taxonID	type	description	language	source
03AD87B9FF80FFFEBAE44522DC5EFDBD.taxon	description	Types. Holotype male, ARGENTINA, Province of Misiones, Pindapoy, X. 1935 (MLPA). Paratypes: two males, three females and six specimens with unknown gender, same data as holotype (MLPA); two specimens with unknown gender, Province of Misiones, X. 1900, C. Bruch col (MACN). Description. Measurements, respectively male / female: body length, 3.5 – 3.7 / 4.0 – 4.2 mm; elytral length 2.7 – 2.8 / 3.1 – 3.2 mm; humeral width 1.5 / 1.6 – 1.7 mm. General coloration brown. Body densely covered by short, decumbent, gold pubescence. Head with compact punctation permeated by dense and uniform pilosity; strongly evident longitudinal sulcus reaching from vertex to clypeus; clypeus subtriangular, somewhat shiny, with a row of hairs close to margin and ventrally directed. Antennal tubercles weakly evident, subcontiguous, separated by longitudinal groove; surface subopaque. Labrum broader than long, shiny, with transversal row of four long hairs ventrally directed; apical margin round. Eyes globose, greatest diameter approximately 2.5 x length of gena; erect bristle close to the superior internal margin. Interocular distance equal to about three times space between antennal insertions. Antennae 11 - segmented, brown to dark-brown, except for base of scape, pedicel and antennomeres III to VII yellow-testaceous; pubescence uniform, interspersed by a few larger bristles mainly placed at apex of each antennomere. Scape subconical, enlarged towards apex; antennomere III largest, approximately twice as long as subglobose pedicel; antennomere IV slightly shorter than III; subsequent antennomeres subequal in shape and length, some enlarged if compared to previous antennomeres. Apical antennomere conical. Thorax subrectangular with subcircular lateral margins; greatest width close to the middle, approximately 1.6 x length. Pronotum punctate, interspersed by short pubescence, with the following distribution of depressions: (a) one deeper on each side of middle, not reaching lateral margins, and (b) two shallower, one close to basal margin and another slightly deeper than basal one, subcontiguous with apical border, but not reaching it. Elytra with subparallel sides, brown (in some specimens, humeral region and posterior tubercles darker brown), except for an irregular half-moon-shaped yellow spot posteriorly directed in apical half, this spot extending from region next to lateral margin to base of disc. Elytral surface densely punctate, with punctation more conspicuous on disc right below basal tubercles. Pubescence short, decumbent, irregular, forming in some regions circular patterns similar to swirls; hair color variable, affected by light incidence. Each elytron with the following distribution of tubercles (Fig. 1): (a) two anterior, one on humeral region, the second next to the first, but round and developed, not reaching scutellar region; (b) two posterior at apical third: one round and laterally positioned, and the other, next to it, taller and subtriangular, more evident when observed laterally or posteriorly (Figs 2, 3). Elytral depression only on region where there is a yellow sublunar spot. Intermediate legs of males with a small spur on apex of tibiae; females not bearing spurs. curved ventrally (Fig. 11). Apex of median lobe subtruncated (Fig. 12). Tegmen (Fig. 13) hastiform, bifurcated near apex forming two divergent arms, each arm curved backwards.	en	Moura, Luciano De A. (2016): New species, new combinations and synonymies of Neotropical Galerucini (Coleoptera, Chrysomelidae, Galerucinae). Zootaxa 4066 (1): 63-70, DOI: 10.11646/zootaxa.4066.1.4
