taxonID	type	description	language	source
03A387F2FFCFFFC2D2E7FDFBFD715D4D.taxon	description	(Figs 7, 12)	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFCFFFC2D2E7FDFBFD715D4D.taxon	diagnosis	Diagnosis. An Entomoculia Croissandeau, 1891 ascribable to the nominal subgenus for its robust build, the presence of chitinized thickenings on the male genital segment and for the characters of the male and female genitalia; well-differentiated from its congeners by the shape of the aedeagus and female copulatory plates.	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFCFFFC2D2E7FDFBFD715D4D.taxon	materials_examined	Type locality. SW Sardinia, Medio Campidano prov., Villacidro, Rio Bidda Scema. Type series. Holotype ♂: “ SW Sardinia, Villacidro (Medio Campidano), Rio Bidda Scema, 330 m, 20. IX. 1986, leg. P. Leo ” (MSNG). Paratypes: Villacidro (Medio Campidano prov.) Rio Bidda Scema, 330 m, 20. IX. 1986, leg. P. Leo, 1 ♀ (CLF-PL). Villacidro (Medio Campidano prov.), Foresta Demaniale di Montimannu, 280 m, 8. V. 1986, leg. P. Leo, 2 ♀ (CLF-PL); 8. VI. 1986, leg. P. Leo, 2 ♀ (CLF-PL, CNBFVR); 8. XI. 1986, leg. P. Leo, 1 ♀ (CLF-PL); 13. X. 1991, leg. P. Leo, 1 ♂ (CLF-PL).	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFCFFFC2D2E7FDFBFD715D4D.taxon	description	Description. Total length 1.10 – 1.15 mm. Head subparallel-sided, with short but prominent frontal carinas; microreticulation distinct but superficial; punctuation weak and sparse. Pronotum about as long as wide, shiny, with indistinct microreticulation; punctuation as on head; median longitudinal dimples lacking. Elytra shiny with slight microsculpture and sparse punctuation. Aedeagus as in Fig. 7 with, in lateral view, a short, thin and slightly downcurved sternal lamina; left paramere small, curved. Female genital armature complex, structured as in Fig. 12. Observations. Four species of subgenus Entomoculia were known so far for Sardinia, three from the central-northern part and one from the south (cf. Coiffait 1972; Pace 1996): E. sassariensis Pace, 1978 (type locality: surroundings of Sassari (Sassari prov.); both sexes known), E. sardoa Coiffait, 1959 (type locality: Ozieri (Sassari prov.); known only from females), E. gardinii Pace, 1994 (type locality: surroundings of Cuglieri (Oristano prov.); known only from males), E. iknusa Coiffait, 1959 (type locality: San Sperate (Cagliari prov.); known only from females). The new species is easily distinguishable from the above by the conformation of the male and female genitalia, and shows no obvious affinities with any other species of the genus. Ecological notes. The specimens were collected by washing soil from mixed woods dominated by Quercus ilex. The sampling localities are situated on granitic (Rio Bidda Scema) and argilloschist (Montimannu) Palaeozoic soils. The following endogean beetles were collected in association with the new species: Bathysciola damryi (Abeille de Perrin, 1881) (Leiodidae), Octavius sardous Coiffait, 1965 (Staphylinidae), Langelandia reitteri Belon, 1882 (Zopheridae), Torneuma sp. (Curculionidae), Alaocyba carinulata Perris, 1869, Raymondiellus sardous sardous (Perris, 1869) (Raymondionymidae).	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFCFFFC2D2E7FDFBFD715D4D.taxon	etymology	Etymology. Named after the type locality.	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC9FFC2D2E7FD4BFF325803.taxon	description	(Figs 9, 13)	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC9FFC2D2E7FD4BFF325803.taxon	diagnosis	Diagnosis. An Entomoculia ascribable to the nominal subgenus for its robust build, the presence of chitinized thickenings on the male genital segment and for the characters of the male and female genitalia; welldifferentiated from its congeners by the shape of the aedeagus and female copulatory plates.	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC9FFC2D2E7FD4BFF325803.taxon	materials_examined	Type locality. SW Sardinia, Carbonia-Iglesias prov., Domusnovas, Gutturu di Monte Nieddu. Type series. Holotype ♂: “ SW Sardinia, Domusnovas (Carbonia-Iglesias), Gutturu di Monte Nieddu, 290 m, 25. X. 1985, leg. L. Fancello & P. Leo ” (MSNG). Paratypes: Domusnovas (Carbonia-Iglesias prov.), Gutturu di Monte Nieddu, 290 m, 22. V. 1985, leg. L. Fancello & P. Leo, 4 ♂♂ and 3 ♀ (CLF-PL); 18. IX. 1985, leg. L. Fancello & P. Leo, 4 ♂♂ and 1 ♀ (CLF-PL); 1. X. 1985, leg. L. Fancello & P. Leo, 2 ♂♂ and 1 ♀ (CLF-PL); 25. X. 1985, leg. L. Fancello & P. Leo, 6 ♂♂ and 5 ♀ (1 ♂ and 1 ♀ CCH, 4 ♂♂ and 2 ♀ CLF-PL, 1 ♂ and 1 ♀ CNBFVR, 1 ♀ MSNG); 10. I. 1986, leg. L. Fancello & P. Leo, 1 ♀ (CLF-PL); 18. XII. 1989, leg. L. Fancello & P. Leo, 3 ♂♂ and 3 ♀ (CLF-PL). Domusnovas (Carbonia-Iglesias prov.), Grotta [= cave] San Giovanni, ingresso nord [= northern entrance], 200 m, 28. III. 1987, leg. P. Leo, 1 ♀ (CLF- PL).	