taxonID	type	description	language	source
03A18784FFBDDA3175B2FD1DFDCDF9C7.taxon	description	(Figs 12 – 14, 33)	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFBDDA3175B2FD1DFDCDF9C7.taxon	materials_examined	Holotype: SAUDI ARABIA ♂ 01. iii. 2024 Asir prov., 10 km N Al Habil 18.164 N, 42.246 E, leg. M. Halada (CSE)	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFBDDA3175B2FD1DFDCDF9C7.taxon	diagnosis	Diagnosis: Brachystegus nemkovi is unique among Palearctic Brachystegus in having a flat clypeus, with an inconspicuously delimited apical lamella. In remaining species, the clypeus has a conspicuous cross-carina, or is rectangulary stepped apically. The colour pattern of B. nemkovi is also distinctive, with an all yellow clypeus, an all red antenna, and the mesosoma lacking yellow spots. The terga are characterized by a rich yellow color.	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFBDDA3175B2FD1DFDCDF9C7.taxon	description	Description of male holotype: Body length: 10.2 mm. Color: Clypeus yellow (apart small basal black zone), mandible basally yellow, medially red, apically black. Labrum, antenna and legs light reddish, coxa partly red. Pronotum dark red, remaining mesosoma black. Terga black with yellow spots or bands, lateral spots on T 1, medially narrowed band on T 2, visible parts of T 3 – 6 all yellow. T 7 reddish. S 1 and S 7 with some red. Wing venation black, wings somewhat infumate. Lower face, most parts of mesosoma, basal half of T 1 and S 2 covered with dense, appressed silver pubescence, concealing integument. Femora below with dense silver pubescence. Morphology: Clypeus flat, apically with lamella, with some macro- and many micropunctures, ACM truncate. AS 2 basally narrowed, AS 2 – 3 and AS 12 as long as wide, AS 4 – 11 shorter than wide, AS 13 below as long as AS 11 + 12, conspicuously curved. Lower face with prominent median keel. Lower gena with keel at innerside near mouthparts. Face, upper surface of mesosoma, mesopleuron and T 1 with coarse, comb-like sculpture. Remaining terga with dense macropunctation, less than a diameter apart, interspaces microsculptured. Metasoma with cross-carina, medially interrupted. T 6 laterally with dentiform keel. T 7 tridentate, median tooth larger than lateral tooth. Apical margin of S 2 – 5 medially with long white bristles, laterally each with a yellow lamella. Female: Unknown.	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFBDDA3175B2FD1DFDCDF9C7.taxon	distribution	Distribution: Asir province in Saudi Arabia (Fig. 33).	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFBDDA3175B2FD1DFDCDF9C7.taxon	etymology	Etymology: The species is named in honor of the late Pavel G. Nemkov, a Russian expert on Spheciformes, who revised Brachystegus and other genera of Bembicidae. He supported the scientific work of the first author with exchange and identification of specimens.	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFBDDA3675B2F978FDEFFBA6.taxon	description	(Figs 8 – 11, 33)	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFBDDA3675B2F978FDEFFBA6.taxon	materials_examined	Holotype: OMAN ♀ 25. vi. 2018, Al Mudhaibi, Samad Ashan, Aswareg 22.82 N, 58.15 E. leg. AJ (ZMHU). Paratypes. 2 ♀ same data as holotype, leg. AJ (AJ / CSE); 1 ♂ collected on 16. vii. 2020, same location as holotype leg AJ (AJ).	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFBDDA3675B2F978FDEFFBA6.taxon	diagnosis	Diagnosis: Brachystegus samadashanensis is characterized by a transverse carina of the clypeus, by a black antenna (apex of male antenna red), by yellow spots on the pronotum and the foreleg, and by tergal yellow spots, widely separated from each other. T 5 – 6 in females and T 6 – 7 in males are black. Another Palearctic species with a black antenna and yellow spots on the pronotum is B. scalaris Illiger, 1807 from Europe and western Asia. It differs by an all red foreleg and by a yellow spot on the scutellum. The appressed silver pubescence of the head and mesosoma is denser in B. samadashanensis than in B. scalaris. Brachystegus schneideri, also described from nothern Oman, is another similar species. It is larger on average (see description) and has a red antenna and a mostly red pronotum. The male (the female is unknown) differs also from Brachystegus samadashanensis by continuous or shortly interrupted yellow bands on T 2 – 6.	