identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03AABA6D1809AA6FFF5BFEC7FCD661FD.text	03AABA6D1809AA6FFF5BFEC7FCD661FD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Crenadactylus Dixon & Kluge 1964	<div><p>Crenadactylus Dixon &amp; Kluge, 1964</p><p>Type species. Diplodactylus ocellatus (= Crenadactylus ocellatus) Gray, 1845 by original designation.</p><p>Diagnosis. Based on Dixon &amp; Kluge (1964). Species within the genus Crenadactylus are small (max SVL 35 mm) geckos differing from all other gekkonids by a combination of the following external morphological characteristics; digits with enlarged subdigital lamellae, terminal phalangeal elements forked and claws absent from all digits; outer margins of anterior portion of frontal notched to receive posterior projection of paired nasals. Internal morphological diagnostics include palatines short and broad, atlas fused dorsally; stapes imperforate (stapedial foramen absent); two pairs of sternal ribs, one or two pairs of mesosternal ribs; 28 sacral and presacral vertebrate; sacral diapophyses overlapping and fused (not fused in juveniles); in adults anterior tip of mesoscapula fused to precoracoid process at its union with precoracoid; interclavicle dagger-shaped; fingers 5; toes 5; phalangeal formula of manus 2-3-4-5-3, pes 2-3-4-5-4; 24–26 scleral ossicles; 13 or 15 premaxillary teeth; 29–31 maxillary teeth; 33–37 mandibular teeth; cloacal bones present in males (Dixon &amp; Kluge 1964).</p><p>Etymology. A combination of the Latin word crena meaning ‘notch’ and Greek word daktylos meaning ‘finger’ in reference to the forked terminal phalanges.</p><p>Content. Crenadactylus horni (Lucas &amp; Frost, 1895), Crenadactylus naso Storr, 1978, Crenadactylus occidentalis sp. nov., Crenadactylus ocellatus (Gray, 1845), Crenadactylus pilbarensis sp. nov., Crenadactylus rostralis Storr, 1978, Crenadactylus tuberculatus sp. nov.</p></div>	https://treatment.plazi.org/id/03AABA6D1809AA6FFF5BFEC7FCD661FD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Doughty, Paul;Ellis, Ryan J.;Oliver, Paul M.	Doughty, Paul, Ellis, Ryan J., Oliver, Paul M. (2016): Many things come in small packages: Revision of the clawless geckos (Crenadactylus: Diplodactylidae) of Australia. Zootaxa 4168 (2): 239-278, DOI: 10.11646/zootaxa.4168.2.2
03AABA6D1809AA6CFF5BFC14FE966746.text	03AABA6D1809AA6CFF5BFC14FE966746.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Crenadactylus ocellatus (Gray 1845) Gray 1845	<div><p>Crenadactylus ocellatus (Gray, 1845)</p><p>South-western clawless geckos Figs. 4 A, 5, 6, 7</p><p>Diplodactylus ocellatus Gray, 1845</p><p>Synonymy: Diplodactylus bilineatus Gray, 1845 (fide Günther 1867) [‘South-west’ of Oliver et al. (2010)]</p><p>Holotype. BMNH 1947.3.6.68 from Western Australia. Specimen obtained from the Buchanan collection from WA, collection date not specified.</p><p>Synonomy. Diplodactylus bilineatus . Syntypes (2), Gray 1845, BMNH 1947.3.6.69 (2 specimens) from Houtman’s Abrolhos, Western Australia. Collected by ‘Mr. Gilbert’, collection date not specified.</p><p>Diagnosis. A moderately large (to 35.5 mm SVL) species of Crenadactylus with wide head (HW/HL 0.52– 0.70) and short trunk length (ILL/SVL 0.37–0.50). Rostral in full contact with nostril, internasal (if present) not extending beyond supranasal, 1 or 2 granular postmentals, dorsal scales homogeneous with smooth to weak keels, usually no pre-cloacal pores visible but some males with 2, when present pore-bearing scales in contact at midline, no enlarged tubercles on original tails. Ground colour tan and dark brown; dorsal pattern comprised of poorlydefined longitudinal pale and dark stripes, lateral zones pale with irregular dark stippling, pattern heavily overlain with intermixed pale and dark scales giving an irregular appearance, pale spots or ocelli comprised of 3–6 pale scales usually present in dorsolateral zone.</p><p>Description. A moderately built, small-bodied gecko (to 35.5 mm SVL); body elongate, oblong in crosssection; head relatively wide (HeadW/HeadL 0.52–0.71), moderately long (HeadL/SVL 0.27–0.33), moderately depressed (HeadD/HeadL 0.27–0.35); in dorsal view, widest at ear openings; head not distinct from neck with neck only weakly constricted; loreal region convex; snout moderately short (SnL/HeadL 0.32–0.40), tapering to rounded tip; head depth ~1.5–2 times eye diameter; eye moderately large; rostral ~2.5 times wider than high, in broad contact with nostril, dorsal edge concave to accommodate supranasals, rostral groove extends from dorsal edge of rostral to ~50% of rostral height; 0 or 1 small internasals; supranasals horizontally elongate, 2–3 times wider than long; nostrils small, similar in size to postnasals, directed laterally, in contact with rostral, supranasal, 2–4 postnasals and first supralabial; supralabials 7–9, antero-dorso edge of first supralabial in contact with nostril; second supralabial rectangular, wider than high, length equal to first supralabial; supralabials 3–5 slightly smaller than first and second, gradually decreasing in size, all wider than high; scales on crown small and slightly rounded, ~one-third the size of dorsal scales, increasing in size anteriorly onto snout and supraocular regions; scales on snout flat and slightly enlarged, 1.5–2 times larger than scales on crown; supraocular scales enlarged; 1 or 2 soft spines on posterior eyelid; mental triangular to trapezoidal in shape, widest anteriorly, not extending past ventral edge of first infralabial; postmental not enlarged relative to other scales adjacent to infralabials; gulars small near infralabials, decreasing to granular, ~0.1–0.2 the size of scales on ventrum, juxtaposed and slightly rounded; gulars flattened from mental to eye, becoming more rounded from eye to neck; infralabials 7–9, first largest and squareshaped, gradually decreasing in size posteriorly.</p><p>Limbs short, distance between limbs close to half of SVL (ILL/SVL 0.37–0.50); scales on anterior surfaces of forelimbs elongate, broad and rounded, slightly imbricate, decreasing in size distally, ~2 times the size of scales elsewhere on forelimb; scales on dorsal surface of hind limbs rounded and slightly raised, decreasing in size and becoming more rounded distally; scales on anterior surface of thigh flat and circular, slightly larger in area than scales on hindlimbs elsewhere; fingers 5; toes 5; claws absent from all digits, digits short; fourth finger lamellae in 6 rows; fourth toe lamellae in 8 rows; apical plates much wider than digit; two pairs of dorsal apical plates; one pair of ventral apical plates, round to ovate.</p><p>Scales on dorsum and lateral surfaces small and homogeneous, smooth to weakly keeled, ~2.5 times wider than deep, juxtaposed; scales slightly raised, highest point at posterior edge of scale, widest at midpoint of scale; scales on ventrum homogeneous, flat, imbricate, ~equal to 1.5 times the size of dorsal scales in area, widest at midpoint, with rounded posterior edge.</p><p>Tail moderately long, maximum width one-quarter to one-third along tail, gradually tapering to a fine point, round in cross-section; scales on dorsal surface of tail ~2–3 times size compared to dorsum, homogeneous, flattened, imbricate, square to oblong in shape, arranged in transverse rows; ventral tail scales flattened, ~2–3 times the size of dorsal tail scales; single enlarged cloacal spur per side, ~3–4 times the size of surrounding scales; no visible pre-cloacal pores (except on 2 males with 2 pores).</p><p>Colouration and pattern. Dorsal background colouration tan and dark brown; light brown dorsolateral lines edged with dark brown emanate from back of eye to tail, continues forwards past eye as pale canthal stripe, terminating above nostril; poorly-defined silver-grey vertebral stripe discernible in some specimens, 2–3 scales wide, bordered by poorly defined dark brown paravertebral lines or dark edges of dorsolateral lines; dorsolateral zone with 7–12 widely spaced ocelli between eye and tail, variably defined ocelli cluster comprised of 3–6 pale scales, often lacking or loosely bordered by darker scales; lateral zone with poorly-defined lateral line that extends from postero-ventral edge of eye above limbs to tail, continuing forward as dark loreal stripes to nostril; below lateral stripes on flanks a mixture of darkly pigmented scales with grey and pale scales; crown and snout a mixture of dark brown, brown and pale grey scales; supralabials pale with dark stippling, infralabials moderately pigmented; ventral edge of orbit dark brown; limbs have same intermixture of dark and pale scales with numerous scattered small pale ocelli, weak on front limbs, more prominent on rear; top of digits have a network of dark lines; ventrum pale with fine to heavy dark brown flecking, not forming conspicuous lines; heavier flecking on rear limbs and cloacal region and tail; tail continues dorsal pattern including ocelli on dorso-lateral stripes, dark vertebral zone and light brown paravertebral zone.</p><p>Distribution. This species occurs in the south-west of Australia (Fig. 2). It is absent from the mesic southwestern corner, extending only as far south-west as Waroona (WAM R88483) then extending eastwards including records from Kojanup, Tambelup and Fitzgerald River National Park. The western-most extent of the distribution is from an old record in Israelite Bay at Cape Arid National Park (WAM R14176) and a more recent record from Balladonia at the edge of the Nullarbor Plain (WAM R157901). It extends into the wheatbelt and to the arid interzone, with records from near Mount Manning and Bowgada Nature Reserves, and an isolated record from Walganna Rock in the north-east (WAM R135103, genotyped as C. ocellatus and shown in Fig. 3 A of Oliver et al. 2010). Occurs on most of the Houtman Abrolhos Islands (off Geraldton) on which surveys have been conducted. Habitat. Found in open woodland habitats throughout its range including areas dominated by Xanthorrhea grass trees, Eucalyptus (jarrah, mallee, marri, tuart, wandoo) and spinifex grass ( Triodia) on a wide variety of soft (sandy and loamy) and hard stony substrates (e.g. laterite). Collectors’ notes record specimens have been found under ground cover such as fallen and rotten logs, woodpiles, leaf litter, granite boulders, limestone slabs, sheets of tin, railway sleepers and rubbish. One specimen (WAM R72276) was found one meter high under the bark of a gimlet tree, and other records mention ‘under bark’ indicating some climbing ability.