taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03AB87BA2D08A202E41FFA987D00FE55.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050785/files/figure.png	https://doi.org/10.5281/zenodo.5050785	FIGURE 2. Shells of Pseudotricula species. A, Pseudotricula eberhardi, holotype, C.165052, Black Curtain Streamway, Cueva Blanca; B–E, Pseudotricula expandolabra, B, holotype, C.439673, Persephone Streamway, Bauhaus; C, paratype, C. 439674, Persephone Streamway, Bauhaus; D, C.201811, Persephone Pot, Bauhaus; E, C201495, Persephone Streamway, Bauhaus; F, Pseudotricula auriforma, holotype, C.439672, main streamway, Damper Cave. Scales: A F, 500 m.	FIGURE 2. Shells of Pseudotricula species. A, Pseudotricula eberhardi, holotype, C.165052, Black Curtain Streamway, Cueva Blanca; B–E, Pseudotricula expandolabra, B, holotype, C.439673, Persephone Streamway, Bauhaus; C, paratype, C. 439674, Persephone Streamway, Bauhaus; D, C.201811, Persephone Pot, Bauhaus; E, C201495, Persephone Streamway, Bauhaus; F, Pseudotricula auriforma, holotype, C.439672, main streamway, Damper Cave. Scales: A F, 500 m.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D08A202E41FFA987D00FE55.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050787/files/figure.png	https://doi.org/10.5281/zenodo.5050787	FIGURE 3: Radulae of Pseudotricula species. A, B, Pseudotricula eberhardi, C.165053, Black Curtain Streamway, Cueva Blanca; C–H, Pseudotricula expandolabra, C, D, C.201495, Persephone Streamway, Bauhaus, E, G, H, C.201811, Persephone Pot, Bauhaus, F, C.201822, main streamway, Bauhaus; I, Pseudotricula arthurclarkei, C.203671, paratype, Quetzalcoatl Conduit; J–L, Pseudotricula conica, C.203676, paratype, Cane Toad Abuse Streamway, Damper Cave. A, C, G, K, central, lateral inner and outer marginal teeth; B, E, L, central teeth; D, outer marginal teeth; F, J, lateral and inner marginal teeth; H, lateral tooth; I, central and lateral teeth. Scales: A, B, E, H–J, L 10 m; C, F, G, K, 20 m; D, 5 m.	FIGURE 3: Radulae of Pseudotricula species. A, B, Pseudotricula eberhardi, C.165053, Black Curtain Streamway, Cueva Blanca; C–H, Pseudotricula expandolabra, C, D, C.201495, Persephone Streamway, Bauhaus, E, G, H, C.201811, Persephone Pot, Bauhaus, F, C.201822, main streamway, Bauhaus; I, Pseudotricula arthurclarkei, C.203671, paratype, Quetzalcoatl Conduit; J–L, Pseudotricula conica, C.203676, paratype, Cane Toad Abuse Streamway, Damper Cave. A, C, G, K, central, lateral inner and outer marginal teeth; B, E, L, central teeth; D, outer marginal teeth; F, J, lateral and inner marginal teeth; H, lateral tooth; I, central and lateral teeth. Scales: A, B, E, H–J, L 10 m; C, F, G, K, 20 m; D, 5 m.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D08A202E41FFA987D00FE55.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050791/files/figure.png	https://doi.org/10.5281/zenodo.5050791	FIGURE 4. Dorsal views of penes of Pseudotricula and Nanocochlea species. A,. Pseudotricula eberhardi; B, Pseudotricula expandolabra; C, Nanocochlea stylesae; D, Nanocochlea damperensis; E, Pseudotricula conica. Scales A, B, E, 0.2 mm; C, D, 0.1 mm. Abbreviations: ds, distal swelling; pd, penial duct.	FIGURE 4. Dorsal views of penes of Pseudotricula and Nanocochlea species. A,. Pseudotricula eberhardi; B, Pseudotricula expandolabra; C, Nanocochlea stylesae; D, Nanocochlea damperensis; E, Pseudotricula conica. Scales A, B, E, 0.2 mm; C, D, 0.1 mm. Abbreviations: ds, distal swelling; pd, penial duct.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D08A202E41FFA987D00FE55.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050781/files/figure.png	https://doi.org/10.5281/zenodo.5050781	FIGURE 1. Schematic plan of mapped caves (black outline) at Precipitous Bluff showing hydrologic relationships (arrows indicate direction of stream flow paths), sample sites (numbered), cave entrances (black dots), and locations mentioned in text. Adapted from Hume (1989). Inserts: A, Tasmania with location of Precipitous Bluff (PB in boxed area), Ida Bay (IB), Hastings (H), Vanishing Falls (VF) and Cracroft (C); B, Precipitous Bluff and New River Lagoon showing surface sample sites (numbered) and location of the caves (boxed area) in relation to dolines (open circles) with sinking streams that feed Damper Cave and Quetzalcoatl Conduit, which becomes Damper Creek upon resurging at the surface. Also shown (in B) is Precipitous Bluff. The numbers shown in B are explained below. The arrows in insert B and the main figure show the direction of water flow. In the main map, the “PB” prefixed numbers refer to cave entrance numbers used by the Australian Speleological Federation. Sample sites shown in both the main map and in insert B are numbered as follows: 1 Bauhaus Cave (PB6) base of aven (stn PB6­1) and main streamway at downclimb (PB6­1A); 2 Bauhaus Cave (Persephone Pot PB17) lower streamway at first contact with stream stn 7; 3 Bauhaus Persephone middle streamway stn 8; 4 Bauhaus Persephone middle streamway stn 9; 5 Bauhaus Persephone upper streamway stn 10; 6 Bauhaus Screaming Stals streamway stn 11; 7 Bauhaus Screaming Stals streamway stn 12; and tributary seep near stn 12; 8 Bauhaus Screaming Stals streamway stn 13; 9 Cueva Blanca (PB4), Black Curtains streamway (=Inundation Passage); 10 Damper Cave (PB1) main streamway ca. 50 m inside entrance; 11 Damper Cave main streamway 100 m into cave; 12 Damper Cave, Honey and Cream; 13 Damper Cave seep near The Keg; 14 Damper Cave, Cane Toad Abuse; 15 Quetzalcoatl Conduit (PB3), 30 m into cave, stn PB3­ 1C; 16 Quetzalcoatl Conduit, beyond Pendulum Palace, stn PB3­2; 17 Damper Creek, 10 m downstream of Damper Cave entrance; 18 Damper Creek, first crossing; 19 Damper Creek at New River Lagoon; 20 Malani Creek; 21 Limestone Creek near New River Lagoon.	FIGURE 1. Schematic plan of mapped caves (black outline) at Precipitous Bluff showing hydrologic relationships (arrows indicate direction of stream flow paths), sample sites (numbered), cave entrances (black dots), and locations mentioned in text. Adapted from Hume (1989). Inserts: A, Tasmania with location of Precipitous Bluff (PB in boxed area), Ida Bay (IB), Hastings (H), Vanishing Falls (VF) and Cracroft (C); B, Precipitous Bluff and New River Lagoon showing surface sample sites (numbered) and location of the caves (boxed area) in relation to dolines (open circles) with sinking streams that feed Damper Cave and Quetzalcoatl Conduit, which becomes Damper Creek upon resurging at the surface. Also shown (in B) is Precipitous Bluff. The numbers shown in B are explained below. The arrows in insert B and the main figure show the direction of water flow. In the main map, the “PB” prefixed numbers refer to cave entrance numbers used by the Australian Speleological Federation. Sample sites shown in both the main map and in insert B are numbered as follows: 1 Bauhaus Cave (PB6) base of aven (stn PB6­1) and main streamway at downclimb (PB6­1A); 2 Bauhaus Cave (Persephone Pot PB17) lower streamway at first contact with stream stn 7; 3 Bauhaus Persephone middle streamway stn 8; 4 Bauhaus Persephone middle streamway stn 9; 5 Bauhaus Persephone upper streamway stn 10; 6 Bauhaus Screaming Stals streamway stn 11; 7 Bauhaus Screaming Stals streamway stn 12; and tributary seep near stn 12; 8 Bauhaus Screaming Stals streamway stn 13; 9 Cueva Blanca (PB4), Black Curtains streamway (=Inundation Passage); 10 Damper Cave (PB1) main streamway ca. 50 m inside entrance; 11 Damper Cave main streamway 100 m into cave; 12 Damper Cave, Honey and Cream; 13 Damper Cave seep near The Keg; 14 Damper Cave, Cane Toad Abuse; 15 Quetzalcoatl Conduit (PB3), 30 m into cave, stn PB3­ 1C; 16 Quetzalcoatl Conduit, beyond Pendulum Palace, stn PB3­2; 17 Damper Creek, 10 m downstream of Damper Cave entrance; 18 Damper Creek, first crossing; 19 Damper Creek at New River Lagoon; 20 Malani Creek; 21 Limestone Creek near New River Lagoon.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D0CA21BE41FFDED7C72FD6D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050785/files/figure.png	https://doi.org/10.5281/zenodo.5050785	FIGURE 2. Shells of Pseudotricula species. A, Pseudotricula eberhardi, holotype, C.165052, Black Curtain Streamway, Cueva Blanca; B–E, Pseudotricula expandolabra, B, holotype, C.439673, Persephone Streamway, Bauhaus; C, paratype, C. 439674, Persephone Streamway, Bauhaus; D, C.201811, Persephone Pot, Bauhaus; E, C201495, Persephone Streamway, Bauhaus; F, Pseudotricula auriforma, holotype, C.439672, main streamway, Damper Cave. Scales: A F, 500 m.	FIGURE 2. Shells of Pseudotricula species. A, Pseudotricula eberhardi, holotype, C.165052, Black Curtain Streamway, Cueva Blanca; B–E, Pseudotricula expandolabra, B, holotype, C.439673, Persephone Streamway, Bauhaus; C, paratype, C. 439674, Persephone Streamway, Bauhaus; D, C.201811, Persephone Pot, Bauhaus; E, C201495, Persephone Streamway, Bauhaus; F, Pseudotricula auriforma, holotype, C.439672, main streamway, Damper Cave. Scales: A F, 500 m.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D0CA21BE41FFDED7C72FD6D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050787/files/figure.png	https://doi.org/10.5281/zenodo.5050787	FIGURE 3: Radulae of Pseudotricula species. A, B, Pseudotricula eberhardi, C.165053, Black Curtain Streamway, Cueva Blanca; C–H, Pseudotricula expandolabra, C, D, C.201495, Persephone Streamway, Bauhaus, E, G, H, C.201811, Persephone Pot, Bauhaus, F, C.201822, main streamway, Bauhaus; I, Pseudotricula arthurclarkei, C.203671, paratype, Quetzalcoatl Conduit; J–L, Pseudotricula conica, C.203676, paratype, Cane Toad Abuse Streamway, Damper Cave. A, C, G, K, central, lateral inner and outer marginal teeth; B, E, L, central teeth; D, outer marginal teeth; F, J, lateral and inner marginal teeth; H, lateral tooth; I, central and lateral teeth. Scales: A, B, E, H–J, L 10 m; C, F, G, K, 20 m; D, 5 m.	FIGURE 3: Radulae of Pseudotricula species. A, B, Pseudotricula eberhardi, C.165053, Black Curtain Streamway, Cueva Blanca; C–H, Pseudotricula expandolabra, C, D, C.201495, Persephone Streamway, Bauhaus, E, G, H, C.201811, Persephone Pot, Bauhaus, F, C.201822, main streamway, Bauhaus; I, Pseudotricula arthurclarkei, C.203671, paratype, Quetzalcoatl Conduit; J–L, Pseudotricula conica, C.203676, paratype, Cane Toad Abuse Streamway, Damper Cave. A, C, G, K, central, lateral inner and outer marginal teeth; B, E, L, central teeth; D, outer marginal teeth; F, J, lateral and inner marginal teeth; H, lateral tooth; I, central and lateral teeth. Scales: A, B, E, H–J, L 10 m; C, F, G, K, 20 m; D, 5 m.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D0CA21BE41FFDED7C72FD6D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050791/files/figure.png	https://doi.org/10.5281/zenodo.5050791	FIGURE 4. Dorsal views of penes of Pseudotricula and Nanocochlea species. A,. Pseudotricula eberhardi; B, Pseudotricula expandolabra; C, Nanocochlea stylesae; D, Nanocochlea damperensis; E, Pseudotricula conica. Scales A, B, E, 0.2 mm; C, D, 0.1 mm. Abbreviations: ds, distal swelling; pd, penial duct.	FIGURE 4. Dorsal views of penes of Pseudotricula and Nanocochlea species. A,. Pseudotricula eberhardi; B, Pseudotricula expandolabra; C, Nanocochlea stylesae; D, Nanocochlea damperensis; E, Pseudotricula conica. Scales A, B, E, 0.2 mm; C, D, 0.1 mm. Abbreviations: ds, distal swelling; pd, penial duct.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D0CA21BE41FFDED7C72FD6D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050793/files/figure.png	https://doi.org/10.5281/zenodo.5050793	FIGURE 5. Shells of Pseudotricula species. A–F, Pseudotricula expandolabra, A–C, C.201489, Screaming Stals streamway, Bauhaus, D–F, Persephone Pot, Bauhaus, D, F, C.201811, E, C.201485, Persephone Pot, Bauhaus; G–I, Pseudotricula auriforma G, H, streamway near entrance, Damper Cave, I, main streamway, Damper Cave. Scales: 1 mm.	FIGURE 5. Shells of Pseudotricula species. A–F, Pseudotricula expandolabra, A–C, C.201489, Screaming Stals streamway, Bauhaus, D–F, Persephone Pot, Bauhaus, D, F, C.201811, E, C.201485, Persephone Pot, Bauhaus; G–I, Pseudotricula auriforma G, H, streamway near entrance, Damper Cave, I, main streamway, Damper Cave. Scales: 1 mm.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D0CA21BE41FFDED7C72FD6D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050797/files/figure.png	https://doi.org/10.5281/zenodo.5050797	FIGURE 6. Protoconchs and teleoconch microsculpture of Pseudotricula and Nanocochlea species. A–D, Pseudotricula expandolabra, A, B, D, C.201493, paratype, main streamway, Bauhaus, C, C.201495, Persephone Streamway, Bauhaus; E–G, Pseudotricula conica, E, F, C.201464, Cane Toad Abuse Streamway, Damper Cave, G, Persephone Pot, Bauhaus; Honey and Cream Streamway, Damper Cave; H, Nanocochlea exigua, C.201475; I, J, Nanocochlea stylesae, C.201476, paratypes, Malani Creek; K–M, Nanocochlea damperensis, K, L, C.203685, paratypes, first crossing, Damper Creek, M, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; N, O, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. A, B, E, F, H, K–N, protoconch; C, D, G, I, J, O, microsculpture. Scales: A, C–E, J, K, M, O, 200 m; B, F–I, L, N, 100 m.	FIGURE 6. Protoconchs and teleoconch microsculpture of Pseudotricula and Nanocochlea species. A–D, Pseudotricula expandolabra, A, B, D, C.201493, paratype, main streamway, Bauhaus, C, C.201495, Persephone Streamway, Bauhaus; E–G, Pseudotricula conica, E, F, C.201464, Cane Toad Abuse Streamway, Damper Cave, G, Persephone Pot, Bauhaus; Honey and Cream Streamway, Damper Cave; H, Nanocochlea exigua, C.201475; I, J, Nanocochlea stylesae, C.201476, paratypes, Malani Creek; K–M, Nanocochlea damperensis, K, L, C.203685, paratypes, first crossing, Damper Creek, M, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; N, O, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. A, B, E, F, H, K–N, protoconch; C, D, G, I, J, O, microsculpture. Scales: A, C–E, J, K, M, O, 200 m; B, F–I, L, N, 100 m.