03AD87B9FF80FFFEBAE44522DC5EFDBD.taxon	distribution	Distribution. Known only from the type locality, Province of Misiones, Argentina.	en	Moura, Luciano De A. (2016): New species, new combinations and synonymies of Neotropical Galerucini (Coleoptera, Chrysomelidae, Galerucinae). Zootaxa 4066 (1): 63-70, DOI: 10.11646/zootaxa.4066.1.4
03AD87B9FF80FFFEBAE44522DC5EFDBD.taxon	etymology	Etymology. The specific name, cabrerae, pays homage to Nora Cabrera (Museo de La Plata, La Plata, Argentina), an expert who dedicated her life to the study of Neotropical chrysomelids.	en	Moura, Luciano De A. (2016): New species, new combinations and synonymies of Neotropical Galerucini (Coleoptera, Chrysomelidae, Galerucinae). Zootaxa 4066 (1): 63-70, DOI: 10.11646/zootaxa.4066.1.4
03AD87B9FF80FFFEBAE44522DC5EFDBD.taxon	discussion	Remarks. Many of the members of Yingaresca have four tubercles on each elytron: two basal — one next to the humerus and another on the disc, close to the scutellum – and two apical, one external and another more on the internal face. Despite the arbitrariness of the division of Yingaresca into groups of species (Wilcox 1971), and the fact that there are no descriptions, to date, defining such groups, we can include Yingaresca cabrerae sp. nov., even though preliminarily, among the Yingaresca of the group difficilis, since it shares an x-shaped pronotum and more evident elytral tubercles. Besides the colored elytral pattern, the new species is the only one in the genus that bears an elevated and conspicuously triangular apical tubercle.	en	Moura, Luciano De A. (2016): New species, new combinations and synonymies of Neotropical Galerucini (Coleoptera, Chrysomelidae, Galerucinae). Zootaxa 4066 (1): 63-70, DOI: 10.11646/zootaxa.4066.1.4
03AD87B9FF82FFFEBAE4449BDDDBFB52.taxon	discussion	Remarks. Jacoby (1886) described and included, although in dubiously, this species in Galerucella. Wilcox (1971, 1974) integrated Galerucella (?) transversicollis in the list of species of Neolochmaea. When revising Neolochmaea, Moura (1998 c) examined the type-material deposited at the BMNH and, since the species does not share the diagnostic characters of the genus, decided to return it to Galerucella. When I more thoroughly analyzed the position of the species in Galerucella as proposed by Moura (1998 c), I verified that it bears (1) small eyes, shorter than the length of the gena, (2) thin antennae, without enlarged antennomeres, and (3) carinae on the elytra; such characteristics are similar to those found in some species of Yingaresca [Y. amazonica (Weise) and Y. batesi (Bowditch)], both comprised in the batesi group according to Wilcox (1971). I therefore transfer G. tranversicollis to this genus. Strengthening the inclusion of the species in Yingaresca, Galerucella is a genus that still does not have a member with Neotropical distribution.	en	Moura, Luciano De A. (2016): New species, new combinations and synonymies of Neotropical Galerucini (Coleoptera, Chrysomelidae, Galerucinae). Zootaxa 4066 (1): 63-70, DOI: 10.11646/zootaxa.4066.1.4
