taxonID	type	description	language	source
03A3D2261F04E73AFF42A21DD88AFCD1.taxon	materials_examined	Material. Sarpang, 8.3 km NW Gelephu, 26.921 ° N, 90.424 ° E, 580 m, 23 – 27.08.2017, Methyl Eugenol, 1 Ơ, FFP 17 BT 077 (CSCA 19 L 046) (BHNPPC). M a l e l u r e. Methyl eugenol. Host plants. Azadirachta indica A. Juss. (Meliaceae) (Drew & Romig, 2013). Distribution. Bhutan, Cambodia, India, Nepal, Malaysia (peninsular), Thailand, Vietnam (David et al., 2017; Drew & Romig, 2013; Hardy, 1973; Leblanc et al., 2016, 2019 a; Norrbom et al., 1999).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F04E73AFF42A34DDD4EFAA6.taxon	description	M a l e l u r e. Cue-lure. Host plants. Xylosma brachystachys Craib (Salicaceae) (Drew & Romig, 2013). Distribution. Bangladesh, Bhutan, Cambodia, China, India (Andaman Island), Laos, Thailand, Vietnam (Drew & Romig, 2013; Leblanc et al., 2014, 2016, 2019 b).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F04E73AFF42A560DA79FA01.taxon	description	D i s t r i b u t i o n. Bhutan, India (Drew & Romig, 2013).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F04E73AFF42A61DDB6EF8CF.taxon	materials_examined	Material. Sarpang, 12.3 km NE Sarpang, 26.953 ° N, 90.346 ° E, 890 m; 24 – 27.08.2017, Malaise trap, 1 Ơ, FFP 17 BT 090 (CSCA 18 L 185); 13 km NW Gelephu, 26.968 ° N, 90.559 ° E, 1200 m, 25 – 27.08.2017, Zingerone, 1 Ơ, FFP 17 BT 099 (CSCA 18 L 216) (BHNPPC). M a l e l u r e. Zingerone. D i s t r i b u t i o n. Bhutan (first record), Vietnam (Leblanc et al., 2018). Note. Transferred from the subgenus Asiadacus Perkins to the subgenus Tetradacus Miyake by Hancock & Drew (2019).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F05E73BFF4AA0A2DB0FFE33.taxon	description	M a l e l u r e. Methyl eugenol. H o s t p l a n t s. A polyphagous pest of fruits, infests at least 91 host plants in 36 families (Allwood et al., 1999; Liquido et al., 2022 a). D i s t r i b u t i o n. Bangladesh, Bhutan (Drew et al., 2007), Cambodia, China (Sichuan, Yunnan), India, Malaysia (peninsular), Myanmar, Nepal, Pakistan, Sri Lanka, Thailand, Vietnam (Drew & Romig, 2013).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F05E73CFF4AA1EBDAFBFB2E.taxon	materials_examined	Material. Wangdue Phodrang, Bajo, ARDC orchards, 27.490 ° N, 89.900 ° E, 1230 m, hand collecting, 15.08.2017, 1 Ơ, 1 Ơ, FFP 17 BT 114 (C. Borkent & Hauser leg.); 15 – 21.08.2017, yeast, 1 Ơ, 1 ♀, FFP 17 BT 012 (CSCA 18 L 158); 1 Ơ, 5 ♀, FFP 17 BT 017; Methyl Eugenol, 147 Ơ, FFP 17 BT 009; Multilure, 11 Ơ, 12 Ơ, FFP 17 BT 013 (BHNPPC); 20.08.2017, Methyl Eugenol, 1 Ơ, FFP 17 BT 136 (CSCA 18 L 228) (C. J. Borkent leg.); 21 – 28.08.2017, Methyl Eugenol, 38 Ơ, FFP 17 BT 014; Multilure, 6 Ơ, 6 ♀, FFP 17 BT 018 (BHNPPC); 5.8 km S Alla, Royal Orchard, 27.298 ° N, 89.968 ° E, 800 m, 18 – 28.08.2017, Methyl Eugenol, 114 Ơ, 1 ♀, FFP 17 BT 056; 18.08.2018, spilled Methyl Eugenol, 16 Ơ, FFP 17 TB 118 (BHNPPC); Sarpang, 7.7 km NW Gelephu, mixed veg farm, 26.910 ° N, 90.422 ° E, 380 m, 23 – 27.08.2017, Zingerone, 2 ♀, FFP 17 BT 087 (CSCA 18 L 164); Methyl Eugenol, 2 Ơ, FF 17 BT 085 (CSCA 18 L 165); 2 Ơ (CSCA 18 L 210); 8.3 km NW Gelephu, Multilure, 26.921 ° N, 90.424 ° E, 580 m, 23 – 27.08.2017, 1 Ơ, FFP 17 BT 082 (CSCA 18 L 172); methyl eugenol, 6 Ơ, FFP 17 BT 077 (CSCA 19 L 046); 6.8 km NW Gelephu, ARDC orchards, 26.91 ° N, 90.433 ° E, 400 m, 23 – 27.08.2017, terpinyl acetate, 1 ♀, FP- P 17 BT 119 (CSCA 18 L 212); methyl isoeugenol, 3 Ơ, 3 ♀, FFP 17 BT 139; methyl eugenol, 17 Ơ, FFP 17 BT 138; 1 f, FFP 17 BT 069; Punakha, 13 km NW Punakha, Methyl Eugenol, 27.667 ° N, 89.775 ° E, 1400 m, 16 – 20.08.2017, 2 Ơ, FFP 17 BT 030 (CSCA 18 L 209) (BHNPPC); Thimphu, NPPC fields, Methyl Eugenol, 27.441 ° N, 89.664 ° E, 2300 m, 14 – 29.08.2017, 67 Ơ, FFP 17 BT 003 (C. Borkent & Hauser leg.). M a l e l u r e. Methyl eugenol, zingerone. Host plants. Polyphagous pest of fruits of at least 500 host plants (Allwood et al., 1999; Liquido et al., 2022 b). D i s t r i b u t i o n. Bangladesh, Bhutan (Drew et al., 2007; Drew & Romig, 2013), Brunei, Cambodia, s. China (Sichuan, Guizhou, Hunan, Fujian, Guangxi, Guangdong, Hong Kong, Hainan, Yunnan), Christmas I., India (Andaman & Nicobar Islands, Andhra Pradesh, Assam, Bihar, Delhi, Gujarat, Haryana, Himachal Pradesh, Jammu and Kashmir, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Manipur, Odisha, Punjab, Rajasthan, Sikkim, Tamil Nadu, Uttar Pradesh, West Bengal), Indonesia (Bali, East & West Nusa Tenggara, Java, Kalimantan, Maluku, Sulawesi, Sumatra), Laos, Malaysia (Peninsular, Sabah), Myanmar, Nepal, Philippines (Cebu, Luzon, Mindanao, Negros, Panay), Singapore, Sri Lanka, Taiwan, Thailand, Vietnam; introduced Australia (Torres Strait Islands), Guam, Hawaii, Indonesia (West Papua), Nauru, Palau, Papua New Guinea (mainland), Angola, Benin, Burkina Faso, Burundi, Cameroon, Central African Republic, Chad, Comoros, Congo (Rep. of), Congo (Dem. Rep.), Equatorial Guinea, Eritrea, Ethiopia, Gabon, Gambia, Ghana, Guinea, Guinea-Bissau, Ivory Coast, Kenya, Liberia, Mali, Mauritania, Mozambique, Namibia, Niger, Nigeria, Oman, Senegal, Sierra Leone, Sudan, Tanzania, Togo, Uganda, Zambia; eradicated from Australia, Japan (Ryukyu Islands) & N. Marianas; brief 1996 outbreak in Mauritius (White, 2006). Note. Bactrocera dorsalis is part of a species complex whose composition and limits continue to be disputed among taxonomists. Bactrocera invadens Drew, Tsuruta & White, 2005; B. papayae Drew & Hancock, 1994; and B. philippinensis Drew & Hancock, 1994 were considered to be synonyms by Schutze et al., 2015), whereas Drew & Hancock (2022) recently resurrected B. invadens and B. papayae, based on morphological structures of the male genitalia. Maneesh et al. (2022) stated that B. invadens has been definitively recorded in the Himalayan region from Bhutan (Drew et al., 2007) and Nepal (Leblanc et al., 2019: as B. dorsalis). In this article, we use the name B. dorsalis because our material does not possess morphological characters given as diagnostic for B. invadens at Drew & Hancock 2022.	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F02E73CFF42A4EBDDAEFA3E.taxon	materials_examined	Material. Wangdue Phodrang, Bajo, ARDC orchards, 27.490 ° N, 89.900 ° E, 1230 m, 21 – 28.08.2017, Cue-lure, 1 Ơ, FFP 17 BT 015 (BHNPPC); 5.8 km S Alla, Royal Orchard, 27.298 ° N, 89.968 ° E, 800 m, 18.08.2017, Cue-lure, 3 Ơ FFP 17 BT 130 (BHNPPC); Hand collecting, 1 Ơ, FFP 17 TB 062 (CSCA 18 L 174) (C. J. Borkent leg.). M a l e l u r e. Cue-lure. D i s t r i b u t i o n. Bhutan, Malaysia (peninsular), Vietnam (Drew & Romig, 2013).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F02E73CFF42A61BDAD5F99A.taxon	description	D i s t r i b u t i o n. Bhutan, northern Vietnam (Drew & Romig, 2013).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F02E73DFF42A6B4DA4EFDAE.taxon	materials_examined	Material. Wangdue Phodrang, 5.8 km S Alla, Royal Orchard, 27.298 ° N, 89.968 ° E, 800 m, 18.08.2018, hand collecting, 1 ♀, CSCA 18 L 163 (FFP 17 TB 061) (M. Hauser leg.); Wangdue Phodrang, Bajo, ARDC orchards, 27.490 ° N, 89.900 ° E, 1230 m, hand collecting, 15.08.2017, 1 ♀ (FFP 17 BT 114) (C. Borkent & Hauser leg.); 21 – 28.08.2017, Multilure, 1 ♀, FFP 17 BT 018 (BHNPPC); Sarpang, 6.8 km NW Gelephu, ARDC orchard, 400 m, 26.910 ° N, 90.433 ° E, 23 – 27.08.2017, Methyl isoeugenol, 1 ♀, FFP 17 BT 139 (C. Borkent & Hauser leg.). Male lure. Latilure (alpha-ionol and cade oil) (McQuate and Peck, 2001). Host plants. Recorded for at least the 59 fruit-bearing plant taxa, belonging to 25 genera in 13 families, with recorded field (Leblanc et al., 2021; McQuate & Liquido, 2022). Significant pest of fruits of Capsicum and Solanum species (Drew & Romig, 2013). D i s t r i b u t i o n. Bangladesh (Leblanc et al., 2013), Bhutan (first record), Brunei, China (Fujian, Hainan, Hong Kong, Yunnan), India (Bihar, Karnataka, Tamil Nadu, West Bengal), Indonesia (Kalimantan, Sulawesi), Laos, Malaysia (peninsular), Myanmar, Pakistan, Singapore, Sri Lanka, Taiwan, Thailand, Vietnam; introduced Hawaii, Japan (Okinawa), Kenya, Tanzania (Drew & Romig, 2013; Vargas et al., 2015).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F03E73EFF4AA331DD20FF0E.taxon	description	Male lure. None known; records from methyl eugenol are regarded as accidental (Hancock & Drew, 2018 a, 2019). Host plants. Fruits of a range of Citrus species (Rutaceae) (Allwood et al., 1999). D i s t r i b u t i o n. Bhutan, Nepal, e. India (Sikkim, West Bengal), China (Jiangsu, Sichuan, Hubei, Hunan, Guangxi, Guizhou, Yunnan) (Drew & Romig, 2013).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F03E73DFF4AA26BDA41FD15.taxon	description	D i s t r i b u t i o n. Bhutan, India (Drew & Romig, 2013).