identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03AD879AF867FFFFFF63FD6D51D4F84E.text	03AD879AF867FFFFFF63FD6D51D4F84E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leptopus riparius Hsiao 1964	<div><p>Leptopus riparius Hsiao, 1964</p><p>Leptopus riparius Hsiao 1964: Acta Zootax. Sinica 1: 289.</p><p>(Figs. 1, 6, 11, 13–15, 29, 34–36, 51, 52, 62)</p><p>Material examined. Holotype: 1 male, CHINA, Sichuan Prov., Leshan city, 22 July 1957, Hao Xu leg. (NKUM) . Paratypes: 1 male, 1 female, CHINA, Sichuan Prov., same data as holotype (NKUM) . Non-type specimens: 2 males, 2 females, CHINA, Hubei Prov., Xianning city, Tongshan county, Jiugongshan mountain, Yinhegu valley, 13 July 2022, Zhen Ye leg. (NKUM); 3 males, 1 female, CHINA, Zhejiang Prov., Ningbo City, Jiufengshan mountain, 11 November 2022, Danyang Zhou leg. (NKUM). 1 female, VIETNAM, Hai Hung Prov., Kheao stream, 23 km. N. of Chi Linh, 125 m., water temp. 17° C., 25 March 2000, CL 4376, J. T. Polhemus and P. Nguyen (BPBM) .</p><p>Diagnosis. Body length 3.7–4.1 (holotype: 4.0), ground color mainly brown (Figs. 1, 6, 11). Head mainly black, clypeus and bucculae yellow; labium short, labiomere I completely hidden beneath bucculae, labiomere II yellowish-white, with four (2+2) spines, labiomere III yellowish-brown to dark-brown, with four (2+2) spines, labiomere IV black; compound eyes distinctly large, with sparse short, spine-like setae; venter of head with three spines beneath each compound eye (Figs. 13–15). Pronotum densely punctured, mainly dark-brown, medially with two yellowish marks, lateral and posterior margins yellowish; scutellum black with a yellowish apex (Figs. 1, 6, 11). Hemelytra mainly brown, with two dark spots on each corium (Fig. 29). Fore femur incrassate, ventrally with 9–10 large spines (i.e., five large spines on outer side and 4–5 on inner side) and two rows of short spines; fore tibia with two rows of spines ventrally, each row with 2–3 large spines (Figs. 34–36). Paramere apically extended with several long hairs (Figs. 51, 52).</p><p>Comparative notes. The differences between L. riparius and L. scitulus Drake, 1954 were noted by Hsiao (1964) (i.e., L. riparius is larger, having a longer scutellum and bearing fewer long spines on the fore tibia). This species is also similar to L. marmoratus Goeze, 1778 due to the body shape and absence of dark fasciae on the hemelytra. However, L. riparius differs from L. marmoratus by the following characteristics: in L. riparius, the dorsal coloration is mainly uniformly brown, without scattered black flecks or patches on the hemelytra, and the femora lack brown annuli (Figs. 1, 6, 11), whereas in L. marmoratus, the hemelytra bear scattered black flecks and patches, the membrane has a distinct black spot basally, and the sub-apical parts of the femora are annulated with brown (Wachmann et al. 2006; Khazaei et al. 2020, see color habitus Fig. 10 in this latter work).</p><p>Remarks. The specimen we report from Vietnam is a female, so it is not possible make a definitive comparison on the basis of the male genitalia, and the identification must therefore be considered provisional. Even so, the specimen falls within the current concept of L. riparius on the basis of body size and shape, coloration, and the basal expansion of the hemelytral embolium. Our recent collections extend the distribution of genus Leptopus Latreille, 1809 eastward into Zhejiang Province, China (approximately 121° E) and southward into Vietnam (new country record) (Fig. 61).