taxonID	type	description	language	source
99254A1C54A67A694A35EE16756A3BD0.taxon	diagnosis	Diagnosis. This is the largest bodied of the three Haleakala species in this group (Fig. 10 A); standardized body length 4.5 - 4.8 mm versus 3.5 - 4.6 mm for the other two species below. Dorsal microsculpture is more developed in this species, with the elytral disc covered with an elongate transverse mesh and parallel lines, and the pronotal disc bearing an evident transverse mesh. Setal formula 2 1 1 (2) 0.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
99254A1C54A67A694A35EE16756A3BD0.taxon	description	Description (n = 3). Head capsule with frontal grooves straight, bordered by a lateral carina and mesal wrinkles; neck flat to slightly concave; eyes convex, largely covering ocular lobe, ocular ratio 1.52 - 1.55, ocular lobe ratio 0.89 - 0.94; labral anterior margin very shallowly emarginate; antennae filiform, antennomeres 2 - 3 covered with sparse pelage of small setae; mentum tooth narrow with acute sides, apex tightly rounded. Pronotum with lateral seta present, basal seta absent; MPW / BPW = 1.52 - 1.66; MPW / PL = 1.11 - 1.23; hind angle right to slightly acute; lateral margin convergent to subparallel anterad hind angle; median base with small punctures, sparse medially, denser laterally; basal margin broadly, slightly convex between laterobasal depressions; median longitudinal impression finely incised, shallow; anterior transverse impression distinct, slightly punctate in middle half; anterior callosity slightly convex, traversed by shallow wrinkles; front angles slightly protruded, tightly rounded; APW / BPW = 1.0 - 1.04; lateral marginal depression moderately narrow, flat near front angle, edge upturned; laterobasal depression a continuation of lateral depression, surface irregularly punctate. Proepisternum with 6 small punctures along hind marginal groove; prosternal process with narrow median impression, lateral margins a narrow bead anterad. Elytra broadly ovoid, convex, suture elevated relative to disc; basal groove nearly straight laterad scutellum, humeral angle subangulate, defined by a hitch at base of lateral depression; humeri broadly rounded, MEW / HuW = 2.11 - 2.14; elytra broad relative to pronotum and head, MEW / MPW = 1.50 - 1.59, MEW / MHW = 2.18 - 2.29; parascutellar seta present; parascutellar striole with 2 - 4 punctures, striole shallow between punctures; juncture of sutural intervals each side upraised, producing a median callus; depth of sutural or first stria subequal to 2 nd stria from base to apex; discal stria finely punctate basally, narrow and smooth apically; lateral striae punctate basally, the punctures small, slightly expanding striae in basal half, punctures absent apically; intervals 2 - 7 moderately convex; all striae finely incised apically; 8 th interval laterad 7 th stria not more convex apically than other intervals; either 1 or 2 dorsal elytral setae, if two, then setae at 0.10 x and 0.28 - 0.33 x elytral length, if one seta, then situated 0.24 - 0.30 x elytral length; apical and subapical setae absent; lateral elytral setae arranged as 7 + 6 (anterior and posterior series); elytral marginal depression narrow, lateral margin upturned; subapical sinuation broadly excavated, shallow, internal plica visible from dorsal view. Mesepisternum punctate, ~ 13 punctures in 2 - 3 vertical rows; metepisternal medial length / maximum width = 1.39; metepisternum / metepimeron suture distinct. Abdomen with irregular longitudinal wrinkles on ventrites 1 - 4, suture between ventrites 2 and 3 complete; males with 2 apical abdominal setae, females with 4 equally spaced setae plus 4 short setae arranged in a median trapezoid. Legs-metatarsomere 1 / metatibial length ratio = 0.195; metatarsomere 4 lobe length 1.5 x medial tarsomere length, subapical and apical setae present; metatarsal dorsolateral sulci very narrow, shallow, evident on mt 1 and mt 2 only. Microsculpture of head capsule transverse, vertex with transverse mesh and fine wrinkles; pronotal disc with evident, reflective transverse-mesh microsculpture; pronotal median base with reflective isodiametric mesh in transverse rows; elytral disc with elongate transverse mesh and parallel lines, apex with evident transverse mesh; metasternum covered with transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse sculpticells. Coloration of vertex brunneous with a slight piceous cast; antennomeres 1 - 3 flavous, 4 - 11 rufobrunneous; pronotal disc rufobrunneous, pronotal margins slightly, broadly paler; proepipleuron rufoflavous, proepisternum brunneous with piceous cast; elytral disc rufobrunneous, darkest behind middle, sutural interval paler, rufous basally, flavous apically; elytral margins concolorous with disc basally, broadly paler apically; elytral apex broadly flavous, flavous coloration extended anterad along suture; elytral epipleuron rufoflavous, metepisternum brunneous; abdominal ventrites 1 - 3 brunneous medially, more apical ventrites flavous; metafemur flavous, metatibia flavous with brunneous cast. Male genitalia (n = 1). Aedeagal median lobe gracile, elongate, distance from parameral articulation to tip 4.7 x depth at midlength (Fig. 11 A); median lobe apex parallel sided with rounded tip, apex evenly downcurved distad ostial opening in lateral view (Fig. 11 A-B), apex curved to right with top expanded as a knob in ventral view (Fig. 11 C); internal sac elongate, flagellar plate small, sac covered with fine spicules only. Female reproductive tract (n = 1). Bursa copulatrix columnar, base broad at vagina, narrower at midlength, bursal length 0.80 mm, breadth 0.40 mm at base (Fig. 12 A); bursal surface translucent, wrinkled, not sclerotized; gonocoxite 1 with 2 - 3 apical fringe setae, 1 small seta at apicomedial angle, 4 - 5 setae along medial surface (Fig. 13 A); gonocoxite 2 subacuminate with lateral panhandle extension; 2 moderately elongate lateral ensiform setae, 0.36 x length of gonocoxite; apical nematiform setae on medial surface at 0.80 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
99254A1C54A67A694A35EE16756A3BD0.taxon	etymology	Etymology. The adjectival species epithet perseveratus is based on the verb perseverate; to repeat insistently or redundantly. Such a name could be appropriately applied for any number of Hawaiian Mecyclothorax, but it is used here as the name shares the first syllable with the following cryptic sibling species.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
65FEC74E490890F8990501713F9D1727.taxon	diagnosis	Diagnosis. Individuals of Mecyclothorax perstriatus and Mecyclothorax superstriatus share reduced microsculpture, the pronotal disc glossy with an obsolete transverse mesh visible over portions of the disc (Fig. 10 B-C). However the elytral microsculpture is more developed in this species, being an evident, shallow transverse mesh versus an obsolete transverse mesh on a glossy surface in Mecyclothorax superstriatus. Beetles of both species also have smaller eyes that cover less of the ocular lobe than do those of Mecyclothorax perseveratus, with Mecyclothorax perstriatus exhibiting an ocular lobe ratio = 0.83 - 0.86. The male genitalia provide certain diagnosis, with the aedeagal median lobe apex of Mecyclothorax perstriatus males extended and downturned (Fig. 11 D-G), versus very short and not at all downturned in Mecyclothorax superstriatus (Fig. 11 H-I). Setal formula 2 1 (2) 1 0; only one individual was observed within which the basal pronotal seta was unilaterally present. Standardized body length 3.5 - 4.6 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
238897C86F8A4AE1BC90DF09DABCBB6B.taxon	diagnosis	Diagnosis. Superficially similar to the preceding two species, but individuals tend to be smaller-standardized body length 3.8 - 4.2 mm-with less developed microsculpture (Fig. 10 C). The pronotal disc and discal elytral intervals are glossy, with only a very shallow transverse mesh discernible over portions of the cuticle. The male aedeagus is small (Figs 11 H-I) with non-projected apex. Setal formula 2 1 1 (0) 0; of 13 individuals assessed, 9 have the anterior dorsal elytral seta present both sides, 3 have the seta present unilaterally, and 1 lacks any dorsal elytral setae.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
43123C1B56A7CC87C35A8AEAFC202DD8.taxon	diagnosis	Diagnosis. This species is easily diagnosed by the pale pronotal and elytral margins contrasted with piceous discal areas and the rugose transverse wrinkles of the pronotal disc (Fig. 15 A). The pronotum is moderately constricted basally-MPW / BPW = 1.44 - 1.59 - and the forebody dorsal surface bears well-developed microsculpture. The vertex is covered with upraised isodiametric sculpticells in transverse rows, and the pronotal disc is covered with a transverse mesh, sculpticell breadth 2 - 3 x length. Conversely, the elytral discal intervals are glossy, with a transverse mesh to transverse lines toward the lateral elytral margins. Setal formula 2 (1 - 2) 2 1 [sae]; of the five individuals scored, two have both lateral and basal pronotal setae, and three have only the lateral pair of setae. Standardized body length 3.4 - 4.1 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
00B683217EA3BCB73F49DC36A5EF36D7.taxon	diagnosis	Diagnosis. Among the pale-margined members of this species group, Mecyclothorax notobscuricornis (Fig. 15 B) stands out due to the basally constricted pronotum; MPW / BPW = 1.57 - 1.61. This is also the largest-bodied species in the group, with standardized body length 3.6 - 4.3 mm. Like Mecyclothorax daptinus, this species is characterized by well-developed forebody microsculpture and glossy elytra. The vertex is covered with well-developed isodiametric and transverse-mesh microsculpture, the pronotal disc by a well-developed transverse mesh, sculpticell breadth 2 - 3 x length, whereas the discal elytral intervals are glossy, their surface covered with reduced transverse lines. Setal formula 2 1 2 1 [sae].	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
00B683217EA3BCB73F49DC36A5EF36D7.taxon	description	Description (n = 4). Head capsule with frontal grooves broad near clypeus, an associated lateral carina extended to anterior supraorbital seta; dorsal impression of neck flat to slightly concave; labral anterior margin broadly, shallowly emarginate; antennomeres 2 - 3 with sparse pelage of short setae; mentum tooth with sides acute, apex tightly rounded. Pronotum with lateral setae present, basal absent; hind angle obtuse to right, to slightly acute, margin rounded posterad angle; lateral margin subparallel to convergent anterad hind angle; median base depressed relative to disc, with fine longitudinal wrinkles; basal margin straight medially, expanded posterad laterobasal depressions; median longitudinal impression moderately deep at midlength, finely incised, extended onto median base; anterior transverse impression broad, shallow, crossed with longitudinal wrinkles, lined with granulate isodiametric microsculpture; anterior callosity slightly convex; front angles slightly projected, tightly rounded, more distant than hind angles, APW / BPW = 1.06 - 1.12; lateral marginal depression narrow throughout length, edge reflexed, beadlike margin only anterad and posterad hind angle; laterobasal depression smooth, continuous with lateral depression. Proepisternum with smooth hind marginal groove; prosternal process medially depressed, sides broadly upraised laterally. Elytra subovoid, disc moderately convex, lateral margins more so; basal groove distinctly recurved to subangulate humeral angle, MEW / HuW = 1.95 - 2.07; parascutellar seta present; parascutellar striole with 3 - 5 punctures, deep, continuous; sutural interval more convex than lateral intervals, sutural juncture elevated; sutural and 2 nd striae of subequal depth from base to apex; all striae deep, associated intervals convex; 7 th and 8 th interval similarly convex near apex; 2 dorsal elytral setae, anterior at 0.29 x, and posterior at 0.58 - 0.61 x elytral length, setae situated in depressions spanning 2 / 3 of interval 3; subapical seta present, apical seta absent; lateral elytral setae arranged in anterior series of 7 setae, posterior series of 5 (6) setae; elytral marginal depression broadly reflexed, translucent posterad humerus, narrowed apically to beadlike margin anterad subapical sinuation; subapical sinuation shallow, more abruptly incurved anteriorly. Mesepisternum with 6 shallow punctures, most in 1 row; metepisternal width to length ratio = 0.82; metepisternum / metepimeron suture distinct. Abdomen with irregular wrinkles laterally on ventrites 1 - 5; suture between ventrites 2 and 3 reduced, effaced laterally; apical male ventrite with 2 apical setae, apical female ventrite with 4 equally spaced setae plus a median trapezoid of 4 small setae. Legs-metatarsomere 1 / metatibial length ratio = 0.19; metatarsomere 4 length along outer lobe 1.25 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci narrow, lateral, median area broad. Microsculpture of vertex well-developed isodiametric mesh in rows; pronotal disc with well-developed transverse mesh, sculpticell breadth 2 - 3 x length; pronotal median base with upraised isodiametric and transverse mesh, transverse sculpticell breadth 2 x length; elytral discal surface glossy, with reduced transverse lines; elytral apex with shallow transverse mesh and lines; metasternum with distinct transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex rufobrunneous; antennomere 1 flavous, antennomeres 2 - 3 rufous, 4 - 11 rufobrunneous; pronotal disc rufopiceous, lateral margins flavous, apex and base rufoflavous; proepipleuron rufoflavous, proepisternum rufobrunneous; elytral disc rufopiceous on intervals 2 - 5 to 2 - 6, base of interval 6 flavous; sutural interval rufobrunneous basally, rufoflavous apically; elytral intervals 7 - 9 pale, flavous, apex flavous to apical juncture of intervals 3 and 4; elytral epipleuron flavous dorsally, rufoflavous ventrally, metepisternum rufopiceous; abdominal ventrites 1 - 5 rufopiceous medially, 3 - 6 flavous laterally; apical half of apical ventrite 6 pale, flavous; metafemur flavous; metatibia rufoflavous with piceous cast. Male genitalia (n = 1). Aedeagal median lobe broad, robust, distance from parameral articulation to tip 2.5 x depth at midlength, apex broadly rounded and little extended beyond ostial opening (Fig. 16 C); slightly curved to the right, the tip broadly rounded in ventral view (Fig. 16 D); internal sac unornamented, large flagellar plate evident inside dorsal surface of median lobe dorsad parameral articulation (Fig. 11 C). Female reproductive tract (n = 1). Bursa copulatrix elongate, broad with rounded apex, length 0.57 mm, breadth at bursal-vaginal juncture 0.23 mm, maximum vagina breadth 0.35 mm (Fig. 17 B); bursal walls translucent, thinly wrinkled; gonocoxite 1 with 3 apical fringe setae, 5 setae on medial surface (Fig. 18 B); gonocoxite 2 subtriangular, apex tightly rounded, base with short broad lateral extension, 2 lateral ensiform setae, the apical seta broader and longer, apical nematiform setae on medioventral surface at 0.70 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
00B683217EA3BCB73F49DC36A5EF36D7.taxon	etymology	Etymology. The adjectival species epithet notobscuricornis is drawn from the converse name obscuricornis, another species in this group. Why Sharp (1903) used that name is unknown, though it too can be derived from the name of a previously described and related species; Mecyclothorax obscuricolor (Blackburn).	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
ED96F4B3CF96ED9799BCDC752ABFB6AE.taxon	diagnosis	Diagnosis. Among the pale-margined species in this group, this species can be diagnosed by the obtuse, rounded pronotal hind angles, the lateral margins only slightly sinuate anterad the hind angles (Fig. 15 C). The elytra are subquadrate, with the lateral margins broadly extended posterad the rounded humeri. The pronotal base is smooth, with the median base moderately depressed and sparsely covered with shallow punctures and longitudinal wrinkles. Setal formula 2120. Standardized body length 3.8 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
ED96F4B3CF96ED9799BCDC752ABFB6AE.taxon	description	Description (n = 1). Head capsule with frontal grooves broad at clypeus, a lateral carina present to anterior supraorbital seta; dorsal impression of neck flat to slightly concave; eyes little convex, ocular ratio = 1.37, ocular lobe ratio = 0.77; labral anterior margin with broad, shallow emargination; antennae filiform, antennomere 2 sparsely setose, antennomere 3 with well-developed pelage of short setae; mentum tooth with sides acute, apex rounded. Pronotum with lateral seta present, basal seta absent; pronotal base little constricted, MPW / BPW = 1.42; basal margin straight medially, expanded posterad laterobasal depressions; median longitudinal impression shallow, finely incised; anterior transverse impression deep, narrow, finely incised; anterior callosity slightly convex, glossy but crossed by indistinct wrinkles; front angles moderately projected, tightly rounded; apical and basal angles at subequal separation, APW / BPW = 0.99; lateral marginal depression narrow but edge reflexed anteriorly, broader at midlength, beaded at hind angle; laterobasal depression smooth, concave, continuous with lateral depression. Proepisternum with 5 minute punctures along hind mar ginal groove; prosternal process medially depressed, broadly upraised laterally. Elytra subquadrate, disc moderately convex, sides slightly more sloped; basal groove moderately recurved to meet rounded humeral angle; parascutellar seta present; parascutellar striole with 3 - 5 punctures, striole shallow between punctures; sutural interval more convex than lateral intervals, sutural juncture elevated; sutural and 2 nd striae of subequal depth from base to apex; striae 1 - 4 distinctly punctate basally, the punctures expanding strial breadth; intervals 2 - 8 convex to apex; 7 th and 8 th interval similarly convex mesad subapical sinuation; 2 dorsal elytral setae at 0.29 x and 0.59 x elytral length, setal impressions spanning 3 rd interval; apical and subapical setae absent; lateral elytral setae arranged as anterior series of 7 setae, and posterior series of 5 - 6 setae; elytral marginal depression narrow at humerus, posteriorly expanded laterally, broad to midlength, a narrow bead at subapical sinuation; subapical sinuation very shallow, nearly obsolete. Mesepisternum with ~ 8 shallow punctures arranged in 2 - 3 rows; metepisternal width to length ratio = 0.80; metepisternum / metepimeron suture distinct; metathoracic flight wing extended to posterior margin of metanotum. Abdomen with irregular lateral wrinkles on ventrites 1 - 4; suture between ventrites 2 and 3 complete; apical ventrite of male with 2 apical setae. Legs-metatarsomere 1 / metatibial length ratio = 0.19; metatarsomere 4 length along outer lobe 1.33 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci narrow, lateral, median area broad. Microsculpture of vertex shallow transverse mesh, sculpticell breadth 2 x length; pronotal disc with shallow transverse mesh, sculpticell breadth 2 x length; pronotal median base with shallow isodiametric and transverse-mesh microsculpture; elytral disc with shallow transverse mesh, sculpticell breadth 2 x length; elytral apex with shallow transverse mesh, sculpticell breadth 2 - 3 x length, plus transverse lines; metasternum with distinct transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex a glossy rufopiceous, antennomere 1 rufoflavous, antennomeres 2 - 3 rufoflavous, 4 - 11 rufobrunneous; pronotal disc rufopiceous, pronotal anterior callosity, lateral margins, and median base rufoflavous; proepipleuron flavous, proepisternum rufobrunneous with piceous cast; elytral disc with intervals 2 - 5 piceous from base to near apex, interval 6 rufous, and 7 - 9 flavous continuous with broadly flavous apex; sutural interval rufoflavous basally, flavous in apical 1 / 3; elytral epipleuron pale creamy ivory, metepisternum rufopiceous with piceous cast; abdominal ventrites 1 - 3 medially, and 4 - 6 mediobasally piceous, flavous laterally and apically; apical 2 / 3 of apical abdominal ventrite 6 flavous; metafemur flavous with piceous cloud covering basal half of anterior face; metatibia rufoflavous with brunneous cast. Male genitalia (n = 1). Aedeagal median lobe robust (Fig. 16 E), but thinner than that of Mecyclothorax notobscuricornis (Fig. 16 C), distance from parameral articulation to tip 3.6 x depth at midlength, apex extended more than its breadth beyond ostial opening, tip tightly rounded; median lobe narrowed apically and curved to the right in ventral view, tip appearing bluntly rounded (Fig. 16 F); internal sac with field of 5 large macrospicules, flagellar plate large and visible in uneverted specimen just basad spicular field.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
ED96F4B3CF96ED9799BCDC752ABFB6AE.taxon	etymology	Etymology. The adjectival epithet mordax means biting, corroding, or pungent, an appropriate name for a carabid beetle. But here it is also a play on the epithet daptinus - used by Sharp (1903) for a species of similar appearance-derived from the Greek dapto; to devour, or gnaw (Brown 1956).	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
3177E944C498DE55EF541D026CB39F3B.taxon	diagnosis	Diagnosis. Of the pale margined species in this group from Haleakala that exhibit a smooth pronotal disc, sinuate lateral pronotal margins and a broader pronotal base-MPW / BPW = 1.42 - 1.52 - this species can be diagnosed by more convex eyes-ocular ratio = 1.44 - 1.55 - and the subparallel pronotal lateral margins and protruded hind angles (Fig. 15 D). The most similar species is Mecyclothorax manducus (Fig. 20 A), which deviates by exhibiting smaller eyes-ocular ratio = 1.40 - 1.43. If a male is available, the aedeagus can settle the matter, with Mecyclothorax mordicus exhibiting a median lobe with a very short rounded apex distad the ostial opening (Fig. 16 G), whereas the median lobe apex of Mecyclothorax manducus is elongate (Fig. 16 I). Setal formula 2 1 2 1 [sae]. Standardized body length 3.6 - 3.9 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
3177E944C498DE55EF541D026CB39F3B.taxon	description	Description (n = 4). Head capsule with frontal grooves broad at clypeus, a lateral carina to anterior supraorbital seta; dorsal impression of neck flat to concave; ocular lobe ratio = 0.75 - 0.77; labral anterior margin broadly, shallowly emarginate; antennae filiform, antennomere with sparse setae, antennomere 3 with pelage of short setae; mentum tooth with sides acute, apex rounded. Pronotum with lateral seta present, basal seta absent; hind angle obtuse, margin rounded posterad; lateral margin subparallel anterad hind angle then divergent; median base moderately depressed, covered with dense longitudinal wrinkles and some punctures; basal margin straight medially, slightly expanded posteriorly mesad laterobasal depressions; median longitudinal impression shallow, finely incised; anterior transverse impression deep, broader medially, lined with granulate isodiametric microsculpture and crossed by longitudinal wrinkles; anterior callosity elevated, flat, crossed by longitudinal wrinkles; front angles little projected, tightly rounded; front angles slightly farther apart than hind angles; APW / BPW = 1.03 - 1.04; lateral marginal depression moderately narrow, edge upturned; laterobasal depression smooth, concave, surface continuous with lateral depression. Proepisternum with 5 minute punctures along hind marginal groove; prosternal process medially depressed, sides broadly upraised. Elytra subquadrate, MEW / HuW = 1.81 - 1.86; disc moderately convex, sides more sloped; basal groove angled anterad from 3 rd stria to meet tightly rounded humeral angle; parascutellar seta present; parascutellar striole with 3 - 5 punctures, striole shallow between punctures; sutural interval more convex than lateral intervals, sutural juncture upraised; sutural and 2 nd striae of subequal depth from base to apex; intervals 2 - 8 convex, associated striae deep, minute irregularities present at base, smooth on disc and at apex; 7 th and 8 th interval of similar convexity mesad subapical sinuation; 2 dorsal elytral setae situated at 0.30 x and 0.60 x elytral length, anterior setal impression spanning interval 3, posterior impression spanning 1 / 2 of interval width; subapical seta present, apical seta absent; lateral elytral setae arranged as anterior series of 7 setae and posterior series of 6 setae; elytral marginal depression moderately narrow at humerus, broader posterad, beadlike at subapical sinuation; subapical sinuation shallow, more abruptly incurved anteriorly. Mesepisternum with ~ 4 shallow punctures in one row; metepisternal width to length ratio = 0.81; metepisternum / metepimeron suture distinct. Abdomen with irregular lateral wrinkles on ventrites 1 - 5; suture between ventrites 2 and 3 complete; apical ventrite of male with 2 setae. Legs-metatarsomere 1 / metatibial length ratio = 0.20; metatarsomere 4 length along outer lobe 1.25 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci narrow, lateral, median area broad. Microsculpture of vertex isodiametric sculpticells in transverse rows; pronotal disc with transverse mesh, sculpticell breadth 2 x length; pronotal median base with transverse mesh, sculpticell breadth 2 x length; elytral disc glossy, transverse mesh to transverse lines present laterally; elytral apex with shallow transverse mesh, sculpticell breadth 2 - 3 x length; metasternum with distinct transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex rufobrunneous; antennomere 1 flavous, antennomeres 2 - 3 rufoflavous, 4 - 11 rufobrunneous; pronotal disc rufopiceous, anterior callosity, lateral margins broadly, and median base rufoflavous; proepipleuron and proepisternum rufoflavous; elytral disc with intervals 2 - 5 piceous from base to apical 5 / 6, and interval 6 piceous on disc; sutural interval rufobrunneous basally and on disc, flavous in apical half; elytral intervals 7 - 9 plus marginal depression flavous, apex broadly flavous to juncture of intervals 3 and 4; elytral epipleuron pale creamy ivory, metepisternum rufobrunneous; abdomen with ventrites 1 - 3 medially and 4 - 6 mediobasally piceous; apical 2 / 3 of apical abdominal ventrite 6 flavous; metafemur flavous; metatibia flavous with brunneous cast. Male genitalia (n = 2). Aedeagal median lobe gracile but with tip very broadly rounded and little extended beyond ostial opening, distance from parameral articulation to tip 4.4 x depth at midlength (Fig. 16 G); median lobe not curved to the right except near bluntly rounded tip in ventral view (Fig. 16 H); internal sac unornamented, lightly sclerotized flagellar plate visible in lateral and ventral views (Figs 16 G-H).	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
3177E944C498DE55EF541D026CB39F3B.taxon	etymology	Etymology. The adjectival mordicus means biting in the mandibular sense, continuing the string of epithets that started with daptinus above.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
3919D20C8CEE17D559E2827E3A4EE6AB.taxon	diagnosis	Diagnosis. Among the pale-margined species of this group, the slightly sinuate pronotal lateral margins associated with the obtuse pronotal hind angles of this species (Fig. 20 A) are intermediate to the non-sinuate lateral margins of Mecyclothorax mordax (Fig. 15 C) and the distinctly sinuate lateral margins of Mecyclothorax mordicus (Fig. 15 D). This species is also characterized by less convex eyes-ocular ratio = 1.40 - 1.43 - versus the more convex eyes of the other two species. Setal formula 2 1 2 (3) 1 [sae]; a third unilaterally present dorsal elytral seta was observed in one individual. Standardized body length 3.6 - 4.1 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
3919D20C8CEE17D559E2827E3A4EE6AB.taxon	description	Description (n = 5). Head capsule with frontal grooves broad near clypeus, a lateral carina present to anterior supraorbital seta; dorsal impression of neck flat to slightly concave; ocular lobe projected posteriorly, eye small, ocular lobe ratio = 0.71 - 0.74; labral anterior margin broadly, shallowly emarginate, antennomere 2 sparsely setose, antennomere with well-developed pelage of short setae; antennae filiform; mentum tooth with sides acute, apex tightly rounded. Pronotum with lateral seta present, basal seta absent; median base moderately depressed relative to disc, minutely punctate, minute wrinkles present on disc; basal margin straight medially, expanded posteriorly mesad laterobasal depressions; median longitudinal impression narrow, shallowly incised, continuous to basal margin; anterior transverse impression deep, narrow, surface behind with granulate isodiametric microsculpture, crossed by fine longitudinal wrinkles; anterior callosity slightly convex, glossy except for longitudinal wrinkles; front angles slightly projected, tightly rounded; APW / BPW = 1.00 - 1.04; lateral marginal depression narrow, edge upturned anteriorly, slightly broader, reflexed near base; laterobasal depression smooth, transversely wrinkled onto disc, continuous with lateral depression. Proepisternum with smooth hind marginal groove; prosternal process medially depressed, sides broadly upraised. Elytra subquadrate, disc moderately convex, sides more so; basal groove gently recurved to rounded humeral angle; MEW / HuW = 1.98 - 2.03; parascutellar seta present; parascutellar striole with three deep punctures, continuous between punctures; sutural interval more convex than lateral intervals, sutural juncture elevated; sutural and 2 nd striae of subequal depth from base to apex; striae 2 - 8 of similar depth, associated intervals convex; striae 1 - 3 with small punctures that expand stria, striae 4 - 5 with slight irregularities; 7 th and 8 th interval similarly convex mesad subapical sinuation; 2 dorsal elytral setae at 0.27 x and 0.63 x elytral length (unilateral third seta at 0.48 x length), setal impressions spanning 2 / 3 of interval 3; subapical seta present, apical seta absent; lateral elytral setae arranged in anterior series of 7 setae, posterior series of 6 setae; elytral marginal depression narrow at humerus, expanded laterally along sides, narrowly beaded at subapical sinuation; subapical sinuation shallow, more abruptly incurved anteriorly. Mesepisternum with ~ 5 very shallow punctures; metepisternal width to length ratio = 0.76; metepisternum / metepimeron suture distinct. Abdomen with irregular lateral wrinkles on ventrites 1 - 5; suture between ventrites 2 and 3 reduced, effaced laterally; apical male ventrite with 2 marginal setae, female ventrite with 4 equally spaced marginal setae plus a median trapezoid of 4 smaller setae. Legs-metatarsomere 1 / metatibial length ratio = 0.18; metatarsomere 4 length along outer lobe 1.25 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci narrow, lateral, median area broad. Microsculpture of vertex a well-developed isodiametric mesh in transverse rows; pronotal disc with well-developed transverse mesh, sculpticell breadth 2 - 3 x length; pronotal median base with well-developed, upraised transverse mesh, sculpticell breadth 2 x length; elytral disc with transverse mesh, sculpticell breadth 2 - 3 x length; elytral apex with mixture of transverse mesh and transverse lines; metasternum with distinct transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex rufobrunneous; antennomere 1 flavous, antennomeres 2 - 3 rufoflavous, 4 - 11 rufobrunneous; pronotal disc rufopiceous, pronotal margins broadly rufobrunneous, median base rufoflavous; proepipleuron rufoflavous, proepisternum rufobrunneous; elytral intervals 2 - 6 piceous from base to juncture of striae 3 and 4, outer intervals and apex flavous; sutural interval rufobrunneous basally, flavous in apical 1 / 3; elytral epipleuron rufoflavous, metepisternum rufobrunneous with piceous cast; abdomen with ventrites 1 - 5 rufopiceous medially, rufoflavous laterally; basal half of apical ventrite 6 rufopiceous, apical half flavous; metafemur flavous; metatibia flavous with brunneous cast. Male genitalia (n = 2). Aedeagal median lobe moderately elongate, distance from parameral articulation to tip 3.7 x depth at midlength (Fig. 16 I); apical extension parallel sided, the tip subangulate at its ventral margin; median lobe apex curved to the right just before blunt tip in ventral view (Fig. 16 J); internal sac unornamented, elongate flagellar plate visible inside dorsal margin of median lobe in lateral view. Female reproductive tract (n = 1). Bursa copulatrix columnar, 0.57 mm long, 0.15 mm broad (Fig. 17 C); bursal walls translucent, thinly wrinkled; gonocoxite 1 with 2 - 5 apical fringe setae, 8 - 9 small setae on medial surface (Fig. 18 C); gonocoxite 2 subtriangular, apex acuminate; base evenly extended from lateral margin and basally curved at apex; 2 gracile lateral ensiform setae, apical nematiform setae on medial surface at 0.70 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
3919D20C8CEE17D559E2827E3A4EE6AB.taxon	etymology	Etymology. This last of the species epithets related to biting, the Latin noun manducus means glutton (Brown 1956).	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
DC0E917ED867C1F65C3A1E1A6E0C1867.taxon	diagnosis	Diagnosis. This species exhibits the most somber coloration in the group, the dorsal surface dark fuscous, and antennomeres 1 - 3 and femora only slightly paler (Fig. 20 B). Elytral striae 1 - 5 are punctate basally, the punctures expanding strial breadth, a characteristic shared with the species triplet, Mecyclothorax waikamoi, Mecyclothorax poouli, and Mecyclothorax ahulili (Fig. 24 B-D), however Mecyclothorax ambulatus exhibits larger body size; standardized body length 4.0 mm versus 2.9 - 3.4 mm in the other three species. Setal formula 2 1 2 0.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
DC0E917ED867C1F65C3A1E1A6E0C1867.taxon	description	Description (n = 1). Head capsule with frontal grooves broad near clypeus, a lateral carina present to anterior supraorbital seta; dorsal impression of neck flat to slightly concave; ocular ratio = 1.43, ocular lobe ratio = 0.81; labral anterior margin broadly, shallowly emarginate; antennae filiform, antennomere 2 sparsely setose, antennomere 3 with well-developed pelage of short setae; mentum tooth with sides acute, apex tightly rounded. Pronotum with lateral seta present, basal seta absent; MPW / BPW = 1.43; hind angle obtuse, apex rounded; lateral margin slightly divergent anterad hind angle, then more divergent anteriorly; median base moderately depressed relative to disc, sparsely covered with minute punctures, minute wrinkles present at juncture with disc; basal margin convexly expanded between laterobasal depressions; median longitudinal impression narrow, shallowly incised; anterior transverse impression narrow, finely incised, shallowest medially, crossed by fine wrinkles; anterior callosity elevated, flat, crossed by fine wrinkles; front angles slightly projected, tightly rounded; APW / BPW = 1.01; lateral marginal depression narrow, edge upturned anteriorly, beadlike at lateral sinuation and posterad laterobasal depressions; laterobasal depression smooth, continuous with lateral depression. Proepisternum with smooth hind marginal groove; prosternal process medially depressed, sides broadly upraised. Elytra broadly subquadrate; disc moderately convex, sides more so; basal groove recurved to subangulate humeral angle defined by a hitch at base of lateral marginal depression; MEW / HuW = 1.93; parascutellar seta present; parascutellar striole with 3 deep punctures, deep, continuous between punctures; sutural interval more convex than lateral intervals, sutural juncture elevated; sutural and 2 nd striae of subequal depth base to apex; striae 1 - 8 deep, interval 8 convex mesad subapical sinuation; 2 dorsal elytral setae at 0.28 - 0.30 x and 0.61 - 0.63 x elytral length, setal impressions moderately deep, spanning 2 / 3 of interval 3; subapical and apical setae absent; lateral elytral setae arranged in an anterior series of 7 setae and a posterior series of 5 setae; elytral marginal depression broad with upraised margin at humerus, gradually narrowed but still evident before subapical sinuation. Mesepisternum with ~ 7 punctures in 2 - 3 irregular rows; metepisternal width to length ratio = 0.80; metepisternum / metepimeron suture distinct. Abdomen with irregular lateral wrinkles on ventrites 1 - 5; suture between ventrites 2 and 3 reduced, effaced laterally; apical ventrite of male with 2 marginal setae. Legs-metatarsomere 1 / metatibial length ratio = 0.19; metatarsomere 4 length along outer lobe 1.33 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci narrow, lateral, median area broad. Microsculpture of vertex a well-developed isodiametric mesh in transverse rows; pronotal disc with well-developed transverse mesh, sculpticell breadth 2 - 3 x length; pronotal median base with well-developed, upraised transverse mesh, sculpticells twice as broad as long; elytral disc with mixture of transverse mesh and transverse lines; elytral apex with mixture of transverse mesh-sculpticell breadth 3 x length-and transverse lines; metasternum with distinct transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex dark rufobrunneous; antennomeres 1 - 3 rufobrunneous, 4 - 11 rufopiceous; pronotal disc and margins concolorous, dark rufobrunneous; proepipleuron rufoflavous, proepisternum rufobrunneous; elytral disc dark rufobrunneous, sutural interval concolorous basally, slightly paler, rufoflavous at apex, elytral marginal depression and apex slightly paler, rufoflavous; elytral epipleuron rufoflavous, metepisternum rufobrunneous; abdomen rufobrunneous, apical 1 / 6 of apical ventrite 6 paler, flavous; metafemur rufobrunneous with piceous cloud covering basal 4 / 4; metatibia rufoflavous with piceous cast. Male genitalia (n = 1). Aedeagal median lobe broad, long, distance from parameral articulation to tip 3.3 x depth at midlength (Fig. 16 K); apex evenly curved, extended for 3 x its breadth beyond ostial opening, tip slightly flattened on dorsal aspect; median lobe distinctly curved to the right just before blunt apex in ventral view (Fig. 16 L); internal sac with dark microspicules over surface, a short flagellar plate visible inside dorsal margin in lateral view.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
DC0E917ED867C1F65C3A1E1A6E0C1867.taxon	etymology	Etymology. The past participle ambulatus means to have travelled or traversed, and is used to signify R. C. L. Perkins' discovery of the single known specimen at 1500 ft. elevation on Haleakala. This low elevation collecting site is interpreted to have been along his walk from Makawao, the home base of his mountain collecting, to Paia, the village where he purchased groceries (Manning 1986).	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
D15B036730820118F01EDFC8C520D221.taxon	diagnosis	Diagnosis. The combination of quadrisetose pronotum, both lateral and basal setae present, subparallel to subovoid elytra, the lateral margins nearly straight along the anterior lateral setal series, and moderate body size, standardized body length, 3.4 - 3.8 mm, will diagnose this species. The pronotal and elytral margins are slightly paler than their respective discs (Fig. 20 C), but not to the degree as in the paler-margined species treated above (Figs 15, 20 A). This is also the only Haleakala species of the group to exhibit both subapical and apical elytral setae; setal formula 2 2 2 2. Nevertheless, beetles of this species lack the parascutellar seta, and exhibit a reduced lateral elytral setal series, with base numbers of six anterior setae and five posterior setae.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
D15B036730820118F01EDFC8C520D221.taxon	description	Description (n = 5). Head capsule with frontal grooves broad near clypeus, a lateral carina to anterior supraorbital seta; dorsal impression of neck flat to slightly concave; ocular lobe protruded, eyes small, ocular ratio = 1.36 - 1.38, ocular lobe ratio = 0.66 - 0.71; labral anterior margin broadly, shallowly emarginate; antennae submoniliform, antennomeres 2 - 3 sparsely setose; mentum tooth with sides acute, apex rounded. Pronotum quadrisetose, both lateral and basal setae present; MPW / BPW = 1.38 - 1.49; hind angle obtuse due to rounded margin behind; lateral margin parallel just anterad angle, then divergent; median base depressed relative to disc, a few large punctures and wrinkles present; basal margin broadly, slightly convex between laterobasal depressions; median longitudinal impression narrowly, shallowly incised, continuous to basal margin, adjoined by curved wrinkles emanating onto disc; anterior transverse impression deep, narrow, finely incised, crossed by fine longitudinal wrinkles; anterior callosity elevated, flat, crossed by fine wrinkles; front angles slightly projected, tightly rounded; APW / BPW = 0.94 - 1.03; lateral marginal depression narrow, edge upturned to beadlike at front, broader, less elevated to base; laterobasal depression surface impunctate with transverse wrinkles, continuous with lateral depression. Proepisternum with ~ 5 minute punctures along hind marginal groove; prosternal process medially depressed, sides broadly upraised. Elytra subparallel to subovoid, disc moderately convex, sides more so; basal groove evenly recurved to tightly rounded humeral angle; MEW / HuW = 1.80 - 2.00; parascutellar seta absent; parascutellar striole smooth, sinuous; sutural interval more convex than lateral intervals, sutural juncture upraised; sutural and 2 nd striae of subequal depth from base to apex; discal striae 1 - 7 with small punctures that cause strial irregularities; 7 th and 8 th interval of similar convexity mesad subapical sinuation; 2 dorsal elytral setae at 0.28 - 0.35 x and 0.59 x elytral length; setal impressions moderate, spanning 2 / 3 of interval 3; apical and subapical setae present; lateral elytral setae arranged in anterior series of 6 setae, and posterior series of 5 setae (rarely 4 or 6 setae); elytral marginal depression narrow from humerus to midlength, gradually narrowed to bead at subapical sinuation; subapical sinuation shallow, more abruptly incurved anteriorly. Mesepisternum with ~ 7 punctures in 2 - 3 rows; metepisternal width to length ratio = 0.80; metepisternum / metepimeron suture distinct; metathoracic flight wing length to width ratio = 2.2, remnant R and M veins present, wing tip extended 2 / 3 distance to hind margin of metanotum. Abdomen with irregular lateral wrinkles on ventrites 1 - 5; suture between ventrites 2 and 3 complete; apical male ventrite with 2 marginal setae, apical female ventrite with 4 equally spaced marginal setae and median trapezoid of 4 - 5 short setae (the 5 th seta, when present, shorter). Legs-metatarsomere 1 / metatibial length ratio = 0.19; metatarsomere 4 length along outer lobe 1.28 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci narrow, lateral, median area broad. Microsculpture of vertex very shallow, transverse, sculpticell breadth 2 x length; pronotal disc with shallow transverse mesh, sculpticell breadth 3 x length; pronotal median base with shallow isodiametric and transverse mesh; elytral disc with shallow transverse mesh, sculpticells 2 - 3 x length; elytral apex with shallow transverse mesh, sculpticell breadth 2 x length; metasternum with distinct transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex glossy rufopiceous; antennomeres 1 - 3 flavous, 4 - 11 only slightly darker; pronotal disc rufopiceous; pronotal lateral margins, anterior callosity and median base rufoflavous to flavous at outer margins; proepipleuron flavous, proepisternum rufobrunneous; elytral intervals 2 - 8 rufopiceous, sutural interval rufoflavous basally, flavous apically; interval 9, marginal depression, and apical margin rufoflavous; elytral epipleuron flavous dorsally, rufoflavous along ventral margin, metepisternum rufobrunneous; abdomen with ventrites 1 - 5 rufopiceous medially, rufoflavous laterally; apical ventrite 6 flavous in apical half; metafemur flavous; metatibia flavous with brunneous cast. Male genitalia (n = 1). Aedeagal median lobe gracile, distance from parameral articulation to tip 4.5 x depth at midlength (Fig. 22 A), apex broad, extended twice its breadth beyond ostial opening, tip flattened on dorsoapical aspect; median lobe slightly curved to the right before rounded tip in ventral view (Fig. 22 B); internal sac covered with evident microspicules, a flagellar plate visible inside dorsal margin in lateral view (Fig. 22 A). Female reproductive tract (n = 1). Bursa copulatrix columnar, nearly as broad as vagina, length 0.54 mm, breadth 0.23 mm (Fig. 17 D); bursal walls translucent, distinctly wrinkled; gonocoxite 1 with 3 - 4 apical fringe setae, 4 - 5 smaller setae on medial surface (Fig. 18 D); gonocoxite 2 subtriangular, apex acuminate, lateral surface distinctly curved, 2 small lateral ensiform setae, apical nematiform setae on medial surface at 0.75 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
D15B036730820118F01EDFC8C520D221.taxon	etymology	Etymology. The adjectival epithet montanus means of the mountains, which aptly fits the habits of this species distributed around the head of Kīpahulu Valley.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
4066E44AF2E7E5D1759693A32A65F70B.taxon	diagnosis	Diagnosis. Of species in this group with concolorous pronotal discs and margins, this species exhibits the smoothest elytral striae, with only minute punctures basally in striae 1 - 7 that cause slight irregularities of the strial surface (Fig. 20 D). The pronotal median base is moderately depressed relative to the disc, with longitudinal punctures and wrinkles producing a rough surface. The dorsal surface bears well-developed microsculpture, an isodiametric mesh on the vertex, and transverse mesh on pronotal and elytral discs with sculpticell breadth 2 - 3 x length. Setal formula 2 1 2 1 [sae]. Standardized body length 3.7 - 4.0 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
0910AB737D011BB7B5846ADCE04F7AE2.taxon	diagnosis	Diagnosis. The combination of standardized body length 3.4 - 3.6 mm, distinctly punctate discal elytral striae (Fig. 24 A), convex lateral elytral margins, and narrowly paler pronotal and elytral margins serves to diagnose this species from others in the group. This species shares with Mecyclothorax obscuricolor well-developed isodiametric and transverse-mesh microsculpture on the vertex, however the pronotal disc has more transverse sculpticells-breadth 3 x length to unconnected transverse lines-and the elytral disc has only a shallow transverse mesh, sculpticell breadth 2 x length, with the surface glossy. In common with Mecyclothorax montanus, the parascutellar seta is lacking. Setal formula 2 1 (2) 2 1 [sae]; the species is scored for rare occurrence of both lateral and basal pronotal setae based on one individual with unilateral presence of the basal seta.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
C6C0660DECC5E6FB1C293E9A60E18619.taxon	diagnosis	Diagnosis. This is the first of three very similar species, all characterized by small body size-standardized body length in this species = 2.9 - 3.4 mm-and subovoid elytra with variously punctate discal striae (Fig. 24 B-D). The three species are all of dark coloration, with rufobrunneous head capsules, rufopiceous pronotal discs, and piceous elytral discs with narrowly paler, rufoflavous to flavous margins. Mecyclothorax waikamoi deviates from the other two in the well-developed punctation of elytral striae 1 - 6 in the basal half of the elytra (Fig. 24 B). The eyes also tend to be less convex in this species-ocular ratio = 1.38 - 1.47 - though that span overlaps the range of ocular ratios of the other two species at 1.43 - 1.51. The male aedeagus (Fig. 26 A) can diagnose the species, with males of this species (Fig. 26 A) exhibiting a broader, more apically flattened apex to the median lobe. Setal formula 2 1 2 0.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
C6C0660DECC5E6FB1C293E9A60E18619.taxon	description	Description (n = 5). Head capsule with frontal grooves broad near clypeus, a lateral carina to anterior supraorbital seta; dorsal impression of neck flat to slightly concave; labral anterior margin broadly, shallowly emarginate; antennae submoniliform, antennomere 2 sparsely setose, antennomere 3 with well-developed pelage of short setae; mentum tooth with sides acute, apex rounded. Pronotum broad, MPW / PL = 1.28 - 1.34, basally constricted, MPW / BPW = 1.47 - 1.54, with sinuate lateral margins anterad slightly obtuse hind angles (obtuse due to rounded basal margin inside angle); median base moderately depressed, sparsely punctate, shallow wrinkles at juncture with disc; basal margin broadly, slightly convex between laterobasal depressions; median longitudinal impression extremely shallow, narrowly incised; anterior transverse impression shallow, narrow, discontinuous medially; anterior callosity slightly convex, smooth; front angles slightly projected, rounded; distance between front and hind angles subequal, APW / BPW = 0.97 - 1.05; lateral marginal depression narrow, edge not upraised from front angle to lateral seta, very narrow and beadlike in basal half; laterobasal depression smooth, narrow, continuous with lateral depression. Proepisternum with ~ 5 minute punctures along hind marginal groove; prosternal process medially depressed, sides broadly upraised. Elytra subovoid and convex, sides and apex depressed relative to disc; basal groove evenly recurved to tightly rounded humeral angle; MEW / HuW = 1.95 - 2.03; parascutellar seta present; parascutellar striole with 3 punctures, discontinuous between punctures; sutural interval more convex than lateral intervals, sutural juncture elevated; sutural and 2 nd striae of subequal depth and punctation from base to apex; striae 1 - 6 and 8 complete, stria 7 shallower, associated intervals convex; 7 th and 8 th interval of similar convexity mesad subapical sinuation; 2 dorsal elytral setae at 0.33 x and 0.50 x elytral length, setal impression small, spanning about half of interval 3; both apical and subapical setae absent; lateral elytral setae arranged as anterior series of 7 (6) setae, and a posterior series of 5 (4) setae; elytral marginal depression slightly broader at humerus, narrowed laterally to a beadlike margin at subapical sinuation; subapical sinuation very shallow, more abruptly incurved anteriorly. Metepisternum with ~ 5 very shallow punctures in 1 row; metepisternal width to length ratio = 0.84; metepisternum / metepimeron suture distinct. Abdomen with irregular lateral wrinkles on ventrites 1 - 5; suture between ventrites 2 and 3 complete; apical male ventrite with 2 marginal setae, female apical ventrite with 4 equally spaced marginal setae and a median trapezoid of 4 - 6 short setae. Legs-metatarsomere 1 / metatibial length ratio = 0.18; metatarsomere 4 length along outer lobe 1.33 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci narrow, lateral, median area broad. Microsculpture of vertex an obsolete transverse mesh, sculpticell breadth 2 - 3 x length; pronotal disc with obsolete transverse mesh, sculpticell breadth 3 x length, surface glossy; pronotal median base glossy with obsolete transverse sculpticells; elytral disc with shallow transverse mesh, sculpticell breadth 2 - 3 x length, transverse lines present on lateral reaches of elytra; elytral apex with transverse sculpticells, breadth 2 - 3 x length; metasternum with dis tinct transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex glossy rufobrunneous to rufopiceous; antennomeres 1 - 3 flavous, 4 - 11 rufobrunneous; pronotal disc rufobrunneous, lateral margins and base slightly paler, rufous; proepipleuron flavous, proepisternum rufoflavous to rufobrunneous; elytral disc rufopiceous on intervals 2 - 9, sutural interval rufoflavous from base, flavous apically; elytral marginal depression narrowly flavous, apex and intervals 8 - 9 near apex flavous; elytral epipleuron flavous dorsally, rufoflavous along ventral margin, metepisternum rufobrunneous; abdomen with ventrites 1 - 3 medially rufopiceous, ventrites 4 - 6 rufobrunneous, apical 1 / 3 of apical ventrite 6 flavous; metafemur flavous, basal half with brunneous to piceous cloud on anterior surface; metatibia rufoflavous with brunneous cast. Male genitalia (n = 3). Aedeagal median lobe distinctly curved dorsally, robust, distance from parameral articulation to tip 2.7 x maximum breadth dorsad ostial opening (Fig. 26 A); dorsal surface expanded at midpoint of ostial opening, apex downturned with apical face flat; internal sac unornamented, lightly spiculated, flagellar plate moderately elongate, length of sclerotized ventral face 0.43 x distance from parameral articulation to apex. Female reproductive tract (n = 1). Bursa copulatrix a very short, broad pouch, length 0.17 mm, breadth at base 0.25 mm (Fig. 23 C); bursal walls thin, transparent, not wrinkled; gonocoxite 1 with 3 apical fringe setae (Fig. 27 A), 4 smaller setae on medial surface; gonocoxite 2 subtriangular, apex pointed, base little extended laterally, 2 short lateral ensiform setae, the apical seta distinctly broader, apical nematiform setae on medial surface at 0.70 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
C6C0660DECC5E6FB1C293E9A60E18619.taxon	etymology	Etymology. The species epithet waikamoi is taken from the Hawaiian Waikamoi, the name of the Maui land section and stream that is translated from Hawaiian as " water of the ruler's taro (Pukui et al. 1974). "	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
B0B2A9BA4EE3B00A70B33D1B0B981659.taxon	diagnosis	Diagnosis. Distinguished from Mecyclothorax waikamoi, above, by the less punctate elytral striae (Fig. 24 B, C) and the slightly larger, more convex eyes; ocular ratio = 1.43 - 1.48, ocular lobe ratio = 0.79 - 0.86. Distinguishable from Mecyclothorax ahulili by the subparallel pronotal lateral margins anterad the obtuse to nearly right hind angles (Fig. 24 C), versus the divergent lateral margins and obtuse rounded hind angles of Mecyclothorax ahulili (Fig. 24 D). The male aedeagal median lobe apex of this species (Fig. 26 B-D) differs distinctively from the very short and broad apex characterizing Mecyclothorax ahulili (Fig. 26 E), and the flagellar plate is larger than observed in Mecyclothorax waikamoi (Fig. 26 A). Setal formula 2 1 2 0. Standardized body length 3.2 - 3.4 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
B0B2A9BA4EE3B00A70B33D1B0B981659.taxon	description	Description (n = 5). [The description of Mecyclothorax waikamoi serves equally well for this species with the following substitutions.] Eyes moderately developed, ocular ratio = 1.43 - 1.48, ocular lobe ratio 0.79 - 0.86; pronotum variably broad, MPW / PL = 1.30 - 1.37; pronotal hind angles obtuse to nearly right, margin rounded behind; pronotal lateral margins subparallel for short distance anterad hind angles, MPW / BPW = 1.49 - 1.53; elytra may be narrower across the humeri in some individuals, MEW / HuW = 1.97 - 2.08; elytral sutural stria with minute irregularities in basal half but without distinct punctures, smooth and deep apically; parascutellar striole with 3 punctures, striole continuous between punctures; lateral elytral setal series arranged as 6 - 7 setae in anterior series, 5 (4) in posterior series; only 4 setae observed in the median trapezoidal setal patch of the female apical abdominal ventrite; metafemora with brunneous cloud on basal anterior surface. Male genitalia (n = 4). Aedeagal median lobe distinctly curved dorsally, robust, distance from parameral articulation to tip 2.7 - 3.0 x maximum breadth dorsad ostial opening (Figs 26 B-D); dorsal surface evenly curved at midpoint of ostial opening, apex downturned with apical face convex to briefly flattened, the flat surface shorter than in Mecyclothorax waikamoi (Fig. 26 A); internal sac unornamented, lightly spiculated, flagellar plate large (Fig. 26 D), length of sclerotized ventral face 0.55 x distance from parameral articulation to apex. Female reproductive tract (n = 1). Bursa copulatrix a triangular tentlike pouch, length 0.32 mm, basal breadth at vagina 0.33 mm (Fig. 23 D); bursal walls thin, transparent; gonocoxite 1 with 2 apical fringe setae, 5 - 6 small setae on medial surface (Fig. 27 B); gonocoxite 2 subtriangular, apex rounded, 2 lateral ensiform setae with apical seta broader and longer, apical nematiform setae on medial surface at 0.72 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
B0B2A9BA4EE3B00A70B33D1B0B981659.taxon	etymology	Etymology. The po'o uli (Melamprosops phaeosoma Casey and Jacoby 1974) was a member of the Hawaiian drepanid finches first discovered in 1973 in the Hanawī rainforest. By 2004 it was extinct (Powell 2008). The Hawaiian species epithet is meant as a memorial to the birds that may have fed on the ancestors of the type series of Mecyclothorax poouli.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
0C93FF2FC07417381362376B1C432B36.taxon	diagnosis	Diagnosis. This species exhibits strial punctation intermediate to Mecyclothorax waikamoi and Mecyclothorax poouli (Fig. 24 B-D) - the sutural stria bears minute punctulae associated with irregularities in the strial orientation-plus the briefest sinuation of the pronotal lateral margin anterad the hind angle. The pronotal lateral marginal depression is also somewhat broader, especially at the front angle where it is almost explanate. The male aedeagal median lobe exhibited by males of this species is very different from those present in males of the other two species, with the apex very brief and ventrally subangulate (Fig. 26 E). The median lobe is also shorter and more robust overall than those seen in males of the other two species. Setal formula 2 1 2 0. Standardized body length 2.9 - 3.2 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
0C93FF2FC07417381362376B1C432B36.taxon	description	Description (n = 4). [As for Mecyclothorax poouli above, the description of Mecyclothorax waikamoi serves for Mecyclothorax ahulili with the following substitutions.] Eyes larger, more convex, ocular ratio = 1.46 - 1.51, ocular lobe ratio = 0.83 - 0.85; pronotum variably broad, MPW / PL = 1.27 - 1.35, basally constricted, MPW / BPW = 1.49 - 1.56; elytra slightly broader across humeri, MEW / HuW = 1.98 - 2.04; 5 setae observed in the median trapezoidal setal patch of the female apical abdominal ventrite; metafemora with brunneous cloud on basal anterior surface. Male genitalia (n = 1). Aedeagal median lobe very broad, robust, distance from parameral articulation to tip 2.2 x depth at midlength (Fig. 26 E); apex very briefly extended beyond ostial opening, length of extension subequal to breadth, apical face slightly flattened; internal sac lightly spiculated, flagellar plate visible in uneverted specimen, plate length 0.49 x distance from parameral articulation to tip.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
0C93FF2FC07417381362376B1C432B36.taxon	etymology	Etymology. The Hawaiian species epithet ahulili is based on the peak Pu'u Ahulili that is near the type locality for this species. ' Ahulili means glowing or dazzling (Pukui et al. 1974), though being a Hawaiian word it is to be treated as a noun.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
9BDF1686B34E577FD434AB7FE5692DEA.taxon	diagnosis	Diagnosis. Among Haleakala species of this group, Mecyclothorax aeneipennis exhibits the most quadrate pronotum, with the lateral margins little sinuate outside the laterobasal depressions (Fig. 28 A); MPW / BPW = 1.17 - 1.29, versus MPW / BPW = 1.34 - 1.57 for all other Haleakala species in this group. The elytral intervals are slightly convex, with discal striae 1 - 5 lined with small but distinct punctures in their basal halves to 2 / 3 of length. At the elytral apex, the 8 th interval is more convex than the fused apical portion of intervals 5 + 7. The vertex is rufobrunneous, elytral disc slightly darker rufopiceous, and elytral disc rufopiceous with a cupreous reflection. The legs are contrastedly paler; femora flavous and tibiae flavous with a brunneous cast. Setal formula 2 2 2 2. Standardized body length 5.4 - 6.3 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
