (Zyras) parageminus PACE, 2010: 319 ff.; preoccupied.</p><p>Zyras (Zyras) neoparageminus HLAVAC, NEWTON &amp; MARUYAMA, 2011: 7; replacement name.</p><p>Zyras (Zyras) subgeminus PACE, 2012a: 339; replacement name; nov.syn.</p><p>C o m m e n t: Zyras parageminus PACE, 2010, which is based on a unique male from Sumatra, is a junior primary homonym of Z. parageminus PACE, 1988 . The name was replaced with the nomen novum Z. neoparageminus by HLAVÁČ et al. (2011). Recording this species from Vietnam based on six females and evidently unaware of this replacement, PACE (2012a) proposed " Z. subgeminus Pace, 2010 nom. n. " [sic] as another nomen novum for Z. parageminus PACE, 2010 . Zyras subgeminus PACE, 2012 is consequently an objective junior synonym of Z. neoparageminus . The record from Vietnam most likely refers to Z. hongkongensis PACE, 1999 .</p></div>	https://treatment.plazi.org/id/03A2856AFF9CA3054B99FA50E2BAFE5C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Assing, Volker;V, North	Assing, Volker, V, North (2015): On the Lomechusini fauna of Vietnam (Coleoptera: Staphylinidae: Aleocharinae). Linzer biologische Beiträge 47 (2): 1257-1282, DOI: 10.5281/zenodo.5282217
03A2856AFF9FA3054B99FC7BE399FAAA.text	03A2856AFF9FA3054B99FC7BE399FAAA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zyras (Zyras) funestus (Dvorak 1996) Assing & V 2015	<div><p>Zyras (Zyras) funestus (DVOŘÁK, 1996), nov.comb. (Figs 56-59)</p><p>Orphnebius (Mesocephalobius) funestus DVOŘAK, 1996: 1 f.</p><p>T y p e m a t e r i a l e x a m i n e d Holotype ♀: " Vietnam, 8.1.1981, Taj Nguen, Bud Lioj, 65 km N Ankche, A. Pokarževskij lgt. / Holotypus / Orphnebius funestus sp. n., det. M. Dvořák.88 / ex coll. M. Dvořák, National Museum Prague, Czech Republic / Zyras funestus (Dvořák), det. V. Assing 2015" (NMP). Paratype ♀: same data as holotype, but " Allotypus " (NMP).</p><p>C o m m e n t: According to the original description, the type material is based on a male holotype, a female allotype, and a male paratype (DVOŘÁK 1996). The holotype and the allotype were located in the collections of the NMP. An examination of these specimens revealed that both of them are females and that they belong to Zyras sensu strictu, clearly not to Orphnebius (Figs 56-59). The species is closely allied to Zyras hirtus (KRAATZ, 1859), Z. setosivestis SCHEERPELTZ, 1965, and related species.</p></div>	https://treatment.plazi.org/id/03A2856AFF9FA3054B99FC7BE399FAAA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Assing, Volker;V, North	Assing, Volker, V, North (2015): On the Lomechusini fauna of Vietnam (Coleoptera: Staphylinidae: Aleocharinae). Linzer biologische Beiträge 47 (2): 1257-1282, DOI: 10.5281/zenodo.5282217
03A2856AFF9FA3054B99FDB3E315FD0E.text	03A2856AFF9FA3054B99FDB3E315FD0E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zyras (Zyras) hongkongensis PACE 2012	<div><p>Zyras (Zyras) hongkongensis PACE, 2012</p><p>C o m m e n t: This species is widespread in the southern East Palaearctic and the Oriental regions. It is present also in Vietnam; primary records will be listed in a study of the Chinese fauna of Zyras sensu strictu (ASSING in press b).</p></div>	https://treatment.plazi.org/id/03A2856AFF9FA3054B99FDB3E315FD0E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Assing, Volker;V, North	Assing, Volker, V, North (2015): On the Lomechusini fauna of Vietnam (Coleoptera: Staphylinidae: Aleocharinae). Linzer biologische Beiträge 47 (2): 1257-1282, DOI: 10.5281/zenodo.5282217