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D0CA21BE41FFDED7C72FD6D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050801/files/figure.png	https://doi.org/10.5281/zenodo.5050801	FIGURE 7. Opercula of Pseudotricula species. A, B, Pseudotricula expandolabra, C.201493, paratypes, main streamway, Bauhaus; C, Pseudotricula auriforma, C.201496, paratype, main streamway, Damper Cave; D, E, Pseudotricula arthurclarkei, C.203671, paratypes, Quetzalcoatl Conduit; F–I, Pseudotricula conica, F, G, C.203680, main streamway, near entrance, Damper Cave; H, I, C.201459, Screaming Stals streamway, Bauhaus; A, D, F, H, outer side; B, C, E, H, I, inner side; Scales: A–I, 200 m.	FIGURE 7. Opercula of Pseudotricula species. A, B, Pseudotricula expandolabra, C.201493, paratypes, main streamway, Bauhaus; C, Pseudotricula auriforma, C.201496, paratype, main streamway, Damper Cave; D, E, Pseudotricula arthurclarkei, C.203671, paratypes, Quetzalcoatl Conduit; F–I, Pseudotricula conica, F, G, C.203680, main streamway, near entrance, Damper Cave; H, I, C.201459, Screaming Stals streamway, Bauhaus; A, D, F, H, outer side; B, C, E, H, I, inner side; Scales: A–I, 200 m.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D0CA21BE41FFDED7C72FD6D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050803/files/figure.png	https://doi.org/10.5281/zenodo.5050803	FIGURE 8. Pallial cavities of male specimens of Pseudotricula and Nanocochlea species. A, Pseudotricula expandolabra, C.201493, paratype, Persephone Streamway, Bauhaus; B, Pseudotricula arthurclarkei, C.203671, paratype, Quetzalcoatl Conduit; C, Pseudotricula auriforma, C.201496, paratype, main streamway, Damper Cave; D, E, Pseudotricula conica, D, C.203680, main streamway near entrance, Damper Cave, E, Cane Toad Abuse Streamway, Damper Cave; F, Nanocochlea exigua, C.203677, paratype, Cane Toad Abuse Streamway, Damper Cave; G, Nanocochlea damperensis, C.203685, paratype, first crossing, Damper Creek; H, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. Scales: A–D, F–H, 500 m, E, 1 mm. Abbreviations: a, anus; ct, ctenidium; ev, efferent vein; os, osphradium; pc, pericardium; pe, pallial edge; ppw, posterior pallial wall; r, rectum; rg, renal gland. The stippled area represents the hypobranchial gland.	FIGURE 8. Pallial cavities of male specimens of Pseudotricula and Nanocochlea species. A, Pseudotricula expandolabra, C.201493, paratype, Persephone Streamway, Bauhaus; B, Pseudotricula arthurclarkei, C.203671, paratype, Quetzalcoatl Conduit; C, Pseudotricula auriforma, C.201496, paratype, main streamway, Damper Cave; D, E, Pseudotricula conica, D, C.203680, main streamway near entrance, Damper Cave, E, Cane Toad Abuse Streamway, Damper Cave; F, Nanocochlea exigua, C.203677, paratype, Cane Toad Abuse Streamway, Damper Cave; G, Nanocochlea damperensis, C.203685, paratype, first crossing, Damper Creek; H, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. Scales: A–D, F–H, 500 m, E, 1 mm. Abbreviations: a, anus; ct, ctenidium; ev, efferent vein; os, osphradium; pc, pericardium; pe, pallial edge; ppw, posterior pallial wall; r, rectum; rg, renal gland. The stippled area represents the hypobranchial gland.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D0CA21BE41FFDED7C72FD6D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050807/files/figure.png	https://doi.org/10.5281/zenodo.5050807	FIGURE 9. Head, penis and stomach of Pseudotricula species. A, D, Pseudotricula conica, C.203680, main streamway near entrance, Damper Cave; C, Pseudotricula expandolabra, C.201493, paratype, Persephone Streamway, Bauhaus; B, E, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. A, B, head and penis; C–E, stomach. Scales: A–E, 500 m. Abbreviations: dgo, opening from stomach to digestive gland; ey, eye; int, intestine; oes, oesophagus, p, penis; pg, penial duck; sn, snout; ss, style sack; st, stomach; tn, tentacle.	FIGURE 9. Head, penis and stomach of Pseudotricula species. A, D, Pseudotricula conica, C.203680, main streamway near entrance, Damper Cave; C, Pseudotricula expandolabra, C.201493, paratype, Persephone Streamway, Bauhaus; B, E, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. A, B, head and penis; C–E, stomach. Scales: A–E, 500 m. Abbreviations: dgo, opening from stomach to digestive gland; ey, eye; int, intestine; oes, oesophagus, p, penis; pg, penial duck; sn, snout; ss, style sack; st, stomach; tn, tentacle.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D0CA21BE41FFDED7C72FD6D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050809/files/figure.png	https://doi.org/10.5281/zenodo.5050809	FIGURE 10. Female reproductive systems of Pseudotricula species. A ­ C, Pseudotricula expandolabra, A, B, C.201493, paratype, Persephone Streamway, Bauhaus, C, C.201815, Persephone Streamway, Bauhaus; D, Pseudotricula auriforma, C.201496, paratype, main streamway, Damper Cave; E, Pseudotricula arthurclarkei, C.203671, paratype, Quetzalcoatl Conduit; F, G, Pseudotricula conica, F, C203680, main streamway near entrance, Damper Cave, G, Cane Toad Abuse Streamway, Damper Cave. Scale: 500 m. Abbreviations: ag, albumen gland; bc, bursa copulatrix; bd, bursal duct; cg, capsule gland; co, coiled oviduct; po, genital opening; pw, pallial wall; sr, seminal receptacle; vc, ventral channel.	FIGURE 10. Female reproductive systems of Pseudotricula species. A ­ C, Pseudotricula expandolabra, A, B, C.201493, paratype, Persephone Streamway, Bauhaus, C, C.201815, Persephone Streamway, Bauhaus; D, Pseudotricula auriforma, C.201496, paratype, main streamway, Damper Cave; E, Pseudotricula arthurclarkei, C.203671, paratype, Quetzalcoatl Conduit; F, G, Pseudotricula conica, F, C203680, main streamway near entrance, Damper Cave, G, Cane Toad Abuse Streamway, Damper Cave. Scale: 500 m. Abbreviations: ag, albumen gland; bc, bursa copulatrix; bd, bursal duct; cg, capsule gland; co, coiled oviduct; po, genital opening; pw, pallial wall; sr, seminal receptacle; vc, ventral channel.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D15A215E41FFC807B18FDF5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050785/files/figure.png	https://doi.org/10.5281/zenodo.5050785	FIGURE 2. Shells of Pseudotricula species. A, Pseudotricula eberhardi, holotype, C.165052, Black Curtain Streamway, Cueva Blanca; B–E, Pseudotricula expandolabra, B, holotype, C.439673, Persephone Streamway, Bauhaus; C, paratype, C. 439674, Persephone Streamway, Bauhaus; D, C.201811, Persephone Pot, Bauhaus; E, C201495, Persephone Streamway, Bauhaus; F, Pseudotricula auriforma, holotype, C.439672, main streamway, Damper Cave. Scales: A F, 500 m.	