03AD87B9FF82FFF9BAE44041DA1BFE9C.taxon	discussion	Remarks. Both Yingaresca limoensis (Bechyné, 1997) comb. nov. and Y. magdalia (Bechyné, 1997) comb. nov. do not share the characteristics originally presented by Wilcox (1965) that define Ophraella, nor in the subsequent revision by LeSage (1986), such as an oval and convex body — generally densely covered by erect hairs — and the presence of longitudinal bands on the elytra of most species. Furthermore, Ophraella has a Nearctic distribution, while Yingaresca occur essentially in the Neotropical region. In relation to host plants, the species of Ophraella are exclusively recorded on Asteraceae (LeSage, 1986). Bechyné (1997) mentioned O. limoensis and O.	en	Moura, Luciano De A. (2016): New species, new combinations and synonymies of Neotropical Galerucini (Coleoptera, Chrysomelidae, Galerucinae). Zootaxa 4066 (1): 63-70, DOI: 10.11646/zootaxa.4066.1.4
03AD87B9FF85FFF9BAE44776DA38F86C.taxon	description	As for Metrogaleruca longula (originally described by Bechyné in Caraguata and here synonymized with M. obscura), Bechyné (1963) distinguished Caraguata longula diminuta only because of its smaller size and the presence of a darker area at each side of the thoracic disc; these characteristics are also observed in the examined specimens of M. obscura and constitute merely intraspecific variation within populations.	en	Moura, Luciano De A. (2016): New species, new combinations and synonymies of Neotropical Galerucini (Coleoptera, Chrysomelidae, Galerucinae). Zootaxa 4066 (1): 63-70, DOI: 10.11646/zootaxa.4066.1.4
03AD87B9FF85FFF9BAE44776DA38F86C.taxon	materials_examined	Examined material. VENEZUELA, Falcon: Chichiriviche, 2 ♀, 29. II. 1964, J. & B. Bechyné col. (MIZA). SURINAME, Paramaribo: Paramaribo (l’Hermitage) (at light), ♂, 1 – 14. XII. 1969, N. Nieser col. (IRCW); Brownsburg (= Brownsberg), ♂, 19. VII. 1975, D. Engleman col. (TAMU); ♀, 20. VII. 1975, D. Engleman col. (TAMU). BRAZIL, Amapá: (Fazendinha), ♀, 03. I. 1980, E. L. Oliveira col. (MPEG); Macapá (Museu C. Lima), ♀, 15. I. 1980, E. L. Oliveira col. (MPEG); Pará: Belém, ♂, 29. XII. 1960, Flávio col. (MPEG); ♂, 02. VI. 1961, J. Bechyné col. (MPEG); (Granja St. Hort. E. F. Bragança), ♂, ♀, II. 1957, Dirings col. (MZSP); (Parque Ambiental), 2 ♂, 20. IV. 2004, A. L. Nunes e equipe col. (sweeping) (MPEG); ♀, 13. X. 2004, A. L. Nunes e equipe col. (MPEG); (IAN), ♀, 26. V. 1961, J. Bechyné col. (MPEG); (Utinga), Metrogaleruca obscura paraensis, holotype ♂, 03. XII. 1960, Bechyné col. (MPEG); ♀, 24. IV. 1961, J. & B. Bechyné col. (MPEG); ♀, 25. IV. 1961, J. Bechyné col. (MPEG); (Marituba), ♀, 01. VI. 1961, J. Bechyné col. (MPEG); ♀, VIII. 1964, E. Dente col. (MCNZ); Benfica, ♂, 15. XII. 1960, J. Bechyné col. (MPEG); Benevides, ♂, 30. XII. 1960, J. Bechyné col. (MPEG); ♂, 15. VI. 1961, J. & B. Bechyné col. (MPEG); 2 ♂, 15. VI. 1961, J. & B. Bechyné col. (MZSP); Maranhão: Igarapé Gurupi-Una (Aldeia Araçu, 50 km E de Canindé), ♂, ♀, II. 1966, Malkin col. (MZSP); Tocantins: Campinas, 2 ♀, XII. 1935, Borgmeier & Lopes col. (MNRJ); Bahia: 3 ♂, 7 ♀, G. Bondar col. (MNRJ); 3 ♂, G. Bondar col. (FMNH); Espírito Santo: Guarapari, ♂, XI. 1961, M. Alvarenga col. (DZUP 059271); Rio de Janeiro: Campos dos Goytacazes, ♀, 14. IV. 1999, D. J. Cavan col. (MCNZ); (Pesagro), ♂, ♀, 29. XI. 1996, M. Hoffmann col. (UENF); Minas Gerais: Belo Horizonte, ♀, X. 1940 (MPEG); Serra do Caraça (Engenho) (800 m), 2 ♂, XI. 1961, Kloss, Lenko, Martins & Silva col. (MZSP); São Paulo: Jaú, ♂ (MCNZ); Santa Catarina: Laguna, ♀, I. 1964, A. Souto col. (MCNZ); Rio Grande do Sul: Torres (Colônia São Pedro), actually Dom Pedro de Alcântara, 2 ♂, 30. IV. 1993, L. Moura col. (MCNZ); Arroio Teixeira, ♀, 28. II. 1989, M. Hoffmann col. (MCNZ). BOLIVIA, Santa Cruz: 9 mi. N. Sta. Cruz, ♂, ♀, 28. III. 1978, O’Brien & Marshall col. (TAMU); ♂, ♀, night GB, 28. III. 1978, Marshall col. (TAMU). PARAGUAY, without data, ♂, ♀ (115.17.2) (MLPA); San Pedro: Carumbé, ♂, 28. I – 10. III. 1965, R. Golbach col. (IMLA); ♀, 01. II. – 08. III. 1966, R. Golbach col. (IMLA). ARGENTINA, Salta: Cerro San Bernardo, 4 ♂, 20. I. 1948, Willink & Monrós col. (IMLA); ♂, Cerro 20 de Febrero, 20. I. 1950, Willink & Monrós col. (IMLA).	