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC9FFC2D2E7FD4BFF325803.taxon	description	Description. Total length 1.00 – 1.10 mm. Head subparallel-sided, with prominant frontal carinas slightly diverging posteriorly; microreticulation obvious and punctuation strong and sparse. Pronotum about as long as wide, shiny, with faint microreticulation and superficial and sparse punctuation, much more sparse than on the head; median longitudinal dimples lacking. Elytra shiny with imperceptible microsculpture and scattered punctuation. Aedeagus as in Fig. 9, with sternal lamina, in lateral view, long and robust, tapering at apex; left paramere long, thin and flexuous. Female genital armature complex, structured as in Fig. 13. Observations. The new species is well-differentiated from the already-known ones and from those described in the present paper; some similarity in the structure of the female genital plates can be found in E. sassariensis from northern Sardinia, but they are easily separable by the shape of the aedeagus (cf. Pace 1996, fig. 87). Ecological notes. This species was collected in woodland in soil samples taken at the base of an old Quercus ilex and under stone, near a natural cave. Sampling localities are situated on Palaeozoic limestone soil. The following endogean beetles were collected in association with the new species: Mayetia sp., Leptotyphlus minator sp. nov., Octavius raymondi Saulcy, 1878 (Staphylinidae), Torneuma sp. (Curculionidae), Alaocyba carinulata, Raymondiellus sardous sardous (Raymondionymidae).	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC9FFC2D2E7FD4BFF325803.taxon	etymology	Etymology. Named after the vegetal coal production that took place in the type locality until the last century.	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFCAFFC1D2E7FF52FAFC5BE2.taxon	description	(Figs 8, 14)	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFCAFFC1D2E7FF52FAFC5BE2.taxon	diagnosis	Diagnosis. An Entomoculia ascribable to the nominal subgenus for its robust build, the presence of chitinized thickenings on the male genital segment and for the characters of the male and female genitalia; welldifferentiated from its congeners by the conformation of the aedeagus and female copulatory plates.	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFCAFFC1D2E7FF52FAFC5BE2.taxon	materials_examined	Type locality. SW Sardinia, Medio Campidano prov., Villacidro, historical city centre. Type series. Holotype ♂: “ SW Sardinia, Villacidro (Medio Campidano), paese [= in town], 260 m, 9. XII. 1989, leg. P. Leo ” (MSNG). Paratypes: Villacidro (Medio Campidano prov.), paese, 260 m, 10. XI. 1985, leg. P. Leo, 2 ♂♂ and 1 ♀ (CLF-PL); 1. I. 1986, leg. P. Leo, 4 ♂♂ and 5 ♀ (1 ♀ CCH, 3 ♂♂ and 3 ♀ CLF- PL, 1 ♂ CNBFVR, 1 ♀, MSNG); 27. IV. 1986, leg. P. Leo, 1 ♀ (CLF-PL); 15. III. 1987, leg. P. Leo, 1 ♀ (CLF- PL).	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFCAFFC1D2E7FF52FAFC5BE2.taxon	description	Description. Total length 1.10 – 1.15 mm. Head subparallel-sided, with prominent frontal carinas slightly diverging posteriorly; microreticulation obvious and punctuation strong and sparse. Pronotum about as long as wide, with obvious microreticulation a bit weaker than on head; punctuation strong and sparse, except for a narrow median line on disc which appears as shiny, free of punctuation and microreticulation; median longitudinal dimples present. Elytra shiny with imperceptible microsculpture and scattered punctuation. Aedeagus as in Fig. 8 with sternal lamina, in lateral view, short, robust and tapering; left paramere robust, bent at a right angle. Female genital armature complex, structured as in Fig. 14. Observations. Entomoculia shardana sp. nov. is clearly separable from all congeners by the genital characters of both sexes. Ecological notes. This species was collected in an abandoned orchard garden of the historical centre of the small town of Villacidro, in soil samples taken at the base of a Ficus carica. The sampling locality is situated on Palaeozoic granitic soil. The following endogean beetles were collected in association with the new species: Bathysciola damryi (Leiodidae), Eudesis aglena Reitter, 1882, Leptotyphlus villacidrinus sp. nov., L. minator sp. nov. (Staphylinidae).	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFCAFFC1D2E7FF52FAFC5BE2.taxon	etymology	Etymology. This new species is named after the mythical Shardana warriors from Nuragic times.	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFCAFFC0D2E7FB30FCAB5E35.taxon	description	(Figs 5, 10 – 11)	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFCAFFC0D2E7FB30FCAB5E35.taxon	diagnosis	Diagnosis. An Entomoculia ascribable to the nominal subgenus for its robust build, the presence of chitinized thickenings on the male genital segment and for the characters of the male and female genitalia; welldifferentiated from its congeners by the conformation of the aedeagus and female copulatory plates.	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFCAFFC0D2E7FB30FCAB5E35.taxon	materials_examined	Type locality. SW Sardinia, Medio Campidano prov., Villacidro, San Sisinnio. Type series. Holotype ♂: “ SW Sardinia, Villacidro (Medio Campidano), San Sisinnio, 250 m, 2. I. 1987, leg. P. Leo ” (MSNG). Paratypes: Villacidro (Medio Campidano prov.), San Sisinnio, 250 m, 8. V. 1986, leg. P. Leo, 3 ♂♂ and 1 ♀ (2 ♂♂ and 1 ♀ CLF-PL, 1 ♂ CNBFVR); 2. I. 1987, leg. P. Leo, 2 ♀ (CLF-PL); 15. III. 1987, leg. P. Leo, 1 ♀ (CLF-PL); 9. XII. 1989, leg. P. Leo, 1 ♂ and 1 ♀ (CLF-PL); 24. IV. 1990, leg. P. Leo, 1 ♂ (CLF-PL).	