FIGURE 2. Shells of Pseudotricula species. A, Pseudotricula eberhardi, holotype, C.165052, Black Curtain Streamway, Cueva Blanca; B–E, Pseudotricula expandolabra, B, holotype, C.439673, Persephone Streamway, Bauhaus; C, paratype, C. 439674, Persephone Streamway, Bauhaus; D, C.201811, Persephone Pot, Bauhaus; E, C201495, Persephone Streamway, Bauhaus; F, Pseudotricula auriforma, holotype, C.439672, main streamway, Damper Cave. Scales: A F, 500 m.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D15A215E41FFC807B18FDF5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050793/files/figure.png	https://doi.org/10.5281/zenodo.5050793	FIGURE 5. Shells of Pseudotricula species. A–F, Pseudotricula expandolabra, A–C, C.201489, Screaming Stals streamway, Bauhaus, D–F, Persephone Pot, Bauhaus, D, F, C.201811, E, C.201485, Persephone Pot, Bauhaus; G–I, Pseudotricula auriforma G, H, streamway near entrance, Damper Cave, I, main streamway, Damper Cave. Scales: 1 mm.	FIGURE 5. Shells of Pseudotricula species. A–F, Pseudotricula expandolabra, A–C, C.201489, Screaming Stals streamway, Bauhaus, D–F, Persephone Pot, Bauhaus, D, F, C.201811, E, C.201485, Persephone Pot, Bauhaus; G–I, Pseudotricula auriforma G, H, streamway near entrance, Damper Cave, I, main streamway, Damper Cave. Scales: 1 mm.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D15A215E41FFC807B18FDF5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050801/files/figure.png	https://doi.org/10.5281/zenodo.5050801	FIGURE 7. Opercula of Pseudotricula species. A, B, Pseudotricula expandolabra, C.201493, paratypes, main streamway, Bauhaus; C, Pseudotricula auriforma, C.201496, paratype, main streamway, Damper Cave; D, E, Pseudotricula arthurclarkei, C.203671, paratypes, Quetzalcoatl Conduit; F–I, Pseudotricula conica, F, G, C.203680, main streamway, near entrance, Damper Cave; H, I, C.201459, Screaming Stals streamway, Bauhaus; A, D, F, H, outer side; B, C, E, H, I, inner side; Scales: A–I, 200 m.	FIGURE 7. Opercula of Pseudotricula species. A, B, Pseudotricula expandolabra, C.201493, paratypes, main streamway, Bauhaus; C, Pseudotricula auriforma, C.201496, paratype, main streamway, Damper Cave; D, E, Pseudotricula arthurclarkei, C.203671, paratypes, Quetzalcoatl Conduit; F–I, Pseudotricula conica, F, G, C.203680, main streamway, near entrance, Damper Cave; H, I, C.201459, Screaming Stals streamway, Bauhaus; A, D, F, H, outer side; B, C, E, H, I, inner side; Scales: A–I, 200 m.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D15A215E41FFC807B18FDF5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050809/files/figure.png	https://doi.org/10.5281/zenodo.5050809	FIGURE 10. Female reproductive systems of Pseudotricula species. A ­ C, Pseudotricula expandolabra, A, B, C.201493, paratype, Persephone Streamway, Bauhaus, C, C.201815, Persephone Streamway, Bauhaus; D, Pseudotricula auriforma, C.201496, paratype, main streamway, Damper Cave; E, Pseudotricula arthurclarkei, C.203671, paratype, Quetzalcoatl Conduit; F, G, Pseudotricula conica, F, C203680, main streamway near entrance, Damper Cave, G, Cane Toad Abuse Streamway, Damper Cave. Scale: 500 m. Abbreviations: ag, albumen gland; bc, bursa copulatrix; bd, bursal duct; cg, capsule gland; co, coiled oviduct; po, genital opening; pw, pallial wall; sr, seminal receptacle; vc, ventral channel.	FIGURE 10. Female reproductive systems of Pseudotricula species. A ­ C, Pseudotricula expandolabra, A, B, C.201493, paratype, Persephone Streamway, Bauhaus, C, C.201815, Persephone Streamway, Bauhaus; D, Pseudotricula auriforma, C.201496, paratype, main streamway, Damper Cave; E, Pseudotricula arthurclarkei, C.203671, paratype, Quetzalcoatl Conduit; F, G, Pseudotricula conica, F, C203680, main streamway near entrance, Damper Cave, G, Cane Toad Abuse Streamway, Damper Cave. Scale: 500 m. Abbreviations: ag, albumen gland; bc, bursa copulatrix; bd, bursal duct; cg, capsule gland; co, coiled oviduct; po, genital opening; pw, pallial wall; sr, seminal receptacle; vc, ventral channel.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D15A215E41FFC807B18FDF5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050803/files/figure.png	https://doi.org/10.5281/zenodo.5050803	FIGURE 8. Pallial cavities of male specimens of Pseudotricula and Nanocochlea species. A, Pseudotricula expandolabra, C.201493, paratype, Persephone Streamway, Bauhaus; B, Pseudotricula arthurclarkei, C.203671, paratype, Quetzalcoatl Conduit; C, Pseudotricula auriforma, C.201496, paratype, main streamway, Damper Cave; D, E, Pseudotricula conica, D, C.203680, main streamway near entrance, Damper Cave, E, Cane Toad Abuse Streamway, Damper Cave; F, Nanocochlea exigua, C.203677, paratype, Cane Toad Abuse Streamway, Damper Cave; G, Nanocochlea damperensis, C.203685, paratype, first crossing, Damper Creek; H, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. Scales: A–D, F–H, 500 m, E, 1 mm. Abbreviations: a, anus; ct, ctenidium; ev, efferent vein; os, osphradium; pc, pericardium; pe, pallial edge; ppw, posterior pallial wall; r, rectum; rg, renal gland. The stippled area represents the hypobranchial gland.	FIGURE 8. Pallial cavities of male specimens of Pseudotricula and Nanocochlea species. A, Pseudotricula expandolabra, C.201493, paratype, Persephone Streamway, Bauhaus; B, Pseudotricula arthurclarkei, C.203671, paratype, Quetzalcoatl Conduit; C, Pseudotricula auriforma, C.201496, paratype, main streamway, Damper Cave; D, E, Pseudotricula conica, D, C.203680, main streamway near entrance, Damper Cave, E, Cane Toad Abuse Streamway, Damper Cave; F, Nanocochlea exigua, C.203677, paratype, Cane Toad Abuse Streamway, Damper Cave; G, Nanocochlea damperensis, C.203685, paratype, first crossing, Damper Creek; H, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. Scales: A–D, F–H, 500 m, E, 1 mm. Abbreviations: a, anus; ct, ctenidium; ev, efferent vein; os, osphradium; pc, pericardium; pe, pallial edge; ppw, posterior pallial wall; r, rectum; rg, renal gland. The stippled area represents the hypobranchial gland.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D1BA216E41FFD187AB4F8DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050813/files/figure.png	https://doi.org/10.5281/zenodo.5050813	FIGURE 11. Shells of Pseudotricula species. A ­ C, Pseudotricula elongata, Persephone Pot, A, holotype, C.166854; B,C, paratypes, B, C.203713, C, C.201809; D–F, Pseudotricula arthurclarkei, C.203671, paratypes, Quetzalcoatl Conduit. Scales: A ­ C, 500 m; D–F, 1 mm.	FIGURE 11. Shells of Pseudotricula species. A ­ C, Pseudotricula elongata, Persephone Pot, A, holotype, C.166854; B,C, paratypes, B, C.203713, C, C.201809; D–F, Pseudotricula arthurclarkei, C.203671, paratypes, Quetzalcoatl Conduit. Scales: A ­ C, 500 m; D–F, 1 mm.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D19A211E41FFED07FA2FA65.