</p><p>Distribution: China (Sichuan, Hubei, Zhejiang) (Hsiao, 1964; new records in Hubei and Zhejiang), Vietnam (new record, Fig. 61).</p></div>	https://treatment.plazi.org/id/03AD879AF867FFFFFF63FD6D51D4F84E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Li, Zihe;Jin, Zezhong;Polhemus, Dan A.;Ye, Zhen	Li, Zihe, Jin, Zezhong, Polhemus, Dan A., Ye, Zhen (2023): Two new species of Valleriola (Hemiptera: Heteroptera: Leptopodidae) and taxonomic notes on the tribe Leptopodini Brullé, 1836 from East and Southeast Asia. Zootaxa 5256 (4): 329-344, DOI: 10.11646/zootaxa.5256.4.2, URL: http://dx.doi.org/10.11646/zootaxa.5256.4.2
03AD879AF865FFF6FF63FAC1557AFEEA.text	03AD879AF865FFF6FF63FAC1557AFEEA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Valleriola buenoi (Usinger 1942)	<div><p>Valleriola buenoi (Usinger, 1942)</p><p>Erianotus buenoi Usinger 1942: Bull. Brooklyn Entomol. Soc. 37: 103–106.</p><p>Valleriola buenoi: Drake &amp; Hottes 1951, J. Kansas Entomol. Soc. 24: 21–22.</p><p>(Figs. 3, 8, 12, 19–21, 31, 40–42, 55, 56)</p><p>Material examined. 2 males, 2 females, PHILIPPINES, Luzon, Aurora, Canile Trail, Bayan ng Maria, 13 December 2019, Zhen Ye leg. (NKUM) .</p><p>Diagnosis. Body length 4.5–4.8, ground color mainly black, with yellowish spots and stripes dorsally (Figs. 3, 8, 12). Head mainly black, clypeus and bucculae yellow; labium short, labiomere I completely hidden beneath bucculae, labiomere II yellowish-white, with four (2+2) spines, labiomeres III and IV black; compound eyes distinctly large, without sparse short, spine-like setae; venter of head with three spines beneath each compound eye (Figs. 19–21). Pronotum densely punctured, mainly black to blackish-brown, posterior margin yellowish; scutellum blackish with yellowish apex (Figs. 3, 8, 12). Hemelytra mainly blackish-brown; clavus medially with a yellowish stripe, apically with a small, yellowish spot; corium with a sub-basal, yellowish spot in cell Sc, usually extend over R+M vein (Fig. 31). Fore femur incrassate, ventrally with six large spines (three spines on outer side and three on inner side) and two rows of dense, small, black spines (Figs. 40–42). Paramere stick-like, simple, apically with dense punctures and interiorly with erect hairs (Figs. 55, 56).</p><p>Comparative notes. Valleriola buenoi is similar to V. javanica, but can be distinguished by the following characters: antennomeres III and IV are shorter (Drake &amp; Hottes 1951); the median part of the pronotum lacks yellowish marks (Figs. 3, 8, 12); the stripe on the clavus does not reach the apex of the clavus (Fig. 31); the fore femur has three large spines on the inner side, with one of these distinctly displaced from the inner row of spines (Figs. 40–42); and the paramere is relatively simple and elongate, without the prominent apical hook characteristic of V. javanica (Figs. 55, 56). In contrast, in V. javanica, antennomeres III and IV are much longer; the median part of the pronotum bears a yellowish spot (Figs. 5, 10); the stripe on the clavus is relatively long, reaching the apex of the clavus (Fig. 33); the fore femur bears four large spines forming a regular row on the inner side (Figs. 46–48); and the paramere is strongly hooked apically (Fig. 60).</p><p>Remarks. We checked arrangement of spines on the fore femur of Valleriola and found that V. asiatica sp. nov., V. javanica and V. bui sp. nov. possess seven large spines on the ventral side (three spines on the inner side and four on the outer side) (Figs. 37–39, 43–48), whereas V. buenoi has only six large spines on the ventral side (three spines on the inner side and three on the outer side), with one large spine on the inner row distinctly displaced from the others (Figs. 40–42).