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F00E73EFF42A10BDA01FC13.taxon	materials_examined	Material. Wangdue Phodrang, Bajo, ARDC orchards, 27.490 ° N, 89.900 ° E, 1230 m, 15 – 21.08.2017, Cue-lure, 1 Ơ, FFP 17 BT 010 (BHNPPC); 5.8 km S Alla, Royal Orchard, 27.298 ° N, 89.968 ° E, 800 m, 18.08.2017, Cue-lure, 1 Ơ, FFP 17 BT 130 (BHNPPC). M a l e l u r e. Cue-lure. Host plants. Fruits of Callicarpa arborea Roxb. (Lamiaceae), Capparis sepiaria L. (Capparaceae), Flueggea virosa (Roxb. ex Willd.) Royle (Phyllanthaceae), Zehneria wallichii (Cucurbitaceae) (Drew & Romig, 2013). D i s t r i b u t i o n. Bangladesh (Leblanc et al., 2014), Bhutan (first record), Cambodia (Leblanc et al., 2016), China (Zhang et al., 2011), Taiwan (Doorenweerd et al., 2019), Thailand (Drew & Romig, 2013), Nepal (Leblanc et al., 2019 a).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F00E73EFF42A10BDA01FC13.taxon	biology_ecology	Host plants. Prunus persica (L.) Batsch (Rosaceae) (Maneesh et al., 2022). D i s t r i b u t i o n. Bhutan, India (Uttar Pradesh), Pakistan (Maneesh et al., 2022). Note. Records of B. nigrofemoralis White & Tsuruta from Bhutan (Drew et al., 2007) were referred to B. divenderi by Maneesh et al. (2022).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F01E73FFF4AA0A2DABCFECE.taxon	description	D i s t r i b u t i o n. Bhutan, Cambodia (Drew & Romig, 2013).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F01E73FFF4AA171DA4BFE30.taxon	description	D i s t r i b u t i o n. Bhutan, China (Drew & Romig, 2013).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F01E73FFF4AA21BDAB0FD65.taxon	description	D i s t r i b u t i o n. Bhutan, Sri Lanka (Drew & Romig, 2013).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F01E73FFF4AA2AED875FBDA.taxon	materials_examined	Material. Sarpang, 7.7 km NW Gelephu, mixed veg farm, 380 m, 26.91 ° N, 90.422 ° E, 23 – 27.08.2017, Cue-lure, 1 Ơ, FFP 17 BT 086 (BHNPPC); Sarpang, 6.8 km NW Gelephu, ARDC orchards, 26.91 ° N, 90.433 ° E, 400 m, 23 – 27.08.2017, Cue-lure, 1 Ơ, FFP 17 BT 123 (BHNPPC). M a l e l u r e. Cue-lure, zingerone. Host plants. Litsea verticillata Hance (Lauraceae) (Liang et al., 1993). D i s t r i b u t i o n. Bangladesh (Leblanc et al., 2013), Bhutan (Drew et al., 2007), China (Wang and Zhao, 1989), India (David et al., 2017), Nepal (Leblanc et al., 2019), Taiwan (Doorenweerd et al., 2019), Sri Lanka (Leblanc et al., 2018), Thailand, Vietnam (Drew & Romig, 2013).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F01E73FFF4AA462DAB5FA2A.taxon	materials_examined	Material. Bhutan: Sarpang, 6.8 km NW, Gelephu, ARDC orchards, 26.91 ° N, 90.433 ° E, 400 m, 23 – 27.08.2017, Zingerone, 1 Ơ, FFP 17 BT 120 (CSCA 18 L 211); Wangdue Phodrang, 5.8 km S Alla, Royal Orchard, 27.298 ° N, 89.968 ° E, 800 m, 18 – 28.08.2017, Zingerone, 7 Ơ, FFP 17 BT 058 (BHNPPC). M a l e l u r e. Zingerone. Host plants. Syzygium jambos (L.) Alston (Myrtaceae) (Tsuruta & White, 2001). D i s t r i b u t i o n. Bangladesh (Leblanc et al., 2019 b), Bhutan (first record), India (David et al., 2017), Indonesia (Doorenweerd et al., 2020), Nepal (Leblanc et al., 2019 a), Sri Lanka (Tsuruta & White, 2001), Vietnam (Leblanc et al., 2018).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F01E730FF4AA5E4D898FB10.taxon	materials_examined	Material. Sarpang, 13 km NW Gelephu, 26.968 ° N, 90.559 ° E, 1200 m, 25 – 27.08.2017, Malaise trap, 1 Ơ, FFP 17 BT 096; Cue-lure, 2 Ơ, FFP 17 BT 098 (BHNPPC); 25.08.2017, hand collecting, 1 Ơ, FFP 17 BT 100 (C. Borkent); 6.8 km NW Gelephu, ARDC orchards, 26.91 ° N, 90.433 ° E, 400 m, 23 – 27.08.2017, Cue-lure, 1 Ơ, FFP 17 BT 123 (BHNPPC); Wangdue Phodrang, Bajo, ARDC orchards, 27.490 ° N, 89.900 ° E, 1230 m, 21 – 28.08.2017, Cue-lure, 2 Ơ, FFP 17 BT 015; 5.8 km S Alla, Royal Orchard, 27.298 ° N, 89.968 ° E, 800 m, 18.08.2017, Cue-lure, 3 Ơ, FFP 17 BT 130 (BHNPPC). M a l e l u r e. Cue-lure. H o s t p l a n t s. Elaeocarpus lanceifolius Roxb. (Elaeocarpaceae) (Drew & Hancock, 1994; Allwood et al., 1999). D i s t r i b u t i o n. Bhutan, Brunei, Cambodia, China (Yunnan), Thailand, Vietnam (Drew & Romig, 2013).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F0EE730FF42A532DADCF9FE.taxon	biology_ecology	Host plants. Pest of cultivated fruit, bred from at least 11 host species, including Mangifera indica L. (mango), Carica papaya L., and plum (Allwood et al., 1999). D i s t r i b u t i o n. Bangladesh, Bhutan (Drew & Romig, 2007; Drew & Romig, 2013), China, India, Myanmar, Nepal, Thailand, Vietnam (Bezzi, 1916; David et al., 2017; Drew et al., 2007; Hardy, 1973; Leblanc et al., 2014, 2019 a; Norrbom et al., 1999).