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050787/files/figure.png	https://doi.org/10.5281/zenodo.5050787	FIGURE 3: Radulae of Pseudotricula species. A, B, Pseudotricula eberhardi, C.165053, Black Curtain Streamway, Cueva Blanca; C–H, Pseudotricula expandolabra, C, D, C.201495, Persephone Streamway, Bauhaus, E, G, H, C.201811, Persephone Pot, Bauhaus, F, C.201822, main streamway, Bauhaus; I, Pseudotricula arthurclarkei, C.203671, paratype, Quetzalcoatl Conduit; J–L, Pseudotricula conica, C.203676, paratype, Cane Toad Abuse Streamway, Damper Cave. A, C, G, K, central, lateral inner and outer marginal teeth; B, E, L, central teeth; D, outer marginal teeth; F, J, lateral and inner marginal teeth; H, lateral tooth; I, central and lateral teeth. Scales: A, B, E, H–J, L 10 m; C, F, G, K, 20 m; D, 5 m.	FIGURE 3: Radulae of Pseudotricula species. A, B, Pseudotricula eberhardi, C.165053, Black Curtain Streamway, Cueva Blanca; C–H, Pseudotricula expandolabra, C, D, C.201495, Persephone Streamway, Bauhaus, E, G, H, C.201811, Persephone Pot, Bauhaus, F, C.201822, main streamway, Bauhaus; I, Pseudotricula arthurclarkei, C.203671, paratype, Quetzalcoatl Conduit; J–L, Pseudotricula conica, C.203676, paratype, Cane Toad Abuse Streamway, Damper Cave. A, C, G, K, central, lateral inner and outer marginal teeth; B, E, L, central teeth; D, outer marginal teeth; F, J, lateral and inner marginal teeth; H, lateral tooth; I, central and lateral teeth. Scales: A, B, E, H–J, L 10 m; C, F, G, K, 20 m; D, 5 m.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D19A211E41FFED07FA2FA65.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050801/files/figure.png	https://doi.org/10.5281/zenodo.5050801	FIGURE 7. Opercula of Pseudotricula species. A, B, Pseudotricula expandolabra, C.201493, paratypes, main streamway, Bauhaus; C, Pseudotricula auriforma, C.201496, paratype, main streamway, Damper Cave; D, E, Pseudotricula arthurclarkei, C.203671, paratypes, Quetzalcoatl Conduit; F–I, Pseudotricula conica, F, G, C.203680, main streamway, near entrance, Damper Cave; H, I, C.201459, Screaming Stals streamway, Bauhaus; A, D, F, H, outer side; B, C, E, H, I, inner side; Scales: A–I, 200 m.	FIGURE 7. Opercula of Pseudotricula species. A, B, Pseudotricula expandolabra, C.201493, paratypes, main streamway, Bauhaus; C, Pseudotricula auriforma, C.201496, paratype, main streamway, Damper Cave; D, E, Pseudotricula arthurclarkei, C.203671, paratypes, Quetzalcoatl Conduit; F–I, Pseudotricula conica, F, G, C.203680, main streamway, near entrance, Damper Cave; H, I, C.201459, Screaming Stals streamway, Bauhaus; A, D, F, H, outer side; B, C, E, H, I, inner side; Scales: A–I, 200 m.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D19A211E41FFED07FA2FA65.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050803/files/figure.png	https://doi.org/10.5281/zenodo.5050803	FIGURE 8. Pallial cavities of male specimens of Pseudotricula and Nanocochlea species. A, Pseudotricula expandolabra, C.201493, paratype, Persephone Streamway, Bauhaus; B, Pseudotricula arthurclarkei, C.203671, paratype, Quetzalcoatl Conduit; C, Pseudotricula auriforma, C.201496, paratype, main streamway, Damper Cave; D, E, Pseudotricula conica, D, C.203680, main streamway near entrance, Damper Cave, E, Cane Toad Abuse Streamway, Damper Cave; F, Nanocochlea exigua, C.203677, paratype, Cane Toad Abuse Streamway, Damper Cave; G, Nanocochlea damperensis, C.203685, paratype, first crossing, Damper Creek; H, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. Scales: A–D, F–H, 500 m, E, 1 mm. Abbreviations: a, anus; ct, ctenidium; ev, efferent vein; os, osphradium; pc, pericardium; pe, pallial edge; ppw, posterior pallial wall; r, rectum; rg, renal gland. The stippled area represents the hypobranchial gland.	FIGURE 8. Pallial cavities of male specimens of Pseudotricula and Nanocochlea species. A, Pseudotricula expandolabra, C.201493, paratype, Persephone Streamway, Bauhaus; B, Pseudotricula arthurclarkei, C.203671, paratype, Quetzalcoatl Conduit; C, Pseudotricula auriforma, C.201496, paratype, main streamway, Damper Cave; D, E, Pseudotricula conica, D, C.203680, main streamway near entrance, Damper Cave, E, Cane Toad Abuse Streamway, Damper Cave; F, Nanocochlea exigua, C.203677, paratype, Cane Toad Abuse Streamway, Damper Cave; G, Nanocochlea damperensis, C.203685, paratype, first crossing, Damper Creek; H, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. Scales: A–D, F–H, 500 m, E, 1 mm. Abbreviations: a, anus; ct, ctenidium; ev, efferent vein; os, osphradium; pc, pericardium; pe, pallial edge; ppw, posterior pallial wall; r, rectum; rg, renal gland. The stippled area represents the hypobranchial gland.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D19A211E41FFED07FA2FA65.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050809/files/figure.png	https://doi.org/10.5281/zenodo.5050809	FIGURE 10. Female reproductive systems of Pseudotricula species. A ­ C, Pseudotricula expandolabra, A, B, C.201493, paratype, Persephone Streamway, Bauhaus, C, C.201815, Persephone Streamway, Bauhaus; D, Pseudotricula auriforma, C.201496, paratype, main streamway, Damper Cave; E, Pseudotricula arthurclarkei, C.203671, paratype, Quetzalcoatl Conduit; F, G, Pseudotricula conica, F, C203680, main streamway near entrance, Damper Cave, G, Cane Toad Abuse Streamway, Damper Cave. Scale: 500 m. Abbreviations: ag, albumen gland; bc, bursa copulatrix; bd, bursal duct; cg, capsule gland; co, coiled oviduct; po, genital opening; pw, pallial wall; sr, seminal receptacle; vc, ventral channel.	FIGURE 10. Female reproductive systems of Pseudotricula species. A ­ C, Pseudotricula expandolabra, A, B, C.201493, paratype, Persephone Streamway, Bauhaus, C, C.201815, Persephone Streamway, Bauhaus; D, Pseudotricula auriforma, C.201496, paratype, main streamway, Damper Cave; E, Pseudotricula arthurclarkei, C.203671, paratype, Quetzalcoatl Conduit; F, G, Pseudotricula conica, F, C203680, main streamway near entrance, Damper Cave, G, Cane Toad Abuse Streamway, Damper Cave. Scale: 500 m. Abbreviations: ag, albumen gland; bc, bursa copulatrix; bd, bursal duct; cg, capsule gland; co, coiled oviduct; po, genital opening; pw, pallial wall; sr, seminal receptacle; vc, ventral channel.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D19A211E41FFED07FA2FA65.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050813/files/figure.png	https://doi.org/10.5281/zenodo.5050813	FIGURE 11. Shells of Pseudotricula species. A ­ C, Pseudotricula elongata, Persephone Pot, A, holotype, C.166854; B,C, paratypes, B, C.203713, C, C.201809; D–F, Pseudotricula arthurclarkei, C.203671, paratypes, Quetzalcoatl Conduit. Scales: A ­ C, 500 m; D–F, 1 mm.	FIGURE 11. Shells of Pseudotricula species. A ­ C, Pseudotricula elongata, Persephone Pot, A, holotype, C.166854; B,C, paratypes, B, C.203713, C, C.201809; D–F, Pseudotricula arthurclarkei, C.203671, paratypes, Quetzalcoatl Conduit. Scales: A ­ C, 500 m; D–F, 1 mm.