</p><p>Distribution: China (Hainan) (Drake &amp; Maa, 1954), Philippines (Luzon) (Usinger, 1942) (Fig. 61).</p></div>	https://treatment.plazi.org/id/03AD879AF865FFF6FF63FAC1557AFEEA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Li, Zihe;Jin, Zezhong;Polhemus, Dan A.;Ye, Zhen	Li, Zihe, Jin, Zezhong, Polhemus, Dan A., Ye, Zhen (2023): Two new species of Valleriola (Hemiptera: Heteroptera: Leptopodidae) and taxonomic notes on the tribe Leptopodini Brullé, 1836 from East and Southeast Asia. Zootaxa 5256 (4): 329-344, DOI: 10.11646/zootaxa.5256.4.2, URL: http://dx.doi.org/10.11646/zootaxa.5256.4.2
03AD879AF86EFFF7FF63FA805200FE02.text	03AD879AF86EFFF7FF63FA805200FE02.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Valleriola javanica Drake & Hottes 1951	<div><p>Valleriola javanica Drake &amp; Hottes, 1951</p><p>Valleriola javanica Drake &amp; Hottes 1951: J. Kansas Entomol. Soc. 24: 22–24.</p><p>(Figs. 5, 10, 25–27, 33, 46–48, 60, 64)</p><p>Material examined. 3 males, 1 female, CHINA, Guangxi Prov., Bobai county, Tianchi Canyon, 28, July, 2019, Zhen Ye leg. (NKUM); 2 males, 2 females, CHINA, Hainan Prov., Baisha county, Yinggeling Nature Reserve, 21 July 2013, Qiang Xie leg. (NKUM); 1 male, CHINA, Hainan Prov., Wuzhishan city, Shuiman village, 10 December 2019, Zhen Ye leg. (NKUM); 1 male, 1 female, CHINA, Taiwan Prov., Pingdong county, Mudan village, 17 June 2013, Zhen Ye leg. (NKUM); 1 female, CHINA, Yunnan Prov., Mengla county, Mengman town, 28 April 2011, Rui Wang leg. (NKUM). 2 males, MALAYSIA, Pahang, Sg. Pua Waterfall, 20 April 2019, Zhen Ye leg. (NKUM); 1 male, LAOS, Vientiane, Vang Veihg, Kaeng Yui Waterfall, 15 November 2018, Zhen Ye leg. (NKUM).</p><p>Diagnosis. Body length 4.8–5.3, ground color mainly blackish-brown, dorsally with yellowish spots and stripes (Figs. 5, 10). Head mainly black, with a distinct orange mark behind ocelli; clypeus and bucculae yellowish; labium short, labiomere I completely hidden beneath bucculae, labiomere II yellowish-white, with four (2+2) spines, labiomere III yellowish-brown, labiomere IV dark-brown to black; compound eyes distinctly large, without sparse, short, spine-like setae, venter of head with three spines beneath each compound eye (Figs. 25–27). Pronotum densely punctured, mainly dark-brown, posterior margin yellowish, median part with a yellowish spot; scutellum black with a yellowish apex (Figs. 5, 10). Hemelytra mainly dark-brown, clavus medially with a long, yellowish stripe, reaching the apex of the clavus and merging with the apical spot (Fig. 33). Fore femur incrassate, with seven long spines (three spines on the outer side and four on the inner side) and two rows of dense, small, black spines (Figs. 46–48). Apex of paramere hook-like, bearing long hairs but without punctures (Fig. 60).</p><p>Comparative notes. Valleriola javanica is most similar to V. bui sp. nov. and V. buenoi; see comparative notes under those species.</p><p>Remarks. This species is widely distributed in Southeast Asia and China and is here newly reported from Laos.</p><p>Distribution: J. Polhemus &amp; D. Polhemus (2012) listed the range of this species as including Indonesia, Malaysia, Thailand, Burma and China (Hong Kong). To this we add Laos and additional provinces in China (Yunnan, Hainan, Guangxi, and Taiwan) (Fig. 61).</p></div>	https://treatment.plazi.org/id/03AD879AF86EFFF7FF63FA805200FE02	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Li, Zihe;Jin, Zezhong;Polhemus, Dan A.;Ye, Zhen	Li, Zihe, Jin, Zezhong, Polhemus, Dan A., Ye, Zhen (2023): Two new species of Valleriola (Hemiptera: Heteroptera: Leptopodidae) and taxonomic notes on the tribe Leptopodini Brullé, 1836 from East and Southeast Asia. Zootaxa 5256 (4): 329-344, DOI: 10.11646/zootaxa.5256.4.2, URL: http://dx.doi.org/10.11646/zootaxa.5256.4.2