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F0EE730FF42A65BDA4DF8D3.taxon	biology_ecology	Host plants. Species of wild Solanum including Solanum verbascifolium L. (Family Solanaceae) (Drew & Hancock, 1994; Allwood et al., 1999). D i s t r i b u t i o n. Bhutan (Drew et al., 2007), India, Indonesia (Java), Malaysia (peninsular), Sri Lanka, Thailand, Vietnam (Drew & Romig, 2013).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F0FE731FF4AA0CEDAFCFE9E.taxon	description	H o s t p l a n t s. Manilkara zapota (Sapotaceae) (White & Elson-Harris, 1992). D i s t r i b u t i o n. Bhutan, India, Sri Lanka (Drew & Romig, 2013).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F0FE731FF4AA1ABDB60FDBA.taxon	description	D i s t r i b u t i o n. Bhutan, India (Karnataka, Kerala, Maharashtra, Tamil Nadu) (Drew et al., 2007; Drew & Romig, 2013).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F0FE731FF4AA284DD56FC43.taxon	biology_ecology	Host plants. Polyphagous fruit pest bred from at least 54 hosts (Allwood et al., 1999; Leblanc et al., 2021; Culliney, 2022). D i s t r i b u t i o n. Bangladesh, Bhutan (Drew et al., 2007), India, Laos, Myanmar, Pakistan, Sri Lanka, Thailand, Vietnam; introduced to Mauritius, Réunion, Egypt, Iran, Israel, Libya, Oman, Saudi Arabia, United Arab Emirates, Yemen (Drew & Romig, 2013).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F0FE731FF4AA3CEDD7BFBBA.taxon	description	D i s t r i b u t i o n. Bhutan, Vietnam (Drew et al., 2007; Drew & Romig, 2013).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F0FE731FF4AA484DA35FAD5.taxon	description	M a l e l u r e. Cue-lure. D i s t r i b u t i o n. Bhutan (Drew & Romig, 2013).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F0FE731FF4AA561D81DF9E5.taxon	description	D i s t r i b u t i o n. Bhutan (Drew et al. 2007) and India (Himachal Pradesh) (Maneesh et al., 2022).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F0FE731FF4AA621D872F8CE.taxon	materials_examined	Material. Sarpang, 8.3 km NW Gelephu, 26.921 ° N, 90.424 ° E, 580 m, 23 – 27.08.2017, Zingerone, 1 Ơ, FFP 17 BT 080 (CSCA 18 L 201); 6.8 km NW Gelephu, ARDC orchards, 26.91 ° N, 90.433 ° E, 400 m, 23 – 27.08.2017, Zingerone, 1 Ơ, FFP 17 BT 120 (CSCA 18 L 211) (BHNPPC). M a l e l u r e. Zingerone. Distribution. Bangladesh (Leblanc et al., 2021), India (David et al., 2020 e); Bhutan (first record).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F0CE732FF42A0A2D86DFE10.taxon	biology_ecology	Host plants. A moderate pest of cucurbits, bred from fruits of Luffa acutangula (L.) Roxb., L. cylindrica (L.) M. Roem., Trichosanthes cucumerina L., and Zehneria wallichii (C. B. Clarke) C. Jeffrey (Cucurbitaceae) (Allwood et al., 1999). D i s t r i b u t i o n. Bangladesh, Bhutan (Drew et al., 2007), Brunei, China (Yunnan), India (Bihar, Karnataka), Indonesia (Bali, Java, Maluku, Sulawesi, West Nusa Tenggara), Laos, Malaysia (Peninsular, Sarawak), Myanmar, Philippines, Thailand, Vietnam (Drew & Romig, 2013).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F0CE732FF42A232DAD8FD73.taxon	description	D i s t r i b u t i o n. Bhutan, Thailand, Vietnam (Drew & Romig, 2013).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F0CE732FF42A2A9DD93FC20.taxon	materials_examined	Material. Sarpang, 10 km NW Gelephu, 26.95 ° N, 90.537 ° E, 790 m, 25 – 27.08.2017, Malaise trap, 1 Ơ, FFP 17 BT 104 (CSCA 18 L 229) (BHNPPC). Ma l e lu re. Cue-lure. D i s t r ibut i on. Bhutan (first record), China (Yunnan), Thailand (Drew & Romig, 2013).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F0DE733FF4AA319D8DCFC03.taxon	description	D i s t r i b u t i o n. Bhutan, China (Gansu), India (Arunachal Pradesh), northern Vietnam (Drew & Romig, 2013).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F0DE733FF4AA419DAC3FB03.taxon	description	D i s t r i b u t i o n. Bhutan, China (Guanxi, Yunnan), India (Arunachal Pradesh), Malaysia (Peninsular, Sarawak), Thailand, Vietnam. (Drew & Romig, 2013).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F0DE733FF4AA519D875FA03.taxon	description	D i s t r i b u t i o n. Bhutan, China (Sichuan, Yunnan), India (West Bengal) (Drew & Romig, 2013).