taxonID	type	description	language	source
03AF87F95279FF9DB463FA45FB6AF840.taxon	description	Fig 1	en	Johanson, Kjell A., Pham, Hong-Thai (2012): Three new species of Helicopsyche (Trichoptera, Helicopsychidae) from northern Vietnam, with a key to Helicopsyche species of Vietnam. European Journal of Taxonomy 6: 1-10, DOI: 10.5852/ejt.2012.6
03AF87F95279FF9DB463FA45FB6AF840.taxon	diagnosis	Diagnosis Helicopsyche melina sp. nov. is most similar to H. admata Malicky & Chantaramongkol, 1992, H. dacklestenensis Schefter & Johanson, 2001 and H. nastia Malicky & Melnitsky (Melnitsky & Malicky 2008) particularly due to the presence of four-segmented maxillary palps in combination with the nearly similar shape of the gonopods and tergum X in lateral view. The new species is easily distinguished from these by the presence of a well-developed transverse plate of segment IX located at mid-height of posterior part of the segment; the primary branch of each gonopod is produced anterad; and the secondary branch of each gonopod is long, straight, and oriented in right angle to the primary branch of the gonopods.	en	Johanson, Kjell A., Pham, Hong-Thai (2012): Three new species of Helicopsyche (Trichoptera, Helicopsychidae) from northern Vietnam, with a key to Helicopsyche species of Vietnam. European Journal of Taxonomy 6: 1-10, DOI: 10.5852/ejt.2012.6
03AF87F95279FF9DB463FA45FB6AF840.taxon	etymology	Etymology Melina, named after the Melinh Station for Biodiversity, near the type locality.	en	Johanson, Kjell A., Pham, Hong-Thai (2012): Three new species of Helicopsyche (Trichoptera, Helicopsychidae) from northern Vietnam, with a key to Helicopsyche species of Vietnam. European Journal of Taxonomy 6: 1-10, DOI: 10.5852/ejt.2012.6
03AF87F95279FF9DB463FA45FB6AF840.taxon	materials_examined	Material examined Holotype ♂ VIETNAM: Vinh Phuc Prov, Me Linh District, stream near Ngoc Thanh Village, Melinh Station for Biodiversity, 21 ° 23 ' 28.2 " N, 105 ° 42 ' 46.7 " E, 233 m, Malaise trap 4 - 12 Apr. 2011, loc # VN 001, leg. K. A. Johanson, P. H. Thai & T. T. Du (NHRS, alcohol) [DNA voucher IP 7]. Paratypes ♂ VIETNAM: 8 ♂♂, Vinh Phuc Prov, Me Linh Distr, stream near Ngoc Thanh Village, 21 ° 23 ' 40.1 " N, 105 ° 42 ' 54.9 " E, light trap 5 Apr. 2011, loc # VN 008, leg. K. A. Johanson, P. H. Thai & T. T. Du (IEBR, alcohol); 241 ♂♂, Vinh Phuc Prov, Me Linh District, stream near Ngoc Thanh Village, 21 ° 23 ' 48.4 " N, 105 ° 42 ' 56.1 " E, 186 m, Malaise trap 5 - 12 Apr. 2011, loc # VN 006, leg. K. A. Johanson, P. H. Thai & T. T. Du (NHRS, alcohol); 20 ♂♂, Vinh Phuc Prov, Me Linh District, stream near Ngoc Thanh Village, 21 ° 23 ' 48.4 " N, 105 ° 42 ' 56.1 " E, 186 m, Malaise trap 5 - 12 Apr. 2011, loc # VN 006, leg. K. A. Johanson, P. H. Thai & T. T. Du (IEBR, alcohol); 34 ♂♂, Vinh Phuc Prov, Me Linh Distr, stream near Ngoc Thanh Village, 21 ° 23 ' 38.9 " N, 105 ° 42 ' 50.7 " E, light trap 4 Apr. 2011, loc # VN 002, leg. K. A. Johanson, P. H. Thai & T. T. Du (NHRS, alcohol); 2 ♂♂, Vinh Phuc Prov, Me Linh Distr, stream near Ngoc Thanh Village, 21 ° 23 ' 58.7 " N, 105 ° 42 ' 49.8 " E, light trap 5 Apr. 2011, loc # VN 007, leg. K. A. Johanson, P. H. Thai & T. T. Du (NHRS, alcohol).	en	Johanson, Kjell A., Pham, Hong-Thai (2012): Three new species of Helicopsyche (Trichoptera, Helicopsychidae) from northern Vietnam, with a key to Helicopsyche species of Vietnam. European Journal of Taxonomy 6: 1-10, DOI: 10.5852/ejt.2012.6