03AD879AF86DFFF3FF63FAB453E8FE26.text	03AD879AF86DFFF3FF63FAB453E8FE26.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Valleriola asiatica Li, Polhemus & Ye 2023	<div><p>Valleriola asiatica Li, Polhemus &amp; Ye, sp. nov.</p><p>(Figs. 2, 7, 16–18, 30, 37–39, 53, 54, 65)</p><p>Material examined. Holotype: male, CHINA, Hainan Prov., Bawangling Nature Reserve, 28 July 2017, Zhen Ye leg. (NKUM); Paratypes: 3 males, 3 females, CHINA, Hainan Prov., same data as holotype (NKUM); 3 males, 3 females, LAOS, Attapu, Sae Pra Waterfall, 18 August 2019, Yanfei Li leg. (NKUM). 1 male, VIETNAM, Lam Dong Prov., stream below Suõi Vàng hydro project (= Gold Stream), 16 km. N. of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.36833&amp;materialsCitation.latitude=11.990556" title="Search Plazi for locations around (long 108.36833/lat 11.990556)">Dalat</a>, 1430 m., 11°59'26"N, 108°22'06"E, 9 May 1998, CL 3078, J. T. Polhemus (USNM); 7 males, 6 females, VIETNAM, Lam Dong Prov., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.265274&amp;materialsCitation.latitude=11.688611" title="Search Plazi for locations around (long 108.265274/lat 11.688611)">Pongour Falls</a>, 62 km. SW of Dalat, 825 m., 11°41'19"N, 108°15'55"E, 13–16 May 1998, CL 3091, J. T. Polhemus (BPBM, USNM); 1 male, 1 female, VIETNAM, Dong Nai Prov., Cat Tien National Park, Dong Nai River, cascade upstream of park HQ, 110 m., 7 May 1998, CL 3073, J. T. Polhemus (USNM); 1 male, 3 females, VIETNAM, Dong Nai Prov., Cat Tien National Park, Dong Nai River, cascade area behind first ranger station, 120 m., 8 May 1998, CL 3077, J. T. Polhemus (BPBM, USNM); 3 males, 7 females, VIETNAM, Lai Châu Prov., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.1925&amp;materialsCitation.latitude=22.147778" title="Search Plazi for locations around (long 103.1925/lat 22.147778)">Nam Na River and Nam Ceung stream trib</a>. 15.5 km. N. of Lai Châu, 200 m., 22°08'52"N, 103°11'33"E, 11 April 2000, 10:00– 13:00 hrs., CL 4409, D. A. Polhemus, J. T. Polhemus and P. Nguyen (BPBM, USNM); 10 males, 11 females, 1 immature, VIETNAM, Gia Lai Prov., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.52473&amp;materialsCitation.latitude=13.767222" title="Search Plazi for locations around (long 108.52473/lat 13.767222)">Ya Rung waterfall</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.52473&amp;materialsCitation.latitude=13.767222" title="Search Plazi for locations around (long 108.52473/lat 13.767222)">Ngoe Ba Riv.</a>, 30 km. S. of An Khe, 345 m., 13°46'02"N, 108°31'29"E, 10 March 2001, CL 4287, J. T. Polhemus &amp; P. Nguyen (BPBM, USNM); 2 males, 4 females, 1 immature, CAMBODIA, Kampong Spoe Prov., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.09028&amp;materialsCitation.latitude=12.412499" title="Search Plazi for locations around (long 104.09028/lat 12.412499)">An Long Rot river</a> at granite rapids, ~ 4.5 km. NW of Chambok village, 95 m., 12°24'45"N, 104°05'025"E, 21 May 2016, 11:00–12:15 hrs., CL 6034, D. A. Polhemus (BPBM).</p><p>Diagnosis. Body length: 6.04–6.50, ground color black with yellowish spots and stripes dorsally. Venter of head and labiomere II with dense, long, white hairs. Femora yellowish with black spots and stripes. Parameres slightly twisted, apical part spoon-like, with long, straight hairs.