taxonID	type	description	language	source
03AD87FA36489D3CFF5DFA6DFD767468.taxon	description	Supra­antennal area ventrally slightly concave, dorsally with distinct longitudinal carina. Occipital carina ending relatively far from hypostomal carina, at a distance of 0.63 – 1.00 the width of mandible base. Pronotum with submarginal lamellar protuberance strongly projected upwards (Fig. 12). Notaulus anterior 0.3 in dorsal view distinct, in lateral view visible up to slightly past center of mesoscutum due to swollen median lobe of mesoscutum. Scuto­scutellar groove smooth and polished, moderately deep. Lateral carina of scutellum near scuto­scutellar groove with circular perforation. Fore wing length 7.9 – 9.7 mm. Fore wing vein M slightly arched, about same length but distinct from Rs. Areolet 1.17 – 2.40 as long as high. Second metasomal tergite 2.21 – 4.25 as long as minimum width, 1.07 – 1.42 as long as maximum width. Ovipositor sheath length / hind tibia length 0.72 – 0.83. Ovipositor slender to thick; laterally slightly to distinctly compressed; apex from blunt to acute (Figs. 3, 6); dorsal valve nodus from vestigial or absent in P. atlantica sp. n. to sharp in P. varians and P. cristata sp. n.; ventral valve apex with 6 – 11 teeth (12 in Fig. 242 of Townes, 1970), 1 st and 2 nd widely spaced, except P. atlantica with all teeth regularly spaced. In his key to genera of Lymeonina Townes (1970) uses, in couplet eight, the character “ upper margin of pronotum strongly swollen, usually with a tubercle­like swelling near upper end of epomia, ” which can be mistaken by the distinct protuberance of P. cristata, taking the species to the wrong section of the key. In this case, other features should be considered, particularly on the propodeum. Although the transverse furrow at the base of the propodeum in P. atlantica is distinctly less wide than in P. varians or P. cristata, its width, associated with the smooth and polished sculpturing, remain characteristic for the genus. The following variation was observed only for southern representatives of the genus: pronotum, mesoscutum and scutellum black; hind trochanter, trochantellus and femur dark brown; propodeum laterally with two large, oval, dark spots; para­ocular white stripe absent from supra­antennal area to posterior orbit (P. atlantica). The characterizing features of Polyphrix also seem to be unique within the subtribe, and since these are not shared, relationships with other genera of Lymeonina remains unapparent.	en	Nogueira, Liana K., Aguiar, Alexandre P. (2005): Generic definition, key, and two new species of Polyphrix Townes (Hymenoptera: Ichneumonidae: Cryptinae) from the Atlantic Forest. Zootaxa 1010 (1): 25-35, DOI: 10.11646/zootaxa.1010.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1010.1.3
03AD87FA364B9D3AFF5DFA70FDBF7798.taxon	etymology	Etymology. The specific epithet atlantica is a reference to the Atlantic Forest, where the holotype was collected.	en	Nogueira, Liana K., Aguiar, Alexandre P. (2005): Generic definition, key, and two new species of Polyphrix Townes (Hymenoptera: Ichneumonidae: Cryptinae) from the Atlantic Forest. Zootaxa 1010 (1): 25-35, DOI: 10.11646/zootaxa.1010.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1010.1.3
03AD87FA364B9D3AFF5DFA70FDBF7798.taxon	description	Description. Holotype female. Length of fore wing: 7.9 mm. Head: antenna with 28 flagellomeres; clypeus 1.8 wider than high, strongly convex, projecting ventrally, without median tooth, sparsely punctate; mandible teeth of equal length, ventral tooth slightly more robust than dorsal tooth; supra­antennal area near ocelli slightly convex, with distinct median longitudinal carina, ventrally concave, without carina; occipital carina moderately high, sharp, uniform; malar space 0.4 longer than basal width of mandible. Mesosoma: epomia absent; pronotum with long, strong wrinkle along posterior margin of collar, fading before reaching ventral corner, latero­centrally with two distinct transverse wrinkles, followed ventrally by short, weak transverse