taxonID	type	description	language	source
C16778878DEE727FEECA8D2B7C3525BA.taxon	diagnosis	Diagnosis. General appearance: wasp-mimetic. Head gibbous, large, as long as wide across eyes, densely covered with long setae on ventral and postocular portions; postantennal tubercles very short to almost imperceptible, acute or rounded; legs elongated, slender; fore femur slightly longer than head and pronotum together, thicker basally. Hemelytra long, surpassing the abdomen by about half of the length of the membrane.	en	Gil-Santana, Helcio R. (2016): First description of the male of Hiranetisatra Stal and new country records, with taxonomic notes on other species of Hiranetis Spinola (Hemiptera, Heteroptera, Reduviidae, Harpactorinae). ZooKeys 605: 91-111, DOI: http://dx.doi.org/10.3897/zookeys.605.8797, URL: http://dx.doi.org/10.3897/zookeys.605.8797
2F19147F271E5A598E79B1407A5B9A0B.taxon	materials_examined	Material examined. One male and one female, each with a green label with the same information: " Museum Paris, Costa Rica, Paul Serre, 1920 " (MNHN). One female, labels: Hiranetis atra Stal / Balzapamba, (Ecuad.), R. Haensch S. / Hiranetis atra Stal, Breddin det. / k [?] olle v. [green label] (ZMHB).	en	Gil-Santana, Helcio R. (2016): First description of the male of Hiranetisatra Stal and new country records, with taxonomic notes on other species of Hiranetis Spinola (Hemiptera, Heteroptera, Reduviidae, Harpactorinae). ZooKeys 605: 91-111, DOI: http://dx.doi.org/10.3897/zookeys.605.8797, URL: http://dx.doi.org/10.3897/zookeys.605.8797
2F19147F271E5A598E79B1407A5B9A0B.taxon	diagnosis	Diagnosis. Hiranetis atra can be readily separated from other species of the genus by its general coloration, which is mostly blackish, especially the hemelytra, which are completely dark (Figs 1 - 3, 5 - 6), while the other species have the pattern of yellowish or straw-colored hemelytra, with a median transverse band and dark apex.	en	Gil-Santana, Helcio R. (2016): First description of the male of Hiranetisatra Stal and new country records, with taxonomic notes on other species of Hiranetis Spinola (Hemiptera, Heteroptera, Reduviidae, Harpactorinae). ZooKeys 605: 91-111, DOI: http://dx.doi.org/10.3897/zookeys.605.8797, URL: http://dx.doi.org/10.3897/zookeys.605.8797
2F19147F271E5A598E79B1407A5B9A0B.taxon	description	Description. MALE. Figures 6 - 17. Measurements (mm): Total length: to tip of abdomen: 12.1; to tip of hemelytra: 16.2; head: total length (lateral view): 1.9; maximum width across eyes: 1.9; interocular space: 1.0; antennal segments: I: 5.5; II: 1.7; III (very bent; approximately): 6.9; IV: 2.0; labium segments: II [first visible]: 1.4; III: 1.1; IV: 0.3. Thorax: pronotum: fore lobe length: 0.7; hind lobe: length: 2.0; width at posterior margin: 2.8. Legs: fore legs: femur: 5.4; tibia: 5.5; tarsus: 0.7; mid legs: femur: 4.8; tibia: 6.2; tarsus: 0.7; hind legs: femur: 6.5; tibia: 9.1; tarsus: 0.8. Abdomen: length: 6.3; maximum width: 2.3. Coloration: general coloration black (Fig. 6). Head, including antennae and labium, blackish, eyes brownish-black. Thorax blackish, with exception of metanotum, which is reddish-brown. Hemelytra blackish. Legs mostly blackish; fore femur with dorsal surface, except at base and extreme apex, pale yellowish, and with a lighter-colored subbasal portion ventrally; mid and hind femora with yellowish annulus situated somewhat distally to their midportion (Fig. 6). Abdominal segments II and III (first two visible) reddish; sternite IV almost completely reddish, except on posterolateral portion, including connexivum at this area, where it is blackish; sternite V mostly reddish but blackish on posterior and lateral portion, including connexivum. Tergites IV and V, and remaining segments, including pygophore and parameres, blackish. Structure and vestiture: Integument mostly shiny, smooth. Head gibbous, large, as long as wide across eyes; integument shiny, with sparse long and short, straight or somewhat curved blackish setae; the latter much denser, forming pubescence of long blackish thick setae on postocular portion and gula; almost completely glabrous between eyes. Labium curved, with scattered and somewhat curved blackish setae. Antennal segments I and II straight, the former approximately three times longer than head, with shiny and smooth integument and sparse short darkened setae; segments II-IV with opaque and somewhat rugose integument; segment