</p><p>Comparative notes. Valleriola asiatica sp. nov. is the largest species of Leptopodini known from Southeast Asia, with the body length of both sexes exceeding 6.0 mm. In comparison to the similar V. cicindeloides from India, V. asiatica sp. nov. may be distinguished by its darker grey ground color, and a shorter antennal segment III in relation to segment II, the ratio of segment II/segment III being 1.00/2.00– 2.20 in V. asiatica sp. nov., versus 1.00/3.00 in V. cicindeloides . Based on figures made by the late J. T. Polhemus of the holotype of V. cicindeloides, housed in the Natural History Museum, London, the male parameres of the two species are rather similar in form, including the presence of a small, angular projection on the basal third of the paramere when seen in inner ventral view. The third author (DP) has in his collection additional species of similar size and appearance to V. asiatica sp. nov. from the Indonesian islands of Timor, Sumba, Celebes and Sumatra, as well as Nepal. All of these examples have relatively large body sizes (over 5.5 mm); dark-brown to dark-blackish-grey dorsal ground coloration; a pair of pale spots arranged horizontally across the posterior portion of the corium on each hemelytron; and the legs and lateral pronotum set with numerous erect, pale setae that are much longer than the diameters of the leg segments on which they arise. Within these populations, however, there are geographically correlated differences in paramere shape in comparison to the Indochinese material, and they are therefore not presently considered conspecific. Instead, they will be dealt with in a separate work already partially completed.</p><p>Description. Male. Color: ground color black (Figs. 2, 7). Head mainly black, clypeus yellowish with black marks; bucculae yellowish; labiomere I completely hidden beneath bucculae, labiomere II yellowish-white with black apex, labiomeres III–IV black; compound eyes and ocelli black to dark-red; antennomere I yellowish with black stripes, antennomeres II–IV dark-brown (Figs. 2, 7, 16–18). Pronotum mainly blackish-brown, collar and callus black, median part of collar with a yellow spot, posterior lobe of pronotum blackish-brown, posterior margin yellowish, median part with a yellowish stripe, posterior angles of pronotum shinning black; scutellum blackish, with a yellowish apex (Figs. 2, 7). Hemelytra mainly blackish-brown, clavus with a yellowish stripe in middle and a yellowish spot on apex, lateral margin of hemelytra pale, spots and stripes on corium and membrane yellowish, middle part of corium with two spots on both sides of R+M vein, sometimes the outer one absent; apical part of sc cell and membrane cell IV with yellowish spots, sometimes with a small spot in r cell (Fig. 30). Legs yellowish, femora with black stripe and spots; tibiae and tarsi blackish-brown, except fore tibia and tarsus yellowish-brown (Figs. 2, 37–39). Abdominal segments black, with white sutures. Paramere yellowish, infuscated apically (Figs. 53, 54). Structural characteristics: body elongated, bearing sparse, short, pale setae dorsally (Figs. 2, 7). Head short and wide, dorsally bearing dense, short pubescence except two circular areas, venter of head bearing dense, long, white hairs, with three spines beneath compound eyes on each side; compound eyes distinctly large, without sparse, short, spine-like setae; bucculae clearly visible; labium short, labiomere I completely