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D19A211E41FFED07FA2FA65.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050815/files/figure.png	https://doi.org/10.5281/zenodo.5050815	FIGURE 12. Shells of Pseudotricula species. A, Pseudotricula arthurclarkei, holotype, C.439656, Quetzalcoatl Conduit; B ­ D, Pseudotricula conica, B, holotype, Cane Toad Abuse Streamway, Damper Cave; C, C.203680, main streamway near entrance, Damper Cave; D, C.201813, Persephone Pot, Bauhaus. Scales: A–D, 500 m.	FIGURE 12. Shells of Pseudotricula species. A, Pseudotricula arthurclarkei, holotype, C.439656, Quetzalcoatl Conduit; B ­ D, Pseudotricula conica, B, holotype, Cane Toad Abuse Streamway, Damper Cave; C, C.203680, main streamway near entrance, Damper Cave; D, C.201813, Persephone Pot, Bauhaus. Scales: A–D, 500 m.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D1FA22DE41FF9887B18FC95.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050787/files/figure.png	https://doi.org/10.5281/zenodo.5050787	FIGURE 3: Radulae of Pseudotricula species. A, B, Pseudotricula eberhardi, C.165053, Black Curtain Streamway, Cueva Blanca; C–H, Pseudotricula expandolabra, C, D, C.201495, Persephone Streamway, Bauhaus, E, G, H, C.201811, Persephone Pot, Bauhaus, F, C.201822, main streamway, Bauhaus; I, Pseudotricula arthurclarkei, C.203671, paratype, Quetzalcoatl Conduit; J–L, Pseudotricula conica, C.203676, paratype, Cane Toad Abuse Streamway, Damper Cave. A, C, G, K, central, lateral inner and outer marginal teeth; B, E, L, central teeth; D, outer marginal teeth; F, J, lateral and inner marginal teeth; H, lateral tooth; I, central and lateral teeth. Scales: A, B, E, H–J, L 10 m; C, F, G, K, 20 m; D, 5 m.	FIGURE 3: Radulae of Pseudotricula species. A, B, Pseudotricula eberhardi, C.165053, Black Curtain Streamway, Cueva Blanca; C–H, Pseudotricula expandolabra, C, D, C.201495, Persephone Streamway, Bauhaus, E, G, H, C.201811, Persephone Pot, Bauhaus, F, C.201822, main streamway, Bauhaus; I, Pseudotricula arthurclarkei, C.203671, paratype, Quetzalcoatl Conduit; J–L, Pseudotricula conica, C.203676, paratype, Cane Toad Abuse Streamway, Damper Cave. A, C, G, K, central, lateral inner and outer marginal teeth; B, E, L, central teeth; D, outer marginal teeth; F, J, lateral and inner marginal teeth; H, lateral tooth; I, central and lateral teeth. Scales: A, B, E, H–J, L 10 m; C, F, G, K, 20 m; D, 5 m.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D1FA22DE41FF9887B18FC95.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050791/files/figure.png	https://doi.org/10.5281/zenodo.5050791	FIGURE 4. Dorsal views of penes of Pseudotricula and Nanocochlea species. A,. Pseudotricula eberhardi; B, Pseudotricula expandolabra; C, Nanocochlea stylesae; D, Nanocochlea damperensis; E, Pseudotricula conica. Scales A, B, E, 0.2 mm; C, D, 0.1 mm. Abbreviations: ds, distal swelling; pd, penial duct.	FIGURE 4. Dorsal views of penes of Pseudotricula and Nanocochlea species. A,. Pseudotricula eberhardi; B, Pseudotricula expandolabra; C, Nanocochlea stylesae; D, Nanocochlea damperensis; E, Pseudotricula conica. Scales A, B, E, 0.2 mm; C, D, 0.1 mm. Abbreviations: ds, distal swelling; pd, penial duct.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D1FA22DE41FF9887B18FC95.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050797/files/figure.png	https://doi.org/10.5281/zenodo.5050797	FIGURE 6. Protoconchs and teleoconch microsculpture of Pseudotricula and Nanocochlea species. A–D, Pseudotricula expandolabra, A, B, D, C.201493, paratype, main streamway, Bauhaus, C, C.201495, Persephone Streamway, Bauhaus; E–G, Pseudotricula conica, E, F, C.201464, Cane Toad Abuse Streamway, Damper Cave, G, Persephone Pot, Bauhaus; Honey and Cream Streamway, Damper Cave; H, Nanocochlea exigua, C.201475; I, J, Nanocochlea stylesae, C.201476, paratypes, Malani Creek; K–M, Nanocochlea damperensis, K, L, C.203685, paratypes, first crossing, Damper Creek, M, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; N, O, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. A, B, E, F, H, K–N, protoconch; C, D, G, I, J, O, microsculpture. Scales: A, C–E, J, K, M, O, 200 m; B, F–I, L, N, 100 m.	FIGURE 6. Protoconchs and teleoconch microsculpture of Pseudotricula and Nanocochlea species. A–D, Pseudotricula expandolabra, A, B, D, C.201493, paratype, main streamway, Bauhaus, C, C.201495, Persephone Streamway, Bauhaus; E–G, Pseudotricula conica, E, F, C.201464, Cane Toad Abuse Streamway, Damper Cave, G, Persephone Pot, Bauhaus; Honey and Cream Streamway, Damper Cave; H, Nanocochlea exigua, C.201475; I, J, Nanocochlea stylesae, C.201476, paratypes, Malani Creek; K–M, Nanocochlea damperensis, K, L, C.203685, paratypes, first crossing, Damper Creek, M, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; N, O, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. A, B, E, F, H, K–N, protoconch; C, D, G, I, J, O, microsculpture. Scales: A, C–E, J, K, M, O, 200 m; B, F–I, L, N, 100 m.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D1FA22DE41FF9887B18FC95.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050801/files/figure.png	https://doi.org/10.5281/zenodo.5050801	FIGURE 7. Opercula of Pseudotricula species. A, B, Pseudotricula expandolabra, C.201493, paratypes, main streamway, Bauhaus; C, Pseudotricula auriforma, C.201496, paratype, main streamway, Damper Cave; D, E, Pseudotricula arthurclarkei, C.203671, paratypes, Quetzalcoatl Conduit; F–I, Pseudotricula conica, F, G, C.203680, main streamway, near entrance, Damper Cave; H, I, C.201459, Screaming Stals streamway, Bauhaus; A, D, F, H, outer side; B, C, E, H, I, inner side; Scales: A–I, 200 m.	FIGURE 7. Opercula of Pseudotricula species. A, B, Pseudotricula expandolabra, C.201493, paratypes, main streamway, Bauhaus; C, Pseudotricula auriforma, C.201496, paratype, main streamway, Damper Cave; D, E, Pseudotricula arthurclarkei, C.203671, paratypes, Quetzalcoatl Conduit; F–I, Pseudotricula conica, F, G, C.203680, main streamway, near entrance, Damper Cave; H, I, C.201459, Screaming Stals streamway, Bauhaus; A, D, F, H, outer side; B, C, E, H, I, inner side; Scales: A–I, 200 m.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D1FA22DE41FF9887B18FC95.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050803/files/figure.png	https://doi.org/10.5281/zenodo.5050803	FIGURE 8. Pallial cavities of male specimens of Pseudotricula and Nanocochlea species. A, Pseudotricula expandolabra, C.201493, paratype, Persephone Streamway, Bauhaus; B, Pseudotricula arthurclarkei, C.203671, paratype, Quetzalcoatl Conduit; C, Pseudotricula auriforma, C.201496, paratype, main streamway, Damper Cave; D, E, Pseudotricula conica, D, C.203680, main streamway near entrance, Damper Cave, E, Cane Toad Abuse Streamway, Damper Cave; F, Nanocochlea exigua, C.203677, paratype, Cane Toad Abuse Streamway, Damper Cave; G, Nanocochlea damperensis, C.203685, paratype, first crossing, Damper Creek; H, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. Scales: A–D, F–H, 500 m, E, 1 mm. Abbreviations: a, anus; ct, ctenidium; ev, efferent vein; os, osphradium; pc, pericardium; pe, pallial edge; ppw, posterior pallial wall; r, rectum; rg, renal gland. The stippled area represents the hypobranchial gland.	