51AC4A78A8C66A8BA80A572B46C7282A.taxon	diagnosis	Diagnosis. Like Mecyclothorax aeneipennis in the quadrate elytra with broad humeri-MEW / HuW = 1.76 - but with cordate pronotum, the lateral margins slightly convergent anterad the projected, right hind angles (Fig. 28 B). The eyes are relatively flat, ocular ratio = 1.35, and the ocular lobes project abruptly from the gena; a slotlike impression at the juncture of gena and ocular lobe. The single specimen, which does not appear teneral, has the pronotal and elytral lateral marginal depressions paler than the discs, the pronotal margins rufobrunneous versus the rufopiceous disc, and the elytral marginal depression narrowly rufoflavous versus the rufopiceous disc. Moreover, this is the only Haleakala species in the group to be characterized by absence of the parascutellar seta. Setal formula 2 2 2 2. Standardized body length 4.5 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
51AC4A78A8C66A8BA80A572B46C7282A.taxon	description	Description (n = 1). Head capsule with frontal grooves broad near clypeus, a lateral carina to anterior supraorbital seta, dorsal surface of neck slightly concave; labral anterior margin broadly, shallowly emarginate; antennae filiform, antennomeres 2 - 3 with sparse pelage of short setae; mentum tooth with sides acute, apex tightly rounded. Pronotum moderately transverse, MPW / PL = 1.30, base moderately constricted, MPW / BPW = 1.38; median base depressed relative to disc, surface strigose with fine longitudinal wrinkles; basal margin slightly convex between the laterobasal depressions; median longitudinal impression shallow, very finely incised at depth, joined by irregular transverse wrinkles; anterior transverse impression deep, finely incised at depth, separate from discal intervals; anterior callosity moderately convex, crossed by shallow wrinkles; front angles slightly projected, apex tightly rounded; APW / BPW = 0.99; lateral marginal depression slightly broader at front angle, moderately narrow behind, edge upturned; laterobasal depression smooth, laterally elevated to projected lateral margin. Proepisternum with smooth marginal groove; prosternal process medially depressed, upraised laterally to narrow bead anterad coxa. Elytra with convex disc, sides progressively sloped laterad; basal groove recurved medially, straight laterally to subangulate humerus; parascutellar striole shallow, with 3 - 4 punctures; sutural interval flat basally, convex at suture from disc to apex; sutural and 2 nd striae of subequal depth from base to apex; intervals 2 - 5 moderately convex, associated striae with minute punctulae causing strial irregularities, all striae smooth and deep apically; 7 th and 8 th intervals of similar convexity mesad subapical sinuation; 2 dorsal elytral setae at 0.25 x and 0.61 - 0.63 x elytral length, setal impressions extended over 2 / 3 of interval 3; both apical and subapical setae present; lateral elytral setae arranged as an anterior series of 7 setae and a posterior series of 6 setae; elytral marginal depression moderately broad throughout length until reduced to beadlike margin from subapical sinuation to apex; subapical sinuation shallow, more abruptly incurved anteriorly. Mesepisternum with ~ 8 - 9 punctures in 2 - 3 rows; metepisternal width to length ratio = 0.83. Abdomen with irregular lateral wrinkles on abdominal ventrites 1 - 5; suture between ventrites 2 and 3 complete; apical ventrite of male with 2 marginal setae. Legs-metatarsomere 1 / metatibial length ratio = 0.20, metatarsomere 4 length along outer lobe 1.2 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci very broad, median area rough to carinate. Microsculpture of vertex isodiametric to slightly transversely stretched in rows; pronotal disc with transverse mesh, sculpticell breadth 2 - 3 x length; pronotal median base with isodiametric to transverse mesh; elytral disc with transverse mesh, sculpticell breadth 3 x length, to transverse lines; elytral apex with shiny transverse mesh, sculpticell breadth 2 - 3 x length; metasternum with distinct transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex rufobrunneous; antennomere 1 flavous, antennomeres 2 - 3 rufoflavous, 4 - 11 rufobrunneous; pronotal disc rufopiceous; pronotal margins rufobrunneous in depression, lateral bead darker, rufopiceous; proepipleuron rufoflavous, proepisternum dark rufobrunneous; elytral disc rufopiceous; sutural interval basally rufobrunneous, apically rufoflavous to flavous; elytral margins and apex narrowly rufoflavous; elytral epipleuron rufoflavous laterally, rufobrunneous ventrally, metepisternum rufopiceous; abdomen medially rufopiceous, laterally rufobrunneous; abdominal apical ventrite with narrowly paler margin, rufobrunneous; metafemur flavous with piceous cloud on basal 2 / 3; metatibia rufobrunneous. Male genitalia (n = 1). Aedeagal median lobe gracile, distance from parameral articulation to tip 4 x median breadth (Fig. 29 D), apex elongate, with narrow extension beyond ostial opening and dorsoventrally expanded, spoonlike tip; internal sac without apparent microtrichial patches, moderately elongate flagellar plate visible dorsad parameral articulation in uneverted specimen, flagellar plate length 0.42 x distance from parameral articulation to tip.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
51AC4A78A8C66A8BA80A572B46C7282A.taxon	etymology	Etymology. The Latin adjective affinis - meaning related to or neighboring-is used for this species to signify its close relationship to Mecyclothorax cognatus, a species named with the Latin word that means kindred or related (Brown 1956).	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
A82125154158892C12BAC6FD998202AC.taxon	diagnosis	Diagnosis. Among the mid-sized species in this group-standardized body length 4.7 - 5.7 mm-this species exhibits the broadest body in both pronotal and elytral dimensions. The pronotum is transverse, MPW / PL = 1.26 - 1.33, and basally broad, MPW / BPW = 1.31 - 1.42. This species is broadly sympatric in the Waikamoi area with Mecyclothorax consanguineus, the species most similar in appearance and thus likely to cause confusion. Mecyclothorax cognatus can be distinguished by the broad elytra (Fig. 28 C), with broad humeri, MEW / HuW = 1.83 - 1.94 versus MEW / HuW = 1.91 - 2.04 for Mecyclothorax consanguineus (Fig. 33 B). The pronotum of Mecyclothorax cognatus also exhibits more broadly explanate lateral margins just before the hind angles, that area broadly extended from the deep, smooth laterobasal depressions. Mecyclothorax consanguineus conversely exhibits less explanate lateral margins at that position. If a male is available, the aedeagal tip is absolutely diagnostic even if extended only slightly from the specimen. Those of Mecyclothorax cognatus males have an angulate apex with subangulate tip (Fig. 29 E-G), whereas those of Mecyclothorax consanguineus males have an elongate apex with a mucronate tip (Fig. 34 A-I). Sharp (1903) described Atelothrus optatus based his interpretation that the lateral pronotal setae were missing in the unique type specimen. The left lateral seta is indeed absent, but the articulatory socket for the right lateral seta is evident, resulting in the species setal formula being scored here as 2 2 (1) 2 2.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
29760A70A3810A10D93C631DE936EEA6.taxon	diagnosis	Diagnosis. This species is easily diagnosed by the distinctly cordate pronotum, MPW / BPW = 1.57, with glabrous hind angles. The pronotal lateral margins are distinctly convergent anterad the acute and acuminately projected hind angles (Fig. 33 A). The head, pronotum, and elytra are a uniformly dark rufous. The eyes are moderately convex, ocular ratio = 1.43, but rather small, covering only 3 / 4 of the little protruded ocular lobe; ocular lobe ratio = 0.76. The elytra are broadly subovoid, with the humeri narrowly rounded; MEW / HuW = 2.0. The discal elytral intervals are covered with irregular transverse-line microsculpture, the lines not joined into a mesh. Setal formula 2 1 2 1 [sae]. Standardized body length 5.1 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
29760A70A3810A10D93C631DE936EEA6.taxon	description	Description (n = 1). Head capsule with frontal grooves broad near clypeus, a lateral carina to anterior supraorbital seta; dorsal surface of neck slightly concave; labral anterior margin broadly, shallowly emarginate, antennae filiform, sparse pelage of short setae present on antennomeres 2 - 3; mentum tooth with sides acute, apex tightly rounded. Pronotum moderately transverse, MPW / PL = 1.26; median base depressed relative to disc, with small punctures and lateral wrinkles; basal margin straight, slightly indented posterad laterobasal depressions; median longitudinal impression shallow, very finely incised at depth; anterior transverse impression very shallow, narrow, crossed medially by longitudinal wrinkles; anterior callosity nearly flat, crossed by indistinct wrinkles; front angles slightly projected, tightly rounded; anterior width greater than basal width, APW / BPW = 1.07; lateral marginal depression narrow throughout, margin upturned to finely beaded before sinuation; laterobasal depression smooth, laterally elevated to projected lateral margin. Proepisternum with smooth hind marginal groove; prosternal process medially depressed, a broad lateral marginal bead that is narrowed anteriorly. Elytra with convex disc, sides depressed; basal groove slightly recurved to broadly rounded humeral angle; parascutellar seta present; parascutellar striole continuous, with 4 - 5 punctures; sutural interval more convex than intervals 2 - 4, sutural juncture upraised; sutural and 2 nd striae of subequal depth from base to apex; striae 1 - 8 complete, stria 7 slightly shallower, associated intervals moderately convex; discal striae with slightly irregular punctulae basally, smooth and deep apically; 7 th and 8 th intervals of similar convexity mesad subapical sinuation; 2 dorsal elytral setae at 0.27 x and 0.49 - 0.52 x elytral length; setal impressions extended over 2 / 3 width of interval 3; subapical seta present, apical seta absent; lateral elytral setae arranged in anterior series of 7 setae and posterior series 6 setae; elytral marginal depression narrow throughout, margin slightly upraised at humerus; subapical sinuation shallow, broad. Mesepisternum with ~ 5 punctures in 1 row; metepisternal width to length ratio = 0.81; metepisternum / metepimeron suture distinct. Abdomen with irregular lateral wrinkles on ventrites 1 - 5; suture between ventrites 2 and 3 complete; apical ventrite of female with 4 equally spaced marginal setae and a median trapezoid of 4 setae, the basal pair longer. Legs-metatarsomere 1 / metatibial length ratio = 0.22; metatarsomere 4 length along outer lobe 1.37 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci very broad, median area rough to carinate. Microsculpture of vertex with isodiametric to slightly stretched isodiametric sculpticells in transverse rows; pronotal disc with transverse mesh, sculpticell breadth 3 - 4 x length; pronotal median base with isodiametric to transverse sculpticells; elytral disc with irregular transverse lines, apex with more regular transverse lines; metasternum with distinct transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of antennomere 1 flavous, antennomeres 2 - 3 rufoflavous, 4 - 11 rufobrunneous; proepipleuron and proepisternum rufobrunneous; elytral apex paler than disc, rufoflavous to position of subapical seta; elytral epipleuron rufoflavous laterally, rufobrunneous ventrally, metepisternum rufobrunneous; abdomen medially rufobrunneous, laterally rufoflavous; abdominal apical ventrite 6 with apical 1 / 3 paler, flavous; metafemur flavous with piceous cloud covering basal half; metatibia rufoflavous with brunneous cast. Female reproductive tract. The lone female holotype was not dissected.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
29760A70A3810A10D93C631DE936EEA6.taxon	etymology	Etymology. The species epithet anchisteus is Greek for next of kin (Jaeger 1955), signifying this species' close affinities to Mecyclothorax cognatus, Mecyclothorax affinis, and Mecyclothorax consanguineus.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
09A66087094E8068D60C18CC145DB566.taxon	diagnosis	Diagnosis. Of species in this group characterized by broader, shorter elytra and a basally constricted, quadrisetose pronotum (Figs 28 B-C, 33 B), this species can be diagnosed by the narrowed humeri, MEW / HuW = 1.91 - 2.04. As in the morphologically similar and partially sympatric Mecyclothorax cognatus, the pronotum is moderately constricted basally-MPW / BPW = 1.37 - 1.60 - and the discal elytral striae are finely impressed and lined with minute punctures, the associated intervals slightly convex. The male aedeagal median lobe configuration is unique in the mucronate apex (Fig. 34 A-I). Setal formula 2 2 (1) 2 2; the basal pronotal seta may be unilaterally present, though at least one seta was observed in all examined specimens. Standardized body length 4.1 - 5.3 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
09A66087094E8068D60C18CC145DB566.taxon	description	Description (n = 5). Head capsule with frontal grooves broad near clypeus, a lateral carina to anterior supraorbital seta; dorsal surface of neck flat to slightly concave; eyes moderately developed, ocular ratio = 1.39 - 1.51, ocular lobe ratio = 0.72 - 0.77; labral anterior margin with subangulate emargination, excavated 0.2 x length medially; antennae filiform, antennomeres 2 - 3 with sparse pelage of short setae; mentum tooth with sides acute, apex tightly rounded. Pronotum moderate ly narrow, MPW / PL = 1.15 - 1.25, and moderately to rather constricted basally, MPW / BPW = 1.37 - 1.60; hind angle right to slightly acute, projected, the lateral margin convergent to parallel just anterad the angle; median base very depressed relative to disc, shallow longitudinal wrinkles and small punctures covering surface; basal margin straight medially, slightly indented posterad laterobasal depressions; median longitudinal impression very shallow, indistinct; anterior transverse impression very shallow, broad, crossed by longitudinal wrinkles; anterior callosity nearly flat with wrinkles on the posterior half; front angles projected, tightly rounded; front and basal pronotal angles variably subequal, APW / BPW = 0.92 - 1.03; lateral marginal depression slightly broader at front angle, moderately narrow behind, edge upturned; laterobasal depression smooth with median extension from disc as a tubercle. Proepisternum with 6 punctures along hind marginal groove; prosternal process medially depressed, with broad lateral marginal bead. Elytra with moderately narrow humeri, the disc convex and side moderately sloped; basal groove slightly recurved to hitched humeral angle at base of elytral lateral depression; parascutellar seta present; parascutellar striole shallow, smooth anteriorly with 3 punctures in posterior portion; sutural interval equally convex as intervals 2 - 4 basally, more convex apically; sutural and 2 nd striae of subequal depth from base to apex; sutural stria finely impressed, irregularly punctate basally, smooth and deep apically, striae 2 - 5 with minute punctulae on disc, striae slightly irregular along length; 7 th and 8 th interval of similar convexity mesad subapical sinuation; two dorsal elytral setae at 0.26 - 0.28 x and 0.54 - 0.56 x elytral length, setal impressions small, extended over 1 / 2 width of interval 3; apical and subapical setae present; lateral elytral setae arranged as an anterior series of 7 setae, a posterior series of 6 setae; elytral marginal depression moderately broad at humerus, gradually narrowed to beadlike margin at subapical sinuation; subapical sinuation shallow, nearly obsolete. Mesepisternum with ~ 8 - 9 punctures in 1 - 2 rows; metepisternal width to length ratio = 0.84; metepisternum / metepimeron suture distinct. Abdomen with irregular lateral wrinkles on ventrites 1 - 5; suture between ventrites 2 and 3 complete; apical ventrite of male with 2 marginal setae, apical ventrite of female with 4 equally spaced marginal setae plus a median trapezoid of 4 smaller, subequal setae. Legs-metatarsomere 1 / metatibial length ratio = 0.21; metatarsomere 4 length along outer lobe 1.4 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci deep, broad, median area strigose to carinate. Microsculpture of vertex a transverse mesh, sculpticell breadth 2 x length; pronotal disc with reduced transverse mesh, sculpticell breadth 2 - 3 x length; pronotal median base with distinct isodiametric and transverse sculpticells; elytral disc with transverse mesh, sculpticell breadth 3 - 4 x length; elytral apex with transverse mesh, sculpticell breadth 3 x length, to transverse lines; metasternum with distinct transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex rufobrunneous; antennomere 1 flavous, antennomeres 2 - 3 rufoflavous, 4 - 11 rufobrunneous; pronotal disc rufobrunneous, pronotal apical and lateral margins rufoflavous; proepipleuron rufoflavous, proepisternum rufobrunneous; elytral disc rufobrunneous, sutural interval concolorous to rufoflavous basally, rufoflavous to flavous apically; elytral marginal depression concolorous with disc to paler, rufoflavous basally, apex slightly paler, rufoflavous to position of subapical seta; elytral epipleuron and metepisternum rufobrunneous; abdomen with ventrites medially rufopiceous, ventrites 3 - 6 rufoflavous laterally; abdominal apical ventrite with apical half paler, flavous; metafemur flavous; metatibia rufobrunneous. Male genitalia (n = 15). Aedeagal median lobe distinctly curved, variably robust, with ratio of distance from parameral articulation to tip versus depth at midlength ranging 3.1 - 4.2 (Fig. 34 A, H), but always with apex narrowly extended well beyond ostial opening, and tip denticulate; a brief indentation along apicoventral surface (Fig. 34 A-B, D-I); median lobe straight in ventral view, right margin slightly concave, left margin incurved before apparently blunt tip (Fig. 34 C); internal sac generally lightly spiculated, but with indistinct ventral microtrichial patch comprised of slightly larger microtrichia (Fig. 34 A); flagellar plate relatively large, length 0.45 x distance from parameral articulation to tip. Female reproductive tract (n = 1). Bursa copulatrix columnar with apex rounded, bursa narrowed apically, length 1.05 mm, apical breadth 0.29 mm, basal breadth 0.36 mm equal to vagina breadth (Fig. 30 C); bursal walls translucent with thick wrinkles; gonocoxite 1 with 3 - 4 apical fringe setae and 6 - 7 small setae on medial surface (Fig. 31 C); gonocoxite 2 narrowly subtriangular with broad apex, base broadly extended laterally, 2 lateral ensiform setae with apical seta broader and longer, apical nematiform setae on medioventral surface at 0.74 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
09A66087094E8068D60C18CC145DB566.taxon	etymology	Etymology. The Latin adjectival consanguineus means related by blood, or kindred (Brown 1956), and signifies this species' close relationship to the previous three species.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
6A9F4D8BBD0B9B0429FC46ADA05CE2D7.taxon	diagnosis	Diagnosis. The narrow body and moderately convex elytral intervals with nearly smooth striae set this species apart. Individuals of this species can be diagnosed by the narrow pronotum-MPW / PL = 1.15 - 1.19 - with its lateral margins convergent for 0.1 x the pronotal length anterad the acute, projected hind angles (Fig. 33 C). The elytra are narrowly ellipsoid, with distinctly recurved basal grooves leading to proximate, subangulate humeri; MEW / HuW = 2.04 - 2.15. The elytral disc is rufobrunneous, with a cupreous reflection enhanced by a distinct granulate isodiametric mesh. At the elytral apex, the 8 th interval is more convex than the fused apical portion of intervals 5 + 7. Setal formula 2 2 2 2. Standardized body length 5.7 - 5.8 mm for specimens available to the author, though Britton (1948 b) listed the size range as 4.8 - 5.8 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
7149DFFCB6A1E7EA12E6EF14DD5209CC.taxon	diagnosis	Diagnosis. This species can be diagnosed by the broad elytra with sides subparallel at midlength (Fig. 33 D), a configuration shared with Mecyclothorax robustus and Mecyclothorax haydeni (Fig. 38 C-E), however the discal elytral striae are smooth to only minutely punctate instead of distinctly punctate, and the associated intervals are only moderately convex instead of convex. The pronotum has the lateral margin parallel to slightly convergent anterad the right hind angles, and the pronotal median base is covered with fine punctures. Beetles of the other two species have pronota with parallel to divergent basal lateral margins. The male aedeagal median lobe of Mecyclothorax antaeus males has the apex elongate and parallel sided with a downturned tip (Fig. 34 J-K), versus the apically flattened median lobe of Mecyclothorax robustus (Fig. 41) and the elongate, evenly curved apex observed in males of Mecyclothorax haydeni (Fig. 43). Setal formula 2 2 2 2. Standardized body length 4.6 - 5.9 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
7149DFFCB6A1E7EA12E6EF14DD5209CC.taxon	description	Description (n = 5). Head capsule with frontal grooves broad near clypeus, a lateral carina to anterior supraorbital seta; dorsal surface of neck flat to slightly concave; eyes moderately convex, ocular ratio = 1.46 - 1.50, ocular lobe distinctly protruded from gena, ocular lobe ratio = 0.71 - 0.78, an abruptly depressed slot at juncture of lobe and gena; labral anterior margin broadly shallowly emarginate; antennae filiform, antennomeres 2 - 3 with sparse pelage of short setae; mentum tooth with sides acute, apex tightly rounded. Pronotum subcordate, quadrate to slightly transverse, MPW / BPW = 1.38 - 1.45, MPW / PL = 1.18 - 1.25; hind angle right, projected, the margin behind convex; basal margin slightly convex between hind angles; median longitudinal impression shallow, finely incised; anterior transverse impression broad, shallow, crossed by longitudinal wrinkles, lined with granulate isodiametric microsculpture; anterior callosity slightly elevated, covered with dense longitudinal wrinkles, strigose; front angles slightly projected, rounded; base broader than distance between front angles, APW / BPW = 0.86 - 0.98; lateral marginal depression moderately broad, explanate, edge upturned; laterobasal depression broad, smooth, with median tubercle. Proepisternum with 6 indistinct punctures along hind marginal groove; prosternal process medially depressed, with broad lateral marginal bead. Elytra subquadrate, humeri moderately narrow, rounded, sides subparallel at midlength; elytral disc convex, sides progressively sloped laterally; basal groove recurved to subangulate humeri, the lateral margin upraised, and humeral angle defined by hitch at base of lateral marginal depression, MEW / HuW = 2.02 - 2.13; parascutellar seta present; parascutellar striole deep with 5 punctures; sutural interval more convex than intervals 2 - 4, sutural juncture upraised; sutural and 2 nd striae of subequal depth from base to apex; sutural and lateral discal striae 2 - 6 minutely punctate, striae slightly irregular along length; 7 th and 8 th intervals of similar convexity mesad subapical sinuation; 2 dorsal elytral setae at 0.29 - 0.34 x and 0.55 - 0.60 x elytral length, setal impressions small, extended over 1 / 2 width of interval 3; apical and subapical setae present; lateral elytral setae arranged in an anterior series of 7 (6) setae and a posterior series of 6 setae; elytral marginal depression moderately broad in anterior half, narrowed anterad subapical sinuation; subapical sinuation shallow, broad. Mesepisternum with ~ 14 punctures in 2 - 3 rows; metepisternal width to length ratio = 0.81; metepisternum / metepimeron suture distinct. Abdomen with irregular lateral wrinkles on ventrites 1 - 5; suture between ventrites 2 and 3 complete; apical ventrite of male with 2 marginal setae, apical ventrite of female with 4 equally spaced setae and median patch of 4 (5) short, subequal setae. Legs-metatarsomere 1 / metatibial length ratio = 0.20; metatarsomere 4 length along outer lobe 1.33 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci deep, broad, median area strigose to carinate. Microsculpture of vertex an isodiametric mesh; pronotal disc with transverse mesh, sculpticell breadth 2 - 3 x length; pronotal median base with distinct to granulate isodiametric mesh; elytral disc and apex with transverse mesh, sculpticell breadth 3 x length, to transverse lines; metasternum with distinct transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex rufobrunneous with a purplish reflection; antennomere 1 flavous, 2 - 3 rufoflavous, 4 - 11 rufobrunneous; pronotal disc rufobrunneous with purplish reflection; pronotal margins rufoflavous inside front angles, otherwise concolorous with disc; proepipleuron rufoflavous, proepisternum rufobrunneous; elytral disc rufobrunneous with purplish reflection; sutural interval concolorous with disc basally, rufoflavous apically; elytral lateral marginal depression rufoflavous, apex slightly paler than disc, rufoflavous to position of subapical seta; elytral epipleuron rufoflavous laterally, rufobrunneous ventrally, metepisternum rufobrunneous; abdomen with ventrites medially rufobrunneous, marginally rufoflavous; apical half of ventrite 6 paler, flavous; metafemur flavous; metatibia rufoflavous. Male genitalia (n = 4). Aedeagal median lobe gracile, distance from parameral articulation to tip 4.1 x depth at midlength (Fig. 34 J-K), apex narrowly extended beyond ostial opening, tip downturned with flat apical face; median lobe broadly curved rightward in ventral view, the apex parallel sided and tip blunt (Fig. 34 L); internal sac tubular, elongate, with two ventral microtrichial patches, a larger basal patch and a smaller apical patch situated just distad a median constriction (Fig. 34 J); flagellar plate of moderate size, length of sclerotized internal face 0.38 x distance from parameral articulation to tip. Female reproductive tract (n = 1). Bursa copulatrix columnar with rounded apex, bursa narrower in apical half, length 1.10 mm, apical breadth 0.32 mm, basal breadth 0.51 mm, broader than width of vagina (Fig. 30 E); bursal walls translucent basally with think wrinkles, apical surface shagreened but not wrinkled; gonocoxite 1 with 3 - 4 apical fringe setae and 6 - 9 smaller setae on medial surface (Fig. 37 A); gonocoxite 2 narrowly subtriangular with subacuminate apex, base extended laterally as a thin extension, 2 lateral ensiform setae, the apical seta slightly broader, apical nematiform setae on medioventral surface at 0.74 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
7149DFFCB6A1E7EA12E6EF14DD5209CC.taxon	etymology	Etymology. Antaeus was a " giant Libyan wrestler whose strength was renewed when he touched the earth (Brown 1956), " an apt epithet for this close relative of Mecyclothorax robustus. The name is treated as a noun.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
948A44B26BAFC05E4D3ADC9C08F7E15A.taxon	diagnosis	Diagnosis. Along with Mecyclothorax cymindulus, the smallest-bodied beetles in the group (Fig. 38 A-B) excepting the smallest individuals of Mecyclothorax antaeus (Fig. 33 D); standardized body length for this species 4.1 - 4.7 mm. This and Mecyclothorax cymindulus are also the most narrow-bodied species, with subquadrate elytra and subangulate, laterally extended humeri (Figs 38 A-B). This species deviates from Mecyclothorax cymindulus by the distinctly punctate discal elytral striae, the punctures in basal portions of striae 1 - 5 expanding strial breadth, and by the less transverse microsculpture; The elytral sutural interval is covered with isodiametric microsculpture, with the lateral intervals bearing a transverse mesh, sculpticell breadth 3 - 4 x length, versus the transverse mesh on the sutural interval and transverse lines laterally observed in individuals of Mecyclothorax cymindulus. The eyes also tend to be more convex in this species-ocular ratio = 1.41 - 1.45 - versus the ocular ratio = 1.34 - 1.43 of Mecyclothorax cymindulus. Setal formula 2 2 2 2.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
94975DB64113FC5B6FB8B49F1CF82F9B.taxon	diagnosis	Diagnosis. Differing from its most similar species group member Mecyclothorax cymindicus (Fig. 38 A-B) by: 1, pronotal hind angles obtuse due to rounded basal margin posterad angle, the pronotal lateral margin subparallel anterad angle; 2, broad, minutely punctate discal elytral striae over basal half of length; 3, elytra with transverse-mesh microsculpture on the sutural interval, and parallel-lined microsculpture laterally on disc and at apex. If a male is available, the aedeagal median lobe of Mecyclothorax cymindulus has a more flattened apex with a subangulate, ventrally angled tip (Fig. 34 N), versus the more rounded apex with ventrally expanded tip of male Mecyclothorax cymindicus (Fig. 34 M). Setal formula 2 2 2 2. Standardized body length 4.1 - 4.6 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
94975DB64113FC5B6FB8B49F1CF82F9B.taxon	description	Description (n = 5). Head capsule with frontal grooves straight, a lateral carina to anterior supraorbital seta; dorsal surface of neck flat to slightly concave; ocular lobe moderately extended from gena, eyes not covering posterior portion of lobe, ocular ratio = 1.34 - 1.41, ocular lobe ratio = 0.70 - 0.76; labral anterior margin subangulately excavated medially to 0.2 x length; antennae broader in apical half, submoniliform, antennomeres 2 - 3 with sparse pelage of short setae; mentum tooth with sides acute, apex tightly rounded. Pronotum moderately transverse, MPW / PL = 1.21 - 1.25, base broad, MPW / BPW = 1.41 - 1.49; median base depressed relative to disc, covered with irregular punctures and longitudinal wrinkles; basal margin slightly convex between laterobasal depressions; median longitudinal impression shallow, very finely incised at depth; anterior transverse impression very shallow, broad, crossed by fine longitudinal wrinkles; anterior callosity slightly convex, crossed by numerous wrinkles; front angles not projected, rounded; anterior and basal widths subequal, APW / BPW = 0.95 - 1.04; lateral marginal depression broader, beaded at front angle, evenly expanded along midlength, moderately broad and upturned toward basal angle; laterobasal depression smooth, broad, with or without convex median extension from disc. Proepisternum with 6 punctures along hind marginal groove; prosternal process medially depressed, a broad marginal bead laterally. Elytra with convex disc, sides more sloped; basal groove straight from sutural stria laterally to subangulate humeri; elytral slightly narrowed basally, subquadrate, MEW / HuW = 1.83 - 1.95; parascutellar seta present; parascutellar striole with 4 - 5 punctures, continuous between punctures; sutural interval more convex than lateral intervals; sutural and 2 nd striae of subequal depth from base to apex; discal striae 1 - 5 and base of 6 minutely punctate, striae slightly irregular, associated intervals slightly convex; 7 th and 8 th interval of similar convexity mesad subapical sinuation; 2 dorsal elytral setae at 0. 25 - 0.27 x and 0.57 - 0.59 x elytral length, setal impressions extended over 2 / 3 width of interval 3; apical and subapical setae present; lateral elytral setae arranged as anterior series of 7 setae, posterior series of 6 setae; ely tral marginal depression broadest at humerus, gradually narrowed to a beadlike margin at subapical sinuation; subapical sinuation shallow, abruptly incurved anteriorly. Mesepisternum with ~ 10 punctures in 2 rows; metepisternal width to length ratio = 0.79; metepisternum / metepimeron suture distinct. Abdomen with irregular lateral wrinkles on ventrites 1 - 5; suture between ventrites 2 and 3 reduced, effaced laterally; apical male ventrite with 2 marginal setae, apical female ventrite with 4 equally spaced marginal setae and a median trapezoid of 4 subequally short setae. Legs-metatarsomere 1 / metatibial length ratio = 0.17; metatarsomere 4 length along outer lobe 1.35 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci deep, broad, basal tarsomeres medially carinate. Microsculpture of vertex transverse, sculpticell breadth 2 x length; pronotal disc with transverse mesh, sculpticell breadth 3 - 4 x length; pronotal median base with distinct isodiametric and transverse sculpticells; metasternum with distinct transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex rufobrunneous; antennomeres 1 - 3 flavous, 4 - 11 rufobrunneous; pronotal disc rufobrunneous, pronotal margins slightly paler; proepipleuron and proepisternum rufoflavous; elytral disc rufobrunneous, metallic reflection present due to microsculpture; sutural interval rufous basally, flavous apically; elytral marginal depression narrowly flavous, apex flavous to juncture of intervals 3 and 4; elytral epipleuron flavous, metepisternum rufoflavous; abdomen with ventrites 1 - 6 medially rufoflavous, laterally flavous, ventrite 6 with apical half flavous; metafemur flavous; metatibia rufoflavous. Male genitalia (n = 1). Aedeagal median lobe gracile, distance from parameral articulation to tip 4.0 x depth at midlength (Fig. 34 N); apex moderately extended beyond ostial opening, apical face flat, tip distinctly angulate at juncture of apical face and ventral margin; internal sac with lightly sclerotized dorsal microtrichial patch, and ventral face covered with fine spicules; flagellar plate length 0.47 x parameral articulation-tip distance. Female reproductive tract (n = 1). Bursa copulatrix columnar with rounded apex, narrowed along midlength, length 0.93 mm, medial breadth 0.31 mm, basal breadth at vagina 0.43 mm (Fig. 39 A); bursal walls translucent with thin wrinkles; gonocoxite 1 with 2 - 3 apical fringe setae and 6 - 7 smaller setae on medial surface (Fig. 37 C); gonocoxite 2 narrowly subtriangular with broad apex, tip tightly rounded, 1 - 2 lateral ensiform setae, apical nematiform setae on medioventral surface at 0.76 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
94975DB64113FC5B6FB8B49F1CF82F9B.taxon	etymology	Etymology. This epithet is taken from the Latin noun cymindis, meaning night hawk, modified with the diminutive ending - ule. Beetles of this species are the same size as those of Mecyclothorax cymindicus, but use of the common stem for the epithet is meant to connote the two species' affinities; a convention used by Perrault (1984, 1986, 1988, 1989) to deal with the rampant Mecyclothorax diversity of Tahiti.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
C04DE0182A8A30823362012029BB69EC.taxon	diagnosis	Diagnosis. This and Mecyclothorax haydeni represent the two larger bodied species in this group with subquadrate elytra, the discal elytral striae lined with distinct, round punctures (Figs 38 C-E). Of the two species, the discal elytral striae are more distinctly and regularly punctate in Mecyclothorax robustus, though this characteristic is variable. In aggregate, individuals of Mecyclothorax robustus have larger eyes, ocular ratio = 1.48 - 1.54, that cover most of the ocular lobe, ocular lobe ratio = 0.76 - 0.84, in contrast to beetles of Mecyclothorax haydeni with ocular ratio = 1.41 - 1.49, and ocular lobe ratio = 0.70 - 0.78. Male genitalia are diagnostic, with the aedeagal median lobe of Mecyclothorax robustus males exhibiting an apex with a flattened apical surface and tightly rounded, ventrally directed tip (Fig. 41), versus a median lobe with an evenly downcurved and narrowed apex for Mecyclothorax haydeni (Fig. 43). Setal formula 2 2 2 2. Standardized body length 4.8 - 6.1 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
C7030038E53B3F5227B4A3E6022BF59A.taxon	diagnosis	Diagnosis. Larger bodied beetles within the species group, standardized body length = 5.0 - 6.2 mm, with broad, subquadrate elytra. The discal elytral striae are lined with small though distinct punctures that at most expand strial breadth, or at least introduce irregularities to the strial orientation (Fig. 38 E). The eyes are smaller than those observed in Mecyclothorax robustus individuals (see diagnosis above), though the male genitalic configuration is the only certain means to diagnose the two species. The male median lobe of Mecyclothorax haydeni males exhibits an evenly downcurved apex (Fig. 43), though the degree of narrowing toward the tip and extension beyond the ostial opening varies among individuals (Fig. 43 B versus 43 D). The male aedeagal internal sac is shorter and narrower in males of Mecyclothorax haydeni (Fig. 43 B, F, H) than in males of Mecyclothorax robustus (Fig. 41 B, E, G). Setal formula 2 2 2 2.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
C7030038E53B3F5227B4A3E6022BF59A.taxon	description	Description (n = 5). Head capsule with frontal grooves broad near clypeus, with broad lateral carina to anterior supraorbital seta; dorsal surface of neck flat to slightly concave; labral anterior margin subangulate, medially excavated 0.1 x length; antennae robustly filiform, antennomeres 2 - 3 with sparse pelage of short setae; mentum tooth with sides acute, apex tightly rounded. Pronotum slightly transverse, MPW / PL = 1.20 - 1.24, variably constricted basally, MPW / BPW = 1.37 - 1.50; hind angle slightly obtuse, basal margin rounded posterad angle, lateral margin slightly divergent anterad hind angle; median base depressed relative to disc, covered with dense punctures and wrinkles, surface strigose; basal margin broadly, moderately convex between laterobasal depressions; median longitudinal impression very shallow, finely incised; anterior transverse impression broad, shallow, crossed by wrinkles and lined by granulate isodiametric microsculpture; anterior callosity slightly elevated, crossed by dense longitudinal wrinkles, strigose; front angles slightly projected, rounded; pronotum broader basally than apically, APW / BPW = 0.90 - 0.95; lateral marginal depression only slightly broader at front angle, moderately narrow, edge upturned; laterobasal depression broad, smooth, with median tubercle. Proepisternum with 6 indistinct punctures along hind marginal groove; prosternal process medially depressed, with broad lateral marginal bead. Elytra subquadrate, disc convex, sides more sloped; basal groove recurved to subangulate humerus, the angle defined by hitch in groove at juncture with lateral marginal depression, MEW / HuW = 1.81 - 2.08; parascutellar seta present; parascutellar striole continuous, with 5 small punctures or irregularities along length; sutural interval more convex than lateral intervals, sutural juncture upraised; sutural and 2 nd striae of subequal depth from base to apex; discal striae minutely punctured in basal 1 / 3, smooth and deep apically, associated intervals convex; 7 th and 8 th intervals of similar convexity mesad subapical sinuation; 2 dorsal elytral setae at 0.31 x and 0.63 x elytral length, setal impressions small, extended over 1 / 2 width of interval 3; apical and subapical setae present; lateral elytral setae arranged as anterior series of 7 setae and posterior series of 6 setae; elytral marginal depression moderately narrow throughout length, flat bottomed until subapical sinuation; subapical sinuation evident, moderately deep. Mesepisternum with ~ 14 punctures in two rows; metepisternal width to length ratio = 0.79; metepisternum / metepimeron suture distinct. Abdomen with irregular lateral wrinkles on ventrites 1 - 5; suture between ventrites 2 and 3 complete; apical male ventrite with 2 marginal setae, and apical female ventrite with 4 equally spaced setae and a median trapezoid of 4 subequal, short setae. Legs-metatarsomere 1 / metatibial length ratio = 0.19; metatarsomere 4 length along outer lobe 1.4 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci deep, broad, median area medially strigose. Microsculpture of vertex an isodiametric mesh; pronotal disc with transverse mesh, sculpticell breadth 2 - 3 x length; pronotal median base with distinct to granulate isodiametric mesh; elytral disc and apex with transverse mesh, sculpticell breadth 3 x length, to transverse lines unconnected into a mesh, the apex with more transverse lines; metasternum with distinct transverse mesh; laterobasal abdominal ventrites swirling isodiametric and transverse microsculpture. Coloration of vertex rufobrunneous; antennomere 1 flavous, 2 - 3 rufoflavous, 4 - 11 rufobrunneous; pronotal disc rufobrunneous, margins and apex slightly paler, base concolorous; proepipleuron rufoflavous, proepisternum rufobrunneous; elytral disc rufobrunneous; sutural interval rufous basally, rufoflavous apically; elytral marginal depression slightly paler, elytral apex concolorous with disc; elytral epipleuron rufoflavous, metepisternum rufobrunneous; abdomen with ventrites 1 - 6 medially rufobrunneous, laterally rufoflavous; apical abdominal ventrite 6 with apical half paler, flavous; metafemur flavous; metatibia rufoflavous. Male genitalia (n = 26). Aedeagal median lobe slender, elongate, distance from parameral articulation to tip 4.1 - 5.5 x depth at midlength (Fig. 43 E, H), but apical extension always smoothly curved relative to median lobe shaft, the apex evenly narrowed to a rounded tip; median lobe broadly curved rightward, the apex blunt in ventral view (Fig. 43 C, G); internal sac of variable length (Fig. 43 B, F, H), with consistent presence of two ventral microtrichial patches-basal and apical as in Mecyclothorax antaeus (Fig. 34 J) - the apical patch of variable size; flagellar plate relatively small, length of sclerotized ventral face 0.33 - 0.36 x parameral articulation-tip distance. Female reproductive tract (n = 1). Bursa copulatrix columnar with narrow, rounded apex, length 1.17 mm, apical breadth 0.31 mm, basal breadth 0.47 mm subequal to vagina breadth (Fig. 39 C); bursal walls translucent with thin wrinkles basally, apical surface shagreened but not wrinkled; gonocoxite 1 with 2 - 4 apical fringe setae, a moderately sized seta just basad medioapical angle and 7 - 8 small setae on medial surface (Fig. 37 E); gonocoxite 2 narrowly subtriangular with broad apex and tightly rounded tip, 2 lateral ensiform setae, apical nematiform setae on medioventral surface at 0.78 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
C7030038E53B3F5227B4A3E6022BF59A.taxon	etymology	Etymology. This species is named to honor the contributions of Dr. James E. Hayden, Jr. to this project, wherein he dissected numerous male specimens of Hawaiian Mecyclothorax in order to delineate species boundaries.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
B00EC2B0ECCA1FD2334AAB1155859128.taxon	diagnosis	Diagnosis. This is the broadest-bodied species in the Haleakala Mecyclothorax fauna, exhibiting a large prothorax and broadly subquadrate elytra, MEW / MPW = 1.30 - 1.32 (Fig. 44 A). The pronotal lateral margins are only slightly sinuate anterad the obtuse-rounded hind angles; and the linear laterobasal depression is separated from the pronotal lateral margin by a broad tubercle. The elytral striae are nearly regular, though striae 2 and 3 may approach unilaterally. The dorsal elytral setal impressions are foveate anteriorly, crossing interval 3 at the anterior seta, and smaller posteriorly, impressing half the interval width at the posterior seta. The dorsal body coloration is uniformly rufobrunneous, with the legs paler, rufoflavous. Setal formula 2 2 2 2. Standardized body length 5.2 - 5.3 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
B00EC2B0ECCA1FD2334AAB1155859128.taxon	distribution	Distribution. Mecyclothorax integer is known only from the lectotype and paralectotype collected by Perkins in his lots 622 and 680; " Haleakala 4000 ft., v- 1896 ", and " Haleakala 4000 + ft., x- 1896 " (Anonymous N D), respectively (Fig. 48). Of the October visit, Perkins (1896 b) wrote: " In October 1896 I camped for a considerable time at about 5,000 ft. on Haleakala and did a good deal of work at various points Eastward in the windward forest entering this at various points from the upper edge towards the forest that lies above Wailua (p. 1). " This locality would lie about 200 ft. elevation below the Ukulele Camp (site) of the USGS (1983) Kilohana, Hawaii quadrangle. Recent collections near this site did not result in rediscovery of this species.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
D3B946421709CE2054582994FAD96E19.taxon	diagnosis	Diagnosis. This species (Fig. 44 B), Mecyclothorax anthracinus (Fig. 44 E), and Mecyclothorax inconscriptus (Fig. 50 A) represent the three smallest-bodied species in this group, with all individuals equal to or less than 4.1 mm length. Of these, Mecyclothorax bradycelloides is the only species without fused elytral striae, though the impressions of the dorsal elytral setae are large, crossing most of the width of interval 3. The pronotum is also the most constricted basally, with MPW / BPW = 1.56 versus a collective span of 1.42 - 1.53 for the other two species. The elytral margins are straight and nearly parallel at elytral midlength, though the tightly rounded humeral angles are narrowly separated relative to elytral width due to the broadly rounded elytral margins laterad the angles; MEW / HuW = 2.0. Setal formula 2 2 2 2. Standardized body length 4.1 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
D3B946421709CE2054582994FAD96E19.taxon	description	Description (n = 1). Head capsule with frontal grooves deep and broad near clypeus, narrowed toward anterior supraorbital seta; dorsal surface of neck slightly convex; ocular ratio = 1.49, ocular lobe ratio = 0.81; antennae filiform, antennomere 3 with sparse pelage of short setae; mentum tooth with sides acute, apex tightly rounded. Pronotum cordate, hind angle obtuse, rounded behind; lateral margin sinuate for short distance anterad hind angle; median base covered with punctures and wrinkles isolated by granulate microsculpture; basal margin nearly straight, slightly convex medially; median longitudinal impression shallow, finely incised; anterior transverse impression deeply incised, complete, crossed by longitudinal wrinkles; anterior callosity broadly, slightly convex; front angles projected, tightly rounded, APW / BPW = 1.07; lateral marginal depression narrow, edge upturned to beaded; laterobasal depression broad, irregular, slightly convex medially. Proepisternum with smooth hind marginal groove; prosternal process medially depressed, with lateral marginal bead. Elytra subquadrate, disc slightly convex; parascutellar seta present on left side, absent on right; parascutellar striole shallow with 6 punctures; sutural interval moderately convex, appearing broader than intervals 2 - 4 due to elevated juncture at suture; sutural and 2 nd striae of subequal depth from base to apex; discal striae with small punctures that cause strial irregularities along length, intervals convex; 7 th and 8 th intervals of similar convexity mesad subapical sinuation; 2 dorsal elytral setae at 0.23 - 0.27 x and 0.60 - 0.63 x elytral length; apical and subapical setae present; lateral elytral setae arranged as anterior series of 7 setae, and posterior series of 4 - 5 setae; elytral marginal depression moderately broad, lined with sculpticells, margin upturned; subapical sinuation very shallow, broad. Mesepisternum with ~ 9 shallow punctures in 1 - 2 rows; metepisternal width to length ratio = 0.81; metepisternum / metepimeron suture distinct. Abdomen with nearly smooth ventrites; suture between ventrites 2 and 3 complete; apical male ventrite with 2 marginal setae. Legs-metatarsomere 1 / metatibial length ratio = 0.17; metatarsomere 4 length along outer lobe 1.2 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci broad, deep, median area carinate. Microsculpture of vertex isodiametric sculpticells arranged in transverse rows; pronotal disc with transverse mesh, the microsculpture parallel in part; pronotal median base with granulate isodiametric sculpticells; elytral disc with irregular elongate transverse mesh and parallel lines; elytral apex with upraised transverse mesh; metasternum with indistinct transverse mesh; laterobasal abdominal ventrites with indistinct transverse mesh. Coloration of vertex rufobrunneous; antennomeres 1 - 3 flavous, 4 - 11 rufobrunneous; pronotal disc rufobrunneous, margins narrowly paler, rufoflavous; proepipleuron flavous, proepisternum brunneous; elytral disc rufobrunneous, sutural interval concolorous basally, rufoflavous apically, margins narrowly rufoflavous in lateral depression, apex broadly flavous; elytral epipleuron flavous with rufous cast, metepisternum rufobrunneous; abdominal ventrites 1 - 5 rufobrunneous, 6 basally rufoflavous, flavous in apical half; metafemur flavous; metatibia flavous with brunneous cast. Male genitalia (n = 1). Aedeagal median lobe robust, distance from parameral articulation to tip 3 x depth at midlength (Fig. 45 B); apex narrowly extended twice its depth beyond ostial opening, tip subangulate where flattened apical face and ventral margin meet; median lobe straight in ventral view, the left margin distinctly incurved to meet blunt tip (Fig. 45 C); internal sac with well-developed, heavily sclerotized dorsal ostial microtrichial patch (based on position near apex of ostium (Fig. 45 B), and separate ventral ostial microtrichial patch (Fig. 45 C); flagellar plate well sclerotized, visible just inside dorsal margin of lobe (Fig. 45 B).	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
D3B946421709CE2054582994FAD96E19.taxon	etymology	Etymology. The adjectival epithet bradycelloides is based on Moloka'i's Mecyclothorax bradycellinus Sharp, with the - oides suffix signifying the similarity between the two species.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
12C6BFC617113F90347CDBBC4EB5F435.taxon	diagnosis	Diagnosis. The irregularly anastomosed elytral striae 2 - 6, combined with the bicolored dorsal surface-piceous head, testaceous pronotum, plus testaceous elytral convexities versus piceous striae (Fig. 44 C) - uniquely diagnose this species within the Hawaiian Mecyclothorax fauna. The ventral body coloration is equally distinctive, with the piceous genae, thoracic ventrites, metacoxae, and mediobasal portions of the basal abdominal ventrites distinctly contrasted to the pale mentum and gula, pronotal and elytral epipleura, and abdominal apex (Fig. 44 D). Setal formula 2 2 2 2. Standardized body length 4.7 - 5.2 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
38D92CD0438683BAFE7889960297E4A5.taxon	diagnosis	Diagnosis. The small, dark-bodied beetles that comprise this species look ever so like small bits of anthracite coal, their dorsal body surface a reflective black (Fig. 44 E). Also, this is the only species in the group that is characterized by absence of the parascutellar seta. The pronotum is narrow and basally constricted, MPW / PL = 1.18 - 1.24, MPW / BPW = 1.42 - 1.50, the disc covered with well-developed transverse wrinkles. The elytral intervals are convex and striation irregular, with striae 5 and 6 fused near the basal groove, and the dorsal setal impressions foveate and of diameter equal to the width of interval 3, these impressions associated with longitudinal irregularities of striae 2 and 3. The femora are flavous apically and covered with a piceous cast over their basal third; tibiae piceous. Setal formula 2 2 2 2. Standardized body length 3.6 - 4.0.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
38D92CD0438683BAFE7889960297E4A5.taxon	description	Description (n = 5). Head capsule with frontal grooves deep near clypeus, straight with external carina to anterior supraorbital seta; dorsal surface of neck convex, eyes moderately convex, ocular ratio = 1.42 - 1.50, ocular lobe ratio 0.77 - 0.85; labral anterior margin broadly, shallowly emarginate, antennae filiform, antennomere 3 with sparse pelage of short setae; mentum tooth with orthogonal sides, apex pointed. Pronotum with lateral margin subparallel to convergent anterad right to acute hind angle; median base depressed relative to disc, covered with rugose wrinkles; basal margin nearly straight, slightly convex medially; median longitudinal impression shallow, finely incised, crossed by wrinkles; anterior transverse impression broad, evident, bordered anteriorly by slightly convex anterior callosity that is crossed by fine wrinkles; front angles slightly projected, rounded; anterior width subequal to broader than basal width, APW / BPW = 1.00 - 1.06; lateral marginal depression moderately narrow, edge upturned to beaded; laterobasal depression broad, depressed with wrinkled surface. Proepisternum with 6 minute punctures along hind marginal groove; prosternal process medially depressed, without marginal bead. Elytra subellipsoid, disc flat, sides moderately sloped; basal groove slightly recurved to tightly rounded humeral angle; humeri narrow, MEW / HuW = 2.06 - 2.10; parascutellar striole finely incised, continuous; sutural interval moderately convex, slightly more upraised than intervals 2 - 4; sutural and 2 nd striae of subequal depth from base to apex; discal striae 1 - 8 complete and deep to apex, smooth with minute irregularities along striae suggesting punctulae; 7 th and 8 th intervals of similar convexity mesad subapical sinuation; 2 dorsal elytral setae at 0.32 x and 0.65 - 0.73 x elytral length; apical and subapical setae present; lateral elytral setae arranged in anterior series of 7 setae and posterior series of 4 (5) setae; elytral marginal depression narrow, edge upturned; subapical sinuation very shallow, nearly obsolete. Mesepisternum with 8 shallow punctures in 2 rows; metepisternal width to length ratio = 0.71; metepisternum / metepimeron suture distinct, metathoracic flight wing an ovoid flap, length 2.5 x breadth, with reduced R and M veins, the flap extended 2 / 3 distance to hind margin of metanotum. Abdomen with indistinct lateral wrinkles on ventrites 1 - 3; suture between ventrites 2 and 3 complete; apical male ventrite with 2 marginal setae, apical female ventrite with 4 equally spaced marginal setae and a median trapezoid of 4, subequal short setae. Legs-metatarsomere 1 / metatibial length ratio = 0.17; metatarsomere 4 length along outer lobe 1.2 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci broad, shallow. Microsculpture of vertex of granulate isodiametric sculpticells; pronotal disc covered with distinct transverse mesh, median base with granulate isodiametric sculpticells; elytral disc covered with well-developed transverse mesh, apex with well-developed isodiametric mesh in transverse rows; metasternum with upraised transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex granulate rufopiceous; antennomere 1 flavous, antennomeres 2 - 3 with piceous cast, 4 - 11 piceous; pronotal disc granulate rufopiceous, margins narrowly paler, rufobrunneous; proepipleuron rufobrunneous with piceous upper margin, proepisternum rufobrunneous with piceous cast; elytral disc rufopiceous, sutural interval paler, dark rufous throughout, margins narrowly paler basally, concolorous with disc apically; elytral epipleuron rufobrunneous, metepisternum rufopiceous; abdomen rufopiceous across width of ventrites 1 - 5, apical ventrite 6 with apical 1 / 3 paler, rufobrunneous. Male genitalia (n = 1). Aedeagal median lobe curved, gracile, distance from parameral articulation to tip 4 x depth at midlength (Fig. 45 F); apex angularly narrowed to tightly rounded tip formed at juncture of flat apical face and ventral margin; median lobe sinuously recurved left then right in ventral view (Fig. 45 G), tip tightly rounded; internal sac with apparent dorsal ostial microtrichial patch (based on uneverted specimen; Fig. 45 F), sac surface covered with microspicules; flagellar plate evident just inside dorsal margin of median lobe. Female reproductive tract (n = 1). Bursa copulatrix a narrow digitiform lobe attached to broader vagina, lobe length 0.26 mm, lobe apical breadth 0.10 mm, vagina breadth 0.25 mm (Fig. 46 B); bursal walls thin, transparent; gonocoxite 1 with 3 apical fringe setae and 5 - 6 smaller setae on medial surface (Fig. 47 B); gonocoxite 2 falcate, narrow apically with base broadly extended laterally, 2 lateral ensiform setae with apical seta broader and longer, apical nematiform setae on medial surface at 0.75 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
38D92CD0438683BAFE7889960297E4A5.taxon	etymology	Etymology. The shiny coal black color of the dorsal surface of these beetles begs for use of the Latin adjective anthracinus.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
87A27FED88931FEB9391480A777FD7E1.taxon	diagnosis	Diagnosis. This species can be diagnosed by the narrow pronotum relative to the elytra, MEW / MPW = 1.64 - 1.67, and the irregularly anastomosing striae 2 - 7, resulting in massive convex warts that may or may not be bilaterally symmetrical (Fig. 49 A-B). These characters set this species apart from all others from Haleakala except Mecyclothorax medeirosi below. That species differs by a relatively broader pronotum (Fig. 49 C); MEW / MPW = 1.47. A third species, Mecyclothorax oppenheimeri Liebherr from West Maui, shares the warty elytral condition of Mecyclothorax arthuri and Mecyclothorax medeirosi while exhibiting a pronotum of relative width intermediate to that of those two species; i. e. MEW / MPW = 1.52 - 1.57 (Liebherr 2011, fig. 36). Setal formula: 2 1 2 0 (1 - 2) [sae]. The variation in the apical elytral setae is distributed as: 5 individuals with both setae absent; 1 individual with apical seta absent and subapical setae present; and 1 individual with both apical and subapical setae present. Standardized body length 4.0 - 4.4 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