hidden beneath bucculae, labiomere II bearing dense, long, white hairs, with two spines on each side; antennomere I short and stout, antennomeres II–IV extremely long and slender (Figs. 2, 7, 16–18). Pronotum sub-triangular, densely punctured except callus; callus relatively rough, distinctly swollen, with a deep impression on median line; posterior angles of pronotum glossy, without punctures; scutellum triangular, medially with a sub-circular depression, laterally serrated, with a distinct process on each side (Figs. 2, 7). Hemelytra elongated, clavus and outer cell of corium with dense punctures; membrane opaque, with four closed cells (Fig. 30). Legs long and slender, fore coxa strong, slightly elongated, with two long spines anteriorly directed; fore femur incrassate, ventrally with seven large spines (three spines on the outer side and four on the inner side) and two rows of dense, small, black spines; fore tibia ventrally with a row of dense, brown spines (Figs. 37–39). Middle and hind legs distinctly elongated. Abdomen slender, tubular-like. Paramere slightly twisted, with a spoon-like apex, covered by dense punctures and long, straight hairs (Figs. 53, 54).</p><p>Female. Color and structural characteristics similar to male. Genital segments of female hidden in abdominal segment VII, not visible in vitro.</p><p>Measurements. Male. Body length: 6.26–6.50 (holotype: 6.50), body width across hemelytra: 1.70–1.84. Head width: 1.31–1.40, head length: 0.78–0.83, head width about 1.69 times head length; antenna about 1.05 times as long as body, length of antennomeres I–IV: 0.41, 1.36, 2.69, 2.38. Pronotum width across collar: 0.57–0.59, pronotum width across posterior angles: 1.63–1.64, pronotum length: 1.24–1.29; scutellum width: 0.74–0.77, scutellum length: 0.68–0.72; length of femur, tibia and tarsus (tarsomeres I+II+III): fore leg: 2.28, 1.69 and 0.60 (0.16+0.24+0.20), middle leg: 2.44, 2.26 and 0.62 (0.16+0.21+0.25), hind leg: 2.57, 3.52 and 0.82 (0.14+0.31+0.37). Female. Body length: 6.04–6.35, body width across hemelytra: 1.67–1.79. Head width: 1.32–1.35, head length: 0.77–0.82, head width about 1.06 times head length; antenna about 1.18 times as long as body, length of antennomeres I–IV: 0.37, 1.21, 2.74, 2.42. Pronotum width across collar: 0.59–0.62, pronotum width across posterior angles: 1.61–1.68; pronotum length: 1.20–1.26; scutellum width: 0.78–0.81, scutellum length: 0.75–0.79; length of femur, tibia and tarsus (tarsomeres I+II+III): fore leg: 2.21, 1.66 and 0.57 (0.13+0.26+0.18), middle leg: 2.50, 2.43 and 0.69 (0.14+0.33+0.22), hind leg: 2.66, 3.66 and 0.77 (0.17+0.36+0.24).</p><p>Etymology. The specific epithet “ asiatica ” refers to the wide distribution of this species in Southeast Asia; an adjective.</p><p>Distribution. China (Hainan), Laos, Vietnam, Cambodia (Fig. 61).</p></div>	https://treatment.plazi.org/id/03AD879AF86DFFF3FF63FAB453E8FE26	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Li, Zihe;Jin, Zezhong;Polhemus, Dan A.;Ye, Zhen	Li, Zihe, Jin, Zezhong, Polhemus, Dan A., Ye, Zhen (2023): Two new species of Valleriola (Hemiptera: Heteroptera: Leptopodidae) and taxonomic notes on the tribe Leptopodini Brullé, 1836 from East and Southeast Asia. Zootaxa 5256 (4): 329-344, DOI: 10.11646/zootaxa.5256.4.2, URL: http://dx.doi.org/10.11646/zootaxa.5256.4.2