FIGURE 8. Pallial cavities of male specimens of Pseudotricula and Nanocochlea species. A, Pseudotricula expandolabra, C.201493, paratype, Persephone Streamway, Bauhaus; B, Pseudotricula arthurclarkei, C.203671, paratype, Quetzalcoatl Conduit; C, Pseudotricula auriforma, C.201496, paratype, main streamway, Damper Cave; D, E, Pseudotricula conica, D, C.203680, main streamway near entrance, Damper Cave, E, Cane Toad Abuse Streamway, Damper Cave; F, Nanocochlea exigua, C.203677, paratype, Cane Toad Abuse Streamway, Damper Cave; G, Nanocochlea damperensis, C.203685, paratype, first crossing, Damper Creek; H, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. Scales: A–D, F–H, 500 m, E, 1 mm. Abbreviations: a, anus; ct, ctenidium; ev, efferent vein; os, osphradium; pc, pericardium; pe, pallial edge; ppw, posterior pallial wall; r, rectum; rg, renal gland. The stippled area represents the hypobranchial gland.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D1FA22DE41FF9887B18FC95.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050807/files/figure.png	https://doi.org/10.5281/zenodo.5050807	FIGURE 9. Head, penis and stomach of Pseudotricula species. A, D, Pseudotricula conica, C.203680, main streamway near entrance, Damper Cave; C, Pseudotricula expandolabra, C.201493, paratype, Persephone Streamway, Bauhaus; B, E, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. A, B, head and penis; C–E, stomach. Scales: A–E, 500 m. Abbreviations: dgo, opening from stomach to digestive gland; ey, eye; int, intestine; oes, oesophagus, p, penis; pg, penial duck; sn, snout; ss, style sack; st, stomach; tn, tentacle.	FIGURE 9. Head, penis and stomach of Pseudotricula species. A, D, Pseudotricula conica, C.203680, main streamway near entrance, Damper Cave; C, Pseudotricula expandolabra, C.201493, paratype, Persephone Streamway, Bauhaus; B, E, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. A, B, head and penis; C–E, stomach. Scales: A–E, 500 m. Abbreviations: dgo, opening from stomach to digestive gland; ey, eye; int, intestine; oes, oesophagus, p, penis; pg, penial duck; sn, snout; ss, style sack; st, stomach; tn, tentacle.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D1FA22DE41FF9887B18FC95.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050809/files/figure.png	https://doi.org/10.5281/zenodo.5050809	FIGURE 10. Female reproductive systems of Pseudotricula species. A ­ C, Pseudotricula expandolabra, A, B, C.201493, paratype, Persephone Streamway, Bauhaus, C, C.201815, Persephone Streamway, Bauhaus; D, Pseudotricula auriforma, C.201496, paratype, main streamway, Damper Cave; E, Pseudotricula arthurclarkei, C.203671, paratype, Quetzalcoatl Conduit; F, G, Pseudotricula conica, F, C203680, main streamway near entrance, Damper Cave, G, Cane Toad Abuse Streamway, Damper Cave. Scale: 500 m. Abbreviations: ag, albumen gland; bc, bursa copulatrix; bd, bursal duct; cg, capsule gland; co, coiled oviduct; po, genital opening; pw, pallial wall; sr, seminal receptacle; vc, ventral channel.	FIGURE 10. Female reproductive systems of Pseudotricula species. A ­ C, Pseudotricula expandolabra, A, B, C.201493, paratype, Persephone Streamway, Bauhaus, C, C.201815, Persephone Streamway, Bauhaus; D, Pseudotricula auriforma, C.201496, paratype, main streamway, Damper Cave; E, Pseudotricula arthurclarkei, C.203671, paratype, Quetzalcoatl Conduit; F, G, Pseudotricula conica, F, C203680, main streamway near entrance, Damper Cave, G, Cane Toad Abuse Streamway, Damper Cave. Scale: 500 m. Abbreviations: ag, albumen gland; bc, bursa copulatrix; bd, bursal duct; cg, capsule gland; co, coiled oviduct; po, genital opening; pw, pallial wall; sr, seminal receptacle; vc, ventral channel.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D1FA22DE41FF9887B18FC95.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050815/files/figure.png	https://doi.org/10.5281/zenodo.5050815	FIGURE 12. Shells of Pseudotricula species. A, Pseudotricula arthurclarkei, holotype, C.439656, Quetzalcoatl Conduit; B ­ D, Pseudotricula conica, B, holotype, Cane Toad Abuse Streamway, Damper Cave; C, C.203680, main streamway near entrance, Damper Cave; D, C.201813, Persephone Pot, Bauhaus. Scales: A–D, 500 m.	FIGURE 12. Shells of Pseudotricula species. A, Pseudotricula arthurclarkei, holotype, C.439656, Quetzalcoatl Conduit; B ­ D, Pseudotricula conica, B, holotype, Cane Toad Abuse Streamway, Damper Cave; C, C.203680, main streamway near entrance, Damper Cave; D, C.201813, Persephone Pot, Bauhaus. Scales: A–D, 500 m.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D1FA22DE41FF9887B18FC95.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050817/files/figure.png	https://doi.org/10.5281/zenodo.5050817	FIGURE 13. Shells of Pseudotricula species. A ­ F, Pseudotricula conica, A, C.203694, Black Curtain streamway, Cueva Blanca, B, Screaming Stals streamway, Bauhaus; C, C.201813, Persephone Pot, Bauhaus, D, C.203464, Cane Toad Abuse Streamway, Damper Cave, E, C.201488, main streamway, Damper Cave, F, C.203692, Damper Cave. Scales: 1 mm.	FIGURE 13. Shells of Pseudotricula species. A ­ F, Pseudotricula conica, A, C.203694, Black Curtain streamway, Cueva Blanca, B, Screaming Stals streamway, Bauhaus; C, C.201813, Persephone Pot, Bauhaus, D, C.203464, Cane Toad Abuse Streamway, Damper Cave, E, C.201488, main streamway, Damper Cave, F, C.203692, Damper Cave. Scales: 1 mm.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D23A22BE41FFCB87AB0F9F2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050797/files/figure.png	https://doi.org/10.5281/zenodo.5050797	FIGURE 6. Protoconchs and teleoconch microsculpture of Pseudotricula and Nanocochlea species. A–D, Pseudotricula expandolabra, A, B, D, C.201493, paratype, main streamway, Bauhaus, C, C.201495, Persephone Streamway, Bauhaus; E–G, Pseudotricula conica, E, F, C.201464, Cane Toad Abuse Streamway, Damper Cave, G, Persephone Pot, Bauhaus; Honey and Cream Streamway, Damper Cave; H, Nanocochlea exigua, C.201475; I, J, Nanocochlea stylesae, C.201476, paratypes, Malani Creek; K–M, Nanocochlea damperensis, K, L, C.203685, paratypes, first crossing, Damper Creek, M, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; N, O, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. A, B, E, F, H, K–N, protoconch; C, D, G, I, J, O, microsculpture. Scales: A, C–E, J, K, M, O, 200 m; B, F–I, L, N, 100 m.	