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFBDDA3675B2F978FDEFFBA6.taxon	description	Description of female holotype: Body length: 8.5 mm. Color: All black, apical half of mandible, tegula, wing base and legs reddish, apex of mandible dark red, coxa black except apex, forefemur on outer surface in apical half and entire foretibia with yellow spot / band. First AS dark red. Pronotum laterally with yellow spot. T 1 – 4 with large lateral spots, black space inbetween as wide or wider than spots. Face and mesosoma covered with dense silver pubescence. Morphology: Clypeus in apical forth with conspicuous edge, area under edge at right angle to basal clypeus surface, somewhat longer than diameter of lateral ocellus. ACM medially truncate. Lower face with prominent median keel. Sculpture of face and mesosoma comb-like, hidden under pubescence. Metanotum with cross-carina. T 1 with dense medium-sized punctures, interspaces shiny, remaining terga with grainlike dense fine punctation. T 6 with some rugae. Description of male: Body length: 8.0 mm. Agreeing in general aspects with female and in morphology also with B. schneideri. Antennal apex red, antenna below somewhat reddish, black above. Cross-carina of clypeus hidden under long silver pilosity, difficult to recognize. Spots on T 5 small, black space inbetween 2.5 × as wide as spot diameter. T 6 – 7 black, T 7 with narrow lateral tooth, and wide medial tooth, apically rounded, nearly truncate.	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFBDDA3675B2F978FDEFFBA6.taxon	distribution	Distribution: Northern Oman (Fig. 33).	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFBDDA3675B2F978FDEFFBA6.taxon	etymology	Etymology: The species is named after its collecting locality, Samad Ashan, a town in Northern Oman.	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFBDDA3675B2F978FDEFFBA6.taxon	discussion	Remark: The type locality is placed south of the Alhajer (= Al Hajar) mountain in the gravel desert, whereas all collecting places of the similar B. schneideri are located north of the Alhajer mountain on the coastal plain, partly with sandy places. This indicates that the two species have different ecological requirements and explains why they occur so close together geographically.	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFBADA3675B2FBADFC49F822.taxon	description	(Figs 3 – 7, 33)	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFBADA3675B2FBADFC49F822.taxon	materials_examined	Holotype: OMAN 1 ♂ 25. iii. 2023 Hazam, 23 km NNE Ar Rustaq 23.558 N, 57.496 E, leg. M. Halada (CSE). Paratypes: 1 ♂ 21. iii. 2023 20 km SE Sur 22.454 N, 59.625 E; 1 ♂ 26. iii. 2023 Al Ajal, 30 km SW Seeb 23.531 N, 57.975 E, leg. M. Halada (CSE).	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFBADA3675B2FBADFC49F822.taxon	diagnosis	Diagnosis: The species is unique among Palearctic Brachystegus in having a red antenna, red legs and in lacking yellow spots on the mesosoma.	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFBADA3675B2FBADFC49F822.taxon	description	Description of male holotype: Body length: 10.0 mm. Color: Black with the following parts reddish: Basal half of mandible, antenna, apical half of pronotum, upper part of mesopleuron, apico-lateral corners of mesoscutum, parts of scutellum, tegula, wing base, legs (coxae only apically), S 1 laterally. T 1 – 3 with large lateral yellow spots, T 4 – 5 with narrow yellow bands. Wing venation black, wings slightly infumate. Clypeus and most of face, dorsal surface of propodeum laterally, underside of mesosoma and S 2 with dense appressed silver pilosity, remaining mesosoma with more sparse pubescence. Morphology: Clypeus in apical forth with conspicuous edge, area under edge at right angle to basal clypeus surface, approximately as long as diameter of lateral ocellus. ACM with two teeth. Lower face with prominent median keel. Face, upper surface of mesosoma and mesopleuron with coarse, comb-like sculpture. Terga with dense macropunctation, punctures less than diameter apart, interspaces microsculptured. Metasoma with indistinct cross-carina. T 6 laterally with dentiform keel. T 7 tridentate, median tooth small and rounded. Apical margins of S 2 – 5 medially with long white bristles, laterally each with narrow and reddish lamella. Paratypes agree with holotype, one paratype measures only 8.5 mm.	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFBADA3675B2FBADFC49F822.taxon	distribution	Distribution: Northern Oman (Fig. 33).	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFBADA3675B2FBADFC49F822.taxon	etymology	Etymology: The species is named in honor to Dr. Jörg Schneider from Rheinsbach, Germany, a friend and insect photographer, who also accompanied us on a collecting trip to Oman.	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFB8DA3475B2FF74FDD1FB82.taxon	description	(Figs 15 – 17, 36)	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFB8DA3475B2FF74FDD1FB82.taxon	materials_examined	Holotype: OMAN ♀ 27. ii. 2020 Al Suwayq, Hail al Ghafa 23.769 N, 57.342 E, leg. AJ (ZMHU). Paratype: 1 ♀ same data as holotype (AJ).	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFB8DA3475B2FF74FDD1FB82.taxon	diagnosis	Diagnosis: Nysson waleedi is unique among Arabian Nyssonini in having a shiny body surface in combination with a reddish colour and a sparse punctation with large interspaces on most parts of the mesosoma. The remainig species of the genus even in the Mediterreanean area are conspicuously sculptured or densely punctate, and always with parts of the body black.	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFB8DA3475B2FF74FDD1FB82.taxon	description	Description of female holotype: Body length: 3.5 mm. Color: Orange red, face and mesoscutum somewhat darker, labrum, parts of antenna, pronotal tubercle and pronotum somewhat lighter than remaining body. Tip of mandible black. Hindtibia with yellow band on outer surface. T 1 – 3 with thin yellow bands, shortly interrupted medially on T 3. Tergal margins of T 4 – 5 somewhat yellowish. Face laterally and mesopleuron below with inconspicuous, lateral part of propodeal dorsum with dense, silver pubescence. Wing venation brown, basally red. Wings transparent. Morphology: ACM straight, laterally with small rectangular corner. Clypeus, face, mesoscutum, scutellum and mesopleuron shiny and polished, with scattered punctation, punctures mostly several diameters apart. Vertex behind and gena rounded, with narrow occipital carina separating mouth parts from gena. Propodeal dorsum striate, propodeal posterior surface punctatorugose. Punctation of T 1 – 2 larger and denser than on mesoscutum, punctures approximately one diameter apart, punctation of remaining terga somewhat denser. T 6 with confluent larger punctures, for shape see Fig. 16. Antenna claviform, AS 2 – 3 as long as wide, AS 3 – 11 shorter than wide, AS 12 as long as both previous AS combined. Male: Unknown.	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFB8DA3475B2FF74FDD1FB82.taxon	distribution	Distribution: Northern Oman (Fig. 36).	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFB8DA3475B2FF74FDD1FB82.taxon	etymology	Etymology: The species is named in honor of Waleed Mubark Al-Busaidi from Muscat, Oman, a specialist of plant science at Sultan Qaboos University.	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFB8DA3A75B2FB48FD23FF4E.taxon	description	(Figs 18 – 20, 34)	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFB8DA3A75B2FB48FD23FF4E.taxon	materials_examined	Holotype: SAUDI ARABIA ♀ 09. iii. 2024 Riyad prov., 100 km NW Riyadh, Ulya 25.281 N, 46.021 E, leg. M. Halada (CSE). Paratypes: ♀ same data as holotype (CSE). Oman 2 ♀ 27. ii. 2020 Al Suwayq, Hail al Ghafa 23.769 N, 57.342 E, leg. AJ (AJ / CSE).	