</p><p>Etymology. The specific name ocellatus refers to the pale scattered blotches or ‘ocelli’ on the dorsum.</p><p>Remarks. Our experience with this species indicates they are reasonably common compared to other Crenadactylus species. Within the genus, C. ocellatus is exceptional for its dorsal patterning, with its intermixture of dark and pale scales and less well-defined longitudinal stripes (Fig. 7). In contrast, most other Crendactylus species have relatively ‘clean’ longitudinal lines with possibly the exception of C. tuberculatus sp. nov. (Fig. 4 D). It is likely this broken colour pattern may be beneficial for camouflage owing to this species’ ground-dwelling habits. In contrast, the striped patterns in other Crenadactylus taxa are likely beneficial to species that climb in spinifex clumps and are likely convergent with the phasmid geckos in the genus Strophurus Fitzinger which also occur in spinifex.</p></div>	https://treatment.plazi.org/id/03AABA6D1809AA6CFF5BFC14FE966746	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Doughty, Paul;Ellis, Ryan J.;Oliver, Paul M.	Doughty, Paul, Ellis, Ryan J., Oliver, Paul M. (2016): Many things come in small packages: Revision of the clawless geckos (Crenadactylus: Diplodactylidae) of Australia. Zootaxa 4168 (2): 239-278, DOI: 10.11646/zootaxa.4168.2.2
03AABA6D180AAA69FF5BFA9DFB3E6323.text	03AABA6D180AAA69FF5BFA9DFB3E6323.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Crenadactylus horni (Lucas & Frost 1895) Lucas & Frost 1895	<div><p>Crenadactylus horni (Lucas &amp; Frost, 1895)</p><p>Central Uplands clawless geckos Figures 3 D, 4B, 8, 9</p><p>Ebenavia horni Lucas &amp; Frost, 1895</p><p>[‘Central Ranges’ of Oliver et al. (2010)]</p><p>Holotype. NMV D7533, adult female, Camp 4 of the Horn Expedition, near Charlotte Waters, Northern Territory fide Dixon and Kluge (1964), donated to National Museum Victoria, Melbourne, Australia (NMV) by W.B. Spencer, April 1897. Stored in 70% ethanol at NMV.</p><p>Diagnosis. A moderately large (to 34.8 mm SVL), robust species of Crenadactylus with wide (HW/HL 0.48– 0.63) and deep (HD/HL 0.28–0.42) head with a long snout (SnL/HL 0.34–0.42). Rostral in full contact with nostril, enlarged internasal extending beyond supranasal, 2 slightly enlarged postmentals, dorsal scales homogeneous, smooth or feebly keeled, 6 pre-cloacal pores, innermost pore-bearing scales separated by an intervening scale, no enlarged tubercles on original tails. Ground colour tan and light brown; dorsal pattern consists of well-defined pale and dark longitudinal stripes, lateral zones pale tan with irregular dark brown stippling, occasionally forming 1 or 2 broken or weakly defined lateral lines; ventrum pale off-white with moderate to weak stippling.</p><p>Description. The holotype is in very poor condition (shrivelled, blackened, front limbs broken off; Fig. 8) so the following description is based on other specimens in good condition. A moderately built, small-bodied gecko (to 34.8 mm SVL); body elongate, oblong in cross-section; head slightly wide (HeadW/HeadL 0.48–0.63), moderately long (HeadL/SVL 0.27–0.32), slightly deep (HeadD/HeadL 0.28–0.42); in dorsal view, widest at ear openings; head distinct from neck with neck slightly constricted; canthal-loreal region slightly convex; snout moderately long (SnL/HeadL 0.34–0.43), tapering to rounded tip; eye moderately large; pupil vertical with four scallops; rostral ~3 times wider than tall, in broad to point contact with nostril, dorsal edge below supranasals concave to accommodate supranasals, rostral groove extends from posterior edge of rostral to ~50% of rostral height; supranasals oblong, ~1.5 times wider than long; single enlarged internasal circular, dorsal edge extends beyond dorsal edge of supranasals, occasionally a tiny additional internasal between enlarged supranasal and rostral; nostrils small, slightly smaller than loreals, directed laterally, in contact with rostral, supranasal, 3 postnasals and first supralabial; supralabials 7–9, antero-dorso edge of first supralabial in contact with nostril, postero-dorso edge notched by second postnasal and first loreal, other infralabials rectangular, wider than high, gradually decreasing in size, to corner of mouth; scales on crown small and rounded, ~one-half the size of dorsal scales, increasing in size anteriorly onto snout and supraocular regions; scales on snout flat and slightly enlarged, 2–3 times larger than scales on crown; 1 or 2 soft spines on posterior eyelid; mental triangular to trapezoidal in shape, widest anteriorly, not extending past ventral edge of first infralabial; 2–3 postmentals not slightly enlarged relative to other scales adjacent to infralabials; gulars small near infralabials, decreasing to granular, ~0.2 the size of scales on ventrum; infralabials 7–9, first largest and square-shaped, gradually decreasing in size posteriorly; ear opening small and round to elliptical in shape.</p><p>Limbs short, distance between limbs close to half of SVL (ILL/SVL 0.37–0.50); scales on anterior surfaces of forelimbs broad and rounded, slightly imbricate, ~2 times the size of scales elsewhere on forelimb; scales on dorsal surface of hind limbs rounded and slightly raised; scales on anterior surface of thigh flat and circular, slightly larger in area than scales on hindlimb and elsewhere; fingers 5; toes 5; digits short, terminating in 2 enlarged apical plates, claw absent; fourth finger lamellae in 6 rows; fourth toe lamellae in 8 rows; apical plates much wider than digit.</p><p>Scales on dorsum and lateral surfaces homogeneous and smooth and rounded (lacking defined keels) to feebly keeled, ovoid in cross-section, juxtaposed; scales slightly raised, highest point at posterior edge of scale, widest at midpoint of scale; scales on ventrum homogeneous, flat, imbricate, equal to 2 times the size of dorsal scales in area, widest at midpoint, with rounded posterior edge.</p><p>Tail moderately long, maximum width 1/4 to 1/3 along tail, gradually tapering to a fine point, round in crosssection; scales on dorsal surface of tail ~2–3 times size compared to dorsum, homogeneous, flattened, imbricate, square to oblong in shape, arranged in travnserse rows; subcaudal scales flattened, ~2–3 times the size of dorsal tail scales; post-cloacal scales 1.5 times larger than typical ventral scales; single enlarged cloacal spur per side, ~3–4 times the size of surrounding scales; no visible pre-cloacal pores (except on 2 males, with six pores).</p><p>Colouration and pattern. Ground colour tan to dark brown; a narrow silvery-grey vertebral stripe usually visible, 2–4 scales wide; vertebral stripe bordered by medium brown paravertebral stripes with darker brown margins; pale dorsolateral stripe 3–5 scales wide, narrowing on neck to ~6 smaller scales, continuing to posterodorsal edge of eye and through eye as pale canthal stripe; wide, dark dorso-lateral stripe up to 9 scales wide, medium brown with dark brown borders, on some individuals, dark borders are highly contrasting with central lighter zone giving the appearance of two separate stripes, continues anteriorly through eye as dark loreal stripe; pale lateral zone with poorly-defined brown lateral stripe; ventral edge of orbit dark brown; labials stippled with dark brown; ventrum pale with light to medium stippling. In one specimen (SAMA R22245), the dorsolateral stripe is narrow and poorly-defined.</p><p>Distribution. Apparently isolated populations in the ranges of central Australia, with most records from the Northern Territory near Alice Springs and West MacDonnell and Finke Gorge National Parks (Fig. 2). As far north as Wilora (AMS R52216), and east to near the Dulcie Ranges National Park (NTM R14120). Also occurring in north-central South Australia near the Ngarutjara and Pukatja homelands. All recent and reliable records are confined to rocky areas, separated by less suitable habitat, indicating populations are confined to these areas. The presumed type locality of Charlotte Waters in the Northern Territory (indicated by a ‘?’ in Fig. 2) is to the southeast of all other records, and the habitat is unlike that of the rest of this species’ records. The collection location is likely to be approximate based on the available landmarks that the Horn Expedition visited.</p><p>Habitat. Collection records for several specimens mention they were taken from spinifex clumps or from under rocks near spinifex, with records also from a rocky gully, rock platforms or outcrops.</p><p>Etymology. Named for William A. Horn, financer and early participant of the Horn Scientific Expedition to central Australia in 1894.</p><p>Remarks. The subspecies C. ocellatus horni used to refer to the central ranges species plus Crenadactylus from the Pilbara, Cape Range and west coast owing to their strongly striped appearance relative to the type (sub)species, C. ocellatus . Along with C. ocellatus, C. horni is a more large-bodied species within Crendactylus. Individuals also possesses this species’ unique diagnostic character of an enlarged supranasal scale (Fig. 3 D) and rounded (not keeled) dorsal scales, otherwise, it is very similar to other striped species in the genus.</p></div>	https://treatment.plazi.org/id/03AABA6D180AAA69FF5BFA9DFB3E6323	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Doughty, Paul;Ellis, Ryan J.;Oliver, Paul M.	Doughty, Paul, Ellis, Ryan J., Oliver, Paul M. (2016): Many things come in small packages: Revision of the clawless geckos (Crenadactylus: Diplodactylidae) of Australia. Zootaxa 4168 (2): 239-278, DOI: 10.11646/zootaxa.4168.2.2