03A2856AFF9FA3054B99FD00E2ADFC1B.text	03A2856AFF9FA3054B99FD00E2ADFC1B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zyras (Zyras) trapeziceps DVORAK 1996	<div><p>Zyras (Zyras) trapeziceps DVOŘÁK, 1996 (Fig. 53)</p><p>Zyras (Zyras) trapeziceps DVORAK, 1996: 5 f.</p><p>T y p e m a t e r i a l e x a m i n e d: Holotype ♀ [with worker of Leptogenys sp. attached to the same pin]: " Vietnam sept. 1990, Sa-Pa, 11-19. VI., (Hoang Lien Son), Mir. Dvořák legit. / Holotypus / Zyras trapeziceps sp. n., det. M. Dvořák. 90" (NMP).</p><p>C o m m e n t: The original description is based on a unique female holotype; its forebody is illustrated in Fig. 53. According to MARUYAMA (e-mail 7 October, 2015), this species is correctly placed in Zyras sensu strictu.</p></div>	https://treatment.plazi.org/id/03A2856AFF9FA3054B99FD00E2ADFC1B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Assing, Volker;V, North	Assing, Volker, V, North (2015): On the Lomechusini fauna of Vietnam (Coleoptera: Staphylinidae: Aleocharinae). Linzer biologische Beiträge 47 (2): 1257-1282, DOI: 10.5281/zenodo.5282217
03A2856AFF9FA3044B99FAECE170FD71.text	03A2856AFF9FA3044B99FAECE170FD71.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zyras (Zyras) quasar DVORAK 1996	<div><p>Zyras (Zyras) quasar DVOŘÁK, 1996</p><p>Zyras (Pella) quasar DVOŘAK, 1996: 6 f.</p><p>Zyras (Zyras) quasar: HLAVAC et al. (2011) .</p><p>T y p e m a t e r i a l e x a m i n e d Holotype 3 [dissected prior to present study]: " Vietnam, 20- 30.12.1980, Taj Nguen, Bud Lioj, 65 km N Ankche, A. Pokarževskij legit. / Holotypus / Zyras quasar sp. n., det. M. Dvořák,90 / ex coll. M. Dvořák, National Museum Prague, Czech Republic " (NMP).</p><p>C o m m e n t: According to the original description, the unique holotype is a female (DVOŘÁK 1996). An examination of the specimen, however, revealed that it is a male. The species has been reported also from Burma, Thailand, and Malaysia, but, owing to frequent misidentifications of Zyras sensu strictu species, all the records from countries other than Vietnam require confirmation.</p><p>Zyras (Zyras) alboantennatus PACE, 1986</p><p>M a t e r i a l e x a m i n e d: Vietnam: 16 exs., Tuyen Quang province, Na Hang reserve [22°16- 31'N, 105°22-29'E], 360 m, 6.-24.VI.1997, leg. Peck (NMP, cAss).</p><p>C o m m e n t: This recently described species was previously known only from Burma. It is dealt with in detail in a revision of Zyras sensu strictu from China (ASSING in press b).</p><p>Zyras (Zyras) sichuanorum PACE, 2012</p><p>M a t e r i a l e x a m i n e d: Vietnam: 16 exs., Tuyen Quang province, Na Hang reserve [22°16- 31'N, 105°22-29'E], 360 m, 6.-24.VI.1997, leg. Peck (NMP, cAss).</p><p>C o m m e n t: This recently described species was previously known only from Sichuan, China. It is dealt with in detail in a revision of Zyras sensu strictu from China (ASSING in press b).</p></div>	https://treatment.plazi.org/id/03A2856AFF9FA3044B99FAECE170FD71	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Assing, Volker;V, North	Assing, Volker, V, North (2015): On the Lomechusini fauna of Vietnam (Coleoptera: Staphylinidae: Aleocharinae). Linzer biologische Beiträge 47 (2): 1257-1282, DOI: 10.5281/zenodo.5282217