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFB8DA3A75B2FB48FD23FF4E.taxon	diagnosis	Diagnosis: Nysson woodi is characterized by the color pattern. The head and the mesosoma are all black, with four white spots or a complete band on the pronotum and a white / red spot on the scutellum (lacking in holotype), and with an all red metasoma (except some black on T 6) with white tergal bands. The vertex and the gena apically have a small carina along whole length.	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFB8DA3A75B2FB48FD23FF4E.taxon	description	Description of female holotype: Body length: 4.5 mm. Color: Head and mesosoma black, mandible basally yellowish, apex of clypeus and first antennal segments below dark reddish. Pronotal lobe and two spots on pronotum medio-apically yellowish-white. Tegula and wing base red, wing venation black, wings transparent. Legs red (trochanter only apically), outer surface of tibiae and forefemur with yellow spot or band. Metasoma red, terga with lateral white spots, medial red part smaller than spots, T 6 black, laterally and apically red (in paratype all red). Face, mesopleuron and lateral part of propodeal dorsum with dense, silver appressed pubescence. Morphology: ACM straight, clypeus medially bulged. AS 3 somewhat longer than wide, AS 4 – 11 as long as wide, AS 12 1.4 × as long as wide. Frons, mesoscutum and S 2 with punctures 0.5 – 1.5 diameters apart, interspaces shiny, scutellum and mesopleuron more coarsely punctate. Propodeal dorsum coarsely striate. Terga with sparse macro- and fine and dense micropuntation inbetween. T 6 with pygidial plate surrounded by carina, with coarse punctation, interspaces shiny. Variation in paratypes: Females from Oman are characterized by a more extended yellow color (pronotum all yellow apically, all tibiae with yellow band on whole outer surface, scutellum with red / yellow spot, pronotum and propodeum laterally red). The paratype female from Saudi Arabia is intermediate between the Omani specimens and the holotype, with a medially interrupted band on the pronotum and a red scutellum, and shorter yellow bands on all tibiae. Male: Unknown.	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFB8DA3A75B2FB48FD23FF4E.taxon	distribution	Distribution: Northern Oman, Central Saudi Arabia (Fig. 34).	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFB8DA3A75B2FB48FD23FF4E.taxon	etymology	Etymology: The species is named after Thomas J. Wood from the Naturalis Biodiversity Center in Leiden, Netherlands, a friend and specialist of bees.	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFB6DA3A75B2FE5CFB47FCEA.taxon	description	(Fig, 21, 36). Nysson arabicus Gadallah and Edmardash in Abu el-Ghiet, Gadallah, Gasib, Al-Fifi, and Edmardash, 2023: 155, ♂. Holotype: ♂, Saudi Arabia: Sala Mountain: Abdul Rahman Al Jabri farm at 17 ° 04 ' 10.5 '' N 43 ° 07 ' 43.3 '' E (Efflatoun Bey collection, Entomology Department, Cairo University, Giza, Egypt (depository not given in the original description, taken from Pulawski, 2025).	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFB6DA3A75B2FE5CFB47FCEA.taxon	discussion	Remark: The species was described in Nysson, but belongs clearly to Synnevrus, which is restored from synonymy here. See introduction for details. Recognition: The male of Synnevrus arabicus can be recognized by the key characters, mainly by the colour pattern. It is the only Arabian species of the genus with a black mesosoma (with some red only) and with yellow spots or bands on T 1 - 5.	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFB6DA3A75B2FE5CFB47FCEA.taxon	distribution	Distribution: Only known from the type area in the Jazan Province in Saudi Arabia (Fig. 36).	