03AD879AF86BFFF0FF63FDC05220FBE6.text	03AD879AF86BFFF0FF63FDC05220FBE6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Valleriola bui Li, Jin & Ye 2023	<div><p>Valleriola bui Li, Jin &amp; Ye, sp. nov.</p><p>(Figs. 4, 9, 22–24, 32, 43–45, 49, 50, 57–59, 63)</p><p>Meterial examined. Holotype: male, LAOS, Vientiane, Vang Veihg, Kaeng Yui Waterfall, 15 November 2018, Zhen Ye leg. (NKUM) . Paratypes: 1 male, 2 females, LAOS, Vientiane, same data as holotype (NKUM); 3 males, 3 females, THAILAND, Nong Bua Lamphu, Na Klang, 13 August 2018, Zhen Ye leg. (NKUM) .</p><p>Diagnosis. Body length: 4.75–4.90, ground color dark-brown with yellowish spots and stripes dorsally. Head mainly yellowish except the frons and the posterior area around compound eyes black. Abdominal segments black in male but yellowish-white in female.</p><p>Comparative notes. Valleriola bui sp. nov. appears to be closely related to V. javanica . However, V. bui sp. nov. can be distinguished by its remarkable coloration of head: most of the head of V. bui sp. nov. is yellowish except the black frons and the posterior area around compound eyes, whereas the head of V. javanica is mainly blackish and only the clypeus and post-ocellar spot are yellowish (Figs. 22–27). Moreover, V. bui sp. nov. has other characteristics different from V. javanica: the stripes and spots on the sub-basal area of the hemelytra are relatively ambiguous, without distinct margins as in V. javanica (Fig. 32, 33); the stripe on the clavus is isolated and not merged with the apical spots (Fig. 32); the paramere is mushroom-like and apically extended with a small process on the inner side, without hook-like structure (Figs. 57–59).</p><p>Description. Male. Color: ground color dark-brown (Figs. 4, 9). Head mainly yellowish, clypeus orange, venter of head yellowish, frons and posterior area around compound eyes black, area behind ocelli with an orange spot; bucculae yellowish; labiomere I completely hidden beneath bucculae, labiomere II yellowish-white, labiomere III yellowish-brown, labiomere IV brown to blackish-brown; compound eyes and ocelli silvery; antennomere I yellowish, antennomeres II–IV dark-brown (Figs. 4, 9, 22–24). Pronotum mainly dark-brown, collar and callus blackish, median part of collar with an orange spot, merged with the spot behind ocelli, posterior lobe of pronotum dark-brown, posterior margin yellowish, median part with a yellowish stripe, posterior angles of pronotum shinning black; scutellum blackish, with a yellowish apex (Figs. 4, 9). Hemelytra mainly brown, margins of clavus yellowishbrown, clavus with a yellowish stripe in middle and a spot on apex, lateral margin of hemelytra pale, spots and stripes on corium and membrane pale yellowish, middle part of corium with a spot on the inner side of R+M vein, r cell and membrane cell IV with pale yellowish spots (Fig. 32). Legs yellowish, fore femur with a wide, brown stripe on outer side; all tibiae and tarsi infuscated (Figs. 4, 43–45). Abdominal segments black, with white sutures (Fig. 49). Paramere yellowish, with a dark-brown apex (Figs. 57–59). Structural characteristics: body elongated, bearing sparse, short, pale setae (Figs. 4, 9). Head short and wide, dorsally bearing dense, short pubescence except two circular areas, venter of head with three spines beneath each compound eye; compound eyes distinctly large, without sparse, short, spine-like setae; bucculae clearly visible; labium short, labiomere I completely hidden beneath bucculae, labiomere II with two spines on each side; antennomere I short and stout, antennomeres II–IV extremely long and slender (Figs. 4, 9, 22–24). Pronotum sub-triangular, densely punctured except callus; callus relatively rough, distinctly swollen, with a deep impression on median line; posterior angles