03AF87F95279FF9DB463FA45FB6AF840.taxon	description	Description MAXILLARY PALPS. Four-segmented, each segment gradually shorter distally. Pair of interantennal setal warts bean shaped, each about as long as width of individual scapus. Each scapus about as long as each basal segment of maxillary palps. Spur formula 1,2,4; each foreleg spur slightly shorter than each first tarsomere. LENGTH. Forewing 2.8 - 3.6 mm, hind wing 2.3 - 2.9 mm (N = 10). MALE ABDOMEN AND GENITALIA. (Fig. 1) Sternal process VI about one-fifth as long as sternum VI, straight, oriented posteroventrad; slightly tapering in lateral view (Fig. 1 A); with slightly club-shaped apex in ventral view (Fig. 1 B). Segment IX about as long as high in lateral view (Fig. 1 C); each anterior lobe of segment IX (Fig. 1 C) narrowly and symmetrically ellipsoid in lateral view, oriented anterad; anterodorsal and anteroventral margins almost straight (Fig. 1 C); in dorsal view, inner margin forming narrowly ellipsoid cavity (Fig. 1 D); in ventral view, without central posterior process (Fig. 1 E); inner margin forming nearly triangular cavity; lateral apodemes nearly horizontal (Fig. 1 C), widening anteriorly, meeting anterior margin, sub-marginal line absent; tergal transverse apodemes well-developed; sternal transverse apodeme absent. Transverse plate of segment IX located at mid-height of posterior margin of segment IX; in lateral view almost club-shaped, curving ventrad (Fig. 1 C); in ventral view forming broadly heart-shaped plate above basal part of gonopods (Fig. 1 E). Tergum X, in lateral view, oriented posterad (Fig. 1 C), almost straight, tapering along its length; apex strongly pointed (Fig. 1 D); in dorsal view (Fig. 1 D), deeply divided into pair of tapering, divergent branches, with about 3 pairs of equally long megasetae in apical group, starting opposite to apex of gonopods in lateral view (Fig. 1 C). Superior appendages tubular (Fig. 1 D), slightly curving dorsally. Primary branch of gonopods, in lateral view (Fig. 1 C), oriented dorsally, broadened distad to mid-height; anterior margin smooth, strongly concave; dorsal and posterior margins edged, undulating, posteroventral margin almost straight (Fig. 1 C); apices produced mesad into rounded lobes above basolateral part of tergum X. Secondary branch of gonopods nearly as long as secondary branch of gonopods; slender, almost straight in lateral view, except apex slightly curving dorsad; with few setae; curving mediad in ventral view (Fig. 1 E). Basomesal lobes absent. Basal plate, in lateral view (Fig. 1 C), nearly straight, narrowing along its length, slightly produced beyond anteroventral margin of segment IX in lateral view (Fig. 1 C); narrow in ventral view (Fig. 1 E). Phallic apparatus, lateral view, tube-shaped along its length (Fig. 1 F), narrowest shortly after basis, apex about as broad as base; nearly straight after mid-length; in ventral view, with basis narrow (Fig. 1 G); widest immediately after mid-length; endotheca weakly produced, posteroventral part not sclerotized, except pair of weakly sclerotized pair of triangular processes distally of phallic sclerite.	en	Johanson, Kjell A., Pham, Hong-Thai (2012): Three new species of Helicopsyche (Trichoptera, Helicopsychidae) from northern Vietnam, with a key to Helicopsyche species of Vietnam. European Journal of Taxonomy 6: 1-10, DOI: 10.5852/ejt.2012.6
03AF87F9527CFF9BB468FED6FB2DFE19.taxon	description	Fig. 2	en	Johanson, Kjell A., Pham, Hong-Thai (2012): Three new species of Helicopsyche (Trichoptera, Helicopsychidae) from northern Vietnam, with a key to Helicopsyche species of Vietnam. European Journal of Taxonomy 6: 1-10, DOI: 10.5852/ejt.2012.6