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFB6DA3A75B2FC60FBF9F9FC.taxon	description	(Figs 22 – 24, 34)	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFB6DA3A75B2FC60FBF9F9FC.taxon	materials_examined	Material examined: OMAN 2 ♀ 19. iii. 2019; 1 ♀ 03. v. 2019; 1 ♀ 05. vii. 2019 Al Mudhaibi, Samad Ashan, Al Akhdhar 22.84 N, 58,17 E, leg. AJ (AJ / CSE); 4 ♀ 11. iii. 2023 Hamman, 13 km SW Muscat 23.470 N, 58,319 E, leg. M. Halada (CSE); 1 ♀ 21. iii. 2023 20 km SE Sur 22.454 N, 59.625 E, leg. M. Halada (CSE). SAUDI ARABIA: 2 ♂ 3 ♀ Jizan Pr., 18 km N Baish 17.52 N 42.56 E, leg M. Halada (CSE). UNITED ARAB EMIRATES: 1 ♀ 15 – 18. iv. 1984; 1 ♂ 26 – 30. iv. 1984 Dubai / al-Awir, 1 ♀ 16. vi. 2009 Wadi Wurayah, leg. T. v. Harten (CSE). Recognition: Synnevrus barrei can be easily recognized by the red mesosoma. This is predominantly black in the other species, except for the yellow markings. Two males have each a yellow spot on the clypeus laterally, the third male has an all black clypeus.	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFB6DA3A75B2FC60FBF9F9FC.taxon	distribution	Distribution: Algeria, Azerbaijan, Central Asia, Iran, Oman, Saudi Arabia, UAE (Fig. 34). The records from Oman and Saudi Arabia are the first records for these countries.	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFB6DA3A75B2FC60FBF9F9FC.taxon	discussion	Remark: The species was formerly placed in Synnevrus, which was synonymized with Nysson by Nemkov & Leley (2013), and the corresponding species were consequently placed in Nysson (Nemkov 2017). However, we consider Synnevrus to be a valid genus and the species is again listed under this genus. See introduction for details. The specimens from the UAE were already published by Schmid-Egger (2011, 2014).	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFB6DA3875B2F95CFE0AFCA3.taxon	description	(Figs 25 – 32, 34)	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFB6DA3875B2F95CFE0AFCA3.taxon	materials_examined	Holotype: SAUDI ARABIA ♀ 25. ii. 2024 Jazan prov., 3 km N Ayban 17.300 N, 43,056 E, leg. M. Halada (CSE). Paratypes: 2 ♀ same data as holotype (CSE); 1 ♂ 21. x. 2024 Mekkah pr., 13 km W Sabt Al Alayah 19.559 N, 41.831 E, leg. M. Halada (CSE)	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFB6DA3875B2F95CFE0AFCA3.taxon	diagnosis	Diagnosis: Synnevrus mekkahensis is characterized by the sharp cross-carina of the clypeus (indistinct in male), which forms a sharp right angle with the lower part of the clypeus. It is rounded in S. barrei and lacks in the other Synnevrus from Arabia. Colour pattern is also unique and distinctive: mesosoma is mostly black with a short median yellow band on the pronotum, T 1 - 2 with yellow band or spots only.	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFB6DA3875B2F95CFE0AFCA3.taxon	description	Description of female holotype: Body length: 6.5 mm. Color: Black, dark red are: mandible (except for black apex), basal half of antenna, pronotum, tegula, propodeum laterally, part of legs, S 2, T 5 – 6. Pronotum with short yellow band medially. T 1 with lateral yellow spots, widely separated, T 2 with continuous narrow yellow band, medially emarginate. Face, mesopleuron and propodeal lateral surface with dense, silver pubescence. Wing venation dark, wings transparent. Morphology: Clypeus in apical forth with conspicuous edge, area under edge at right angle to the basal clypeus surface, shiny, approximately as long as diameter of lateral ocellus. ACM with two medial teeth. Gena in lower half with large carina, and a second smaller continuous carina at inner border of gena and vertex. Keel between antennal insertions V-shaped above. Face, mesoscutum and mesopleuron with coarse, comb-like punctation. Pronotum laterally with obtuse angle as seen in dorsal view. T 1 – 5 and sterna with macropunctation, punctures approximately 0.5 – 1.5 diameter, interspaces dull with fine micropunctation. T 6 with lateral carina, surface somewhat shiny, with confluent punctation. Description of male: Body length: 6.0 mm. Agreeing in general aspects and color pattern with female. Differs by the following characters: clypeal cross-carina and space below less developed, barely visible in the dense silver pubescence. Hindtibia apically on outer surface with small yellow spot. T 3 – 7 partly or all reddish. T 6 laterally with spine. T 7 laterally with black spine, medially with widely rounded tooth.	