03AABA6D180FAA76FF5BFE71FC3063B3.text	03AABA6D180FAA76FF5BFE71FC3063B3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Crenadactylus occidentalis	<div><p>Crenadactylus occidentalis sp. nov.</p><p>Western clawless geckos</p><p>Figs. 4 C, 10, 11</p><p>[‘Carnarvon Basin’ of Oliver et al. (2010)]</p><p>Holotype. WAM R113683, adult male, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=113.2&amp;materialsCitation.latitude=-26.0" title="Search Plazi for locations around (long 113.2/lat -26.0)">Dirk Hartog Homestead</a>, Dirk Hartog Island, WA (26°00'S, 113°12'E), collected by B. Maryan and R. Browne-Cooper, 19 April 1992. Fixed in 10% formalin, stored in 70% ethanol at WAM.</p><p>Paratypes (6). WAM R57525, 40 km north-east of Yuna, WA (28°06'S, 115°15'E); WAM R96676, 10 km north-west of Wandina Homestead, WA (27°56'S, 115°33'E); WAM R 120779, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=114.58917&amp;materialsCitation.latitude=-27.056665" title="Search Plazi for locations around (long 114.58917/lat -27.056665)">Nerren Nerren Station</a>, WA (27°03'24"S, 114°35'21"E) ; WAM R124891, 38 km west south-west Hamelin Homestead, WA (26°35'34"S, 113°53'22"E); WAM R131376, 70 km south of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=114.11667&amp;materialsCitation.latitude=-22.579723" title="Search Plazi for locations around (long 114.11667/lat -22.579723)">Exmouth</a>, WA (22°34'47"S, 114°07'00"E) ; WAM R135497, False Entrance Well, WA (26°23'S, 113°19'E).</p><p>Diagnosis. A medium-sized (to 32.6 mm SVL) species of Crenadactylus with wide (HW/HL 0.50–0.60) head. Rostral in full contact with nostril, internasal (if present) not extending beyond supranasal, 2 (occasionally 3) small postmentals, dorsal scales homogeneous and weakly keeled, 3–4 pre-cloacal pores, innermost pore-bearing scales separated by an intervening scale, no enlarged tubercles on original tails. Ground colour tan and light brown; dorsal pattern consists of well-defined pale and dark longitudinal stripes, small pale white to orange spots comprised of 2– 3 scales may be present, lateral zones pale to dark grey with at most faint uniform stippling (not forming lines); ventrum pale off-white with stippling absent to moderate.</p><p>Description of holotype. WAM R113683, a small adult male with the following meristics: SVL 28.0 mm; tail absent, remaining portion ~ 3.1 mm; HeadL 8.1 mm; HeadW 4.6 mm; HeadD 2.7 mm; SnL 2.5 mm; ILL 12.6 mm; SupLab 9 (left), 8 (right); InfLab 9, 8; internasals 0; PCP 4.</p><p>A slender, elongate, small-bodied gecko; body oblong in cross section, ~2.0–2.5 times wider than deep; head narrow (HeadW/HeadL 0.57), elongate (HeadL/SVL 0.29), moderately depressed (HeadD/HeadL 0.33); in dorsal view, widest at ear opening, as wide as widest portion of body; head not distinct from neck, neck weakly constricted; loreal region weakly convex, concave around nasal region; snout elongate (SnL 2.5; SnL/HeadL 0.31), rounded tip; eye moderately large; rostral ~3.0 times wider than high, in broad contact with nostril, dorsal edge concave to accommodate first supranasal, slightly notched by median internasal, rostral groove originates from dorsal edge of scale, ~40% of rostral height; internasals absent; supranasals 2 per side, first oblong, transverselyoriented, ~2.5 times wider than high, in broad contact with nostril; second supranasal small, rounded, ~half the size of first supranasal; nostril small, slightly smaller than second supranasal, directed dorsolaterally, in contact with rostral, first and second supranasals, three postnasals and first supralabial; postnasals 3; supralabials 9 (left), 8 (right), antero-dorso edge of first supralabial in contact with nostril; second supralabial square, narrower than first; supralabials 3–4 square, equal in size, 5–9 rectangular, gradually decreasing in size, all wider than high; scales on crown small and granular, increasing in size anteriorly onto snout; scales on snout flat, enlarged; supraocular scales enlarged; two enlarged conical scales at postero-dorso edge of eye; mental trapezoid-shaped, widest anteriorly, narrowing slightly to posterior edge, posterior edge not extending beyond ventral edge of first infralabials; postmentals 2, round in shape, equal to size of scales posterior to postmentals; gulars rounded, gradually decreasing in size posteriorly to granular scales; infralabials 8 on each side, 1–3 largest, equal in size, square-shaped, slightly higher than wide, 4–8 rectangular, wider than high, gradually decreasing in size posteriorly.</p><p>Limbs short, distance between limbs nearly half of SVL (ILL/SVL 0.45); scales on dorsal surface of forelimbs imbricate, rounded and slightly raised, becoming flatter distally; scales on ventral surface of forelimbs smaller; scales on dorsal surface of hind limbs imbricate, rounded and slightly raised, becoming flatter distally; scales on ventral surface of hind limbs similar to scales on dorsal surface of hind limbs, becoming granular distally; fingers 5 (fifth finger missing from left forelimb); toes 5; claws absent from all digits, digits moderately long; fourth finger lamellae in 8 rows; fourth toe lamellae in 10 rows; in dorsal view, enlarged scale between apical plates ~2 times the size of adjacent lateral scales, apical plates much wider than digit; two pairs of dorsal apical plates; one pair of ventral apical plates, ovate.</p><p>Scales on dorsum small and homogeneous, round to oblong, juxtaposed; scales weakly keeled, highest point at posterior edge of scale, widest at midpoint of scale, becoming flatter and imbricate laterally; scales on ventrum homogeneous, flat, imbricate, ~2–3 times the size of dorsal scales in area, widest at midpoint, rounded posteriorly.</p><p>Tail detached, remaining portion ~ 3.1 mm, tapers gradually from widest point posterior to vent; round in crosssection; scales similar to scales on dorsum, rounded and slightly raised, imbricate; ventral tail scales flattened and imbricate, ~1.5–2.0 times the size of scales on dorsal surface of tail; single enlarged raised post-cloacal spur on both sides of cloaca, ~10 times the size of surrounding scales; pre-cloacal pores 4, 2 innermost pore-bearing scales separated by a single intervening poreless scale, pore in notch on posterior edge of scale.</p><p>Colouration and pattern. In preservative, dorsum with longitudinal series of stripes, yellowish-brown ground colour; vertebral stripe silvery-grey, 3 scales wide; 2 dark brown paravertebral stripes, 3 scales wide; pale wide dorsal stripes, light yellow-brown with scattered dark scales, 4 scales wide, terminates at postero-dorsal edge of eye; brown latero-dorsal stripes 2–3 scales wide, originates at posterior edge of eye, over ear and above limbs, continuing to base of tail.</p><p>Variation. Most specimens were similar to the holotype in body size and proportions and scalation. Variation in the snout tip area usually took the form of from 3–6 scales in contact with first supranasals, including presence or absence of a small internasal between the first supranasals (Fig. 3 C). Pre-cloacal pores were 4 with the exception of WAM R 120789 with 3. Males possessed enlarged cloacal spurs whereas in females, the spurs are smaller and protrude away from body less. Original tails with large scales (~4–6 times the size of dorsal scales), scales rectangular with rounded posterior edge, slightly imbricate, arranged in regular rows; subcaudal scales ~1.5 times larger than dorsal tail scales, flattened, imbricate; regenerated tails with smaller, flatter scales compared to original scales; hemipenes bifid.</p><p>Colouration and pattern. Highly contrasting dark brown and pale yellowish-brown longitudinal stripes; ground colour a dull tan or light yellow-brown; vertebral stripe pale silvery-grey, 3 scales wide, extending to crown; vertebral stripe enclosed by dark brown paravertebral stripes, 3 scales wide, that meet between hindlimbs and base of tail, extending anteriorly to crown, becoming intermixed with pale scales on crown; wide pale tan to light yellow-brown dorsolateral stripe, 4 scales wide, extends from upper part of eye to tail, extending forwards through eye to form pale canthal stripe, edged with dark brown in some individuals; above canthal stripes a dark brown triangular patch on snout; rostral and labial scales pale, darkly stippled to varying degrees, posterior edge of orbit with dark brown mark; dark brown dorsolateral stripes, 2–3 scales wide, emanate from posterior edge of eye and continue to tail, extending forwards through eye to form dark brown loreal stripe; pale dorsolateral zone occasionally flecked with small orange clusters of scales; dark paravertebral and dorsolateral stripes peppered with fine clusters or single pale scales in some individuals; lateral zone pale silvery-white, almost continuous to ventrum, thin weakly-defined brown lateral stripe usually present, below lateral stripe, scattered dark flecks may be present. Ventrum ranges from an almost immaculate, pale silvery-grey, to lightly stippled with black, occasionally forming short streaks. Limbs variably marked, a mixture of pale, light brown and dark brown scales with scattered clusters or ocelli of 3–8 pale scales, darker scales forming irregular dark variegations. Original tails have a continuation of the dorsal pattern, with dark brown paravertebral stripes on dorsum merging to form single dark vertebral stripe; regenerated tails have a mixture of dark and pale scales, not forming a regular pattern.