of pronotum glossy, without punctures; scutellum triangular, medially with a sub-circular depression, laterally serrated, with a distinct process on each side (Figs. 4, 9). Hemelytra elongated, clavus and sc cell of corium with dense punctures; membrane opaque, with four closed cells (Fig. 32). Legs long and slender, fore coxa strong, slightly elongated, with two long spines anteriorly directed; fore femur incrassate, ventrally with seven large spines (three spines on outer side and four on inner side) and two rows of dense, small, black spines (Figs. 43–45); fore tibia ventrally with a row of dense, brown spines. Middle and hind legs slender, distinctly elongated. Abdomen slender, tubular-like (Fig. 49). Paramere small, apically with a mushroom-like extension, covered by dense punctures and long, curved hairs, with a spine-like process on inner side (Figs. 57–59).</p><p>Female. Color and structural characteristics similar to male, but abdominal segments of female are yellowishwhite except abdominal segment VII mainly brown (Fig. 50). Genital segments of female hidden in abdominal segment VII, not visible in vitro.</p><p>Measurements. Male. Body length: 4.75–4.90 (holotype: 4.80), body width across hemelytra: 1.25–1.35. Head width: 1.25–1.30, head length: 0.65–0.68, head width about 1.92 times head length; antenna about 1.31 times as long as body, length of antennomeres I–IV: 0.38, 0.90, 1.90, 2.88. Pronotum width across collar: 0.49–0.51, pronotum width across posterior angles: 1.15–1.20, pronotum length: 0.98–1.00; scutellum width: 0.53–0.56, scutellum length: 0.54–0.55; length of femur, tibia and tarsus (tarsomeres I+II+III): fore leg: 1.85, 1.48 and 0.42 (0.06+0.23+0.13), middle leg: 1.88, 2.10 and 0.50 (0.09+0.25+0.16), hind leg: 2.30, 3.00 and 0.52 (0.10+0.23+0.19). Female. Body length: 4.90–5.00, body width across hemelytra: 1.30–1.35. Head width: 1.29–1.31, head length: 0.63–0.64, head width about 2.06 times head length; antenna about 1.18 times as long as body, length of antennomere I–IV: 0.38, 0.86, 1.85, 2.75. Pronotum width across collar: 0.50–0.54, pronotum width across posterior angles: 1.20–1.30; pronotum length: 0.98–1.05; scutellum width: 0.56–0.60, scutellum length: 0.55–0.56; length of femur, tibia and tarsus (tarsomeres I+II+III): fore leg: 1.85, 1.48 and 0.38 (0.09+0.16+0.13), middle leg: 1.98, 2.10 and 0.53 (0.09+0.24+0.20), hind leg: 1.01, 1.60 and 0.57 (0.09+0.25+0.23).</p><p>Etymology. This species is named in honor of Prof. Wenjun Bu for his outstanding contributions to the study of taxonomy, phylogeny and biogeography of Heteroptera, on the occasion of his 60th birthday.</p><p>Remarks. Valleriola bui sp. nov. is sexually dimorphic in appearance, with the male having a black abdomen whereas that of females is yellowish-white (Figs. 49, 50). The third author (DP) has seen additional undescribed species from Thailand and Vietnam, to be described in a forthcoming work, that also share this interesting sexual dimorphism in regard to abdominal pigmentation.</p><p>Distribution. Laos, Thailand (Fig. 61).</p></div>	https://treatment.plazi.org/id/03AD879AF86BFFF0FF63FDC05220FBE6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Li, Zihe;Jin, Zezhong;Polhemus, Dan A.;Ye, Zhen	Li, Zihe, Jin, Zezhong, Polhemus, Dan A., Ye, Zhen (2023): Two new species of Valleriola (Hemiptera: Heteroptera: Leptopodidae) and taxonomic notes on the tribe Leptopodini Brullé, 1836 from East and Southeast Asia. Zootaxa 5256 (4): 329-344, DOI: 10.11646/zootaxa.5256.4.2, URL: http://dx.doi.org/10.11646/zootaxa.5256.4.2