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFCAFFC0D2E7FB30FCAB5E35.taxon	description	Description. Total length 1.00 – 1.10 mm. Head shiny, with diverging edges, distinctly widening posteriorly, and with inconspicuous frontal carinas; microreticulation faint, punctuation strong and sparse. Pronotum about as long as wide, shiny, with imperceptible microreticulation; punctuation slightly thinner than on head and slightly sparser; median longitudinal dimples lacking. Elytra shiny with imperceptible microsculpture and scattered punctuation. Aedeagus as in Fig. 10, with sternal lamina strongly bending downward, short, robust and tapering at apex. Female genital armature complex, structured as in Fig. 11. Observations. Entomoculia melonii sp. nov. is clearly separable from all congeners by the genital characters of both sexes. Ecological notes. The type specimens were found in samples taken at the base of old olive trees. The sampling locality is situated on argilloschist Palaeozoic soil. No other endogean beetles were collected together with the new species.	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFCAFFC0D2E7FB30FCAB5E35.taxon	etymology	Etymology. We dedicate this species to our friend Carlo Meloni from Cagliari, an expert of the Sardinian insect fauna with whom we have shared countless excursions.	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFCBFFCFD2E7FE63FAD85FCB.taxon	description	(Figs 16, 18 – 20)	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFCBFFCFD2E7FE63FAD85FCB.taxon	diagnosis	Diagnosis. A Leptotyphlus Fauvel, 1874 of medium size, with rather poor punctuation and superficial microreticulation. Differs from its congeners by the characters of the copulatory organ, but is undoubtedly akin to Leptotyphlus s. str. species of the L. sardous Coiffait, 1955 a-group (sensu Pace 1996) and to some peninsular taxa of the L. lessinicus Pace, 1973 - group, belonging to subgenus Stigmotyphlus Coiffait, 1955 b.	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFCBFFCFD2E7FE63FAD85FCB.taxon	materials_examined	Type locality. SW Sardinia, Carbonia-Iglesias prov., Iglesias, Nebida. Type series. Holotype ♂: “ SW Sardinia, Iglesias (Carbonia-Iglesias), Nebida, 120 m, 25. I. 1987, leg. L. Fancello & P. Leo ” (MSNG). Paratypes: Carbonia (Carbonia-Iglesias prov.), Barbusi, 130 m, 11. XI. 1986, leg. L. Fancello & P. Leo, 2 ♂♂ and 2 ♀ (CLF-PL). Domusnovas (Carbonia-Iglesias prov.), Gutturu di Monte Nieddu, 290 m, 12. I. 1990, leg. L. Fancello & P. Leo, 1 ♂ and 1 ♀ (CLF-PL); 13. IV. 1990, leg. L. Fancello & P. Leo, 2 ♂♂ and 11 ♀ (1 ♀ CCH, 2 ♂♂ and 9 ♀ CLF-PL, 1 ♀ CNBFVR). Iglesias (Carbonia-Iglesias prov.), Nebida, 120 m, 25. I. 1987, leg. L. Fancello & P. Leo, 2 ♂♂ and 3 ♀ (2 ♂♂ and 2 ♀ CLF-PL, 1 ♀ MSNG).	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFCBFFCFD2E7FE63FAD85FCB.taxon	description	Description. Total length 1.10 – 1.20 mm. Body of pale testaceous colour, clearly shiny, with superficial microsculpture on all integuments. Head slightly wider than pronotum, with moderately rounded sides and only slightly outlined frontal carinas; punctuation poor; labrum (Fig. 18) with three weakly developed median teeth; mandibles (Fig. 19) bifid with robust retinaculum and premolar tooth; antennae relatively short, just longer than head. Pronotum slightly longer than wide, with two longitudinal superficial grooves, slightly converging anteriorly; punctuation minute and sparser than on head. Elytral punctuation imperceptible. Convexity of abdominal tergites normal. Aedeagus as in Fig. 16, lying on the left side of the abdomen, with a robust and arched dorsal copulatory piece; sternal lamina distally divided into two branches; tip of ventral branch in lateral view with a distinct downturned lobiform appendix. Female genital armature asymmetrical, structured as in Fig. 20. Observations. As hinted in the diagnosis, the phyletic relationships of the new species seem complex. This forced us to describe the new species and the following two without subgeneric assignment. Besides, the subdivision of Leptotyphlus is still debated: Herman (2001) considers all described subgenera simply as synonyms of Leptotyphlus, whereas Smetana (2004 a, 2004 b) continues to follow the classic arrangement of the genus proposed by Coiffait (1972) and Pace (1996). In our opinion the characters proposed by Coiffait (1972) and Pace (1996) sometimes cause the placement of clearly allied species within separate subgenera, like in the emblematic case of the new species described above. Leptotyphlus nardii sp. nov. should indeed be ascribed to subgenus Stigmotyphlus for the asymmetrical female genital armature and for the aedeagus with a large coil-shaped dorsal copulatory piece; however, from further examination it appears that