</p><p>Distribution. Occurs on the Carnarvon coast of Western Australia from areas inland of Geraldton in the south, extending north through Kalbarri, Shark Bay and to the southern part of the North West Cape at Yardie Creek (where it is narrowly sympatric with C. tuberculatus sp. nov.) (Fig. 2). Also collected from Bernier and Dorre islands in Shark Bay and off Carnarvon. Furthest inland occurrences near Wandana Nature Reserve at the southern part of its distribution (~ 100 km inland) and Nerren Nerren further north (~ 70 km inland) (both with genotyped specimens), otherwise most records occur within 50 km of the coastline.</p><p>Habitat. Collection records indicate a preference for spinifex and coastal dune vegetation such as shrubs, Acacia and Banksia . Occasionally collected from under tin and limestone slabs.</p><p>Etymology. The specific name occidentalis (Latin) refers to the western distribution of this species.</p><p>Remarks. This species’ distribution is closely associated with the western coast and its furthest inland extent is also the most southern, possibly indicating a low tolerance for extremely dry conditions. This area receives variable rainfall during the winter months (May–August), with decreasing rain furthest north where C. occidentalis is only found in a narrow band along the coast.</p><p>The sister species of C. occidentalis is C. tuberculatus sp. nov., with an estimate of divergence around 5 mya (Oliver et al. 2010). Unlike the rather typical Crenadactylus morphology possessed by C. occidentalis, however, C. tuberculatus sp. nov. has a highly derived morphology and pattern and is entirely restricted to the rugged limestone hills and gorges of the Cape Range. The two species come into contact at the northern edge of the distribution of C. occidentalis, including sympatry where the two species meet at Yardie Creek.</p></div>	https://treatment.plazi.org/id/03AABA6D180FAA76FF5BFE71FC3063B3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Doughty, Paul;Ellis, Ryan J.;Oliver, Paul M.	Doughty, Paul, Ellis, Ryan J., Oliver, Paul M. (2016): Many things come in small packages: Revision of the clawless geckos (Crenadactylus: Diplodactylidae) of Australia. Zootaxa 4168 (2): 239-278, DOI: 10.11646/zootaxa.4168.2.2
03AABA6D1810AA73FF5BFDFCFE0663CF.text	03AABA6D1810AA73FF5BFDFCFE0663CF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Crenadactylus tuberculatus	<div><p>Crenadactylus tuberculatus sp. nov.</p><p>Cape Range clawless geckos Figs. 3 B, 4D, 12, 13</p><p>[‘Cape Range’ of Oliver et al. (2010)]</p><p>Holotype. WAM R174304, adult male, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=114.010994&amp;materialsCitation.latitude=-22.090689" title="Search Plazi for locations around (long 114.010994/lat -22.090689)">Charles Knife Road</a>, Cape Range National Park, North West Cape, Western Australia (22°05'26.48"S, 114°00'39.57"E) by P. Doughty, A.M. Bauer and R.J. Ellis, 19 August 2014. Fixed in 10% formalin, stored in 70% ethanol at WAM . Liver sample stored in 100% ethanol at WAM.</p><p>Paratypes (6). WAM R61203, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=113.816666&amp;materialsCitation.latitude=-22.333334" title="Search Plazi for locations around (long 113.816666/lat -22.333334)">Yardie Creek</a>, WA (22°20'S, 113°49'E) ; WAM R134295, WAM R134627, WAM R134945, 1.5 km SW Vlamingh Head lighthouse, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=114.101944&amp;materialsCitation.latitude=-21.808332" title="Search Plazi for locations around (long 114.101944/lat -21.808332)">North West Cape</a>, WA (21°48'30"S, 114°06'07"E) ; WAM R132481, WAM R132483, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=114.01667&amp;materialsCitation.latitude=-22.05" title="Search Plazi for locations around (long 114.01667/lat -22.05)">Shothole Canyon</a>, Cape Range National Park, WA (22°03'S, 114°01'E) .</p><p>Diagnosis. A very small (to 27.7 mm SVL) species of Crenadactylus with long (HL/SVL 0.30–0.35) and narrow (HW/HL 0.43–0.50) head. Rostral in full contact with nostril, internasal (if present) not extending beyond supranasal, 2 small postmentals, dorsal scales heterogeneous with scattered enlarged keeled tubercles, males with 4 pre-cloacal pores with innermost pore-bearing scales separated by a small intervening scale, no enlarged tubercles on original tails. Ground colour light to dark greyish-brown, dorsal pattern consists of weakly-contrasting pale and dark longitudinal stripes, small pale spots may be present in dorsolateral zone, lateral zones grey with at most faint uniform stippling (not forming irregular lines), ventrum pale off-white with stippling absent to weak.</p><p>Description of holotype. WAM R174304, an adult male with the following meristics (in mm): SVL 24.8; TailL 11.3 (original); HeadL 7.9; HeadW 3.9; HeadD 2.3; SnL 2.2; ILL 11.2; SupLab 8 (left), 8 (right); InfLab 9, 9; internasals 1; PCP 4.</p><p>Slender, small-bodied with elongate body, round to oblong in cross section, ~1.5 times wider than deep; head narrow (HeadW/HeadL 0.49), elongate (HeadL/SVL 0.32), moderately depressed (HeadD/HeadL 0.29); in dorsal view, widest just posterior to eye, slightly wider than widest portion of body; head not distinct from neck, neck weakly constricted; loreal region weakly convex, weakly concave around nasal region; snout elongate (SnL 2.2; SnL/HeadL 0.40), round tip; eye moderately large; rostral ~3 times wider than high, in broad contact with nostril, dorsal edge concave to accommodate first supranasal, rostral groove ~60% of rostral length from dorsal edge below internasal notch; internasal tiny, ~one-fifth the size of first supranasal in area; supranasals two per side; first supranasal (in contact with rostral) oblong, transversely oriented, ~2.5 times wider than high, in broad contact with nostril; second (upper) supranasal small, rounded, ~one-third the size of first supranasal; nostril small, ~one-third size of first supranasal, directed dorsolaterally, in contact with rostral, first and second supranasals, three postnasals, and first supralabial; postnasals 3; supralabials 8 on both sides; antero-dorso edge of first supralabial in contact with nostril, postero-dorso edge notched by third postnasal and first loreal; second supralabial rectangular, wider than high, length equal to length of first supralabial; supralabials 3–8 smaller than first and second, gradually decreasing in size, all wider than high; scales on crown small and flattened, ~equal to size of smaller dorsal scales, ~one-sixth the size of enlarged dorsal tubercles, increasing in size anteriorly onto snout; scales on snout flat, enlarged, ~1.5–2.0 times the size of scales on crown; supraocular scales enlarged; two enlarged conical scales at postero-dorso edge of eye; mental trapezoid-shaped, widest anteriorly, narrowing slightly to posterior edge, terminates just beyond ventral edge of first infralabials; postmentals two, round in shape, ~equal to size of scales posterior to postmental; gulars granular, rounded; infralabials 9 on each side, first largest, square-shaped, gradually decreasing in height and width posteriorly, 1–3 square, 4–9 rectangular, wider than tall.</p><p>Limbs short, distance between limbs nearly half of SVL (ILL/SVL 0.45), nearly equal to tail length (ILL/TailL 0.99); scales on dorsal surface of forelimbs flattened and rounded, decreasing in size distally; scales on ventral surface of forelimbs similar to dorsal; scales on dorsal surface of hind limbs rounded and slightly raised, decreasing in size and becoming flattened and more rounded distally; scales on ventral surface of hind limbs flat and circular, imbricate, increasing in size distally; fingers 5; toes 5; claws absent from all digits; digits moderately long; fourth finger lamellae in 6 rows; fourth toe lamellae in 8 rows; in dorsal view, enlarged scale between apical plates ~2 times the size of adjacent lateral scales; apical plates much wider than digit; one pair of ventral apical plates, circular, as wide as long.</p><p>Scales on dorsum heterogeneous: background of small homogeneous scales with scattered enlarged scales with prominent keeling; small dorsal scales homogeneous, slightly raised; enlarged tubercles scattered on dorsum and nuchal region, loosely aligned longitudinally, ~4–6 times the size of surrounding smaller scales in diameter, separated from one another by 1–4 smaller scales, enlarged tubercles have raised midline forming a moderate keel, tallest at posterior edge of scale; some enlarged tubercles in dorso-lateral zone, absent on lateral surfaces; scales on ventrum homogeneous, rounded and flattened, ~2 times the size of smaller scales present on dorsum, all similar in size and shape.</p><p>Tail original, nearly half of SVL (TailL/SVL 0.45), tapers gradually from widest point posterior to vent to narrow rounded point; round in cross-section; dorsal scales somewhat rounded and imbricate, variable in size and shape, ~0.25–0.75 the size of enlarged tubercles on dorsum; ventral tail scales similar to those of dorsal tail scales but more flattened; single enlarged post-cloacal spur on both sides of cloaca, ~6 times the size of surrounding scales; pre-cloacal pores 4, forming a partial chevron, pore-bearing scales notched on posterior edge to accommodate pore, two pores on each side separated by a single intervening poreless scale.