87A27FED88931FEB9391480A777FD7E1.taxon	description	Description (n = 5). Head capsule with frontal grooves deep near clypeus, straight, with external carina; dorsal surface of neck flat to convex; eyes moderately convex, ocular ratio = 1.43 - 1.47, covering 3 / 4 of slightly protruded ocular lobe, ocular lobe ratio = 0.72 - 0.80; labral anterior margin broadly, moderately deeply emarginate; antennae filiform, antennomeres 2 - 3 with sparse pelage of short setae; mentum tooth with sides acute, apex rounded. Pronotum bisetose, lateral seta present, basal seta absent, basally constricted, MPW / BPW = 1.46 - 1.53; hind angle acute, apex acuminate, lateral margin broadly convergent anterad angle; median base depressed relative to disc, with sparsely distributed punctures near basal margin, longitudinal wrinkles at juncture with disc; basal margin straight between laterobasal depressions; median longitudinal impression shallow, indistinct, to deep, distinct, always crossed by transverse wrinkles emanating onto disc; anterior transverse impression deeply incised, complete, short wrinkles extended from impression posteriorly onto disc; anterior callosity broadly convex, smooth; front angles slightly produced, tightly rounded; width between front angles greater than basal width, APW / BPW = 1.01 - 1.08; lateral marginal depression narrow, edge beaded; laterobasal depression broad, a depressed expansion of lateral depression. Proepisternum with 6 minute punctures along hind marginal groove; prosternal process medially depressed, with broad marginal bead. Elytra broadly subquadrate, lateral margins convex from humerus to subapical sinuation; basal groove incrementally recurved, bordering 4 basal convexities mesad tightly rounded to subangulate humeral angle; humeri narrow relative to broadest portion of elytra behind midlength; MEW / HuW = 2.07 - 2.16; parascutellar seta present; parascutellar striole narrow, deep, directly connected to isolated basal portion of sutural stria; sutural interval as convex as interval 2, though less convex than warty protuberances associated with intervals 3 - 7; discal striae lined with sculpticells, smooth; sutural and 2 nd striae of subequal depth in apical half of elytra; discal striae 2 - 6 joined irregularly at positions of dorsal elytral setae, larger convexities associated with striae 2 - 4, smaller more irregular convexities laterad; warty convexities vary among individuals, as well as bilaterally (Fig. 49 A-B); 7 th and 8 th interval of similar convexity mesad subapical sinuation; 2 dorsal elytral setae at 0.31 x and 0.69 x elytral length, setal impressions very small, shallow, not distinctly associated with a depressed stria; lateral elytral setae arranged in anterior series of 5 setae, an isolated intermediate seta, and 4 posterior setae; elytral marginal depression broad laterally, narrow behind; subapical sinuation shallow, broad. Mesepisternum with 8 shallow punctures in 2 rows; metepisternal width to length ratio = 0.72; metepisternum / metepimeron suture distinct; metathoracic flight wing an ovoid flap 3.3 x long as broad, remnant R and M veins present, flap extended to hind margin of metanotum. Abdomen with indistinct lateral wrinkles on ventrites 1 - 3; suture between ventrites 2 and 3 complete; apical male ventrite with 2 marginal setae, apical female ventrite with 4 equally spaced marginal setae and median trapezoid of 4 subequal, short setae. Legs-metatarsomere 1 / metatibial length ratio 0.20; metatarsomere 4 length along outer lobe 1.2 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci broad, shallow. Microsculpture of vertex a granulate isodiametric mesh; pronotal disc with shallow transverse mesh median base with isodiametric sculpticells and glossy portions; elytral disc and apex with very shallow isodiametric sculpticells in transverse rows; metasternum with distinct transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex rufobrunneous with piceous cast; antennomere 1 rufoflavous, antennomeres 2 - 11 rufopiceous; pronotal disc rufobrunneous, pronotal margins broadly paler, rufoflavous; proepipleuron flavous, proepisternum rufous; elytral disc rufobrunneous, sutural interval concolorous basally, rufous apically; elytral marginal depression narrowly rufoflavous, apex concolorous with disc; elytral epipleuron rufoflavous, metepisternum rufous; abdomen rufobrunneous mediobasally, all ventrites flavous laterally, the apical ventrite with apex broadly flavous; metafemur flavous with piceous basal cloud; metatibia rufobrunneous, piceous cast medially. Male genitalia (n = 1). Aedeagal median lobe slender, distance from parameral articulation to tip 5.1 x depth at midlength (Fig. 45 H); apex extended 4 x its depth beyond ostial opening, gradually narrowed to narrowly rounded tip; median lobe constricted laterally toward apex in ventral view, right margin distinctly concave, left margin more gradually narrowed, tip blunt (Fig. 45 J); internal sac with bulbous ventral lobe near midlength, apex broadly rounded with small sclerotized flagellar plate visible along dorsal surface of apical lobe (Fig. 45 H-I), sac surface uniformly covered with microspicules. Female reproductive tract (n = 1). Bursa copulatrix very short, broad, little extended from broad vaginal base, length 0.23 m, breadth 0.40 mm (Fig. 46 C); bursal walls translucent, thinly wrinkled; gonocoxite 1 with 3 - 4 apical fringe setae in oblique series, 4 - 6 smaller setae on medial surface (Fig. 47 C); gonocoxite 2 subtriangular, apex evenly narrowed, base broadly extended laterally, 2 narrow lateral ensiform setae and broad dorsal ensiform seta with rounded apex, apical nematiform setae on medioventral surface at 0.75 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
87A27FED88931FEB9391480A777FD7E1.taxon	etymology	Etymology. Patronyms used in this revision honor the contributions of colleagues to the work in hand. The immense contributions of Dr. Arthur Medeiros in teaching the author how to conduct operative science in the Hawaiian rainforest made this entire work possible. Thus this is the first of two patronyms to honor him.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
5EA7889D8CF66FA926A835EAD3DF26A6.taxon	diagnosis	Diagnosis. Like Mecyclothorax arthuri in exhibiting tortuously anastomosed elytral striae, though in this species all striae from the sutural to 7 th stria are involved (Fig. 49 C). This species also differs in the relatively broader pronotum; MEW / MPW = 1.47 versus a ratio of 1.64 - 1.67 for the former species. The pronotum itself is broader, more transverse, MPW / PL = 1.38, versus MPW / PL = 1.21 - 1.26 in Mecyclothorax arthuri. The elytra are also broader basally in this species, with the lateral elytral margins rounded more broadly laterad the angulate humeri (Fig. 49 C) than observed in individuals of Mecyclothorax arthuri (Fig. 49 A-B); MEW / HuW = 1.98 versus values of 2.07 - 2.16 for the other species. Finally, elytral setation differs between the species, with Mecyclothorax medeirosi characterized by three dorsal elytral setae and presence of both apical and subapical setae, producing a setal formula of 2 1 3 2. Standardized body length 4.35 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
5EA7889D8CF66FA926A835EAD3DF26A6.taxon	description	Description (n = 1). Head capsule with frontal grooves sinuous, broad near clypeus and with lateral carina posteriorly; dorsal surface of neck slightly concave; eyes convex, ocular lobe protruded from gena, ocular ratio = 1.48, ocular lobe ratio = 0.75 - 0.79; labral anterior margin broadly, moderately emarginate, antennae filiform, antennomeres 2 - 3 with sparse pelage of short setae; mentum tooth with sides acute, apex rounded. Pronotum bisetose, hind angles glabrous; base moderately constricted, MPW / BPW = 1.51; hind angle right to obtuse with rounded apex, lateral margin subparallel to slightly divergent anterad angle; median base smooth medially, 5 - 6 punctures mesad laterobasal depressions; basal margin slightly convex medially; median longitudinal impression deep, broad, medially incised, crossed by transverse wrinkles; anterior transverse impression deeply incised, complete, slightly irregular; anterior callosity broadly convex, crossed by many fine wrinkles; front angles slightly projected, tightly rounded; apical and basal widths subequal, APW / BPW = 1.02; lat eral marginal depression moderate, edge upturned to beaded; laterobasal depression broad, flat to slightly upraised by low tubercle. Proepisternum with 5 minute punctures along hind marginal groove; prosternal process medially depressed, with narrow marginal bead. Elytra broadly subquadrate, disc moderately convex along entire length; basal groove distinctly recurved to subangulate humeral angle; parascutellar seta present; parascutellar striole narrow, deep, isolated from sutural stria; sutural and 2 nd striae of subequal depth from base to apex; adjacent striae from the sutural stria to stria 7 confusedly fused, producing numerous, approximately symmetrical wartlike protuberances, all intervals convex, the warts incorporating portions of more than one interval; discal striae smooth, lined with sculpticells only; 7 th and 8 th interval of similar convexity mesad subapical sinuation; 3 dorsal elytral setae at 0.29 x, 0.51 x, and 0.64 x elytral length, setal impressions very small, shallow, but associated with depressed discal strial fusions; apical and subapical setae present; lateral elytral setae arranged as anterior series of 7 setae and a posterior series of 5 - 6 setae, with an isolated intermediate seta present on left side; elytral marginal depression broad along anterior setal series, moderate at midlength, narrower behind; subapical sinuation shallow, broad. Mesepisternum with ~ 9 shallow punctures in 1 - 2 rows; metepisternum nearly quadrate, width to length ratio = 0.93; metepisternum / metepimeron suture distinct. Abdomen with indistinct lateral wrinkles on ventrites 1 - 3; suture between ventrites 2 and 3 complete; apical female ventrite with 4 equally spaced marginal setae, a median trapezoid of 4 setae, the basal setae shorter, a 5 th seta on right side. Legs-metatarsomere 1 / metatibial length ratio = 0.15; metatarsomere 4 length along outer lobe 1.67 x me dian base, the tarsomere broad, robust, apical and subapical setae present; metatarsal dorsolateral sulci broad, shallow. Microsculpture of vertex a granulate isodiametric mesh; pronotal disc with granulate isodiametric mesh, some sculpticells in transverse rows; pronotal median base with granulate isodiametric mesh; elytral disc and apex with very shallow isodiametric sculpticells in shallow rows; metasternum with upraised transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex rufobrunneous with piceous cast; antennomere 1 flavous, antennomeres 2 - 11 piceous; pronotal disc rufobrunneous, margins broadly paler, rufoflavous; proepipleuron flavous, proepisternum piceous (Fig. 49 D); elytral disc with rufobrunneous intervals and darker brunneous depressions and striae, sutural interval concolorous with lateral intervals; elytral margins rufoflavous, apex broadly flavous; elytral epipleuron flavous, metepisternum piceous; abdomen broadly brunneous, a piceous cast mediobasally, abdominal apical ventrite broadly flavous (Fig. 49 D); metafemur flavous with indistinct median rufous cloud; metatibia rufobrunneous. Female reproductive tract. The lone female holotype was not dissected.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
5EA7889D8CF66FA926A835EAD3DF26A6.taxon	etymology	Etymology. This is the second patronym to honor the contributions of Dr. Art Medeiros, most specifically for field collecting efforts validated in this revision, and more broadly for his leadership in Hawaiian conservation biology (e. g., Perkins et al. 2014). Both Mecyclothorax arthuri and Mecyclothorax medeirosi occupy distributional ranges in Kīpahulu Valley and the Hana Bogs region of Haleakala (Fig. 51). Based on the species known to science to date, the species' hypothesized sister-taxon relationship can be signified by the convention, Mecyclothorax arthuri + Mecyclothorax medeirosi.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
DF660E80CC9C82AC791D6542E178C239.taxon	diagnosis	Diagnosis. This is one of the three species in the group with body lengths of 4.1 mm or less, accompanying Mecyclothorax bradycelloides and Mecyclothorax anthracinus in that distinction, however Mecyclothorax inconscriptus (Fig. 50 A) incongruently differs from those species by sharing with Mecyclothorax foveolatus and Mecyclothorax interruptus below (Fig. 50 B-D), the state of elytral striae 2 - 4 fused in association with the dorsal elytral setae. The pronotum is basally constricted, MPW / BPW = 1.49 - 1.52, versus values of 1.38 - 1.47 for Mecyclothorax foveolatus and Mecyclothorax interruptus. Pronotal setation also differs from all four species mentioned above in that the basal setae are absent from a slight majority of available specimens, and in the minority of specimens that exhibit the basal setae, they are small and underdeveloped; therefore setal formula 2 1 (2) 2 2. Standardized body length 3.7 - 4.0 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
DF660E80CC9C82AC791D6542E178C239.taxon	description	Description (n = 5). Head capsule with frontal grooves deep near clypeus, straight with lateral carina; dorsal surface of neck convex; ocular lobe obtusely projected from genal surface, eyes little convex, ocular ratio = 1.36 - 1.44, ocular lobe ratio = 0.72 - 0.77; labral anterior margin broadly, shallowly emarginate; antennae filiform, antennomere 3 with sparse pelage of short setae; mentum tooth with sides acute, apex tightly rounded. Pronotum transverse, MPW / PL = 1.31 - 1.37; hind angle obtuse rounded to denticulate, lateral margin straight for short distance to immediately sinuate anterad angle; median base with dense elongate punctures and wrinkles; basal margin nearly straight, slightly convex medially; median longitudinal impression shallow, finely incised, crossed by fine wrinkles; anterior transverse impression shallow, broad, crossed by fine wrinkles; anterior callosity slightly convex, crossed by fine wrinkles; front angles slightly projected, tightly rounded; apical width subequal to slightly larger than basal width, APW / BPW = 1.0 - 1.04; lateral marginal depression moderate, edge upturned to beaded; laterobasal depression broad, surface wrinkled. Proepisternum with 6 minute punctures along hind marginal groove; prosternal process medially depressed with broad marginal bead. Elytra subquadrate, disc flat, sides moderately sloped; basal groove slightly recurved to rounded humeral angle, the humeri moderately narrowed relative to greatest width behind midlength, MEW / HuW = 1.93 - 2.05; parascutellar seta present; parascutellar striole fine, continuous, with 6 small punctures; sutural interval moderately convex, slightly more elevated than intervals 2 - 4; sutural and 2 nd striae of subequal depth from base to apex; discal striae deep, impunctate, continuous to apex with exception of strial fusions; strial fusions include those of striae 3 and 4 in association with dorsal elytral setae, striae 5 and 6 posterad humerus, and irregularly striae 3 and 4, 4 and 5, and 5 and 6 near apex (Fig. 50 A); 7 th and 8 th interval of similar convexity mesad subapical sinuation; 2 dorsal elytral setae at 0.29 x and 0.61 - 0.65 x elytral length, setal impressions foveate, spanning interval 3; apical and subapical setae present; lateral elytral setae arranged in anterior series of 7 setae and posterior series of 5 setae; elytral marginal depression narrow, lateral margin upturned; subapical sinuation very shallow, nearly obsolete. Mesepisternum with 8 shallow punctures in 2 rows; metepisternal width to length ratio = 0.86; metepisternum / metepimeron suture distinct; metathoracic flight wing an ovoid flap 2.5 x long as wide, remnant R plus M veins present, vestige extends 3 / 4 distance to posterior margin of metanotum. Abdomen with indistinct lateral wrinkles on ventrites 1 - 3; suture between ventrites 2 and 3 complete; apical male ventrite with 2 marginal setae, apical female ventrite with 4 equally spaced marginal setae and median trapezoid of 4 subequal, short setae. Legs-metatarsomere 1 / metatibial length ratio = 0.17; metatarsomere 4 length along outer lobe 1.3 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci broad, shallow. Microsculpture of vertex granulate iso diametric; pronotal disc with granulate transverse mesh, median base with granulate isodiametric mesh; elytral disc with elongate transverse mesh, apex with well-developed isodiametric mesh in transverse rows; metasternum with upraised transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex rufobrunneous; antennomere 1 flavous, antennomeres 2 - 3 rufopiceous, 4 - 11 piceous; pronotal disc rufobrunneous with piceous cast, margins concolorous; proepipleuron rufoflavous, proepisternum rufobrunneous with piceous cast; elytral disc rufopiceous, sutural interval paler, rufous throughout length, lateral marginal depression paler at base, 8 th stria and depression paler at apex; elytral epipleuron rufoflavous, metepisternum rufobrunneous; abdomen rufopiceous across width of ventrites 1 - 5, apical ventrite 6 with apical 1.3 paler, rufobrunneous; metafemur flavous with piceous cast in basal 1 / 3; metatibia flavous with piceous cast. Male genitalia (n = 1). Aedeagal median lobe short, robust, distance from parameral articulation to tip 2.2 x depth at midlength (Fig. 45 K); apex narrowly extended beyond ostial opening 3 x its depth, tip tightly rounded at juncture of flattened apical face and downturned ventral margin; median lobe straight in ventral view, right and left margins convergent to blunt tip (Fig. 45 L); internal sac unornamented, flagellar plate large, sclerotized plate visible in lateral view, plate length 0.58 x parameral articulation-tip distance. Female reproductive tract (n = 1). Bursa copulatrix very broad basally, narrowed to a nipplelike apical lobe, overall length 0.51 mm, apical lobe width 0.09 mm, basal width at vagina 0.44 mm (Fig. 46 D); bursal walls transparent at base, wrinkled in extension apicad juncture with common oviduct, apical lobe translucent, thinly wrinkled; gonocoxite 1 with 2 subequal apical fringe setae, 4 smaller setae on medial surface (Fig. 47 D); gonocoxite 2 broadly subtriangular, apex tightly rounded, base broadly extended, 2 lateral ensiform setae with apical seta broader and longer, apical nematiform setae on medioventral surface at 0.72 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
DF660E80CC9C82AC791D6542E178C239.taxon	etymology	Etymology. The Latin participle conscriptus is used as the stem of this epithet, with the converse inconscriptus signifying this species' membership in the Mecyclothorax interruptus species group.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
0FA27A045FE37B5D49375FA24B1EE57B.taxon	diagnosis	Diagnosis. This and the following species, Mecyclothorax interruptus, are cryptic sibling species with only subtle differences both externally (Fig. 50 B, D) and in the male genitalia (Fig. 45 M-P), but nevertheless they may be consistently diagnosed morphologically. This species differs from its sibling in its more upraised cuticular microsculpture; 1, vertex covered by a granulate isodiametric mesh, the surface appearing coriaceous, matte and non-reflective; 2, pronotal disc with coriaceous isodiametric and transverse sculpticells in transverse rows, the pronotal base with granulate isodiametric mesh, again the surface with a matte finish. The corresponding microsculpture in Mecyclothorax interruptus individuals is: 1, vertex with isodiametric mesh, the sculpticell surfaces shiny in part; 2, pronotal disc with mixture of isodiametric and transverse sculpticells, the surface iridescent, and pronotal base with flattened isodiametric sculpticells, the surface shiny in part. The discal elytral striae are also smoother in this species, with minute punctulae at the deepest parts of the striae associated with longitudinal irregularities in the strial orientation, whereas in Mecyclothorax interruptus individuals the striae, especially the sutural and 2 nd stria, are minutely but clearly punctate in their deepest portions, with those small punctures slightly expanding the stria at its deepest point. The male median aedeagal lobe apex in this species has a distinct obtuse tooth on its dorsal surface (Fig. 45 M-N) versus the lobe apex of Mecyclothorax interruptus males with a rounded dorsal projection (Fig. 45 O-P). The median lobe tip is also flattened apically in this species, versus more evenly rounded in Mecyclothorax interruptus. Setal formula 2 2 2 2. Standardized body length 4.8 - 5.9 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
0FA27A045FE37B5D49375FA24B1EE57B.taxon	description	Description (n = 5). Head capsule with frontal grooves deep and broad near clypeus, sinuously directed to terminus mesad anterior supraorbital seta; dorsal surface of neck flat to convex; eyes moderately convex, ocular ratio = 1.42 - 1.50, ocular lobe distinctly projected from gena; ocular lobe ratio 0.79 - 0.88; labral anterior margin with broad, moderately deep emargination; antennae filiform, antennomere 3 with sparse pelage of short setae; mentum tooth with side acute, apex broadly rounded. Pronotum moderately transverse, MPW / PL = 1.19 - 1.29, moderately constricted basally, MPW / BPW = 1.38 - 1.46; hind angle variably obtuse, to right, to slightly acute, apex tightly rounded, lateral margin subparallel anterad angle; median base with dense elongate punctures and longitudinal wrinkles; basal margin nearly straight, slightly convex medially; median longitudinal impression broad, shallow, crossed by wrinkles; anterior transverse impression deeply incised, complete, crossed by longitudinal wrinkles that extend across broadly, slightly convex anterior callosity; front angles slightly projected, rounded; pronotal apical and basal widths subequal, APW / BPW = 0.95 - 1.06; lateral marginal depression moderate, edge broadly upturned; laterobasal depression broad, smooth, with median tubercle. Proepisternum with 6 small, elongate punctures along hind marginal groove; prosternal process broad, medially depressed with lateral marginal bead. Elytra subquadrate, disc slightly convex; basal groove evenly recurved to join lateral marginal depression at rounded humerus; MEW / HuW = 1.79 - 2.02; parascutellar seta present (on 1 specimen near base of stria 2); parascutellar striole deep, continuous; sutural interval moderately convex, convexity similar to that of lateral intervals of similar breadth; sutural and 2 nd striae of subequal depth from base to apex; discal striae 2 - 4 fused in association with dorsal elytral setae, striae 5 and 6 may be fused behind humerus, intervals moderately convex to convex; 7 th and 8 th intervals of similar convexity mesad subapical sinuation; 2 dorsal elytral setae at 0.29 x and 0.60 x elytral length, setal impressions foveate, placed within depressed areas associated with strial fusions; apical and subapical setae present; lateral elytral setae arranged as anterior series of 7 setae (or anterior series of 6 with isolated 7 th intermediate seta) and a posterior series of 6 setae; elytral marginal depression narrow, lateral margin upturned; subapical sinuation broad and very shallow. Mesepisternum with ~ 22 distinct punctures 3 - 4 rows; metepisternal width to length ratio = 0.76; metepisternum / metepimeron suture distinct; metathoracic flight wing configuration an ovoid flap, length 2.1 x breadth, remnant R and M veins present, the flap extended 1 / 3 length beyond posterior margin of metanotum. Abdomen with indistinct lateral wrinkles on ventrites 1 - 3; suture between ventrites 2 and 3 complete; apical male ventrite with 2 marginal setae, apical female ventrite with 4 equally spaced marginal setae and median trapezoid of 4 short setae, the basal pair slightly longer. Legs-metatarsomere 1 / metatibial length ratio = 0.22; metatarsomere 4 length along outer lobe 1.4 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci broad, deep, median area irregular. Microsculpture of elytral disc an elongate transverse mesh, apex with upraised transverse mesh; metasternum with obsolete transverse mesh; laterobasal abdominal ventrites with elongate transverse mesh and glossy areas. Coloration of vertex brunneous; antennomere 1 flavous, antennomeres 2 - 3 flavous with a piceous cast, 4 - 11 piceous; pronotal disc brunneous with piceous cast, margins broadly rufoflavous; proepipleuron flavous with rufous cast, proepisternum rufopiceous; elytral disc rufobrunneous with piceous cast, sutural interval concolorous, margins concolorous to slightly darker; elytral epipleuron flavous with rufous cast, metepisternum piceous; abdominal ventrites 1 - 6 piceous medially, flavous laterally, the apical ventrite flavous in apical half; metafemur flavous; metatibia flavous with brunneous cast. Male genitalia (n = 2). Aedeagal median lobe slender, distance from parameral articulation to tip 6.6 x depth at midlength (Fig. 45 M), shaft slightly curved basally, ventral margin straight apically, apex with acute dorsal projection, tip tightly rounded at juncture of flat apical face and ventral margin; in ventral view median lobe slightly curved rightward toward apex, right margin concave, left margin more incurved to apex, apical denticle visible to left of rounded tip (Fig. 45 N); internal sac without ornamentation. Female reproductive tract (n = 1). Bursa copulatrix elongate with broad base at vagina, length 0.60 mm, apical width 0.23 mm, basal width 0.34 mm (Fig. 46 E); bursal walls translucent, thinly wrinkled; gonocoxite 1 with 4 apical fringe setae and 4 - 6 smaller setae on medial surface (Fig. 47 E); gonocoxite 2 subtriangular, apex acuminate, base evenly extended from lateral margin, 2 parallel-sided lateral ensiform setae, apical nematiform setae on medioventral surface at 0.79 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
0FA27A045FE37B5D49375FA24B1EE57B.taxon	etymology	Etymology. The foveae surrounding dorsal elytral setae of this species are the basis for use of the Latin adjectival foveatus as this species' epithet.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
A7BF1BB6412B56B8CDE51A92415A1D6C.taxon	diagnosis	Diagnosis. This species (Fig. 50 D) can be diagnosed from its cryptic sibling species, Mecyclothorax foveolatus (Fig. 50 B), using the criteria presented under that species (above). These two are the only Haleakala species exhibiting: 1, strial fusion implicating striae 2 - 4 at the positions of the dorsal elytral setae; and 2, body sizes above 5 mm. Setal formula 2 2 2 2. Standardized body length 5.0 - 5.4 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
0AFAA04A04BC94A1CD9B7852EE5B0EEA.taxon	diagnosis	Diagnosis. This species is uniquely diagnosed by the presence of 1 - 3 setae in interval 5 accompanying the usual two dorsal elytral setae in interval 2, and broad subquadrate elytra with intervals of consistent breadth that bear regular, transverse-mesh microsculpture. When only one supplementary 5 th interval seta is present, it is situated behind the position of the posterior dorsal elytral seta of the 3 rd interval (Fig. 52 A). When two extra 5 th interval setae are present, they include the posterior trailing seta, plus a 2 nd seta situated between the positions of the anterior and posterior dorsal elytral setae. A third 5 th interval seta may be present, situated at a position immediately posterad or anterad the anterior dorsal elytral seta of interval 3. Setal formula 2 2 2 2. Standardized body length 4.5 - 5.5 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
C28C1C647F113DD63438CE6D12740587.taxon	diagnosis	Diagnosis. Individuals of this species are instantly recognizable by the lustrous, shimmery dorsal surface resulting from the well-developed isodiametric and transverse microsculpture, and the presence of elytral setae on intervals 3, 5, and 7 (Fig. 52 B). The dorsal surface appears similar in reflective pattern to that of the fire-adapted Sericoda species (Liebherr 1991). In Mecyclothorax inaequalis, there are 4 - 6 setae on elytral interval 3, 3 - 5 setae on interval 5, and 3 on interval 7, each seta associated with partial convergence of the adjacent elytral striae. The dorsal microsculpture is arrayed as: 1, vertex with upraised isodiametric mesh, the sculpticells in transverse rows on the neck; 2, pronotal disc with upraised, slightly transversely stretched sculpticells in transverse rows, the median base with granulate isodiametric sculpticells, some in rows; 3, elytral disc with irregular, upraised isodiametric sculpticells plus a transverse mesh, sculpticell breadth 2 x length, the apex with a shiny transverse mesh, sculpticell breadth 2 - 3 x length. Setal formula 2 2 4 2. Standardized body length 3.9 - 4.7 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
6F4881621516CB39BCA1DB2ED72398E9.taxon	diagnosis	Diagnosis. It is the narrow, parallel-sided body shape (Fig. 52 C) that diagnoses this species. The quadrate, elongate elytra exhibit only slightly convex lateral margins and relatively broad humeri, MEW / HuW = 1.83 - 1.93, and the pronotum is very broad relative to the elytra, MEW / MPW = 1.33 - 1.36. In this group, only Mecyclothorax consobrinus (Fig. 57 B) and Mecyclothorax sobrinus (Fig. 57 C) exhibit such quadrate elytra, but their body sizes are much larger; standardized body length for these two species spans 5.5 - 6.6 mm versus a standardized body length of 5.1 - 5.3 for Mecyclothorax longulus. Setal formula 2 2 2 2.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
2C761CCB2203FD18C61C47BEEE420E94.taxon	diagnosis	Diagnosis. Of the beetles comprising this taxonomic group, only individuals of this species simultaneously exhibit: 1, two dorsal elytral setae on interval 3 but no additional setae on intervals 5 or 7; 2, laterally convex elytra with the greatest width behind midlength in combination with slightly narrowed humeri, MEW / HuW = 1.98; and 3, moderately smaller body size, standardized body length 4.5 - 4.9 mm. The dorsal body surface is reflective due to largely transverse microsculpture: 1, vertex and pronotal disc covered with a distinct transverse mesh, sculpticell breadth subequal to twice sculpticell length; and 2, elytra with a transverse mesh, sculpticell breadth 2 - 3 x length. Only the pronotal median base exhibits the upraised isodiametric sculpticells-irregularly swirling based on the orientation of the cuticular surface-characteristic of other species in the group. Setal formula 2 2 2 2.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
2AE947CE9547457E252F5658F8C9F172.taxon	diagnosis	Diagnosis. This along with Mecyclothorax sobrinus and Mecyclothorax consobrinus represent the large-bodied species of the group; in this species standardized body length 5.6 mm. This species differs from the other two by the basally more constricted elytra (Fig. 57), and more punctate discal elytral striae, the striae themselves deeper and more regular. The pronotal median base is more discretely punctate, with ~ 6 distinct, rounded punctures plus 3 - 4 longitudinal wrinkles each side. The male aedeagal median lobe (Fig. 53 H) lacks the robust, accessory subapical projection shared by males of Mecyclothorax consobrinus and Mecyclothorax sobrinus (Fig. 58). Setal formula 2 2 2 2.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
2AE947CE9547457E252F5658F8C9F172.taxon	description	Description (n = 1). Head capsule with frontal grooves deep near clypeus, angled laterally at midpoint to terminate mesad anterior supraorbital setae; dorsal impression of neck slightly concave; eyes moderate in size and convexity, ocular ratio = 1.44, ocular lobe ratio = 0.76; labral anterior margin moderately emarginate; antennae filiform, antennomeres 2 - 3 with sparse pelage of short setae; mentum tooth with sides right, apex broadly rounded. Pronotum transverse, MPW / PL = 1.33, broad basally, MPW / BPW = 1.36, hind angles obtuse, little projected, with lateral margins subparallel for short distance anterad angles; basal margin straight medially, extended slightly between laterobasal depressions; median longitudinal impression shallow, finely incised at depth, extended onto median base; anterior transverse impression deep, finely incised at depth, anterior callosity elevated, flat, both crossed by longitudinal wrinkles; front angles projected, rounded; pronotal base broader than apex, APW / BPW = 0.90; lateral marginal depression broad at front angle, moderately narrow at midlength, broadened near laterobasal depression, edge upturned; laterobasal depression broad, smooth, with median tubercle. Proepisternum with 5 minute punctures along hind marginal groove; prosternal process medially depressed, margin upraised with bead only anterad procoxae. Elytra subovoid, disc flat, sides abruptly sloped; basal groove slightly recurved to subangulate humeral angle; humeri narrowed, MEW / HuW = 1.90, lateral margin narrowly curved posterad outside angle; parascutellar seta present; parascutellar striole smooth anteriorly, 3 punctures posteriorly near apex; sutural interval of same convexity as lateral intervals basally, upraised as a callous apically; sutural and 2 nd striae of subequal depth from base to apex; 8 th interval convex laterad 7 th stria near subapical sinuation, 7 th stria and nearby portion of 7 th interval depressed just apicad diminution of interval 6 at fusion of intervals 5 + 7; 2 dorsal elytral setae at 0.34 x and 0.64 - 0.67 x elytral length, setae in foveate depressions that span interval 3 plus lateral half of interval 4; apical and subapical setae present; lateral elytral setae arranged in anterior series of 7 setae, posterior series of 6 setae; elytral marginal depression moderately narrow throughout length, beadlike near subapical sinuation. Mesepisternum with ~ 18 punctures in 3 rows; metepisternal width to length ratio 0.86. Abdomen with irregular lateral wrinkles on ventrites 1 - 5; suture between ventrites 2 and 3 complete; apical male ventrite with 2 marginal setae. Legs-metatarsomere 1 / metatibial length ratio = 0.18; metatarsomere 4 length along outer lobe 1.3 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci moderately deep, basal tarsomeres medially convex. Microsculpture of vertex upraised isodiametric sculpticells in transverse rows; pronotal disc with isodiametric sculpticells in transverse rows to a transverse mesh, sculpticell breadth 2 x length, median base with granulate isodiametric mesh; elytral disc with upraised isodiametric sculpticells, the apex with upraised transverse mesh, sculpticell breadth 2 x length; metasternum with distinct transverse mesh; laterobasal abdominal ventrites with swirling transverse and isodiametric microsculpture. Coloration of vertex dark rufobrunneous; antennomere 1 flavous, antennomeres 2 - 3 rufoflavous, 4 - 11 rufobrunneous; pronotal disc dark rufobrunneous, margins rufoflavous mesad front angles, lateral marginal depressions and median base rufobrunneous; proepipleuron rufoflavous, proepisternum rufopiceous; elytral disc rufobrunneous, sutural interval concolorous basally, slightly paler, rufoflavous apically; elytral marginal depression and apex slightly paler than disc, rufoflavous; elytral epipleuron flavous dorsally, rufobrunneous on ventral margin, metepisternum piceous; abdomen piceous, apical 1 / 5 of apical ventrite 6 flavous; metafemur with ground color flavous, basal half with broad piceous cloud; metatibia rufoflavous with piceous cast. Male genitalia (n = 1). Aedeagal median lobe (Fig. 53 H) much like a more robust version of Mecyclothorax longulus (Fig. 53 E), gracile, distance from parameral articulation to tip 4.1 x depth at midlength, apex extended 3 x depth beyond apex of ostial opening, expanded dorsoventrally at tip, with apical face of tip obliquely flattened; internal sac unornamented, flagellar plate short, length 0.30 x parameral articulation-tip distance.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
2AE947CE9547457E252F5658F8C9F172.taxon	etymology	Etymology. The Latin adjectival foveopunctatus signifies the foveate depressions surrounding the dorsal elytral setae.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
92E8A563B6A75C6245C1EDE9D8C1A7DD.taxon	diagnosis	Diagnosis. This and Mecyclothorax sobrinus share the conditions of subquadrate elytra with dorsal setae restricted to interval 3 (Fig. 57 B-C), and large body size; in this species standardized body length 5.8 - 6.3 mm. The two species are considered adelphotaxa based on shared possession of a very distinctive male aedeagal median lobe that exhibits a robust, apically directed process situated immediately basad the ostial opening (Fig. 58). Mecyclothorax consobrinus can be diagnosed by the more developed dorsal microsculpture: 1, vertex with upraised isodiametric sculpticells in transverse rows; 2, pronotal disc with upraised slightly transversely stretched sculpticells, pronotal median base with granulate isodiametric and transverse sculpticells; 3, elytral disc with upraised isodiametric mesh, the apex with isodiametric mesh and swirling transverse mesh in large depression associated with 7 th stria. The elytra are also more irregularly depressed in this species, in some individuals due to presence of a third dorsal elytral seta in interval 3 (Fig. 57 B), and in all individuals due to the deep and irregular depression associated with the fused apical portion of striae 5 + 7, this depression in some instances also involving the apical termination of interval 6. Setal formula 2 2 2 - 3 2.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
EEE491352AC3700BAC4BA42A897B7F4A.taxon	diagnosis	Diagnosis. Individuals of this species exhibit overlapping head, prothoracic, and elytral ratios and standardized body length- 5.5 - 6.6 mm-with Mecyclothorax consobrinus, reducing any diagnosis of the two species to qualitative characters associated with elytral setation and punctation, and cuticular microsculpture. The discal elytral striae are deeper and of more uniform depth in this species than in Mecyclothorax consobrinus. Striae 1 - 6 are closely punctured in the basal 2 / 3 of their length. The setal impressions associated with the two dorsal elytral setae are shallower and less broad than seen in Mecyclothorax consobrinus, depressing interval 3 plus less than half of interval 4. Also, the apical depression associated with interval 7 is shallower and more regularly depressed among individuals of this species. Finally, the microsculpture is less upraised overall: 1, vertex with well-developed isodiametric sculpticells in transverse rows; 2, pronotal disc with well-developed slightly transversely stretched mesh, the median base covered with a mixture of isodiametric and transverse sculpticells; 3, elytral disc with a transverse mesh, sculpticell breadth 2 x length, apex with mesh more transverse, breadth 2 - 3 x length. Also, no individuals representing this species have been observed to have more than two dorsal elytral setae. Setal formula 2 2 2 2.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
E37029CE0AA5AED3F2B6D07FF1B4850B.taxon	diagnosis	Diagnosis. This is one of three species in the group that is characterized by bicolored elytra; the lateral elytral intervals flavous, contrasted to the piceous disc (Fig. 60 A-C). In this species intervals 7 - 9 and the apex of the sutural stria are rufobrunneous to flavous, whereas the basal portions of intervals 2 - 5 are rufopiceous. The dorsal surface of the head and the pronotum are rufous with a flavous cast. The pronotum is moderately transverse, MPW / PL = 1.26, and basally constricted, MPW / BPW = 1.58. The dorsal surface of the head bears well-developed isodiametric sculpticells arranged in transverse rows. The setal formula is 2 1 2 0; the other two bicolored species- Mecyclothorax patulus and Mecyclothorax patagiatus - are characterized by presence of the subapical seta. Moreover, this species lacks the parascutellar seta, whereas it is present in the other two species. Standardized body length 3.45 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
E37029CE0AA5AED3F2B6D07FF1B4850B.taxon	description	Description (n = 1). Head capsule with frontal grooves broad near clypeus, convexity present laterad groove, a narrow carina extended to supraorbital seta; dorsal surface of neck flat; eyes large, moderately convex, ocular ratio = 1.46, ocular lobe ratio = 0.84; labral anterior margin very shallowly emarginate medially; antennae filiform, antennomeres 2 - 3 glabrous except for 1 or 2 small setae on shafts; mentum tooth with sides acute, apex tightly rounded. Pronotum with glabrous hind angles, the base constricted, and lateral margins subparallel anterad the right, projected hind angles; median base moderately depressed, strigose due to long punctures and wrinkles; basal margin straight medially, margin expanded posterad laterobasal depressions; median longitudinal impression shallow, finely incised; anterior transverse impression shallow, broad, crossed by indistinct wrinkles; anterior callosity elevated, flat, crossed by indistinct longitudinal wrinkles; front angles projected, tightly rounded; pronotal apex broader than base, APW / BPW = 1.06; lateral marginal depression narrow, edge slightly upturned, broader at front angle, beaded anterad basal sinuation; laterobasal depression smooth, continuous with lateral depression; slight tubercle mesad lateral margin. Proepisternum with 5 minute punctures along hind marginal groove; prosternal process with narrow median impression, lateral margins broadly beaded between coxae. Elytra subellipsoid, disc flat, sides moderately sloped; basal groove slightly recurved to tightly rounded humeral angle; greatest width near midlength, MEW / HuW = 2.09; parascutellar striole discontinuous along length, with 3 - 4 punctures; sutural interval coplanar with lateral striae basally, upraised in apical half; sutural and 2 nd striae of subequal depth from base to apex; discal striae 1 - 5 broad but defined, lined with minute elongate punctulae in basal 1 / 3 of length, 6 - 7 discontinuous, represented by serial punctures, discal intervals moderately convex; 8 th interval slightly more convex than fused apical portion of fused striae 5 + 7; 2 dorsal elytral setae at 0.28 x and 0.64 x elytral length, setal impression small, spanning 1 / 3 - 1 / 2 of interval 3; apical and subapical setae absent; lateral elytral setae arranged in an anterior series of 6 setae and a posterior series of 5 setae; elytral marginal depression narrow, margin little upturned in basal half, narrowly beadlike near subapical sinuation; subapical sinuation very shallow, nearly obsolete. Mesepisternum with ~ 8 punctures in 1 - 2 rows; metepisternal width to length ratio = 0.71; metepisternum / metepimeron suture distinct. Abdomen with irregular lateral wrinkles on ventrites 1 - 5; suture between ventrites 2 and 3 complete; apical female ventrite with 4 equally spaced marginal setae plus median trapezoid of 4 subequal, short setae. Legs-metatarsomere 1 / metatibial length ratio = 0.20; metatarsomere 4 length along outer lobe 1.33 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci narrow, shallow, median surface broad. Microsculpture of pronotal disc transverse mesh, sculpticell breadth 2 x length, median base with mixture of isodiametric and transverse sculpticells; elytral disc with transverse mesh, sculpticell breadth 2 - 4 x length, apex with shallow transverse mesh of the same dimensions; metasternum with shallow transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of antennomere 1 flavous, antennomeres 2 - 3 rufoflavous, 4 - 11 rufobrunneous; proepipleuron and proepisternum rufoflavous; elytral epipleuron rufoflavous, metepisternum rufobrunneous; abdomen with ventrites 1 - 3 medially pale rufobrunneous, lateral portions and ventrites 4 - 6 rufoflavous; femur rufoflavous; tibia rufoflavous with rufous cast. Female reproductive tract. The lone female holotype was not dissected.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
E37029CE0AA5AED3F2B6D07FF1B4850B.taxon	etymology	Etymology. Britton's choice of the Latin adjective subtilis to signify the minute, slender body of this beetle was extremely appropriate, and that choice is hereby validated.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
84ED6BB059E71BCFA9897014E9D2B0F2.taxon	diagnosis	Diagnosis. This second of the three bicolored species (Fig. 60 B) exhibits more narrowly flavous elytral margins-only intervals 8 - 9 - versus piceous discal intervals 2 - 7. The sutural interval is rufous basally, flavous apically. The bisetose pronotum is more transverse than that of Mecyclothorax subtilis; MPW / PL = 1.28, and less constricted basally, MPW / BPW = 1.53. As in the next species below, Mecyclothorax patagiatus, the parascutellar and subapical elytral setae are present, but the elytral humeri are narrower in this species-MEW / HuW = 2.06 versus 2.0 - and the elytra more ellipsoid. The setal impressions of the dorsal elytral setae are larger in beetles of this species, as they span interval 3. Setal formula 2 1 2 1 [sae]. Standardized body length 3.4 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
84ED6BB059E71BCFA9897014E9D2B0F2.taxon	description	Description (n = 1). Head capsule with frontal grooves broad near clypeus, a lateral carina extended to mesad anterior supraorbital seta; dorsal surface of neck flat; eyes moderately convex, ocular ratio = 1.47, not extended onto posterior portion of ocular lobe, ocular lobe ratio = 0.75; labral anterior margin broadly emarginate to 1 / 6 of length; antennae filiform, sparse setae on apex of antennomere 1 and shafts of antennomeres 2 - 3; mentum tooth with sides acute, apex tightly rounded. Pronotum transverse, MPW / PL = 1.28, bisetose, glabrous hind angles obtuse, rounded behind, lateral margin subparallel for short distance anterad angle; base moderately broad, MPW / BPW = 1.53; median base nearly coplanar with disc, ~ 10 sparsely distributed, isolated punctures each side; basal margin convexly expanded between laterobasal depressions; median longitudinal impression shallow, finely incised, crossed by fine longitudinal wrinkles; anterior transverse impression deep, finely incised, minute irregularities in deepest part; anterior callosity convex, glossy surface with minute longitudinal wrinkles; front angles slightly projected, tightly rounded; pronotal apical width greater than basal width, APW / BPW = 1.10; lateral marginal depression narrow, edge upturned, slightly broader at front angle; laterobasal depression with slightly irregular surface, continuous with lateral depression. Proepisternum with 5 minute punctures along hind marginal groove; prosternal process with narrow median impression, lateral margins broadly beaded between coxae. Elytra broadly subellipsoid, disc flat, sides moderately sloped; basal groove evenly and distinctly recurved to tightly rounded humeral angle, MEW / HuW = 2.06; parascutellar seta present; parascutellar striole with 3 - 4 punctures, very shallow between punctures; sutural interval more convex than lateral intervals, sutural juncture upraised; sutural and 2 nd striae of subequal depth from base to disc, 2 nd stria slightly shallower at apex; discal striae 1 - 5 moderately broad, evident, stria 6 shallower and stria 7 shallower still, interrupted along length; striae 1 - 4 with minute elongate punctures, stria 5 with shallower punctures and stria 6 with only irregularities along length; intervals 2 - 5 moderately convex, lateral intervals less so; 8 th interval slightly more convex than fused apical portion of striae 5 + 7; 2 dorsal elytral setae at 0.28 - 0.32 x and 0.62 x elytral length, setal impressions evident, spanning interval 3; apical elytral seta absent, subapical elytral seta present; lateral elytral setae arranged as anterior series of 6 setae and posterior series of 5 setae; elytral marginal depression narrow, edge little upturned in basal half, margin beadlike near subapical sinuation; subapical sinuation very shallow, nearly obsolete. Mesepisternum with ~ 6 punctures in 1 - 2 rows; metepisternal width to length ratio = 0.80; metepisternum / metepimeron suture distinct. Abdomen with irregular lateral wrinkles on ventrites 1 - 5, lateral depressions on ventrites 3 - 6; suture between ventrites 2 and 3 complete; apical female abdominal ventrite with 4 equally spaced setae plus trapezoid of 4 subequal, short setae. Legs-metatarsomere 1 / metatibial length ratio = 0.20; metatarsomere 4 length along outer lobe 1.3 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci narrow, shallow, median surface broad. Microsculpture of vertex shallow isodiametric sculpticells in rows; pronotal disc with shallow transverse mesh, sculpticell breadth 3 x length, to transverse lines not connected into mesh; pronotal median base glossy, obsolete transverse mesh between punctures; elytral disc and apex with shallow transverse mesh, sculpticell breadth 2 - 4 x length; metasternum with transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex rufobrunneous; antennomere 1 flavous, antennomeres 2 - 3 rufoflavous, 4 - 11 rufobrunneous; pronotal disc rufobrunneous with piceous cast, lateral margins, apex, and base rufoflavous; proepipleuron flavous, proepisternum rufobrunneous with piceous cast; elytral epipleuron dorsally flavous, ventrally rufoflavous, metepisternum rufobrunneous with piceous cast; abdomen with ventrites 1 - 2 medially, and 3 - 5 mediobasally rufopiceous, ventrites 3 - 6 apical and marginally rufoflavous; metafemur flavous; metatibia flavous with brunneous cast. Female reproductive tract. The lone female holotype was not dissected.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
84ED6BB059E71BCFA9897014E9D2B0F2.taxon	etymology	Etymology. The Latin adjective patulus means open, spread out, or broad (Brown 1956), signifying the short, broad body of these beetles.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
4F2AE802D11B6929DF8D7C04508B3352.taxon	diagnosis	Diagnosis. This, the third Haleakala Mecyclothorax ovipennis group species to display bicolored elytra (Fig. 60 C), can be diagnosed by the following combination: 1, discal elytral striae 1 - 6 rufobrunneous with piceous cast, intervals 7 - 9 contrastedly rufoflavous; 2, pronotal disc also rufobrunneous with piceous cast, pronotal margins rufoflavous; 3, parascutellar seta present; 4, subapical elytral seta present, apical seta absent; 5, pronotal lateral margins slightly divergent from hind obtuse hind angles; 6, elytral humeri broadly rounded, the elytra broadly subquadrate, MEW / HuW = 2.0. Criteria 1, 5, and 6 diagnose this species from both Mecyclothorax subtilis and Mecyclothorax patulus, whereas criteria 2, 3, and 4 diagnose this species from Mecyclothorax subtilis. The pronotum of the unique holotype has the basal pronotal seta present on the right side, with the left hind angle glabrous (specimen examined at 125 x), obviating use of this character in the diagnosis. Setal formula 2 1 - 2 2 1 [sae]. Standardized body length 3.8 mm (slightly larger than both preceding species).	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
4F2AE802D11B6929DF8D7C04508B3352.taxon	description	Description (n = 1). Head capsule with frontal grooves broad near clypeus, lateral carina to anterior supraorbital seta; dorsal surface of neck flat; eyes moderately convex, ocular ratio = 1.43, ocular lobe ratio 0.78; labral anterior margin broadly emarginate to 1 / 6 of length; antennae filiform, antennomeres 2 - 3 with sparse pelage of short setae; mentum tooth with sides acute, apex tightly rounded. Pronotum transverse, MPW / PL = 1.30, base moderately constricted, MPW / BPW = 1.50; hind angle obtuse, margin behind rounded; median base moderately depressed, ~ 20 punctures or strigose wrinkles each side; basal margin convexly expanded between laterobasal depressions; median longitudinal impression evident, finely incised, joined by fine transverse wrinkles; anterior transverse impression deep, finely incised, minute irregularities in deepest part; anterior callosity elevated, flat, crossed by indistinct longitudinal wrinkles; front angles projected, broadly rounded; apical and basal pronotal widths subequal, APW / BPW = 1.03; lateral marginal depression moderate, edge upturned, broader at front angle; laterobasal depression narrowly concave, continuous with lateral depression. Proepisternum with 5 minute punctures along hind marginal groove; prosternal process narrowly impressed medially, lateral margins broadly beaded between coxae. Elytra with disc flat, sides moderately sloped; basal groove slightly recurved to broadly rounded humerus; parascutellar striole with 3 - 4 punctures, shallow between punctures; sutural interval coplanar with lateral intervals basally, upraised in apical half; sutural and 2 nd striae of subequal depth and breadth from base to apex; discal striae 1 - 5 broad, deep, stria 6 shallower and more irregular, stria 7 shallower still; sutural stria deep, finely punctate basally, deep, narrow, and smooth apically; striae 2 - 4 with minute punctures on disc, punctures shallower in stria 5, linear irregularities in stria 6; discal intervals 2 - 4 convex, lateral intervals less so; 8 th interval slightly more convex than fused apical portion of striae 5 + 7; 2 dorsal elytral setae at 0.27 x and 0.54 x elytral length, setal impressions small, spanning 1 / 2 width of interval 3; lateral elytral setae arranged in anterior series of 6 setae, posterior series of 5 setae; elytral marginal depression narrow, margin little upturned in basal half, beadlike near subapical sinuation; subapical sinuation very shallow, nearly obsolete. Mesepisternum with ~ 6 punctures in 1 - 2 rows; metepisternal width to length ratio = 0.80; metepisternum / metepimeron suture distinct. Abdomen with irregular lateral wrinkles in ventrites 1 - 5, lateral depressions in ventrites 3 - 6; suture between ventrites 2 and 3 complete; apical male ventrite with 2 marginal setae. Legs-metatarsomere 1 / metatibial length ratio = 0.19; metatarsomere 4 length along outer lobe 1.3 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci narrow, shallow, median surface broad. Microsculpture of vertex with evident isodiametric sculpticells in transverse rows; pronotal disc with shallow transverse mesh, sculpticell breadth 3 x length, to transverse lines; pronotal median base with shallow transverse mesh laterally, median area glossy; elytral disc with evident transverse mesh, sculpticell breadth 2 - 4 x length, apex with more distinct transverse mesh of same dimensions; metasternum with transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex rufobrunneous; antennomere 1 flavous, antennomeres 2 - 3 rufoflavous; 4 - 11 rufobrunneous; pronotal disc rufobrunneous with piceous cast, margins rufoflavous; proepipleuron rufoflavous, proepisternum rufobrunneous with piceous cast; elytral sutural interval concolorous with disc basally, rufoflavous apically; elytral epipleuron dorsally flavous, ventrally rufoflavous; metepisternum rufobrunneous; abdomen with ventrites 1 - 2 medially, 3 - 5 mediobasally rufopiceous, 3 - 6 apically and marginally rufoflavous, apical ventrite 6 with apical 3 / 4 paler, flavous; metafemur flavous; metatibia flavous with brunneous cast. Male genitalia (n = 1). Aedeagal median lobe robust, distance between parameral articulation and tip 3.2 x depth at midlength (Fig. 61 A); extension of apex beyond ostial opening parallel sided at base and evenly narrowed dorsoventrally to tightly rounded tip; median lobe curved sinuously leftward toward apex in ventral view, the right margin distinctly concave, left margin convex (Fig. 61 B); internal sac without ornamentation, flagellar plate elongate, length of sclerotized plate 0.54 x parameral articulation-tip distance.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
4F2AE802D11B6929DF8D7C04508B3352.taxon	etymology	Etymology. The Latin stem for this species epithet is patagium; i. e. a gold edging or border (Brown 1956). The adjectival form patagiatus signifies the pale elytral border of beetles of this species.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
0C4ABC92293F23AEADBDB7448130EB2E.taxon	diagnosis	Diagnosis. This species (Fig. 60 D) plus Mecyclothorax ovipennis (Fig. 65 A) and Mecyclothorax flaviventris (Fig. 68 C) comprise the three species in this group with the largest, most convex eyes; ocular ratio = 1.55 - 1.57 in this species. Of these, both Mecyclothorax ovipennis and this species are characterized by impunctate discal striae 1 - 4. This species (Fig. 60 D) can be told from Mecyclothorax ovipennis (Fig. 65 A) by the narrower body with more basally constricted pronotum, MPW / BPW = 1.56 - 1.67, and more narrowly ellipsoid elytra. Moreover, this is the only one of the three with glabrous hind pronotal angles, and without any apical elytral setae; setal formula 2 1 2 0. Standardized body length 4.3 - 4.7 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
0C4ABC92293F23AEADBDB7448130EB2E.taxon	description	Description (n = 2). Head capsule with frontal grooves broad near clypeus, lateral carina to anterior supraorbital seta; dorsal surface of neck flat; ocular lobe distinctly protruded from gena, eyes large, ocular lobe ratio = 0.79 - 0.80; labral anterior margin very shallowly emarginate medially; antennae filiform, antennomeres 2 - 3 with sparse pelage of short setae; mentum tooth with sides acute, apex broadly rounded. Pronotum appearing elongate, MPW / PL = 1.11 - 1.18; hind angle right, lateral margin straight, subparallel to slightly convergent anterad hind angle; median base only slightly depressed, ~ 15 densely distributed punctures each side extended to laterobasal depression; basal margin nearly straight across base; median longitudinal impression very shallow, finely incised; anterior transverse impression very shallow, broad, crossed by longitudinal wrinkles; anterior callosity nearly flat, crossed by indistinct wrinkles; front angles not projected, tightly rounded; pronotal apical width greater than basal width, APW / BPW = 1.09 - 1.16; lateral marginal depression very narrow throughout length, edge tightly upturned; laterobasal depression depressed, punctate surface continuous with median base. Proepisternum with 5 minute punctures along hind marginal groove; prosternal process with narrow median impression, lateral margins broadly beaded between coxae. Elytra with disc flat, sides steeply sloped to depressed lateral margins and apex; basal groove briefly, distinctly recurved to angulate humerus; parascutellar seta present; parascutellar striole with 4 - 5 punctures, shallow, continuous between punctures; sutural interval more convex than intervals 2 - 4, but sutural juncture still depressed; sutural stria shallow, with minute punctulae basally, slightly deeper and more well defined than 2 nd stria on disc, the two of subequal depth apically; discal striae 2 - 4 shallow, smooth, striae 5 - 6 obsolete but traceable, stria 7 absent; discal intervals 2 - 4 only slightly convex to nearly flat, lateral intervals flat; 8 th interval of similar convexity to fused apical portion of striae 5 + 7; 2 dorsal elytral setae at 0.30 - 0.31 x and 0.61 - 0.66 x elytral length, setal impressions very small, spanning 1 / 3 of interval 3; lateral elytral setae arranged in anterior series of 7 setae, and posterior series of 6 setae; elytral marginal depression narrow, margin upturned, beaded only at subapical sinuation; subapical sinuation shallow, more abruptly incurved anteriorly. Mesepisternum with ~ 7 punctures in 2 rows; metepisternal width to length ratio = 0.72; metepisternum / metepimeron suture distinct. Abdomen with irregular lateral wrinkles on ventrites 1 - 3, lateral depressions on ventrites 3 - 6; suture between ventrites 2 and 3 reduced laterally, effaced; apical female ventrite with 4 equally spaced setae plus median trapezoid of 4 setae, the basal pair longer. Legs-metatarsomere 1 / metatibial length ratio = 0.18; metatarsomere 4 length along outer lobe 1.25 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci narrow, shallow, median area broad. Microsculpture of vertex and pronotal disc a transverse mesh, sculpticell breadth 2 - 3 x length; pronotal median base with transverse mesh, sculpticell breadth 2 x length, between punctures; elytral disc and apex with isodiametric to transverse sculpticells, sculpticell breadth 2 x length, in transverse rows; metasternum with distinct transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex rufobrunneous; antennomere 1 - 3 flavous, 4 - 11 darker, more brunneous; pronotal disc rufobrunneous with piceous cast, the lateral margins moderately, and base and apex broadly, rufoflavous; proepipleuron rufoflavous, proepisternum rufobrunneous with piceous cast; elytral disc dark, rufobrunneous to rufopiceous; sutural interval rufobrunneous basally, flavous in apical 1 / 3; elytral intervals 7 - 9 slightly paler than disc, rufoflavous basally, flavous apically; elytral epipleuron flavous, metepisternum rufobrunneous with piceous cast; abdomen with ventrite 1 (plus metepimeron) rufobrunneous, ventrites 2 - 3 and middle of ventrites 4 - 5 rufopiceous; abdominal apical ventrite with apical 2 / 3 flavous; metafemur flavous; metatibia flavous with brunneous cast. Female reproductive tract (n = 1). Bursa copulatrix columnar with apical expansion, length 1.0 mm, apical expansion breadth 0.46 mm, basal breadth 0.34 mm (Fig. 62 A); bursal shaft translucent, thinly wrinkled, apex more transparent, less wrinkled; gonocoxite 1 with 3 apical fringe setae, the middle seta more robust, a larger seta at apicomedial angle and 12 - 13 smaller setae basally on medial surface (Fig. 63 A); gonocoxite 2 falcate with subacuminate apex, base extended laterally as sinuous panhandle, 2 lateral ensiform setae, apical nematiform setae on medial surface at 0.75 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
0C4ABC92293F23AEADBDB7448130EB2E.taxon	etymology	Etymology. The Latin adjectival strigosus, meaning thin, signifies the narrow body shape characteristic of this species.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
7DCB2BB24DE924632869E6EF962B8732.taxon	diagnosis	Diagnosis. Among Haleakala Mecyclothorax (Fig. 65 A), this is most similar to the preceding, Mecyclothorax strigosus (Fig. 60 D) based on the well-developed eyes, ocular ratio = 1.55 - 1.61 and ocular lobe ratio = 0.84 - 0.89, plus basally constricted pronotum and ellipsoid elytra. They can be separated by the setal conformation, with this species characterized by a quadrisetose pronotum, and presence of the apical elytral seta; setal formula 2 2 2 1 [ae]. This species is characterized by the same setal formula as Mecyclothorax flaviventris, but individuals of that species exhibit punctate discal striae (Fig. 68 C). Standardized body length 3.7 - 4.9 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