L. nardii sp. nov. is clearly related to some peninsular species of Stigmotyphlus of the L. lessinicus - group (L. molisensis Pace, 1977 and L. campanus Pace, 1977, extremely similar with regard to all structures of the copulatory organ: cf. Pace 1977, figs 190 – 191), and to Sardo-Corsican species of Leptotyphlus s. str. of the L. sardous - group, also with a coil-shaped dorsal copulatory piece (cf. Pace 1996, figs 163 – 165). Ecological notes. The species was collected by washing soil in both Mediterranean maquis with Pistacia lentiscus dominance and Quercus ilex woodland. Collecting sites are situated on Palaeozoic limestone soils. The following endogean beetles were collected together with the new species: Batysciola damryi (Leiodidae), Langelandia reitteri (Zopheridae), Torneuma sp. (Curculionidae), Alaocyba carinulata, Raymondiellus sardous sardous (Raymondionymidae).	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFCBFFCFD2E7FE63FAD85FCB.taxon	etymology	Etymology. It is our pleasure to dedicate this species to our friend Gianluca Nardi (CNBFVR).	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC4FFCDD2E7FB55FC5D5DB6.taxon	description	(Figs 15, 21 – 23)	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC4FFCDD2E7FB55FC5D5DB6.taxon	diagnosis	Diagnosis. A Leptotyphlus of medium size, with weakly punctuated integuments and superficial microreticulation. Clearly differentiated from all Italian species known so far but undoubtedly related to the North-African species L. tuniseus Normand, 1910, from which it differs essentially in the genital structures.	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC4FFCDD2E7FB55FC5D5DB6.taxon	materials_examined	Type locality. SW Sardinia, Medio Campidano prov., Villacidro, historical city centre. Type series. Holotype ♂: “ SW Sardinia, Villacidro (Medio Campidano), paese [= in town], 260 m, 10. XI. 1985, leg. P. Leo ” (MSNG). Paratypes: Villacidro (Medio Campidano prov.), paese, 260 m, 10. XI. 1985, leg. P. Leo, 2 ♂♂ and 5 ♀ (CLF-PL); 1. I. 1986, leg. P. Leo, 5 ♂♂ and 12 ♀ (1 ♀ CCH, 5 ♂♂ and 9 ♀ CLF- PL, 1 ♀ CNBFVR, 1 ♀ MSNG); 16. III. 1986, leg. P. Leo, 2 ♂♂ and 7 ♀ (CLF-PL); 27. IV. 1986, leg. P. Leo, 3 ♀ (CLF-PL). Villacidro (Medio Campidano prov.), Sa Spendula, 220 m, 9. XII. 1989, leg. P. Leo, 1 ♀ (CLF- PL). Villacidro (Medio Campidano prov.), San Sisinnio, 250 m, 8. V. 1986, 2. I. 1987, leg. P. Leo, 3 ♀ (CLF- PL).	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC4FFCDD2E7FB55FC5D5DB6.taxon	description	Description. Total length 1.10 – 1.20 mm. Body of pale testaceous colour, rather shiny, with sparse microsculpture on all of its surface. Head just wider than pronotum, not markedly rounded at sides, with only slight frontal carinas; punctuation not strong; labrum (Fig. 21) with two symmetrical median teeth; mandibles (Fig. 22) bifid with robust retinaculum and premolar tooth; antennae relatively short, just longer than head. Pronotum about as long as wide, with two superficial longitudinal grooves, slightly converging anteriorly; punctuation even more minute and sparse than on head. Elytral punctuation almost imperceptible. Abdomen with regularly convex tergites. Aedeagus as in Fig. 15, lying to the left side of the abdomen, with a robust and arched dorsal copulatory piece; sternal lamina distally divided into two branches: ventral branch, in lateral view, simple, short and not markedly sinuous; upper branch long, very thin and upturned. Female genital armature asymmetrical, structured as in Fig. 23. Observations. As already mentioned in the diagnosis, the only species close to Leptotyphlus villacidrinus sp. nov. is L. tuniseus from northern Tunisia, which was ascribed by Coiffait (1972) to subgenus Stigmotyphlus. In particular, similarities can be found in the large and strongly arched dorsal copulatory piece, and in the shape and chaetotaxy of the left paramere. Differences between these two species lie in the shape and chaetotaxy of the right paramere and the structure of the other copulatory pieces (cf. Coiffait 1972, fig. 197). Ecological notes. Most specimens of the type series were collected in an abandoned orchard garden of the historical city centre of Villacidro, in soil samples taken at the base of a Ficus carica tree; other specimens were found in woody areas dominated by Quercus ilex. The collecting localities are situated on granitic (Villacidro city centre and Sa Spendula) and argilloschist (San Sisinnio) Palaeozoic soils. The following endogean beetles were found in association with the new species: Bathysciola damryi (Leiodidae), Eudesis aglena, Trimium amplipenne Reitter, 1908, Entomoculia shardana sp. nov., Leptotyphlus minator sp. nov. (Staphylinidae), Torneuma sp. (Curculionidae), Alaocyba carinulata, Raymondiellus sardous sardous (Raymondionymidae).	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC4FFCDD2E7FB55FC5D5DB6.taxon	etymology	Etymology. This new species is named after the type locality.	