</p><p>Colouration and pattern. In life, ground colour light brownish-grey with moderately-contrasting dark brown longitudinal stripes, all stripes with some intermingling of dark and pale scales, some hints of orange on enlarged tubercles. Dark paravertebral stripes enclose a narrow pale vertebral stripe from the crown to posterior to hindlimbs where the paravertebral stripes join; pale dorsolateral stripe from posterior edge of eye to tail, small widely-spaced clusters of orange-tinted enlarged tubercles above hindlimbs and before tail, stripe continuing forwards anterior to eye as pale canthal stripe; above pale canthal stripe a dark triangular patch on snout; crown with a mixture of dark brown and black scales; dark brown dorsolateral stripe from posterior edge of eye to tail, continuing forwards on snout as dark loreal stripe; labial scales pale with dark mottling; lateral surfaces light brown with dark variegations suggesting a lateral line; limbs a mixture of light and dark brown scales with peppering of black scales occasionally forming short lines; ventrum pale off-white with dark stippling on underside of limbs, lower abdomen and between arms, elsewhere lightly stippled; original (see below) tail light to medium brown with scattered black scales. In preservative, colours faded but all stripes discernible.</p><p>Variation. Most specimens examined conformed to the holotype, with the following exceptions. Enlarged dorsal tubercles tended to not occur anterior to the ear openings, but in some individuals they occurred up to the level of the eyes, especially behind the eyes. Most individuals lacked tails (original or regrown), and we believe the holotype tail is an original based on the scalation and darker colouration compared to regenerated tails in our limited sample. Regrown tails tended to be paler with smaller scales compared to the holotype tail. WAM R134627, a male, has an atypical pattern of three pre-cloacal pores in a single row, but the scales in this area were irregularly-shaped. Females differed from males in that they lacked pre-cloacal pores and the cloacal spurs tended to not protrude laterally as much as those of males.</p><p>Colouration and pattern. Of the few individuals photographed in life, most of the variation is in shade—from light tan or brown to dark greyish brown. Other variable features are the small clusters of orange-tinted scales within the pale dorsolateral stripe near the legs and base of tail, ranging from absent to conspicuous. Occasionally the upper lateral zone has a dark edge that forms a weakly-defined additional thin lateral stripe, continuing anteriorly from forelimb to ear opening. In contrast to the holotype, the pale spots on the limbs are more conspicuous in most other individuals.</p><p>Distribution. Restricted to the Cape Range of the North West Cape on the Western Australian coast near Exmouth (Fig. 2). The southernmost records are from Yardie Creek, where it is sympatric with C. occidentalis sp. nov. (Fig. 2). Several records are from Charles Knife Road and Shothole Canyon, with additional records from near Vlamingh Head and the lighthouse at the northern end of the Cape Range. All save one record are from within the boundaries of the Cape Range National Park.</p><p>Habitat. Most habitat notes indicate collection from living or dead spinifex clumps, and usually in limestone gorges but with some records from coastal dunes. One specimen was collected from a dead log (WAM R88600). The holotype was collected from the upper edge of a rocky gorge with scattered shrubs and spinifex. Etymology. The species name tuberculatus (Latin) is in reference to the enlarged tubercles on the dorsum of this species, unique within Crenadactylus .</p><p>Remarks. No other Crenadactylus has heterogeneous dorsal scales as observed in C. tuberculatus sp. nov. (Fig. 3 B). The function of these scales is unknown. This species is also Australia’s smallest gecko species (maximum SVL &lt;28 mm).</p></div>	https://treatment.plazi.org/id/03AABA6D1810AA73FF5BFDFCFE0663CF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Doughty, Paul;Ellis, Ryan J.;Oliver, Paul M.	Doughty, Paul, Ellis, Ryan J., Oliver, Paul M. (2016): Many things come in small packages: Revision of the clawless geckos (Crenadactylus: Diplodactylidae) of Australia. Zootaxa 4168 (2): 239-278, DOI: 10.11646/zootaxa.4168.2.2
03AABA6D1815AA71FF5BFE10FC0B66C3.text	03AABA6D1815AA71FF5BFE10FC0B66C3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Crenadactylus pilbarensis	<div><p>Crenadactylus pilbarensis sp. nov.</p><p>Pilbara clawless geckos</p><p>Figs. 3 F, 4E, 14, 15</p><p>[‘Pilbara’ of Oliver et al. (2010)]</p><p>Holotype. WAM R132630, adult female, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=116.799446&amp;materialsCitation.latitude=-20.59278" title="Search Plazi for locations around (long 116.799446/lat -20.59278)">Burrup Peninsula</a>, Western Australia (20°35'34"S, 116°47'58"E), collected by B. Maryan, 31 May 1998. Fixed in 10% formalin, stored in 70% ethanol at WAM . Liver sample stored at the Evolutionary Biology Unit, South Australian Museum, Adelaide (SAMA) .</p><p>Paratypes (6). WAM R 100988, 80 km south of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=122.083336&amp;materialsCitation.latitude=-22.333334" title="Search Plazi for locations around (long 122.083336/lat -22.333334)">Telfer</a>, WA (22°20'S, 122°05'E) ; WAM R132540, Burrup Peninsula, WA (20°36'45"S, 116°47'37"E); WAM R132627, WAM R132629, Burrup Peninsula, WA (20°35'34"S, 116°47'58"E); WAM R 140394, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=121.00222&amp;materialsCitation.latitude=-21.321945" title="Search Plazi for locations around (long 121.00222/lat -21.321945)">Deepdale Outstation</a>, Robe River Station, WA (21°43'S, 116°11'E) WAM R160059, 55 km east-south-east of Meentheena Outcamp, WA (21°19'19"S, 121°00'08"E) .</p><p>Diagnosis. A small (to 28.4 mm SVL) species of Crenadactylus . Rostral in full contact with nostril, internasal (if present) not extending beyond supranasal, single greatly enlarged postmental, dorsal scales homogeneous and moderately keeled, 6 pre-cloacal pores in males with innermost pore-bearing scales in contact, no enlarged tubercles on original tails. Ground colour light brown; dorsal pattern consists of well-defined pale and dark longitudinal stripes, scattered clusters of 1–3 pale scales, lateral zones light grey to tan with stippling occasionally forming 1 or 2 broken or weakly defined lateral lines; ventrum pale off-white with weak to moderate stippling.</p><p>Description of holotype. WAM R132630, adult female with the following meristics (mm): SVL 27.0 mm; TailL 24.3 mm (partially regenerated); HeadL 8.4; HeadW 4.1; HeadD 2.5; SnL 3.3; ILL 13.0; SupLab 8 (left), 9 (right); InfLab 8, 9; internasals 1; PCP 0.</p><p>A slender, small-bodied gecko; body elongate, oblong in cross section, ~1.5–2.0 times wider than deep; head narrow (HeadW/HeadL 0.49), elongate (HeadL/SVL 0.31), moderately depressed (HeadD/HeadL 0.30); in dorsal view, widest at ear openings, not wider than widest portion of body; head not distinct from neck, neck very weakly constricted; loreal region weakly convex, concave around nasal region; snout elongate (SnL 3.3; SnL/HeadL 0.39), rounded tip; eye moderately large; rostral ~3 times wider than high, in broad contact with nostril, slightly scalloped dorsally to accommodate first supranasal, notched by internasal, rostral groove originates from dorsal edge of scale, ~40% of rostral height; internasal tiny, ~one-fifth the size of first supranasal in area; supranasals 2 per side; first supranasal oblong, transversely oriented, ~2 times wider than high, in broad contact with nostril; second (dorsal) supranasal small, rounded, tapering to blunt point at nostril contact, ~half the size of first supranasal; nostrils small, ~one-third size of first supranasal, directed dorsolaterally, in contact with rostral, first and second supranasals, postnasals, and first supralabial; postnasals 2, equal to size of second supranasal; supralabials 8 on both sides, antero-dorso edge of first supralabial in contact with nostril; second supralabial rectangular, wider than high, length equal to length of first supralabial, supralabials 3–8 smaller than first and second, gradually decreasing in size, all wider than high; scales on crown small and flattened, ~one-third the size of dorsal scales, increasing in size anteriorly onto snout; scales on snout flat, enlarged, ~2–3 times the size of scales on crown; supraocular scales enlarged; single enlarged conical scale at postero-dorso edge of eye; mental trapezoid-shaped, widest anteriorly, narrowing slightly to concave posterior edge, terminates equal to ventral edge of first infralabials; single enlarged postmental, hexagonal in shape, ~2 times area of scales posterior and lateral to enlarged postmental, ~4 times larger than other gular scales; gulars granular, rounded, ~0.25–0.50 the size of scales on ventrum; infralabials 9 on each side, first largest, square-shaped, gradually decreasing in height and width posteriorly, infralabials 1–3 square or rectangular, 4–9 rectangular, wider than high.</p><p>Limbs short, distance between limbs nearly half of SVL (ILL/SVL 0.48), slightly over half the length of tail (ILL/TailL 0.54); scales on dorsal surface of forelimbs imbricate, flattened and rounded, decreasing in size distally; scales on ventral surface of forelimbs similar to dorsal; scales on dorsal surface of hind limbs rounded and slightly raised, decreasing in size and becoming more rounded distally; scales on ventral surface of hindlimbs flat and circular, slightly larger in area than dorsal hindlimb scales; fingers 5; toes 5; claws absent from all digits; digits moderately long; fourth finger lamellae in 6 rows; fourth toe lamellae in 8 rows; in dorsal view, enlarged scale between apical plates 1.5 times the size of adjacent lateral scales; apical plates much wider than digit; two pairs of dorsal apical plates; one pair of ventral apical plates, circular.