03A2856AFF9EA3044B99FB8DE170FA82.text	03A2856AFF9EA3044B99FB8DE170FA82.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zyras (Zyras)	<div><p>Zyras (Zyras) sp. 2</p><p>M a t e r i a l e x a m i n e d: Vietnam: 1♀, 10 km NW Sa Pa, 22°22'N, 103°45'E, 1850 m, moist stream valley with deciduous trees, litter and roots between rocks sifted, 8.VIII.2013, leg. Wunderle (cWun).</p><p>C o m m e n t: The above specimen is characterized by a completely black body. It is, however, not conspecific with any of the similarly coloured species recorded from China (ASSING in press b).</p></div>	https://treatment.plazi.org/id/03A2856AFF9EA3044B99FB8DE170FA82	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Assing, Volker;V, North	Assing, Volker, V, North (2015): On the Lomechusini fauna of Vietnam (Coleoptera: Staphylinidae: Aleocharinae). Linzer biologische Beiträge 47 (2): 1257-1282, DOI: 10.5281/zenodo.5282217
03A2856AFF9EA3044B99FCA5E2D5FB89.text	03A2856AFF9EA3044B99FCA5E2D5FB89.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zyras (Zyras)	<div><p>Zyras (Zyras) sp. 1</p><p>M a t e r i a l e x a m i n e d: Vietnam: 1♀, 6 km NW Sa Pa, 22°22'N, 103°47'E, 1810 m, margin of pasture with bushes and small trees, litter and roots sifted, 7.VIII.2013, leg. Assing (cAss).</p><p>C o m m e n t: The above female probably represents an undescribed species closely allied to Zyras setosivestis SCHEERPELTZ, 1965, a species described and known only from Burma. It differs from Z. funestus, which belongs to the same species group, by distinctly larger size, darker coloration, much longer and more slender antennae, and by the distinctly finer and shallower punctation of the forebody.</p></div>	https://treatment.plazi.org/id/03A2856AFF9EA3044B99FCA5E2D5FB89	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Assing, Volker;V, North	Assing, Volker, V, North (2015): On the Lomechusini fauna of Vietnam (Coleoptera: Staphylinidae: Aleocharinae). Linzer biologische Beiträge 47 (2): 1257-1282, DOI: 10.5281/zenodo.5282217
03A2856AFF9EA3024B99FA94E136FB28.text	03A2856AFF9EA3024B99FA94E136FB28.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zyras (Zyras) fansipanicus Assing & V 2015	<div><p>Zyras (Zyras) fansipanicus nov.sp. (Figs 15-18, 33-34, 62)</p><p>T y p e m a t e r i a l: Holotype 3: "N-Vietnam - 7 km NW Sa Pa, 22°22'47''N, 103°47'10''E, 1850 m, decid. forest, 31.VII.2013, V. Assing [5+2] / Holotypus 3 Zyras fansipanicus sp. n., det. V. Assing 2015" (cAss). Paratype ♀: "N-Vietnam - pass 8 km NW Sa Pa, 22°21'10''N, 103°46'01''E, 2010 m, second. forest, 12.VIII. 2013, V. Assing [7b+2]" (cAss).</p><p>E t y m o l o g y: The specific epithet is an adjective derived from Fansipan, the highest peak of Vietnam, near which the type locality is situated.</p><p>D e s c r i p t i o n: Rather large species; body length 7.0-8.0 mm; length of forebody 3.4-3.7 mm. Coloration: head and pronotum blackish; elytra bicoloured, with the anterior third to two-fifths and the suture reddish-yellow and the posterior portion extensively blackish; abdomen blackish, with the paratergites and the posterior margins of segments III-VI narrowly, those of segments VII and VIII more broadly reddish; legs and maxillary palpi yellowish; antennae dark-brown, with the basal 2-3 antennomeres pale-reddish and antennomere XI reddish to dark-reddish.</p><p>Head (Fig. 15) distinctly transverse, narrowly to broadly impunctate along middle; punctation in lateral dorsal portions moderately dense and rather fine. Eyes slightly longer than postocular region in dorsal view. Antenna (Fig. 16) rather massive and 2.7- 2.9 mm long; antennomeres IV-VII as long as broad or weakly oblong, VIII-IX very weakly transverse, X much less than 1.5 times as broad as long, and XI approximately as long as the combined length of IX and X.