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F0DE734FF4AA619D8ACFDAE.taxon	materials_examined	Material. Wangdue Phodrang, Bajo, ARDC orchards, 27.490 ° N, 89.900 ° E, 1230 m, 15 – 21.08.2017, Cue-lure, 1 Ơ, FFP 17 BT 010 (BHNPPC); 21 – 28.08.2017, Cue-lure, 15 Ơ, FFP 17 BT 015 (BHNPPC); 5.8 km S Alla, Royal Orchard, 27.298 ° N, 89.968 ° E, 800 m, 18.08.2017, Cue-lure, 2 Ơ, FFP 17 BT 130 (BHNPPC). M a l e l u r e. Cue-lure, zingerone. Host plants. At least 136 hosts in 30 families (Allwood et al., 1999; Leblanc et al., 2021; McQuate et al., 2022). Infests numerous cucurbit crops. D i s t r i b u t i o n. Afghanistan, Bangladesh, Bhutan, Brunei, Cambodia, s. China (widespread; Hainan, Yunnan), India, Indonesia (Java, Kalimantan, East & West Nusa Tenggara, Sulawesi, Sumatra), Laos, Malaysia, Myanmar, Nepal, Pakistan, Philippines, Singapore, Sri Lanka, Taiwan, Thailand, Vietnam; eradicated Japan (Ryukyu Islands), Kiribati; introduced Benin, Burkina Faso, Gambia, Ghana, Guinea, Ivory Coast, Kenya, Mali, Mauritius, Nigeria, Réunion, Senegal, Seychelles, Tanzania, Togo, Uganda, Egypt, Iran, Saudi Arabia, United Arab Emirates, Indonesia (North Maluku, Papua), Papua New Guinea (mainland, Bougainville, New Britain, New Ireland), Australia (Torres Strait Islands), Christmas I., Guam, Nauru, Northern Mariana Islands, Hawaii, Solomon Islands (Drew & Romig, 2013; Vargas et al., 2015).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F0AE734FF42A29DDDE8FC5A.taxon	materials_examined	Material. Wangdue Phodrang, Bajo, ARDC orchards, 27.490 ° N, 89.900 ° E, 1230 m, 21 – 28.08.2017, Cue-lure, 1 Ơ, FFP 17 BT 015 (BHNPPC). M a l e l u r e. Cue-lure. D i s t r i b u t i o n. Bhutan, China (Guangxi, Hainan, Sichuan, Yunnan), Indonesia (Java, Sumatra), Malaysia (peninsular), Taiwan, Thailand, Vietnam, India (Drew & Romig, 2013).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F0AE735FF42A3E9D891FE7E.taxon	materials_examined	Material. Wangdue Phodrag, Bajo, ARDC orchards, 27.490 ° N, 89.900 ° E, 1230 m, 15 – 21.08.2017, Multilure, 3 Ơ, 3 ♀, FFP 17 BT 013; Yeast, 1 Ơ, 1 ♀, FFP 17 BT 017; 21 – 28.08.2017, 1 ♀ (FFP 17 BT 014); Multilure, 6 Ơ, 9 ♀, FFP 17 BT 018 (BHNPPC). Ma l e lu re. Cue-lure. Host plants. Bred from flowers of nine species of Cucurbitaceae (Allwood et al., 1999). It is also known to secondarily feed on and damage cucurbit fruits. This species is also known to feed on 13 other hosts in 10 genera and four families (Leblanc et al., 2021). D i s t r i b u t i o n. Bangladesh, China, India, Nepal, Pakistan, Sri Lanka, Thailand, Vietnam (Coquillett, 1904; Drew & Romig, 2013; Kabir et al., 1991; Norrbom et al., 1999), Bhutan (first record).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F0BE735FF4AA28ADD10FC7A.taxon	materials_examined	Material. Wangdue Phodrang, Bajo, ARDC orchards, 27.490 ° N, 89.900 ° E, 1230 m, 21 – 28.08.2017, Cue-lure, 1 Ơ, FFP 17 BT 015 (BHNPPC). M a l e l u r e. Cue-lure. D i s t r i b u t i o n. Bhutan (first record), Thailand (Drew & Romig, 2013).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F0BE735FF4AA1DADA35FD8E.taxon	description	D i s t r i b u t i o n. Bhutan (Drew & Romig, 2013).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F0BE735FF4AA3D7DAC0FB95.taxon	description	D i s t r i b u t i o n. Bhutan, China, Thailand (Drew & Romig, 2013).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F08E736FF42A0BDDAD8FE45.taxon	biology_ecology	Host plants. Zehneria wallichii (= Mukia wallichii); Z. maysorensis (Cucurbitaceae) (Drew & Romig, 2013). D i s t r i b u t i o n. Bhutan, Thailand, Vietnam (Drew & Romig, 2013).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F08E736FF42A2A7DA2CFC85.taxon	description	D i s t r i b u t i o n. Bhutan (Drew & Romig, 2013).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F08E736FF42A660DAF1F8CA.taxon	biology_ecology	Host plants. Bred from flowers of species of Cucurbitaceae (Allwood et al., 1999). D i s t r i b u t i o n. Bhutan, India, Thailand (Drew & Romig, 2013).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F08E736FF42A1C9DA2CFD63.taxon	description	D i s t r i b u t i o n. Bhutan (Drew & Romig, 2013).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F08E736FF42A389DD3DFAC0.taxon	materials_examined	Material. Wangdue Phodrang, Bajo, ARDC orchards, 27.490 ° N, 89.900 ° E, 1230 m, 21 – 28.08.2017, Cue-lure, 6 Ơ, FFP 17 BT 015 (BHNPPC). M a l e l u r e. Cue-lure. Host plants. Bred from flowers of nine species of Cucurbitaceae (Allwood et al., 1999). D i s t r i b u t i o n. Bhutan, China (Guangxi, Yunnan), India (Arunachal Pradesh, Bihar, Himachal Pradesh, Karnataka, Meghalaya, Uttarakhand, West Bengal), Malaysia (peninsular), Myanmar, Nepal, Pakistan, Thailand, Vietnam (Drew & Romig, 2013).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F08E736FF42A55DDA0DF9A5.taxon	biology_ecology	Host plants. Bred from flowers of species of Cucurbitaceae (Allwood et al., 1999). D i s t r i b u t i o n. Bhutan, China, Japan, Korea, Malaysia (peninsular), Taiwan, Thailand, Vietnam (Drew & Romig, 2013; Norrbom, 2022).