wrinkles; dorsal margin of pronotum with submarginal, low, transverse protuberance near mid­length (Figs. 1, 11); mesoscutum and scutellum glabrous, polished, shiny; mesoscutum subcircular; notaulus delicate, faint, in dorsal view distinct approximately in the anterior 0.3, in lateral view swollen median lobe of mesoscutum makes notaulus visible up to slightly past center of mesoscutum; scutoscutellar groove smooth, polished; mesopleuron, smooth, shiny; sternaulus upcurved, incomplete, reaching about 0.4 the distance to middle coxa; epicnemial carina restricted to ventral 0.70 of mesopleuron, almost straight, mesopleuron dorsad of carina giving rise to short, transverse strigulation; mesopleural suture straight, narrow and crenulate; metapleuron with sparse punctures with hairs; all tibiae with numerous bristles; all pre­apical tarsi weakly bilobed; hind femur 7.5 times as long as wide. Propodeum: transverse furrow at base laterally with two longitudinal, short wrinkles; distance from anterior transverse carina to anterior margin 1.36 longer than width of transverse furrow at base, measured centrally; scarcely pilose, its anterior margin laterally regular, centrally weakly concave; anterior transverse carina strong, straight, complete, area in front of it smooth, shiny, area behind it with about 13 strong, regular, sharp, transverse wrinkles (Figs. 2, 17); posterior transverse carina and apophyses absent; spiracle about twice as long as wide. Fore wing crossveins cu­a and Cu 1 b vertical, straight; vein 2 ­ Cu 1 0.2 length of crossvein Cu 1 b; ramellus absent; areolet pentagonal, 1.17 wider than high (Fig. 9); 3 rs­m as long as 2 rs­m, both vertical; bulla of fore wing crossvein 3 rs­m on posterior 0.70 of vein. Hind wing vein 1 ­ Cu 1 about same length as crossvein cu­a; vein 2 ­ 1 A reaching about 0.7 the distance to posterior margin. Metasoma: first tergite smooth, shiny, spiracle at center, moderately prominent; metasomal tergite 1 about 4.8 longer than wide; tergite 2 smooth, shiny, with sparse pilosity, apex 2.3 as wide as base; thyridium longer than wide; tergites 3 – 8 smooth, shiny, pilose; ovipositor sheath 0.83 as long as hind tibia; ovipositor strong, thick, sparsely punctate, basal half almost cylindrical, apical half laterally weakly compressed; dorsal valve without nodus, ventral valve with seven apical teeth (Fig. 3). Color. Head, pronotum and mesoscutum generally black, remainder of mesosoma deep yellow (239,194,014), metasoma mostly black (Fig. 14). Head: scape dark brown (045,030,030), with inner apical subcircular white spot; pedicel, flagellomeres dark brown, except 9 and 16 dorsal half and 10 – 15, white; supraclypeal area, clypeus, mouth parts, base of mandible and malar space, white; apex of mandible dark brown. Mesosoma: pronotum black, collar from middle to ventral corner, white; mesoscutum, scutellum dorsally, and postscutellum, black; scutellum laterally yellow (255,216,063); tegula brownish, marginally yellow; basal and lateral areas of propodeum, deep yellow; propodeum laterally, behind anterior transverse carina, with two lateral, oval, large, undefined dark brown spots; fore and mid coxae yellow, hind coxa deep yellow; fore and mid femora, mid trochantellus outer side brownish; hind trochanter, trochantellus and femur, dark brown; hind tarsi pale yellow (215,210,155), with apical tarsomere light brown (155,135,060). Metasoma: black; first tergite basal 0.26 yellow; tergites 1 – 7 posterior margin white, widest on T 2 and T 7; ovipositor dark red (095,055,045); sheaths dark brown; wings hyaline.	en	Nogueira, Liana K., Aguiar, Alexandre P. (2005): Generic definition, key, and two new species of Polyphrix Townes (Hymenoptera: Ichneumonidae: Cryptinae) from the Atlantic Forest. Zootaxa 1010 (1): 25-35, DOI: 10.11646/zootaxa.1010.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1010.1.3