II, except at basis, covered with very numer ous darkened short setae, with some longer intermixed setae and some very thinner elements at distal portions (interpreted as trichobothria); segment III thickened in basal half, curved; III and IV covered with dense, very short and somewhat lighter-colored pubescence, with short darkened setae scattered on segment III and few of these on segment IV; the latter is thinner than the other segments and moderately curved. Postantennal tubercles small and somewhat acuminate. Eyes globose, glabrous, projecting laterally, prominent in dorsal view, reaching dorsal margin of head at interocular sulcus in approximately its midportion; not reaching ventral margin of head, which is far from inferior margin of the eye. Interocular sulcus thin and moderately deep. Ocelli elevated, much closer to eyes than to each other. Collum thin. Thorax with shiny integument; prothorax covered with very numerous blackish thick setae on forelobe, anterior portions of propleura and hind lobe; the latter with sparse long setae at dorsal portion or, almost glabrous, except on midline, where thinner, somewhat shorter and light yellowish to whitish setae form a faint midlongitudinal line on hind lobe. Transverse sulcus not very deep, interrupted before middle by a pair of submedian shallow carina. Midlongitudinal sulcus on forelobe of pronotum becoming abruptly deeper at transverse sulcus to form a depression; posteriorly to the latter, a blunt short rounded prominence; disc of hind lobe smooth; lateral longitudinal sulci well marked at posterior half to posterior two-thirds of hind lobe of pronotum. Humeral angle elevated, rounded at lateral margin; median portion of posterior margin of pronotum with some long thin darkened setae. Scutellum elevated at disc, pointed posteriorly, with scattered thin dark long setae. Posterior portion of propleura, mesopleura, metapleura and thoracic sterna with long darkened setae, which are shorter and thinner at center of mesosternum and metasternum. Legs: coxae with numerous long dark setae on distal half, ventrally, and some longer thinner light-colored elements, while the basal third and lateral portions are almost completely glabrous; trochanters densely covered with long setae ventrally and with some scattered even longer thinner setae, which are lighter-colored on forelegs and dark on mid and hind legs. Fore femur subequally longer than head and pronotum together; all femora thicker basally and slightly subapically too, covered with scattered few long and strong dark setae and with a dense group of long and thick setae and some thinner and even longer setae on ventral portion of the basal enlarged portion of femora; these setae are lighter on fore femora and darker on the others. Fore tibiae somewhat curved, with uniform thickness, except at apex, which is somewhat enlarged, and where there is a dorsal spur and a mesal comb. Mid and hind tibiae straight and somewhat thickened at basal half. All tibiae with scattered long thick blackish setae; fore and mid tibiae covered with shorter dark setae on ventral surface, which become progressively more numerous towards apex, where they also covers lateral and dorsal surfaces; hind tibiae, except at base, densely covered with short dark setae, which are somewhat longer in the slightly enlarged basal half. Tarsi with moderately long dark setae. Hemelytra long, surpassing abdomen by about half length of membrane; corium with curved scattered adpressed short dark setae, which are much more numerous over costal and subcostal veins, becoming less numerous on distal half of corium, including over those veins; membrane glabrous. Abdomen: elongate; spiracles rounded; sternites with shiny integument and sparse long thin setae, which are light on reddish portions and dark on the blackish segments, and thicker, longer and more numerous on parts adjacent to genitalia and on the latter too. There is also a fusiform grouping of whitish minute short setae on midlateral portions of sternite V. MALE GENITALIA (Figs 7 - 17): pygophore: blackish, subpentagonal in ventral view, with a subtriangular rounded apex (medial process) (Fig. 7); lateral to the latter, a somewhat deep and rounded emargination (Fig. 8); between anterior and genital opening, a very well sclerotized bridge that has a conspicuous median dorsal rounded prominence; long, thick and dark setae ventrally (on