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F09E737FF4AA0BDDAA8FEB5.taxon	description	D i s t r i b u t i o n. Bhutan, Vietnam (Drew & Romig, 2013).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F09E737FF4AA190DB87FBF5.taxon	materials_examined	Material. Punakha, 13 km NW Punakha; 27.667 ° N, 89.775 ° E, 1400 m, 16 – 20.08.2017, Cue-lure, 3 Ơ, CSCA 19 L 109 (BHNPPC); Wangdue Phodrang; Bajo, ARDC orchards, 27.490 ° N, 89.900 ° E, 1230 m, 15 – 21.08.2017, Cue-lure, 5 Ơ FFP 17 BT 010 (BHNPPC); 21 – 28.08.2017, Cue-lure, 26 Ơ FFP 17 BT 015 (BHNPPC); 5.8 km S Alla, Royal Orchard, 27.298 ° N, 89.968 ° E, 800 m, 18.08.2017, Cue-lure, 6 Ơ, FFP 17 BT 130 (BHNPPC). M a l e l u r e. Cue-lure. Host plants. Bred from flowers of nine species of Cucurbitaceae (Allwood et al., 1999). It is also known to secondarily feed on and damage cucurbit fruits. This species is also known to feed on 13 other hosts in 10 genera and four families (Leblanc et al., 2021; Liquido et al., 2022 c). D i s t r i b u t i o n. Bangladesh, Bhutan, Brunei, Cambodia, China (Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hubei, Hunan, Jiangxi, Sichuan, Xizang, Yunnan, Zhejiang), India (Andaman & Nicobar Islands, Arunachal Pradesh, Bihar, Haryana, Himachal Pradesh, Jammu & Kashmir, Karnataka, Kerala, Maharashtra, Meghalaya, Punjab, Sikkim, Tamil Nadu, Uttarakhand, West Bengal), Indonesia (Bali, Java, Nusa Tenggara, Sumatra), Laos, Malaysia (Peninsular, Sabah, Sarawak), Myanmar, Singapore, Sri Lanka, Taiwan, Thailand, Vietnam (Drew & Romig, 2013).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F09E737FF4AA549D892F9BF.taxon	materials_examined	Material. Wangdue Phodrang, Bajo, ARDC orchards, 27.490 ° N, 89.900 ° E, 1230 m, 21 – 28.08.2017, Cue-lure, 2 Ơ, FFP 17 BT 015 (BHNPPC). M a l e l u r e. Cue-lure. D i s t r i b u t i o n. Bhutan, India, Nepal, Thailand, Vietnam (Drew & Romig, 2013; Hancock & Drew, 2018 b).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F09E728FF4AA69AD8DFFDF5.taxon	description	D i s t r i b u t i o n. Bhutan, India, Myanmar, Nepal, Pakistan, Sri Lanka, Vietnam (Drew & Romig, 2013). Note. Possible synonym of Z. tau (Drew & Romig, 2013). Dacinae: Gastrozonini Biology. Most species of the tribe Gastrozonini feed in bamboo culms mainly as saprophages. The Gastrozonini comprise 148 described species and several undescribed species (Kovac et al., 2006). They are regarded as a monophyletic entity, although to date there is no consistent morphological synapomorphy that would permit proper definition of clades (Dohm et al., 2014).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F16E728FF42A250D86AFC85.taxon	materials_examined	Material. Sarpang, 6.8 km NW, Gelephu, ARDC orchards, 26.910 ° N, 90.433 ° E, 400 m, 23.08.2017, hand collecting, 1 Ơ, FFP 17 BT 071 (CSCA 18 M 271) (M. Hauser leg.). H o s t p l a n t s. Acroceratitis larvae infest shoots of bamboo (Poaceae). Bambusa bambos (Kapoor, 1993), B. tulda, Dendrocalamus asper, D. membranaceus, D. strictus (Allwood et al., 1999). D i s t r i b u t i o n. Bangladesh, China (Yunnan), India (Assam, Bihar, Jharkhand, Uttarakhand, Uttar Pradesh, West Bengal), Myanmar, Thailand (Norrbom, 2022); Bhutan (first record).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F16E729FF42A3B0D872FED0.taxon	materials_examined	Material. Sarpang, 6.8 km NW Gelephu, ARDC orchards, 26.910 ° N, 90.433 ° E, 400 m, 23.08.2017, hand collecting, 2 ♀, FFP 17 BT 071 (CSCA 18 M 271) (M. Hauser leg.). H o s t p l a n t s. Shoots of Dendrocalamus pendulus Ridley, Gigantochloa scortechinii Gamble (Dohm et al., 2014). D i s t r i b u t i o n. Southern Thailand, Malaysia (mainland) (Norrbom, 2022); Bhutan (first record).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F17E729FF4AA147DA4BFC5A.taxon	materials_examined	Material. Sarpang, 6.8 km NW Gelephu, ARDC orchards, 26.910 ° N, 90.433 ° E, 400 m, 23.08.2017, hand collecting, 2 Ơ, 5 Ơ, FFP 17 BT 071 (CSCA 18 M 271) (M. Hauser leg.); 23 – 27.08.2017, Malaise trap, 1 ♀, FFP 17 BT 068 (BHNPPC); 23.08.2017, hand collecting, 1 ♀ (M. Hauser leg.) FFP 17 BT 071; 23.08.2017, hand collecting, 1 Ơ, 3 ♀, FFP 17 BT 072 (C. Borkent). Host plants. It has been bred from shoots of Bambusa dolichoclada, B. glaucescens, B. oldhami, B. shimadai, B. stenostachya, B. tulda, B. tuldoides, B. vulgaris var. vulgaris, Dendrocalamus asper, D. giganteus, D. hamiltonii, D. latiorus, Gigantochloa auriculata, G. nigrociliata, Phyllostachys mackinoi, P. nigripes, P. pubescens and Thyrostachys siamensis (Norrbom, 2022). D i s t r i b u t i o n. Bangladesh, Burma, China, India, Indonesia (Sumatra), Laos, Malaysia (peninsular), Taiwan, Thailand, Vietnam (Hancock & Drew, 1999); Bhutan (first record). Phytalmiinae: Acanthonevrini Biology. Most species of the subfamily have saprophagous larvae living under the bark of fallen trees or inside bamboo culms (Kovac & Streit, 1996; Dohm & Kovac, 2001; Korneyev & Ovtshinnikova, 2004; Dohm et al., 2008, 2014).