en	Moura, Luciano De A. (2016): New species, new combinations and synonymies of Neotropical Galerucini (Coleoptera, Chrysomelidae, Galerucinae). Zootaxa 4066 (1): 63-70, DOI: 10.11646/zootaxa.4066.1.4
03AD87B9FF84FFFBBAE44130DC01F84C.taxon	description	Types. Holotype male, BRAZIL, state of Rio Grande do Sul, Itaúba, 07. IV. 1978, H. Bischoff col., MCN 27212 (MCNZ). Paratype female, ARGENTINA, Province of Chaco, Resistencia, XI. 1935 (MLPA). Description. Measurements, respectively male / female: body length, 5.9 / 6.9 mm; elytral length 4.5 / 5.6 mm; humeral width 2.3 / 2.7 mm. Head surface rugosely punctate, with short hairs sparsely distributed; antennal tubercles weakly developed, separated by longitudinal sulcus that reaches base of subtriangular clypeus; a weakly developed tubercle present above each antennal tubercle, close to superior internal margin of eye. Eyes globose, small, greatest diameter subequal to length of gena. Labrum yellow, shiny, slightly emarginate medially, with four hairs directed toward ventral margin. Insertion of each antenna positioned at level of inferior portion of the eyes. Interocular distance approximately three times as great as space between antennal insertions. Antennae black, except for yellowtestaceous base and the ventral face of the scape, pedicel, and base and ventral basal half of antennomere III (in paratype ♀, also yellow-testaceous on ventral face of X and XI). Hairs sparsely distributed on first four antennomeres and, beginning with antennomere V, pubescence short and dense, interspersed with longer, erect hairs. Scape somewhat dilated towards apex, twice as long as pedicel; antennomere III longest; antennomere IV slightly longer than the V; antennomeres VI to X subequal in length with V, mildly enlarged in comparison with previous antennomeres. Antennomere XI slightly longer than X, acuminate. Thorax yellow-testaceous, with the following pattern of black maculae on pronotum: one central spot with irregular shape (in female paratype, this spot Y-shaped, Fig. 9); on each side of central spot, a thin, long, curved stripe; a macula on lateral margin at mid-length. Surface of pronotum densely, rugosely punctate, with deep depression on each side of middle, with shallower medial impression next to the anterior margin. Lateral prothoracic margins sub-rounded; width almost twice as great as length, with greatest width at mid-length. One seta present on each anterior and posterior angle of pronotum. Short hairs present, sparsely distributed. Borders of the anterior, lateral and posterior margins enlarged. Anterior coxal cavities closed. Procoxae subcontiguous, not separated by visible invisible prosternal process. Mesosternum subtrapezoidal, with pubescence uniformly distributed, reaching middle of intermediate coxae; metasternum with shiny integument coated by dense pubescence. Femora fusiform, subequal in length to tibiae; tibiae somewhat dilated apically. Legs yellow with black median ring, apex of femora, and base and apical half of tibiae and tarsi. Male with small, acute tubercle apically on intermediate tibia and a small basal tubercle on the ventral face of protarsomere I; female without such