exposed surface), somewhat more numerous on lateroapical portions. Parameres symmetrical, rod-like in shape; somewhat curved in basal half and straight towards apices, which are rounded, blackish, glabrous in basal two-thirds and with long setae in apical third; those at apicomedial margins even longer (Fig. 9). Phallus somewhat elongate when not inflated (Fig. 10); articulatory apparatus with basal plate arms and bridge narrow, forming a subrectangular set, except in apical portion, where the arms are curved (Fig. 11); basal plate extension (pedicel) moderately short, slightly expanded towards apex and somewhat more sclerotized than the arms and basal bridge (Fig. 11). Dorsal phallothecal plate weakly sclerotized, flat, suboval in shape, with numerous longitudinal thin grooves at apical half; apical margin almost transverse, straight (Fig. 12). Struts with curved lateral arms and parallel somewhat curved median arms which are expanded at apex into a pair of asymmetrical sub oval / subsquared structures; there is a medial bridge joining the bases of the latter (Figs 12 - 13). Endosoma wall mostly minutely spiny, with a small smooth semi-oblong dorsal prolongation at midportion (Figs 14 - 15). After endosoma extension, seven processes were observed: 1 - a larger U to M-shaped basal process formed by diffuse thickening (Fig. 14); 2 - a median subspherical process, situated between the upper arms of the basal process, formed by minute tooth-like thickenings (Figs 14 - 16); 3 - a pair of elongate apical-median flat, longitudinally striated and somewhat curved and moderately sclerotized processes, wrapped in a smooth portion (not minutely spiny, but with fine longitudinal grooves) of endosoma wall, all of which lying dorsally to the other subapical processes described next (Fig. 17); 4 - a pair of small sclerotized thickened curved processes, located below the next process (Fig. 15); 5 - a transverse thickening above the pair of processes described previously (Fig. 15). The spiny endosoma wall above the latter process has larger and more sclerotized elements (Fig. 15). FEMALE (from Costa Rica): Measurements (mm): Total length: to tip of abdomen: 16.5; to tip of hemelytra: 21.0; head: total length (lateral view): 2.3; maximum width across eyes: 2.2; interocular space: 1.2; antennal segments: I: 6.3; II: 2.3; III-IV: absent; labium segments: II [first visible]: 1.6; III: 1.3; IV: 0.5. Thorax: pronotum: fore lobe length: 0.9; hind lobe: length: 2.5; width at posterior margin: 3.5. Legs: fore legs: femur: 6.0; tibia: 6.0; tarsus: 0.7; mid legs: femur: 4.9; tibia: 6.5; tarsus: 0.7; hind legs: femur: 7.2; tibia: 10.0; tarsus: 0.9. Abdomen: length: 9.5; maximum width: 3.8. Similar to male (Fig. 5). Anterior half of stridulitrum lighter-colored, reddish; sternite IV completely reddish; sternite V almost completely reddish, except on posterolateral portion, including connexivum in this area, where it is blackish; mid-anterior portion of sternite VI somewhat reddish.	en	Gil-Santana, Helcio R. (2016): First description of the male of Hiranetisatra Stal and new country records, with taxonomic notes on other species of Hiranetis Spinola (Hemiptera, Heteroptera, Reduviidae, Harpactorinae). ZooKeys 605: 91-111, DOI: http://dx.doi.org/10.3897/zookeys.605.8797, URL: http://dx.doi.org/10.3897/zookeys.605.8797
2F19147F271E5A598E79B1407A5B9A0B.taxon	distribution	Distribution. Colombia (Stal 1872, Maldonado 1990).	en	Gil-Santana, Helcio R. (2016): First description of the male of Hiranetisatra Stal and new country records, with taxonomic notes on other species of Hiranetis Spinola (Hemiptera, Heteroptera, Reduviidae, Harpactorinae). ZooKeys 605: 91-111, DOI: http://dx.doi.org/10.3897/zookeys.605.8797, URL: http://dx.doi.org/10.3897/zookeys.605.8797