03AF87F9527CFF9BB468FED6FB2DFE19.taxon	diagnosis	Diagnosis Helicopsyche meander sp. nov. is most similar to H. lamnata sp. nov., H. anaktangga Malicky, 1995, H. anaksaku Malicky, 1995, H. minyas Malicky & Nawvong (Malicky et al. 2004) and H. cymodoce Schmid, 1993, particularly due to the presence of two-segmented maxillary palps in combination with the nearly similar shape of the gonopods in lateral view. The new species is distinguished from H. anaktangga, H. anaksaku, H. minyas and H. cymodoce by the pointed apex of tergum X in lateral view; and from H. lamnata sp. nov., H. anaksaku and H. minyas by the more strongly S-shaped gonopods. It is furthermore separated from H. lamnata sp. nov. by the absence of a central posterior process of segment IX.	en	Johanson, Kjell A., Pham, Hong-Thai (2012): Three new species of Helicopsyche (Trichoptera, Helicopsychidae) from northern Vietnam, with a key to Helicopsyche species of Vietnam. European Journal of Taxonomy 6: 1-10, DOI: 10.5852/ejt.2012.6
03AF87F9527CFF9BB468FED6FB2DFE19.taxon	etymology	Etymology Meander, named after the shape of the gonopods, resembling a meandering river.	en	Johanson, Kjell A., Pham, Hong-Thai (2012): Three new species of Helicopsyche (Trichoptera, Helicopsychidae) from northern Vietnam, with a key to Helicopsyche species of Vietnam. European Journal of Taxonomy 6: 1-10, DOI: 10.5852/ejt.2012.6
03AF87F9527CFF9BB468FED6FB2DFE19.taxon	materials_examined	Material examined Holotype ♂ VIETNAM: Vinh Phuc Prov, Me Linh District, stream near Ngoc Thanh Village, Melinh Station for Biodiversity, 21 ° 23 ' 28.2 " N, 105 ° 42 ' 46.7 " E, 233 m, Malaise trap 4 - 12 Apr. 2011, loc # VN 001, leg. K. A. Johanson, P. H. Thai & T. T. Du (NHRS, alcohol) [DNA voucher IP 7].	en	Johanson, Kjell A., Pham, Hong-Thai (2012): Three new species of Helicopsyche (Trichoptera, Helicopsychidae) from northern Vietnam, with a key to Helicopsyche species of Vietnam. European Journal of Taxonomy 6: 1-10, DOI: 10.5852/ejt.2012.6
03AF87F9527CFF9BB468FED6FB2DFE19.taxon	description	Description MAXILLARY PALPS. Two-segmented, with distal segment slightly longer than basal segment. Pair of interantennal setal warts semi-spherical, elevated. Each scapus about as long as each basal segment of maxillary palps. Spur formula 1,2,4; each foreleg spur slightly shorter than each first tarsomere. LENTH. Forewing 3.7 mm, hind wing 2.8 mm (N = 1). MALE ABDOMEN AND GENITALIA. (Fig. 2) Sternal process VI about one-eighth as long as sternum VI, straight, oriented posteroventrad; slightly tapering in lateral view (Fig. 2 A); pointed in ventral view (Fig. 2 B). Segment IX about 1.5 x taller than long in lateral view (Fig. 2 C); each anterior lobe of segment IX (Fig. 2 C) irregularly narrowing anteriorly in lateral view, asymmetrical, oriented anterad; anterodorsal margin almost straight; anteroventral margin concave (Fig. 2 C); in dorsal view, inner margin forming narrowly ellipsoid cavity (Fig. 2 D); in ventral view, without central posterior process (Fig. 2 E); inner margin forming wide, almost rectangular cavity; lateral apodemes nearly horizontal (Fig. 2 C), each parallel-sided along their length, meeting anterior margin; sub-marginal line present above lateral apodemes; tergal transverse apodemes present, fading immediately above superior appendages; sternal transverse apodeme present along posterior margin of segment IX (Fig. 2 C). Transverse plate absent. Tergum X, in lateral view, oriented posteroventrad (Fig. 2 C), slightly sigmoid, tapering towards mid-length, apex strongly club-shaped, slightly curving dorsally (Fig. 2 C); in dorsal view (Fig. 2 D), deeply and widely divided into pair of parallel-sided and parallel-running branches, with 3 pairs of equally long