tubercles. Pubescence of legs sparsely distributed, denser towards apex of tibiae. Tarsal claws bifid, with internal tooth somewhat convergent and slightly shorter than external tooth. Scutellum yellow-testaceous; punctation weakly marked; short hairs sparsely distributed. Elytra oval, convex, yellow-testaceous with small, black, circular spots throughout (Fig. 9); dense punctation well-marked, interspersed with short, sparse pubescence. Epipleural margin cariniform, somewhat elevated, extending from the humerus to apex; epipleura well developed, subconcave, apically almost reaching sutural margin, covered by short, sparse hairs. Apical sutural margin of elytra round in male, with small, acute tooth in female. Abdomen light-brown, with pubescence sparsely distributed; sternum I about twice as long as II; II to V subequal in length; V with deep, subtriangular, central emargination in male. Aedeagus with median lobe sclerotized, curved in lateral view, with basal spurs thickened, well developed (Fig. 11). Apex of median lobe pointed (Fig. 12). Tegmen (Fig. 13) hastiform and bifurcated near extremity forming two divergent arms, each arm with apex directed downward.	en	Moura, Luciano De A. (2016): New species, new combinations and synonymies of Neotropical Galerucini (Coleoptera, Chrysomelidae, Galerucinae). Zootaxa 4066 (1): 63-70, DOI: 10.11646/zootaxa.4066.1.4
03AD87B9FF84FFFBBAE44130DC01F84C.taxon	distribution	Distribution. Brazil (state of Rio Grande do Sul) and Argentina (province of Chaco).	en	Moura, Luciano De A. (2016): New species, new combinations and synonymies of Neotropical Galerucini (Coleoptera, Chrysomelidae, Galerucinae). Zootaxa 4066 (1): 63-70, DOI: 10.11646/zootaxa.4066.1.4
03AD87B9FF84FFFBBAE44130DC01F84C.taxon	etymology	Etymology. The specific name, from the Latin onca (onça in Portuguese), means jaguar and refers to the pattern of round maculae on the elytra, similar to the fur of a jaguar.	en	Moura, Luciano De A. (2016): New species, new combinations and synonymies of Neotropical Galerucini (Coleoptera, Chrysomelidae, Galerucinae). Zootaxa 4066 (1): 63-70, DOI: 10.11646/zootaxa.4066.1.4
03AD87B9FF84FFFBBAE44130DC01F84C.taxon	discussion	Remarks. Caraguata onca sp. nov. bears a small basal tubercle on the ventral face of protarsomere I in males (females lack this structure) and an acute tooth at the posterior sutural angle of the elytra of females (round in males). To date, there have been no records of sexual dimorphism for these structures in the section Coelomerites, which includes the genus Caraguata. Such sexual dimorphism has however been observed in the section Schematizites: protarsomere I has a ventral tubercle in males of Yingaresca holosericea (Bowditch, 1923), Y. difficilis (Bowditch, 1923), Y. amazonica (Weise, 1921), Schematiza flavofasciata (Klug, 1829), and Brucita marmorata (Jacoby, 1886) (Wilcox 1965; Bechyné & Bechyné 1969; L. de A. Moura, unpublished data); and females have acuminate sutural angles of the elytra in the species of Neolochmaea (Moura 1998 c), Iucetima (Moura 1998 b), and Chlorolochmaea (Bechyné & Bechyné 1969; Moura 1998 a). Caraguata onca sp. nov. differs from all other species of the genus by the maculate pattern of the elytra, and by the sexually dimorphic characters of the anterior tarsi of males and the elytral apices of females.	en	Moura, Luciano De A. (2016): New species, new combinations and synonymies of Neotropical Galerucini (Coleoptera, Chrysomelidae, Galerucinae). Zootaxa 4066 (1): 63-70, DOI: 10.11646/zootaxa.4066.1.4