823C3188649E7618C170C2C5120DD4B6.taxon	materials_examined	Material examined. Myocoris braconiformis, female, " typus ", labels: 2777 / Braconiformis, N., Stoll. Cim. t. 21. f. 147 [green label] / Para, Sieber [green label] / Typus [red label]; Myocoris pompilodes, female, " typus ", labels: 2771 / Pompilodes, N. [green label] / Cameta, Sieber [green label] / Typus [red label]; Myocoris pompilodes, male, " allotypus ", labels: 2771 / * Hiranetis pompilodes Burm., ♂, Allotypus / Cameta, Sieber [green label] / Allo-Typus [red label] (ZMHB). The female " typus " of Hiranetis braconiformis (Burmeister), described from " Para " (Burmeister 1835, 1838), is deposited in ZMHB (Figs 18 - 20). This region (" Para ") is today a state in the northern region of Brazil, within the Amazonian region of South America. The female " typus " and a male " allotypus " of Hiranetis pompilodes (Burmeister), from " Cameta ", are also deposited in ZMHB (Figs 21 - 24). Because the original description (Burmeister 1838) did not designate a holotype, it is better to consider all the type specimens to be syntypes. In the male type, the distal portion of the abdomen is missing (Fig. 23). Although Burmeister (1838) had mentioned that " Cameta " was in " South Brazil ", the only locality with this name in Brazil is the municipality of " Cameta " in the same northern state of Para, from which Hiranetis braconiformis was described. It is possible that all these specimens were collected in the same region (Para) and even during the same period, since on all the labels, the name " Cameta " was followed by the name " Sieber " and at least those of the female syntype were apparently handwritten by the same person (Figs 20, 22). As a matter of fact, Friedrich Wilhelm Sieber was a servant and preparator of Johann Centurius Count von Hoffmannsegg, who obtained permission from the King of Portugal to send him to Brazil to collect insects. Leaving Lisbon in 1801, Sieber went to the province of Para, where he remained 12 years, collecting in different parts of this province, including Cameta (Papavero 1971). Friedrich W. Sieber did not collect in other regions of Brazil and remained in Amazonia throughout this period (Papavero 1971), which reinforces the preceding assertion. All of these points may be important in ascertaining the type locality of these taxa and are particularly relevant because Hiranetis pompilodes was subsequently considered by Champion (1898) to be a junior synonym of Hiranetis braconiformis. With the exception of the mention of the length, the descriptions of Hiranetis braconiformis and Hiranetis pompilodes emphasized only their coloration (Burmeister 1835, 1838). Stal (1872) stated that the two taxa were very similar and Hiranetis pompilodes differed from Hiranetis braconiformis through the coloration of the thorax (blackish, except at its margin), coxae, trochanters and basal portion of fore femora (yellowish and not blackish). Champion (1898) considered Hiranetis pompilodes to be a junior synonym of Hiranetis braconiformis. He reported that he had examined " a long series " of Hiranetis braconiformis, stating that it varied in " the colour of the pronotum and also to a certain extent in that of the femora. In many of the specimens the pronotum is entirely rufo-testaceous (braconiformis, Burm.); but in others (...) it is partly or entirely black, the basal margin or a subtriangular patch on the disc behind being pale in some examples (pompilodes, Burm.). " Champion (1898) also recorded variation in coloration of the femora, which are sometimes narrowly (fore femora) to broadly (mid and hind femora) black basally; " the hind pair have the apex broadly, and rarely a median ring, fuscous or black ", while the mid femora are often infuscate apically. In his figures of a pair of this species, he highlighted the variation in color amongst specimens from the same locality. The fact that Champion (1898) recorded the color variation among specimens from the same locality, which had been attributed by Burmeister (1838) and Stal (1872) to Hiranetis braconiformis and Hiranetis pompilodes, may be considered to be arguments in favor of both the historical evidence that the types of these taxa must have been collected in the same region (Brazilian state of Para) and the assumption that they belong to the same species as stated by Champion (1898). On the other hand, Champion (1898) apparently did not examine any type specimens of these taxa, or any specimen from Brazil. He also did not mention how many specimens formed his " long series ", or whether there might be any other sexual differences besides the third antennal segment thickened at its base. Moreover, he did not take into account any features other than coloration when commenting on the synonymy between Hiranetis braconiformis and Hiranetis pompilodes. Subsequently, Wygodzinsky (1949) still listed Hiranetis pompilodes as a valid species in his catalogue, while Maldonado (1990) considered it to be a junior synonym of Hiranetis braconiformis.	en	Gil-Santana, Helcio R. (2016): First description of the male of Hiranetisatra Stal and new country records, with taxonomic notes on other species of Hiranetis Spinola (Hemiptera, Heteroptera, Reduviidae, Harpactorinae). ZooKeys 605: 91-111, DOI: http://dx.doi.org/10.3897/zookeys.605.8797, URL: http://dx.doi.org/10.3897/zookeys.605.8797