03AD87FA364B9D3AFF5DFA70FDBF7798.taxon	discussion	Comments. Similar to P. varians, from which it can be differentiated by the shorter fore wing (7.9 – 8.3 mm vs. 9.0 – 9.7 mm); para­ocular white stripe interrupted from supraclypeal area to posterior orbit (vs. complete); pronotum with submarginal protuberance weak (vs. small and sharp); anterior margin of propodeum weakly sinuous (vs. centrally clearly concave); transverse furrow at base of propodeum centrally narrower than width of propodeum between anterior margin and anterior transverse carina (vs. distinctly wider) (compare Figs. 17 and 19); pronotum, mesoscutum and scutellum black (vs. dark orange); hind trochanter, trochantellus and femur dark brown (vs. yellowish); ovipositor strong, thick, dorsal valve without nodus. Variability. Paratype specimen with fore wing length 8.3 mm; areolet approximately rectangular, 1.33 wider than high (Fig. 10), crossvein 2 rs­m almost opposite 2 m­cu; scape dark brown, laterally dusty white. Distance from anterior transverse carina to anterior margin of propodeum 1.31 longer than width of transverse furrow at base of propodeum, measured centrally. First tergite yellow on basal 0.30. Male. Unknown.	en	Nogueira, Liana K., Aguiar, Alexandre P. (2005): Generic definition, key, and two new species of Polyphrix Townes (Hymenoptera: Ichneumonidae: Cryptinae) from the Atlantic Forest. Zootaxa 1010 (1): 25-35, DOI: 10.11646/zootaxa.1010.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1010.1.3
03AD87FA364B9D3AFF5DFA70FDBF7798.taxon	biology_ecology	Host. Unknown.	en	Nogueira, Liana K., Aguiar, Alexandre P. (2005): Generic definition, key, and two new species of Polyphrix Townes (Hymenoptera: Ichneumonidae: Cryptinae) from the Atlantic Forest. Zootaxa 1010 (1): 25-35, DOI: 10.11646/zootaxa.1010.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1010.1.3
03AD87FA364B9D3AFF5DFA70FDBF7798.taxon	materials_examined	Type material. Holotype female: “ Brazil: ES, Santa Teresa, Estação Biológica Santa Lúcia, 17 – 20. VI. 1997, Yellow Pans, WAHoffmann & RRibeiro. ” Hind right tarsomeres 2 – 5 missing; metasoma crumpled; otherwise in good shape. Paratype female: same locality and collector, 28 – 30. IV. 1997. Right hind coxa, posterior apical tarsus, and apex of ventral valve of ovipositor missing; otherwise in good shape. Holotype and paratype in the collection of the Museu de Zoologia da Universidade de São Paulo, Brazil.	en	Nogueira, Liana K., Aguiar, Alexandre P. (2005): Generic definition, key, and two new species of Polyphrix Townes (Hymenoptera: Ichneumonidae: Cryptinae) from the Atlantic Forest. Zootaxa 1010 (1): 25-35, DOI: 10.11646/zootaxa.1010.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1010.1.3
03AD87FA364B9D3AFF5DFA70FDBF7798.taxon	distribution	Distribution. Brazil (ES).	en	Nogueira, Liana K., Aguiar, Alexandre P. (2005): Generic definition, key, and two new species of Polyphrix Townes (Hymenoptera: Ichneumonidae: Cryptinae) from the Atlantic Forest. Zootaxa 1010 (1): 25-35, DOI: 10.11646/zootaxa.1010.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1010.1.3
03AD87FA364C9D38FF5DFEE8FDBF763F.taxon	etymology	Etymology. The specific epithet cristata is from the Latin crista, meaning ridge, a reference to the large, laminar dorso­lateral projection on the pronotum.	en	Nogueira, Liana K., Aguiar, Alexandre P. (2005): Generic definition, key, and two new species of Polyphrix Townes (Hymenoptera: Ichneumonidae: Cryptinae) from the Atlantic Forest. Zootaxa 1010 (1): 25-35, DOI: 10.11646/zootaxa.1010.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1010.1.3
03AD87FA364C9D38FF5DFEE8FDBF763F.taxon	description	Description. Holotype female. Length of fore wing: 8.5 mm. Head: clypeus 1.4 wider than high, strongly convex, projecting ventrally, without median tooth, sparsely weakly punctate; mandible teeth of equal length, ventral tooth slightly more robust than dorsal tooth; supra­antennal area near ocelli slightly convex, with distinct median longitudinal carina, ventrally concave; occipital carina moderately high, sharp, uniform; malar space 0.5 longer than basal width of mandible. Mesosoma: epomia absent; pronotum with long, strong wrinkle along posterior margin of collar, fading