</p><p>Pronotum (Fig. 15) rather large in relation to head, distinctly transverse, 1.14-1.15 times as broad as long, and approximately 1.36 times as broad as head; punctation fine to moderately fine and moderately dense, more or less equally distributed; interstices on average broader than diameter of punctures; midline narrowly impunctate; lateral margins each with two moderately long setae, one at anterior angles and one at anterior third. Elytra (Fig. 15) 0.76-0.77 times as long as pronotum; punctation moderately dense and not very coarse, equally distributed. Hind wings fully developed. Metatarsomere I longer than the combined length of II-IV.</p><p>Abdomen (Fig. 17) approximately as broad as elytra, with rather shallow anterior impressions on tergites III-V; anterior impressions of tergites III-V with sparse and fine non-setiferous punctures; disc of tergite III with a lateral puncture on either side and with four punctures at posterior margin; tergite IV with a lateral marginal puncture on either side and with eight punctures at posterior margin; tergite V with a lateral puncture on either side, with a median pair of punctures, and with 10-12 punctures at posterior margin; tergite VI with a transverse band of non-setiferous punctures near anterior margin, with a lateral puncture on either side, with a transverse series of eight punctures in posterior portion, and with approximately twelve punctures at posterior margin; tergite VII (Fig. 18) with a transverse band of dense non-setiferous punctures anteriorly and with three partly somewhat irregular transverse series of setiferous punctures in posterior half; tergite VIII (Fig. 18) in posterior portion with punctation bearing long black setae; integument without microsculpture and glossy; posterior margin of tergite VII with palisade fringe; posterior margin of tergite VIII truncate or weakly concave in the middle.</p><p>3: sternite VIII somewhat longer than tergite VIII, its posterior margin convex; median lobe of aedeagus 0.95 mm long and shaped as in Figs 33-34; paramere of the usual condition, with moderately long apical lobe.</p><p>♀: posterior margin of sternite VIII weakly concave in the middle.</p><p>C o m p a r a t i v e n o t e s: The new species belongs to a very speciose group of species with a similar coloration pattern (ASSING in press b). In both external and the male sexual characters, it is most similar to Z. gilvipalpis ASSING, in press from East Yunnan. It differs from that species particularly by the less dense and equally distributed punctation of the elytra and by the different shape of the ventral process of the aedeagus (more slender in lateral and in ventral view, and with a less acute apex in ventral view.</p><p>D i s t r i b u t i o n a n d n a t u r a l h i s t o r y: The type specimens were collected in two localities at and near a pass to the northwest of Sa Pa, North Vietnam, at altitudes of 1850 and 2010 m. They were sifted from leaf litter and roots in secondary deciduous forests. The type locality is illustrated in Fig. 62. An association with ants was not observed.</p></div>	https://treatment.plazi.org/id/03A2856AFF9EA3024B99FA94E136FB28	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Assing, Volker;V, North	Assing, Volker, V, North (2015): On the Lomechusini fauna of Vietnam (Coleoptera: Staphylinidae: Aleocharinae). Linzer biologische Beiträge 47 (2): 1257-1282, DOI: 10.5281/zenodo.5282217