823C3188649E7618C170C2C5120DD4B6.taxon	distribution	Distribution. Brazil (state of Para, Amazonian region) (Burmeister 1835, 1838), Mexico, Guatemala, Costa Rica, Panama and Guyana (Champion 1898).	en	Gil-Santana, Helcio R. (2016): First description of the male of Hiranetisatra Stal and new country records, with taxonomic notes on other species of Hiranetis Spinola (Hemiptera, Heteroptera, Reduviidae, Harpactorinae). ZooKeys 605: 91-111, DOI: http://dx.doi.org/10.3897/zookeys.605.8797, URL: http://dx.doi.org/10.3897/zookeys.605.8797
9D10A60C925B90CF98F3BA1B59719AF9.taxon	materials_examined	Material examined. Myocoris barbipes, female, " typus ", labels: 2772 / barbipes, two unrecognizable markings, &, ♀. / Bras. r. Olf. [green label] / Typus [red label] (ZMHB). The description of Hiranetis mentioned some structural features, and also that their tibiae are all hairy (Spinola 1840). This was followed by the description of Hiranetis membranacea, based on one or more females and males from Brazil, without ascribing any specific locality to the specimens described. The antennae, body and legs of this species were recorded as black; the hemelytra as entirely membranous, blackish, often slightly darker at their base to their end, but all transparent: a large yellow spot, on three-quarters of their length on outer edge, and a smaller, hyaline, also on the external borders. Single measurements were attributed to the species (" m. long 9. lign. Larg. 2. Lign. "; approximately 20.3 and 4.5 mm, respectively). After making this short description, Spinola (1840) commented that Hiranetis membranacea did not seem to be rare in South America and often showed variation: 1 - in the coloration of the thorax and abdomen, which were black, brown or even testaceous; 2 - in the legs, which could have yellowish annulus or be entirely yellowish; 3 - in the coloration of the hemelytra, which could be lighter-colored or hyaline, even in the basal portion, in some specimens; 4 - in the size, which could be half of or a third smaller. However, he concluded by stating that the intermediary specimens that he had at hand left no doubt in his own mind regarding the unity of the species. Unfortunately, it was not possible to locate any type specimen of Hiranetis membranacea. The material described by Massimiliano Spinola (1780 - 1857) is in his collection, which is deposited in the " Museo Regionale di Scienze Naturali ", Turin, Italy (Schuh and Slater 1995). More than a decade ago, when looking for a type of other species described by M. Spinola (see Forero and Gil-Santana 2003), I contacted its [former] curator, Dr Mauro Daccordi, who kindly donated the catalogue of Spinola's hemipterological collection (Casale 1981), clarifying that all extant specimens were listed there. There is no reference to any specimens of Hiranetis membranacea in it. It is worth mentioning that after M. Spinola's death (1857), his hemipterological collection remained in Tassarolo Castle until its acquisition by Museo Regionale di Scienze Naturali of Turin in 1979 (Casolari and Moreno 1980, Casale 1981). Taking into account " the precarious conditions the Collection was exposed for over a century " (Casale 1981), the types of Hiranetis membranacea can be considered lost. Nonetheless, at the end of 2015, I contacted the current curator of the Museum, Dr Marinella Garzena, who also kindly confirmed that no specimens of this species are present there. Therefore, it must be assumed that no type specimens of Hiranetis membranacea Spinola exist anymore. Myocoris barbipes was considered to be the largest species among several other species that were included in Myocoris Burmeister, 1835, at that time (Burmeister 1838). This species was recorded as coming from " Rio Janeiro " (Burmeister 1838). Its female " typus " is deposited in ZMHB (Figs 25 - 27). Regarding its type locality, " Rio Janeiro " (Burmeister 1838) may correspond to the current municipality of Rio de Janeiro or, because of the historical scenario at the beginning of the nineteenth century, more likely it should be extended to the state of Rio de Janeiro or even