megasetae in apical group, starting well before apex of gonopods in lateral view (Fig. 2 C). Superior appendages club-shaped (Fig. 2 C), oriented ventrad. Primary branch of gonopods, in lateral view (Fig. 2 C), with basal part oriented posterad before strongly bending dorsad, angling posterad at half-length, apical part produced mesad above tergum X; almost parallel-sided along their length; anterior margin smooth, strongly concave; dorsal and posterior margins undulating. Secondary branch of gonopods absent. Basomesal lobes long, tuboid, each with pair of apical setae; sigmoid in lateral view (Fig. 2 D), parallel-sided with diverging apex in ventral view (Fig. 2 E). Basal plate, in lateral view (Fig. 2 C), curving anteroventrad, narrowing along its length from posterior to anterior end, slightly produced beyond anteroventral margin of segment IX in lateral view (Fig. 2 C); widely triangular in ventral view (Fig. 2 E). Phallic apparatus, lateral view, irregularly tube-shaped along its length (Fig. 2 F), apex about as broad as base; nearly straight after mid-length; in ventral view, with basis narrow (Fig. 2 G); widest after mid-length; endotheca weakly produced, posteroventral part not sclerotized, except single, weakly sclerotized triangular processes distally of phallic sclerite.	en	Johanson, Kjell A., Pham, Hong-Thai (2012): Three new species of Helicopsyche (Trichoptera, Helicopsychidae) from northern Vietnam, with a key to Helicopsyche species of Vietnam. European Journal of Taxonomy 6: 1-10, DOI: 10.5852/ejt.2012.6
03AF87F9527EFF96B46BFED6FAA2F9BA.taxon	description	Fig. 3	en	Johanson, Kjell A., Pham, Hong-Thai (2012): Three new species of Helicopsyche (Trichoptera, Helicopsychidae) from northern Vietnam, with a key to Helicopsyche species of Vietnam. European Journal of Taxonomy 6: 1-10, DOI: 10.5852/ejt.2012.6
03AF87F9527EFF96B46BFED6FAA2F9BA.taxon	diagnosis	Diagnosis Helicopsyche lamnata sp. nov. is most similar to H. vongsombathi Johanson & Malm, 2007, from which it is easily distinguished by the presence of a central posterior process of segment IX and the distal part of the gonopods are more strongly produced posteriorly. The species also resembles H. meander sp. nov., particularly by the nearly club-shaped apex of tergum X in lateral view, but in H. lamnata sp. nov. the VIth sternal process is much longer and the gonopods are less sharply sigmoid.	en	Johanson, Kjell A., Pham, Hong-Thai (2012): Three new species of Helicopsyche (Trichoptera, Helicopsychidae) from northern Vietnam, with a key to Helicopsyche species of Vietnam. European Journal of Taxonomy 6: 1-10, DOI: 10.5852/ejt.2012.6
03AF87F9527EFF96B46BFED6FAA2F9BA.taxon	etymology	Etymology Lamnata, plate-like in Latin, refers to the process on the posterior margin of sternite IX.	en	Johanson, Kjell A., Pham, Hong-Thai (2012): Three new species of Helicopsyche (Trichoptera, Helicopsychidae) from northern Vietnam, with a key to Helicopsyche species of Vietnam. European Journal of Taxonomy 6: 1-10, DOI: 10.5852/ejt.2012.6
03AF87F9527EFF96B46BFED6FAA2F9BA.taxon	materials_examined	Material examined Holotype ♂ VIETNAM: Vinh Phuc Prov, Me Linh District, stream near Ngoc Thanh Village, Melinh Station for Biodiversity, 21 ° 23 ' 28.2 " N, 105 ° 42 ' 46.7 " E, 233 m, Malaise trap 4 - 12 Apr. 2011, loc # VN 001, leg. K. A. Johanson, P. H. Thai & T. T. Du (NHRS, alcohol) [DNA voucher IP 9]. Paratype Same data as holotype, except (IEBR, alcohol).	en	Johanson, Kjell A., Pham, Hong-Thai (2012): Three new species of Helicopsyche (Trichoptera, Helicopsychidae) from northern Vietnam, with a key to Helicopsyche species of Vietnam. European Journal of Taxonomy 6: 1-10, DOI: 10.5852/ejt.2012.6