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC6FFCCD2E7FCECFE395EE3.taxon	description	(Figs 17, 24 – 26)	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC6FFCCD2E7FCECFE395EE3.taxon	diagnosis	Diagnosis. A Leptotyphlus of large size, with robust punctuation and microreticulation, close to L. solarii Coiffait, 1957, but strongly differing from the latter in the shape of the male and female genitalia.	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC6FFCCD2E7FCECFE395EE3.taxon	materials_examined	Type locality. SW Sardinia, Carbonia-Iglesias prov., Iglesias, Nebida. Type series. Holotype ♂: “ SW Sardinia, Iglesias (Carbonia-Iglesias), Nebida, 25. I. 1987, 120 m, leg. L. Fancello & P. Leo ” (MSNG). Paratypes: Carbonia (Carbonia-Iglesias prov.), Monte Sirai, 120 m, 22. XII. 1990, leg. L. Fancello & P. Leo, 1 ♂ (CLF-PL). Domusnovas (Carbonia-Iglesias prov.), Gutturu di Monte Nieddu, 290 m, 1. X. 1985, leg. L. Fancello & P. Leo, 2 ♂♂ and 2 ♀ (1 ♂ and 2 ♀ CLF-PL, 1 ♂ CNBFVR). Fluminimaggiore (Carbonia-Iglesias prov.), near grotta Su Mannau, 200 m, 19. III. 1989, leg. P. Leo, 1 ♂ (CLF-PL). Narcao (Carbonia-Iglesias prov.), 310 m, 5. I. 1990, leg. L. Fancello & P. Leo, 1 ♀ (CLF- PL). Iglesias (Carbonia-Iglesias), Nebida, 25. I. 1987, 120 m, leg. L. Fancello & P. Leo 1 ♂ (CLF-PL). Villacidro (Medio Campidano prov.), paese [= in town], 260 m, 27. IV. 1986, leg. P. Leo, 1 ♀ (CLF-PL). Villacidro (Medio Campidano prov.), San Sisinnio, 250 m, 15. III. 1987, leg. P. Leo, 1 ♀ (CLF-PL).	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC6FFCCD2E7FCECFE395EE3.taxon	description	Description. Total length 1.35 – 1.50 mm. Body robust, of reddish colour, slightly shiny, with obvious microreticulation throughout. Head large, wider than pronotum distally, slightly rounded at sides, with frontal carinas slightly diverging posteriorly; punctuation strong and deep; labrum bifid (Fig. 25); mandibles (Fig. 26) with a few indistinct small teeth between retinaculum and premolar tooth; antennae somewhat elongated, much longer than head. Pronotum about as wide as long, with two longitudinal grooves from base of pronotum to anterior ¼; punctuation large, as on head. Elytra with sparse, robust punctuation, less strong than on head and pronotum. First five abdominal segments slightly flattened on disc. Aedeagus as in Fig. 17, lying to the left of abdomen and with a strongly developed basal bulb carrying an obvious chitinized nodule on upper surface; sternal lamina in lateral view strongly arched upwards. Female genital armature weakly chitinized, structured as in Fig. 24. Observations. Leptotyphlus solarii (described from Aritzo, central Sardinia) is the only species that show some affinity with Leptotyphlus minator sp. nov.: this species, ascribed by Coiffait (1972) and Pace (1996) to subgenus Subhesperotyphlus Coiffait, 1972, can be easily distinguished by its smaller size and the shape of the male copulatory organ and female genital armature (cf. Coiffait 1972: fig. 172; Pace 1996, fig. 176). Ecological notes. The new species, relatively widespread but always rare in the study area, was found in soil samples taken from a wide range of habitats: Mediterranean maquis with Pistacia lentiscus dominance, Quercus ilex woodland, degraded areas at the base of Ficus carica and Olea europaea trees. The sampling localities are situated on a range of soil types: Palaeozoic granites, limestones and argilloschists, and Cenozoic basaltic trachytes. The following endogean beetles were collected in association with the new species: Bathysciola damryi (Leiodidae), Eudesis aglena, Mayetia sp., Entomoculia carbonaria sp. nov., E. shardana sp. nov., Leptotyphlus villacidrinus sp. nov. (Staphylinidae), Torneuma sp. (Curculionidae), Alaocyba carinulata, Raymondiellus sardous sardous (Raymondionymidae).	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC6FFCCD2E7FCECFE395EE3.taxon	etymology	Etymology. Named after the mining activities which took place across the centuries in the area of occurrence of the new species.	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC7FFCCD2E7FD43FBC75C8B.taxon	description	Literature data. Iglesias (Carbonia-Iglesias prov.), Mamenga, 1. III. 2006, leg. L. Fancello, 1 ex, 1 rs (Magrini et al. 2006). Iglesias (Carbonia-Iglesias prov.), Parco Monte Marganai, loc. Mamenga (Magrini & Fancello 2007).	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC7FFCCD2E7FD43FBC75C8B.taxon	distribution	Distribution. Species endemic of the Iglesiente region, known only from the type locality (Iglesias, Passo Genna Bogai) and the above-cited site. Notes. Collected with the Berlese method in a Quercus ilex woodland.	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC7FFCCD2E7FB89FBC75ACE.taxon	description	Literature data. Iglesias (Carbonia-Iglesias prov.), Mamenga, 430 m, 28. XI. 2006, 20. I. 2007, 2. III. 2007, leg. L. Fancello, 11 ex (Magrini & Fancello 2007).	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC7FFCCD2E7FB89FBC75ACE.taxon	distribution	Distribution. Species endemic of the Iglesiente region; the nominal subspecies and ssp. leoi are only known from the respective type localities: Fluminimaggiore (Carbonia-Iglesias prov.), Sant’Angelo (Magrini 2003) and the above-cited site. Notes. Collected with the Berlese method in a Quercus ilex woodland.	