</p><p>Scales on dorsum small and homogeneous, round to oblong, juxtaposed; scales slightly raised, moderately keeled, highest point at posterior edge of scale, widest at midpoint of scale, becoming flatter and imbricate laterally; scales on ventrum homogeneous, flat, imbricate, equal to 1.5 times the size of dorsal scales in area, widest at midpoint, tapering to a rounded or flattened posterior edge.</p><p>Tail partially regenerated, last 40% of tail regenerated, slightly shorter than SVL (TailL/SVL 0.90), tapers gradually from widest point posterior to vent, more abruptly in posterior quarter of tail; round in cross-section; scales homogeneous, imbricate and flat, square to oblong in shape; subcaudal scales flattened, ~1.5–2 times the size of dorsal tail scales; single enlarged spur on both sides of cloaca, ~3–4 times the size of surrounding scales.</p><p>Colouration and pattern. In preservative, pattern comprised of alternating light and dark longitudinal stripes on dorsum and sides; light brown-tan vertebral stripe narrow, 2–4 scales wide, scales lightly stippled with darker brown, originating on back of head, extending length of body, terminates at base of tail where paravertebral stripes merge; paravertebral stripes wider than vertebral, 3–5 scales wide, medium brown, originating from postero-dorsal edge of eye, curving around posterior of head to vertebral stripe, continuing parallel with vertebral stripe for length of body, tapering from hindlimbs to merge and form single broken line on tail; light brown-tan dorso-lateral stripe 3–4 scales wide, extends from postero-dorsal edge of eye, above forelimbs, parallel with vertebral stripe for length of body, over hindlimbs and onto tail, faded and broken on tail; dorsolateral stripe narrow, 3–4 scales wide, ~equal width to paravertebral stripe, dark brown, same as paravertebral, extends from posterior edge of eye above ear opening, above forelimbs along length of body, above hindlimbs and onto tail, ~one-half of tail length; pale lateral zone below dorsolateral stripe with narrow, solid, dark brown lateral stripe between limbs, 2–3 scales wide; weak stripe below lateral stripe on ventro-lateral edge, extending anteriorly from forelimb to ventral edge of ear opening; ventrum pale, heavily stippled with brown flecks; dark triangular patch on snout, crown a mixture of light and dark scales, dark-edged by extension of paravertebral stripes to eyes, top of eyes purplish, dark brown scales below eye on orbit; labials pale, heavily stippled with brown flecks; pale canthal stripe, extension of pale dorsolateral stripe from eye to rostral scale, below a dark loreal stripe, extension of dark dorsolateral stripe; dorsal surface of limbs with mixed pale, light brown and dark brown scales, scattered clusters of pale scales resulting in some spotting, extending onto digits; dorsal stripes (with exception of vertebral) continue to tail but broken up.</p><p>Variation. Most specimens conformed to the holotype, with some of the following exceptions. Males possessed 6 pre-cloacal pores (one individual with 4) in a continuous row. Pores formed by a notch in the posterior edge of scale. Females lacked pre-cloacal pores. In males the cloacal spurs are larger than those of females and protrude further than for females. Regrown tails tended to be paler with smaller scales compared to the tail of the holotype.</p><p>The diagnostic character of a greatly enlarged postmental was unambiguous in all specimens examined, although the shape tended to vary from round to oblong (Fig. 3 F). Keeling on the dorsal scales also varied from weak keels to moderately strong.</p><p>Colouration and pattern. In life, light to dark brown ground colour with contrasting dark and pale longitudinal stripes. In preservative, thin vertebral stripe pale tan-grey, paravertebral stripes to either side emanating from top of eye and joining just before base of tail; pale dorsolateral stripe from antero-dorso portion of eye to tail, continuing anteriorly from eye as pale canthal stripe; dark brown triangular patch on snout continuous with dark brown crown, intermixed with paler scales; dark dorsolateral stripe from ventro-dorso portion of eye to above limbs to tail, anterior to eye forming dark loreal stripes; labials pale, heavily stippled with dark brown; lateral surfaces pale with thin dark brown stripes, uppermost more defined, lower lateral stripe can be extensively broken and indistinct; stippling on pale ventrum variable; limbs a mixture of dark and light brown scales with small clusters of pale scales apparent; regenerated tails a mixture of dark and pale brown scales, forming short lines or showing no pattern.</p><p>Distribution. Restricted to the Pilbara region of Western Australia (Fig. 2). Most records are from the Pilbara craton, although many specimens have also been collected from the Burrup Peninsula. Absent from offshore islands except Dolphin Island that is separated from the Burrup Peninsula by a channel only several hundred metres wide. Northern records are from Shay Gap and Meentheena Conservation Reserve. The southern-most records are from the Barlee Range in the south-west and Kumarina in the south-east. Outlying records to the east are from Karlamilyi (Rudall River) National Park and Jiggalong.</p><p>Habitat. Collection records indicate this species occurs in spinifex clumps on rocky hills and in gullies on the mainland, but sandy substrates for the Dolphin Island specimens.</p><p>Etymology. Named for the Pilbara region where this species occurs.</p></div>	https://treatment.plazi.org/id/03AABA6D1815AA71FF5BFE10FC0B66C3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Doughty, Paul;Ellis, Ryan J.;Oliver, Paul M.	Doughty, Paul, Ellis, Ryan J., Oliver, Paul M. (2016): Many things come in small packages: Revision of the clawless geckos (Crenadactylus: Diplodactylidae) of Australia. Zootaxa 4168 (2): 239-278, DOI: 10.11646/zootaxa.4168.2.2
03AABA6D1816AA7EFF5BFF72FC32676E.text	03AABA6D1816AA7EFF5BFF72FC32676E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Crenadactylus rostralis Storr 1978	<div><p>Crenadactylus rostralis Storr, 1978</p><p>South-west Kimberley clawless geckos Figs. 3 H, 4F, 16, 17</p><p>Crenadactylus ocellatus rostralis Storr, 1978 [‘Kimberley A’ of Oliver et al. (2010, 2012b)]</p><p>Holotype. WAM R32154, adult male, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=125.65&amp;materialsCitation.latitude=-18.116667" title="Search Plazi for locations around (long 125.65/lat -18.116667)">Geikie Gorge</a>, Western Australia (18°07'S, 125°39'E), collected by W.H. Butler, April 1968. Fixed in 10% formalin, stored in 70% ethanol at WAM.</p><p>Paratypes (17). WAM R23058*, 43 km SE <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=127.25&amp;materialsCitation.latitude=-18.45" title="Search Plazi for locations around (long 127.25/lat -18.45)">Halls Creek</a>, WA (18°27'S, 127°15'E) ; WAM R26824–26, Napier Range, WA (17°20'S, 124°50'E); WAM R27388–92, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=123.933334&amp;materialsCitation.latitude=-18.033333" title="Search Plazi for locations around (long 123.933334/lat -18.033333)">Mount Anderson</a>, WA (18°02'S, 123°56'E) ; WAM R46058– 59*, near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=126.75&amp;materialsCitation.latitude=-18.716667" title="Search Plazi for locations around (long 126.75/lat -18.716667)">Wolfe Creek Meteorite Crater</a>, 122 km SW Halls Creek, WA (18°43'S, 126°45'E) ; WAM R46115*, 11 km E <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=126.96667&amp;materialsCitation.latitude=-18.633333" title="Search Plazi for locations around (long 126.96667/lat -18.633333)">Margaret River Homestead</a>, WA (18°38'S, 126°58'E) ; WAM R56841–45*, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.75&amp;materialsCitation.latitude=-16.25" title="Search Plazi for locations around (long 128.75/lat -16.25)">Lake Argyle</a>, WA (16°15'S, 128°45'E) .</p><p>Some registration numbers presented in the original description are in error, with WAM R56441–45 associated with different species. The error originates from a minor numerical error in the registration numbers presented by Storr (1978), and underlined here. The correct numbers associated with the paratype specimens are WAM R56841– 45. Nine paratypes (marked with an asterisk) designated by Storr (1978) fall within the distribution of lineages associated with the C. naso complex identified by Oliver et al. (2012b), and are not true C. rostralis .</p><p>Diagnosis. A moderate-sized (to 32.7 mm SVL) species of Crenadactylus . Rostral not in contact with nostril, 2–3 internasals present, not extending beyond supranasal, postmentals granular, dorsal scales homogeneous and weakly keeled, 4–6 pre-cloacal pores, innermost pore-bearing scales in contact, no enlarged tubercles on original tails. Ground colour light greyish brown; dorsal pattern consists of well-defined pale and dark longitudinal stripes, lateral zone light grey with usually two moderate to well-defined dark lines between limbs, stippled with single pale scales; ventrum pale grey to white with moderate stippling forming longitudinal lines.</p><p>Description of holotype. WAM R32154, adult male with the following meristics (in mm): SVL 29.8; TailL 27.5 (original); HeadL 9.3; HeadW 4.6; HeadD 2.8; SnL 3.3; ILL 14.5; SupLab 9 (left), 9 (right); InfLab 9, 9; internasals 3.