BAD02644D7C728D485E515D8FB6BD3A9.taxon	diagnosis	Diagnosis. Within the Mecyclothorax ovipennis group, this species (Fig. 65 B) forms part of a triplet of species also including Mecyclothorax apicalis (Fig. 65 C) and Mecyclothorax parapicalis (Fig. 65 D) that are collectively diagnosed by glabrous, obtuse pronotal hind angles, shallow impunctate discal elytral striae, and concolorous elytral disc and margins. Of these, Mecyclothorax takumiae is characterized by the smallest body size, standardized body length 3.6 mm, and least developed microsculpture. The pronotal disc and median base are glossy with obsolete transverse-mesh microsculpture and parallel lines over portions of the cuticle, whereas the elytral disc has a shallow mesh of isodiametric and transverse sculpticells, breadth 2 - 3 x length. Both Mecyclothorax apicalis and Mecyclothorax parapicalis have well-developed sculpticells covering those areas. Setal formula 2 1 2 (1) 0; the right elytron has both dorsal elytral setae, whereas the left has only a seta at the anterior position of the right elytron. As the bisetose condition is shared with Mecyclothorax apicalis and Mecyclothorax parapicalis, the unisetose elytron is considered a variant.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
BAD02644D7C728D485E515D8FB6BD3A9.taxon	description	Description (n = 1). Head capsule with frontal grooves broad near clypeus, lateral carina to anterior supraorbital seta; dorsal surface of neck flat; eyes moderately developed, ocular ratio = 1.45, ocular lobe ratio = 0.80; labral anterior margin medially emarginate, medially excavated 1 / 6 of length; antennae filiform, antennomeres 2 - 3 with only 1 or 2 short setae on shafts; mentum tooth with sides acute, apex rounded. Pronotum slightly transverse, MPW / PL = 1.21, moderately constricted basally, MPW / BPW = 1.53; hind angle obtuse, margin behind rounded; median base markedly depressed versus disc, ~ 9 punctures each side, surface glossy between; basal margin convexly expanded between laterobasal depressions; median longitudinal impression shallow, finely incised, crossed by fine transverse wrinkles; anterior transverse impression narrow, shallowly incised laterally, obsolete medially; anterior callosity slightly convex, smooth, glossy; front angles not to slightly projected, tightly rounded; pronotal apical and basal width equal, APW / BPW = 1.0; lateral marginal depression narrow through apical 1 / 3 of length, edge upturned, widened in basal 1 / 3; laterobasal depression narrow, surface irregular, continuous with lateral depression. Proepisternum with 5 minute punctures along hind margin; prosternal process with narrow median impression, lateral margins broadly beaded between coxae. Elytra subquadrate, disc flat, sides distinctly sloped; basal groove evenly recurved to subangulate (left) to tightly rounded (right) humerus, MEW / HuW = 2.04; parascutellar seta present; parascutellar striole with 4 punctures, shallow between punctures; sutural interval slightly more convex than lateral intervals, upraised at sutural juncture; sutural stria shallow, continuous basally, punctate on disc, smooth and deep apically; striae 2 - 6 progressively shallower, smooth, stria 7 obsolete; stria 1 subequal to slightly deeper than stria 2 at elytral apex; elytral interval 2 slightly convex, intervals 3 - 7 progressively flatter; 8 th interval subcarinate laterad fused apical portion of striae 5 + 7; 2 dorsal elytral setae at 0.31 x and 0.63 x elytral length (right elytron), setal impressions moderate, spanning 2 / 3 of interval 3; apical and subapical setae absent; lateral elytral setae arranged in anterior series of 6 setae and posterior series of 6 setae; elytral marginal depression slightly broader at humerus, narrowed laterally, beadlike at subapical sinuation; subapical sinuation shallow, more abruptly incurved anteriorly. Mesepisternum with ~ 8 punctures in 1 - 2 rows; metepisternal width to length ratio = 0.65; metepisternum / metepimeron suture distinct. Abdomen with irregular lateral wrinkles on ventrites 1 - 6; suture between ventrites 2 and 3 complete; apical female ventrite with 4 equally spaced setae plus a median trapezoid of 4 subequal, short setae. Legs-metatarsomere 1 / metatibial length ratio = 0.18; metatarsomere 4 length along outer lobe 1.33 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci narrow, shallow, median area broad. Microsculpture of vertex a shallow transverse mesh, sculpticell breadth 2 - 3 x length; elytral apex with transverse mesh, sculpticell breadth 2 - 3 x length; metasternum with transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex rufobrunneous with piceous cast; antennomeres 1 - 3 flavous, 4 - 11 darker, more brunneous; pronotal disc rufopiceous, lateral margins concolorous, base and apex paler, rufoflavous; proepipleuron rufoflavous, proepisternum rufobrunneous with piceous cast; elytral disc rufopiceous, sutural interval rufous in basal half, flavous in apical half; 9 th elytral interval and lateral marginal depression rufous, apex of intervals 8 and 9 flavous; elytral epipleuron dorsally flavous, ventrally rufoflavous, metepisternum rufopiceous; abdomen with ventrites 1 - 5 and the base of 6 rufopiceous, apical 1 / 3 of apical ventrite 6 flavous; metafemur flavous; metatibia flavous with brunneous cast. Female reproductive tract. The lone female holotype was not dissected.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
BAD02644D7C728D485E515D8FB6BD3A9.taxon	etymology	Etymology. Raina Takumi Kahaloa'a has provided numerous specimens for this revision, most often from difficult to access natural areas, and from months of the year with very few other records. Thus it is a pleasure to honor her contributions to Mecyclothorax diversity by naming this species in her honor.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
FAEC3618C715613BA3114BA5B5C8E8C8.taxon	diagnosis	Diagnosis. This species and Mecyclothorax parapicalis share: 1, evident microsculpture, isodiametric to transverse, on the vertex and pronotal disc; 2, divergent pronotal lateral margins anterad obtuse, glabrous hind angles; 3, shallow, impunctate discal elytral intervals that are of similar color to lateral elytral intervals 7 - 9. The pronotum is more basally constricted in individuals of this species-MPW / BPW = 1.46 - 1.54 - and more transverse-MPW / PL = 1.26 - 1.33 - than in beetles comprising Mecyclothorax parapicalis (Fig. 65 C-D). The elytra are also narrower basally, with the lateral margins little extended laterally behind the tightly rounded humeri (Fig. 65 C). The male aedeagal median lobe is distinctively different from that of Mecyclothorax parapicalis, with the apex sinuously recurved with an expanded, spoonlike tip (Figs 61 F, G). Individuals of this species vary in the degree of punctation in the discal elytral striae. The striae vary from smooth, impunctate, to more irregular due to the presence of elongate punctulae (Fig. 65 C). This variation is taken into account in the dichotomous key above. Setal formula 2 1 2 0. Standardized body length 4.0 - 4.2 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
716C0F2FF5931E7B5E76D44F579C5B21.taxon	diagnosis	Diagnosis. Like Mecyclothorax apicalis, but the body is broader at the elytral base, and the elytral lateral margins are slightly extended laterally behind the humeri, the humeral angle defined by a slight hitch at the base of the moderately broad lateral marginal depression (Fig. 65 D). The microsculpture on the forebody is more transverse, with: 1, a shallow transverse mesh, sculpticell breadth 2 - 3 x length on the vertex; 2, transverse mesh, sculpticell breadth 3 x length, to transverse lines on the pronotal disc; and 3, evident isodiametric and transverse sculpticells over the pronotal median base. The male aedeagal median lobe exhibits a rhomboidal apex, with the apical and ventral margins meeting at an acute angle (Fig. 61 I). Setal formula 2 1 2 0. Standardized body length- 3.8 - 3.9 mm-is slightly smaller than that of Mecyclothorax apicalis; length 4.0 - 4.2 for that species.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
716C0F2FF5931E7B5E76D44F579C5B21.taxon	description	Description (n = 3). Head capsule with frontal grooves broad near clypeus, a broad lateral carina to anterior supraorbital seta; dorsal surface of neck flat to slightly concave; eyes moderately convex but covering much of ocular lobe, ocular ratio = 1.41 - 1.46, ocular lobe ratio = 0.82 - 0.84; labral anterior margin medially emarginate 1 / 6 of length; antennae submoniliform, antennomeres 2 - 3 with sparse pelage of short setae; mentum tooth with sides acute, apex tightly rounded. Pronotum transverse, MPW / PL = 1.28 - 1.36, broad basally, MPW / BPW = 1.41 - 1.45; hind angle slightly obtuse, margin rounded behind or not, lateral margin subparallel or slightly divergent immediately anterad angle; median base slightly depressed relative to disc, smoother medially, laterally punctate, longitudinal wrinkles lining juncture with disc; basal margin broadly, slightly convex between laterobasal depressions; median longitudinal impression very shallow, finely incised; anterior transverse impression deep, finely incised, shallower at midline; anterior callosity slightly convex, smooth, glossy; front angles slightly produced, tightly rounded; pronotal apical width slightly narrower than basal width, APW / BPW = 0.93 - 0.97; lateral marginal depression narrow throughout, edge beaded except where slightly broader at front angles; laterobasal depression broadly convex between median base and hind angle. Proepisternum with 5 minute punctures along hind marginal groove; prosternal process with narrow median impression, margins beaded between coxae. Elytra subquadrate, disc flat, sides distinctly sloped; basal groove distinctly recurved inside humeral angle; elytra broadest behind midlength and humeri extended laterally in concert with broad pronotal base, MEW / HuW = 1.95 - 2.0; parascutellar seta present; parascutellar striole with 5 punctures, shallow but continuous between punctures; sutural interval flat in basal half, progressively elevated along suture to apex; sutural and 2 nd striae of subequal depth from base to apex; discal striae 1 - 4 smooth, shallow, striae 5 - 6 shallower, traceable, stria 7 absent; discal elytral intervals 2 - 6 only slightly convex to nearly flat on lateral intervals; 8 th interval slightly more convex than apical fused portion of striae 5 + 7; 2 dorsal elytral setae at 0.32 x and 0.67 x elytral length, setal impressions small, spanning 1 / 2 width of interval 3; apical and subapical setae absent; lateral elytral setae arranged in anterior series of 7 setae and posterior series of 6 setae; elytral marginal depression of moderate breadth at humerus, narrowed laterad posterior setal series, margin upturned except beaded at subapical sinuation; subapical sinuation shallow, more abruptly incurved anteriorly. Mesepisternum with ~ 6 punctures in 1 - 2 rows; metepisternal width to length ratio = 0.75; metepisternum / metepimeron suture distinct. Abdomen with irregular lateral wrinkles on ventrites 1 - 5, and lateral depressions on ventrites 3 - 6; suture between ventrites 2 and 3 reduced laterally, effaced; apical male ventrite with 2 marginal setae. Legs-metatarsomere 1 / metatibial length ratio = 0.19; metatarsomere 4 length along outer lobe 1.33 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci narrow, shallow, median area broad. Microsculpture of elytral disc consisting of distinct isodiametric and transverse sculpticells arranged in a mesh, elytral apex with transverse mesh, sculpticell breadth 2 x length; metasternum with distinct transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex rufobrunneous; antennomeres 1 - 3 rufoflavous, 4 - 11 darker, brunneous; pronotal disc rufobrunneous with piceous cast, lateral margins, base, and apex narrowly rufobrunneous; proepipleuron rufoflavous, proepisternum rufobrunneous; elytral disc rufobrunneous with slightly metallic reflection, sutural interval concolorous basally, rufoflavous apically; intervals 7 - 8 inside humeral angle plus lateral marginal depression rufoflavous; elytral apex contrastedly flavous from apical terminus of interval 4; elytral epipleuron dorsally rufoflavous, ventrally rufobrunneous, metepisternum rufobrunneous; abdomen with ventrites 1 - 6 medially rufopiceous, ventrites 3 - 6 laterally rufoflavous, abdominal apical ventrite with apical 1 / 6 paler, flavous; metafemur flavous; metatibia flavous with brunneous cast. Male genitalia (n = 1). Aedeagal median lobe robust, distance between parameral articulation and tip 3.8 x depth at midlength (Fig. 61 I); apex trapezoidal, with flat apical face and angled ventral margin meeting at tightly rounded tip; internal sac with diffusely developed dorsal ostial microtrichial patch, flagellar plate short, length 0.3 x parameral articulation-tip distance.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
716C0F2FF5931E7B5E76D44F579C5B21.taxon	etymology	Etymology. The similarity of this species to Mecyclothorax apicalis makes the species epithet parapicalis appropriate; the adjectival epithet meaning like apicalis	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
AEC1E67F27CFF21C12ED9A02FE519A92.taxon	diagnosis	Diagnosis. Among the assemblage of Mecyclothorax ovipennis group species with punctate discal elytral striae and concolorous elytral intervals, this and the next species, Mecyclothorax subternus, stand out due to their broadly ellipsoid to obovoid elytra (Fig. 68 A-B). The pronotum characterizing these species is also very constricted basally-MPW / BPW = 1.58 - 1.72 for this species-with a minutely punctate, glossy median base. Mecyclothorax mauiae can be diagnosed from Mecyclothorax subternus by the better developed microsculpture: 1, vertex with evident transverse mesh, sculpticell breadth 2 x length; and 2, pronotal and elytral discs and elytral apex with transverse mesh, sculpticell breadth 2 - 3 x length. The eyes also tend to be less convex and slightly smaller than in Mecyclothorax subternus, with ocular ratio = 1.43 - 1.50, and ocular lobe ratio = 0.75 - 0.79 in this species. The final arbiter for any identification involving a male specimen is the apex of the aedeagal median lobe, with the apex always rounded in male of this species (Fig. 69) versus acuminate in Mecyclothorax subternus (Fig. 70 A-D). The degree of strial development varies among individuals of this species, potentially allowing specimens of this species to be confused with those of Mecyclothorax nanunctus of the Mecyclothorax palustris group. That group is characterized by a less depressed apex on elytral stria 2; a character exhibiting some infraspecific variation. Specimens of Mecyclothorax mauiae always exhibit broader pronotal lateral marginal depressions, and more ellipsoid elytra than observed in individuals of Mecyclothorax nanunctus. As with Mecyclothorax subternus, examination of male genitalia will finalize the diagnosis, as the male aedeagal median lobe of Mecyclothorax nanunctus is much more elongate and gracile, with the apex terminated in narrowly projected tip (Fig. 153 D-J). Setal formula 2 1 2 0. Standardized body length 3.4 - 4.5 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
AEC1E67F27CFF21C12ED9A02FE519A92.taxon	description	Description (n = 5). Head capsule with frontal grooves broad near clypeus, lateral carina to supraorbital seta; dorsal surface of neck flat to slightly concave; labral anterior margin very shallowly emarginate medially; antennae filiform, antennomeres 2 - 3 with sparse pelage of short setae; mentum tooth with sides acute, apex rounded. Pronotum cordate, moderately transverse, MPW / PL = 1.23 - 1.27; hind angle right to slightly acute, lateral margin convergent just before angle; median base almost coplanar to slightly depressed relative to disc, ~ 12 sparsely distributed punctures each side; basal margin convexly expanded between laterobasal depressions; median longitudinal impression shallow, finely incised, crossed by fine transverse wrinkles; anterior transverse impression narrow, shallowly incised laterally, obsolete medially; anterior callosity slightly convex, smooth to slightly irregular due to fine wrinkles, surface glossy; front angles slightly projected, rounded; pronotal apical width greater than basal width, APW / BPW = 1.03 - 1.11; lateral marginal depression moderately narrow, edge beaded anterad lateral seta, broader at front angle, edge little upturned in basal half; laterobasal depression narrow and deep, continuous with lateral depression. Proepisternum with 5 minute punctures along hind marginal groove; prosternal process with narrow median impression, lateral margins broadly beaded between coxae. Elytra broadly obovoid, disc narrowly flat medially, sides steeply sloped; basal grooves briefly recurved to proximate humeral angles, the tightly rounded to subangulate angles defined by hitch at the base of the lateral marginal depression, MEW / HuW = 2.28 - 2.51; parascutellar seta present; parascutellar striole with 4 isolated punctures, striole may be discontinuous between adjacent punctures; sutural interval slightly more convex than lateral intervals, sutural juncture upraised; sutural stria shallow between deep, round punctures basally, smooth, moderately deep apically, 2 nd stria shallower but also with rounded punctures on disc, shallower and broader apically, the two striae of subequal depth at elytral apex; discal striae 3 - 4 very shallow, punctate on disc, stria 5 very shallow, traceable, striae 6 - 7 obsolete, associated inner intervals slightly convex, lateral intervals flat; 8 th interval convex, though striae are obsolete in that area of elytra; 2 dorsal elytral setae at 0.31 x and 0.56 - 0.58 x elytral length, setal impressions shallow, spanning 1 / 2 to 2 / 3 width of interval 3; apical and subapical setae absent; lateral elytral setae arranged in anterior series of 7 setae and posterior series of 6 setae; elytral marginal depression slightly broader at humerus, narrowed laterally and posteriorly to beadlike at subapical sinuation; subapical sinuation shallow, concavity symmetrical. Mesepisternum with ~ 8 punctures in 1 - 2 rows; metepisternal width to length ratio = 0.70; metepisternum / metepimeron suture distinct. Abdomen with irregular lateral wrinkles on ventrites 1 - 5, lateral depressions on ventrites 3 - 6; suture between ventrites 2 and 3 complete; apical male ventrite with 2 marginal setae; apical female ventrite with 4 equally spaced marginal setae and median trapezoid of 4, subequal, short setae. Legs-metatarsomere 1 / metatibial length ratio = 0.18; metatarsomere 4 length along outer lobe 1.4 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci narrow, shallow, median area broad. Microsculpture of pronotal median base obsolete medially, the surface glossy, a transverse mesh present laterally between punctures; metasternum with shallow transverse mesh; laterobasal abdominal ventrites with swirl ing isodiametric and transverse microsculpture. Coloration of vertex rufobrunneous; antennomere 1 flavous, antennomeres 2 - 3 brunneous, 4 - 11 with piceous cast; pronotal disc rufobrunneous, lateral margins narrowly, and base and apex rufous; proepipleuron rufoflavous, proepisternum rufobrunneous; elytral disc rufobrunneous basally, apical half the same with piceous cast, sutural interval rufous throughout, intervals 8 - 9 rufoflavous, paler apically; elytral epipleuron dorsally rufoflavous, ventrally darker, metepisternum rufobrunneous; abdomen with ventrites 1 - 3 rufobrunneous, ventrites 4 - 5 darker, rufopiceous, apical ventrite 6 rufoflavous mediobasally, apical half flavous; metafemur flavous; metatibia flavous with brunneous cast. Male genitalia (n = 50). Aedeagal median lobe variably robust, distance between parameral articulation and tip 2.2 - 3.3 x depth at midlength (Fig. 69 E, M), but apex always little extended beyond ostial opening, with tip broadly rounded (Fig. 69); median lobe symmetrical in ventral view, broadest at midlength of lobe shaft, evenly narrowed to blunt tip which lies on right side of apex (Fig. 69 B); internal sac broad, of variable length (Fig. 69 H-J), sac length from ostium to base of flagellar plate 0.77 - 1.1 x parameral articulation-tip distance, surface unornamented except denser microspicules may occur on ventral surface (Fig. 69 J); flagellar plate very large, length 0.60 - 0.67 x parameral articulation-tip distance. Female reproductive tract (n = 2). Bursa copulatrix columnar, elongate, length 1.08 - 1.16 mm, breadth 0.33 - 0.36 mm, an apical lobe offset from basal portion by curved constriction, apical lobe 0.49 mm long (Fig. 62 D-E); bursal walls translucent with thick wrinkles; gonocoxite 1 with 3 - 4 apical fringe setae, the outermost setae of series smallest, and 5 - 8 smaller setae on medial surface (Fig. 63 D); gonocoxite 2 falcate with tightly rounded tip, base with lateral panhandle extension with 90 ° bend at terminus, 2 lateral ensiform setae with apical seta longer and broader, apical nematiform setae on medial surface at 0.68 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
AEC1E67F27CFF21C12ED9A02FE519A92.taxon	etymology	Etymology. This species' similarity to Mecyclothorax molokaiae of Moloka'i, suggests use of mauiae as the species epithet, the first declension genitive singular form to mean Maui's Mecyclothorax.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
47A1C538FE83EA7E715FCB979FE2F656.taxon	diagnosis	Diagnosis. Individuals of this species (Fig. 68 B) are nearly identical externally to those of Mecyclothorax mauiae (Fig. 68 A), though the eyes tend to be larger and more convex; ocular ratio = 1.48 - 1.54, ocular lobe ratio = 0.79 - 0.82, and the dorsal surface is less sculptured; 1, vertex, pronotal disc and median base, and elytral disc glossy, microsculpture obsolete; 2, elytral apex with shallow to obsolete transverse mesh, the surface glossy; 3, metasternum with shallow transverse-mesh microsculpture. The male aedeagal median lobe certifies the identification, as the apex terminates in a pointed tip (Fig. 70 A-C) instead of the broadly rounded apex of Mecyclothorax mauiae (Fig. 69). Setal formula 2 1 2 0. Standardized body length 3.4 - 4.2 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
47A1C538FE83EA7E715FCB979FE2F656.taxon	description	Description (n = 5). The description of Mecyclothorax mauiae also serves for this species with the following substitutions. Pronotum slightly broader basally and more transverse than that of Mecyclothorax mauiae; MPW / BPW = 1.57 - 1.65, MPW / PL = 1.23 - 1.32. Pronotal lateral margin narrow, beaded, not wider at front angles. Discal elytral striae shallow between deep, rounded and nearly isolated punctures; discal elytral interval 2 slightly convex, intervals 3 - 4 nearly flat. Mesepisternum smooth, with 3 - 4 punctures arrayed in 1 row. Coloration pale (though single topotypic type series may include exclusively somewhat teneral individuals); vertex rufous; antennomere 1 flavous, antennomeres 2 - 3 rufoflavous, 4 - 11 brunneous; pronotal disc and margins pale rufous; elytral disc basally rufous, apically rufobrunneous, sutural interval basally rufoflavous, apically flavous; abdominal ventrites 1 - 5 mediobasally rufous, laterally and apically rufobrunneous; metafemur flavous with medial brunneous cloud. Male genitalia (n = 2). Aedeagal median lobe robust, distance between parameral articulation and tip 2.5 x depth at midlength (Fig. 70 A-C); apex little extended beyond ostial opening, tip acutely pointed; median lobe symmetrical in ventral view, right margin broadly concave, left margin incurved to blunt tip which is on right side of apex (Fig. 70 D); internal sac very broad basally, with oblong ventral ostial microtrichial patch on right side (Fig. 70 B); flagellar plate very large, length 0.67 x parameral articulation-tip distance. Female reproductive tract (n = 1). Bursa copulatrix a broad sac, length 1.23 mm, maximum breadth in apical half 0.65 mm, breadth at vagina 0.34 mm (Fig. 62 F); bursal walls translucent with thick wrinkles; gonocoxite 1 with 3 apical fringe setae, middle seta of series largest, 8 smaller setae along medial margin (Fig. 63 E); gonocoxite 2 falcate with acuminate tip, base with sinuous lateral extension, 2 lateral ensiform setae with apical seta broader and longer, apical nematiform setae on medial surface at 0.72 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
47A1C538FE83EA7E715FCB979FE2F656.taxon	etymology	Etymology. The adjectival species epithet subternus, meaning that which is underneath (Jaeger 1955), signifies the geographical range of this species being at the lower elevational bounds of its closely related, and much more broadly distributed relative, Mecyclothorax mauiae (Fig. 71).	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
D24661784E33B53780D235F1D49E487A.taxon	diagnosis	Diagnosis. This species is the most gracile-bodied Mecyclothorax ovipennis group species with cordate pronotum, narrowed elytral humeri, and punctate discal elytral intervals (Fig. 68 C). In body conformation it is most similar to Mecyclothorax ovipennis (Fig. 65 A), and both species share the setal formula 2 2 2 1 [ae]. However individuals of this species lack the parascutellar seta. Also, this species is uniquely characterized among species in the group by the apically flavous abdomen, with the flavous ventrites 4 - 6 contrasted to the rufobrunneous basal ventrites (Fig. 68 D). Standardized body length 3.7 - 4.1 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
D24661784E33B53780D235F1D49E487A.taxon	description	Description (n = 5). Head capsule with frontal grooves broad near clypeus, lateral carina to supraorbital seta; dorsal surface of neck flat to slightly concave; eyes large and moderately convex, ocular ratio = 1.51 - 1.56, ocular lobe ratio = 0.82 - 0.88; labral anterior margin deeply angulate medially, emarginated 1 / 3 of length; antennae filiform; antennomeres 2 - 3 with only a few short setae along shafts; mentum tooth with sides acute, apex tightly rounded. Pronotum quadrisetose, distinctly constricted basally, MPW / BPW = 1.49 - 1.58, with lateral margins convergent before the right to slightly acute, projected hind angles; pronotum appearing narrow, but actually slightly transverse, MPW / PL = 1.13 - 1.21; median base broadly depressed relative to disc, rugose medially, longitudinally strigose laterally; basal margin very slightly convex between laterobasal depressions; median longitudinal impression evident, finely incised, continued onto median base; anterior transverse impression broad, surface with granulate microsculpture and deep, dense longitudinal wrinkles; anterior callosity depressed, covered with longitudinal wrinkles from transverse impression; front angles not projected, tightly rounded; pronotal apical width greater than basal width, APW / BPW = 1.06 - 1.09; lateral marginal depression moderately narrow, edge upturned, slightly broader inside front angle; laterobasal depression slightly transversely wrinkled, continuous with lateral depression. Proepisternum with 4 - 5 minute punctures along hind marginal groove; prosternal process with narrow median impression, lateral margins broadly beaded between coxae. Elytra distinctly ovoid with maximal width behind midlength, disc narrowly flat medially, sides steeply sloped; basal groove gently recurved to subangulate humerus that is defined by hitch at base of lateral marginal depression, MEW / HuW = 2.10 - 2.18; parascutellar striole very shallow, smooth, difficult to trace; sutural interval slightly more convex than lateral intervals, but depressed at suture; sutural and 2 nd striae of subequal depth from base to apex, striae 2 - 6 progressively shallower on disc, stria 7 discontinuous, obsolete, discal striae 1 - 4 with small punctures restricted to deepest parts of striae; 8 th interval slightly more convex than apical fused portion of striae 5 + 7; 2 dorsal elytral setae at 0.30 - 0.31 x and 0.61 - 0.66 x elytral length, setal impressions shallow, spanning 1 / 2 width of interval 3; apical elytral seta present near medial apex of interval 3, subapical seta absent; lateral elytral setae arranged in anterior series of 7 setae and posterior series of 6 setae; elytral marginal depression moderately narrow, edge upraised until beaded at subapical sinuation; subapical sinuation shallow, more abruptly incurved anteriorly. Mesepisternum with ~ 8 punctures in 1 - 2 rows; metepisternal width to length ratio = 0.71; metepisternum / metepimeron suture distinct. Abdomen with irregular lateral wrinkles on ventrites 1 - 5, lateral depressions on ventrites 3 - 6 (Fig. 68 D); suture between ventrites 2 and 3 reduced laterally, effaced; apical male ventrite with 2 marginal setae, apical female ventrite with 4 equally spaced setae plus median trapezoid of 4 subequal, short setae. Legs-metatarsomere 1 / metatibial length ratio = 0.21; metatarsomere 4 length along outer lobe 1.33 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci narrow, shallow, median area broad. Microsculpture of vertex and pronotal disc a transverse mesh, sculpticell breadth 2 - 3 x length; pronotal median base with granulate isodiametric and transverse mesh, sculpticell breadth 2 x length; elytral disc with shallow isodiametric and transverse mesh, sculpticell breadth 2 x length, in transverse rows; elytral apex with shallow isodiametric sculpticells in transverse rows; metasternum with upraised transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex rufobrunneous with piceous cast; antennomeres 1 - 3 flavous, 4 - 11 darker, more brunneous; pronotal disc rufopiceous, lateral margins, base, and apex narrowly rufoflavous; proepipleuron rufoflavous, proepisternum rufobrunneous; elytral disc dark rufobrunneous, sutural interval narrowly rufobrunneous basally, rufoflavous apically; elytral marginal depression and 9 th interval rufoflavous, elytral apex narrowly flavous; elytral epipleuron dorsally flavous, ventrally rufoflavous, metepisternum rufopiceous; abdomen with ventrites 1 - 2 rufobrunneous, ventrite 3 medially rufobrunneous, laterally and apically flavous, ventrites 3 - 6 flavous; metafemur flavous; metatibia flavous with a brunneous cast. Male genitalia (n = 5). Aedeagal median lobe variably gracile, distance between parameral articulation and tip 3.7 - 4.2 x depth at midlength (Fig. 70 E, H, I); apex extended twice its depth beyond apex of ostial opening, tip slightly downturned, rounded; median lobe curved rightward toward apex in ventral view (Fig. 70 F), right margin concave, left margin incurved to blunt tip; internal sac elongate, length from ostial margin to base of flagellar plate 0.79 x parameral articulation-tip distance, sac surface unornamented but with microspicules more developed over entire ventral surface (Fig. 70 G); flagellar plate moderately short, length 0.36 x parameral articulation-tip distance. The male from Kīpahulu Valley, 1500 m elevation (Fig. 70 H) exhibits a broader apex, though the internal sac in its uneverted condition looks identical to the configuration of the Kīpahulu Valley, 1845 m elevation male (Fig. 70 E). All of these specimens exhibit the flavous abdominal ventrites that represent an autapomorphy for the species, and if the lower elevation Kīpahulu Valley population is determined to be distinct, the additionally recognized species will be the sister to Mecyclothorax flaviventris. A present, with only a single specimen from the lower Kīpahulu Valley locality, such recognition is deemed premature, and the aedeagal conformations are assumed to represent infraspecific variability. Female reproductive tract (n = 1). Bursa copulatrix columnar with rounded, slightly expanded apex, length 0.68 mm, breadth 0.29 mm (Fig. 62 G); bursal walls translucent with thin wrinkles; gonocoxite 1 with 0 - 1 apical fringe setae and 7 - 8 smaller setae on medial surface (Fig. 63 F); gonocoxite 2 falcate with tightly rounded tip, base with long, thin lateral extension with 90 ° bend at terminus, 2 lateral ensiform setae, the apical seta longer and broader, apical nematiform setae on medial surface at 0.69 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
D24661784E33B53780D235F1D49E487A.taxon	etymology	Etymology. The Latin genitive singular flaviventris signifies the flavous abdominal ventrites 4 - 6 contrasted to the piceous abdominal base.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
9D0A643D321CA75A65D9521313C4BDC6.taxon	diagnosis	Diagnosis. The rufobrunneous dorsal body color, narrow, cordate pronotum, and well-developed dorsal microsculpture-isodiametric on the elytra-will allow individuals of this species to be identified in the field using a hand lens (Fig. 73 A). More microscopic characters amply diagnosis this species, including: 1, elytral parascutellar seta absent; 2, dorsal elytral setae absent, though a single seta may be present in rare instances; 3, vertex with shallow transverse mesh, sculpticell breadth 2 x length, and pronotal disc with transverse mesh, sculpticell breadth 2 - 3 x length. Setal conformation is highly variable among individuals of this species, with the basal pronotal setae present or absent, and either both apical and subapical elytral setae, or just the subapical seta present. Setal formula 2 1 - 2 0 (1) 1 - 2 [sae]. Standardized body length 3.9 - 4.6 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
F79E9183EA59A7B4A65D03840C3D9BA7.taxon	diagnosis	Diagnosis. This species (Fig. 73 B), Mecyclothorax cordaticollaris (Fig. 73 C), and Mecyclothorax subconstrictus (Fig. 73 D) are the only three Mecyclothorax ovipennis group species to exhibit the 2 1 1 0 setal formula (at least for some of the individuals of the latter species). Mecyclothorax cordaticollis and Mecyclothorax subconstrictus share shallow, minutely punctate discal elytral intervals and associated slightly convex intervals, plus a narrowly constricted pronotum; MPW / BPW = 1.55 - 1.61 in this species. The discal elytral striae are much more punctate in Mecyclothorax cordaticollaris, and the associated intervals are much more convex. Mecyclothorax cordaticollis and Mecyclothorax subconstrictus can be definitively diagnosed using dorsal microsculpture. The pronotal disc of Mecyclothorax cordaticollis bears transverse-mesh microsculpture, sculpticell breadth 2 - 3 x length, and the elytral disc is covered with a transverse mesh, sculpticell breadth 2 - 4 x length. For Mecyclothorax subconstrictus, the pronotal disc is glossy medially, and laterally covered with an obsolete transverse mesh, whereas the elytral disc bears a mixture of isodiametric and transverse sculpticells. Standardized body length 3.6 - 4.1 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
77E6782E578FFEB1D7CE02BD15452171.taxon	diagnosis	Diagnosis. Like Mecyclothorax cordaticollis (Fig. 73 B) or Mecyclothorax subconstrictus (Fig. 73 D) in the setal formula- 2 2 1 0 - but with much deeper elytral striae, the discal striae 1 - 5 distinctly punctate with the punctures expanding strial breadth (Fig. 73 C). The pronotal median base is minutely punctate, ~ 15 distinct punctures each side, with the punctures elongate at the juncture with the disc. The pronotal disc is glossy, with obsolete transverse-mesh microsculpture, sculpticell breadth 3 x length, and the median base is glossy medially, and with an isodiametric mesh present laterally between the punctures. The elytra are relatively broader basally than in Mecyclothorax cordaticollis or Mecyclothorax subconstrictus, with the basal groove distinctly recurved to meet the angulate humerus, the angle defined by a hitch at the base of the lateral marginal depression. Standardized body length 3.9 - 4.4 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
77E6782E578FFEB1D7CE02BD15452171.taxon	description	Description (n = 4). Head capsule with frontal grooves broad near clypeus, straight, lateral carina to anterior supraorbital seta; dorsal surface of neck flat to slightly concave; eyes moderately convex, ocular ratio = 1.48 - 1.50, ocular lobe ratio = 0.78 - 0.82; labral anterior margin very shallow emarginate medially; antennae filiform, antennomeres 2 - 3 with sparse pelage of short setae; mentum tooth with sides right, apex rounded. Pronotum transverse, MPW / PL = 1.24 - 1.27, constricted basally, MPW / BPW = 1.47 - 1.55; hind angle right, lateral margins subparallel to slightly convergent anterad projected angles; basal margin slightly, evenly convex between laterobasal depressions; median longitudinal impression shallow, finely incised; anterior transverse impression moderately deep, smooth, finely incised; anterior callosity slightly convex, smooth, glossy; front angles very slightly produced, broadly rounded; apical and basal pronotal widths subequal, APW / BPW = 0.95 - 1.0; lateral marginal depression narrow, edge upturned anterad seta, slightly broader at front angle, beadlike margin from midlength to basal sinuation; laterobasal depression smooth laterad median base, broadly raised in explanate lateral margin. Proepisternum with 5 minute punctures along hind margin; prosternal process with narrow median impression, lateral margins broadly beaded between coxae. Elytra broadly ovoid, disc moderately convex, sides more so; MEW / HuW = 2.11 - 2.16; parascutellar seta present; parascutellar striole with 4 punctures, continuous between punctures; sutural interval only slightly more convex than lateral intervals in basal half, more convex apically; sutural and 2 nd striae of subequal depth from base to apex; elytral intervals 2 - 4 moderately convex, lateral intervals flatter; 8 th interval slightly more convex than fused apical portion of striae 5 + 7; one dorsal elytral seta at 0.23 - 0.28 x elytral length, setal impression small, spanning 1 / 2 width of interval 3; apical and subapical setae absent; lateral elytral setae arranged in anterior series of 7 setae and posterior series of 6 setae; elytral marginal depression narrow, margin upturned, beadlike near subapical sinuation; subapical sinuation shallow, more abruptly incurved anteriorly. Mesepisternum with ~ 10 punctures in 2 - 3 rows; metepisternal width to length ratio = 0.65; metepisternum / metepimeron suture distinct. Abdomen with irregular lateral wrinkles on ventrites 1 - 5 and lateral depressions on ventrites 3 - 6; suture between ventrites 2 and 3 reduced laterally, effaced; apical male ventrite with 2 marginal setae, apical female ventrite with 4 equally spaced marginal setae plus median trapezoid of 4 subequal, short setae. Legs-metatarsomere 1 / metatibial length ratio = 0.21; metatarsomere 4 length along outer lobe 1.25 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci shallow, narrow, median area broad. Microsculpture of vertex distinct, transversely stretched, sculpticell breadth 2 - 3 x length; elytral disc with shallow transverse mesh, sculpticell breadth 3 - 4 x length, apex with more developed transverse mesh of same dimensions; metasternum with shallow transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex rufobrunneous; antennomeres 1 - 3 flavous, 4 - 11 darker, more brunneous; pronotal disc rufobrunneous with piceous cast, lateral margins, base, and apex rufoflavous; proepipleuron rufoflavous, proepisternum rufobrunneous with piceous cast; elytral disc rufobrunneous, sutural interval paler rufous basally, rufoflavous apically; elytral lateral marginal depression narrowly rufoflavous, apex contrastedly flavous from apical terminus of interval 4; elytral epipleuron rufoflavous, metepisternum rufopiceous; abdominal ventrites 1 - 2 rufopiceous, ventrites 3 - 5 medially rufopiceous, laterally paler, apical ventrite with apical half flavous; metafemur flavous; metatibia flavous with brunneous cast. Male genitalia (n = 1). Aedeagal median lobe extremely slender, apically narrowed, needlelike, distance from parameral articulation to tip 5.9 x depth at midlength (Fig. 76 C); apex elongate, very narrow, angled slightly downward about half way along apical extension, with tip narrowly rounded; median lobe straight in ventral view, right and left margins approaching each other for 1 / 3 lobe length in this view, tip narrowly pointed (Fig. 76 D); internal sac without apparent microtrichial fields in uneverted specimen, flagellar plate length estimated to be 0.35 x parameral articulation-tip distance. Female reproductive tract (n = 1). Bursa copulatrix broad basally at vagina, with elongate, digitiform apical lobe, overall bursal length 0.74 mm, with apical lobe 0.51 mm long x 0.19 mm broad, and basal bulb at vagina 0.23 mm long x 0.41 mm broad (Fig. 62 J); bursal walls smooth, only lightly wrinkled, the walls of apical lobe thinner, less stained than broad base with darker staining and thicker wrinkles; gonocoxite 1 with 4 apical fringe setae, medial surface lined with 7 - 10 smaller setae (Fig. 74 C); gonocoxite 2 falcate with subacuminate apex, base broadly extended by short panhandle with curved terminus, 2 short lateral ensiform setae with apical seta longer and broader, apical nematiform setae on medial surface at 0.68 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
77E6782E578FFEB1D7CE02BD15452171.taxon	etymology	Etymology. This species' great similarity to Mecyclothorax cordaticollis leads to use of the similar epithet cordaticollaris. As in the former name, this adjectival epithet is meant to signify the cordate pronotum.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
BCFA3D9E18206B6CD09A44E2D6890269.taxon	diagnosis	Diagnosis. This, the third species of the group to exhibit setal formula 2 1 1 0 for at least some individuals, can be diagnosed from the other two- Mecyclothorax cordaticollis (Fig. 73 B) and Mecyclothorax cordaticollaris (Fig. 73 C) - by the following combination: 1, pronotal hind angles obtuse, lateral margin divergent from angle (Fig. 73 D), not parallel before angle; 2, elytra broader relative to pronotal width, MEW / MPW = 1.53 - 1.66, versus values of 1.47 - 1.52 observed in specimens of the other two species; 3, elytral disc with mixture of isodiametric and transverse sculpticells, sculpticell breadth 2 x length, versus more transverse meshes in the other two species. Setal formula 2 1 1 (2) 0; individuals have a posterior dorsal elytral seta present unilaterally in rare instances. Standardized body length 3.7 - 4.3 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
2F1CF157AC8D98BD71E200D5764F0AB0.taxon	diagnosis	Diagnosis. This species is uniquely characterized by the very broad pronotum and the remarkably narrow, subparallel elytra (Fig. 78 A); MEW / MPW = 1.31 - 1.36. The only other Haleakala species to approach this conformation is Mecyclothorax pusillus (Fig. 78 C). The pronotum of Mecyclothorax nubicola also distinctively diagnoses the species, with the hind angles represented by a slight widening of the lateral marginal bead, with the lateral margin only slightly sinuate anterad the very obtuse angle. Also, this species is composed of small beetles; standardized body length 3.4 - 3.5 mm. The setal formula of 2 2 2 2 is shared with the following three species; Mecyclothorax krushelnyckyi, Mecyclothorax pusillus, and Mecyclothorax rusticus. All four species also lack the parascutellar seta.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
EE21C42545CB97FC2BE02F590142A49D.taxon	diagnosis	Diagnosis. This species (Fig. 78 B) can be diagnosed from the other three small-bodied Mecyclothorax ovipennis group species with setal formula 2 2 2 2 - Mecyclothorax nubicola (Fig. 78 A), Mecyclothorax pusillus (Fig. 78 C), and Mecyclothorax rusticus (Fig. 78 D) - by: 1, the more narrowly constricted pronotal base; MPW / BPW = 1.54; 2, subellipsoid elytra, with the margin broadly rounded posterad the humerus, and the lateral margins evenly convex laterally; 3, elytra broad relative to forebody, with MEW / MPW = 1.51 and MEW / MHW = 2.15; and 4, the glossy upper body surface with little-developed microsculpture. The vertex is glossy, with obsolete transverse sculpticells, breadth 2 x length in depressions. The pronotal disc is glossy, with obsolete transverse microsculpture visible in angled light; the pronotal base with obsolete transverse microsculpture over portions of the cuticle. The elytral disc is also glossy, with a transverse mesh, sculpticell breadth 2 - 3 x length, restricted to the lateral and apical areas. Standardized body length 3.95 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
EE21C42545CB97FC2BE02F590142A49D.taxon	description	Description (n = 1). Head capsule with frontal grooves broad near clypeus, lateral carina to anterior supraorbital seta; dorsal surface of neck slightly concave; eyes moderately developed, ocular ratio = 1.46, ocular lobe ratio = 0.78; labral anterior margin very shallowly emarginate apically; antennae filiform, antennomeres 2 - 3 with only 1 - 2 setae each along shafts; mentum tooth with sides acute, apex rounded. Pronotum transverse, MPW / PL = 1.28; hind angle sharply obtuse, lateral margin subparallel for short distance anterad angle; median base slightly depressed, ~ 10 punctures each side, punctures more elongate at juncture with disc; basal margin convexly expanded between laterobasal depressions; median longitudinal impression shallow, finely incised; anterior transverse impression deep, finely incised, area behind impression depressed relative to disc; anterior callosity convexly upraised, glossy; front angles slightly projected, rounded; pronotal apical width slightly greater than basal width, APW / BPW = 1.04; lateral marginal depression very narrow, edge appearing beaded except at front angle where margin is slightly broader; laterobasal depression narrow, continuous with lateral depression. Proepisternum with 5 minute punctures along hind marginal groove; prosternal process with narrow median impression, lateral margins broadly beaded between coxae. Elytra with disc flat, sides distinctly sloped; basal groove distinctly recurved to tightly rounded humeri, base relatively broad, MEW / HuW = 1.98; parascutellar seta absent; parascutellar striole with 4 punctures, striole shallow between punctures; sutural interval coplanar with lateral intervals basally, upraised apically; sutural and 2 nd striae of subequal depth from base to apex; discal striae 1 - 5 minutely punctate, the punctures joined by depressed stria medially, striae 4 - 5 composed of isolated punctures, inner intervals slightly convex, lateral intervals flat; 8 th interval slightly more convex than fused apical portion of striae 5 + 7; 2 dorsal elytral setae at 0.24 - 0.30 x and 0.57 x elytral length, setal impressions moderate, spanning 2 / 3 of interval 3; apical and subapical setae present; lateral elytral setae arranged in anterior series of 7 setae and posterior series of 6 setae; elytral marginal depression narrow from humerus to midlength, gradually reduced to beadlike margin at subapical sinuation; subapical sinuation shallow, more abruptly incurved anteriorly. Mesepisternum smooth; metepisternal width to length ratio 0.70; metepisternum / metepimeron suture distinct. Abdomen with irregular lateral wrinkles on ventrites 1 - 5; suture between ventrites 2 and 3 complete; apical female ventrite with 4 equally spaced setae plus a median trapezoid of 4 subequal, short setae. Legs-metatarsomere 1 / metatibial length ratio = 0.19; metatarsomere 4 length along outer lobe 1.33 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci shallow, narrow, median area broad. Microsculpture of metasternum a shallow transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex rufobrunneous with piceous cast; antennomeres 1 - 3 flavous, 4 - 11 darker, more brunneous; pronotal disc rufopiceous; pronotal lateral margins very narrowly and base and apex rufoflavous; proepipleuron flavous, proepisternum rufopiceous; elytral disc rufopiceous, sutural interval rufous in basal half, rufoflavous in apical half, lateral marginal depression and apex of interval 9 rufoflavous; elytral epipleuron dorsally flavous, ventrally rufoflavous, metepisternum rufopiceous; abdomen with ventral ventrites 1 - 5 and base of 6 rufopiceous, apical 1 / 3 of ventrite 6 flavous; metafemur flavous; metatibia flavous, only tibial spines brunneous. Female reproductive tract. The lone female holotype was not dissected.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
EE21C42545CB97FC2BE02F590142A49D.taxon	etymology	Etymology. This species is named to honor Dr. Paul Krushelnycky's numerous, important discoveries of Mecyclothorax and Blackburnia species in the high elevation habitats of Haleakala.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
F59A5A056D69454D62F872C36927E06B.taxon	diagnosis	Diagnosis. Of the four Mecyclothorax ovipennis group species from Haleakala with setal formula 2 2 2 2 - Mecyclothorax nubicola (Fig. 78 A), Mecyclothorax krushelnyckyi (Fig. 78 B), Mecyclothorax rusticus (Fig. 78 D), and this species (Fig. 78 C) - Mecyclothorax pusillus can be diagnosed by: 1, pronotal base moderately broad, MPW / BPW = 1.39 - 1.48; 2, pronotal hind angles projected, obtuse, the lateral margin subparallel for twice the distance of the basal articulatory socket; 3, elytra narrow relative to head, MEW / MHW = 1.85 - 1.93, with sides subparallel, MEW / HUW = 1.84 - 1.90, the humerus tightly rounded; 4, frons and vertex with evident shallow transverse-mesh microsculpture, sculpticell breadth 2 - 3 x length. Standardized body length 3.3 - 3.9 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
BF6C28AEBF39C4863A6FFBEE9114FBBC.taxon	diagnosis	Diagnosis. Individuals of this species (Fig. 78 D) are most like those of Mecyclothorax pusillus (Fig. 78 C), and they are best diagnosed by the broader, more convexly margined elytra. The basal groove is recurved evenly on the rounded humeri, not tightly rounded to subangulate as in Mecyclothorax pusillus. Also, the head is narrow relative to the elytra; MEW / MHW = 2.02 - 2.07, with slightly more elongate antennomeres. The discal elytral striae are deeper and more continuous than in Mecyclothorax pusillus, though the lateral striae are more similar between the species; striae 5 - 6 discontinuous, stria 7 a series of minute punctures. The pronotal median base is little depressed relative to the disc, and is covered with ~ 10 isolated, elongate punctures each side, the surface glossy between the punctures. Finally, the vertex is glossy, with only an obsolete transverse mesh visible in angled light, sculpticell breadth 2 - 3 x length. Setal formula 2 2 2 2. Standardized body length 3.7 - 4.3 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
89859D9006B69871108F2B9945CC6501.taxon	diagnosis	Diagnosis. This species is easily distinguished by the very small, little convex eyes; ocular ratio = 1.29 - 1.35, ocular lobe ratio = 0.68 - 0.69, in combination with the granulate isodiametric elytral microsculpture (Fig. 81 A). Setal formula 2 2 2 2. Among species of the Mecyclothorax argutor group, the beetles are of moderate size, standardized body length 4.8 - 5.5 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
89859D9006B69871108F2B9945CC6501.taxon	description	Description (n = 2). Head capsule with frontal grooves broad near clypeus, curved laterally near clypeal juncture, a straight ridgelike lateral carina extended from dorsal mandibular articulation to mesad anterior supraorbital seta; dorsal surface of neck flat to slightly convex; compound eye with approximately 12 - 13 ommatidia across the maximal horizontal diameter; labral anterior margin medially emarginate 0.2 x length; antennae filiform, antennomere 3 with sparse, short setae; mentum tooth with sides acute, apex pointed. Pronotum slightly transverse, MPW / PL = 1.17 - 1.27, constricted basally, MPW / BPW = 1.51 - 1.58; hind angle right, apex rounded, lateral margin parallel anterad angle; median base depressed relative to disc, covered with rugose wrinkles and round punctures; basal margin broadly, medially extended between laterobasal depressions; median longitudinal impression fine, shallow, crossed by shallow wrin kles; anterior transverse impression broad, shallow, crossed by longitudinal wrinkles; anterior callosity slightly convex, crossed by longitudinal wrinkles from anterior impression; front angles not projected, rounded; pronotal anterior width slightly greater than basal width, APW / BPW = 1.02 - 1.05; lateral marginal depression narrow, margin upturned to beaded; laterobasal depression moderately broad with low oblique ridge terminated at hind angle. Proepisternum with 6 punctures along hind marginal groove; prosternal process medially concave, margin smoothly upraised. Elytra subquadrate, narrow basally, the basal groove meeting lateral marginal depression at subangulate humerus, MEW / HuW = 1.81 - 1.94; disc flat, sides moderately depressed; parascutellar seta present; parascutellar striole shallow, smooth, continued onto base; sutural interval slightly more convex than lateral intervals; sutural and 2 nd striae of subequal depth from base to apex; discal striae 1 - 5 continuous, slightly irregular but impunctate, deeper and smooth apically, stria 6 reduced, stria 7 traceable, very shallow, associated intervals broadly, slightly convex; 8 th interval slightly convex laterad 7 th stria, principally due to deep apical portion of 7 th stria; 2 dorsal elytral setae at 0.30 x and 0.59 - 0.60 x elytral length, setal impressions shallow, about breadth of interval 3; apical and subapical setae present; lateral elytral setae arranged as anterior series of 7 setae and posterior series of 6 setae; elytral marginal depression moderately broad, lined with sculpticells, the margin upturned; subapical sinuation evident, abrupt and brief. Mesepisternum with ~ 14 punctures in 2 - 3 rows, metepisternal width to length ratio = 0.67; metepisternum / metepimeron suture distinct. Abdomen with shallow indistinct wrinkles on ventrites 1 - 3, suture between ventrites 2 and 3 complete; apical female ventrite with 4 equally spaced setae and median trapezoid of 4 (- 5 unilaterally) subequal, short setae. Legs-metatarsomere 1 / metatibial length ratio = 0.19; metatarsomere 4 length along outer lobe 1.4 x medial tarsomere length, with apical and subapical setae; metatarsal dorsolateral sulci broad, shallow. Microsculpture of vertex an upraised isodiametric mesh in transverse rows; pronotal disc with transverse mesh, median base with granulate isodiametric mesh; elytral apex with distinct transverse mesh; metasternum with distinct transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex rufobrunneous; antennomere 1 flavous, 2 - 11 rufoflavous; pronotal disc rufobrunneous, lateral margins narrowly paler in lateral depression, rufoflavous; proepipleuron rufoflavous, proepisternum rufobrunneous with piceous cast; elytral disc reflective rufobrunneous, sutural interval basally rufous, apically flavous, elytral margins broadly rufoflavous, apex broadly rufoflavous to flavous; elytral epipleuron flavous, metepisternum rufobrunneous with piceous cast; abdomen rufobrunneous with piceous cast, apical half of ventrite 6 broadly paler, flavous; metafemur flavous; metatibia rufoflavous. Female reproductive tract. The holotype and paratype females were not dissected.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
89859D9006B69871108F2B9945CC6501.taxon	etymology	Etymology. The species epithet compounds the Greek words plax, flat round plate, and ommatos, the genitive of omma, i. e. eye. As Greek, ommatoplax is treated as a noun. The epithet refers to the very flat eyes characteristic of this species.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
7ADA59E73D51965CE6C4EED79ACDC072.taxon	diagnosis	Diagnosis. This (Fig. 81 B) and the following Mecyclothorax refulgens (Fig. 81 C) synapomorphously exhibit pale, flavous elytral margins that contrast with the piceous elytral disc. Beetles of this species have broader bodies, with the pronotum more transverse, MPW / PL = 1.21 - 1.32, and the elytra more broadly ellipsoid, the greatest width just behind midlength. Cuticular microsculpture is less developed in this species, the vertex covered with shallow isodiametric sculpticells in transverse rows, the elytral disc covered with a shallow, elongate transverse mesh, sculpticell breadth 2 - 3 x length. Also, beetles of this species are larger than those of Mecyclothorax refulgens; standardized body length 5.1 - 5.4 mm. Setal formula 2 2 2 2.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
7ADA59E73D51965CE6C4EED79ACDC072.taxon	description	Description (n = 5). Head capsule with frontal grooves broad and moderately deep near clypeus, bordered laterally by broad carina to supraorbital seta; dorsal surface of neck flat to convex; eyes convex, of moderate size, ocular ratio = 1.39 - 1.47, ocular lobe ratio = 0.76 - 0.80; labral anterior margin broadly, shallowly emarginate; antennae filiform, antennomere 3 sparsely setose; mentum tooth with sides acute, apex tightly rounded. Pronotum cordate, lateral margins distinctly sinuate basally, hind angles acute, projected, MPW / BPW = 1.53 - 1.61; median base depressed relative to disc, with ~ 12 rounded punctures and sinuous depressions each side; basal margin slightly convex medially; median longitudinal impression fine, shallow, anterior transverse impression moderately deep, narrow, longitudinal wrinkles fore and aft; anterior callosity slightly convex, with indistinct longitudinal wrinkles; front angles slightly projected, tightly rounded; pronotal apex variably as broad as base, APW / BPW = 0.98 - 1.11; lateral marginal depression narrow throughout, margin upraised to beaded; laterobasal depression narrow, with low oblique ridge terminated at hind angle. Proepisternum with 6 punctures along hind marginal groove; prosternal process medially concave, margin smoothly upraised. Elytra moderately convex, disc narrowly flattened medially; basal groove recurved at tightly rounded humeral angle, MEW / HuW = 2.0 - 2.22; parascutellar seta present; parascutellar striole with 4 punctures, shallow between punctures; sutural interval of same convexity as intervals 2 - 3; sutural and 2 nd striae of subequal depth from base to apex; discal striae 1 - 3 shallow with small but evident punctures, lateral striae 4 - 6 shallower, traceable as isolated serial punctures, stria 7 obsolete except at elytral apex where it is broad and shallow; 8 th interval as convex as fused apical portion of striae 5 + 7; 2 dorsal elytral setae at 0.23 - 0.27 x and 0.57 - 0.69 x elytral width, setal impressions shallow, spanning 1 / 2 width of interval 3; apical and subapical setae present; lateral elytral setae arranged in anterior series of 7 setae and posterior series of 6 setae; elytral marginal depression moderately broad, lined with sculpticells, margin upturned; subapical sinuation shallow, nearly obsolete. Mesepisternum with ~ 9 punctures in 2 - 3 rows; metepisternal width to length ratio = 0.78; metepisternum / metepimeron suture distinct. Abdomen with shallow indistinct lateral wrinkles on ventrites 1 - 3; suture between ventrites 2 and 3 complete; apical male ventrite with 2 marginal setae and apical female ventrite with 4 equally spaced setae plus median trapezoid of 4 subequal, short setae. Legs-metatarsomere 1 / metatibial length ratio = 0.18; metatarsomere 4 length along outer lobe 1.4 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci broad, shallow. Microsculpture of pronotal median base of distinct isodiametric sculpticells in transverse rows; elytral apex with distinct transverse mesh; metasternum with shiny transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex rufobrunneous; antennomeres 1 - 4 flavous, 5 - 11 rufobrunneous; pronotal disc rufobrunneous, margins narrowly rufoflavous; proepipleuron rufoflavous, proepisternum rufobrunneous; elytral disc reflective rufopiceous, sutural interval rufous basally, flavous apically; elytral margins broadly paler, with intervals 7 - 9 or 8 - 9 rufoflavous near interval 6 to flavous laterally, apex broadly flavous apicad subapical sinuation; elytral epipleuron flavous with rufous cast, metepisternum rufobrunneous with piceous cast; abdomen with ventrites 1 - 2 piceous, 3 - 5 rufoflavous, 6 broadly flavous; metafemur flavous, metatibia rufoflavous. Male genitalia (n = 1). Aedeagal median lobe robust, distance from parameral articulation to tip 3.2 x depth at midlength (Fig. 82 A); apex narrowed distad ostial opening, tip expanded dorsally, knoblike; median lobe distinctly curved rightward near apex in ventral view (Fig. 82 B), left margin indented before parallel-sided, bluntly rounded tip; internal sac with small dorsal ostial microtrichial patch (left patch in Fig. 82 B) and larger ventral ostial patch; flagellar plate of moderate size, length 0.38 x parameral articulation-tip distance. Female reproductive tract (n = 1). Bursa copulatrix columnar with rounded apex, length 0.82 mm, breadth 0.37 mm (Fig. 83 A); bursal walls translucent, smooth with a few wrinkles; gonocoxite 1 with 4 apical fringe setae, the middle two setae larger, 1 small seta basad apicomedial angle, 5 - 7 smaller setae on medial surface (Fig. 84 A); gonocoxite 2 broadly falcate, base broadly extended laterally, 2 stout, apically narrowed lateral ensiform setae, apical nematiform setae on medial surface at 0.80 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