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC7FFCCD2E7F9D4FAC958B3.taxon	description	(Fig. 4) Leo et al. 2005: 181 Literature data. Villacidro (Medio Campidano prov.), San Sisinnio, 250 m (type locality), 27. IV. 1986, 15. III. 1987, 24. IV. 1990, 2 ex, 2 rs (Leo et al. 2005). Unpublished data. Villacidro (Medio Campidano prov.), San Sisinnio, 250 m, 4. IV. 2007, leg. L. Fancello & P. Leo, 1 ex (CLF-PL).	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC7FFCCD2E7F9D4FAC958B3.taxon	distribution	Distribution. Sardinian endemic, known only from the above localities. Notes. Due to a typing error in the species description, it was not clearly stressed that the holotype is of male sex. Collected by hand and with the Berlese method in a Quercus ilex - dominated woodland.	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC0FFCBD2E7FF24FB8A5CD1.taxon	description	Literature data. Domusnovas (Carbonia-Iglesias prov.), 27. X. 1985, leg. L. Fancello & P. Leo, 4 ex. Villacidro (Medio Campidano prov.), 10. XI. 1985, 1. I. 1986, 16. III. 1986, 27. IV. 1986, 20. IX. 1986, leg. P. Leo, 22 ex. Villacidro (Medio Campidano prov.), Foresta di Montimannu, 8. XI. 1986, leg. L. Fancello & P. Leo, 14 ex (Giachino 1990; Zoia & Latella 2006). Iglesias (Carbonia-Iglesias prov.), Mamenga (Magrini et al. 2006; Fancello & Magrini 2007; Casale et al. 2009). Unpublished data. Domusnovas (Carbonia-Iglesias prov.), Gutturu di Monte Nieddu, 290 m, 10. I. 1986, leg. L. Fancello & P. Leo, 5 ex (CLF-PL). Iglesias (Carbonia-Iglesias prov.), Mamenga, 430 m, 1. III. 2006, leg. L. Fancello, 6 ex (CLF-PL). Villacidro (Medio Campidano prov.), paese [= in town], 260 m, 15. III. 1987, 9. XII. 1989, leg. P. Leo, 6 ex (CLF-PL). Villacidro (Medio Campidano prov.), Foresta Demaniale di Montimannu, 280 m, 8. V. 1986, 8. VI. 1986, leg. P. Leo, 11 ex (CLF-PL). Villacidro (Medio Campidano prov.), San Sisinnio, 250 m, 27. IV. 1986, 8. V. 1986, leg. P. Leo, 8 ex (CLF-PL).	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC0FFCBD2E7FF24FB8A5CD1.taxon	distribution	Distribution. Sardinian endemic, widespread throughout the island (Giachino 1990, Zoia & Latella 2006). Notes. Collected in an abandoned orchard garden of the small town of Villacidro and in wooded areas and Mediterranean maquis with the Berlese method, direct collecting and buried bait.	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC0FFCBD2E7FB89FCA8597D.taxon	description	Unpublished data. Domusnovas (Carbonia-Iglesias prov.), Sa Duchessa, 350 m, 10. I. 1986, leg. L. Fancello & P. Leo, 1 ex (CLF-PL). Iglesias (Carbonia-Iglesias prov.), Mamenga, 430 m, 7. IV. 2000, leg. L. Fancello, 1 ex (CLF-PL). Villacidro (Medio Campidano prov.), paese, 260 m, 1. I. 1986, leg. P. Leo, 3 ex (CLF-PL).	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC0FFCBD2E7FB89FCA8597D.taxon	distribution	Distribution. Sardinian endemic, known so far only from few localities in western Sardinia: Gonnesa, Fluminimaggiore, Asuni and Campeda (Binaghi 1948). Its occurence in Corsica, still mentioned in some recent works (Sánchez-Terròn 2004, with doubt; Vít 2004) should be refused (cf. Binaghi 1948). Notes. Collected with the Berlese method in various habitats: Quercus ilex woodland, Mediterranean maquis and in an orchard garden at the base of a Ficus carica.	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC0FFCBD2E7F99BFC0A58B3.taxon	description	Unpublished data. Domusnovas (Carbonia-Iglesias prov.), Gutturu di Monte Nieddu, 290 m, 22. V. 1985, 13. IV. 1990, leg. L. Fancello & P. Leo, 5 ex (CLF-PL). Iglesias (Carbonia-Iglesias prov.), Mamenga, 430 m, 7. IV. 2000, leg. L. Fancello, 1 ex (CLF-PL).	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC0FFCBD2E7F99BFC0A58B3.taxon	distribution	Distribution. Species seemingly endemic of SW Sardinia, previously known only from the type locality: Fluminimaggiore (Carbonia-Iglesias prov.) (Binaghi 1948). Notes. Collected with the Berlese method in a Quercus ilex wood.	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC1FFCAD2E7FDF8FBC75CFB.taxon	description	Literature data. Villacidro (Medio Campidano prov.) (Poggi 1992; Poggi & Sabella 2006). Unpublished data. Villacidro (Medio Campidano prov.), San Sisinnio, 250 m, 8. V. 1986, 24. IV. 1990, 7. V. 1991, leg. P. Leo, 10 ex (CLF-PL).	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC1FFCAD2E7FDF8FBC75CFB.taxon	distribution	Distribution. Sardinian endemic, widespread throughout Sardinia (Poggi 1992, Poggi & Sabella 2006). Notes. Collected with the Berlese method in a Quercus ilex woodland.	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC1FFCAD2E7FC19FA935AC2.taxon	description	Literature data. Domusnovas (Carbonia-Iglesias prov.) (Poggi 1992). Villacidro (Medio Campidano prov.) (Poggi 1992). Unpublished data. Domusnovas (Carbonia-Iglesias prov.), Gutturu di Monte Nieddu, 290 m, 10. I. 1986, leg. L. Fancello & P. Leo, 1 ex (CLF-PL). Domusnovas (Carbonia-Iglesias prov.), Valle di Oridda, 570 m, 12. II. 1992, leg. L. Fancello & P. Leo, 1 ex (CLF-PL). Iglesias (Carbonia-Iglesias prov.), Mamenga, 430 m, 1. III. 2006, leg. L. Fancello, 1 ex (CLF-PL).	