03AF87F9527EFF96B46BFED6FAA2F9BA.taxon	description	Description MAXILLARY PALPS. Two-segmented, all segments sub-equal in length. Pair of interantennal setal warts very large, spherical, elevated, densely covered by long setae. Each scapus slightly longer than individual segments of maxillary palps. Spur formula 1,2,4; each foreleg spur nearly as long as each first tarsomere. LENGTH. Forewing 4.0 mm, hind wing 3.1 mm (N = 1). MALE ABDOMEN AND GENITALIA. (Fig. 3) Sternal process VI about two-fifths as long as sternum VI, straight, oriented posteroventrad; parallel-sided in lateral view (Fig. 3 A); almost parallel-sided in ventral view (Fig. 3 B). Segment IX slightly taller than long in lateral view (Fig. 3 C); each anterior lobe of segment IX (Fig. 3 C) smoothly narrowing anteriorly in lateral view, asymmetrical, oriented anterodorsad; anterodorsal margin almost straight; anteroventral margin concave (Fig. 3 C); in dorsal view, inner margin forming narrowly ellipsoid cavity (Fig. 3 D); large central posterior process located below basis of gonopods, sharply triangular in lateral view (Fig. 3 C); in ventral view widely trapezoid, with slightly concave posterior margin. Lateral apodemes nearly horizontal (Fig. 3 C), parallel-sided along their length, located at mid-height of segment IX, ending in sub-marginal line; tergal transverse apodemes absent; sternal transverse apodeme oriented in right angle to lateral apodeme, located along posterior margin of segment IX (Fig. 3 C). Transverse plate absent. Tergum X, in lateral view, oriented posteroventrad (Fig. 3 C), strongly tapering towards mid-length, with single, dorsolateral, stout seta at mid-length; distal half almost parallel-sided, apex slightly club-shaped, orienting posteroventrally (Fig. 3 C); in dorsal view (Fig. 3 D), deeply and narrowly divided into pair of parallel-sided and parallelrunning branches; each with 4 pairs equally long megasetae in apical group, and two pairs stout, posteradoriented spines (Fig. 3 C). Superior appendages club-shaped (Fig. 3 C), oriented posteroventrad. Primary branch of gonopods, in lateral view (Fig. 3 C), with basal part oriented posterad before strongly bending posterodorsad, curving posterad after mid-length, apical part rounded in dorsal view; produced mesad into pointed plates above tergum X in dorsal view (Fig. 3 D); slightly widening along their length; basal part of anterior margin smooth; dorsal and posterior margins undulating. Secondary branch of gonopods absent. Basomesal lobes long, slightly broadening before apex, each with pair of apical setae; straight in lateral view (Fig. 3 C), diverging in ventral view (Fig. 3 E). Basal plate, in lateral view (Fig. 3 C), straight, strongly narrowing anteriorly towards mid-length, anterior half very slender, not produced beyond anteroventral margin of segment IX in lateral view (Fig. 3 C); almost parallel-sided in ventral view (Fig. 3 E), with rounded anterior apex. Phallic apparatus, lateral view, irregularly tube-shaped along its length (Fig. 3 F), apex about as broad as base; nearly straight after mid-length; irregular in ventral view (Fig. 3 G); widest after mid-length; endotheca weakly produced, posteroventral part not sclerotized, except long, single, weakly sclerotized triangular processes distally of phallic sclerite.	en	Johanson, Kjell A., Pham, Hong-Thai (2012): Three new species of Helicopsyche (Trichoptera, Helicopsychidae) from northern Vietnam, with a key to Helicopsyche species of Vietnam. European Journal of Taxonomy 6: 1-10, DOI: 10.5852/ejt.2012.6