7ADA59E73D51965CE6C4EED79ACDC072.taxon	etymology	Etymology. The adjectival species epithet semistriatus refers to the medially striate and laterally smooth elytra.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
0A75D4202EA58D94F28C87D2F1649010.taxon	diagnosis	Diagnosis. This and Mecyclothorax semistriatus, comprise the pair of Mecyclothorax argutor group species characterized by contrastingly pale elytral margins and shallow elytral striae. This species (Fig. 81 C) can be told from Mecyclothorax semistriatus (Fig. 81 B) by the broadly paler pronotal and elytral lateral margins, and the basally narrower elytra, the elytral shape distinctly ovoid with the greatest width well behind midlength. Also, beetles of this species exhibit more well-developed microsculpture, with the elytral disc subiridescent due to the well-developed transverse and isodiametric sculpticells. The discal elytral striae are less punctate in this species, with only the sutural stria punctate basally, striae 2 - 5 at most slightly irregular. The sutural and 2 nd striae are of subequal depth only at the extreme apex of the elytra. More basally the sutural stria is broader and deeper. Setal formula 2 2 2 2. Standardized body length 4.8 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
0A75D4202EA58D94F28C87D2F1649010.taxon	description	Description (n = 1). Head capsule with frontal grooves broad near clypeus, a sinuous lateral carina extended to anterior supraorbital seta; dorsal surface of neck flat; eyes moderately convex, ocular lobe moderately protruded from gena, ocular ratio = 1.46, ocular lobe ratio = 0.79; labral anterior margin broadly, shallowly emarginate; antennae filiform, antennomere 3 sparsely setose; mentum tooth with sides acute, apex tightly rounded. Pronotum cordate, little transverse, MPW / PL = 1.19, lateral margins distinctly sinuate before very slightly acute, projected hind angles; pronotal base narrow, MPW / BPW = 1.63; median base depressed relative to disc, 12 - 14 punctures laterally plus wrinkles present along juncture with disc; basal margin straight; median longitudinal impression fine, very shallow; anterior transverse impression broad, shallow medially, crossed by longitudinal wrinkles, deep, continuous in lateral 2 / 3; anterior callosity slightly convex, longitudinal wrinkles indistinct; front angles slightly projected, tightly rounded; pronotal apical width greater than basal width, APW / BPW = 1.10; lateral marginal depression narrow throughout, margin upturned to beaded; laterobasal depression moderately broad, slightly raised medially. Proepisternum with 6 punctures along hind marginal groove; prosternal process medially concave, margin smoothly upraised. Elytra convex, sides depressed; basal groove meeting lateral marginal depression at subangulate humerus; elytra narrow basally, MEW / HuW = 2.10; parascutellar seta present; parascutellar striole shallow, smooth, with 1 - 2 punctures; sutural interval more convex than lateral intervals, the sutural juncture up raised; sutural stria deeper and more punctate than 2 nd stria on disc, discal striae 3 - 5 traceable, smooth, stria 6 obsolete, stria 7 absent, associated intervals broadly, slightly convex on disc, flat laterally; 8 th interval laterad position of 7 th stria not more convex than more medial elytral surface; 2 dorsal elytral setae at 0.23 - 0.29 x and 0.66 x elytral length, setal impressions shallow, broad, spanning interval 3; apical and subapical setae present; lateral elytral setae arranged in anterior series of 7 setae and posterior series of 6 setae; elytral marginal depression narrow, margin upturned; subapical sinuation shallow, broad. Mesepisternum with ~ 8 punctures in 2 - 3 rows; metepisternal width to length ratio = 0.79; metepisternum / metepimeron suture distinct. Abdomen with shallow, indistinct lateral wrinkles on ventrites 1 - 3; suture between ventrites 2 and 3 complete; apical female ventrite with 4 equally spaced setae and median pair of 2 small setae along ventrite margin. Legs-metatarsomere 1 / metatibial length ratio = 0.18; metatarsomere 4 length along outer lobe 1.4 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci narrow, canaliculated, median carina broad. Microsculpture of vertex isodiametric sculpticells in transverse rows; pronotal disc with transverse mesh and parallel lines, median base with distinct isodiametric sculpticells; elytral apex covered with reflective transverse mesh; metasternum with reflective transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex rufobrunneous with piceous cast; antennomeres 1 - 3 flavous, 4 - 11 rufobrunneous; pronotal disc rufopiceous, lateral margins and pronotal base broadly rufoflavous; proepipleuron rufoflavous, proepisternum rufobrunneous with piceous cast; elytral disc reflective rufopiceous, sutural interval rufous basally, flavous apically; elytral margins broadly paler, intervals 8 - 9 rufoflavous to flavous at marginal depression, elytral apex broadly rufoflavous to flavous apicad subapical sinuation; elytral epipleuron flavous, metepisternum rufobrunneous with piceous cast; abdomen with ventrites 1 - 2 and base of 3 piceous, apex of 3 rufoflavous, ventrites 4 - 6 flavous; metafemur flavous; metatibia rufoflavous. Female reproductive tract. The lone female specimen was not dissected.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
0A75D4202EA58D94F28C87D2F1649010.taxon	etymology	Etymology. The dorsal body surface of these beetles is quite reflective, leading to use of the present participle of the Latin verb refulgeo-to shine-as the species epithet. The nominative singular participle refulgens maintains its ending regardless of gender.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
799E729EC7FF6698207C79937DE8565E.taxon	diagnosis	Diagnosis. This species (Fig. 81 D) and Mecyclothorax argutuloides (Fig. 87 C) comprise a second sister-species pair within the Haleakala Mecyclothorax argutor group. Both species can be told by the broad, robust pronotum that is little constricted basally, MPW / BPW = 1.38 - 1.40. Of the two, this species displays a more robust appearance, with the pronotal lateral margins broad, and elytra broadly expanded laterally with broad, translucent lateral marginal depressions. The eyes of Mecyclothorax argutulus beetles are also less convex, ocular ratio = 1.33, though covering about as much of the ocular lobe as observed in Mecyclothorax argutuloides; ocular lobe ratio for this species = 0.69, ratios of Mecyclothorax argutuloides individuals span 0.68 - 0.73. Setal formula 2 2 2 - 4 2; the single specimen has two dorsal elytral setae on the left elytron, four on the right. Standardized body length 6.1 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
799E729EC7FF6698207C79937DE8565E.taxon	description	Description (n = 1). Head capsule with frontal grooves broad near clypeus, a broad, straight lateral carina extended to anterior supraorbital seta; dorsal surface of neck slightly concave; ocular lobe broadly expanded from gena, outer surface of eye not extended beyond curvature of lobe; horizontal diameter of eye crosses 18 - 19 ommatidia; labral anterior margin broadly, shallowly emarginate; antennae filiform, antennomere 3 sparsely setose; mentum tooth with sides acute, apex pointed. Pronotum slightly transverse, MPW / PL = 1.22, the lateral margins distinctly sinuate before the acute, very projected hind angles, the pronotal base broad so that MPW / BPW = 1.40; median base depressed relative to disc, rugose due to dense wrinkles and punctures; basal margin broadly extended medially between laterobasal depressions; median longitudinal impression fine, shallow, crossed by shallow transverse wrinkles; anterior transverse impression deeply incised, smooth, anterior callosity broadly convex, smooth; front angles well projected, rounded; pronotal apical width subequal to broad basal width, APW / BPW = 0.98; lateral marginal depression broad, margin broadly upraised; laterobasal depression broad, slightly upraised tubercle present. Proepisternum with 6 punctures along hind marginal groove; prosternal process medially concave, margin smoothly upraised. Elytra subquadrate, broadest behind middle, MEW / MPW = 1.38 elytral disc flat, sides moderately sloped; basal groove recurved at tightly rounded humerus, MEW / HuW = 1.83; parascutellar seta present; parascutellar striole deep, smooth, with 5 elongate punctures; sutural interval slightly convex, similar to lateral intervals; sutural and 2 nd striae of subequal depth from base to apex; discal striae 1 - 5 minutely punctate, continuous, stria 6 reduced, very shallow, stria 7 traceable only; 8 th interval laterad 7 th stria of similar convexity to more medial intervals; the 2 paired dorsal elytral setae at 0.24 - 0.27 x and 0.54 - 0.56 x elytral length, extra dorsal setae on the right elytron at 0.45 x and 0.82 x elytral length, setal impressions shallow, spanning about 1 / 2 width of interval 3; apical and subapical setae present; lateral elytral setae arranged in anterior series of 7 setae and a posterior series of 6 setae; elytral marginal depression moderately broad, translucent and lined with sculpticells, margin upturned; subapical sinuation shallow, broad. Mesepisternum with ~ 6 shallow punctures on a glossy surface; metepisternal width to length ratio = 0.89; metepisternum / metepimeron suture a broad, incomplete depression; metathoracic flight wing a triangular flap, 1.7 x long as broad, remnant R and M veins present, the flap extended to hind margin of metanotum. Abdomen with shallow indistinct lateral wrinkles on ventrites 1 - 3; suture between ventrites 2 and 3 complete; apical male ventrite with 2 marginal setae. Legs-metatarsomere 1 / metatibial length ratio = 0.17; metatarsomere 4 length along outer lobe 1.25 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci broad, shallow. Microsculpture of vertex shallow isodiametric sculpticells in transverse rows; pronotal disc with shallow transverse mesh, median base with distinct isodiametric sculpticells; elytral disc with an elongate transverse mesh, the sculpticells more distinct on the elytral apex; metasternum with distinct transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex brunneous with piceous cast; antennomeres 1 - 3 and base of 4 rufoflavous, apical portion with piceous cast; pronotal disc rufopiceous, lateral marginal depression broadly rufoflavous; proepipleuron rufoflavous with piceous cast, proepisternum rufopiceous; elytral disc reflective rufobrunneous with piceous cast, sutural interval concolorous basally, rufoflavous apically; elytral margins narrowly rufoflavous in lateral depression, apex slightly, broadly paler; elytral epipleuron rufoflavous, metepisternum rufopiceous; abdomen rufopiceous, slightly paler laterally, apical half of apical ventrite broadly paler, rufoflavous; metafemur and metatibia rufoflavous. Male genitalia (n = 1). Aedeagal median lobe very robust, distance from parameral articulation to tip 3.6 x depth at midlength (Fig. 82 C); apex narrowly extended beyond ostial opening, tip angled downward, tightly rounded; median lobe curved rightward near apex in ventral view (Fig. 82 D), median shaft very broad, expanded medially, maximum breadth 0.37 x parameral articulation-tip distance; internal sac with very large dorsal ostial microtrichial patch (right patch in Fig. 82 D), plus well-developed ventral ostial microtrichial patch; flagellar plate moderately short, length 0.33 x parameral articulation-tip distance (estimated from shadow of sclerotized plate inside dorsal margin; Fig. 82 C).	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
799E729EC7FF6698207C79937DE8565E.taxon	etymology	Etymology. The adjectival species epithet argutulus is the diminutive of argutus, clear, bright, or sharp (Brown 1956).	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
85C0163BCB5BAC862620F76313C021AB.taxon	diagnosis	Diagnosis. This species (Fig. 87 A) and Mecyclothorax planatus (Fig. 87 B) comprise the third sister-species pair within the Haleakala Mecyclothorax argutor group fauna. Their adelphotaxon status is supported by the broad depressions of the elytra; an anterior depression centered each side on the anterior dorsal elytral seta, and a posterior depression implicating intervals 1 - 5 and lying posterolaterad of the posterior dorsal elytral seta. A second synapomorphy for the pair is the presence of four marginal setae on the male apical abdominal ventrite, a character not observed in any other Hawaiian Mecyclothorax species. Mecyclothorax planipennis can be differentiated from Mecyclothorax planatus by: 1, elytral intervals slightly convex near anterior dorsal elytral seta, not flat as in Mecyclothorax planatus; 2, pronotum tending to be relatively less constricted basally, MPW / BPW = 1.45 - 1.50 versus values of 1.49 - 1.60 in Mecyclothorax planatus, that basal constriction leading to a diagnostic difference in apical versus basal pronotal widths, APW / BPW = 1.02 - 1.05 for Mecyclothorax planipennis versus APW / BPW = 1.06 - 1.15 for Mecyclothorax planatus. Although these species are exceedingly similar in outward appearance, the male aedeagal median lobes differ distinctively. The median lobe tip of Mecyclothorax planipennis males is pointed (Fig. 82 E-H), whereas the median lobe tip of Mecyclothorax planatus males is bluntly rounded (Fig. 82 I-L). Setal formula 2 2 2 2. Standardized body length 5.2 - 5.7 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
85C0163BCB5BAC862620F76313C021AB.taxon	description	Description (n = 5). Head capsule with frontal grooves broad near clypeus, a broad lateral carina posterad to anterior supraorbital seta; dorsal surface of neck flat to convex; eyes moderately convex, ocular lobe obtusely extended from gena, ocular ratio = 1.41 - 1.51, ocular lobe ratio = 0.72 - 0.80; labral anterior margin broadly, shallowly emarginate; antennae filiform, antennomere 3 sparsely setose; mentum tooth with sides acute, apex tightly rounded. Pronotum transverse, MPW / PL = 1.30 - 1.39; hind angle right, margin behind rounded, lateral margin subparallel to slightly convergent for short distance anterad angle; median base depressed relative to disc, covered with rugose wrinkles and punctures; basal margin broadly convex medially; median longitudinal impression fine, shallow; anterior transverse impression broad, bordered anteriorly by depressed to slightly convex anterior callosity, the callosity crossed by numerous, densely packed longitudinal wrinkles; front angles slightly projected, rounded; lateral marginal depression moderately narrow, edge upturned; laterobasal depression broadly concave. Proepisternum with 6 punctures along hind marginal groove; prosternal process medially concave, margins broadly upraised. Elytra subquadrate, disc flattened due to anterior elytral depressions, sides moderately sloped; basal groove evenly curved to subangulate juncture with lateral marginal depression, the lateral marginal depression broader than basal groove, MEW / HuW = 1.83 - 1.92; parascutellar seta present; parascutellar striole shallow, with 6 - 7 punctures; sutural interval more convex than lateral intervals, sutural juncture upraised; sutural and 2 nd striae of subequal depth from base to apex; discal striae 1 - 4 punctate in basal 2 / 3 of length, stria 5 continuous, striae 6 - 7 obsolete laterally, striae 1 - 5 and 7 evident at elytral apex; 8 th interval convexly elevated immediately laterad 7 th stria; 2 dorsal elytral setae at 0.26 - 0.29 x and 0.54 - 0.60 x elytral length, setal impression spanning interval 3; apical and subapical setae present; lateral elytral setae arranged in anterior series of 7 setae and posterior series of 6 setae; elytral marginal depression moderately broad, lined with sculpticells, margin upturned; subapical sinuation shallow, broad. Mesepisternum with ~ 14 punctures in 2 - 3 rows; metepisternal width to length ratio = 0.75; metepisternum / metepimeron suture distinct. Abdomen with 1 - 2 longitudinal, lateral wrinkles on ventrites 1 - 4; suture between ventrites 2 and 3 complete; apical male ventrite with 4 equally spaced marginal setae (a 5 th unilateral seta in one individual), apical female ventrite with 4 equally space marginal setae and median trapezoid of 4 subequal, short setae. Legs-metatarsomere 1 / metatibial length ratio = 0.19; metatarsomere 4 length along outer lobe 1.3 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci narrow, canaliculated, median carina broad. Microsculpture of vertex upraised isodiametric sculpticells in transverse rows; pronotal disc with isodiametric sculpticells in transverse rows, median base with upraised isodiametric mesh; elytral disc with upraised isodiametric sculpticells in transverse rows, apex with upraised isodiametric sculpticells; metasternum with transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex brunneous with piceous cast; antennomeres 1 - 3 rufoflavous, 4 - 11 darker, with piceous cast; pronotal disc rufobrunneous with piceous cast, margins paler, rufoflavous in lateral depression; proepipleuron rufoflavous, proepisternum rufobrunneous with piceous cast; elytral disc reflective rufobrunneous, sutural interval rufous basally, rufoflavous apically; elytral margins narrowly rufoflavous in lateral depression, apex slightly, broadly paler; elytral epipleuron rufoflavous, metepisternum rufopiceous; abdomen uniformly dark, rufopiceous, apical abdominal ventrite very narrowly paler, rufoflavous; metafemur with basal half rufopiceous, apex flavous; metatibia rufobrunneous with piceous cast. Male genitalia (n = 3). Aedeagal median lobe robust, distance from parameral articulation to tip 3.2 x depth at midlength (Fig. 82 E, G-H); apex briefly extended beyond ostial opening, apical face broadly flattened, tip acutely angulate, slightly downturned; median lobe straight in ventral view (Fig. 82 F), right margin indented before rounded tip in this view, left margin slightly curved; internal sac with ventral surface covered with denser microspicules (Fig. 82 G-H), a ventral ostial microtrichial patch either present (Fig. 82 H) or absent (Fig. 82 G) near sac base; flagellar plate large, length 0.56 - 0.61 x parameral articulation-tip distance. Female reproductive tract (n = 1). Bursa copulatrix columnar, elongate, length 0.97 mm, breadth 0.34 mm (Fig. 83 B); bursal walls moderately opaque, thickly wrinkled; gonocoxite 1 with 2 - 3 apical fringe setae and 3 - 4 smaller setae on medial surface (Fig. 84 B); gonocoxite 2 falcate, tip pointed, base broadly extended laterally, 2 stout, apically narrowed lateral ensiform setae, apical nematiform setae on medial surface at 0.73 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
85C0163BCB5BAC862620F76313C021AB.taxon	etymology	Etymology. The species epithet planipennis compounds the Latin planus, or even, flat, with the plural of penna, or wing; therefore planipennis, flat wings. The species epithet is treated as a noun in apposition.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
C9E38541CA8E7897DD0BC7B63E8780A6.taxon	diagnosis	Diagnosis. This is the adelphotaxon to Mecyclothorax planipennis, treated immediately above, with the two species synapomorphically united by the anterior and posterior dorsal elytral depressions (Fig. 87 A-B), and by the males possessing four marginal setae on the apical abdominal ventrite. The pronotal base exhibited by members of this species is narrower relative to maximum pronotal width; MPW / BPW = 1.49 - 1.60, and APW / BPW = 1.06 - 1.15. The discal elytral intervals within the anterior elytral depression are flatter in this species (Fig. 87 B), and the male aedeagal median lobe is bluntly rounded (Figs 82 I-L). Setal formula 2 2 2 2. Standardized body length 5.2 - 5.7 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
C9E38541CA8E7897DD0BC7B63E8780A6.taxon	description	Description (n = 5). As in the taxonomic treatments of other cryptic sibling species pairs in this revision, this description provides only those attributes and measurements that deviate from the description provided above for the sibling species, Mecyclothorax planipennis. Eyes tending to be more convex than in Mecyclothorax planipennis; ocular ratio = 1.47 - 1.53, ocular lobe ratio = 0.72 - 0.80. Pronotum transverse, MPW / PL = 1.30 - 1.37. Elytra subquadrate, but narrower basally than in Mecyclothorax planipennis, MEW / HuW = 1.91 - 2.02; discal strial punctures small, little expanding strial breadth within anterior elytral depression; lateral marginal depression extremely narrow at humerus, the lateral margin depression and basal groove similarly upraised laterad and mesad humeral angle. Male genitalia (n = 5). Aedeagal median lobe robust, distance from parameral articulation to tip 3.2 x depth at midlength (Fig. 82 I, K-L); apex little extended beyond ostial opening, flattened on apical face, tip acutely rounded; median lobe curved rightward near apex, right and left margins subparallel near (Fig. 82 J); internal sac with small dorsal ostial microtrichial patch and larger, heavily spiculated and projected ventral ostial microtrichial patch (Fig. 82 K); flagellar plate very large, length 0.67 x parameral articulation-tip distance. Female reproductive tract (n = 1). Bursa copulatrix columnar, length 0.84 mm, breadth 0.46 mm (Fig. 83 C); bursal walls translucent, thickly wrinkled; gonocoxite 1 with 4 apical fringe setae and 3 smaller setae on medial surface (Fig. 84 C); gonocoxite 2 falcate, base extended laterally, 2 stout, apically narrowed lateral ensiform setae, apical nematiform setae on medial surface at 0.75 x length of gonocoxite.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
C9E38541CA8E7897DD0BC7B63E8780A6.taxon	etymology	Etymology. The adjectival species epithet planatus is derived from the Latin planus, flat or level, and signifies the flattened qualities of beetles of this species.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
2F9430F132390F84290FB41439A25978.taxon	diagnosis	Diagnosis. This species (Fig. 87 C) looks like a smaller bodied, allometrically more conservative version of its adelphotaxon, Mecyclothorax argutulus (Fig. 81 D). Both species share a little transverse, basally broad pronotum; MPW / PL = 1.19 - 1.22, MPW / BPW = 1.38 - 1.40 for this species; values that subsume the ratios derived from the unique Mecyclothorax argutulus holotype. However the body proportions of this species are narrower overall, leading to a more gracile appearance. The pronotal lateral marginal depression is narrow at midlength, and although the elytra are narrower in this species than in its robust relative, the elytra are still broader relative to the pronotum; MEW / MPW = 1.39 - 1.47 versus MEW / MPW = 1.38 in Mecyclothorax argutulus. The elytra tend to have more dorsal elytral setae present, though presence of these setae is also unstable in this species. Of the four specimens available, the setal counts for left plus right elytra are: 4 + 4, 4 + 3, 2 + 3, and 3 + 4; setal formula 2 2 3 - 4 (2) 2. Standardized body length 5.3 - 5.6 mm, versus 6.2 mm for its closest relative.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
2F9430F132390F84290FB41439A25978.taxon	description	Description (n = 4). Head capsule with frontal grooves broad near clypeus, sinuous, broad convex carina to mesad anterior supraorbital seta; dorsal surface of neck slightly concave; eyes small, little convex, ocular lobe obtusely extended from ocular lobe, ocular ratio = 1.36 - 1.42, ocular lobe ratio 0.68 - 0.73; labral anterior margin broadly shallowly emarginate; antennomeres 2 - 3 with sparse pelage of short setae; mentum tooth with sides acute, apex rounded. Pronotum with lateral margin slightly convergent before acute hind angle, the angle acute even with convexly rounded margin behind angle; median base depressed relative to disc, rugose due to elongate wrinkles and punctures; basal margin broadly extended medially between laterobasal depressions, also convexly expanded posteriorly behind depressions to hind angle; median longitudinal impression finely incised, shallow; anterior transverse impression deeply incised, smooth, anterior callosity broadly convex, the callosity smooth with indistinct longitudinal wrinkles; front angles projected, tightly rounded; apical and basal pronotal widths subequal, APW / BPW = 0.96 - 1.02; lateral marginal depression narrow at midlength, broader at front angles, expanded in basal sinuation, margin upturned; laterobasal depression broadly concave, median tubercle present in some individuals. Proepisternum with 6 punctures along hind marginal groove; prosternal process medially concave, margin smoothly upraised. Elytra subquadrate, disc flat, sides moderately sloped; basal groove subangulate at base of sutural stria, extended to tightly angled humerus, the lateral marginal depression broader, its margin more upraised laterad humerus, MEW / HuW 1.75 - 1.85; parascutellar seta present; parascutellar striole deep, with 3 - 4 elongate punctures, smooth between punctures; sutural interval convexity only slightly more convex than lateral intervals, sutural juncture upraised at midlength, depressed basally and apically; sutural and 2 nd striae of subequal depth from base to apex; discal striae 1 - 5 present, the median 4 striae continuous, stria 5 partially interrupted, stria 6 reduced and stria 7 only traceable; discal strial punctures minute, only slightly expanding strial breadth and associated with strial irregularities; 8 th interval laterad 7 th stria of similar convexity to more mesal intervals; up to 4 dorsal elytral setae, at 0.22 - 0.24 x, 0.38 - 0.46 x, 0.52 - 0.52 x, and 0.67 - 0.74 x elytral length when all are present, setal impressions shallow, spanning about 1 / 2 width of interval 3; apical and subapical setae present; lateral elytral setae arranged in anterior series of 7 setae and posterior series of 6 setae; elytral marginal depression moderately broad, lined with sculpticells, margin upturned; subapical sinuation shallow, broad. Mesepisternum with ~ 7 shallow punctures on glossy surface; metepisternal width to length ratio = 0.80; metepisternum / metepimeron suture distinct; the metepisternal ratio is 0.88, and the suture indistinct in the robust adelphotaxon, Mecyclothorax argutulus. Abdomen with shallow indistinct lateral wrinkles on ventrites 1 - 3; suture between ventrites 2 and 3 complete; apical male ventrite with 2 setae, apical female ventrite with 4 equally spaced setae and median trapezoid of 4 subequal, short setae. Legs-metatarsomere 1 / metatibial length ratio = 0.18; metatarsomere 4 length along outer lobe 1.4 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci shallow, broad. Microsculpture of vertex a shallow isodiametric mesh in transverse rows; pronotal disc with shallow transverse mesh, median base with distinct isodiametric sculpticells; elytral disc with elongate transverse mesh, apex with reflective transverse mesh; metasternum with reflective transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse sculpticells. Coloration of vertex rufobrunneous; antennomeres 1 - 3 and base of 4 rufoflavous, balance of antennae with a piceous cast; pronotal disc rufobrunneous with a piceous cast, margins concolorous, depression rufoflavous at front angles; proepipleuron rufoflavous, proepisternum rufobrunneous with piceous cast; elytral disc reflective rufobrunneous, sutural interval concolorous basally, slightly paler apically; elytral margins narrowly rufoflavous in lateral depression, apex broadly paler, rufoflavous; elytral epipleuron rufoflavous, metepisternum rufobrunneous with piceous cast; abdomen rufobrunneous, slightly paler laterally, apical half of apical abdominal ventrite broadly paler, rufoflavous; metafemur rufoflavous; metatibia rufoflavous with brunneous cast. Male genitalia (n = 1). Aedeagal median lobe robust, dorsal margin flat, distance from parameral articulation to tip 3.7 x depth at midlength (Fig. 88 A); apex elongate, with groove extended from ostial opening to downwardly curved, acutely rounded tip; median lobe apex slightly curved rightward in ventral view (Fig. 88 B); median shaft broadly expanded medially, maximum breadth 0.32 x parameral articulation-tip distance; internal sac with large dorsal ostial microtrichial patch (right patch in Fig. 88 B), and smaller ventral ostial microtrichial patch; flagellar plate of moderate size, length 0.33 x parameral articulation-tip distance. Female reproductive tract (n = 1). Bursa copulatrix very elongate, slightly broader in apical half, length 1.37 mm, breadth 0.43 mm (Fig. 83 D); bursal walls moderately opaque, thickly wrinkled; gonocoxite 1 with 3 - 4 apical fringe setae, 2 of the setae larger, the balance smaller (Fig. 84 D), 7 smaller setae on medial surface; gonocoxite 2 narrowly falcate, base broadly, moderately extended laterally, 2 broad lateral ensiform setae (apices worn), apical nematiform setae on medial surface at 0.90 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
2F9430F132390F84290FB41439A25978.taxon	etymology	Etymology. The adjectival species epithet argutuloides modifies the epithet argutulus proposed above for a closely related species, leading to a name that signifies " like the small shining " Mecyclothorax argutulus. Again Perrault's (1984, 1986, 1988, 1989) convention of using a common stem for epithets of closely related species is employed.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
41C9F90325A79510B1F39817A0959EDC.taxon	diagnosis	Diagnosis. This, the most commonly encountered Mecyclothorax species on Haleakala, can be diagnosed by the distinctly punctate elytral striae (Fig. 87 D), little upraised sutural interval, and transverse pronotum with constricted base, MPW / PL = 1.25 - 1.38, MPW / BPW = 1.46 - 1.56. In body proportions this species approaches the Mecyclothorax planipennis - Mecyclothorax planatus species pair, but the cuticular microsculpture of Mecyclothorax cordithorax beetles is much smoother and more transverse, with the pronotal disc, for example, covered with a mixture of shallow transverse mesh and shallow transverse lines. Setal formula 2 2 2 2. Standardized body length 4.4 - 6.2 mm (vast majority of beetles> 4.6 mm length).	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
EC43334412F8C24DB478C4A63C860012.taxon	diagnosis	Diagnosis. With standardized body length 4.5 mm, this is the largest-bodied species of the Mecyclothorax microps group residing across all islands. Like Mecyclothorax minor (Fig. 90 B) and Mecyclothorax angusticollis (Fig. 90 C), the elytra are parallel sided and narrow relative to the pronotum (Fig. 90 A); MEW / MPW = 1.42 for this species. The cuticle is more pallid than observed in Mecyclothorax angusticollis, and the pronotum is more constricted basally than in Mecyclothorax minor. Also, the elytra have a single dorsal elytral seta at the anterior position, resulting in the setal formula 2 2 1 0 unique to this species within the species group.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
EC43334412F8C24DB478C4A63C860012.taxon	description	Description (n = 1). Head capsule with frontal grooves convexly curved from clypeus to mesad anterior supraorbital seta, broad carina laterad convex groove; dorsal impression of neck slightly concave; eyes little convex, ocular lobe very obtusely extended from gena, ocular ratio = 1.31; ocular lobe ratio = 0.73; 10 - 11 ommatidia across horizontal diameter of eye; labral anterior margin angularly emarginate medially, impressed 1 / 7 length of labrum; antennomeres 2 - 3 with sparse pelage of short setae; antennae submoniliform; mentum tooth with sides acute, apex pointed. Pronotum little transverse, MPW / PL = 1.19, base moderately constricted, MPW / BPW = 1.49, with lateral margin straight to sinuate anterad obtuse hind angle; median base depressed relative to disc, covered with minute punctures basally and minute wrinkles at juncture with disc; basal margin slightly convex between laterobasal depressions; median longitudinal impression finely incised, shallow; anterior transverse impression broad, moderately deep laterally, obsolete medially; anterior callosity coplanar with disc medially, crossed by distinct wrinkles; front angles moderately produced, tightly rounded; pronotal apical and basal widths subequal, APW / BPW = 0.99; lateral marginal depression moderately broad with upturned margin before lateral seta, narrow with upraised margin to hind angle; laterobasal depression with irregular surface, continuous with lateral marginal depression. Proepisternum with 5 - 6 minute punctures along hind margin; prosternal process broadly depressed medially. Elytra subparallel, disc medially depressed, sides markedly depressed to margins; basal groove broadly recurved on rounded humeri, MEW / HuW = 1.84; parascutellar seta absent; parascutellar striole discontinuous, with 4 - 5 small punctures; sutural interval coplanar with lateral intervals basally, slightly upraised in apical half; sutural interval deeper and more steeply sided than 2 nd stria throughout most of length, striae of subequal depth at elytral apex; discal striae 1 - 4 shallow, punctate, striae 5 - 6 a series of isolated punctures, stria 7 absent near midlength; discal intervals 2 - 4 flat; 8 th interval laterad 7 th stria of same convexity as more mesal intervals; 1 dorsal elytral seta at 0.23 - 0.26 x elytral length, setal impressions shallow, spanning 1 / 2 width of interval 3; apical and subapical setae absent; lateral elytral setae arranged in anterior series of 7 setae and posterior series of 5 setae; elytral marginal depression moderately broad with margin upraised laterad humerus, beadlike from midlength to subapical sinuation; subapical sinuation shallow, more abruptly incurved anteriorly. Mesepisternum smooth; metepisternal width to length ratio = 0.58; metepisternum / metepimeron suture distinct; metathoracic flight wing length 2.33 x breadth, remnant R and M veins present, apex extended 0.75 x wing remnant's length beyond hind margin of metanotum (all estimated by viewing through elytra). Abdomen with indistinct lateral wrinkles on ventrites 1 - 5; suture between ventrites 2 and 3 reduced laterally, effaced; apical male ventrite with 2 marginal setae. Legs-metatarsomere 1 / metatibial length ratio = 0.20; metatarsomere 4 length along outer lobe 1.5 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci broad, basal tarsomeres medially subcarinate. Microsculpture of vertex an obsolete transverse mesh, sculpticell breadth 2 x length; pronotal disc with obsolete transverse mesh, sculpticell breadth 2 - 4 x length, portions of disc glossy, median base with mixture of isodiametric and transverse sculpticells, breadth 2 x length; elytral disc with shallow isodiametric mesh in transverse rows, apex with same mesh obsolete; metasternum with distinct transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex rufoflavous; antennomere 1 flavous, 2 - 7 rufoflavous (outer antennomeres broken off); pronotal disc and margins rufoflavous; proepipleuron flavous, proepisternum rufoflavous; elytral disc and sutural interval rufoflavous, marginal depression narrowly flavous; elytral apex and apices of intervals 7 - 9 flavous apicad terminus of interval 4; elytral epipleuron flavous dorsally, rufoflavous ventrally, metepisternum rufoflavous; abdomen rufoflavous, apical half of apical ventrite 6 flavous; metafemur rufoflavous; metatibia rufoflavous with brunneous cast. Male genitalia (n = 1). Aedeagal median lobe slender, straight at midlength, shaft abruptly curved ventrally at base and apex (Fig. 91 A); distance from parameral articulation to tip 5.4 x depth at midlength; apex broadly extended beyond ostial opening, tip blunt with dorsal angle rounded and ventral angle denticulate; median lobe abruptly curved rightward ear apex in ventral view (Fig. 91 B), blunt tip appearing concave in this view; internal sac with broad fields of dark microspicules but no apparent macrospicules (assessed in uneverted type specimen); right paramere parallel sided, broad nearly to apex (Fig. 91 A).	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
EC43334412F8C24DB478C4A63C860012.taxon	etymology	Etymology. The Latin adjectival epithet major, i. e. larger, is used for this species in order to contrast it with the following Mecyclothorax minor.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
9898FBFD6011515317F23E75203990D3.taxon	diagnosis	Diagnosis. This species (Fig. 90 B) and Mecyclothorax angusticollis (Fig. 90 C) represent the two species of the Mecyclothorax microps group characterized by: 1, a quadrate, basally broad pronotum, MPW / BPW = 1.30 - 1.35; 2, both anterior and posterior dorsal elytral setae present; and 3, elytra that are narrow relative to the forebody, MEW / MPW = 1.35 - 1.44, MEW / MHW = 1.93 - 2.12. Britton distinguished his Mecyclothorax minor from Mecyclothorax angusticollis based on the paler body color, and the broader pronotal base with non-sinuate lateral margins anterad the obtuse hind angles. Setal formula 2 2 2 0. Standardized body length 3.7 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
AB3AF089FAB47392412E9994EEEA1D38.taxon	diagnosis	Diagnosis. This species is distinguished from all others in the group by the transverse pronotum (Fig. 90 C), MPW / PL = 1.21 - 1.28, that is broad relative to the elytra, MEW / MPW = 1.35 - 1.39. Most similar to Mecyclothorax minor, Mecyclothorax angusticollis can also be distinguished from that species by the distinct isodiametric sculpticells in transverse rows on the elytral disc and apex. Body coloration is darker as well, with the pronotal disc rufobrunneous to rufopiceous, and the elytra rufobrunneous, in some instances with an apical rufopiceous cloud. Setal formula 2 2 2 0. Standardized body length 3.2 - 3.7 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
4362419E4FC5EF0C538A2A107EFE0824.taxon	diagnosis	Diagnosis. This species (Fig. 90 D) plus the following 2 species, Mecyclothorax orbiculus (Fig. 90 E), and Mecyclothorax contractus (Fig. 90 F), comprise the only species of the Mecyclothorax microps group that can be characterized by presence of both supraorbital setae and both pronotal setae, but much reduced elytral setation; no dorsal elytral setae or subapical and apical setae present; base setal formula 2 2 0 0. As the single known specimen of this species unilaterally lacks the anterior supraorbital seta on the right side, this species' setal formula is scored 2 (1) 2 0 0. As in all Mecyclothorax microps group species, the parascutellar seta is also absent. This setal configuration relegates the lateral elytral setae to be the sole setiform sensory organs of the dorsal hindbody. Mecyclothorax xestos exhibits the most reduced elytral striation of the three species, with: 1, only the sutural and 2 nd striae shallow, evident; 2, striae 3 - 4 traceable only as irregular, shallow grooves; 3, striae 5 - 6 very shallow, obsolete and interrupted; and 4, stria 7 absent basally, traceable only near elytral apex. Standardized body length 3.8 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
4362419E4FC5EF0C538A2A107EFE0824.taxon	description	Description (n = 1). Head capsule with frontal grooves narrow, a broad convexity laterad groove and fine carinae mesad eye; dorsal impression of neck broadly, slightly concave; eyes flat externally and small, ocular ratio = 1.28, ocular lobe ratio = 0.67; 9 - 10 ommatidia across horizontal diameter of eye; labral anterior margin angularly emarginate medially 1 / 7 of length; antennae nearly moniliform, antennomeres 2 - 3 with few very short setae; mentum tooth with sides acute, apex tightly rounded. Pronotum slightly transverse, MPW / PL = 1.20, basally constricted, MPW / BPW = 1.52; hind angle right to slightly acute, lateral margin subparallel to convergent for short distance anterad angle; median base depressed relative to disc, covered with longitudinal strigae; basal margin slightly convex between laterobasal depressions; median longitudinal impression finely incised, shallow but evident; anterior transverse impression broad, shallow, obsolete medially; anterior callosity coplanar with disc medially, slightly raised laterally, smooth, glossy; front angles not projected, tightly rounded; apical and basal pronotal widths subequal, APW / BPW = 0.98; lateral marginal depression narrow, edge upraised throughout length, slightly wider at front angle; laterobasal depression with irregular surface, continuous with lateral depression. Proepisternum with numerous small irregularities along hind marginal groove due to microsculpture; prosternal process broadly depressed medially. Elytra broadly subellipsoid, disc flat medially, moderately sloped at sides and apex; basal groove evenly curved anterad to angulate humeral juncture with lateral marginal depression, MEW / HuW = 1.90; parascutellar seta absent; parascutellar striole very shallow, smooth, hard to trace; sutural interval slightly upraised throughout length; discal intervals 2 - 4 flat; 8 th interval laterad 7 th stria more convex than more mesal intervals; elytral marginal depression moderately broad, margin upraised laterad humerus, beadlike from midlength to subapical sinuation; subapical sinuation shallow, more abruptly incurved anteriorly. Mesepisternum with ~ 9 punctures in 2 - 3 rows; metepisternal width to length ratio = 0.68; metepisternum / metepimeron suture distinct. Abdomen with indistinct lateral wrinkles on ventrites 1 - 3, circular depressions on ventrites 3 - 6; suture between ventrites 2 and 3 reduced laterally, effaced; apical female ventrite with 4 equally spaced marginal setae plus median trapezoid of 4 subequal, short setae. Legs-metatarsomere 1 / metatibial length ratio = 0.20; metatarsomere 4 length along outer lobe 1.3 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci broad, basal tarsomeres medially subcarinate. Microsculpture of vertex a very shallow isodiametric and transverse mesh, sculpticell breadth 2 x length mesad eyes; pronotal disc with shallow transverse mesh, sculpticell breadth 2 - 4 x length, median base with mixture of isodiametric and transverse sculpticells, breadth 2 x length; elytral disc with regular transverse mesh, sculpticell breadth 2 x length on disc, 3 - 4 x length laterally, apex with transverse mesh, sculpticell breadth 2 - 3 x length; metasternum with distinct transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex rufous, clypeus rufoflavous; antennomeres 1 - 3 flavous, 4 - 11 rufoflavous; pronotal disc rufous with brunneous cast, lateral margins, base, and apex broadly rufoflavous; proepipleuron flavous, proepisternum rufobrunneous; elytral disc rufobrunneous with reflective sheen, sutural interval rufoflavous basally, flavous api cally; elytral lateral marginal depression and interval 9 rufoflavous, elytral apex and intervals 7 - 9 flavous apicad terminus of interval 4; elytral epipleuron flavous dorsally, rufoflavous ventrally, metepisternum rufobrunneous; abdominal ventrite 1 rufobrunneous, ventrites 2 - 3 medially rufopiceous, laterally rufoflavous, ventrites 4 - 5 and base of 6 rufoflavous, apical half of ventrite 6 flavous; metafemur flavous; metatibia flavous with rufoflavous cast. Female reproductive tract. The lone female specimen was not dissected.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
4362419E4FC5EF0C538A2A107EFE0824.taxon	etymology	Etymology. This species epithet is the Greek xestos, or scraped, planed, smoothed, or polished (Brown 1956).	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
5DCEA90FE67F7149B6EF776C010655D1.taxon	diagnosis	Diagnosis. Among the triplet of Mecyclothorax microps group species without dorsal elytral setae, this species stands out due to the subellipsoid elytra, the lateral margins markedly convex from the narrowed humeri to the apex (Fig. 90 E). The eyes are more convex than in either of the other species in the triplet, with ocular ratio = 1.38; versus 1.28 - 1.33 in the other two species. The elytral microsculpture also differs, being a distinct isodiametric mesh versus transverse mesh to transverse lines in the other two species. Setal formula 2 2 0 0. Standardized body length 4.1 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
5DCEA90FE67F7149B6EF776C010655D1.taxon	description	Description (n = 1). Head capsule with frontal grooves broad, shallow, a broad low convexity laterally before the eye and a very low, rounded carina mesad anterior supraorbital seta; dorsal impression of neck broadly, slightly concave; 14 - 15 ommatidia across horizontal diameter of eye; labral anterior margin angularly emarginate to 1 / 6 of labral length; antennae nearly moniliform, antennomeres 2 - 3 with sparse pelage of short setae; mentum tooth with sides acute, apex pointed. Pronotum slightly transverse, MPW / PL = 1.22, moderately constricted basally, MPW / BPW = 1.49; hind angle obtuse due to convex margin behind angle; lateral margins slightly convergent anterad sharply projected hind angles; median base moderately depressed relative to disc, ~ 14 punctures each side; basal margin slightly convex between laterobasal depressions; median longitudinal impression finely incised, shallow but evident; anterior transverse impression broad, shallow, obsolete medially; anterior callosity co planar with disc medially, slightly raised laterally, smooth, glossy; front angles slightly projected, tightly rounded; apical pronotal width greater than basal width, APW / BPW = 1.05; lateral marginal depression narrow, the edge beadlike to upturned near broader area at front angle; laterobasal depression with irregular surface, continuous with lateral depression. Proepisternum with 5 - 6 minute punctures along hind marginal groove; prosternal process broadly depressed medially. Elytra with narrow humeri, basal groove distinctly curved to meet lateral marginal depression at subangulate humerus, MEW / HuW = 1.91; parascutellar striole shallow, with 4 punctures; sutural interval slightly upraised relative to lateral intervals throughout its length; sutural and 2 nd striae of subequal depth from base to apex; discal striae 1 - 6 progressively shallower, stria 7 traceable though interrupted repeatedly; discal striae 1 - 5 shallowly punctate, minute elongate punctulae in the base of stria 6, all striae smooth and complete on elytral apex; elytral intervals slightly convex medially, nearly flat laterally; 8 th interval laterad 7 th stria subcarinate, more convex than more mesal intervals; lateral elytral setae arranged in anterior series of 7 setae and posterior series of 6 setae; elytral marginal depression broader, margin slightly upraised at humerus, narrow, beadlike from midlength to subapical sinuation; subapical sinuation shallow, more abruptly incurved anteriorly. Mesepisternum with ~ 10 punctures in 2 - 3 rows; metepisternal width to length ratio = 0.72; metepisternum / metepimeron suture distinct. Abdomen with indistinct lateral wrinkles on ventrites 1 - 3 and circular lateral depressions on ventrites 3 - 6; suture between ventrites 2 and 3 reduced laterally, effaced; apical female ventrite with 4 equally spaced setae and median trapezoid of 4 subequal, short setae. Legs-metatarsomere 1 / metatibial length ratio = 0.19; metatarsomere 4 length along outer lobe 1.3 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci broad, basal tarsomeres medially subcarinate. Microsculpture of vertex a shallow isodiametric mesh, transverse sculpticells, breadth 2 x length mesad eyes; pronotal disc with transverse mesh, sculpticell breadth 2 - 4 x length, median base with mixture of isodiametric and transverse sculpticells, breadth 2 x length; elytral apex with mixture of isodiametric and transverse sculpticells in transverse rows; metasternum with distinct transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex rufobrunneous with rufoflavous clypeus; antennomeres 1 - 3 flavous, 4 - 11 rufoflavous; pronotal disc rufous with brunneous cast, lateral margins, base, and apex broadly rufoflavous; proepipleuron rufoflavous, proepisternum rufobrunneous; elytral disc rufobrunneous with indistinct purplish sheen, sutural interval rufoflavous basally, flavous in apical 1 / 3, elytral lateral margin depression, 9 th interval and apex rufoflavous; elytral epipleuron flavous dorsally, rufoflavous ventrally, metepisternum rufobrunneous; abdomen with abdominal ventrite 1 rufobrunneous, ventrites 2 - 3 medially rufopiceous, ventrites 4 - 6 rufoflavous, with apical half of ventrite 6 flavous; metafemur flavous; metatibia flavous with rufoflavous cast. Female reproductive tract. The lone female specimen was not dissected.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
5DCEA90FE67F7149B6EF776C010655D1.taxon	etymology	Etymology. The species epithet orbiculus is the diminutive of orbis, or circle, and signifies the small eyes characteristic of this species.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
CF7C9164651E8B609BBA21A6906463EA.taxon	diagnosis	Diagnosis. This third species of the triplet lacking dorsal elytral setae can be diagnosed by the more quadrate pronotum (Fig. 90 F); MPW / PL = 1.07 - 1.13. The pronotum is relatively less constricted basally in association with the quadrate shape; MPW / BPW = 1.42 - 1.43. The elytra are subquadrate, the lateral margins extended laterally outside the distinctly angulate humeri, and broader relative to the pronotum than in other Mecyclothorax microps group species; MEW / MPW = 1.56 - 1.59. Setal formula 2 2 0 (1) 0; 1 of the 3 specimens has the anterior dorsal elytral seta present. Standardized body length 4.0 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
CF7C9164651E8B609BBA21A6906463EA.taxon	description	Description (n = 3). Head capsule with frontal grooves broad at clypeus, sinuous, with broad convexity laterad groove and thin carina mesad supraorbital seta; dorsal impression of neck slightly concave; ocular lobe little projected, eyes small, little convex, vertical dimension greater than length, ocular ratio = 1.29 - 1.33, ocular lobe ratio = 0.73 - 0.78, about 10 ommatidia across horizontal diameter of eye; labral anterior margin angularly emarginate medially, impressed 1 / 6 length; antennomere 2 with sparse setae at apex, antennomere 3 sparsely setose along shaft; mentum tooth with sides acute, apex pointed. Pronotum with lateral margins broadly sinuate anterad slightly obtuse, projected hind angles, margin behind angle convex; median base depressed relative to disc, covered with punctures and longitudinal strigae; basal margin slightly convex between laterobasal depressions; median longitudinal impression finely incised, shallow but evident, continued onto median base; anterior transverse impression broad, shallow, surface slightly irregular; anterior callosity coplanar with disc medially, slightly raised laterally, smooth, glossy; front angles not projected, rounded; apical and basal pronotal widths subequal, APW / BPW = 0.97 - 1.04; lateral marginal depression broadest inside front angles, narrow with beadlike margins in anterior half, margin upraised basally; laterobasal depression with irregular surface, continuous with lateral depression. Proepisternum with 5 - 6 minute punctures along hind marginal groove; prosternal process broadly depressed medially. Elytra subquadrate, the disc flat medially, moderately sloped on sides and apex; the humeral angles are proximate relative to elytral width, MEW / HuW = 1.95 - 2.0; parascutellar striole with 3 punctures, the striole interrupted between punctures; sutural interval slightly more upraised throughout its length than lateral intervals; sutural striae slightly deeper than 2 nd stria basally, of similar depth apically; discal striae 1 - 4 moderately impressed, minutely punctate, intervals 5 - 6 shallow but evident with elongate irregularities along length, stria 7 obsolete, interrupted; 8 th interval laterad 7 th stria of same convexity as more mesal intervals; 1 dorsal elytral seta present in 1 of 3 specimens at 0.23 - 0.26 x elytral length, setal impression shallow, spanning 1 / 2 width of interval 3; lateral elytral setae arranged in anterior series of 6 (7) setae and posterior series of 5 (6) setae; elytral marginal depression moderately broad with upraised margin outside humerus, beadlike from midlength to subapical sinuation; subapical sinuation shallow, more abruptly incurved anteriorly. Mesepisternum with ~ 7 punctures in 2 rows; metepisternal width to length ratio = 0.88; metepisternum / metepimeron suture distinct; metathoracic flight wing 2.33 x long as broad, with remnant R and M veins, the vestigium extended for 0.75 x its length beyond hind margin of metanotum. Abdomen with indistinct lateral wrinkles on ventrites 1 - 5; suture between ventrites 2 and 3 reduced laterally, effaced; apical female ventrite with 4 equally space marginal setae plus median trapezoid of 4 subequal, short setae. Legs-metatarsomere 1 / metatibial length ratio = 0.19; metatarsomere 4 length along outer lobe 1.5 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci broad, basal tarsomeres medially subcarinate. Microsculpture of vertex very shallow isodiametric and transverse mesh, sculpticell breadth 2 x length mesad eyes; pronotal disc with shallow transverse mesh, sculpticell breadth 2 - 4 x length, median base with mixture of isodiametric and transverse sculpticells, breadth 2 x length, apex with transverse mesh, sculpticell breadth 4 x length; metasternum with distinct transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex rufous; antennomeres 1 - 3 flavous, 4 - 11 rufoflavous; pronotal disc rufous, lateral margins, base, and apex broadly rufoflavous; proepipleuron rufoflavous, proepisternum rufobrunneous; elytral disc rufobrunneous with reflective sheen, sutural interval rufoflavous basally, flavous apically; elytral marginal depression and 9 th interval rufoflavous, intervals 7 - 9 and apex apicad terminus of interval 4 tending toward flavous; elytral epipleuron flavous dorsally, rufoflavous ventrally, metepisternum rufobrunneous; abdominal ventrite 1 and 4 - 6 rufoflavous, ventrites 2 - 3 rufobrunneous, apical half of apical ventrite 6 flavous; metafemur flavous; metatibia flavous with rufoflavous cast. Female reproductive tract (n = 1). Bursa copulatrix columnar with rounded apex, length 0.67 mm, breadth 0.23 mm (Fig. 92 B); bursal walls translucent with thin wrinkles; gonocoxite 1 with 3 small apical fringe setae and 5 - 6 smaller setae on medial surface (Fig. 93 B); gonocoxite 2 narrowly subtriangular, apex rounded, base narrowly extended laterally, 2 lateral ensiform setae with apical seta broader and longer, apical nematiform setae on medial surface at 0.75 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
CF7C9164651E8B609BBA21A6906463EA.taxon	etymology	Etymology. The adjectival species epithet contractus means drawn together, or made narrow, and refers to the basally constricted pronotum of beetles in this species.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
15752D58EB84F8A04ADECDCF0953224D.taxon	diagnosis	Diagnosis. Although known only from two specimens collected by R. C. L. Perkins in 1894 and 1896, new specimens of this species would be instantly recognizable by the presence of only the posterior supraorbital seta, glabrous pronotum with carinate lateral margins, minimally punctate pronotal median base (Fig. 95 A), and very large body size; standardized body length 7.4 mm (paratype female). Setal formula 1 0 2 0, and the parascutellar seta is present. Britton (1948 b) records the body length as 7.2 - 7.8 mm based on his measurements of holotype and paratype females.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
AA0162AC5C9D3B0DE9114025DB7FF539.taxon	diagnosis	Diagnosis. This species (Fig. 95 B) looks much like a smaller version of Mecyclothorax molops (Fig. 95 A), but the pronotal hind angles are projected, distinctly obtuse, and the pronotal lateral margin clearly sinuate anterad the angle. The pronotal median base is more punctate, with 20 or more punctures each side that are isolated by glossy cuticle. The sutural stria is easily followed to the basal groove, though it is smooth near the elytral base. The parascutellar seta is absent. The setal formula is 1 0 2 0; anterior supraorbital seta absent. Among Haleakala species of the group this formula is shared only with Mecyclothorax macrops and Mecyclothorax scarites. At standardized body length 6.1 - 6.4 mm, these beetles are diagnostically smaller than the former, and diagnostically larger than the latter.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
ADA177B06916054F54B8FB1878AA6C3A.taxon	diagnosis	Diagnosis. This species (Fig. 95 C) shares a glabrous pronotum with three other members of the Mecyclothorax scaritoides group: Mecyclothorax molops (Fig. 95 A), Mecyclothorax macrops (Fig. 95 B), and Mecyclothorax scarites (Fig. 95 D). Among them this species deviates by exhibiting two supraorbital setae; setal formula 2 0 2 0. The discal elytral striae are more distinctly punctate in this species, with both the sutural stria and parascutellar striole punctate-in some instances very indistinctly so-to the elytral basal groove. In beetles of the most similar species, Mecyclothorax scarites, the sutural stria is continued to the basal groove, but it is smooth, and the parascutellar striole is little impressed with the punctures taking the form of elongate irregularities. The elytra are also more broadly subellipsoid in this species, with the tightly rounded to subangulate humeri more proximate; MEW / HuW = 2.04 - 2.15 versus values of 1.93 - 2.0 for specimens of Mecyclothorax scarites. Standardized body length 5.0 - 5.8 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
B7F2080CAC8EC5164AE116238F12CFBC.taxon	diagnosis	Diagnosis. This species (Fig. 95 D) is assigned to the setal formula 1 0 2 0 along with two other Haleakala species- Mecyclothorax molops (Fig. 95 A) and Mecyclothorax macrops (Fig. 95 B) - yet it is more similar in pronotal configuration, overall body proportions, and elytral striation to Mecyclothorax scaritoides (Fig. 95 C), a species characterized by presence of both supraorbital setae. In addition to the supraorbital setae, Mecyclothorax scarites can be distinguished from Mecyclothorax scaritoides by the shallow but distinct transverse microsculpture on the elytral disc that is composed of sculpticells 3 - 4 x broad as long plus transverse lines not joined into a mesh. The discal elytral striae are not punctate, with the sutural stria irregularly impressed along its discal portion, and striae 2 - 4 variable impressed among individuals, but smooth to slightly irregular across the disc in all individuals. The elytra are slightly more ovoid than those of Mecyclothorax scaritoides, with the humeral angles more distant relative to overall elytral width; MEW / HuW = 1.93 - 2.0. Standardized body length 5.3 - 5.5 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