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC1FFCAD2E7FC19FA935AC2.taxon	distribution	Distribution. Sardo-Corsican endemic, widespread in Sardinia (Poggi 1992; Löbl & Besuchet 2004; Poggi & Sabella 2006). Notes. Collected with the Berlese method and by hand in wooded and Mediterranean maquis areas.	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC2FFC9D2E7FBECFCAC5BCA.taxon	description	(Figs 9, 13) Species known only from the study area and described above.	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC2FFC9D2E7FAC8FCAC5AEE.taxon	description	(Figs 5, 10 – 11) Species known only from the study area and described above.	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC2FFC9D2E7FA34FCAC5902.taxon	description	(Figs 8, 14) Species known only from the study area and described above.	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC2FFC9D2E7F910FCAC5826.taxon	description	(Figs 7, 12) Species known only from the study area and described above.	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC2FFC9D2E7FE2AFB1B5CB6.taxon	description	Unpublished data. Domusnovas (Carbonia-Iglesias prov.), Gutturu di Monte Nieddu, 290 – 320 m, 25. X. 1985, 10. I. 1986, leg. L. Fancello & P. Leo, 5 ex (CLF-PL). Domusnovas (Carbonia-Iglesias prov.), Valle di Oridda, 570 m, 12. II. 1992, leg. L. Fancello & P. Leo, 1 ex (CLF-PL). Villacidro (Medio Campidano prov.), Foresta Demaniale di Montimannu, 280 m, 8. V. 1986, leg. P. Leo, 1 ex (CLF-PL). Villacidro (Medio Campidano prov.), San Sisinnio, 250 m, 9. XII. 1989, leg. P. Leo, 1 ex (CLF-PL).	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC2FFC9D2E7FE2AFB1B5CB6.taxon	distribution	Distribution. Sardinian endemic (Löbl & Besuchet 2004) recorded only from three localities of centralsouthern Sardinia: Oristano (Oristano prov.), Aritzo (Nuoro prov.) and Fluminimaggiore (Carbonia-Iglesias prov.) (Orousset 1980). Notes. Collected with the Berlese method in wooded areas with Quercus ilex dominance.	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC3FFC8D2E7FF52FBB95E5B.taxon	description	(Figs 17, 24 – 26) Species known only from the study area and adjacent localities, and described above.	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC3FFC8D2E7FEB9FBB95D7F.taxon	description	(Figs 16, 18 – 20) Species known only from the study area and adjacent localities, and described above.	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC3FFC8D2E7FD9AFCAC5D93.taxon	description	(Figs 15, 21 – 23) Species known only from the study area and described above.	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC3FFC8D2E7FC81FB4C5BD6.taxon	description	Unpublished data. Domusnovas (Carbonia-Iglesias prov.), Gutturu di Monte Nieddu, 310 m, 10. I. 1986, 280 m, leg. L. Fancello & P. Leo, 2 ex (CLF-PL).	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC3FFC8D2E7FC81FB4C5BD6.taxon	distribution	Distribution. Corsica?, Sardinia, Algeria and Tunisia (Smetana 2004 a, 2004 c); in Sardinia was previously known only from the type locality in the north of the Island: Bonnari (Sassari prov.) (Coiffait 1984). Notes. Collected with the Berlese method in a wood, at the base of a Quercus ilex.	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC3FFC8D2E7FACCFB415937.taxon	description	Unpublished data. Villacidro (Medio Campidano prov.), Foresta Demaniale di Montimannu, 280 m, 8. V. 1986, 8. XI. 1986, leg. P. Leo, 9 ex (CLF-PL).	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC3FFC8D2E7FACCFB415937.taxon	distribution	Distribution. Sardinian endemic (Smetana 2004 a, 2004 c), previously known only from the type locality in central Sardinia: Aritzo (Nuoro prov.) (Coiffait 1984). Notes. Collected with the Berlese method in a Quercus ilex - dominated woodland.	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC3FFD7D2E7F934FE625EE3.taxon	description	Unpublished data. Domusnovas (Carbonia-Iglesias prov.), Gutturu di Monte Nieddu, 290 – 320 m, 22. V. 1985, 12. I. 1990, 13. IV. 1990, leg. L. Fancello & P. Leo, 5 ex (CLF-PL). Domusnovas (Carbonia-Iglesias prov.), Valle di Oridda, 570 m, 12. II. 1992, 20. III. 1992, leg. L. Fancello & P. Leo, 4 ex (CLF-PL). Iglesias (Carbonia- Iglesias prov.), Mamenga, 430 m, 14. I. 2001, leg. L. Fancello, 1 ex (CLF-PL). Villacidro (Medio Campidano prov.), Rio Bidda Scema, 330 m, 20. IX. 1986, leg. P. Leo, 6 ex (CLF-PL). Villacidro (Medio Campidano prov.), San Sisinnio, 250 m, 7. V. 1991, leg. P. Leo, 1 ex (CLF-PL).	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFC3FFD7D2E7F934FE625EE3.taxon	distribution	Distribution. A West-Mediterranean species (Ś lipi ṅski & Schuh 2008) overlooked in the Italian checklist (Angelini et al. 1995) but already quoted from Sardinia by Dajoz (1977, as Langelandia reitteri reitteri). Notes. Collected by hand and with the Berlese method, in open habitats and in woodland and Mediterranean maquis.	