to some of the contiguous states in southeastern Brazil as they are currently delimited. In fact, the handwritten de scription on the green label attached to the type seems to read " Bras. r. Olf. " (Fig. 27). It is known that in 1816, Ignaz Franz Werner Maria von Olfers arrived in Rio de Janeiro with the Legation of Prussia to study Brazilian nature. He collected extensively in trips from the state of Rio de Janeiro to the contiguous states of Minas Gerais and Sao Paulo, and back to Rio de Janeiro, in the years 1818 to 1820. His collection, including insects, was then sent to museums in Vienna and Berlin (Papavero 1971). Herrich-Schaeffer (1848) provided a figure (habitus) and a short diagnosis of Hiranetis membranacea (as Myocoris membranaceus). The diagnosis referred only to color features: [general coloration] red; antenna, head, femora apices, tibiae and tarsi black; hemelytra pale yellowish with a median band and apex dark. He then commented on the variation in coloration and size, as had previously been recorded by Spinola (1840) for this species. In a footnote, Stal (1859) mentioned Hiranetis barbipes (" = membranaceus Spin.; H. Sh. "). On the other hand, in Stal (1872), Myocoris barbipes was set as a junior synonym of Hiranetis membranacea. There was no mention of the reasons for attributing synonymy to these two species. However, with exception of Walker (1873), this was adopted in all the subsequent catalogues (Lethierry and Severin 1896, Wygodzinsky 1949, Maldonado 1990). Stal (1872) recorded features of structure and vestiture in his diagnosis of the species of Hiranetis. However, it is unlikely that these will be helpful in ascertaining better characteristics of Hiranetis membranacea, so as to remove doubts regarding the validity of the synonymy that he proposed, and / or to provide better knowledge about the diagnostic features of all taxa discussed here. Firstly because he reported that he had examined a single specimen of Hiranetis membranacea from " Brasilia " [i. e., country of Brazil], which he stated was deposited in the Museum of Stockholm [" Mus. Holm. "]. Taking into consideration all the historical data on types of Hiranetis membranacea, there is no evidence that this specimen could be a type. In this case, it becomes clear that he did not examine the type of Myocoris barbipes that is still extant and is deposited in Berlin (ZMHB; Figs 25 - 27), even though he placed Myocoris barbipes as a junior synonym of Hiranetis membranacea. Similarly, regarding Hiranetis braconiformis, he also cited " Mus. Holm. ", thus denoting that he probably used other specimen (s) but not the type (also deposited in ZMHB; Figs 18 - 20) to define the features of the latter species. Secondly, among those features, some are common to other species of Hiranetis and coincide with the diagnosis of the genus, or may even be common to species of other genera. Some other features are known to vary among specimens and the possibility of inter-individual variation was probably not taken into consideration at that time. Thirdly, as discussed below, the recorded variations in Hiranetis membranacea (Spinola 1840) and H. [cf.] braconiformis (Champion 1848), the similarities in coloration between them and the absence of records of other or better features of each of them when they were originally described (Spinola 1840, Burmeister 1835, 1838) make any identification imprecise. This compromises the diagnosis of Hiranetis membranacea and Hiranetis braconiformis furnished by Stal (1872), because it seems that he did not examine any type specimens of these species.	en	Gil-Santana, Helcio R. (2016): First description of the male of Hiranetisatra Stal and new country records, with taxonomic notes on other species of Hiranetis Spinola (Hemiptera, Heteroptera, Reduviidae, Harpactorinae). ZooKeys 605: 91-111, DOI: http://dx.doi.org/10.3897/zookeys.605.8797, URL: http://dx.doi.org/10.3897/zookeys.605.8797
9D10A60C925B90CF98F3BA1B59719AF9.taxon	distribution	Distribution. Brazil (Spinola 1840, Burmeister 1838, Herrich-Schaeffer 1848, Maldonado 1990).	en	Gil-Santana, Helcio R. (2016): First description of the male of Hiranetisatra Stal and new country records, with taxonomic notes on other species of Hiranetis Spinola (Hemiptera, Heteroptera, Reduviidae, Harpactorinae). ZooKeys 605: 91-111, DOI: http://dx.doi.org/10.3897/zookeys.605.8797, URL: http://dx.doi.org/10.3897/zookeys.605.8797