B7F2080CAC8EC5164AE116238F12CFBC.taxon	description	Description (n = 5). Head capsule with frontal grooves broad, sinuous, a broad lateral convexity before eye, groove nearly joined to postocular groove anterad posterior supraorbital seta; dorsal impression of neck slightly concave; eyes moderately convex, covering much of ocular lobe, ocular ratio = 1.47 - 1.52, ocular lobe ratio = 0.84 - 0.88; labral anterior margin very shallowly emarginate medially; antennae filiform, antennomeres 1 - 3 glabrous; mentum tooth with sides acute, apex tightly rounded. Pronotum moderately transverse, MPW / PL = 1.20 - 1.22, base moderately constricted, MPW / BPW = 1.47 - 1.55, lateral margin sinuate for 0.1 x pronotal length anterad obtuse hind angles, their apex rounded; median base coplanar with disc medially, ~ 14 small, isolated punctures each side; basal margin moderately convex between hind angles; median longitudinal impression very shallowly incised, traceable to obsolete; anterior transverse impression broad, shallow medially, narrow and finely incised laterally; anterior callosity slightly convex, smooth; front angles slightly projected, tightly rounded; pronotal apical and basal widths subequal, APW / BPW = 0.94 - 1.05; lateral marginal depression very narrow, edge beaded even at front angle, disc very convex; laterobasal depression smooth, convex with U-shaped groove along lateral and basal margins of convexity. Proepisternum with 5 minute punctures along hind marginal groove; prosternal process with very shallow depression medially. Elytra ovoid, greatest width slightly behind midlength; disc very convex, slightly depressed along suture, sides distinctly sloped to near vertical inside marginal depression; basal groove distinctly curved to subangulate humerus; parascutellar seta present; parascutellar striole shallow, with 3 - 4 elongate punctures, smooth between punctures; sutural interval slightly more convex than intervals 2 - 4 throughout their length; sutural stria narrow, moderately deep apically, 2 nd stria obsolete, traceable at apex; 8 th interval laterad 7 th stria of similar convexity to fused apical portion of striae 5 + 7; 2 dorsal elytral setae at 0.19 - 0.23 x and 0.52 - 0.53 x elytral length, setal impressions very small, crossing 1 / 3 of interval 3; lateral elytral setae arranged in anterior series of 7 setae and posterior series of 6 setae; elytral marginal depression very narrow throughout, a low bead present at humerus, a more distinct bead near subapical sinuation; subapical sinuation very shallow, less concave than width of marginal bead. Mesepisternum with ~ 10 punctures in 2 - 3 rows; metepisternal width to length ratio = 0.73; metepisternum / metepimeron suture distinct. Abdomen with irregular lateral wrinkles on ventrites 1 - 6, lateral depressions on ventrites 3 - 6; suture between ventrites 2 and 3 effaced; apical male ventrite with 2 marginal setae and apical female ventrite with 4 equally spaced setae plus median trapezoid of 4 subequal, short setae. Legs-metatarsomere 1 / metatibial length ratio = 0.20; metatarsomere 4 length along outer lobe 1.5 x medial tarsomere length, apical and subapical setae present, the lateral subapical seta short; metatarsal dorsolateral sulci broad, shallow, medially subcarinate on metatarsomere 1 only. Microsculpture of vertex obsolete transverse mesh, sculpticell breadth 2 x length, surface glossy; pronotal disc with obsolete transverse mesh, sculpticell breadth 3 - 4 x length, most parts glossy, median base glossy, indistinct transverse cells laterally; elytral apex with shallow transverse mesh, sculpticell breadth 2 - 3 x length; metasternum with obsolete transverse mesh, glossy; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex rufous with a piceous cast; antennomeres 1 - 3 rufoflavous, 4 - 11 rufobrunneous; pronotal disc rufous, concolorous basally and later ally, apex slightly paler; proepipleuron pale rufobrunneous, proepisternum rufobrunneous; elytral disc rufous, sutural interval concolorous basally, rufoflavous apically; elytral marginal depression and 9 th interval rufoflavous apically; elytral epipleuron pale rufobrunneous, metepisternum rufobrunneous; abdomen rufobrunneous, ventrites 1 - 3 rufoflavous laterally, apical 1 / 6 of apical ventrite flavous; metafemur rufoflavous; metatibia rufoflavous with brunneous cast. Male genitalia (n = 2). Aedeagal median lobe slender, distance from parameral articulation to tip 5.0 x depth at midlength (Fig. 96 D); apex distinctly expanded dorsally and broadly expanded ventrally producing an apical knob, the outline of the tip broadly rounded; internal sac elongate, sac length from ventral margin of ostial opening to base of flagellar plate 0.78 x parameral articulation-tip distance, without ornamentation (Fig. 96 E); flagellar plate very short, nearly as broad as long, length 0.24 x parameral articulation-tip distance. Female reproductive tract (n = 1). Bursa copulatrix columnar with narrower apex, broader base, length 0.91 mm, apical breadth 0.23 mm, basal breadth 0.41 mm (Fig. 97 C); bursal walls translucent, thinly wrinkled; gonocoxite 1 with 3 - 4 apical fringe setae (Fig. 98 C), a thick, curved seta at medioapical angle and 10 - 11 smaller setae along medial surface; gonocoxite 2 falcate with acuminate tip, base broadly extended laterally with curved terminus, 2 broad lateral ensiform setae, apical nematiform setae on medioventral surface at 0.66 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
B7F2080CAC8EC5164AE116238F12CFBC.taxon	etymology	Etymology. The cylindrical body shape of Mecyclothorax scaritoides appears to be the basis for Blackburn (1878 b) using a species epithet derived from the Greek skaritis, that stem used by Fabricius (1775) to name the large-bodied Scarites beetles of the Holarctic. This species is given the species epithet scarites to complete the cycle with the generic name Scarites F.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
C00D3371A039A5DC97119203BF1577BA.taxon	diagnosis	Diagnosis. Of the five Haleakala species of this group with the lateral pronotal seta present (Fig. 100), this species (Fig. 100 A) can be diagnosed by the elongate convergence of the pronotal lateral margins anterad the acute hind angles. Like Mecyclothorax crassuloides (Fig. 100 B) and Mecyclothorax crassulus (Fig. 100 C), all discal striae save the sutural are reduced so much that they can at best be traced by very shallow serial punctures; the 8 th interval remains well developed, deep and broad along the lateral margin of the elytron. This species can be told from the above two by the relatively broader pronotal base; MPW / BPW = 1.63, versus values of 1.67 - 1.79. Setal formula 2 1 2 0. Standardized body length 4.9 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
C00D3371A039A5DC97119203BF1577BA.taxon	description	Description (n = 1). Head capsule with frontal grooves deep, directly divergent from clypeus to mesad anterior supraorbital seta, broad convexity laterally; dorsal impression of neck slightly concave; eyes large, moderately convex, ocular ratio = 1.54, ocular lobe ratio 0.85; labral anterior margin angularly emarginate medially to 1 / 6 of length; antennae filiform, antennomeres 2 - 3 with sparse pelage of short setae; mentum tooth with sides acute, apex tightly rounded. Pronotum slightly transverse, MPW / PL = 1.16; lateral margins convergent for 0.2 x pronotal length anterad hind angles; median base depressed relative to disc, ~ 20 small, isolated punctures each side; basal margin straight medially, margin anteriorly curved laterad laterobasal depressions; median longitudinal impression very finely incised within slight depression of disc, crossed by indistinct transverse wrinkles; anterior transverse impression very shallow, broad, obsolete medially, smooth but crossed by minute longitudinal wrinkles; anterior callosity slightly convex, smooth, glossy; front angles moderately produced, tightly rounded; pronotal apical width slightly greater than basal width, APW / BPW = 1.05; lateral marginal depression narrow, edge upturned, slightly broader at front angle; laterobasal depression smooth, glossy surface continuous with lateral depression. Proepisternum with 5 minute punctures along hind marginal groove; prosternal process narrow, evenly depressed medially. Elytra ovoid, disc convex overall, slightly depressed along suture, distinctly sloped laterally to marginal depression; basal groove briefly, angularly curved to subangulate humerus, MEW / HuW = 2.42; parascutellar seta present; parascutellar striole brief with 3 - 4 punctures, deep and continuous between punctures; sutural interval convex throughout length, but crossing depressed medial portion of disc; discal intervals 2 - 4 and beyond laterally flat, difficult to demark; sutural stria continuous from base to apex, minutely punctate basally, smooth and deeper apically; 8 th interval laterad 7 th stria slightly more convex than fused apical portion of striae 5 + 7; 2 dorsal elytral setae at 0.20 x and 0.45 - 0.48 x elytral length, setal impressions shallow, spanning 1 / 2 width of interval 3; lateral elytral setae arranged in anterior series of 7 setae and posterior series of 6 setae; elytral marginal depression broadest at humeral angle, margin upturned in basal half, beaded near subapical sinuation; subapical sinuation shallow, more abruptly incurved anteriorly. Mesepisternum with ~ 10 large punctures in 2 - 3 rows; metepisternal width to length ratio = 0.68; metepisternum / metepimeron suture distinct. Abdomen with irregular lateral wrinkles on ventrites 1 - 5, lateral depressions on ventrites 3 - 6; suture between ventrites 2 and 3 partially effaced; apical male ventrite with 2 marginal setae. Legs-metatarsomere 1 / metatibial length ratio = 0.20; metatarsomere 4 length along outer lobe 1.3 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci broad, shallow, basal tarsomere medially subcarinate. Microsculpture of vertex a transverse mesh, sculpticell breadth 2 - 3 x length; pronotal disc with shallow transverse mesh to transverse lines, the surface glossy in part, median base glossy, with indistinct transverse sculpticells laterally; elytral disc with shallow transverse mesh, sculpticell breadth 2 x length, apex with such a mesh mixed with isodiametric sculpticells; metasternum with transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex rufous with a piceous cast; antennomeres 1 - 3 rufoflavous, 4 - 11 rufobrunneous; pronotal disc and margins rufous; proepipleuron rufoflavous, proepisternum rufoflavous dorsally, rufobrunneous ventrally; elytral disc and margins rufous, sutural interval concolorous basally, rufoflavous apically; elytral apex gradually paler, rufoflavous apicad subapical sinuation; elytral epipleuron and metepisternum rufoflavous; abdominal ventrites 1 - 3 rufobrunneous, ventrites 4 - 6 rufoflavous; metafemur rufoflavous; metatibia rufoflavous with brunneous cast. Male genitalia (n = 1). Aedeagal median lobe gracile, distance from parameral articulation to tip 4.6 x depth at midlength (Fig. 96 F); apex narrowly and sinuously extended 3 x its depth beyond ostial opening, the tip narrowly rounded; internal sac broad, with projected, pineapplelike lobe bearing sclerotized macrospicules at midlength on dorsal surface, and dense, brownish microspicules covering ventral surface; flagellar plate short, length 0.33 x parameral articulation-tip distance.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
C00D3371A039A5DC97119203BF1577BA.taxon	etymology	Etymology. This species honors the many entomological contributions of Phillip H. Timberlake, including the collection of the only known specimen of this species in 1919. Mr. Timberlake specialized in Hymenoptera, especially bees, and was an associate entomologist with the Hawaiian Sugar Planter's Association from 1914 - 1924 before completing his long and distinguished career at the Citrus Experiment Station, Riverside, CA (Hurd et al. 1982).	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
37DA52E25DCE3293735808C6A36FD65B.taxon	diagnosis	Diagnosis. This species (Fig. 100 B) and Mecyclothorax crassulus (Fig. 100 C) are cryptic, sibling species, and based on synapomorphous configurations of the male aedeagal median lobes (Fig. 96 G-J), they are sister species. They can be diagnosed externally by microsculpture only, with this species characterized by the elytra bearing indistinct isodiametric sculpticells in transverse rows on the disc, those sculpticells visible only outside any areas of reflected microscope light. On the elytral apex, the isodiametric sculpticells are more visible, but they are flat without upraised centers. The male aedeagal median lobe is much more robust in this species, broader dorsoventrally with a broader apex (Fig. 96 I). The aedeagal internal sac is also larger, in keeping with the lobe dimensions, and more heavily spiculated. Setal formula 2 1 2 0. Standardized body length 4.5 - 5.1 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
37DA52E25DCE3293735808C6A36FD65B.taxon	description	Description (n = 5). Head capsule with frontal grooves moderately deep, broad, the convexity laterad groove broad and low, groove terminated at short carina mesad anterior supraorbital seta; dorsal impression of neck slightly concave; eyes convex, ocular ratio = 1.54 - 1.58, moderately large, ocular lobe ratio = 0.74 - 0.80; labral anterior margin angularly emarginate 1 / 8 length; antennae filiform, antennomeres 2 - 3 with sparse pelage of short setae; mentum tooth with sides acute, apex tightly rounded. Pronotum moderately transverse, MPW / PL = 1.20 - 1.30, distinctly constricted basally, MPW / BPW = 1.67 - 1.80; hind angle acute, lateral margin subparallel for 0.1 x pronotal length; median base depressed relative to disc, ~ 13 small, isolated punctures each side; basal margin straight, extended posteriorly near hind angles; median longitudinal impression very finely incised, within slight depression of disc, crossed by indistinct transverse wrinkles; anterior transverse impression broad, deep, may be crossed by distinct longitudinal wrinkles; anterior callosity slightly convex, smooth or with minute longitudinal wrinkles; front angles slightly produced, tightly rounded; apical pronotal width much greater than basal width, APW / BPW = 1.12 - 1.20; lateral marginal depression narrow, edge upturned, slightly broader at front angle; laterobasal depression with slightly irregular surface as on median base, continuous with lateral depression. Proepisternum with 5 minute punctures along hind marginal groove; prosternal process narrowly, evenly depressed medially. Elytra distinctly ovoid, widest behind midlength; disc slightly depressed along suture, sides distinctly sloped to marginal depression; basal groove briefly curved to subangulate humerus, the humeral angles proximate relative to elytral width, MEW / HuW = 2.31 - 2.43; parascutellar seta present; parascutellar striole brief, with 3 - 4 punctures, deep, continuous between punctures; sutural interval convex throughout length, disc depressed each side at sutural stria; discal striae 2 - 4 obsolete, partially traceable by very small punctulae or by cuticular pigment dots associate with strial development, lateral striae 5 - 7 absent (again their course visible only by serial dots of pigment associated with strial development); both 8 th interval laterad 7 th stria and fused apical portion of striae 5 + 7 slightly, broadly convex; 2 dorsal elytral setae at 0.26 x and 0.54 - 0.65 x elytral length, setal impressions very small, shallow, spanning 1 / 4 of interval 3; lateral elytral setae arranged in anterior series of 7 setae and posterior series of 6 setae; elytral marginal depression broadest outside humerus, margin upturned in basal half, beaded near subapical sinuation; subapical sinuation shallow, more abruptly incurved anteriorly. Mesepisternum with ~ 10 large punctures in 2 - 3 rows; metepisternal width to length ratio = 0.63; metepisternum / metepimeron suture distinct. Abdomen with irregular lateral wrinkles on ventrites 1 - 5, lateral depressions on ventrites 3 - 6; suture between ventrites 2 and 3 partially effaced; apical male ventrite with 2 marginal setae and apical female ventrite with 4 equally spaced setae and median trapezoid of 4 subequal, short setae. Legs-metatarsomere 1 / metatibial length ratio = 0.20; metatarsomere 4 length along outer lobe 1.3 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci broad, shallow, basal tarsomere medially subcarinate. Microsculpture of vertex a transverse mesh, sculpticell breadth 2 - 3 x length; pronotal disc with shallow transverse mesh to transverse lines, glossy in part, median base glossy with indistinct transverse sculpticells laterally; metasternum with transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex rufous with a piceous cast; antennomere 1 rufoflavous, 2 - 11 rufobrunneous in fully melanized individuals, antennomeres 1 - 3 rufoflavous in less melanized beetles; pronotal disc rufous, lateral margins concolorous, base and apex rufopiceous; proepipleuron rufoflavous, proepisternum rufoflavous dorsally, rufobrunneous ventrally; elytral disc rufous with a dark brunneous cast, sutural interval rufous basally, rufoflavous apically; elytral interval 9 rufopiceous, marginal depression rufoflavous; elytral apex rufoflavous only on interval 8 and along apical margin; elytral epipleuron flavous, metepisternum rufoflavous; abdomen rufoflavous; metafemur flavous with smoky brunneous apex; metatibia rufobrunneous with piceous cast. Male genitalia (n = 2). Aedeagal median lobe very broad basally, attenuated apically with sinuous ventral surface, distance from parameral articulation to tip 2.4 x basal depth (Fig. 96 G, I), apex thick at distal end of ostial opening, sinuously narrowed to acutely pointed tip; base of median lobe nearly symmetrical in ventral outline (Fig. 96 H) but left margin broadly curved 90 ° rightward before bluntly rounded tip in this view; internal sac with broadly convex dorsal lobe near base of sac, a suggested dorsal ostial microtrichial patch on right side at base, and ventral surface broadly covered with dark macro- and microspicules (Fig. 96 I); flagellar plate elongate, length 0.50 x parameral articulation-tip distance. Female reproductive tract (n = 1). Bursa copulatrix columnar with rounded apex, broader base, length 1.0 mm, apical breadth 0.39 mm, basal breadth 0.48 mm (Fig. 97 D); bursal walls translucent with shagreened surface and very fine wrinkles; gonocoxite 1 with 2 - 4 apical fringe setae, a thick, curved seta at medioapical angle and 6 - 8 smaller setae along medial surface (Fig. 98 D); gonocoxite 2 subtriangular with subacuminate tip, 2 lateral ensiform setae, apical nematiform setae on medioventral surface at 0.78 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
37DA52E25DCE3293735808C6A36FD65B.taxon	etymology	Etymology. The adjectival species epithet crassuloides is derived from the Latin crassus; thick, heavy (Jaeger 1955). This species and Mecyclothorax crassulus below are named to follow Mecyclothorax crassus of West Maui (Sharp 1903), a closely related species. As crassulus is the diminutive of crassus, then the epithet crassuloides used here means resembling a diminutive crassus.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
CE13A9CAFDFBFE8C227D7A1BF3C21593.taxon	diagnosis	Diagnosis. This adelphotaxon (Fig. 100 C) to Mecyclothorax crassuloides (Fig. 100 B) can be diagnosed externally by the elytral microsculpture, with the discal intervals covered with a shallow isodiametric mesh, and the lateral intervals with more transverse sculpticells, the microsculpture of all intervals visible in areas of reflected microscope light. The elytral apex bears a distinctly isodiametric mesh, with the sculpticell centers upraised, as opposed to the flat tiling of the isodiametric sculpticells on the elytral apex of Mecyclothorax crassuloides. As for this species' sister taxon, the male aedeagal median lobe provides a certain diagnosis, with the lobe of Mecyclothorax crassulus more gracile throughout its length, and the apex narrower (Fig. 96 J). The internal sac differs also, beings smaller and bearing less melanized spicules. Setal formula 2 1 2 0. Standardized body length 4.6 - 4.9 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
CE13A9CAFDFBFE8C227D7A1BF3C21593.taxon	description	Description (n = 5). As the microsculptural characters above are the only reliable external anatomical characters yet found to diagnoses these two species, the description of Mecyclothorax crassuloides can serve for this species as well, allowing the following substitutions involving non-diagnostic mensural and qualitative characters. Head capsule with eyes large, variably convex, ocular ratio = 1.47 - 1.62, ocular lobe ratio 0.77 - 0.85; labral anterior margin nearly straight; mentum tooth with sides right, apex rounded. Pronotum moderately transverse, MPW / PL = 1.22 - 1.31, distinctly constricted basally, MPW / BPW = 1.67 - 1.79; hind angle slightly obtuse, lateral margin subparallel for 0.12 x pronotal length; median base depressed relative to disc, with ~ 16 small isolated punctures each side; basal margin slightly convex between hind angles, not extended posteriorly near angles; pronotal apical width nondiagnostically narrower relative to basal width in this species versus Mecyclothorax crassulus, APW / BPW = 1.08 - 1.17. Elytra tending to be narrower (Fig. 100 C), more subellipsoid than in Mecyclothorax crassulus (Fig. 100 B), MEW / HuW = 2.17 - 2.39; discal striae 2 - 4 often more evident, traceable as interrupted linear series of very small punctures. Coloration of available specimens extremely variable, ranging from sclerotized but not fully melanized individuals (as in Fig. 100 C), to darker, more melanic individuals (e. g. Mecyclothorax crassuloides of Fig. 100 B). Darker specimens share the smoky brunneous femoral apex on a flavous femoral base observed in specimens of Mecyclothorax crassuloides. Male genitalia (n = 2). Aedeagal median lobe robust, broadest basally, distance from parameral articulation to tip 2.5 x basal depth (Fig. 96 J), apex elongate, sinuously extended 5 x its depth to obliquely expanded tip with blunt dorsal projection, flattened apical face, and acutely rounded tip (median lobe appears much like a more slender version of that seen in Mecyclothorax crassuloides, Fig, 96 I); internal sac with distinct field of macrospicules on right surface near base, otherwise covered with fine microspicules; flagellar plate very large, length 0.67 x parameral articulation-tip distance. Female reproductive tract (n = 1). Bursa copulatrix columnar with rounded apex, length 0.61 mm, breadth 0.30 mm, bursal walls translucent, thickly wrinkled (Fig. 97 E); gonocoxite 1 with 2 apical fringe setae, the medial seta smaller, and 7 smaller setae along medial surface (Fig. 98 E); gonocoxite 2 subtriangular with subacuminate tip, 2 lateral ensiform setae, apical nematiform setae on medioventral surface at 0.77 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
CE13A9CAFDFBFE8C227D7A1BF3C21593.taxon	etymology	Etymology. This adjectival species epithet crassulus lies intermediate along the Mecyclothorax crassus - Mecyclothorax crassuloides linguistic cline. The adjectival crassulus is the diminutive of crassus - thick, heavy-used by Sharp (1903) for Mecyclothorax crassus of West Maui (Liebherr 2011).	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
1AAF713F3353808B14129C8140668802.taxon	diagnosis	Diagnosis. This is by far the smallest-bodied Haleakala species in the Mecyclothorax scaritoides group; standardized body length 3.8 mm. The pronotum is basally constricted (Fig. 100 D), MEW / BPW = 1.59, and the elytra are ellipsoid. These body proportions and size are similar to those of the O'ahu Mecyclothorax scaritoides group species, Mecyclothorax simiolus (Blackburn) and Mecyclothorax pelops Liebherr (Liebherr 2009 a). However, beetles of the O'ahu species lack pronotal setae, thereby fitting setal formula 2 0 2 0, and Mecyclothorax gracilicollis is characterized by a bisetose pronotum and therefore this species matches the 2 1 2 0 setal formula of the other Haleakala species in the group.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
1AAF713F3353808B14129C8140668802.taxon	description	Description (n = 1). Head capsule with frontal grooves broad near clypeus, only slightly sinuous, a broad convexity laterad groove before eye, and low carina mesad anterior supraorbital seta; dorsal impression impression of neck slightly concave concave; eyes little convex, ocular ratio = 1.39, but covering much of ocular lobe, ocular lobe ratio = 0.80; labral anterior margin angularly emarginate 1 / 6 labral length; antennae filiform, antennomere 2 with 1 short seta on shaft, antennomere 3 with 2 such setae; mentum tooth with sides acute, apex tightly rounded. Pronotum little transverse, MPW / PL = 1.19; hind angle sharp, obtuse due to curved basal margin inside hind angle, the lateral margin subparallel anterad angle; median base slightly depressed, ~ 18 small punctures each side; basal margin moderately convex between hind angles; median longitudinal impression shallow, narrowly defined, crossed by indistinct transverse wrinkles; anterior transverse impression very shallow, broad, obsolete medially, smooth; anterior callosity slightly convex, smooth, glossy; front angles slightly projected, tightly rounded; pronotal anterior width greater than basal width, APW / BPW = 1.05; lateral marginal depression narrow, edge upturned throughout length except beaded at front angle where depression is slightly broader; laterobasal depression with irregular surface continued from median base. Proepisternum with 5 minute punctures along hind marginal groove; prosternal process narrowly, evenly depressed medially. Elytra with disc slightly convex medially, side distinctly sloped to margin; basal groove curved to subangulate humerus, humeral angle defined by hitch in depression caused by juncture of narrow basal groove and broader marginal depression, MEW / HuW = 2. 17; parascutellar seta present; parascutellar striole with 4 - 5 punctures, deep, continuous between punctures; sutural interval more convex than slightly convex interval 2, sutural juncture upraised; sutural stria shallow, broad, minutely punctate basally, finely incised, deep, and smooth apically; striae 2 - 3 very shallow, striae 4 - 6 traceable and stria 7 obsolete on disc, apically striae 2 - 4 and 7 very shallow, incomplete, striae 5 - 6 obsolete; elytral intervals 2 - 3 slightly convex on disc, intervals 4 - 5 flat; 8 th interval laterad 7 th stria slightly more convex than fused apical portion of striae 5 + 7; 2 dorsal elytral setae at 0.29 x and 0.56 x elytral length, setal impressions very small, spanning 1 / 2 width of interval 3; lateral elytral setae arranged in anterior series of 7 setae and posterior series of 4 - 5 setae; elytral marginal depression broader laterad humerus to midlength, beadlike near subapical sinuation; subapical sinuation evident, more abruptly incurved anteriorly. Mesepisternum with ~ 10 punctures in 2 - 3 rows; metepisternal width to length ratio = 0.87; metepisternum / metepimeron suture distinct. Abdomen with irregular lateral wrinkles in ventrites 1 - 3, circular lateral depressions in ventrites 3 - 6; suture between ventrites 2 and 3 partially effaced; apical female ventrite with 4 equally spaced setae plus median trapezoid of 4 subequal, short setae. Legs-metatarsomere 1 / metatibial length ratio = 0.19; metatarsomere 4 length along outer lobe 1.3 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci shallow, broad, median area broad. Microsculpture of vertex very shallow, transverse, sculpticell breadth 2 - 3 x length; pronotal disc with obsolete transverse lines in part, glossy between sculpticells, median base glossy, indistinct transverse cells laterally; elytral disc with regular, distinct transverse mesh, sculpticell breadth 2 - 4 x length, apex with shallow transverse mesh, sculpticell breadth 2 - 4 x length; metasternum with shallow transverse mesh, sculpticell breadth 3 - 4 x length; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex rufobrunneous; antennomeres 1 - 3 rufoflavous, 4 - 11 rufobrunneous; pronotal disc rufobrunneous, lateral margins, base, and apex narrowly rufoflavous; proepipleuron rufoflavous, proepisternum rufobrunneous; elytral disc rufobrunneous with iridescent sheen, sutural interval rufoflavous basally and apically, rufous on disc; elytral lateral marginal depression narrowly rufoflavous, apex rufoflavous apicad subapical sinuation; elytral epipleuron rufoflavous, metepisternum rufobrunneous; abdominal ventrites 1 - 6 rufobrunneous, ventrites 2 - 3 slightly paler laterally, apical 1 / 3 of apical ventrite flavous; metafemur rufoflavous; metatibia rufobrunneous. Female reproductive tract. The lone female specimen was not dissected.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
1AAF713F3353808B14129C8140668802.taxon	etymology	Etymology. This species epithet combines the Latin adjective gracilis, slender or thin, with the Latin noun collis, or hill. The noun gracilicollis is used here to signify the basally constricted pronotum of beetles of this species.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
D06086B629D200A8DC6362C3E425DFBE.taxon	diagnosis	Diagnosis. Most similar (Fig. 100 E) to the preceding Mecyclothorax gracilicollis (Fig. 100 D) in elytral striation but larger bodied; standardized body length 4.75 mm. Also differing in the more broadly based pronotum-MPW / BPW = 1.51 - and the glossy elytra with only patches of transverse-mesh microsculpture, sculpticell breadth 2 - 3 x length, between the glossy portions. The unique holotype is slightly teneral-the left elytron failed to fully inflate-but the legs exhibit the smoky brunneous femoral apex that is shared with Mecyclothorax crassuloides (Fig. 100 B) and Mecyclothorax crassulus (Fig. 100 C). Setal formula 2 1 2 0.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
D06086B629D200A8DC6362C3E425DFBE.taxon	description	Description (n = 1). Head capsule with frontal grooves deep and broad near clypeus, sinuously curved toward eye and terminated mesad a thin, low carina at anterior supraorbital seta; dorsal impression of neck slightly concave; eyes moderately convex, ocular ratio = 1.47, large, covering much of ocular lobe, ocular lobe ratio = 0.88; labral anterior margin angularly emarginate medially to 1 / 8 of labral length; antennae filiform, antennomeres 2 - 3 with sparse pelage of short setae; mentum tooth with sides acute, apex rounded. Pronotum appearing longer than broad, but MPW / PL = 1.18; hind angle right, margin slightly rounded behind, lateral margin subparallel for 0.1 x length before angle; median base slightly depressed relative to disc, ~ 8 isolated punctures each side and minute longitudinal wrinkles along margin; basal margin slightly convex between laterobasal depressions; median longitudinal impression very finely incised, situated in slight depression of disc, crossed by indistinct longitudinal wrinkles; anterior transverse impression broad, moderately deep, smooth, narrowly incised at front angles; anterior callosity slightly convex, smooth, glossy; front angles slightly produced, rounded; apical and basal pronotal widths equal; lateral marginal depression very narrow, edge beadlike, slightly broader at front angle; laterobasal depression with slightly irregular surface, broadest apically and narrowed to hind angle. Proepisternum with 5 minute punctures along hind marginal groove; prosternal process narrowly medially depressed. Elytra subquadrate to slightly ovoid (left elytron slightly deformed), disc slightly depressed along suture, sides distinctly sloped to lateral marginal depression; basal groove distinctly curved to angulate humerus at juncture with lateral marginal depression, MEW / HuW = 2.17; parascutellar seta present; parascutellar striole with 4 punctures, deep, continuous; sutural interval more convex than 2 nd interval throughout length, disc depressed at sutural interval; sutural stria broad, shallow, and smooth basally, deeper and more finely incised apically; discal striae 2 - 6 very shallow but traceable, associate intervals very slightly convex; 8 th interval laterad 7 th stria slightly more convex than apical fused portion of striae 5 + 7; 2 dorsal elytral setae at 0.30 x and 0.59 x elytral length, setal impressions very small, spanning 1 / 2 width of interval 3; lateral elytral setae arranged in anterior series of 7 setae and posterior series of 5 - 6 setae; elytral marginal depression broadest at humeral angle, margin upturned in basal half, beaded toward subapical sinuation; subapical sinuation deep, abruptly incurved anteriorly. Mesepisternum with ~ 9 punctures in 1 - 2 rows; metepisternal width to length ratio = 0.7; metepisternum / metepimeron suture distinct. Abdomen with irregular lateral wrinkles in ventrites 1 - 5, lateral depressions in ventrites 3 - 6; suture between ventrites 2 and 3 partially effaced; apical female ventrite with 4 equally spaced setae and median trapezoid of 4 subequal, short setae. Legs-metatarsomere 1 / metatibial length ratio = 0.19; metatarsomere 4 length along outer lobe 1.3 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci broad, shallow, median area subcarinate. Microsculpture of vertex an obsolete transverse mesh, sculpticell breadth 2 x length, surface glossy; pronotal disc with obsolete microsculpture, surface glossy, median base glossy with indistinct transverse sculpticells laterally; elytral apex with shallow transverse mesh, sculpticell breadth 2 - 3 x length; metasternum with shallow transverse mesh, sculpticell breadth 3 - 4 x length; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex rufous with piceous cast (based on estimate from teneral type); antennomere 1 rufoflavous, antennomeres 2 - 3 rufobrunneous, 4 - 11 with piceous cast; pronotal disc rufous to rufobrunneous, lateral margins concolorous, base and apex rufoflavous; proepipleuron rufoflavous, proepisternum rufoflavous dorsally, rufobrunneous ventrally; elytral disc rufous with brunneous cast, sutural interval rufous basally, rufoflavous apically; elytral interval 9 rufopiceous, marginal depression rufoflavous; elytral apex rufoflavous only on interval 8 and along apical margin; elytral epipleuron flavous, metepisternum rufoflavous; abdominal ventrites 1 - 3 rufobrunneous, 4 - 6 rufoflavous, apical half of apical ventrite 6 flavous; metafemur flavous with smoky brunneous base and apex; metatibia rufobrunneous with piceous cast. Female reproductive tract. The lone female specimen was not dissected.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
D06086B629D200A8DC6362C3E425DFBE.taxon	etymology	Etymology. The adjectival species epithet dispar means different, or unequal in Latin (Brown 1956), and is used here to represent the unusual body proportions that characterize this species.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
B50E3A33BA1501FD28CB299A1855279A.taxon	diagnosis	Diagnosis. The quadrisetose pronotum with deep transverse wrinkles crossing the disc and with the median base distinctly rugose (Fig. 102 A) can diagnose this species from others in the group. The elytra are ovoid with narrow humeri, but the humeri are not as narrow relative to maximum elytral width as in the other two most similar species, Mecyclothorax reiteratus (Fig. 102 B) and Mecyclothorax splendidus (Fig. 102 C); MEW / HuW = 2.23 - 2.38 for this species versus values of 2.39 - 2.56 for measured individuals of the other two species. If the aedeagal median lobe projects even slightly from the abdominal apex of a male specimen, the broadened apex with dorsal projection (Fig. 103) can diagnose the specimen from those representing Mecyclothorax reiteratus (Fig. 107) or Mecyclothorax splendidus (Fig. 108). Setal formula 2 2 2 1 [sae]. Standardized body length 4.7 - 6.0 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
273C4955C2024376E82D1D37677CB9CE.taxon	diagnosis	Diagnosis. This species (Fig. 102 B) and Mecyclothorax splendidus (Fig. 102 C) represent the only species in the group characterized by a bisetose pronotum, the lateral seta present and hind angles glabrous. In beetles of these species both supraorbital setae are present, and in this species both apical and subapical setae are absent; setal formula 2 1 2 0. The discal elytral intervals 2 - 4 are flat to even slightly concave, in agreement with Mecyclothorax iteratus but diagnostically different from Mecyclothorax splendidus, which exhibits slightly convex discal intervals. Elytral proportions vary among individuals, but the humeri-defined by the angle at the juncture of the basal groove and lateral marginal depression-tend to be more proximate in these beetles; MEW / HuW = 2.45 - 2.56 versus MEW / HuW = 2.39 - 2.49 in Mecyclothorax splendidus. The male aedeagal median lobe is very diagnostic, with males of this species exhibiting a rounded tip with a short " crochet hook " dorsal projection (Fig. 107), not the broadly expanded lobe apex of Mecyclothorax iteratus (Fig. 103) nor the slightly dorsoventrally expanded apex of Mecyclothorax splendidus (Fig. 108). Standardized body length 4.5 - 5.4 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
273C4955C2024376E82D1D37677CB9CE.taxon	description	Description (n = 5). Head capsule with frontal grooves slightly sinuous, broad medial face transversely wrinkled, narrow lateral carina mesad anterior supraorbital seta; dorsal impression of neck broadly concave, visible dorsally; eyes convex, ocular ratio = 1.56 - 1.71, covering much of protruded ocular lobe, ocular lobe ratio = 0.82 - 0.90; labral anterior margin shallowly emarginate to 1 / 7 labral length; antennae filiform, antennomeres 2 - 3 with sparse pelage of short setae; mentum tooth with sides acute, apex rounded, margins beaded. Pronotum appearing longer than broad, but MPW / PL = 1.08 - 1.11, base constricted, MPW / BPW = 1.53 - 1.62; hind angles slightly acute, sharply angled, lateral margins subparallel to convergent for 0.1 x length; median base depressed relative to disc, ~ 25 elongate punctures each side separated by smooth cuticle; basal margin straight between hind angles; median longitudinal impression finely incised, broader on front of median base, crossed by indistinct transverse wrinkles; anterior transverse impression deeply incised, crossed by distinct longitudinal wrinkles medially, narrower and smoother mesad front angle; anterior callosity flat, crossed by dense wrinkles of bordering impression; front angles slightly projected, tightly rounded; apical pronotal width subequal to greater than basal width, APW / BPW = 0.99 - 1.08; lateral marginal depression very narrow, beaded laterally, slightly broader with upturned edge at front angle, upraised outside laterobasal depression; laterobasal depression an irregular continuation of lateral depression, margin beaded laterally. Proepisternum with 5 distinct punctures along hind marginal groove, small carinae between some punctures; prosternal process with very broad median depression, lateral margins broadly upraised. Elytra elongate obovoid, disc flat medially, sides distinctly sloped to margins; basal groove slightly curved to angulate humerus at juncture with lateral marginal depression, humeri proximate, lateral margins narrowly rounded, MEW / HuW = 2.45 - 2.56; parascutellar seta present; parascutellar striole with 5 isolated punctures, striole interrupted between punctures; sutural interval nearly coplanar with lateral intervals throughout length; sutural stria marked by proximate isolated punctures, very shallowly impressed, striae 2 - 5 with punctures progressively more isolated; discal intervals 2 - 4 flat to concave, intervals 5 - 8 following curvature of elytron; 7 th and 8 th intervals laterad apex of 7 th stria of same convexity; 2 dorsal elytral setae at 0.30 - 0.31 x and 0.66 - 0.70 x elytral length, setal impressions shallow, spanning 1 / 2 width of interval 3; apical and subapical setae absent; lateral elytral setae arranged in anterior series of 7 setae and posterior series of 6 setae; elytral marginal depression moderately narrow, edge upturned at humerus, depression lined with isodiametric sculpticells in anterior half, narrowed in apical 1 / 3 to subapical sinuation; subapical sinuation shallow, symmetrical. Mesepisternum with ~ 14 punctures in 2 - 3 rows; metepisternal width to length ratio = 0.75; metepisternum / metepimeron suture distinct. Abdomen with irregular longitudinal wrinkles laterally on ventrites 1 - 6, rounded lateral depressions on ventrites 3 - 6; suture between ventrites 2 and 3 effaced; apical male ventrite with 2 marginal setae and apical female ventrite with 4 equally spaced marginal setae and median trapezoid of 4 subequal, short setae. Legs-metatarsomere 1 / metatibial length ratio = 0.20; metatarsomere 4 length along outer lobe 1.4 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci narrow, lateral, median surface granulate. Microsculpture of vertex a transverse mesh, sculpticell breadth 2 x length, on frons, sculpticells 3 - 4 x broad as long on neck; pronotal disc with indistinct transverse mesh to transverse lines; pronotal median base with isodiametric sculpticells medially, transverse cells laterally; elytral disc with isodiametric mesh medially, transverse sculpticells on lateral intervals, apex with shallow isodiametric mesh; metasternum with transverse lines to transverse mesh, the sculpticell breadth 2 x length; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex rufous with piceous cast in frontal grooves; antennomere 1 rufoflavous, antennomeres 2 - 11 rufobrunneous; pronotal disc rufopiceous, lateral margins narrowly, and base and apex broadly, rufous; proepipleuron rufoflavous, proepisternum rufobrunneous; elytral disc rufous with piceous cast, sutural interval pale rufous laterad scutellum to apex, elytral lateral margins slightly paler, rufobrunneous, apex broadly rufoflavous; elytral epipleuron rufoflavous, metepisternum rufobrunneous; abdominal ventrites 1 - 5 rufous, laterally and apically on ventrite 6 rufoflavous; metafemur flavous, apex with a rufobrunneous cast that matches rufobrunneous metatibia. Male genitalia (n = 6). Aedeagal median lobe moderately robust, distance from parameral articulation to tip 3.4 - 3.8 x depth at midlength (Fig. 107 A, C-D); apex extended for 1.5 x its depth beyond ostial opening, dorsal surface expanded into a bluntly acute tooth, surface of tip rounded apicad tooth; median lobe curved rightward nearly 90 ° apically in ventral view, right margin evenly concave, left margin sinuous-convex then slight concave-before rounded, slightly expanded tip (Fig. 107 B); internal sac with discrete, ovoid dorsal ostial microtrichial patch comprised of macrospicules situated near sac base, and broadly diffuse field of dark microspicules covering ventral surface (Fig. 107 C); flagellar plate large and robust, length 0.55 x parameral articulation-tip distance. Female reproductive tract (n = 1). Bursa copulatrix columnar, narrow and elongate, apex rounded, length 1.14 mm, rounded apex breadth 0.45 mm, basal breadth 0.34 mm (Fig. 104 B); bursal walls translucent, thickly wrinkled; gonocoxite 1 with 3 - 5 apical fringe setae, a thick seta at medioapical angle and 5 - 6 setae along medial surface (Fig. 105 B); gonocoxite 2 broadly subtriangular, tip tightly rounded, base thinly extended laterally, 2 elongate lateral ensiform setae, apical nematiform setae on medioventral surface at 0.73 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
273C4955C2024376E82D1D37677CB9CE.taxon	etymology	Etymology. Beetles of this species are similar to those of Mecyclothorax iteratus, and so use of the same stem plus a prefix can signify that similarity. The epithet reiteratus represents a redundancy of iteratus - to repeat-so this species repeats again a similar habitus with differences most pronounced in one setational and more male genitalic characters.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
8904C5952B904B859DE133BB7A54776E.taxon	diagnosis	Diagnosis. Extremely similar to Mecyclothorax reiteratus, with bisetose pronotum and elytral humeri nearly as narrow; MEW / HuW = 2.39 - 2.49. The discal elytral intervals are more convex than those of Mecyclothorax reiteratus (Figs 102 B-C), and the pronotal lateral margins are slightly more convergent before the hind angles; lateral margin convergent for 0.12 x pronotal length. The dorsal microsculpture is more developed on beetles of this species, with the pronotal median base bearing a distinct isodiametric mesh across its width, and the elytral disc with a well-developed isodiametric mesh. The configuration of the male aedeagal median lobe is the best arbiter for identification, with the rounded lobe apex more or less symmetrically expanded dorsoventrally in males of this species, even given the infraspecific variation present in this structure (Fig. 108), versus a median lobe apex with a dorsal " crochet hook " expansion in males of Mecyclothorax reiteratus (Fig. 107). Setal formula 2 1 2 0 (1) [sae]; the subapical elytral seta is most commonly absent, but present in rare instances. Standardized body length 4.4 - 5.3 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
8904C5952B904B859DE133BB7A54776E.taxon	description	Description (n = 5). [The above description of Mecyclothorax reiteratus can serve to describe this species with the following substitutions.] Head with well-developed eyes, ocular ratio = 1.58 - 1.67, ocular lobe ratio = 0.84 - 0.90. Pronotum appearing longer than broad, but MPW / PL = 1.04 - 1.10, base moderately constricted, MPW / BPW = 1.46 - 1.54; median longitudinal impression finely incised, broadened on median base, crossed by moderately deep transverse wrinkles; anterior transverse impression shallow, broad, surface obscured by deep, dense longitudinal wrinkles; apical pronotal width slightly greater than basal width, APW / BPW = 1.02 - 1.05. Microsculpture of vertex an indistinct transverse mesh, sculpticell breadth 2 - 3 x length, cuticle glossy in part; elytral apex with evident isodiametric microsculpture; metasternum with shallow transverse lines to transverse mesh, sculpticell breadth 2 x length. Male genitalia (n = 18). Aedeagal median lobe gracile, distance from parameral articulation to tip 4.2 - 4.7 x depth at midlength (Fig. 108 A, C-I); apex extended 2 - 3 x depth beyond ostial opening, slightly expanded dorsoventrally at rounded tip; in ventral view, median lobe slightly curved rightward in apical half, the right and left margins convergent to rounded tip (Fig. 108 B); internal sac with broad 1 - part (Fig. 108 E), or 2 - 3 - part (Fig. 108 F, H) dorsal ostial microtrichial patch on right side at base, a transverse band of macrospicules on left side of sac base also present in some individuals (Fig. 108 H), ventral sac surface covered with shaggy pelage of microsetae; flagellar plate moderately large, length 0.40 - 0.44 x parameral articulation-tip distance. Female reproductive tract (n = 1). Bursa copulatrix columnar, elongate, apex rounded, length 1.03 mm, rounded apex breadth 0.50 mm, basal breadth 0.32 mm (Fig. 104 C); bursal walls translucent, apex thinly wrinkled, base more thickly wrinkled; gonocoxite 1 with 6 apical fringe setae, a thick seta at medioapical angle and 5 - 6 setae along medial surface (Fig. 105 C); gonocoxite 2 broadly subtriangular, tip subacuminate, base broadly extended laterally, 2 elongate lateral ensiform setae, apical nematiform setae on medioventral surface at 0.70 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
8904C5952B904B859DE133BB7A54776E.taxon	etymology	Etymology. Though hardly unique in its splendidness, this species is given the adjectival epithet splendidus to carry the flag for all species of this hyperdiverse Hawaiian radiation that evolved in splendid isolation (Simpson 1980, Zevon 1989).	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
8BEE2E9AD49244AAFAB2453A319CB0C5.taxon	diagnosis	Diagnosis. This species plus Mecyclothorax haleakalae and Mecyclothorax simpulum comprise the second triplet of Haleakala's Mecyclothorax haleakalae group species; this triplet characterized by the glabrous pronotum. In addition to the external key characters of: 1, minutely punctate pronotal median base; and 2, glossy elytral surface without evident microsculpture, this species (Fig. 110 A) is easily diagnosed from Mecyclothorax haleakalae (Fig. 110 B) and Mecyclothorax simpulum (Fig. 110 C) by the foreshortened, more transverse pronotum, MPW / PL = 1.15 - 1.21. The elytral microsculpture is also less developed in beetles of this species, with the discal surface of the elytra glossy, the surface slightly irregular but without microsculpture. Setal formula 1 (2) 0 2 0; the anterior supraorbital seta is present in rare instances. Standardized body length 4.9 - 5.9 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
8BEE2E9AD49244AAFAB2453A319CB0C5.taxon	description	Description (n = 5). Head capsule with frontal groove sinuously curved laterally from deep juncture with clypeus to position mesad supraorbital seta; dorsal impression of neck broadly concave, visible dorsally; protruded ocular lobe largely covered by convex eye; ocular ratio = 1.51 - 1.61, ocular lobe ratio = 0.84 - 0.91; labral anterior margin shallowly emarginate to 1 / 8 labral length; antennae filiform, antennomeres 2 - 3 with extremely short setae to glabrous; mentum tooth narrow, sides acute, apex tightly rounded. Pronotum with basal margin slightly convex medially; median longitudinal impression very finely incised, obsolete on disc, continued deeply on front of median base; anterior transverse impression broad, shallow medially, well incised in lateral halves of each side to front angle; anterior callosity slightly convex, surface minutely irregular due to densely lined longitudinal wrinkles; front angles slightly projected, broadly expanded before tightly rounded margin; pronotal apical width greater than basal width, APW / BPW = 1.05 - 1.12; lateral marginal depression very narrow, beaded laterally, broader with margin upturned at front angle and outside laterobasal depression; laterobasal depression very narrow, surface irregular, continuation of lateral depression. Proepisternum with 5 distinct punctures along hind marginal groove, small carinae between some of the punctures; prosternal process with broad median depression, lateral margins broadly upraised. Elytra elongate subovoid, disc flat medially, sides distinctly sloped to margins; basal groove curved to angulate humerus defined by hitch at base of marginal depression, MEW / HuW = 2.26 - 2.44; parascutellar seta present; parascutellar striole with 7 isolated punctures, interrupted between punctures; sutural interval slightly elevated basally, more so apically; discal striae 1 - 2 shallow, punctate, striae 3 - 5 a series of punctures, and striae 6 - 7 absent, sutural stria deep and fine apically, stria 2 obsolete on apex; discal intervals slightly convex on inner intervals, flat laterally; 8 th interval laterad 7 th stria of same convexity as apical fused portion of striae 5 + 7; 2 dorsal elytral setae at 0.28 - 0.30 x and 0.58 - 0.60 x elytral length, setal impressions spanning 1 / 2 width of interval 3; lateral elytral setae arranged in anterior series of 7 setae and posterior series of 6 setae; elytral marginal depression moderately narrow with upturned edge at humerus, depression lined with isodiametric sculpticells in anterior half, narrowed in apical 1 / 3 to subapical sinuation; subapical sinuation shallow, symmetrical, internal plica visible from dorsal view. Mesepisternum with ~ 11 punctures in 2 - 3 rows; metepisternal width to length ratio = 0.70; metepisternum / metepimeron suture distinct. Abdomen with irregular longitudinal wrinkles laterally on ventrites 1 - 6, round lateral depressions on ventrites 4 - 6; suture between ventrites 2 and 3 effaced; apical male ventrite with 2 marginal setae and apical female ventrite with 4 equally spaced marginal setae plus median trapezoid of 4 subequal, short setae. Legs-metatarsomere 1 / metatibial length ratio = 0.20; metatarsomere 4 length along outer lobe 1.3 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci narrow, lateral, median surface granulate. Microsculpture reduced on vertex, surface glossy with indistinct transverse sculpticells in shallow depressions of the cuticle; pronotal disc glossy, indistinct transverse sculpticells over parts of the surface, median base glossy between the isolated punctures; elytral apex glossy, with indistinct transverse sculpticells along margin; metasternum with shallow transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex rufous with a piceous cast; antennomere 1 rufoflavous, 2 - 11 rufobrunneous; pronotum rufopiceous, marginal depression narrowly rufoflavous; proepipleuron rufobrunneous, proepisternum rufopiceous; elytral disc rufopiceous with cupreous reflection, sutural interval concolorous basally, rufoflavous apically; elytral marginal depression rufobrunneous, apex narrowly rufobrunneous apicad subapical sinuation; elytral epipleuron dorsally rufoflavous, ventrally rufobrunneous, metepisternum rufopiceous; abdominal ventrites 1 - 5 medially rufopiceous, ventrites 3 - 6 rufoflavous marginally, apical ventrite 6 rufoflavous in apical half; metafemur rufoflavous, apex with rufobrunneous cast; metatibia rufobrunneous. Male genitalia (n = 5). Aedeagal median lobe gracile, distance from parameral articulation to tip 4.0 - 4.3 x depth at midlength (Fig. 111 A-C, E); apex elongate, extended 6 x its minimum depth beyond ostial opening, terminated in a spoonlike expanded tip; median lobe apex distinctly curved rightward at 45 ° angle in ventral view (Fig. 111 D), right margin distinctly concave, and left side slightly convex before apex which terminates in a chiseled tip in this view; internal sac with ovoid macrospicular field on right side near base, the ventral surface broadly and diffusely covered with a pelage of microspicules (Fig. 111 E); flagellar plate moderately large, length 0.47 x parameral articulation-tip distance. Female reproductive tract (n = 1). Bursa copulatrix columnar, narrow and elongate, apex rounded, length 1.20 mm, breadth 0.34 mm (Fig. 104 D); bursal walls translucent, thinly wrinkled; gonocoxite 1 with 4 - 5 apical fringe setae, a thicker seta at medioapical angle and 3 - 6 smaller setae along medial surface (Fig. 105 D); gonocoxite 2 subtriangular, apex tightly rounded, base broadly extended laterally, 2 broad and elongate lateral ensiform setae, apical nematiform setae on medioventral surface at 0.75 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
8BEE2E9AD49244AAFAB2453A319CB0C5.taxon	etymology	Etymology. The species epithet bacrionis, or Latin for ladle with a long handle (Brown 1956), is here used as a noun in apposition. The epithet references the conformation of the male aedeagal median lobe (Fig. 111).	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
8F41B61056E6FF1BE283842E6E07E6D1.taxon	diagnosis	Diagnosis. This species (Fig. 110 B) and Mecyclothorax simpulum (Fig. 110 C) both are characterized by a glabrous quadrate pronotum-MPW / PL = 1.07 - 1.14 for this species-with a densely punctate median base. Specimens of Mecyclothorax haleakalae can be told by the well-developed and regular isodiametric mesh covering the elytral disc. The anterior supraorbital seta is absent from specimens of this species, whereas nearly all specimens examined of Mecyclothorax simpulum have both anterior and posterior pairs of these setae. The male aedeagal median lobe apex is broader dorsoventrally and flattened apically in Mecyclothorax haleakalae males (Figs 113 A, C-D), and the internal sac bears two small ventral ostial microtrichial patches (Fig. 113 C) which are absent from Mecyclothorax simpulum males (Fig. 113 G). Setal formula 1 0 2 0. Standardized body length 5.2 - 6.0 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
CBD29086EB192F9E93F571B90A65A4A7.taxon	diagnosis	Diagnosis. This species (Fig. 110 C) can be easily diagnosed from the other two in this triplet- Mecyclothorax bacrionis (Fig. 110 A) and Mecyclothorax haleakalae (Fig. 110 B) by the presence of both anterior and posterior supraorbital setae (in 1 of 22 specimens, the anterior supraorbital setae is unilaterally absent; setal formula 2 (1) 0 2 0). The pronotum of this species is of exceedingly similar to that of Mecyclothorax haleakalae, however there are fewer punctures on the median base; ~ 25 each side. The sutural stria is shallower also, though discal stria 2 is about as impressed as in individuals of Mecyclothorax haleakalae. The male aedeagal median lobe has a slightly narrower apex, assessed dorsoventrally, with a more rounded tip (Fig. 113 E, G) versus the dorsoventrally broader, apically flattened median lobe apex of Mecyclothorax haleakalae males (Fig. 113 A, C-D). The male aedeagal internal sac also lacks any discrete ventral ostial microtrichial patches (Fig. 113 G), though a dorsal ostial microtichial patch is present. Standardized body length 5.3 - 6.3 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
CBD29086EB192F9E93F571B90A65A4A7.taxon	description	Description (n = 5). [The above description of Mecyclothorax bacrionis can serve to describe this species with the following substitutions.] Eyes large and convex, ocular ratio = 1.51 - 1.61, ocular lobe ratio = 0.84 - 0.91. Pronotum appearing elongate, but MPW / PL = 1.11 - 1.12, basally constricted, MPW / BPW = 1.59 - 1.72; hind angles slightly obtuse, apex sharp, projected; basal margin slightly convex between laterobasal de pressions, margins posterad laterobasal depressions separately convex; anterior callosity slightly convex, crossed by dense though indistinct longitudinal wrinkles; pronotal apical width variably subequal to greater than basal width, APW / BPW = 1.01 - 1.14, laterobasal depression narrow, surface punctate, continuous with lateral depression. Elytra narrowly ovoid but humeral angles situated posterolaterad pronotal hind angles, therefore MEW / HuW = 2.19 - 2.38 (values marginally less than those recorded for Mecyclothorax bacrionis); discal striae 1 - 2 impressed, punctate, the punctures more closely spaced on sutural stria; striae 3 - 5 punctate but less impressed than inner striae, punctures therefore more isolated; 2 dorsal elytral setae at 0.22 - 0.27 x and 0.66 - 0.69 x elytral length. Mesepisternum with ~ 11 punctures in 2 - 3 rows. Microsculpture of vertex an indistinct transverse mesh, sculpticell breadth 2 - 3 x length, surface glossy in parts; pronotal disc with indistinct transverse lines on a glossy surface, median base glossy medially, indistinct transverse sculpticells laterally; elytral disc with very small, indistinct isodiametric sculpticells in partially transverse rows, apex with shallow isodiametric mesh in transverse rows. Coloration of elytral disc rufopiceous, but without cupreous reflection associated with glossy elytral surface of Mecyclothorax bacrionis. Male genitalia (n = 5). Aedeagal median lobe robust, distance from parameral articulation to tip 3.2 x depth at midlength (Fig. 113 E, G); apex narrowly extended 2 - 3 x its depth beyond ostial opening, tip slightly oblique, dorsoapical face flattened, tip slightly expanded ventrally; median lobe parallel sided along shaft, right margin broadly concave before slightly offset parallel-sided apex with blunt tip (Fig. 113 F); internal sac with transverse dorsal ostial microtrichial patch at base of right side (Fig. 113 G), ventral surface covered with shaggy pelage of long microtrichia; flagellar plate large, robust, length 0.56 x parameral articulation-tip distance. Female reproductive tract (n = 1). Bursa copulatrix columnar with a rounded apical expansion, length 1.6 mm, apical lobe width 0.72 mm, basal constriction at vagina 0.36 mm broad (Fig. 104 F); bursal walls translucent, thickly wrinkled; gonocoxite 1 with 5 - 6 apical fringe setae, 1 - 2 thick setae at medioapical angle and 12 - 14 smaller setae along medial surface (Fig. 105 F); gonocoxite 2 subacuminate, lateral extension at base with curved terminus, 2 lateral ensiform setae, apical nematiform setae on medioventral surface at 0.75 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
CBD29086EB192F9E93F571B90A65A4A7.taxon	etymology	Etymology. As in Mecyclothorax bacrionis, the conformation of the male aedeagal median lobe is the most reliable means to diagnose this species. Here the Latin simpulum, small ladle, is used as a noun in apposition to signify the shape of the male median lobe (Figs 113 E, G).	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
0CFAF5AD9AA6ABF3257DBA7C90DCFCFA.taxon	diagnosis	Diagnosis. Beetles of this species display the most vivid bluish metallic reflection of any species in the group (Fig. 115 A-B). That characteristic in combination with the more transverse, glabrous pronotum, MPW / PL = 1.18 - 1.23 assures facile identification even in the field. The elytra vary in shape, with the humeral area broader (Fig. 115 A) or narrower (Fig. 115 B); MEW / HuW = 2.12 - 2.30. Setal formula 2 0 2 0. Standardized body length 4.9 - 5.6 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