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFB6DA3875B2F95CFE0AFCA3.taxon	distribution	Distribution: Jazan and Mekkah province in Saudi Arabia (Fig. 34).	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFB6DA3875B2F95CFE0AFCA3.taxon	etymology	Etymology: The species is named after the Saudi Arabian town and province of Mekkah, where one of the paratypes was collected.	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFB4DA3875B2FCA9FF4BF86B.taxon	description	(Figs 31 – 32, 35)	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFB4DA3875B2FCA9FF4BF86B.taxon	materials_examined	Material examined: OMAN 5 ♂ 2 29. viii. 2019 Dhofar, Wadi Darbat near Highway 17.065 N, 54.431 E, leg. CSE (CSE); 1 ♀ 30. viii. 2019 Dhofar, 2 km W Taqah 17.033 N, 54.357 E, leg. AJ (AJ); 5 ♂ 30. viii. 2019 Dhofar, 6 km E Taqah 17.049 N, 54.454 E, leg. CSE (CSE); 1 ♂ 31. viii. 2019 Dhofar, 18 km W Raysut, Ayn Ishat 16,997 N, 53,821 E, leg. CSE (CSE); 3 ♂ 04. ix. 2019 Dhofar, 13 km N Mirbat, Samhan mountain 17.111 N, 54.711 E, leg. CSE (CSE); 1 ♂ 27. ii. 2020 Dhofar, 2 km W Taqah 17.033 N, 54.357 E, leg. AJ (AJ); 1 ♂ 03. iii. 2020 Nemer, Wadi Ronub 18.529 N, 56.347 E, leg. AJ (CSE); 1 ♀ 12. x. 2021 20 km W Salalah, Ayn Humran 17.097 N, 54.284 E, leg.; 11 ♂ 7 12. x. 2021 80 km SSW Salalah 16,857 N, 53,420 E; 1 ♀ 15. x. 2021 75 km W Salalah 16,842 N, 53,434 E; 1 ♂ 1 16. iii. 2023 4 km S Jamma 2 km N Mansur 23.521 N, 57.553 E; 1 ♂ 21. iii. 2023 20 km SE Sur 22.454 N, 59.625 E; 1 ♂ 21. ix. 2023 15 km W Mirbat 17.033 N, 54.592 E, M; 6 ♂ 1 24. ix. 2023 5 km NE Magsail 16,910 N, 53,803 E; 3 ♂ 2 30. ix. 2023 6 km NW Thalkut 16.751 N, 53.204 E, leg. M. Halada (CSE). SAUDI ARABIA 1 ♂ 25. ii. 2024 Jazan prov., Wadi Jawra, Ayban 17.45 N, 43.07 E; 1 ♀ 28. ii. 2024 Jazan prov., 25 km N Baysh 17.586 N, 42.624 E; 1 ♀ 21. x. 2024 Jizan pr., 12 km W Al Khawbah 16.788 N, 43.114 E; 1 ♀ 22. x. 2024; 1 ♂ 22. x. 2024 Makkah pr., QarnHuthail, wadi 19.252 N, 41.794 E; 3 ♂ 24. x. 2024 Asir pr., 30 km NNW Muhayil 18.773 N, 41.960 E; 1 ♀ 26. x. 2024 Jizan pr., N of Ad Darb, W. Reem 18.841 N, 42.267 E; 2 ♀ 03. xi. 2024; 4 ♂ 03. xi. 2024 Jizan pr., 18 km N Baish, 17.526 N, 42.587 E; 1 ♀ 08. xi. 2024 Jizan pr., 20 km SE Abu Arish, Iskan 16.829 N, 42.971 E; 8 ♂ 09. xi. 2024 Jizan pr., 12 km W Al Khawbah 16.788 N, 43.114 E, leg. M. Halada (CSE), UNITED ARAB EMIRATES 1 ♂ 26. iv. 1984 Dubai / Al Awir 24.93 N, 55.16 E, leg. Sudgen (ZMHU, type specimen). Recognition: Synnevrus ohli is unique among Arabian Nyssonni in having the combination of a continuous white band on the pronotum, a white basal spot on the scutellum and white lateral spots on T 1 – 3 only (in rare cases also on T 4). The base of T 1 is usually black, but may be red in males and females.	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFB4DA3875B2FCA9FF4BF86B.taxon	distribution	Distribution: Oman, Saudi Arabia and UAE (Fig. 35). It is by far the commonest species of Nyssonini on the Arabian Peninsula. The specimens from the UAE were already published by Schmid-Egger (2011, 2014). The records from Oman are the first records for that country.	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
03A18784FFB4DA3875B2FCA9FF4BF86B.taxon	discussion	Remark: Synnevrus ohli was described in Synnevrus. This genus was synonymized with Nysson by Nemkov & Lelej (2013), and the corresponding species were consequently placed in Nysson by Nemkov (2017). However, we consider Synnevrus to be a valid genus, and the species is again listed under this genus. See introduction for details.	en	Schmid-Egger, Christian, Al-Jahdhami, Ali A. (2025): Nyssonini of the Arabian Peninsula with description of six new species (Hymenoptera, Spheciformes, Bembicidae). Zootaxa 5696 (2): 247-259, DOI: 10.11646/zootaxa.5696.2.4, URL: https://doi.org/10.11646/zootaxa.5696.2.4