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F17E72AFF4AA3F4D8DEFE9A.taxon	description	FFP 17 BT 096 (BHNPPC). D i s t r i b u t i o n. Myanmar (Hering, 1941); Bhutan (first record). Note. Males of this species have feathered fore femur and fore tarsi as in other species of Lenitovena Ito, and can be identified with the key by Hancock (2011). Its wing pattern is similar to that of L. varipes (Chen, 1948) differing from the latter by the presence of the second hyaline spot in the medial portion of cell dm (in L. varipes, only one hyaline spot is present near the apex of dm).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F14E72BFF42A1B7D891FF0E.taxon	materials_examined	Material. Sarpang, 6.8 km NW, Gelephu, ARDC orchards, 26.91 ° N, 90.433 ° E, 400 m, 24.08.2017, from banana, 1 ♀ (Hauser & C. Borkent leg.) FFP 17 BT 124 (CSCA 18 L 167); 23.08.2017, hand collecting, 1 Ơ (M. Hauser leg.) FFP 17 BT 071 (CSCA 18 M 271); 23 – 27.08.2017, Malaise trap, 1 m, 1 ♀ (BHNPPC) FFP 17 BT 068; 24 – 27.08.2017, 6 m MT, 1 Ơ (BHNPPC) FFP 17 BT 069. Host plants. Larvae saprophagous in the internodes of Bambusa vulgaris, Dendrocalamus asper, D. membranaceus, D. strictus (Permkam, 2005), Gigantochloa scortechinii, Schizostachyum cf. grande (Dohm et al., 2014). D i s t r i b u t i o n. Bangladesh, Brunei, Cambodia (Korneyev & Phauk, 2019), China (Fujian, Yunnan), India (Assam), Indonesia (Java, Kalimantan, Sulawesi, Maluku), Laos, Malaysia (mainland, Sarawak), Myanmar, Taiwan, Thailand, Vietnam (Norrbom, 2022); Bhutan (first record).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F15E72BFF4AA10AD891FDDE.taxon	materials_examined	Material. Sarpang, 6.8 km NW Gelephu, ARDC orchards, 26.91 ° N, 90.433 ° E, 400 m, 24 – 27.08.2017, 6 m MT, 4 Ơ, 1 ♀ (BHNPPC) FFP 17 BT 069; 23 – 27.08.2017, yeast, 2 Ơ, 1 Ơ (BHNPPC) FFP 17 BT 122. H o s t p l a n t s. Bambusa tuldoides, B. vulgaris, Cephalostachyum pergracile, Dendrocalamus nudus (Norrbom, 2022). D i s t r i b u t i o n. Bangladesh, China (Yunnan), India (Arunachal Pradesh, West Bengal), Indonesia (Java, Sumatra), Malaysia, Myanmar, Taiwan, Thailand (Norrbom, 2022); Bhutan (first record).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F15E72BFF4AA27ADA4BFCB0.taxon	materials_examined	Material. Punakha, 13 km NW Punakha, 27.667 ° N, 89.775 ° E, 1400 m, 16 – 20.08.2017, yeast, 2 Ơ, 1 ♀ (BHNPPC) FFP 17 BT 033. Distribution. China (Hainan, Heilongjiang, Sichuan, Yunnan), India (Arunachal Pradesh), Japan (Hokkaido to Ryukyu Islands), Korea, Laos, Myanmar, e. Russia, Taiwan, Thailand, Vietnam (Norrbom, 2022); Bhutan (first record).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F15E72CFF4AA36DDDA7FC3D.taxon	materials_examined	Material. Sarpang, 6.8 km NW Gelephu, ARDC orchards, 26.91 ° N, 90.433 ° E, 400 m, 24.08.2017, hand collecting, 1 ♀ (C. Borkent) FPP 17 BT 074 (CSCA 18 L 214); 23 – 27.08.2017, yeast, 3 Ơ (BHNPPC) FFP 17 BT 122. H o s t p l a n t s. Bambusa blumeana, B. polymorpha, Dendrocalamus pendulus, Gigantochloa scortechinii (Norrbom, 2022). D i s t r i b u t i o n. Bangladesh, China (Yunnan), India (Assam), Indonesia, Malaysia (peninsular), Myanmar, Thailand, Vietnam (Norrbom, 2022); Bhutan (first record).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F12E72DFF42A418D8DAFB6C.taxon	materials_examined	Material. Sarpang, 7.7 km NW Gelephu, mixed veg farm, 380 m, 26.91 ° N, 90.422 ° E, 23 – 27.08.2017, Cue-Lure, 1 ♀ (BHNPPC) FFP 17 BT 086; 8.3 km NW Gelephu, 26.921 ° N, 90.424 ° E, 580 m, 23 – 27.08.2017, Cue-lure, 1 Ơ (BHNPPC) FFP 17 BT 079. D i s t r i b u t i o n. China (Yunnan), India (Assam, Bengal, Karnataka) (Norrbom, 2022); Bhutan (first record). Tephritinae: Cecidocharini	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F13E72DFF4AA4B1D8EFF9C8.taxon	materials_examined	Material. Wangdue Phodrang, 3.5 km E Bajo, 27.486 ° N, 89.936 ° E, 1480 m, 17 – 21.08.2017, hand collecting, 1 ♀ (C. Borkent) FFPBT 052 (CSCA 18 L 220); Malaise trap, 1 ♀ (BHNPPC) FFPBT 050 (CSCA 18 L 