</p><p>A slender, small-bodied gecko; body elongate, oblong in cross section, ~1.5–2 times wider than deep; head narrow (HeadW/HeadL 0.50), elongate (HeadL/SVL 0.31), moderately depressed (HeadD/HeadL 0.30); in dorsal view, widest between eye and ear openings, narrower than widest portion of body; head not distinct from neck, neck weakly constricted; loreal region weakly convex, weakly concave around nasal region; snout elongate (SnL 3.3; SnL/HeadL 0.36), rounded tip; eye moderately large; rostral ~3 times wider than high, excluded from nostril contact by first supranasal, dorsal edge concave to accommodate first supranasal, slightly notched by median internasal, rostral groove originates from dorsal edge of scale below internasal, ~40% of rostral height; internasals 3, median internasal tiny; supranasals 2 per side; first oblong, transversely-oriented, ~2.0–2.5 times wider than high, in broad contact with nostril; second supranasal small, rounded, ~size of first supranasal; nostril small, ~1/3 size of first supranasal, directed dorso-laterally, in contact with both supranasals, 3 postnasals and first supralabial; postnasals 3, first and third ~half the size of first supranasal, second small, ~1/3 of first and third; supralabials 9 on both sides, antero-dorso edge of first supralabial in contact with nostril; second supralabial rectangular, wider than high, supralabials 3–5 smaller than first and second, all wider than high, sixth supralabial ~equal to first supralabial, supralabials 7–9 smaller than first two, gradually decreasing in size posteriorly; scales on crown small and granular, ~half the size of dorsal scales, becoming flattened and increasing in size anteriorly onto snout; scales on snout flat, enlarged, ~1.5–4.0 times the size of scales on crown; supraocular scales enlarged; 3 enlarged conical scales at postero-dorso edge of eye; mental trapezoidal, widest anteriorly, narrowing slightly to rounded posterior edge; gulars granular, ~1/3 to 1/4 the size of scales on ventrum; infralabials 9 on each side, first trapezoidal, slightly smaller than second, infralabials 2–4 square, gradually decreasing in size posteriorly, infralabials 5–9 rectangular, wider than high, gradually decreasing in height and width posteriorly.</p><p>Limbs short, distance between limbs nearly half of SVL (ILL/SVL 0.48), just over half of tail length (ILL/ TailL 0.52); scales on dorsal surface of forelimbs flat and rounded, imbricate, decreasing in size and becoming more granular distally; scales on ventral surface of forelimbs similar to dorsal; scales on dorsal surface of hind limbs rounded and slightly raised, imbricate, decreasing in size and becoming more granular distally; scales on ventral surface of hind limbs circular and slightly raised, increasing in size and becoming flatter and imbricate distally, granular on feet; fingers 5; toes 5; claws absent from all digits; digits moderately short; fourth finger lamellae 6 rows; fourth toe lamellae 8 rows; in dorsal view, enlarged scale between apical plates ~equal to 2 times the size of adjacent lateral scales; apical plates much wider than digit; two pairs of dorsal apical plates; one pair of ventral apical plates, circular.</p><p>Scales on dorsum homogeneous, small, feebly keeled, juxtaposed, scales slightly raised with highest point at posterior edge of scale, widest at midpoint of scale, becoming flatter and imbricate laterally; scales on ventrum homogeneous, flat, imbricate, equal to 1.5–2.0 times the size of dorsal scales in area, widest at midpoint, rounded posteriorly.</p><p>Tail original, nearly equal to SVL (TailL/SVL 0.92), tapers gradually from widest point posterior to vent to narrow rounded point; round in cross-section; scales homogeneous, imbricate and flat, variable in size and shape, ~3–4 times the size of scales on dorsum; ventral tail scales similar to dorsal tail scales, flattened and imbricate, ~1.5–3 times the size dorsal tail scales; single enlarged spur on both sides of cloaca, ~5–6 times the size of surrounding scales; pre-cloacal pores 6, forming a slight curve, median innermost pore-bearing scales in contact, pore-bearing scales notched on posterior edge to accommodate pore.</p><p>Coloration and pattern. In preservative, colour faded to light brown. A thin pale vertebral stripe 2–3 scales wide, bracketed by medium brown paravertebral stripes 3–4 scales wide; a light brown dorsolateral zone 6–8 scales wide, with several widely-spaced pale scales, continuing forwards through eye and onto snout as pale canthal stripe; dark brown dorsolateral stripes 3–4 scales wide, in contact with dorsal edge of hindlimb, continuing forwards to eye and onto snout as loreal stripe; light brown lateral zone below dark dorsolateral stripe, 3–4 scales wide; ventro-lateral stripe medium brown, weakly defined, 3–5 scales wide; crown and snout medium brown, bordered by light brown stripe (extension of dorsolateral zone); supralabials stippled with dark brown, continuing as weakly-defined dark brown stripe to arm; ventrum pale with 4 longitudinal stripes, outer pair darker, 2–3 scales wide, inner pair narrower, 1–2 scales wide; arms with dark variegations on dorsal surface, legs with weaklydefined dark brown longitudinal lines, ventral surfaces of limbs with irregular stippling; basal portion of tail (original) with dark brown stripes continuing from paravertebral and dorsolateral stripes on body, ventral portion of tail with 2 dark brown stripes.</p><p>Variation. Other specimens generally corresponded to the holotype specimen, with the following exceptions. The degree of keeling on the dorsal scales ranged from feebly keeled to smooth and rounded. Females lacked pre-cloacal pores and the spurs were smaller and less projecting.</p><p>Colouration and pattern. In life (based on recent photographs by PMO), background colour grey-brown with a series of alternating pale and dark alternating longitudinal stripes; pale grey vertebral stripe bordered by medium dark brown paravertebral stripes; pale to medium grey dorsolateral zone, continuing forwards through eye as pale canthal stripe; dark brown lateral stripe, continuing through eye on snout as poorly-defined loreal stripe; light grey ventrolateral zone with occasionally poorly-defined thin dark stripe near ventrolateral edge of body; top of head dark brown with some pale mottling; labials pale, stippled with dark brown. Dark brown paravertebral and lateral stripes usually with scattered pale scales. Lateral stripe usually darker than paravertebral stripes, but can also be of similar shade. Ventrolateral zone can be much paler than dorsolateral zone. Limbs dark brown with variable small pale spots; original tails continue dorsal pattern, regenerated tails with a mixture of pale and dark scales not forming conspicuous lines.</p><p>In preservative, pattern consists of alternating dark and light stripes. Silvery-grey vertebral stripe, narrow, 2–3 scales wide; medium grey-brown paravertebral stripes, bordered by dark brown edges, 6–7 scales wide; pale dorsolateral stripe, 4–5 scales wide, starts at postero-dorsal edge of eye, continues above limbs to tail; dark brown dorso-lateral stripe extends posteriorly from back of eye to top of hindlimb, anteriorly extends to top of ear opening; below dorsolateral line are two thin, ventro-lateral stripes; pale lateral zone divided by thin dark lateral stripe between limbs, forming pale line below dorsolateral that extends from angle of jaw across top of forelimbs and through hindlimb. Ventrum lightly stippled, one or two weakly defined brown lines towards outer edge of ventrum. Pale canthal and dark loreal stripes weakly defined, continuation of pale and dark dorsolateral stripes; top of eyes purplish; labials pale and heavily stippled with brown flecks; posterior edge of jaw heavily stippled, continuing as lateral line below ear opening to forelimb. Limbs dark, back of thigh has strong pale stripe, diffuse with small clusters of pale scales.</p><p>Distribution. Genotyped specimens were all collected from the Devonian Reefs Ranges in the south-western Kimberley region (Fig. 2), extending from the Napier Ranges in the north-west to the Pillara Range on Gogo Station in the south-east. Additional (not genotyped) records of Crenadactylus from Mt Anderson in the south-west (~ 40 km from Derby) and from the south-east Kimberley are here considered to belong to C. rostralis pending genetic data to resolve their status (Fig. 2). All samples formerly regarded as this species from elsewhere in the Kimberley are here considered part of the C. naso species complex.</p><p>Habitat. The few records with collection details all mention occurring in spinifex, including a number of recent samples that were raked from large clumps on and around limestone ranges (PMO, pers. obs.).</p><p>Etymology. The name rostralis is from the Latin rostrum, meaning beak or snout, in reference to this species’ distinctive configuration of having the rostral scale separated from the nostril.</p></div>	https://treatment.plazi.org/id/03AABA6D1816AA7EFF5BFF72FC32676E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Doughty, Paul;Ellis, Ryan J.;Oliver, Paul M.	Doughty, Paul, Ellis, Ryan J., Oliver, Paul M. (2016): Many things come in small packages: Revision of the clawless geckos (Crenadactylus: Diplodactylidae) of Australia. Zootaxa 4168 (2): 239-278, DOI: 10.11646/zootaxa.4168.2.2