before reaching ventral corner, latero­centrally with three weak transverse wrinkles, dorsally with submarginal lamellar protuberance clearly exceeding dorsal margin mid­length (Figs. 4, 12), projecting upwards (Fig. 15); mesoscutum and scutellum glabrous, polished, shiny; mesoscutum subcircular; notaulus delicate, faint, in dorsal view distinct approximately in the anterior 0.3, in lateral view swollen median lobe of mesoscutum makes notaulus visible up to slightly past center of mesoscutum; scuto­scutellar groove smooth, polished; mesopleuron smooth, shiny; sternaulus upcurved, incomplete, reaching about 0.4 the distance to middle coxa; epicnemial carina restricted to ventral 0.60 of mesopleuron, almost straight, mesopleuron dorsad of carina giving rise to short, transverse strigulation; mesopleural suture straight, narrow and crenulate; metapleuron smooth, shiny, with sparse hairs; all tibiae with numerous bristles; fore and middle pre­apical tarsi weakly bilobed; hind femur 7.2 times as long as wide. Propodeum: transverse furrow at base smooth, shiny; distance from anterior transverse carina to anterior margin 1.00 longer than width of transverse furrow at base, measured centrally; scarcely pilose, its anterior margin laterally prominent, centrally concave; anterior transverse carina strong, straight, complete, area in front of it smooth, shiny, with sparse hairs, area behind it with about 10 strong, complete or incomplete, sharp, transverse wrinkles (Figs. 5, 18); posterior transverse carina and apophyses absent; spiracle about twice as long as wide. Fore wing crossveins cu­a and Cu 1 b vertical, straight; vein 2 ­ Cu 1 0.2 length of crossvein Cu 1 b; ramellus absent; areolet rectangular, 1.80 wider than high; 3 rs­m as long as 2 rs­m, both vertical; bulla of fore wing crossvein 3 rs­m on posterior 0.75 of vein. Hind wing vein 1 ­ Cu 1 about 0.8 length of crossvein cu­a; vein 2 ­ 1 A reaching about 0.7 the distance to posterior margin. Metasoma: first tergite smooth, shiny, with very sparse pilosity, spiracle at center, prominent; metasomal tergite 1 about 5.8 longer than wide; tergite 2 smooth, shiny, with sparse pilosity, apex 2.50 as wide as base; thyridium longer than wide; tergites 3 – 8 smooth, shiny, pilose; ovipositor sheath 0.75 as long as hind tibia; ovipositor slender, sparsely punctate, laterally strongly compressed; dorsal valve with nodus, ventral valve with nine apical teeth, the last 3 – 4 faint (Fig. 6). Color. Head, pronotum and mesoscutum generally black, remainder of mesosoma deep yellow (239,194,014), metasoma mostly black (Fig. 15). Head: scape dark brown; pedicel, flagellomeres dark brown (045,030,030), except 9 apical and 17 dorsal half and 10 – 16, white; supraclypeal area, clypeus, mouth parts, base of mandible and malar space, white; apex of mandible dark brown. Mesosoma: pronotum black, collar and the distance from middle to ventral corner, white; mesoscutum, scutellum dorsally, and postscutellum, black; scutellum apex and lateral 0.3, and tegula, pale yellow (215,210,155); axillary trough of mesoscutum with ventral, subcircular, dark spot near scutellum; area in front of anterior carina of propodeum brownish, except centrally narrowly yellowish, area behind anterior carina with large, brownish, undefined spot on each side; fore and mid coxae yellow (255,216,063), hind coxa deep yellow; hind trochanter, trochantellus and femur, dark brown; hind tarsi pale yellow; all apical tarsomeres light brown (155,135,060). Metasoma: black; first metasomal tergite basal 0.23 yellowish, remainder black, apical margin narrowly white; tergites 1 – 7 posterior margin white, tergite 8 black; ovipositor dark red (095,055,045); sheaths dark brown; wings hyaline.	en	Nogueira, Liana K., Aguiar, Alexandre P. (2005): Generic definition, key, and two new species of Polyphrix Townes (Hymenoptera: Ichneumonidae: Cryptinae) from the Atlantic Forest. Zootaxa 1010 (1): 25-35, DOI: 10.11646/zootaxa.1010.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1010.1.3