223). Host plants. Chromolaena odorata (L.) R. M. King & H. Rob. (Norrbom, 2022) D i s t r i b u t i o n. Argentina, Brazil, Colombia, Venezuela; introduced to Ghana, Guam, India, Indonesia, Ivory Coast, Palau, Papua New Guinea, Thailand (Norrbom, 2022); Bhutan (first record). Note. Introduced into various countries in the Afrotropical, Australasian and Oriental Regions as an agent to control the “ triffid weed ” Chromolaena odorata. Tephritinae: Tephritini	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F13E72EFF4AA62DD8A5FE9A.taxon	description	Malaise trap, 1 Ơ, 3 ♀ (BHNPPC) FFP 17 BT 060 (CSCA 18 L 877). Host plants. Bidens biternata, B. macroptera, B. pilosa, B. tripartita, Coreopsis basalis, C. grandiflora, Cosmos caudatus, Dahlia pinnata, Guizotia abyssinica, Lactuca sativa, Lycopodium clavatum (Norrbom, 2022). D i s t r i b u t i o n. Azores, Canary Islands, Madeira, North Africa (Algeria, Tunisia); Spain; Western Asia (Saudi Arabia, Turkey, Yemen); India, Nepal, Thailand, China, Taiwan, S to Southern Africa & Australia, New Caledonia, Samoa, Niue, Cook Islands, French Polynesia, Fiji, Hawaii (Norrbom, 2022); Bhutan (first record). Note. Subcosmopolitan species associated with some Asteraceae: Heliantheae plants spread by humans.	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F10E72EFF42A37BDDB2FBE3.taxon	materials_examined	Material. Punkha, 1 km S Mendrelgan, 21 – 28.08.2017, 27.519 ° N, 89.832 ° E, 1625 m, Malaise trap, 1 Ơ (BHNPPC) FFP 17 BT 127 (CSCA 18 L 323). D i s t r i but i on. China: Guangxi, Taiwan (Norrbom, 2022); Bhutan (first record).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F10E72EFF42A1B7D891FCAC.taxon	materials_examined	Material. Punakha, 1 km S Mendrelgang, 27.519 ° N, 89.832 ° E, 1625 m, 16 – 21.08.2017, Malaise trap, 1 Ơ (BHNPPC) FFP 17 BT 043; Wangdue Phodrang, 5.8 km S Alla, Royal Orchard, 27.298 ° N, 89.968 ° E, 800 m, 18 – 28.08.2017, Malaise trap, 1 Ơ, 1 ♀ (BHNPPC) FFP 17 BT 060 (CSCA 18 L 877). H o s t p l a n t s. Acmella caulirhiza, Ageratum conyzoides, Brachyscome squamatus, Ceruana pratensis, Helichrysum scorpioides, Leptorhynchos squamatus, Matricaria aurea, Tithonia diversifolia, Xerochrysum subundulatum (Norrbom, 2022). D i s t r i b u t i o n. Egypt, China, Saudi Arabia, Yemen; Taiwan, India, Thailand, Indonesia; widespread in Afrotropical, Oriental, & Australasian Regions (Norrbom, 2022); Bhutan (first record). Trypetinae: Trypetini	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F1EE720FF47A4BBDD6AFAA7.taxon	materials_examined	Material. Wangdue Phodrang, 8.5 km ENE Bajo, 1600 m, 27.532 ° N, 89.976 ° E, 15.08.2017, hand collecting, 1 Ơ (C. Borkent & Hauser leg.) FFP 17 BT 025. D i s t r i b u t i o n. Malaysia (Hancock & Drew, 1994 a); Bhutan (first record).	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
03A3D2261F1CE723FF42A0A2DAA0FCBB.taxon	materials_examined	Material. Thimphu, 3 km NNE Dochula, 27.518 ° N, 89.755 ° E, 2360 m, 19.08.2017, hand collecting, 2 Ơ (M. Hauser leg.) FFP 17 BT 065 (CSCA 19 L 110). D i s t r i b u t i o n. India (Hendel, 1915; Kapoor, 1993); Bhutan (first record). Note. This species has short ocellar setae, 3 slightly proclinate frontal setae, and a wide apical crossband, characters shared with other species related to T. zoe Meigen, 1826. In the latter species, the wing pattern is sexually dimorphic, with the preapical crossband better developed in females and usually reduced in males; on the contrary, males usually have an extensive irregular dark area on and posterbasal to the pterostigma also covering the radial fork. The specimens on hand are males, which have an entirely brownish- or reddish-yellow mesonotum without black spots on the mediotergite as in the holotype female of T. indica, but the female wing pattern (fig. 21, d) is conspicuously different from the male wing pattern (fig. 21, a) in having complete discal crossband; however, similar dimorphism occurs in other species of the zoe group. We consider these males to belong to T. indica. The locality in Bhutan is only ca. 150 km from Dargeeling, its type locality.	en	Korneyev, S. V., Hauser, M., Borkent, C., Maples, B. K., Roubtsova, T. V., Zangpo, T., Dorji, S., Chophel, S., Dorji, N., Tsomo, Dendup, U., Dawa, K., Dorji, L., Dhimal, C. Mani, Kinley, R., Dorji, U., Dema, Y., Korneyev, V. A., Gaimari, S. D. (2023): The Fruit Flies (Diptera, Tephritidae) In Bhutan: New Faunistic Records And Compendium Of Fauna. Zoodiversity 57 (2): 93-124, DOI: 10.15407/zoo2023.02.093UDC595.773.4