B919BC2DD9DE47C82EFBF71987E1764D.taxon	diagnosis	Diagnosis. Beetles of this species (Fig. 115 C) exhibit a dorsal body surface that is as glossy as that of Mecyclothorax vitreus (Fig. 115 A-B), but the pronotum is much less transverse, MPW / PL = 1.08 - 1.10, and the lateral pronotal seta is present. The sutural and 2 nd stri ae are slightly impressed on the disc in association with a slightly convex interval 2. The elytra are rufobrunneous, with the sutural interval paler, rufous basally and rufoflavous apically. In keeping with the narrow elytral base, the humeri are more proximate than in any other member of the group; MEW / HuW = 2.42 - 2.53. Setal formula 2 1 2 0. Standardized body length 5.2 - 5.3 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
D193F9093C2425EAF3094FCFA9DACEC5.taxon	diagnosis	Diagnosis. The four newly described species of the Mecyclothorax vitreus group complex-this species (Fig. 115 D-E), Mecyclothorax kipahulu (Fig. 122 A), Mecyclothorax kaumakani (Fig. 122 B-C), and Mecyclothorax kuiki (Fig. 122 D-E) - were sorted for description principally by the configuration of the male aedeagus. Thus any external characters used here and in the key above are at best guideposts on the way to an identification, with a dissected male necessary for an authoritative determination, especially in the geographic area of extreme sympatry surrounding Kīpahulu Valley (Fig. 121). Nevertheless, based on available specimens, beetles of this species have the elytra more broadly ovoid relative to the width of the head across the eyes than observed in specimens of the other species; MEW / MHW = 2.18 - 2.21 (note: the span of these values for Mecyclothorax kuiki = 2.08 - 2.18). Pronotal shape, elytral striation, and microsculpture, usually of great help for species-level identification, are so variable infraspecifically that consulting the aedeagus is the only means to put a name on a specimen of this species with any confidence. That said, the male aedeagal median lobe is distinctive and relatively stable morphologically across this species' range (Fig. 120), with the apex broad dorsoventrally and blunt apically. The aedeagal internal sac bears a ventral ostial microtrichial patch, and the lobe is only partially divided into an apical versus a basal lobe (Fig. 120 G). Setal formula 2 1 2 0. Standardized body length 3.8 - 5.1 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
D193F9093C2425EAF3094FCFA9DACEC5.taxon	description	Description (n = 5). Head capsule with frontal grooves deep apically near clypeus, sinuous laterally to mesad anterior supraorbital seta, separated from seta by low carina; dorsal impression of neck broad, shallow, visible dorsally; ocular lobe moderately protruded from gena, eyes moderately convex, ocular ratio = 1.46 - 1.53, ocular lobe ratio = 0.80 - 0.89; labral anterior margin broadly shallowly emarginate; antennae filiform, antennomeres 2 - 3 with sparse pelage of short setae; mentum tooth with sides slightly acute, apex tightly rounded. Pronotum slightly, variably transverse, MPW / PL = 1.07 - 1.20, variably constricted basally, MPW / BPW = 1.49 - 1.63; hind angle slightly acute, lateral margin slightly convergent to evenly concave (in the same specimen, Fig. 115 E) anterad angle; median base with 9 - 14 isolated punctures each side; basal margin straight medially, slightly expanded posterad behind laterobasal depressions; median longitudinal impression very shallow, indistinct; front angles slightly projected, rounded; pronotal apex broader than base, APW / BPW = 1.05 - 1.14; lateral marginal depression moderately narrow, edge upturned; laterobasal depression narrow, smooth, laterally upraised to margin. Proepisternum with 6 minute punctures along hind marginal groove; prosternal process medially depressed with a low lateral marginal bead. Elytra convex, sides depressed; basal groove evenly and briefly curved to tightly rounded to obtusely angulate humerus at juncture of basal groove and lateral marginal depression; MEW / HuW = 2.32 - 2.44; parascutellar seta present; parascutellar striole with 4 isolated punctures; sutural interval slightly convex, lateral intervals 1 - 4 flat; sutural stria deep and distinct apically, whereas 2 nd stria shallow, broad there; 8 th interval laterad 7 th stria as convex as fused apical portion of striae 5 + 7; 2 dorsal elytral setae at 0.25 x and 0.59 - 0.66 x elytral length, setal impressions spanning 1 / 2 width of interval 3; lateral elytral setae arranged in anterior series of 7 setae and a posterior series of 6 setae; elytral marginal depression narrow, lateral margin upturned; subapical sinuation shallow, broad. Mesepisternum with ~ 16 isolated punctures in 3 rows; metepisternal width to length ratio = 0.72; metepisternum / metepimeron suture distinct; metathoracic flight wing a narrow strap 2.1 x long as broad, remnant R and M veins present, strap not reaching hind margin of metanotum. Abdomen with irregular lateral wrinkles on ventrites 1 - 3; suture between ventrites 2 and 3 complete; apical male ventrite with 2 marginal setae and apical female ventrite with 4 equally spaced marginal setae plus median trapezoid of 4 short setae, the basal pair longer. Legs-metatarsomere 1 / metatibial length ratio = 0.20; metatarsomere 4 length along outer lobe 1.5 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci broad, shallow. Microsculpture of vertex a transversely stretched mesh; pronotal disc with indistinct elongate transverse mesh, median base with isodiametric sculpticells near punctures, glossy surface in spaces between punctures; elytral disc with indistinct to obsolete isodiametric mesh in transverse rows-sculpticells visible only in small patches; elytral apex glossy with patches of indistinct isodiametric sculpticells; metasternum covered with swirling transverse mesh; laterobasal abdominal ventrites with swirling isodiametric sculpticells. Coloration of vertex a glossy brunneous; antennomeres 1 - 3 flavous, 4 - 11 rufobrunneous; pronotal disc brunneous, lateral depression and edge of disc rufoflavous; proepipleuron flavous, proepisternum rufobrunneous with piceous cast; elytral disc glossy rufobrunneous, sutural interval concolorous basally, rufoflavous apically, margins narrowly rufoflavous in lateral depression, apex rufoflavous; elytral epipleuron flavous, metepisternum rufobrunneous with piceous cast; abdomen piceous basally, lateral margins of ventrites 3 - 6 flavous; metafemur flavous; metatibia flavous with brunneous cast. Male genitalia (n = 38). Aedeagal median lobe squat, evenly curved, dorsal margin very convex along left margin of ostial opening, distance from parameral articulation to tip 2.4 - 2.8 x depth at midlength (Fig. 120 A, C-I); apex only briefly extended beyond ostial opening, obliquely blunt with apical face nearly flat to slightly convex, and ventral tip acutely rounded; median lobe straight overall in ventral view, apex curved slightly to left before rounded tip, convex dorsal margin visible to right of curved tip (Fig. 120 B); internal sac broad and squat, a variably developed ventral ostial microtrichial patch (Fig. 120 D-E, G-I), and a variably developed basal lobe (Fig. 120 D, G) present; flagellar plate very small, length 0.20 - 0.30 x parameral articulation-tip distance. Female reproductive tract (n = 1). Bursa copulatrix broad, saclike with rounded apex, length 0.74 mm, breadth 0.46 mm (Fig. 117 C); bursal walls translucent, thinly wrinkled; gonocoxite 1 with 5 apical fringe setae of 2 distinct sizes, 10 - 13 small setae across ventral surface (more may occur on dorsal surface but if so, they are impossible to discern) (Fig. 118 C); gonocoxite 2 broadly falcate, apex subacuminate, base broadly extended laterally, 2 lateral narrow ensiform setae, apical nematiform setae on medioventral surface at 0.70 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
D193F9093C2425EAF3094FCFA9DACEC5.taxon	etymology	Etymology. This species is named to commemorate the efforts of the first person who sorted to species the many Haleakala Mecyclothorax specimens collected during the initial years of this project. Prof. Kipling W. Will was the first to separate specimens of this species now called Mecyclothorax kipwilli.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
6ACDF31F5DA2370F9FF27DBF5814C96E.taxon	diagnosis	Diagnosis. Individual of this species can be diagnosed from those of Mecyclothorax kaumakani and Mecyclothorax kuiki by the less developed elytral microsculpture. The elytral intervals are glossy, with only indistinct transverse lines, at most, visible over portions of the glossy cuticle. The male aedeagal median lobe has a short apex that is narrowed from the ostial opening to a rounded tip (Fig. 123 A-E). The median lobe internal sac is divided into a shorter, broader basal lobe and a longer, thinner apical lobe (Fig. 123 C). Setal formula 2 0 1 0. Standardized body length 4.1 - 5.2 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
6ACDF31F5DA2370F9FF27DBF5814C96E.taxon	description	Description (n = 5). [The above description of Mecyclothorax kipwilli can serve to describe this species with the following substitutions.] Eyes more convex than in Mecyclothorax kipwilli; ocular ratio = 1.55 - 1.59, ocular lobe ratio = 0.79 - 0.85. Pronotum slightly transverse, MPW / PL = 1.16 - 1.19, variably constricted basally, MPW / BPW = 1.56 - 1.69; median base bearing 13 - 16 large, isolated punctures each side. Elytra narrowly ellipsoid, lateral margins somewhat projected laterad humeri, MEW / HuW = 2.17 - 2.36 (a conformation that coincides with the elytral shape of Mecyclothorax kaumakani, MEW / HuW = 2.10 - 2.27, and is broader basally, though not diagnostically so, than the elytra of Mecyclothorax kipwilli, MEW / HuW = 2.32 - 2.44, and Mecyclothorax kuiki, MEW / HuW = 2.30 - 2.39). Mesepisternum with ~ 8 - 10 punctures in 2 rows; metathoracic flight wing a narrow strap 1.9 x long as broad, remnant M vein present; strap not reaching hind margin of metanotum. Coloration of vertex rufous with piceous cast; pronotal disc rufopiceous, margins narrowly paler, rufous; elytral disc glossy rufopiceous, sutural interval rufous basally and apically, concolorous on disc. Male genitalia (n = 14). Aedeagal median lobe moderately robust, shaft curved, dorsal margin convex near apex of ostial opening, distance from parameral articulation to tip 3.0 - 3.4 x depth at midlength (Fig. 123 A, C-E); apical extension parallel sided, the tip evenly rounded; median lobe nearly straight in ventral view, the tip curved rightward, dorsal margin visible behind and to the left of curved tip (Fig. 123 B), internal sac with large broad basal lobe, and narrower, more elongate apical lobe bearing the flagellar plate (Figs 123 C, 124), sac surface covered with variously pigmented microspicules, but without distinct microtrichial patches; flagellar plate small, length 0.29 x parameral articulation-tip distance. Based on aedeagal conformation, Mecyclothorax kipwilli (Fig. 120), Mecyclothorax kipahulu (Fig. 123 A-E), and Mecyclothorax kaumakani (Fig. 123 F-L) comprise a triplet of closely related species. All have males with median lobes that are robust in breadth, distinctly curved, and have dorsoventrally broad, truncate apices. Males of all three species possess aedeagal internal sacs with both apical and basal lobes (Figs 120 G, 123 C, G), however in Mecyclothorax kipahulu and Mecyclothorax kaumakani males, the basal lobe approaches or exceeds the size of the plesiomorphically present apical lobe that bears the flagellar plate, supporting adelphotaxon status for these species. Female reproductive tract (n = 1). Bursa copulatrix columnar with rounded apex, length 0.84 mm, apical breadth 0.34 mm, maximum breadth 0.43 mm (Fig. 117 D); bursal walls thickly wrinkled at midlength, more finely wrinkled apically; gonocoxite 1 with 2 - 4 apical fringe setae, a curved seta at medioapical angle and 5 - 6 smaller setae along medial surface (Fig. 118 D); gonocoxite 2 falcate, apex subacuminate, base broadly extended laterally with curved terminus, 2 elongate lateral ensiform setae, apical nematiform setae on medioventral surface at 0.69 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
6ACDF31F5DA2370F9FF27DBF5814C96E.taxon	etymology	Etymology. This species is given the epithet kipahulu in reference to Kīpahulu Valley, home to Laka, a god of canoe makers (Pukui et al. 1974), and within whose boundaries this species lives (Fig. 121).	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
F7CBEF916CA31099805495197895D4DF.taxon	diagnosis	Diagnosis. The pronotal median base is more punctate in this species versus Mecyclothorax kipahulu and Mecyclothorax kuiki, with ~ 16 - 20 punctures each side (Fig. 122 B-C). The basal margin of the pronotum is smoothly curved, without dense longitudinal wrinkles as in Mecyclothorax kuiki (Fig. 122 D-E). Also, the elytra are not so widened in their apical half, leading to a lower ratio of MEW / HuW = 2.10 - 2.27, versus higher ratios among individuals of Mecyclothorax kuiki. The male aedeagal median lobe is most like those of Mecyclothorax kipahulu males, but the dorsal and ventral margins are more parallel on the apex, and the tip is slightly expanded into a knob (Fig. 123 F-L). Setal formula 2 1 2 0. Standardized body length 4.6 - 5.0.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
F7CBEF916CA31099805495197895D4DF.taxon	description	Description (n = 5). [The above description of Mecyclothorax kipwilli can serve to describe this species with the following substitutions.] Eyes more convex than in Mecyclothorax kipwilli, though not diagnostically so, ocular ratio = 1.53 - 1.59, ocular lobe ratio = 0.79 - 0.87. Pronotum little transverse, MPW / PL = 1.10 - 1.15, variably constricted basally, MPW / BPW = 1.46 - 1.65. Elytra narrowly ellipsoid, lateral margins somewhat projected laterad humeri, MEW / HuW = 2.10 - 2.27. Metathoracic flight wing a narrow strap 1.3 x long as broad, remnant M vein present; strap not reaching hind margin of metanotum. Microsculpture of pronotal median base an indistinct to distinct isodiametric mesh across the surface between punctures; elytral disc with shallow, evident transversely stretched isodiametric mesh in transverse rows. Coloration of vertex rufous with piceous cast; pronotal disc rufopiceous; elytral disc glossy rufopiceous, sutural interval concolorous basally rufoflavous apically; elytral epipleura rufoflavous, metepisternum rufopiceous. Male genitalia (n = 11). Aedeagal median lobe moderately robust, curved, dorsal margin distinctly convex, bulging near apex of ostial opening, distance from parameral articulation to tip 3.2 - 3.5 x depth at midlength (Fig. 123 F-K); median lobe straight in ventral view, with apex extended from right side, and bulging dorsal surface visible behind and to the left of apex (Fig. 123 L); internal sac with large, broad basal lobe, and narrow, short apical lobe bearing the flagellar plate, the sac surface covered with microspicules only; flagellar plate very small, length 0.19 - 0.20 x parameral articulation-tip distance. Female reproductive tract (n = 1). Bursa copulatrix broad, saclike, with basal lobe at vagina ventrad common oviduct, length 0.62 mm, breadth 0.36 mm (Fig. 117 E); bursal walls more heavily stained and thickly wrinkled at basal lobe, more translucent and not wrinkled near apex; gonocoxite 1 with 3 apical fringe setae and 8 smaller setae along medial surface (Fig. 118 E); gonocoxite 2 falcate, apex subacuminate, base broadly extended laterally with curved terminus, 2 lateral ensiform setae, apical nematiform setae on medioventral surface at 0.72 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
F7CBEF916CA31099805495197895D4DF.taxon	etymology	Etymology. The distribution of this species embraces Kīpahulu Valley, with specimens collected on Kaumakani mountain to the east of Kīpahulu Valley, and along the western rim of Kīpahulu Valley ESE of Kuiki (Fig. 121). Kaumakani is taken as the species epithet for this species, allowing the next species of this complex to be named after Kuiki.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
AAF15F9C212D09B6E1FC31E5361DC6F4.taxon	diagnosis	Diagnosis. This fourth species of the Mecyclothorax vitreus group complex can be diagnosed by the pronotal median base with large punctures traversing the midline of the base, and small longitudinal wrinkles dissecting or at least disturbing the basal margin (Fig. 122 D-E). The median base also has well-developed isodiametric microsculpture between the punctures. The elytra are ellipsoid, the base narrow with the margins laterad the humeri barely extended, resulting in high ratios of MEW / HuW = 2.30 - 2.39 that are mirrored in Mecyclothorax kipahulu (Fig. 122 A) and Mecyclothorax kipwilli (Fig. 115 D-E), but greater than those derived from Mecyclothorax kaumakani (Fig. 122 B-C). The male aedeagal median lobe is of the most plesiomorphic configuration within this species complex, as the median lobe is gracile and moderately curved, and apex is narrowly extended beyond the ostial opening, the tip narrowly rounded and slightly downturned (Fig. 125 A-C, F-G). The male aedeagal internal sac corroborates the median lobe's relative plesiomorphy, as it is unilobate, long, and generally of the configuration observed across the genus (Fig. 125 B, F, G). Setal formula 2 1 2 0. Standardized body length 4.4 - 5.5 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
AAF15F9C212D09B6E1FC31E5361DC6F4.taxon	description	Description (n = 5). [The above description of Mecyclothorax kipwilli can serve to describe this species with the following substitutions.] Eyes of similar convexity to Mecyclothorax kipwilli; ocular ratio = 1.46 - 1.58, ocular lobe ratio = 0.76 - 0.81. Pronotum appearing elongate but slightly transverse, MPW / PL = 1.10 - 1.15, variably constricted basally, MPW / BPW = 1.46 - 1.65; median base bearing 10 - 13 isolated punctures across midlength of base, plus fine longitudinal wrinkles along the basal margin. Metathoracic flight wing an ellipsoid strap 1.4 x long as broad, remnant R + M veins present; strap not reaching hind margin of metanotum. Microsculpture of elytral disc shallow, evident, comprising a transversely stretched isodiametric mesh arranged in transverse rows, elytral apex with evident isodiametric mesh. Coloration of vertex rufous with piceous cast; pronotal disc rufobrunneous with piceous cast, margins narrowly paler, rufous; elytral disc rufobrunneous, sutural interval rufous basally and apically, concolorous on disc. Male genitalia (n = 11). Aedeagal median lobe robust, dorsal surface broadly convex, distance from parameral articulation to tip 2.3 x depth at midlength (Fig. 125 A-C, E-G); apex very narrowly extended, the tip obliquely rounded with dorsoapical face slightly flattened; median lobe sinuously curved in ventral view, right margin slightly concave, left margin briefly convex before narrow, bluntly rounded tip (Fig. 125 D); internal sac very broad, cylindrical (Fig. 125 B, F-G), length variable, from 1.0 - 1.2 x parameral articulation-tip distance; a small basal lobe may (Fig. 125 B) or may not (Fig. 125 F, G) be present on dorsal surface of sac; ventral sac surface covered with shaggy pelage of longer microtrichia; flagellar plate robust, very large, length 0.61 - 0.67 x parameral articulation-tip distance (plate size is positively associated with sac length, Fig. 125 B, F-G). Female reproductive tract (n = 1). Bursa copulatrix columnar, broad and elongate, length 1.48 mm, breadth 0.57 mm (Fig. 117 F); bursal walls translucent, surface more thickly wrinkled near base, apex with only thin wrinkles; gonocoxite 1 with 3 - 4 apical fringe setae and 6 - 9 smaller setae along medial surface (Fig. 118 F); gonocoxite 2 falcate, apex subacuminate, basal extension elongate with curved terminus, 2 thin lateral ensiform setae, apical nematiform setae on medioventral surface at 0.71 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
AAF15F9C212D09B6E1FC31E5361DC6F4.taxon	etymology	Etymology. As most specimens of this species have been found in proximity to Kuiki, the summit of Kīpahulu Valley's west rim, that locality name is used as the epithet for this final species in the Mecyclothorax kipwilli + Mecyclothorax kipahulu + Mecyclothorax kaumakani + Mecyclothorax kuiki species quartet.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
3202E58187E7EB4178A9861B9E6AF7BC.taxon	diagnosis	Diagnosis. These handsome beetles are instantly recognizable by: 1, the very large body size, standardized body length 6.9 - 7.4 mm; 2, dark, glossy cuticle; and 3, transverse pronotum-MPW / PL = 1.23 - 1.27 - with projected, glabrous hind angles (Fig. 126 A). The discal elytral striae are deep and distinctly punctate, the associated intervals moderately convex. The only other Hawaiian Mecyclothorax of similar body size is the very different-looking Mecyclothorax molops (Fig. 95 A). Although only possible to test using first-hand experience, the elytral cuticle is so hard and brittle that use of an insect mounting pin to prepare a fully sclerotized specimen will shatter the elytron (much as would happen during such a pinning exercise for a specimen of Clivina Latreille). Setal formula 2 1 2 1 [sae].	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
3202E58187E7EB4178A9861B9E6AF7BC.taxon	description	Description (n = 4). Head capsule with frontal grooves broad near clypeus, shallow, lateral convexity very low; dorsal surface of neck flat to convex; ocular lobe obtusely projected from gena, ocular ratio = 1.48 - 1.56, ocular lobe ratio = 0.79 - 0.84; labral anterior margin broadly, shallowly emarginate; antennae filiform, antennomeres 2 - 3 glabrous except for apical setae; mentum tooth with sides acute, apex rounded. Pronotum broad, moderately constricted basally, MPW / BPW = 1.20 - 1.27; hind angle obtuse, margin behind angle convex, lateral margin anterad angle subparallel for short distance before diverging; median base smooth with minute, isolated punctures; basal margin convex between hind angles; median longitudinal impression shallow, finely incised; anterior transverse impression broad, deep, continuous medially, finely incised at front angles; anterior callosity broadly convex, smooth; front angles subangulate, not protruded; pronotal apex much narrower than basal width, APW / BPW = 0.79 - 0.83; lateral marginal depression narrow, edge beaded at midlength, slightly broader at front angle, broadly explanate at hind angle; laterobasal depression smooth, deep, continuous with lateral depression. Proepisternum and proepimeron with ~ 6 large punctures along hind marginal grooves; prosternal process broadly depressed on ventral surface, low marginal bead between coxae. Elytra subparallel with broadly convex apical margins, disc convex, sides depressed; basal groove absent from base of parascutellar interval, angled anteriorly at lateral edge of sutural interval and subangulate humerus at base of stria 6; elytral base broad, MEW / HuW = 1.63 - 1.70; parascutellar seta present; parascutellar striole shallow, 3 - 4 isolated punctures along length; sutural interval more convex than lateral intervals, sutural juncture upraised; sutural and 2 nd striae of subequal depth on disc, stria 2 evanescent at apex whereas sutural stria finely incised, deep, and smooth; all striae except sutural and 8 th absent from elytral apex, elytral surface evenly convex between those striae; 2 dorsal elytral setae at 0.25 x and 0.55 x elytral length, setal impressions small, spanning about 1 / 3 of interval 3; apical absent, subapical setae present in melanized track of reduced stria 7; lateral elytral setae arranged in anterior series of 7 setae and posterior series of 6 setae; elytral marginal depression narrow, lateral margin upturned; subapical sinuation very shallow, internal plica visible from dorsal viewpoint. Mesepisternum with ~ 20 punctures in 2 - 3 rows; metepisternal width to length ratio 0.83; metepisternum / metepimeron suture distinct; metathoracic flight wing an elongate strap, 3.7 x long as broad, remnant R and M veins present, the strap extended beyond posterior margin of metanotum for 0.4 x its length. Abdomen with anterior margin of abdominal ventrite 2 depressed along suture; abdominal ventrites 1 - 6 smooth; suture between ventrites 2 and 3 very shallow but traceable; apical male ventrite with 2 marginal setae and apical female ventrite with 4 equally spaced setae and median trapezoid of 4 subequal, short setae. Legs-metatarsomere 1 / metatibial length ratio = 0.17; metatarsomere 4 length along outer lobe 1.2 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci broad, shallow. Microsculpture of vertex, pronotal disc and base obsolete, surfaces glossy; elytral disc with obsolete, elongate transverse mesh, apex without microsculpture, surface glossy; metasternum and laterobasal abdominal ventrites with glossy surfaces. Coloration of vertex rufopiceous; antennomere 1 rufoflavous, antennomeres 2 - 11 with piceous cast; pronotal disc and margins rufopiceous; proepipleuron rufobrunneous with piceous upper margin, proepisternum rufobrunneous with piceous cast; elytral disc rufopiceous, sutural interval concolorous basally to rufobrunneous apically, margins concolorous with disc; elytral epipleuron and metepisternum rufous; abdomen rufous with a piceous cast, apex of apical ventrite only slightly paler; metafemur rufoflavous with median piceous cloud; metatibia rufopiceous with tibial setae golden. Male genitalia (n = 2). Aedeagal median lobe large, moderately gracile, distance from parameral articulation to tip 3.9 x depth at midlength (Fig. 127 A); apex extended 2.5 x depth beyond apex of ostial opening, curved dorsally near tip that is slightly expanded dorsally, and broadly convex apically; median lobe straight in ventral view, lateral margins evenly convergent to knifelike apex that is slightly curved toward left at tip (Fig. 127 B); internal sac narrow, stalklike, with a broad dorsal ostial microtrichial patch at base, and ventral surface broadly covered with shaggy, melanic, microspicules (Fig. 127 C); flagellar plate elongate, as long as membranous internal sac, and 0.64 x as long as parameral articulation-tip distance. Female reproductive tract (n = 1). Bursa copulatrix columnar with rounded apex and basal constriction, length 1.48 mm, breadth 0.74 mm, basal constriction 0.45 mm broad (Fig. 128 A); bursal walls translucent, thickly wrinkled; gonocoxite 1 with 3 apical fringe setae, the middle seta of series largest, a curved seta at medioapical angle and 8 - 10 setae on medial surface (Fig. 129 A); gonocoxite 2 narrowly subtriangular with tightly rounded apex, base only moderately extended laterally, 2 lateral ensiform setae with apical seta broader and longer than basal seta, apical nematiform setae on medioventral surface at 0.70 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
3202E58187E7EB4178A9861B9E6AF7BC.taxon	etymology	Etymology. At 7.4 mm, the largest individuals of this species are among the largest Mecyclothorax beetles in Hawai'i, and so the species epithet rex - Latin for king-seems an appropriate epithet to apply.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
4181F60800CF35FB237458C5D109A788.taxon	diagnosis	Diagnosis. This is one of two Haleakala species (Fig. 126 B-C) in this group that exhibit the full complement of supraorbital, pronotal, and dorsal and apical / subapical elytral setae; setal formula 2 2 2 2. However Mecyclothorax montivagus beetles are of moderate to large size for Hawaiian Mecyclothorax - standardized body length 4.9 - 6.9 mm-in contrast to those of Mecyclothorax micans; standardized body length 3.6 - 4.3 mm. If a qualitative character is desired to separate the species, one may use the presence of the parascutellar seta in individuals of this species, and the lack thereof in individuals of Mecyclothorax micans.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
BA33DD3A3F475534071AA5121E093B38.taxon	diagnosis	Diagnosis. This species (Fig. 126 C) shares with Mecyclothorax montivagus (Fig. 126 B): 1, a broad subquadrate elytra with broadly rounded humeri, MPW / HuW = 1.91 - 2.05; and 2, a transverse pronotum-MPW / PL = 1.36 - 1.32 - with briefly sinuate lateral margins anterad the obtuse hind angles. The parascutellar seta is absent. The cuticular microsculpture is similarly to that of Mecyclothorax montivagus: 1, vertex and pronotal disc with obsolete transverse mesh; 2, pronotal median base glossy; 3, elytral disc with obsolete elongate transverse mesh, apex with isodiametric sculpticells in transverse rows; 4, metasternum glossy; and 5, laterobasal abdominal ventrites with swirling isodiametric and transverse sculpticells. Setal formula 2 2 2 2. Standardized body length 3.6 - 4.3 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
797EAABEBA92A13FE190E9D5859BAB5C.taxon	diagnosis	Diagnosis. This is the only Haleakala species of the group that consistently exhibits a glabrous pronotum (Fig. 133 A). Mecyclothorax aquilus is otherwise most similar in appearance to Mecyclothorax insolitus (Fig. 133 B) and Mecyclothorax invisitatus (Fig. 133 C) sharing the broadly flavous apical elytral margins. However beetles of this species are larger, standardized body length 5.1 - 5.6 mm, versus a range of 4.2 - 5.1 mm for the other two species. Setal formula 2 0 2 0.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
797EAABEBA92A13FE190E9D5859BAB5C.taxon	description	Description (n = 2). Head capsule with frontal grooves deep and broad near clypeus, a broad lateral convexity before eye that continues posterad enveloping anterior supraorbital seta; dorsal impression of neck slightly concave; ocular lobe moderately protruded from gena, eyes moderately convex, ocular ratio = 1.48 - 1.49, ocular lobe ratio = 0.80 - 0.81; labral anterior margin shallowly emarginate to 1 / 8 of labral length; antennae filiform, antennomeres 2 - 3 with sparse pelage of short setae; mentum tooth with sides acute, apex tightly rounded. Pronotum slightly transverse, MPW / PL = 1.11 - 1.16, the disc broadly convex; hind angles obtuse but sharp apically, lateral margins evenly divergent anterad, MPW / BPW = 1.52 - 1.56; median base slightly depressed relative to disc, ~ 20 minute, isolated punctures each side in glossy surface; basal margin distinctly convex between hind angles; median longitudinal impression very finely incised, broader in front of median base; anterior transverse impression very shallow, broad, incised only immediately inside front angle; anterior callosity flat medially, surface glossy; front angles slightly projected, tightly rounded; pronotal apical width variably subequal to basal width, APW / BPW = 0.97 - 1.06; lateral marginal depression obsolete, narrowly beaded laterally and basally, bead thicker only at hind angle, depression narrow only inside upraised margin of front angle; laterobasal depression deep, sloping from disc, margined by narrow U-shaped depression. Proepisternum with 5 minute punctures along hind marginal groove; prosternal process with broad median depression, lateral margins broadly upraised between coxae. Elytra subovoid, domed, the sides distinctly sloped to a vertical juncture with lateral marginal depression; basal groove evenly and briefly curved to angulate humerus at juncture with broader lateral marginal depression, MEW / HuW = 2.21 - 2.38; parascutellar striole with 3 - 5 isolated punctures, the striole shallow between punctures; sutural interval coplanar with lateral intervals basally, upraised in apical half; sutural and striae 2 - 4 of equal development on disc, but sutural stria the only one complete to base and apex, the basal part shallow and punctate, the apical portion finely incised, smooth and deep; stria 5 traceable only as a series of shallow punctures in basal half, striae 6 - 7 absent; elytral apex evenly and broadly convex, the lateral surface overlying the deep 8 th stria; 2 dorsal elytral setae at 0.33 - 0.36 x and 0.54 - 0.60 x elytral length, setal impressions small, spanning 1 / 4 of interval 3; lateral elytral setae arranged in an anterior series of 7 setae and a posterior series of 6 setae; elytral marginal depression narrow throughout, edge upturned at humerus, beaded laterally to subapical sinuation; subapical sinuation shallow, symmetrical. Mesepisternum with ~ 11 punctures in 2 - 3 rows; metepisternal width to length ratio = 0.71; metepisternum / metepimeron suture distinct. Abdomen with smooth glossy ventrites, round lateral depressions on ventrites 3 - 6; suture between ventrites 2 and 3 effaced; apical male ventrite with 2 marginal setae and apical female ventrite with 4 equally spaced marginal setae plus trapezoid of 4 subequal, short setae. Legs-metatarsomere 1 / metatibial length ratio = 0.20; metatarsomere 4 length along outer lobe 1.5 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci narrow, lateral, upper surface granulate without carina. Microsculpture of vertex an obsolete transverse mesh, sculpticell breadth 2 x length; pronotal disc and base glossy; elytral disc glossy, indistinct transverse lines visible through the reflective sheen; elytral apex glossy, surface irregular but no sculpticells visible; metasternum glossy; laterobasal abdominal ventrites glossy, indistinct sculpticells in lateral depressions. Coloration of vertex rufous with a piceous cast; antennomeres 1 - 4 rufoflavous, 5 - 11 slightly darker; pronotal disc rufous with piceous cast, lateral margins narrowly rufous, and base and apex broadly rufous; proepipleuron rufoflavous, proepisternum rufopiceous; elytral disc rufopiceous, sutural interval broadly rufous basally, rufoflavous to flavous apically; elytral margins with intervals 7 - 8 pale rufous basally, margin more broadly flavous in apical 1 / 3 where apex of interval 2 is also broadly flavous; elytral epipleuron flavous dorsally, darker ventrally, metepisternum rufopiceous; abdomen with ventrite 1 - 5 medially rufopiceous, ventrites 3 - 6 flavous marginally, the apical half of ventrite 6 flavous; metafemur flavous; metatibia flavous with brunneous cast. Male genitalia (n = 1). Aedeagal median lobe gracile, distance from parameral articulation to tip 4.5 x depth at midlength (Fig. 134 A); apex sinuously extended to a dorsoventrally expanded knoblike tip; median lobe curved to right beyond ostial opening in ventral view; internal sac with small dorsal ostial microtrichial patch based on uneverted specimen; flagellar plate short, length 0.32 x parameral articulation-tip distance. Female reproductive tract. The female allotype was not dissected.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
797EAABEBA92A13FE190E9D5859BAB5C.taxon	etymology	Etymology. The Latin adjective aquilus - dark-colored, blackish, dun, swarthy (Brown 1956) - is used to signify the dark body coloration representative of this species.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
8FE2FBA987E64C9453987AB5A4F0D933.taxon	diagnosis	Diagnosis. This species (Fig. 133 B) and Mecyclothorax invisitatus (Fig. 133 C) comprise a cryptic sibling species pair that can be best diagnosed externally by the breadth of elytra relative to the pronotum. In this species the elytra are broader, subovoid, with MEW / MPW = 1.63 - 1.69, whereas in Mecyclothorax invisitatus recorded values are 1.52 - 1.62. The pronotal disc bears obsolete transverse-mesh and transverse-line microsculpture in this species, whereas the disc surface is glossy in Mecyclothorax invisitatus. As there are no other reliable external characters that can separate these species, their recognition lies heavily on the differing configurations of the male aedeagal median lobe. Males of this species exhibit a median lobe with the apex expanded both dorsally and ventrally, and more flattened apically (Fig. 134 F-K). The apex may be displaced dorsoventrally relative to the more basal portion of the shaft giving it a " prehensile proboscis " appearance. Conversely, the median lobe of Mecyclothorax invisitatus males is expanded only dorsally into a " crochet hook " apex, with the apical margin evenly rounded; a conformation also observed in males of Mecyclothorax reiteratus (Fig. 107). In the specimens available, the elytral apex is less broadly flavous in Mecyclothorax insolitus (Fig. 133 B) compared to Mecyclothorax invisitatus (Fig. 133 C), but the diagnostic value of this feature should be confirmed through examination of the male median lobe apex. Setal formula 2 1 2 0. Standardized body length 4.2 - 5.1 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
218B4F6635EB611FBB9CB5854B7F6B37.taxon	diagnosis	Diagnosis. Among the Haleakala members of the Mecyclothorax ducalis group, this species is diagnosable by: 1, pronotum with lateral seta present; 2, elytra with apical margins flavous, contrasted to the rufopiceous disc; 3, pronotal disc glossy, microsculpture absent; 4, elytra narrowly subovoid, MEW / MPW = 1.52 - 1.62. Externally the cryptic sibling species, Mecyclothorax insolitus, differs in the last two criteria, with male aedeagal median lobe configuration offering an additional diagnostic difference between the species. Males of this species have the median lobe apex expanded into a dorsal hook, with the apex rounded ventrally (Fig. 138 A-C), whereas Mecyclothorax insolitus males have the median lobe apex expanded both dorsally and ventrally, with the apical margin straighter (Fig. 134 F-K). Setal formula 2 1 2 0. Standardized body length 4.3 - 5.1 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
218B4F6635EB611FBB9CB5854B7F6B37.taxon	description	Description (n = 4). Head capsule with frontal grooves broad and deep near clypeus, a broad lateral convexity anterad eye, and low carina mesad anterior supraorbital seta; dorsal impression of neck broad, shallow, the concavity visible in dorsal view; ocular lobe protruded obtusely from gena, eyes moderately convex, ocular ratio = 1.50 - 1.55, ocular lobe ratio 0.80 - 0.84; labral anterior margin moderately, angularly emarginate 1 / 6 of labral length; antennae filiform, antennomeres 2 - 3 with sparse pelage of short setae; mentum tooth with sides acute, apex rounded. Pronotum moderately transverse, MPW / PL = 1.15 - 1.24, constricted basally with lateral margins briefly sinuate anterad right to obtuse hind angles, MPW / BPW = 1.53 - 1.62; median base slightly depressed medially relative to disc, more so laterally, ~ 20 punctures each side isolated in glossy surface; basal margin distinctly convex between hind angles; median longitudinal impression finely incised, crossed by indistinct transverse wrinkles; anterior transverse impression shallow, broad, smooth, incised only mesad front angle; anterior callosity slightly convex, smooth; front angles slightly projected, tightly rounded; pronotal apical width subequal to slightly broader than basal width, APW / BPW = 1.0 - 1.05; lateral marginal depression obsolete, narrowly beaded except slightly broader at front angle and evenly elevated from laterobasal depression to apex of projected hind angle; laterobasal depression deep, narrow, continuous with lateral depression. Proepisternum with 5 minute punctures along hind marginal groove; prosternal process slightly convex medially but with broad upraised margin. Elytra convex, sides distinctly sloped to vertical juncture with lateral marginal depression; basal groove briefly curved to proximate, subangulate humerus at juncture with broader lateral marginal depression, MEW / HuW = 2.22 - 2.50; parascutellar striole with 5 punctures, striole shallow between punctures; sutural interval slightly more convex than lateral intervals basally, more upraised apically; discal striae 1 - 6 distinctly punctate, progressively shallower laterally, stria 7 a series of shallow punctures at midlength; sutural stria deep and smooth on elytral apex, striae 2 - 3 broad and very shallow there, lateral striae not visible; discal elytral intervals 2 - 4 moderately convex, outer intervals follow curvature of elytron; 2 dorsal elytral setae at 0.25 x and 0.51 - 0.53 x elytral length, setal impressions shallow, crossing 1 / 4 of interval 3; lateral elytral setae arranged in anterior series of 7 setae and posterior series of 6 setae; elytral marginal depression narrow throughout length, edge upturned at humerus, beaded laterally to subapical sinuation; subapical sinuation shallow, symmetrical. Mesepisternum with ~ 14 punctures in 2 - 3 rows; metepisternal width to length ratio = 0.79; metepisternum / metepimeron suture distinct. Abdomen with irregular longitudinal wrinkles on ventrites 1 - 6, round lateral impressions on ventrites 3 - 6; suture between ventrites 2 and 3 effaced; male apical ventrite with 2 marginal setae and female apical ventrite with 4 equally spaced setae and median trapezoid of 4 subequal, short setae. Legs-metatarsomere 1 / metatibial length ratio = 0.20; metatarsomere 4 length along outer lobe 1.3 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci broad, shallow, median area subcarinate. Microsculpture of vertex an obsolete transverse mesh, sculpticell breadth 2 x length; pronotal median base glossy medially, indistinct transverse sculpticells present laterally; elytral disc and apex glossy, microsculpture obsolete; metasternum with a transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex rufous; antennomeres 1 - 2 rufoflavous, 3 - 11 darker, rufobrunneous; pronotal disc rufous, lateral margins, base, and apex narrowly rufoflavous; proepipleuron flavous, proepisternum rufous with piceous cast; elytral disc rufopiceous, sutural interval rufobrunneous basally, rufoflavous apically, elytral margins with intervals 7 - 9 or 8 - 9 contrastedly paler, rufoflavous to flavous, apex (up to apical 0.2 x length) flavous; elytral epipleuron flavous dorsally, piceous ventrally, metepisternum rufopiceous; abdomen with all ventrites piceous mediobasally, flavous laterally, apical ventrite with apical 3 / 4 flavous; metafemur flavous; metatibia rufoflavous with brunneous cast. Male genitalia (n = 2). Aedeagal median lobe gracile, distance from parameral articulation to tip 3.8 x depth at midlength (Fig. 138 A, C); apex hooklike with blunt dorsal projection and evenly convex tip, the ventral margin not expanded ventrally as in Mecyclothorax insolitus (Fig. 134 D, F, G, K); median lobe apically curved rightward at approximately 60 ° angle in ventral view (Fig. 138 C, E, J), tip narrowly rounded in this view; internal sac with heavily spiculated dorsal ostial microtrichial patch at base of sac, and ventral ostial microtrichial patch at midlength (Fig. 138 C, F, H); flagellar plate large, length 0.51 x parameral articulation-tip distance. Female reproductive tract (n = 1). Bursa copulatrix columnar, narrow with rounded apex, length 1.14 mm, apical width 0.40 mm, minimal width 0.30 mm (Fig. 136 B); bursal walls more translucent basally, more thickly wrinkled apically; gonocoxite 1 with 3 short apical fringe setae, a curved seta at medioapical angle and 4 - 5 setae on medial surface (Fig. 137 B); gonocoxite 2 short and broad basally, triangular with lateral margin evenly expanded to base, 2 small lateral ensiform setae, apical nematiform setae on medioventral surface at 0.80 x gonocoxite length (short, broad apex may be due to wear).	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
218B4F6635EB611FBB9CB5854B7F6B37.taxon	etymology	Etymology. This close relative of Mecyclothorax insolitus is given the name Mecyclothorax invisitatus to signify the two species' affinity. The adjective insolitus is translated as unusual, uncommon, or strange (Brown 1956), whereas invisitatus can be translated as strange, new, uncommon (Brown 1956). Thus the new species name proposed here owes its origin to David Sharp's (1903) use of insolitus for the former species.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
CE4B7A76CC9D95C766F140662AA21072.taxon	diagnosis	Diagnosis. This large bodied species, standardized body length 6.0 - 6.7 mm, also stands out from other species in the group by the dark glossy, rufopiceous body color and contrasting flavous femora and infuscated tibiae. The pronotum is less constricted basally than in the other Haleakala species of the group, MPW / BPW = 1.43 - 1.46, and the elytra are broadly subquadrate, with laterally extended basal margins outside the tightly rounded humeri (Fig. 133 D); MEW / HuW = 2.06 - 2.15. Given these beetles' large body size, the elytral striae are not deeply impressed, with only the sutural and 2 nd striae impressed on the disc, and lateral striae 3 - 5 indicated by increasingly isolated punctures. Setal formula 2 1 (0) 2 0; the lectotype lacks pronotal lateral setae, a second specimen lacks the left seta, with the right present, and the other two examined specimens have both right and left lateral setae present.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
12E3170F416CA172CF528AF7E8EBBE95.taxon	diagnosis	Diagnosis. This species (Fig. 140 A-B) and the following, Mecyclothorax brevidux (Fig. 140 C-D) are together diagnosed among Haleakala species of the Mecyclothorax ducalis group by: 1, the bisetose pronotum; 2, dark rufous to rufopiceous elytra with concolorous apex; 3, basally constricted pronotum, MPW / BPW = 1.40 - 1.55 in this species; and 4, moderate body size, standardized body length 5.5 - 5.7 mm for this species. The elytral striae in individuals of this species are much more distinctly punctate, with the punctures distributed across the discal striae 1 - 5 where they expand strial breadth. The metafemur has the basal half of the anterior face covered with a piceous cloud, contrasted to the flavous medial coloration of the femur. In contrast, the metafemur in Mecyclothorax brevidux has only the basal 1 / 5 to 1 / 3 with piceous infuscation. In both species the femoral apex is darkened to match the rufobrunneous to rufopiceous tibial coloration. The male aedeagal median lobe confirms the value of these characters for diagnosis, with the lobe apex of Mecyclothorax longidux males narrow and elongate, with a slightly downturned tip (Fig. 141 A-B, D), versus a shorter broader lobe apex in males of Mecyclothorax brevidux (Fig. 141 F-I). The spiculation of the internal sac is also more developed in males of this species, with a bipartite dorsal ostial microtrichial patch present as well as a large, heavily spiculated ventral ostial microtrichical patch (Fig. 141 D-E). Setal formula 2 1 2 0.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
12E3170F416CA172CF528AF7E8EBBE95.taxon	description	Description (n = 5). Head capsule with frontal grooves deep, broad near clypeus, low broad lateral convexity anterad eye, terminated posteriorly mesad low ridge inside anterior supraorbital seta; dorsal impression of neck broad, shallow, visible in dorsal view; eyes large, moderately convex, ocular ratio = 1.53 - 1.63, ocular lobe ratio = 0.81 - 0.84; labral anterior margin angularly emarginate to 0.2 x labral length; antennae filiform, antennomeres 2 - 3 with sparse pelage of short setae; mentum tooth with sides acute, apex tightly rounded. Pronotum little transverse, MPW / PL = 1.08 - 1.20; lateral margins subparallel to slightly convergent for 0.1 x pronotal length anterad slightly obtuse, projected hind angles; median base moderately depressed relative to disc, ~ 25 minute, isolated punctures each side; basal margin distinctly convex between hind angles; median longitudinal impression finely incised, crossed by indistinct longitudinal wrinkles; anterior transverse impression shallow, broad, finely incised immediately mesad front angles; anterior callosity flat medially, smooth; front angles very slightly produced, tightly rounded; apical width subequal to basal width, APW / BPW = 0.99 - 1.02; lateral marginal depression obsolete, narrowly beaded laterally, broadly beaded basally, narrowly expanded inside front an gle; laterobasal depression with irregularly punctured surface, margined by narrow U-shaped depression. Proepisternum with 5 minute punctures along hind marginal groove; prosternal process with broad median depression, lateral margins broadly upraised. Elytra subovoid, disc convex, slides sloped to nearly vertical juncture with lateral marginal depression; basal groove slightly curved to subangulate humerus at juncture with broader lateral marginal depression, MEW / HuW = 2.23 - 2.43; parascutellar striole with 4 - 5 elongate punctures, striole continuous between punctures; sutural interval more convex than lateral intervals 2 - 4, sutural juncture upraised; sutural and 2 nd striae of subequal depth and similar punctation on disc, 2 nd stria reduced in depth and punctation both basally and in apical 1 / 3 of length; discal striae 2 - 7 progressively shallower, impressed portions shorter, striae 6 - 7 reduced to a series of punctures, or stria 7 absent; striae 3 - 7 absent from elytral apex, surface evenly convex between striae 2 and 8; 2 dorsal elytral setae at 0.26 x and 0.50 - 0.53 x elytral length, setal impressions moderate, shallow, spanning 1 / 2 width of interval 3; lateral elytral setae arranged in anterior series of 7 setae and posterior series of 6 setae; elytral marginal depression; broad, explanate laterad humerus, narrowed and beaded laterally to subapical sinuation; subapical sinuation very shallow, nearly obsolete. Mesepisternum with ~ 11 punctures in 2 - 3 rows; metepisternal width to length ratio = 0.74; metepisternum / metepimeron suture distinct. Abdomen with irregular lateral wrinkles on ventrites 1 - 6, round lateral depressions on ventrites 3 - 6; suture between ventrites 2 and 3 effaced; apical male ventrite with 2 marginal setae and apical female ventrite with 4 equally spaced setae plus median trapezoid of 4 subequal, short setae. Legs-metatarsomere 1 / metatibial length ratio = 0.19; metatarsomere 4 length along outer lobe 1.3 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci narrow, lateral, upper surface granulate, broadly convex. Microsculpture of vertex an obsolete transverse mesh, sculpticell breadth 2 x length, sculpticells most visible in depressed areas of cuticle; pronotal disc an obsolete transverse mesh, sculpticell breadth 2 - 3 x length traceable over depressed portions of disc but surface mostly glossy; elytral disc and apex glossy, microsculpture obsolete; metasternum with shallow transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex rufous with a piceous cast; antennomere 1 rufoflavous, antennomeres 2 - 11 rufobrunneous; pronotal disc dark rufous with piceous cast, lateral margins, base and apex rufopiceous; proepipleuron rufobrunneous, proepisternum rufous; elytral disc dark rufous, sutural interval rufous basally, rufoflavous apically; elytral lateral marginal depression piceous basally, rufoflavous toward apex; elytral apex rufoflavous apicad subapical sinuation; elytral epipleuron rufoflavous, metepisternum rufobrunneous; abdominal ventrites 1 - 6 medially rufous with piceous cast, laterally rufoflavous, apical 1 / 4 of ventrite 6 paler, rufoflavous. Male genitalia (n = 4). Aedeagal median lobe gracile, distance from parameral articulation to tip 3.5 - 3.8 x depth at midlength (Fig. 141 A-B); apex narrowly extended 3 x its depth beyond ostial opening, tip slightly downturned with apical face variably flattened (Fig. 141 A-B, D); median lobe straight at midlength along shaft, apex offset toward right so that convex left margin is apical in ventral view (Fig. 141 C); internal sac broadly expanded ventrally, a bipartite dorsal ostial microtrichial patch and ovoid ventral ostial microtrichial patch, each composed of heavily sclerotized macrospicules (Fig. 141 D-E); flagellar plate of moderate size, length 0.35 x parameral articulation-tip distance. Female reproductive tract (n = 1). Bursa copulatrix sac-shaped, slightly constricted basally, length 1.14 mm, maximum breadth 0.64 mm, basal constriction 0.43 mm broad (Fig. 136 D); bursal walls thick, broadly wrinkled; gonocoxite 1 with 3 - 5 apical fringe setae, a curved seta at medioapical angle and 5 - 7 setae on medial surface (Fig. 137 D); gonocoxite 2 falcate with long basal panhandle, apex subacuminate, 2 narrow, moderately elongate lateral ensiform setae, apical nematiform setae on medioventral surface at 0.64 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
12E3170F416CA172CF528AF7E8EBBE95.taxon	etymology	Etymology. This and the following Mecyclothorax brevidux represent cryptic sibling species best diagnosed by the length of the male aedeagal median lobe (Fig. 141). As both are members of the Mecyclothorax ducalis species group, the Latin stem dux-a leader-is used for both species. In this species dux is combined with the Latin longus to form the noun longidux.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
5F8632C4D63A63806B66339A2A25D229.taxon	diagnosis	Diagnosis. This species is extremely similar to Mecyclothorax longidux (Fig. 140 A-B), but can be diagnosed from that species by the less punctate discal striae, with small punctures present in impressed striae 1 - 2 to 1 - 3, and striae 4 - 5 reduced to linear series of small punctures (Fig. 140 C-D). The discal elytral intervals are less convex than in Mecyclothorax longidux, though the sutural interval is as upraised as in that species. The metafemur is darkened in its basal 1 / 5 to 1 / 3 of length, with the middle of the femur flavous, and apex narrowly piceous to match the darkened metatibia. The male aedeagal median lobe apex is shorter and broader (Fig. 141 F-I) than in males of Mecyclothorax longidux (Fig. 141 A-B, D). Setal formula 2 1 2 0. Standardized body length 5.2 - 5.9 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
5F8632C4D63A63806B66339A2A25D229.taxon	description	Description (n = 5). [The above description of Mecyclothorax longidux can serve to describe this species with the following substitutions.] Eyes with ocular ratio = 1.50 - 1.58, ocular lobe ratio = 0.81 - 0.84. Pronotum with lateral margins subparallel to slightly divergent for 0.1 x pronotal length anterad slightly obtuse, projected hind angles; MPW / PL = 1.15 - 1.21, MPW / BPW = 1.51 - 1.63, APW / BPW = 1.0 - 1.07. Elytra subovoid, MEW / HuW = 2.20 - 2.49. Microsculpture of elytral disc and apex minimally developed, with indistinct isodiametric and transverse sculpticells visible through the glossy reflection. Male genitalia (n = 4). Aedeagal median lobe gracile, distance from parameral articulation to tip 3.3 - 3.6 x depth at midlength (Figs 141 F-H); apex broad, extended twice maximum depth beyond ostial opening, apical face flattened, tip ventrally expanded so that ventral margin of lobe curves downward; internal sac short, broad, with lightly melanized dorsal ostial microtrichial patch and ventral ostial microtrichial patch (Fig. 141 I); flagellar plate of moderate size, length 0.39 x parameral articulation-tip distance (larger ratio of plate size relative to Mecyclothorax longidux due to shorter median lobe in Mecyclothorax brevidux; compare Fig. 141 D, I) Female reproductive tract (n = 1). Bursa copulatrix short, sac-shaped, length 1.0 mm, maximum width 0.51 mm, basal constriction 0.40 mm broad (Fig. 136 E); bursal walls thick, broadly wrinkled; gonocoxite 1 with 4 apical fringe setae, a curved seta at medioapical angle and 5 setae on medial surface (Fig. 137 E); gonocoxite 2 subacuminate, apex tightly rounded, basal extension curved at lateral terminus, 2 moderately elongate lateral ensiform setae, apical nematiform setae on medioventral surface at 0.74 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
5F8632C4D63A63806B66339A2A25D229.taxon	etymology	Etymology. This cryptic sister sibling species of Mecyclothorax longidux is given the epithet brevidux - short leader-in reference to the foreshortened aedeagal median lobe apex (Figs 141 F-I).	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
E421D27F035B49E65D29C9DACA8F5490.taxon	diagnosis	Diagnosis. Beetles of this species are larger than all others assigned to this species group, standardized body length 5.7 - 6.0 mm (Fig. 143 A), and are the only ones with a quadrisetose pronotum. Body coloration is uniformly dark, rufopiceous to rufobrunneous, with the elytral margins only narrowly rufoflavous. The discal elytral striae are distinctly punctate, with striae 1 - 3 impressed on the disc, and striae 4 - 5 indicated by more isolated punctures. Both the apical and subapical elytral setae are present. Setal formula 2 2 2 2.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
E421D27F035B49E65D29C9DACA8F5490.taxon	description	Description (n = 5). Head capsule with frontal grooves deep near clypeus, divergent mesad broad lateral convexity anterad eye, parallel mesad anterior supraorbital seta; dorsal impression of neck distinct, concave, visible in dorsal view; ocular lobe obtusely protruded from gena, eyes moderately convex, ocular ratio = 1.44 - 1.51, not covering posterior portion of ocular lobe, ocular lobe ratio = 0.76 - 0.81; labral anterior margin broadly emarginate to 1 / 6 of labral length; antennae filiform, antennomeres 2 - 3 with only a few setae basad apical rings; mentum tooth with sides acute, apex tightly rounded. Pronotum moderately transverse, MPW / PL = 1.20 - 1.25, lateral margins subparallel laterad basal pronotal setal articulatory sockets, hind angles obtuse, apex tightly rounded; median base depressed relative to disc,> 20 distinct punctures each side, the punctures elongate along juncture with disc; basal margin broadly convex between laterobasal depressions; median longitudinal impression very fine, obsolete on disc, distinct just anterad median base; anterior transverse impression finely incised, deep, small punctures in deepest part; anterior callosity convex, smooth; front angles not produced, rounded posterad front margin; pronotal apical width subequal to pronotal basal width, APW / BPW = 0.99 - 1.07; lateral marginal depression moderately broad, reflexed, broader with minute bead at front angle; laterobasal depression deep, surface irregular, continuous with lateral depression. Proepisternum with 5 distinct punctures along hind marginal groove; prosternal process with broad median depression, lateral margins broadly beaded between coxae. Elytra subquadrate, disc moderately convex, sides evenly sloped to lateral marginal depression; basal groove subangulate at sutural stria, extended laterally to rounded humerus defined by a hitch at the juncture of narrow basal groove and broader lateral depression, MEW / HuW ratio = 2.06 - 2.12; parascutellar seta present; parascutellar striole with 5 - 7 punctures, striole shallow to coplanar with disc between punctures; sutural interval more convex than lateral intervals throughout length; sutural stria traceable to basal groove, deep and narrow apically, 2 nd stria obsolete basally, shallow and broad apically; discal striae 3 - 5 progressively shallower and less extended apically, all absent from base, stria 6 represented by a few very shallow punctures or absent, stria 7 absent; 8 th interval convex 7 th stria at position of subapical elytral seta; 2 dorsal elytral setae at 0.29 x and 0.62 - 0.64 elytral length, setal impressions evident but spanning only 1 / 2 width of interval 3; apical and subapical setae present; lateral elytral setae arranged in anterior series of 7 setae and posterior series of 5 - 7 setae; elytral marginal depression narrow, edge slightly upraised at humerus, narrowed to bead at subapical sinuation; subapical sinuation shallow, more abruptly incurved apically. Mesepisternum with ~ 12 punctures in 2 - 3 rows; metepisternal width to length ratio = 0.69; metepisternum / metepimeron suture distinct. Abdomen with irregular lateral wrinkles on ventrites 1 - 6, lateral depressions on ventrites 3 - 6; suture between ventrites 2 and 3 complete; apical male ventrite with 2 marginal setae and apical female ventrite with 4 equally spaced setae and median trapezoid of 4 subequal, short setae. Legs-metatarsomere 1 / metatibial length ratio = 0.17; metatarsomere 4 length along outer lobe 1.3 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci narrow, lateral, median area broad. Microsculpture of vertex a shallow transverse mesh, sculpticell breadth 2 - 3 x length; pronotal disc with indistinct elongate transverse mesh to transverse lines, transverse sculpticell breadth 2 - 4 x length, median base with distinct transverse mesh between punctures, sculpticell breadth 2 - 3 x length; elytral disc and apex with distinct, regular transverse mesh, sculpticell breadth 2 - 3 x length; metasternum with obsolete transverse mesh, the surface glossy; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex rufobrunneous with piceous cast; antennomeres 1 - 3 rufoflavous, 4 - 11 rufobrunneous; pronotal disc rufopiceous, lateral margins narrowly, and base and apex more broadly rufobrunneous; proepipleuron rufobrunneous dorsally, rufoflavous along ventral margin, proepisternum dark rufobrunneous; elytral disc rufopiceous, sutural interval narrowly rufous along suture in basal 2 / 3, narrowly rufoflavous apically; elytral lateral marginal depression narrowly rufoflavous, elytral apex narrowly rufoflavous apicad subapical sinuation; elytral epipleuron dark rufoflavous, ventrally rufopiceous, metepisternum rufopiceous with cupreous reflection; abdominal ventrites 1 - 6 rufobrunneous mediobasally, rufoflavous on lateral and apical margins except apical 1 / 3 of ventrite 6 paler, rufobrunneous; metafemur rufoflavous with basal piceous cloud; metatibia rufoflavous with brunneous cast. Male genitalia (n = 4). Aedeagal median lobe elongate, gracile, distance from parameral articulation to tip 4.2 x depth at midlength (Fig. 144 A, C); apex elongate, narrowly extended 5 x its minimum depth beyond ostial opening, the tip expanded dorsally as a blunt tooth, and ventrally as rounded tip with oblique, straight, apical face; median lobe straight in ventral view, but apex offset to right side, dorsal (left) margin straight in ventral view, but apex offset to right side, dorsal (left) margin angled to meet thin apex (Fig. 144 B); internal sac broad, breadth about 0.67 x distance from ventral ostial margin to base of flagellar plate, a broad, transverse dorsal ostial microtrichial patch defined by shingled macrospicules present near base; flagellar plate moderately long, length 0.42 x parameral articulation-tip distance. Female reproductive tract (n = 1). Bursa copulatrix columnar with expanded apex, length 1.50 mm, apical breadth 0.68 mm, basal breadth 0.34 mm (Fig. 145 A); bursal walls translucent, thickly wrinkled; gonocoxite 1 with 4 - 5 apical fringe setae, the medial seta smaller, a curved seta at medioapical angle and 6 - 7 smaller setae on medial surface (Fig. 146 A); gonocoxite 2 falcate with subacuminate apex, broad basal extension little curved at lateral terminus, 2 lateral ensiform setae, apical nematiform setae on medioventral surface at 0.71 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