03AD87FA364C9D38FF5DFEE8FDBF763F.taxon	discussion	Comments. Similar to P. atlantica sp. n., from which it can be differentiated by showing para­ocular white stripe complete (vs. interrupted from supraclypeal area to posterior orbit); pronotum with strong and distinct submarginal protuberance (vs. weak); anterior margin of propodeum centrally clearly concave (vs. weakly sinuous); transverse furrow at base of propodeum distinctly wider than width of propodeum between anterior margin and anterior transverse carina (vs. centrally narrower) (compare Figs. 17 and 18); first metasomal tergite 5.8 longer than wide (vs. 4.8); ovipositor slender, dorsal valve with nodus. Male. Unknown.	en	Nogueira, Liana K., Aguiar, Alexandre P. (2005): Generic definition, key, and two new species of Polyphrix Townes (Hymenoptera: Ichneumonidae: Cryptinae) from the Atlantic Forest. Zootaxa 1010 (1): 25-35, DOI: 10.11646/zootaxa.1010.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1010.1.3
03AD87FA364C9D38FF5DFEE8FDBF763F.taxon	biology_ecology	Host. Unknown.	en	Nogueira, Liana K., Aguiar, Alexandre P. (2005): Generic definition, key, and two new species of Polyphrix Townes (Hymenoptera: Ichneumonidae: Cryptinae) from the Atlantic Forest. Zootaxa 1010 (1): 25-35, DOI: 10.11646/zootaxa.1010.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1010.1.3
03AD87FA364C9D38FF5DFEE8FDBF763F.taxon	materials_examined	Type material. Holotype female. “ Brazil: ES, Santa Maria de Jetibá: Fazenda Paulo Seick, 06 – 12. XII. 2002, Malaise, MTTavares & COAzevedo & eq. col. ” Right antenna apical half and left antenna apex, hind right tarsomeres 3 – 5, missing; metasoma crumpled; otherwise in good shape. Deposited in the collection of the Universidade Federal do Espírito Santo, Brazil.	en	Nogueira, Liana K., Aguiar, Alexandre P. (2005): Generic definition, key, and two new species of Polyphrix Townes (Hymenoptera: Ichneumonidae: Cryptinae) from the Atlantic Forest. Zootaxa 1010 (1): 25-35, DOI: 10.11646/zootaxa.1010.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1010.1.3
03AD87FA364C9D38FF5DFEE8FDBF763F.taxon	distribution	Distribution. Brazil (ES).	en	Nogueira, Liana K., Aguiar, Alexandre P. (2005): Generic definition, key, and two new species of Polyphrix Townes (Hymenoptera: Ichneumonidae: Cryptinae) from the Atlantic Forest. Zootaxa 1010 (1): 25-35, DOI: 10.11646/zootaxa.1010.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1010.1.3
03AD87FA36419D36FF5DFEE8FC71748F.taxon	distribution	Distribution. The specimens of P. varians originally studied by Townes, now at the American Entomological Institute, were checked, but none have been collected in Peru. Therefore, the occurrence of P. varians in that country, as registered by Townes (1970) and later cited by Carrasco (1972), is incorrect. The specimens examined by the authors were collected recently in the Amazon Basin, Brazil, at nearly the same latitude and about 4 º W from the type locality.	en	Nogueira, Liana K., Aguiar, Alexandre P. (2005): Generic definition, key, and two new species of Polyphrix Townes (Hymenoptera: Ichneumonidae: Cryptinae) from the Atlantic Forest. Zootaxa 1010 (1): 25-35, DOI: 10.11646/zootaxa.1010.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1010.1.3
03AD87FA36419D36FF5DFEE8FC71748F.taxon	materials_examined	Material examined. Three females: Brazil, PA, Melgaço, Caxiuanã, Trilha Estação Científica Ferreira Pena, 13 – 16. XI. 2003, yellow pans, APAguiar & JDias, and same data except Trilha Igarapé Tijucaquara, 18 – 21. XI. 2003; Brasil, GO, Uruaçu, Serra da Mesa Survey, 14 º 17.0´S 48 º 54.8´W, 27. V. 1996, yellow pans along permanent stream, N. F. Johnson leg. Specimens in the collections of the Museu de Zoologia da Universidade de São Paulo and Museu Paraense Emílio Goeldi, Brazil.	en	Nogueira, Liana K., Aguiar, Alexandre P. (2005): Generic definition, key, and two new species of Polyphrix Townes (Hymenoptera: Ichneumonidae: Cryptinae) from the Atlantic Forest. Zootaxa 1010 (1): 25-35, DOI: 10.11646/zootaxa.1010.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1010.1.3