03AABA6D181BAA7BFF5BFF72FDF66564.text	03AABA6D181BAA7BFF5BFF72FDF66564.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Crenadactylus naso Storr 1978	<div><p>Crenadactylus naso Storr, 1978</p><p>Crenadactylus naso species complex Northern clawless geckos</p><p>Figs. 4 G, 4H, 18, 19</p><p>Crenadactylus ocellatus naso Storr, 1978</p><p>[‘Kimberley B–G’ of Oliver et al. (2010, 2012b)]</p><p>Holotype. WAM R56206, adult male, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=125.78333&amp;materialsCitation.latitude=-14.5" title="Search Plazi for locations around (long 125.78333/lat -14.5)">Crystal Creek</a>, Western Australia (14°30'S, 125°47'E), collected by L.A. Smith and R.E. Johnstone, 2 November 1976. Fixed in 10% formalin, stored in 70% ethanol at WAM.</p><p>Paratypes (8). WAM R41373–74, Heywood Island, Bonaparte Archipelago, WA (15°20'S, 124°20'E); WAM R43220–21, WAM R43224, Mitchell Plateau, WA (14°57'S, 124°20'E); WAM R56185–87, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=125.78333&amp;materialsCitation.latitude=-14.5" title="Search Plazi for locations around (long 125.78333/lat -14.5)">Crystal Creek</a>, WA (14°30'S, 125°47'E) .</p><p>Diagnosis. A small (to 30.7 mm SVL), slender Crenadactylus . Rostral usually in narrow contact with nostril, sometimes excluded (especially in eastern Kimberley), internasals not extending beyond supranasals, gular scales granular, dorsal scales homogeneous and smooth to weakly keeled, 4–8 pre-cloacal pores in contact and forming gradual curve, enlarged tubercles on original tails. Ground colour light grey to dark brown; dorsal pattern consists of weak to well-defined pale and dark longitudinal stripes, dark stripes with scattered single pale scales; lateral zone light grey to tan, ventrum pale grey to off-white, lateral zone and ventrum immaculate to moderately stippled with occasional thin longitudinal lines.</p><p>Description of holotype. WAM R56206, adult male with the following meristics (in mm) and counts: SVL 26.8; TailL 22.2 (detached, partially regenerated); HeadL 7.7; HeadW 4.1; HeadD 2.5; SnL 3.3; ILL 11.6; SupLab10 (left), 9 (right); InfLab 9, 9; internasals 3; PCP 6.</p><p>A slender, small-bodied gecko; body elongate, oblong in cross section, ~1.5–2.0 times wider than deep; head narrow (HeadW/HeadL 0.53), elongate (HeadL/SVL 0.28), moderately depressed (HeadD/HeadL 0.32); in dorsal view, widest at ear openings, not wider than widest portion of body; head not distinct from neck, neck weakly constricted; loreal region weakly convex, concave around nasal region; snout elongate (SnL 3.3; SnL/HeadL 0.43), rounded tip; eye moderately large; rostral ~3.5 times wider than high, in narrow contact with nostril, dorsal edge faintly scalloped to accommodate first supranasals, rostral groove absent; internasals 3, variable in size, ~0.5–0.75 the size of first supranasals in area; supranasals two per side; supranasals on left side, first oblong, transverselyoriented, wider than long, ~1.5 times wider than long, in broad contact with nostril, second round, ~half the size of first, in broad contact with nostril; supranasals on right side, first oblong, transversely oriented, wider than long, ~1.75 times wider than long, in broad contact with nostril, second oblong, transversely-orientated, equal in size to first, in broad contact with nostril; nostrils small, directed dorsolaterally, in contact with rostral, first and second supranasals, postnasals and first supralabial; postnasals 3, oblong in shape, longer than high; supralabials 10 (left), 9 (right), antero-dorso edge of first supralabial in contact with nostril; second supralabial rectangular, wider than high, slightly longer than first supralabial, supralabials 3–10 (left)/9 (right) smaller than first and second, gradually decreasing in size, all wider than high; scales on crown small and flattened, ~half the size of dorsal scales, increasing in size anteriorly onto snout; scales on snout flat, enlarged, ~2–3 times the size of scales on crown; supraocular scales enlarged; two enlarged conical scales at postero-dorso edge of eye; mental trapezoid-shaped, widest anteriorly, narrowing slightly to concave posterior edge, posterior width ~one-third of anterior width; gulars granular, rounded, ~0.50–0.75 the size of scales on ventrum; infralabials 9 on each side, first largest, squareshaped, gradually decreasing in width and height posteriorly.</p><p>Limbs short, distance between limbs nearly half of SVL (ILL/SVL 0.43), slightly over half the length of tail (ILL/TailL 0.52); scales on dorsal surface of forelimbs imbricate, rounded and slightly raised, becoming flatter distally; scales on ventral surface of forelimbs similar to dorsal; scales on dorsal surface of hindlimbs rounded and slightly raised; scales on ventral surface of hind limbs rounded and slightly raised, becoming flatter distally; fingers 5; toes 5; claws absent from all digits; digits moderately long; fourth finger lamellae in 7 rows; fourth toe lamellae 8 rows; in dorsal view, enlarged scale between apical plates ~2 times the size of adjacent lateral scales, apical plates much wider than digit; one pair of ventral apical plates, circular.</p><p>Scales on dorsum small and homogeneous, round to oblong, widest at midpoint of scale, becoming flatter and imbricate laterally, juxtaposed; dorsal scales slightly raised, highest point at posterior edge of scale, weakly keeled; scales on ventrum homogeneous, flat, imbricate, equal to 1.5 times the size of dorsal scales in area, widest at midpoint, rounded posteriorly.</p><p>Tail detached, partially regenerated, last 60% of tail regenerated, slightly shorter than SVL (TailL/SVL 0.82), tapers gradually from widest point posterior to vent; round in cross-section; scales rounded and slightly raised, imbricate, 2 longitudinal rows of enlarged tubercles along dorsolateral edges of tail, similar in area to surrounding scales, but more prominently raised; ventral tail scales flattened and imbricate, ~equal to 1.5 times the size of dorsal tail scales; scales on regenerated portion rounded and slightly raised on dorsal surface, enlarged tubercles absent, ventral tail scales flat and imbricate; single enlarged raised post-cloacal spur on both sides of cloaca, ~1.5– 2 times the size of surrounding scales; pre-cloacal pores 6, arranged in a continuous gradual curve, innermost porebearing scales in contact, pore-bearing scales notched on posterior edge to accommodate pore.</p><p>Colouration and pattern. In preservative, colouration faded to medium brown, pattern comprised of alternating pale and dark longitudinal lines; pale vertebral stripe bordered by dark brown; medium brown paravertebral zone; pale dorsolateral stripes bordered with dark brown; pale and brown lateral stripes somewhat discernible; crown of head pale; top of snout dark brown with pale canthal stripe (extension of pale dorsolateral stripe anterior to eye) and dark brown loreal stripe (extension of paravertebral zone); labial scales pale; arms with feint longitudinal lines; hindlimbs with longitudinal dark and pale stripes; ventral surface light brown; original portion of tail with feint dark and pale stripes, ventral surface light brown; regenerated portion uniform light brown.</p><p>Variation. The holotype specimen is somewhat faded and shrivelled, whereas more recently-collected material has more vivid colouration and is in better condition. Within the complex, body size to 30.7 mm SVL, ILL/SVL 39–54%; the rostral-nostril contact varies from narrow to point contact and also exclude in some individuals; rostral groove varied from 0–40%, internasals usually 2 or 3 (occasionally 1 or 4), SupLab and InfLab 7–10; precloacal pores usually 6 (occasionally 4 or 8), innermost pore-bearing scales in contact. Dorsal scales homogeneous, smooth to weakly keeled.</p><p>In life, light yellowish-grey background colour with alternating pale and dark longitudinal lines; pale vertebral stripe often forks on back of head; pale dorsolateral zone with small spots of 1–2 pale scales dotted at regular intervals and continuing on to tail on enlarged tubercles.</p><p>In preservative, pale vertebral stripe, bordered by medium brown paravertebral stripes with a thin dark brown border, pale dorsolateral zone originates on postero-dorsal edge of eye and continues on to tail, below dorso-lateral zone is a brown lateral zone with dark border, greyish-brown weakly defined lateral stripe between limbs, dorsum has variably expressed small isolated scattered pale scales, sometimes bordered by dark brown scales forming small ocelli, limbs dark medium dark brown with scattered groups of pales scales forming spots, snout dark brown, pale canthal and dark loreal stripes prominent; labials heavily stippled; ventrum pale, finely stippled, especially on chest and neck region, medium peppering of dark brown, sometimes forming weak longitudinal lines; pale enlarged tubercles on tail, occasionally with black scales adjacent.</p><p>Distribution. As defined here, this species complex occurs throughout most of the Kimberley, except for the Devonian Reef ranges in the south-west which are occupied by C. rostralis, and a small number of other records of uncertain status from the southern Kimberley (Fig. 2). We tentatively assign populations from the Northern Territory and Queensland to this taxon as well.</p><p>Habitat. Collected from rocky areas with spinifex, including screes, sandstone outcrops and gorges. Numerous specimens were collected by burning spinifex, with occasional records from vine thickets and more open woodland.</p><p>Etymology. The word naso (Latin) means ‘long-nosed’ in reference to the long snout (Storr et al. 1983).</p><p>Remarks. Despite extensive searches for diagnostic characters among lineages in the C. naso species complex, we could not find non-overlapping characters amongst the few existing voucher specimens. Rather than delay the resolution of the southern and arid zone taxa, by redescribing C. rostralis we have hopefully isolated the remaining taxonomic issues within Crenadactylus to a monophyletic unit.</p><p>The resolution of the C. naso species complex may be possible through examination of internal characters, and would benefit from a larger series of voucher specimens (with photos in life) to better understand patterns of morphological variation in the group.</p></div>	https://treatment.plazi.org/id/03AABA6D181BAA7BFF5BFF72FDF66564	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Doughty, Paul;Ellis, Ryan J.;Oliver, Paul M.	Doughty, Paul, Ellis, Ryan J., Oliver, Paul M. (2016): Many things come in small packages: Revision of the clawless geckos (Crenadactylus: Diplodactylidae) of Australia. Zootaxa 4168 (2): 239-278, DOI: 10.11646/zootaxa.4168.2.2