E421D27F035B49E65D29C9DACA8F5490.taxon	etymology	Etymology. This species resembles Mecyclothorax vulcanus (Blackburn) and Mecyclothorax hephaestus Liebherr of Hawai'i Island (Liebherr 2008 b), leading to use of hephaestoides as the species epithet.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
97B5320043CCBEB0205C4ABC82185510.taxon	diagnosis	Diagnosis. This species is one of the few Hawaiian Mecyclothorax that lacks dorsal elytral setae. Among the Haleakala fauna only Mecyclothorax xestos (Fig. 90 D) of the Mecyclothorax microps group shares this feature. The parascutellar seta is absent, a unique occurrence within the Mecyclothorax palustris group, and the pronotum is broader relative to the elytra (Fig. 143 B), MEW / MPW = 1.44, than in any other species with concolorous elytral disc and margins in the group. The bicolored Mecyclothorax bicoloris and Mecyclothorax bicoloratus exhibit similar somite proportions (Fig. 143 C-D). The elytral microsculpture is a well-developed mesh of isodiametric sculpticells arranged in transverse rows on the inner intervals, the sculpticells more transverse on intervals 7 and 8. Setal formula 2 1 0 1 [sae]. Standardized body length 4.2 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
97B5320043CCBEB0205C4ABC82185510.taxon	description	Description (n = 1). Head capsule with frontal grooves broadest near clypeus, expanded medially inside broad convexity anterad eyes, terminated posteriorly at thin carina mesad supraorbital seta; dorsal impression of neck slightly concave; ocular lobe barely protruded from gena, eyes relatively flat, ocular ratio = 1.43, ocular lobe ratio = 0.81; labral anterior margin angularly emarginate 1 / 6 labral length; antennae filiform, antennomeres 2 - 3 with sparse pelage of short setae; mentum tooth with sides acute, apex tightly rounded. Pronotum with lateral margins convex anterad a deep sinuation at the right hind angles, the lateral margins parallel for 1 / 6 pronotal length, MPW / PL = 1.21, MPW / BPW = 1.61; median base nearly coplanar with disc, shallow strigose wrinkles each side; basal margin very slightly convex between laterobasal depressions; median longitudinal impression very shallow, middle of disc flat; anterior transverse impression broad, shallow medially, more marked laterally, finely incised mesad front angles; anterior callosity moderately convex, smooth; front angles slightly projected, tightly rounded; pronotal apex broader than base, APW / BPW = 1.07; lateral marginal depression narrow, beaded laterally, edge upturned and broader at front angles; laterobasal depression smooth, continuous with lateral depression. Proepisternum with 5 minute punctures along hind marginal groove; prosternal process with broad median depression, lateral margins broadly beaded between coxae. Elytra subquadrate, moderately broad basally, disc flat, sides moderately sloped; basal groove angulate at sutural stria, extended to angulate humerus defined by juncture of basal groove and much broader lateral marginal depression, MEW / HuW = 1.95; parascutellar striole with 3 - 4 punctures, striole very shallow between punctures; sutural interval more convex than lateral intervals, sutural juncture upraised; sutural and 2 nd striae of subequal depth and punctation on disc, sutural stria deeper and more finely incised at apex; discal intervals 2 - 5 slightly convex, striae 2 - 5 progressively shallower and less punctate laterally, stria 6 a series of punctures, and stria 7 marked by sparse elongate irregularities at midlength, shallowly continuous apically; 8 th interval of similar convexity to apical fused portion of striae 5 + 7; subapical seta present in apical portion of stria 7 laterad apex of stria 2; lateral elytral setae arranged in anterior series of 7 setae and posterior series of 6 setae; elytral marginal depression moderately narrow posterad humerus, edge slightly upturned there, narrowed to a bead at subapical sinuation; subapical sinuation shallow, more abruptly incurved anteriorly. Mesepisternum with ~ 13 punctures in 2 - 3 rows; metepisternal width to length ratio = 0.76; metepisternum / metepimeron suture distinct. Abdomen with irregular lateral wrinkles on ventrites 1 - 5 and lateral depressions on ventrites 3 - 6; suture between ventrites 2 and 3 complete; apical female ventrite with 4 equally spaced setae and median trapezoid of 4 subequal, short setae; Legs-metatarsomere 1 / metatibial length ratio = 0.19; metatarsomere 4 length along outer lobe 1.3 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci broad, shallow, basal tarsomere medially subcarinate. Microsculpture of vertex a shallow transverse mesh, sculpticell breadth 2 - 3 x length; pronotal disc with shallow transverse mesh, sculpticell breadth 2 - 4 x length, median base with distinct isodiametric mesh; metasternum with a shallow transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration (lone holotype specimen is teneral) of vertex rufoflavous; antenna rufoflavous basally, dusky rufoflavous apically; pronotal disc rufoflavous, lateral margins narrowly, and base and apex flavous; proepipleuron flavous, proepisternum rufobrunneous with rufoflavous margins; elytral disc rufoflavous, sutural interval flavous basally and apically, concolorous on disc; elytral marginal depression narrowly flavous, apex gradually paler, flavous from apical terminus of interval 4; elytral epipleuron flavous, metepisternum rufobrunneous; abdominal ventrites 1 - 5 rufoflavous, apical half of ventrite 6 flavous; metafemur flavous; metatibia flavous with rufoflavous cast. Female reproductive tract. The lone female specimen was not dissected.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
97B5320043CCBEB0205C4ABC82185510.taxon	etymology	Etymology. The very small eyes characterizing this species are the basis for using oculellus - small eye-as the species epithet. The diminutive oculellus was used instead of ocellus due to the very specific meaning of the latter term in entomology.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
1C4243FBCE24DD0B84BCEF418B6202BE.taxon	diagnosis	Diagnosis. This species (Fig. 143 C) and Mecyclothorax bicoloratus (Fig. 143 D) comprise a cryptic sibling species pair diagnosed by: 1, broadly flavous elytral margins contrasted to a rufopiceous disc; and 2, punctate discal elytral striae 1 - 5. The strial punctures vary in size (Fig. 143 C-D), though unfortunately for the taxonomist, not in tandem with the diagnostic male aedeagal median lobe. The median lobe apex for Mecyclothorax bicoloris males is ventrally pointed (Fig. 144 D, F-G), whereas the lobe apex in Mecyclothorax bicoloratus males is broadly rounded (Fig. 144 H-I). Two subtle external characters allow these two species to be diagnosed, though not with the confidence based on a male dissection. Firstly, the microsculpture of the vertex of Mecyclothorax bicoloris individuals comprises an indistinct transverse mesh, sculpticell breadth 2 - 3 x length, with areas of microsculpture interspersed with glossy areas. The sculpticell margins are not visible in areas of reflected microscope light. Mecyclothorax bicoloratus individuals, in contrast, exhibit a well-developed transverse mesh on the vertex, with the convex surfaces of individual sculpticells discernible, and the entire surface less glossy. Secondly, the elytral lateral marginal groove is narrower basally just laterad the humeral angle in Mecyclothorax bicoloris (Fig. 143 C), whereas it is broader with a more elevated margin in Mecyclothorax bicoloratus (Fig. 143 D). Setal formula 2 1 2 0. Standardized body length 4.1 - 4.6 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
1C4243FBCE24DD0B84BCEF418B6202BE.taxon	description	Description (n = 5). Head capsule with frontal grooves deepest in anterior half of length where they are parallel on the frons, grooves narrower and shallower posterad a lateral jog in their direction, grooves terminated mesad narrow carina inside anterior supraorbital seta; dorsal impression of neck slightly concave; ocular lobe distinctly protruded from gena, eyes moderately convex and large, ocular ratio = 1.48 - 1.53, ocular lobe ratio = 0.85 - 0.91; labral anterior margin angularly emarginate 1 / 8 of labral length; antennae filiform; antennomeres 2 - 3 with sparse pelage of short setae; mentum tooth with sides acute, apex tightly rounded. Pronotum transverse, MPW / PL = 1.15 - 1.29, lateral margins distinctly sinuate for short distance anterad right to slightly acute hind angles, MPW / BPW = 1.56 - 1.70; median base distinctly depressed relative to disc, with ~ 20 small, isolated punctures each side; basal margin slightly convex medially, straight behind laterobasal depressions; median longitudinal impression very shallow, middle of disc flat, crossed by indistinct transverse wrinkles; anterior transverse impression broad, shallow, smooth, obso lete medially, incised immediately mesad front angles; anterior callosity nearly flat, slightly convex, smooth; front angles slightly projected, tightly rounded; pronotal apical width slightly greater than basal width, APW / BPW = 1.01 - 1.09; lateral marginal depression narrow, edge upturned anterad lateral seta, a bit broader at front angle; laterobasal depression narrow with irregular surface, continuous with lateral depression. Proepisternum with 5 minute punctures along hind marginal groove; prosternal process with narrow median impression, lateral margins broadly upraised. Elytra subquadrate, disc slightly convex, sides moderately sloped to margins; basal groove angulate at sutural stria, extended directly to angulate humerus at juncture of basal groove and broader lateral depression, MEW / HuW = 2.04 - 2.12; parascutellar seta present; parascutellar striole with 5 punctures, striole shallow but continuous between punctures; sutural interval coplanar basally, upraised apically; sutural and 2 nd striae of subequal depth and punctation on disc; sutural stria continued to base whereas striae 2 - 5 obsolete basally; at elytral apex sutural stria deep and smooth, striae 2 - 3 and 7 traceable apically but very shallow; discal striae 2 - 6 progressively shallower laterally, punctures more isolated and associated intervals flatter, stria 7 absent at elytral midlength; 8 th interval of similar convexity to apical fused portion of striae 5 + 7; 2 dorsal elytral setae at 0.26 x and 0.65 x elytral length, setal impressions small, shallow, spanning 1 / 2 width of interval 3; apical and subapical setae absent; lateral elytral setae arranged in anterior series of 7 setae and posterior series of 6 setae; elytral marginal depression narrow laterad humerus with upturned margin, narrower and beaded apically at subapical sinuation; subapical sinuation very shallow, symmetri cal. Mesepisternum with ~ 9 punctures in 2 - 3 rows; metepisternal width to length ratio = 0.65; metepisternum / metepimeron suture distinct. Abdomen with irregular lateral wrinkles on ventrites 1 - 5, lateral depressions on ventrites 3 - 6; suture between ventrites 2 and 3 complete; apical male ventrite with 2 marginal setae and apical female ventrite with 4 equally spaced setae and median trapezoid of 4 subequal, short setae. Legs-metatarsomere 1 / metatibial length ratio = 0.19; metatarsomere 4 length along outer lobe 1.3 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci broad, shallow, basal tarsomere medially subcarinate. Microsculpture of pronotal disc shallow transverse mesh to transverse lines between areas of glossy cuticle, median base glossy between punctures, indistinct transverse sculpticells over parts of cuticle; elytral disc with transversely stretched isodiametric sculpticells, sculpticells up to 2 x broad as long, apex with mixture of isodiametric and transverse sculpticells; metasternum with shallow transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse sculpticells. Coloration of vertex rufobrunneous; antennomeres 1 - 3 flavous, 4 - 11 rufoflavous; pronotal disc rufobrunneous, lateral margins, base, and apex rufoflavous; proepipleuron rufoflavous, proepisternum dorsally rufoflavous, ventrally rufobrunneous; elytral disc basally rufobrunneous to rufopiceous, lateral margins 7 - 9 contrastedly flavous, sutural interval rufoflavous basally, flavous apically; elytral apex broadly flavous, the paler area matching or exceeding the breadth of the pale lateral margin; elytral epipleuron flavous dorsally, rufoflavous ventrally, metepisternum rufobrunneous; abdomen with ventrites 1 - 3 rufobrunneous medially, rufoflavous laterally, ventrites 4 - 6 basally rufobrunneous, apically flavous; metafemur flavous; metatibia rufoflavous. Male genitalia (n = 3). Aedeagal median lobe moderately stout, distance from parameral articulation to tip 3.0 x depth at midlength (Fig. 144 D, F); apex extended 2 - 3 x its minimum depth beyond ostial opening, dorsal and ventral margins parallel until tip that is acutely angled and slightly downturned, the apical face convex; median lobe straight, right and left margins similarly convergent in ventral view, the apex with oblique, blunt tip in this view (Fig. 144 E); internal sac about twice as long as broad, with broad, diffuse dorsal ostial microtrichial patch near base, and ventral sac surface covered with brownish microspicules (Fig. 144 G); flagellar plate of moderate size, length 0.40 x parameral articulation-tip distance. Female reproductive tract (n = 1). Bursa copulatrix ovate-like an insect net in profile (Fig. 145 B) - with length 0.72 mm, basal breadth 0.51 mm, midlength breadth 0.34 mm; bursal walls translucent, thickly wrinkled; gonocoxite 1 with 3 - 4 apical fringe setae, a curved seta at medioapical angle and 6 - 7 setae on medial surface (Fig. 146 B); gonocoxite 2 falcate with subacuminate apex, basal extension curved at terminus, 2 lateral ensiform setae, apical nematiform setae on medioventral surface at 0.75 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
1C4243FBCE24DD0B84BCEF418B6202BE.taxon	etymology	Etymology. The species epithet bicoloris refers to the bicolored elytra of members of this species. The epithet is the genitive singular form of bicolor, and its ending does not change with gender.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
DA4DFD566452936E1A7D3A98884470E1.taxon	diagnosis	Diagnosis. This species (Fig. 143 D) shares the bicolored elytra, punctate discal elytral striae, and setal formula 2 1 2 0 with its cryptic sibling species, Mecyclothorax bicoloris (Fig. 143 C). As detailed under that species' treatment, the more intensely microsculptured vertex can externally diagnose individuals of this species, with the isodiametric sculpticells upraised, their individual reflections producing a textured look to the surface. Also, the elytral lateral marginal depression is broader laterad the humerus in this species (Fig. 143 D). The aedeagus of this species has a broadly rounded median lobe apex (Fig. 144 H-I) instead of the ventrally pointed apex of Mecyclothorax bicoloris males (Fig. 144 D-G). Standardized body length 4.1 - 4.8 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
DA4DFD566452936E1A7D3A98884470E1.taxon	description	Description (n = 5). [The above description of Mecyclothorax bicoloris can serve to describe this species. All diagnostic characters are presented above, so only the various recorded ratios are presented below.] Eyes moderately convex, covering much of the protruded ocular lobes, ocular ratio = 1.44 - 1.50, ocular lobe ratio = 0.79 - 0.84. Pronotum transverse, MPW / PL = 1.18 - 1.30, lateral margins distinctly sinuate for short distance anterad right to slightly acute hind angles, MPW / BPW = 1.54 - 1.64. Elytra narrowly subquadrate, MEW / HuW = 2.05 - 2.14. Male genitalia (n = 2). Aedeagal median lobe gracile, distance from parameral articulation to tip 3.7 x depth at midlength (Fig. 144 H-I); apex parallel sided, of variable breadth, extended 1.5 - 2.7 x minimum depth beyond ostial opening, tip with apical face less convex dorsad rounded juncture with ventral margin; internal sac with evident, broadly diffuse dorsal ostial microtrichial patch (uneverted specimen, Fig. 144 H); flagellar plate of moderate size, length 0.35 x parameral articulation-tip distance (uneverted specimens, Fig. 144 H-I). Both figured specimens were collected at State Fence Camp on Helele'ike'oha Stream, and as there are no other features that diagnose these males, variation in the breadth of the median lobe and its apex is considered infraspecific. Female reproductive tract (n = 1). Bursa copulatrix broad basally, with parallel-sided apical lobe with rounded apex, overall length 0.76 mm, apical lobe 0.60 mm long x 0.29 mm broad, base at vagina 0.38 mm broad (Fig. 145 C); bursal walls translucent, thickly wrinkled; gonocoxite 1 with 3 - 4 apical fringe setae, the medial 1 or 2 smaller (Fig. 146 C), a curved seta at apicomedial angle and 9 - 10 setae on medial surface; gonocoxite 2 narrowly subtriangular, with apex curved laterad, lateral margin straight near ensiform setae and base moderately extended laterally, 2 lateral ensiform setae, apical nematiform setae on medioventral surface at 0.76 - 0.79 gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
DA4DFD566452936E1A7D3A98884470E1.taxon	etymology	Etymology. The adjectival species epithet bicoloratus is used to name the second bicolored species in the Mecyclothorax palustris group, adding to the examples of sibling species named with the same word stem.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
FA287ABEA9EA5C31EAAA8878A3202066.taxon	diagnosis	Diagnosis. Among the four species in this group with individuals exhibiting contrastedly paler elytral margins, this species (Fig. 149 A) is diagnosed by the very shallow elytral striae, with only the sutural and 2 nd striae impressed on the disc. The pronotal median base is less punctate than in the two bicolored species with punctate striae; Mecyclothorax bicoloris (Fig. 143 C) and Mecyclothorax bicoloratus (Fig. 143 D). In this species the pronotum has ~ 16 isolated punctures each side, and the surface of the pronotal laterobasal depression is smooth. The male aedeagal internal sac is bilobed in this species, with the basal lobe subequal in length to the apical lobe (Fig. 150 E-F). Setal formula 2 1 2 0. Standardized body length 3.9 - 4.7 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
FA287ABEA9EA5C31EAAA8878A3202066.taxon	description	Description (n = 5). Head capsule with frontal grooves broad and deep at clypeus, straight and subparallel, terminated at a low carina mesad anterior supraorbital seta; dorsal impression of neck slightly concave; ocular lobe obtusely protruded from gena, eyes convex, ocular ratio = 1.54 - 1.60, ocular lobe ratio = 0.78 - 0.81; labral anterior margin angularly emarginate 1 / 6 of labral length; antennae filiform, antennomeres 2 - 3 with sparse pelage of short setae; mentum tooth with sides acute, apex tightly rounded. Pronotum slightly transverse, MPW / PL = 1.15 - 1.20, basally constricted, MPW / BPW = 1.53 - 1.64, lateral margins briefly sinuate anterad acute, projected hind angles; median base depressed relative to disc; basal margin indistinctly trisinuate, slightly convex medially; median longitudinal impression very shallow, broad, crossed by indistinct transverse wrinkles; anterior transverse impression broad, very shallow medially, incised laterally mesad front angles; anterior callosity flat, slightly upraised, smooth; front angles not projected, rounded posterad curve of front margin; pronotal apical width variably subequal to broader than pronotal basal width, APW / BPW = 0.97 - 1.13; lateral marginal depression moderate, edge upturned at hind angle, beaded anterad inside basal sinuation, depression not wider at front angle. Proepisternum with 5 minute punctures along hind marginal groove; prosternal process with broad median depression, lateral margins broadly beaded between coxae. Elytra subovoid, disc convex, sides distinctly sloped to margins; basal groove evenly curved to angulate humerus, MEW / HuW = 2.14 - 2.36; parascutellar seta present; parascutellar striole with 4 isolated punctures; sutural interval coplanar with lateral intervals basally, upraised in apical half; sutural stria deeper than 2 nd stria throughout length, deeper and more punctate on disc, a series of isolated punctures near basal groove, and deep, narrow, and smooth at elytral apex; discal striae 2, or 2 - 3 traceable as linear series of shallow punctures, lateral striae traceable only as disturbances in the microsculpture, or melanic spots in the cuticle; striae 2, 3 and the apex of 7 shallow and traceable at elytral apex, area between striae 2 and 7 evenly convex there; 2 dorsal elytral setae at 0.26 - 0.28 x and 0.49 - 0.57 x elytral length, setal impressions small, shallow, spanning 1 / 2 width of interval 3; apical and subapical setae absent; lateral elytral setae arranged in anterior series of 7 setae and posterior series of 6 setae; elytral marginal depression narrow at humerus, margin upturned, beaded posteriorly near subapical sinuation; subapical sinuation shallow, more abruptly incurved anteriorly. Mesepisternum with 5 punctures in 1 row; metepisternal width to length ratio = 0.68; metepisternum / metepimeron suture distinct. Abdomen with irregular lateral wrinkles on ventrites 1 - 5, lateral depressions on ventrites 3 - 6; suture between ventrites 2 and 3 complete; apical male ventrite with 2 marginal setae and apical female ventrite with 4 equally spaced setae and median trapezoid of 4 subequal, short setae. Legs-metatarsomere 1 / metatibial length ratio = 0.18; metatarsomere 4 length along outer lobe 1.3 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci broad, shallow, median area broad. Microsculpture of vertex a shallow transverse mesh, sculpticell breadth 2 - 3 x length; pronotal disc with obsolete transverse mesh, sculpticell breadth 2 - 4 x length, median base glossy medially, indistinct isodiametric sculpticells laterally; elytral disc and apex with indistinct isodiametric sculpticells in transverse rows; metasternum with shallow transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse sculpticells. Coloration of vertex rufous; antennomeres 1 - 3 flavous, 4 - 11 rufoflavous; pronotal disc rufobrunneous, lateral margins, base, and anterior callosity rufoflavous; proepipleuron flavous, proepisternum rufobrunneous with rufoflavous margins; elytral disc rufopiceous, base rufobrunneous, sutural interval basally rufous, apical half rufoflavous, marginal intervals 7 - 9 contrastedly flavous, apex flavous apicad the posterior seta of the lateral setal series; elytral epipleuron flavous, metepisternum rufobrunneous; abdomen with ventrite 1 rufoflavous, ventrites 2 - 3 rufopiceous with flavous apex, ventrites 4 - 6 basally rufobrunneous, apical 3 / 4 of apical ventrite 6 flavous; metafemur flavous; metatibia flavous with rufoflavous cast. Male genitalia (n = 7). Aedeagal median lobe robust, distance from parameral articulation to tip 2.5 - 2.8 x depth at midlength (Fig. 150 A, C-F); apex a smoothly curved extension of ventral margin, the extension of variable depth (Fig. 150 A versus C-F); median lobe broad, symmetrical in ventral view, the convex dorsal margin (Fig. 150 A) visible as convex apical margin in this view (Fig. 150 B); internal sac with well-developed basal lobe that is subequal in length to apical lobe (Fig. 150 E-F), the lobe varying in breadth; sac with well-developed microspicules on ventral surface, a very diffuse patch of somewhat larger, lightly melanized macrospicules present near base (Fig. 150 E); flagellar plate of moderate size, length 0.38 - 0.40 x parameral articulation-tip distance. Female reproductive tract (n = 1). Bursa copulatrix broad basally, with parallel-sided apical lobe with rounded apex, overall length 0.76 mm, apical lobe 0.56 mm long x 0.34 mm broad, base at vagina 0.55 mm broad (Fig. 145 D); bursal walls translucent, thinly wrinkled; gonocoxite 1 with 3 apical fringe setae, 7 - 8 smaller setae on medial surface (146 D); gonocoxite 2 falcate, with tightly rounded tip, base with long thick lateral extension that curves at terminus, 2 lateral ensiform setae, apical nematiform setae on medial surface at 0.76 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
FA287ABEA9EA5C31EAAA8878A3202066.taxon	etymology	Etymology. The adjectival species epithet bilobatus refers to the bilobed internal sac of the male aedeagus (Figs 150 E-F).	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
F7E0E87C1F9535141C7C6BB08DEC8821.taxon	diagnosis	Diagnosis. This species can be diagnosed by the impressed elytral striae 1 - 3 that are irregular along their length but not distinctly punctate, and the broadly flavous elytral apical, lateral, and sutural margins (Fig. 149 B). The elytral coloration varies infraspecifically. The lateral intervals 7 - 9 are flavous in contrast to the rufopiceous discal intervals 2 - 6 in some individuals. Alternatively, the elytra may exhibit gradually paler lateral margins, with intervals 7 or 7 - 8 rufobrunneous mesad the more flavous 9 th interval and lateral marginal depression. The breadths of the flavous apical and lateral marginal bands are positively associated, with the apex variably pale from beyond the posterior seta of the lateral elytral setal series, or more narrowly pale beyond the apical fusion of striae 3 + 4 (Fig. 149 B). Individuals with differing degrees of pale margination cooeccur within the same collecting series. Individuals of this species are most similar to those of Mecyclothorax tauberorum and Mecyclothorax pau, though both of those species are characterized by darker, less contrasted lateral elytral margins. Individuals of Mecyclothorax palustroides are all larger than those of Mecyclothorax tauberorum; standardized body length for this species = 4.6 - 5.5 mm versus s. b. l. = 3.9 - 4.6 mm for Mecyclothorax tauberorum. Mecyclothorax palustroides can be diagnosed from Mecyclothorax pau by the more upraised sculpticells in the transverse discal elytral microsculpture, imparting an alutaceous sheen to the surface, and by the more broadly paler elytral intervals 8 - 9 posterad the humerus; that pale margin either rufobrunneous or flavous versus the rufopiceous disc. Also, the male aedeagal median lobe can definitively diagnose the 3 species: 1, Mecyclothorax palustroides with the lobe apex broadly rounded (Fig. 152); 2, Mecyclothorax tauberorum with the lobe apex short and bluntly rounded (Fig. 161 A, C-D); and 3, Mecyclothorax pau with the lobe apex elongate, narrowly rounded (Fig. 161 E-F, H-L). Setal formula 2 1 2 0.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
F7E0E87C1F9535141C7C6BB08DEC8821.taxon	description	Description (n = 5). Head capsule with frontal grooves broad near clypeus, triangularly expanded medially, and divergent to terminate mesad fine carina inside anterior supraorbital seta; dorsal impression of neck slightly concave; ocular lobe obtusely protruded from gena, ocular ratio = 1.50 - 1.58, ocular lobe ratio = 0.81 - 0.84; labral anterior margin angularly emarginate to 1 / 9 labral length; antennae filiform, antennomeres 2 - 3 with sparse pelage of short setae; mentum tooth with sides acute, apex tightly rounded. Pronotum slightly transverse, MPW / PL = 1.08 - 1.21, variably constricted basally, MPW / BPW = 1.52 - 1.68; lateral margins convergent for short distance anterad right hind angles, the basal margin convex just mesad hind angle; median base distinctly depressed relative to disc, elongate punctures bordering disc, ~ 19 isolated punctures each side; basal margin trisinuate, slightly convex medially; median longitudinal impression very shallow, middle of disc flat; anterior transverse impression obsolete medially, finely incised laterally, fine longitudinal wrinkles extended from impression across flat anterior callosity; front angles slightly projected, tightly rounded; pronotal apical width variably broader than pronotal basal width, APW / BPW = 1.02 - 1.18; lateral marginal depression narrow, edge upturned laterally, slightly broader at front angle; laterobasal depression narrow, surface irregular, continuous with lateral depression. Proepisternum with 5 minute punctures along hind marginal groove; prosternal process with narrow median impression, lateral margins broadly upraised. Elytra subovoid, lateral margins rounded posterad humeral angles, disc convex, sides distinctly sloped to marginal depression; basal groove evenly curved to subangulate humerus, MEW / HuW = 2.21 - 2.33; parascutellar seta present; parascutellar striole with 4 - 5 punctures, striole very shallow between punctures; sutural interval coplanar with lateral intervals basally, upraised in apical half; sutural and 2 nd striae of similar depth on disc, sutural interval continued as isolated punctures at base, sutural stria deep, smooth and finely incised apically, 2 nd stria broader and irregularly interrupted apically; discal striae 2 - 5 progressively shallower, inner striae irregular, lateral striae represented by isolated punctures, striae 6 - 7 traceable only as series of very shallow punctures at midlength; mesal intervals slightly convex, lateral intervals flat; apex with striae 1, 2, 7, and 8 present, striae 3 - 6 obsolete, though intermittently traceable; 8 th interval slightly convex laterad 7 th stria mesad subapical sinuation; 2 dorsal elytral setae at 0.27 x and 0.54 - 0.64 x elytral length, setal impressions small, shallow, spanning 1 / 2 width of interval 3; lateral elytral setae arranged in anterior series of 7 setae and posterior series of 6 setae; elytral marginal depression moderately broad with edge upturned laterad humerus, narrowed to a beaded margin at subapical sinuation; subapical sinuation shallow, more abruptly incurved anteriorly. Mesepisternum with ~ 9 - 10 punctures in 2 - 3 rows; metepisternal width to length ratio = 0.71; metepisternum / metepimeron suture distinct. Abdomen with irregular lateral wrinkles on ventrites 1 - 5, lateral depressions on ventrites 3 - 6; suture between ventrites 2 and 3 complete; apical male ventrite with 2 marginal setae and apical female ventrite with 4 equally spaced setae and median trapezoid of 4 subequal, short setae. Legs-metatarsomere 1 / metatibial length ratio = 0.20; metatarsomere 4 length along outer lobe 1.3 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci broad, shallow, basal tarsomere medially subcarinate. Microsculpture of vertex a distinct transverse mesh, sculpticell breadth 2 - 3 x length; pronotal disc with shallow transverse mesh, sculpticell breadth 3 - 4 x length, median base with indistinct transverse mesh, sculpticell breadth 2 x length; elytral apex with shallow isodiametric and transverse sculpticells, sculpticell breadth 2 x length; metasternum with shallow transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex rufobrunneous with piceous cast; antennomeres 1 - 3 rufoflavous, 4 - 11 rufobrunneous; pronotal disc rufobrunneous with piceous cast, lateral margins narrowly, and base and apex rufoflavous; proepipleuron rufoflavous, proepisternum dorsally rufoflavous, ventrally rufobrunneous; elytral disc dark rufobrunneous with iridescent sheen, sutural interval rufous in basal half, rufoflavous in apical half; elytral epipleuron rufoflavous, metepisternum rufobrunneous; abdomen with ventrites 1 - 3 rufopiceous medially, rufoflavous laterally, ventrites 4 - 6 basally rufobrunneous, apically flavous, the apical ventrite flavous in apical 3 / 4; metafemur flavous; metatibia flavous with rufoflavous cast. Male genitalia (n = 19). Aedeagal median lobe gracile to slightly robust, dorsal and ventral margins subparallel along median shaft, distance from parameral articulation to tip 3.2 - 4.1 x depth at midlength (Fig. 152 A, C-H); apex broadly and briefly extended, tip slightly curved apically along ventral margin, slightly expanded along dorsal margin, with obliquely convex apical face; median lobe broadly, evenly curved rightward apically in ventral view (Fig. 152 B, F), the concave right margin and convex left margin convergent to blunt, oblique tip; internal sac with very pale, diffuse dorsal ostial microtrichial patch near base, and variably ornamented ventral surface, covered either with shaggy pelage of microspicules (Fig. 152 G), or a ventral ostial microtrichial patch composed of overlapping scaly macrospicules (Fig. 152 H); flagellar plate moderately sized, length 0.40 x parameral articulation-tip distance. Female reproductive tract (n = 1). Bursa copulatrix columnar with basally constricted apical lobe, length 1.08 mm, apical cap 0.40 mm long x 0.29 mm broad, width at midlength 0.40 mm (Fig. 145 E); bursal walls translucent, thickly wrinkled; gonocoxite 1 with 4 apical fringe setae, a curved seta near medioapical angle and 3 - 5 smaller setae on medial surface (Fig. 146 E); gonocoxite 2 falcate, apex broad with sensilla doubled along lateral margin, base broadly extended laterally, 2 lateral ensiform setae, apical nematiform setae on medioventral surface at 0.77 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
F7E0E87C1F9535141C7C6BB08DEC8821.taxon	etymology	Etymology. This species' resemblance to Mecyclothorax palustris (Sharp) leads to the use of palustroides as the species epithet. The stem palustris describes marshy or swampy situations.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
FE2538D37910C181A265F6AD8511E86E.taxon	diagnosis	Diagnosis. Among Haleakala's Mecyclothorax palustris group species, Mecyclothorax filipoides stands out based on the disparate dorsal microsculpture: 1, head and pronotum with glossy surfaces, their microsculpture respectively a shallow transverse mesh, and obsolete transverse mesh to transverse lines; 2, elytral disc and apex with coriaceous surface comprised of isodiametric or slightly transversely stretched sculpticells arranged in irregular transverse rows. The elytral disc and lateral intervals are rufobrunneous with the elytral apex only indistinctly paler. The discal elytral striae are shallow, with the punctures of the inner striae isolated, and the sutural stria the most complete (Fig. 149 C). Setal formula 2 1 2 0. Body size is large among species of the group; standardized body length 4.7 - 5.5 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
FE2538D37910C181A265F6AD8511E86E.taxon	description	Description (n = 5). Head capsule with frontal grooves narrow and deep anteriorly, sinuous laterally and shallower until terminated posteriorly mesad thin carina that borders depression surrounding anterior supraorbital seta; dorsal impression of neck slightly concave; ocular lobes obtusely protruded from gena, eyes large, ocular ratio = 1.52 - 1.59, ocular lobe ratio = 0.82 - 0.88; labral anterior margin angularly emarginate 1 / 7 labral length; antennae filiform, antennomeres 2 - 3 with sparse pelage of short setae; mentum tooth with sides acute, apex tightly rounded. Pronotum appearing quadrate, MPW / PL = 1.12 - 1.19, lateral margins slightly convergent anterad acute hind angles for 1 / 6 of pronotal length, base moderately constricted, MPW / BPW = 1.47 - 1.56; median base slightly depressed medially, more so laterally, ~ 25 isolated punctures each side, those along juncture of base and disc elongate; basal margin trisinuate, slightly convex medially and expanded posteriorly behind laterobasal depressions; median longitudinal impression shallow, finely incised, smooth, continued onto median base; anterior transverse impression broad, very shallow medially, well-marked laterally mesad front angles; anterior callosity flat, gradually elevated from transverse impression to front margin, crossed by minute longitudinal wrinkles; front angles projected, narrowly rounded; pronotal apical width slightly less than pronotal basal width, APW / BPW = 0.95 - 0.99; lateral marginal depression moderate, edge beaded laterally to basal sinuation, more broadly upraised outside laterobasal depressions; laterobasal depression minutely punctate, continuous with lateral depression. Proepisternum with 4 distinct punc tures along hind marginal groove; prosternal process with broad median depression, lateral margins broadly margined between coxae. Elytra subquadrate, disc convex, sides distinctly sloped to margins; basal groove evenly curved to subangulate to angulate humerus defined by hitch in margin at juncture of narrow basal groove and broader lateral marginal depression, MEW / HuW = 2.09 - 2.24; parascutellar seta present; parascutellar striole with 5 large punctures, striole shallow to interrupted between punctures; sutural interval broadly domed to elevated sutural juncture; sutural stria punctate, deeper than 2 nd stria on disc, finely incised, deep, and smooth apically, where 2 nd stria is broader and shallower, and irregularly interrupted; discal striae 2 - 4 progressively shallower laterally with elongate punctures progressively isolated, associated intervals flat, appearing concave on middle of disc; striae 1, 2, and 7 present on elytral apex, striae indicated by series of isolated punctures, 8 th interval slightly more convex than 7 th mesad subapical sinuation; 2 dorsal elytral setae at 0.15 - 0.18 x and 0.53 - 0.60 x elytral length, setal impressions small, shallow, spanning 1 / 2 width of interval 3; lateral elytral setae arranged in anterior series of 7 setae and posterior series of 6 setae; elytral marginal depression moderately broad, margin little upraised even near subapical sinuation; subapical sinuation shallow, concavity symmetrical. Mesepisternum with ~ 18 punctures in 2 - 3 rows; metepisternal width to length ratio = 0.81; metepisternum / metepimeron suture distinct. Abdomen with irregular lateral wrinkles on ventrites 1 - 5, and lateral depressions on ventrites 3 - 6; suture between ventrites 2 and 3 reduced laterally, effaced; apical male ventrite with 2 marginal setae and apical female ventrite with 4 equally spaced setae and median trapezoid of 4 subequal, short setae. Legs-metatarsomere 1 / metatibial length ratio = 0.20; metatarsomere 4 length along outer lobe 1.3 x medial tarsomere length, apical and subapical setae present but short; metatarsal dorsolateral sulci broad, shallow, median area subcarinate. Microsculpture of vertex a shallow transverse mesh, sculpticell breadth 2 - 3 x length; pronotal median base glossy, shallow transverse sculpticells laterally near punctures; metasternum with shallow transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse sculpticells. Coloration of vertex rufobrunneous with piceous cast; antennomeres 1 - 3 rufoflavous, 4 - 11 rufobrunneous; pronotal disc rufobrunneous, lateral margins narrowly, and base and apex rufoflavous; proepipleuron rufoflavous, proepisternum rufobrunneous with piceous cast; elytral disc rufobrunneous, sutural interval basally rufous, concolorous on disc, apically rufoflavous; elytral marginal depression rufoflavous outside humerus, concolorous with disc posteriorly; elytral epipleuron rufoflavous, metepisternum rufobrunneous; abdominal ventrites 1 - 5 rufopiceous medially, rufoflavous laterally, apical 3 / 4 of apical ventrite 6 rufoflavous; metafemur flavous; metatibia rufoflavous. Male genitalia (n = 3). Aedeagal median lobe very slender immediately distad basal bulb, greatly expanded along dorsal margin of ostial opening, distance from parameral articulation to tip 3.2 x greatest depth (Fig. 153 A, C), apex extended as an asymmetrical spoon with blunt ventral projection; median lobe with apex directed rightward in ventral view (Fig. 153 B), bulging dorsal margin (Fig. 153 A) visible as convex apical prominence in this view; internal sac short, narrow, with broad, diffuse dorsal ostial microtrichial patch near base (Fig. 153 C); flagellar plate small, length 0.20 x parameral articulation-tip distance (Fig. 153 C). Female reproductive tract (n = 1). Bursa copulatrix broadly columnar with small apical cap, overall length 0.78 mm, cap 0.29 mm broad x 0.17 mm long, shaft breadth 0.43 mm (Fig. 154 A); apical cap finely wrinkled, shaft translucent with thick wrinkles; gonocoxite 1 with 3 - 4 apical fringe setae, a curved seta just basad medioapical angle and 7 smaller setae on medial surface (Fig. 155 A); gonocoxite 2 falcate with acuminate tip, basal extension straight, 2 lateral ensiform setae, apical nematiform setae on medial surface at 0.68 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
FE2538D37910C181A265F6AD8511E86E.taxon	etymology	Etymology. This species appears not unlike Mecyclothorax filipes (Sharp) of Lana'i (Liebherr 2009 b), leading to filipoides as the species epithet. The adjectival stem filipes refers to the possession of threadlike feet (Jaeger 1955).	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
89C3619FE41B33AE171813A167E0F8FB.taxon	diagnosis	Diagnosis. This species (Fig. 149 D) can be separated from all other Mecyclothorax palustris group species based on: 1, uniformly pale body coloration without contrasted elytral lateral margins; 2, presence of two dorsal elytral setae, setal formula 2 1 2 0; and 3, relatively small size; standardized body length 3.9 - 4.6 mm. Were it to be confused with any other Haleakala Mecyclothorax, the best candidate for confusion would be Mecyclothorax mauiae (Fig. 68 A), as beetles of both species are pale bodied, and the size ranges overlap. Careful attention must be paid to the comparative depth of the sutural and 2 nd striae, as this is a major criterion for group membership. Also, the bodies of Mecyclothorax mauiae are broader, with the elytra slightly obovoid; i. e. with the greatest breadth in the anterior half. Conversely Mecyclothorax nanunctus beetles have the elytra more ovoid, with the greatest width in the apical half. The pronotum in Mecyclothorax mauiae is less transverse, though not diagnostically so, and the pronotal lateral depression is narrower with a less explanate margin. The surest arbiter given a male in the series is the male aedeagal median lobe, which in Mecyclothorax nanunctus males is gracile and elongate (Fig. 153 D-J), and in Mecyclothorax mauiae males is short with a broadly rounded apex (Fig. 69). The two species are broadly sympatric (Figs 71, 157), but based on ecological records from the Waikamoi area, Mecyclothorax nanunctus is a species more at home in mesic forest situations dominated by Acacia koa and ' ōhi'a (Metrosideros polymorpha), whereas Mecyclothorax mauiae is found in wet forest situations that are dominated by ' ōhi'a but which also include ' ōlapa (Cheirodendron trigynum).	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
89C3619FE41B33AE171813A167E0F8FB.taxon	description	Description (n = 5). Head capsule with frontal grooves broad for most of length, terminated mesad low convexity mesad anterior supraorbital seta; dorsal impression of neck slightly concave; ocular lobe obtusely protruded from gena, eyes small and moderately convex, ocular ratio = 1.41 - 1.49, ocular lobe ratio = 0.79 - 0.84; labral anterior margin shallowly emarginate to 1 / 8 labral length; antennae filiform; antennomeres 2 - 3 with sparse pelage of short setae; mentum tooth with sides acute, apex tightly rounded. Pronotum moderately transverse, MPW / PL = 1.18 - 1.25, base narrowly constricted, MPW / BPW = 1.57 - 1.68, lateral margins subparallel for 0.1 x pronotal length anterad right hind angles; median base distinctly depressed relative to convex disc, ~ 19 isolated punctures each side, half of them lining juncture between base and disc; basal margin slightly convex between laterobasal depressions; median longitudinal impression very shallow, middle of disc flat; anterior transverse impression broad, shallow, obsolete medially, crossed by indistinct wrinkles, incised only mesad front angles; anterior callosity slightly convex, crossed by indistinct longitudinal wrinkles; front angles slightly projected, tightly rounded; pronotal apical width greater than basal width, APW / BPW = 1.05 - 1.09; lateral marginal depression very narrow, slightly wider at front angle, thickly upraised outside laterobasal depression; laterobasal depression narrow, surface irregular, continuous with lateral depression. Proepisternum with 5 minute punctures along hind marginal groove; prosternal process with narrow lateral depression, lateral margins broadly upraised. Elytra convex, disc domed, sides distinctly sloped; basal groove evenly curved to tightly rounded humerus defined by juncture of basal groove and broader lateral marginal depression, MEW / HuW = 2.21 - 2.34; parascutellar seta present; parascutellar striole with 4 punctures, striole shallow or interrupted between punctures; sutural interval more convex than lateral intervals, sutural juncture upraised; sutural and 2 nd striae of similar depth and punctation on disc, sutural stria continuous to base, and finely incised, deep, and smooth apically, whereas 2 nd stria is absent from base, shallow and irregularly interrupted apically; discal striae 2 - 3 shallow, punctate, striae 4 - 5 very shallow to obsolete, striae 6 - 7 only shallow isolated punctures, or absent; elytral apex between 2 nd and 8 th striae uniformly convex; 2 dorsal elytral setae at 0.27 - 0.29 x and 0.49 - 0.51 x elytral length, setal impressions small, shallow, spanning 1 / 2 width of interval 3; lateral elytral setae arranged in anterior series of 6 setae and a posterior series of 5 (6) setae; elytral marginal depression broader at humerus, edge little upturned there, gradually narrowed to bead at subapical sinuation; subapical sinuation shallow, more abruptly incurved anteriorly. Mesepisternum with ~ 10 punctures in 2 - 3 rows; metepisternal width to length ratio = 0.68; metepisternum / metepimeron suture distinct. Abdomen with irregular lateral wrinkles on ventrites 1 - 5, and lateral depressions on ventrites 3 - 6; suture between ventrites 2 and 3 complete; apical male ventrite with 2 marginal setae and apical female ventrite with 4 equally spaced setae and median trapezoid of 4 subequal, short setae. Legs-metatarsomere 1 / metatibial length ratio = 0.19; metatarsomere 4 length along outer lobe 1.2 x medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci broad, shallow, median area subcarinate. Microsculpture of vertex a transverse mesh, sculpticell breadth 2 - 3 x length; pronotal disc with shallow transverse mesh to transverse lines, glossy in part, median base with shallow transverse mesh between glossy portions; elytral disc with distinct, regular transverse mesh, sculpticell breadth 2 - 3 x length, apex with less-developed mesh of same dimensions; metasternum with shallow transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex rufous; antennomeres 1 - 3 flavous, 4 - 11 rufoflavous; pronotal disc rufoflavous, base slightly darker, rufobrunneous; proepipleuron flavous, proepisternum dorsally flavous, ventrally rufoflavous; elytral disc rufobrunneous, sutural interval rufoflavous basally, flavous apically, lateral depression and apex of 8 th interval flavous; elytral epipleuron flavous, metepisternum rufoflavous; abdomen with all ventrites medially rufoflavous, marginally flavous, apical half of apical ventrite flavous; metafemur flavous; metatibia flavous with rufoflavous cast. Male genitalia (n = 9). Aedeagal median lobe slender, elongate, distance from parameral articulation to tip 5.6 - 6.0 x maximum depth of shaft (Fig. 153 D-E, G, I-J), apex nar rowly extended 3.0 - 3.5 x depth beyond ostial opening (Fig. 153 E, J); median lobe nearly symmetrical in ventral view (Fig. 153 F, H), with apex curved leftward to pointed tip in this view; internal sac with well-developed microtrichial field on right-ventral face near base (homologous with dorsal patch?), sac surface otherwise covered with fine microspicules (Fig. 153 J); flagellar plate of moderate size, length 0.40 x parameral articulation-tip distance. Female reproductive tract (n = 1). Bursa copulatrix parallel sided, elongate, with slight apical expansion, length 1.06 mm, breadth 0.29 mm (Fig. 154 B); bursal walls translucent, thinly wrinkled; gonocoxite 1 with 3 - 4 apical fringe setae, a small seta near medioapical angle and another ~ 16 small setae on medial surface (Fig. 155 B); gonocoxite 2 falcate with subacuminate apex, lateral margin straight near ensiform setae, terminus of basal extension slightly curved, 2 elongate lateral ensiform setae, apical nematiform setae on medioventral surface at 0.69 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
89C3619FE41B33AE171813A167E0F8FB.taxon	etymology	Etymology. During the initial sorting of specimens for this revision, this species was given the working name " little- unctus. " That name is validated here using the prefixed Latin participle nanunctus, or " small-anointed " (Brown 1956).	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
B0CF5CFC8C1DCD91E276C61C8CBA02E9.taxon	diagnosis	Diagnosis. These beetles exhibit uniformly dark, rufobrunneous to rufopiceous body color (Fig. 158 A-B), with the dorsal surface glossy due to the very transverse microsculpture. They are broad-bodied beetles with pronotum transverse, MPW / PL = 1.22 - 1.33, and elytra basally broad and subquadrate, MEW / HuW = 2.05 - 2.19. This breadth of body distinguishes them from the Mecyclothorax palustroides triad (Figs 149 B, 158 C-E). Nevertheless, this widely distributed species (Fig. 160) exhibits substantial variation in body size (standardized body length 3.7 - 5.2 mm), elytral breadth (Fig. 158 A-B), and elytral striation. The discal striae 1 - 3, 1 - 4, 1 - 5, or 1 - 6 may be traceable by deeply impressed striae medially and linear series of isolated punctures laterally. Setal formula 2 1 (2) 2 0; the basal pronotal setae may be present in rare instances (e. g. Fig. 158 B).	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
AB6AEAEB5B90755C794B2BCDF91975D2.taxon	diagnosis	Diagnosis. This is the second species (Fig. 158 C-D), along with Mecyclothorax palustroides (Fig. 149 B) and Mecyclothorax pau (Fig. 158 E), that comprise a taxonomic triplet characterized by extremely similar external appearance. Individuals of this species exhibit laterally darker elytra, with intervals 8 - 9 gradually paler to rufobrunneous compared to the rufopiceous disc; a condition shared with Mecyclothorax pau. There may be a broader pale area behind the humerus, but at midlength the lateral intervals 7 - 8 are indistinctly paler. The pale margin of the elytral apex is narrow, at most extended to a line traversing the subapical sinuations. The pronotal median base appears less depressed relative to the disc due to its less developed punctation, with ~ 12 small, isolated punctures each side and few or no elongate punctures along the juncture of disc and base. And the elytral microsculpture is an upraised transverse mesh comprising a mixture of isodiametric and transverse sculpticells, their individual surfaces partially upraised so that the surface is alutaceous; a condition shared with Mecyclothorax palustroides. The male aedeagal median lobe is bluntly rounded apically (Fig. 161 A-D). Setal formula 2 1 2 0. Standardized body length 3.9 - 4.5 mm, diagnostically smaller than the body size of the sympatrically distributed Mecyclothorax palustroides; standardized body length 4.6 - 5.5 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
AB6AEAEB5B90755C794B2BCDF91975D2.taxon	description	Description (n = 5). [The above description of Mecyclothorax palustroides can serve to describe this species. All diagnostic characters are presented above, so only the various recorded ratios are presented below.] Eyes moderately convex, ocular ratio = 1.48 - 1.54, ocular lobe ratio = 0.77 - 0.85. Pronotum slightly transverse, MPW / PL = 1.09 - 1.21, variably constricted basally, MPW / BPW = 1.52 - 1.71; pronotal apical width greater than basal width, APW / BPW = 1.07 - 1.15. Elytra subovoid, MEW / HuW = 2.13 - 2.35. Male genitalia (n = 6). Aedeagal median lobe short, squat, distance from parameral articulation to tip 3.2 - 3.3 x depth at midlength (Fig. 161 A, C); apex bluntly rounded and not extended beyond ostial opening; median lobe slightly curved rightward near blunt apex, the right and left margins subparallel in ventral view (Fig. 161 B); internal sac with ventral surface densely covered with pelage of shaggy microtrichia (Fig. 161 D), the sac only 1.5 x as long as flagellar plate; flagellar plate large, length 0.61 x parameral articulation-tip distance. Female reproductive tract (n = 1). Bursa copulatrix vase shaped, parallel sided with basal constriction, overall length 1.14 mm, midlength breadth 0.40 mm, basal constriction 0.22 mm broad (Fig. 154 D): bursal walls translucent, thickly wrinkled; gonocoxite 1 with 4 apical fringe setae and 4 smaller setae on medial surface (Fig. 155 D); gonocoxite 2 falcate with narrowly rounded apex, thick basal extension curved at terminus, 2 elongate lateral ensiform setae, apical nematiform setae on medioventral surface at 0.71 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
AB6AEAEB5B90755C794B2BCDF91975D2.taxon	etymology	Etymology. This species is named to honor Dr. Catherine A. Tauber and Prof. Maurice J. Tauber for their collegiality and friendship, and for their contributions to Hawaiian entomology, evolutionary biology, and the study of Neuroptera worldwide.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
DF9F5F68D211A401E9107F0D0A41763C.taxon	diagnosis	Diagnosis. This (Fig. 158 E), the third and last species of the Mecyclothorax palustroides - based triad to be taxonomically treated, is diagnosed by the following combination: 1, lateral elytral intervals as dark or only gradually paler laterally than the disc, a condi tion shared with Mecyclothorax tauberorum (Fig. 158 C-D); 2, pronotal median base distinctly punctate, with ~ 19 punctures each side, the punctures along the juncture of disc and median base elongate, a condition shared with Mecyclothorax palustroides (Fig. 149 B); and 3, elytral microsculpture a shallow transverse mesh, sculpticells a mixture of isodiametric in transverse rows and transverse, breadth 2 x length, with adjacent sculpticells tiled, their surfaces flat, and thus the surface slightly glossy. Setal formula 2 1 2 0. Standardized body length 4.0 - 5.3 mm.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
DF9F5F68D211A401E9107F0D0A41763C.taxon	description	Description (n = 5). [The above description of Mecyclothorax palustroides can serve to describe this species. All diagnostic characters are presented above, so only the various recorded ratios are presented below.] Eyes moderately convex, ocular ratio = 1.51 - 1.58, ocular lobe ratio = 0.7.8 - 0.86. Pronotum slightly transverse, MPW / PL = 1.16 - 1.21, variably constricted basally, MPW / BPW = 1.59 - 1.69; pronotal apical width subequal to greater than basal width, APW / BPW = 0.99 - 1.13. Elytra subovoid, MEW / HuW = 2.19 - 2.39. Male genitalia (n = 43). Aedeagal median lobe variably robust, distance from parameral articulation to tip 4.4 - 4.1 x depth at midlength (Fig. 161 E, I); apex extended 1.6 - 2.5 x depth beyond ostial opening, the extension narrow to moderately broad (Fig. 161 E, H); median lobe shaft straight in ventral view, apex offset to right side with left (dorsal) margin convex (Fig. 161 G) and tip bluntly rounded; internal sac variable, either a long curved cylinder (Figs 161 H, 162 A-D) or bilobed with a smaller basal lobe and larger apical lobe (Fig. 161 I), the sac surface covered with microspicules only; flagellar plate moderately large, length 0.40 x parameral articulation-tip distance in the individual with bilobed sac (Fig. 161 I), or large, length 0.54 - 0.61 x parameral articulation-tip distance in the balance of everted individuals with cylindrical sac (Figs 161 H, 162). As the sac length can be assessed in uneverted specimens based on the plate's shadow in backlit photographs (e. g. 161 E, F, J-L), it is concluded that only one dissected male (Fig. 161 I) has the bilobed sac condition with the associated small flagellar plate. The individual with the bilobed sac (Fig. 161 I; lot 19980511.01, CUIC) was one of two males collected at State Camp on Helele'ike'oha Stream. A second male collected at that site (Fig. 161 J; lot 19980512.11, NMNH) exhibits the long flagellar plate condition (i. e. plate length 0.55 x parameral articulation-tip distance, Fig. 161 J), Cylindrical-sac, large-plate males are also observed to the north of State Camp at Poouli Cabin (Fig. 161 F, H), and to east and south in Hana Bogs and Kīpahulu Valley (Figs 161 K, 162 A-B), and on the Manawainui Planeze (Figs 161 L, 162 C-D). Because both the cylindrical and bilobed conditions are observed sympatrically at State Camp, they are interpreted as two forms of an infraspecific polymorphism. Various hypotheses for the origin of this polymorphism might include: 1, the bilobed male condition is based on a rare allele primitively shared with the sympatrically distributed Mecyclothorax bilobatus (Fig. 150 E, F); or 2, the bilobed male condition is the product of recent hybridization between the sympatric Mecyclothorax pau and Mecyclothorax bilobatus. Female reproductive tract (n = 1). Bursa copulatrix vase shaped, parallel sided with basal constriction just distad vagina, overall length 1.05 mm, midlength breadth 0.43 mm, basal constriction 0.25 mm broad (Fig. 154 E): bursal walls translucent, thickly wrinkled; gonocoxite 1 with 4 apical fringe setae, a curved seta at medioapical angle and 9 - 10 smaller setae on medial surface (Fig. 155 E); gonocoxite 2 falcate with acuminate apex, the lateral margin slightly expanded near apex, thick basal extension curved at terminus, 2 elongate lateral ensiform setae, apical nematiform setae on medioventral surface at 0.71 x gonocoxite length.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
DF9F5F68D211A401E9107F0D0A41763C.taxon	etymology	Etymology. The epithet pau given to this terminal species in the revision is the useful and definitive Hawaiian word for: finished, ended, completed, over, all done, final (Pukui et al. 1975). Given the intense endemism of Haleakala Mecyclothorax species, there is no doubt that this epithet will be superseded in the future by names of presently undescribed species.	en	Liebherr, James K. (2015): The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1-407, DOI: http://dx.doi.org/10.3897/zookeys.544.6074, URL: http://dx.doi.org/10.3897/zookeys.544.6074