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFDCFFD7D2E7FE31FBC75CD5.taxon	description	Literature data. Domusnovas (Carbonia-Iglesias prov.), Gutturu di Monte Nieddu, 290 m (type locality), 18. IX. 1985, 1. X. 1985, leg. L. Fancello & P. Leo, 3 ex (Fancello & Leo 1991). Unpublished data. Iglesias (Carbonia-Iglesias prov.), Mamenga, 430 m, 20. XI. 2005, leg. L. Fancello, 1 ex (CLF-PL).	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFDCFFD7D2E7FE31FBC75CD5.taxon	distribution	Distribution. Endemic to SW Sardinia, previously known only from the type locality and Villamassargia (Carbonia-Iglesias prov.), Monte Santu Miai (Magrini et al. 2003); known to us also from a neighbouring locality: Iglesias (Carbonia-Iglesias prov.), Masua, 150 m, 13. IV. 2004, leg. L. Fancello, 2 ex (CLF-PL). Notes. Collected with the Berlese method in a Quercus ilex woodland.	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFDCFFD7D2E7FB8AFD9059D3.taxon	description	(Fig. 6) Literature data. Domusnovas (Carbonia-Iglesias prov.), Gutturu di Monte Nieddu, 290 m (Fancello & Leo 1991). Iglesias (Carbonia-Iglesias prov.), Mamenga (Magrini et al. 2006; Magrini & Fancello 2007). Unpublished data. Domusnovas (Carbonia-Iglesias prov.), Gutturu di Monte Nieddu, 290 - 330 m, 18. IX. 1985, 1. X. 1985, 25. X. 1985, 10. I. 1986, 13. IV. 1990, 12. III. 2007, leg. L. Fancello & P. Leo, 20 ex (CLF- PL). Domusnovas (Carbonia-Iglesias prov.), Valle di Oridda, 570 m, 12. II. 1992, leg. L. Fancello & P. Leo, 3 ex (CLF-PL). Iglesias (Carbonia-Iglesias prov.), Mamenga, 430 m, 1. III. 2006, leg. L. Fancello, 1 ex (CLF- PL). Villacidro (Medio Campidano prov.), Foresta Demaniale di Montimannu, 280 m, 8. V. 1986, leg. P. Leo, 1 ex (CLF-PL). Villacidro (Medio Campidano prov.), San Sisinnio, 250 m, 27. IV. 1986, 8. V. 1986, 15. III. 1987, 9. XII. 1989, 7. V. 1991, 4. IV. 2007, leg. P. Leo, 14 ex (CLF-PL).	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFDCFFD7D2E7FB8AFD9059D3.taxon	distribution	Distribution. Probably a Sardinian endemic. Notes. Collected by hand and with the Berlese method, in open habitats and in woodland and Mediterranean maquis. Pending a full revision of the genus (cf. Colonnelli & Osella 2009) it was impossible to identify these specimens to species.	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFDCFFD6D2E7F888FF345DFB.taxon	description	Unpublished data. Domusnovas (Carbonia-Iglesias prov.), Gutturu di Monte Nieddu, 290 - 330 m, 18. IX. 1985, 1. X. 1985, 25. X. 1985, 10. I. 1986, 28. III. 1986, 12. I. 1990, 13. IV. 1990, leg. L. Fancello & P. Leo, 42 ex (CLF-PL). Domusnovas (Carbonia-Iglesias prov.), Valle di Oridda, 570 m, 12. II. 1992, 20. III. 1992, leg. L. Fancello & P. Leo, 4 ex (CLF-PL). Iglesias (Carbonia-Iglesias prov.), Case Marganai, 725 m, 10. IV. 2003, leg. L. Fancello, 3 ex (CLF-PL). Iglesias (Carbonia-Iglesias prov.), Mamenga, 430 m, 20. XI. 2005, leg. L. Fancello, 8 ex (CLF-PL). Villacidro (Medio Campidano prov.), Foresta Demaniale di Montimannu, 280 m, 8. V. 1986, 8. VI. 1986, 8. XI. 1986, 13. X. 1991, leg. P. Leo, 10 ex (CLF-PL). Villacidro (Medio Campidano prov.), San Sisinnio, 250 m, 8. V. 1986, 15. III. 1987, 24. IV. 1990, 7. V. 1991, leg. P. Leo, 11 ex (CLF-PL).	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFDCFFD6D2E7F888FF345DFB.taxon	distribution	Distribution. Sardinian endemic (Osella 1977). Notes. Collected by hand and with the Berlese method in wooded areas (Fig. 3) and Mediterranean maquis.	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFDDFFD6D2E7FD19FF345A05.taxon	description	Literature data. Domusnovas (Carbonia-Iglesias prov.), Gutturu di Monte Nieddu, 290 m, (Fancello & Leo 1991). Iglesias (Carbonia-Iglesias prov.), Mamenga (Magrini et al. 2006; Magrini & Fancello 2007). Unpublished data. Domusnovas (Carbonia-Iglesias prov.), Gutturu di Monte Nieddu, 290 m, 18. IX. 1985, 1. X. 1985, 25. X. 1985, 10. I. 1986, 12. I. 1990, 13. IV. 1990, leg. L. Fancello & P. Leo, 28 ex (CLF- PL). Domusnovas (Carbonia-Iglesias prov.), Valle di Oridda, 570 m, 12. II. 1992, 20. III. 1992, leg. L. Fancello & P. Leo, 3 ex (CLF-PL). Iglesias (Carbonia-Iglesias prov.), Case Marganai, 725 m, 10. IV. 2003, leg. L. Fancello, 2 ex (CLF-PL). Iglesias (Carbonia-Iglesias prov.), Mamenga, 430 m, 20. XI. 2005, leg. L. Fancello, 2 ex (CLF-PL). Villacidro (Medio Campidano prov.), Foresta Demaniale di Montimannu, 280 m, 8. V. 1986, 8. VI. 1986, 8. XI. 1986, leg. P. Leo, 4 ex (CLF-PL). Villacidro (Medio Campidano prov.), San Sisinnio, 250 m, 27. IV. 1986, 8. V. 1986, 15. III. 1987, 7. V. 1991, leg. P. Leo, 6 ex (CLF-PL).	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
03A387F2FFDDFFD6D2E7FD19FF345A05.taxon	distribution	Distribution. Endemic to SW Sardinia, with a subspecies on Isola di San Pietro (Carbonia-Iglesias prov.) (Osella 1977). Notes. Collected by hand and with the Berlese method in wooded areas (Fig. 3) and Mediterranean maquis.	en	Fancello, Luca, Hernando, Carles, Leo, Piero (2009): The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *. Zootaxa 2318: 317-338