FIGURE 6. Protoconchs and teleoconch microsculpture of Pseudotricula and Nanocochlea species. A–D, Pseudotricula expandolabra, A, B, D, C.201493, paratype, main streamway, Bauhaus, C, C.201495, Persephone Streamway, Bauhaus; E–G, Pseudotricula conica, E, F, C.201464, Cane Toad Abuse Streamway, Damper Cave, G, Persephone Pot, Bauhaus; Honey and Cream Streamway, Damper Cave; H, Nanocochlea exigua, C.201475; I, J, Nanocochlea stylesae, C.201476, paratypes, Malani Creek; K–M, Nanocochlea damperensis, K, L, C.203685, paratypes, first crossing, Damper Creek, M, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; N, O, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. A, B, E, F, H, K–N, protoconch; C, D, G, I, J, O, microsculpture. Scales: A, C–E, J, K, M, O, 200 m; B, F–I, L, N, 100 m.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D23A22BE41FFCB87AB0F9F2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050803/files/figure.png	https://doi.org/10.5281/zenodo.5050803	FIGURE 8. Pallial cavities of male specimens of Pseudotricula and Nanocochlea species. A, Pseudotricula expandolabra, C.201493, paratype, Persephone Streamway, Bauhaus; B, Pseudotricula arthurclarkei, C.203671, paratype, Quetzalcoatl Conduit; C, Pseudotricula auriforma, C.201496, paratype, main streamway, Damper Cave; D, E, Pseudotricula conica, D, C.203680, main streamway near entrance, Damper Cave, E, Cane Toad Abuse Streamway, Damper Cave; F, Nanocochlea exigua, C.203677, paratype, Cane Toad Abuse Streamway, Damper Cave; G, Nanocochlea damperensis, C.203685, paratype, first crossing, Damper Creek; H, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. Scales: A–D, F–H, 500 m, E, 1 mm. Abbreviations: a, anus; ct, ctenidium; ev, efferent vein; os, osphradium; pc, pericardium; pe, pallial edge; ppw, posterior pallial wall; r, rectum; rg, renal gland. The stippled area represents the hypobranchial gland.	FIGURE 8. Pallial cavities of male specimens of Pseudotricula and Nanocochlea species. A, Pseudotricula expandolabra, C.201493, paratype, Persephone Streamway, Bauhaus; B, Pseudotricula arthurclarkei, C.203671, paratype, Quetzalcoatl Conduit; C, Pseudotricula auriforma, C.201496, paratype, main streamway, Damper Cave; D, E, Pseudotricula conica, D, C.203680, main streamway near entrance, Damper Cave, E, Cane Toad Abuse Streamway, Damper Cave; F, Nanocochlea exigua, C.203677, paratype, Cane Toad Abuse Streamway, Damper Cave; G, Nanocochlea damperensis, C.203685, paratype, first crossing, Damper Creek; H, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. Scales: A–D, F–H, 500 m, E, 1 mm. Abbreviations: a, anus; ct, ctenidium; ev, efferent vein; os, osphradium; pc, pericardium; pe, pallial edge; ppw, posterior pallial wall; r, rectum; rg, renal gland. The stippled area represents the hypobranchial gland.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D23A22BE41FFCB87AB0F9F2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050807/files/figure.png	https://doi.org/10.5281/zenodo.5050807	FIGURE 9. Head, penis and stomach of Pseudotricula species. A, D, Pseudotricula conica, C.203680, main streamway near entrance, Damper Cave; C, Pseudotricula expandolabra, C.201493, paratype, Persephone Streamway, Bauhaus; B, E, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. A, B, head and penis; C–E, stomach. Scales: A–E, 500 m. Abbreviations: dgo, opening from stomach to digestive gland; ey, eye; int, intestine; oes, oesophagus, p, penis; pg, penial duck; sn, snout; ss, style sack; st, stomach; tn, tentacle.	FIGURE 9. Head, penis and stomach of Pseudotricula species. A, D, Pseudotricula conica, C.203680, main streamway near entrance, Damper Cave; C, Pseudotricula expandolabra, C.201493, paratype, Persephone Streamway, Bauhaus; B, E, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. A, B, head and penis; C–E, stomach. Scales: A–E, 500 m. Abbreviations: dgo, opening from stomach to digestive gland; ey, eye; int, intestine; oes, oesophagus, p, penis; pg, penial duck; sn, snout; ss, style sack; st, stomach; tn, tentacle.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D23A22BE41FFCB87AB0F9F2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050821/files/figure.png	https://doi.org/10.5281/zenodo.5050821	FIGURE 14. Shells of Nanocochlea species and Pseudotricula progenitor. A ­ C, Nanocochlea exigua, A, holotype, C.439393, Cane Toad Abuse Streamway, Damper Cave; B, C, C.201465, main streamway, Damper Cave; D, Nanocochlea stylesae, holotype, C.439398, Malani Creek; E–G, Nanocochlea damperensis, E, C.203685, paratype, first crossing, Damper Creek; F, holotype, C.439399, first crossing, Damper Creek, F, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; H, I, Pseudotricula progenitor, H, paratype, C.439397, Persephone Streamway, Bauhaus; I, holotype, C.439396, Persephone Streamway, Bauhaus. Scales: 500 m.	FIGURE 14. Shells of Nanocochlea species and Pseudotricula progenitor. A ­ C, Nanocochlea exigua, A, holotype, C.439393, Cane Toad Abuse Streamway, Damper Cave; B, C, C.201465, main streamway, Damper Cave; D, Nanocochlea stylesae, holotype, C.439398, Malani Creek; E–G, Nanocochlea damperensis, E, C.203685, paratype, first crossing, Damper Creek; F, holotype, C.439399, first crossing, Damper Creek, F, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; H, I, Pseudotricula progenitor, H, paratype, C.439397, Persephone Streamway, Bauhaus; I, holotype, C.439396, Persephone Streamway, Bauhaus. Scales: 500 m.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D23A22BE41FFCB87AB0F9F2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050823/files/figure.png	https://doi.org/10.5281/zenodo.5050823	FIGURE 15. Shells of Nanocochlea species and Pseudotricula progenitor. A ­ C, Nanocochlea exigua, A, C.203674, outside Damper Cave, Damper Creek B, C.166855, Persephone Pot, Bauhaus, C, C.201465, main streamway, Damper Cave; D–F, Nanocochlea stylesae, D, C.201476, paratype, Malani Creek, E, F, C.203686, first crossing, Damper Creek; G­I, Nanocochlea damperensis, G,H, C.303685, seepages at Bauhaus entrance, I, C.203685, paratype, first crossing, Damper Creek; J–L, Pseudotricula progenitor, J, L, C.201463, paratype, Persephone streamway, Bauhaus, K, C.201454, Persephone, Bauhaus. Scales: 1 mm.	FIGURE 15. Shells of Nanocochlea species and Pseudotricula progenitor. A ­ C, Nanocochlea exigua, A, C.203674, outside Damper Cave, Damper Creek B, C.166855, Persephone Pot, Bauhaus, C, C.201465, main streamway, Damper Cave; D–F, Nanocochlea stylesae, D, C.201476, paratype, Malani Creek, E, F, C.203686, first crossing, Damper Creek; G­I, Nanocochlea damperensis, G,H, C.303685, seepages at Bauhaus entrance, I, C.203685, paratype, first crossing, Damper Creek; J–L, Pseudotricula progenitor, J, L, C.201463, paratype, Persephone streamway, Bauhaus, K, C.201454, Persephone, Bauhaus. Scales: 1 mm.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D23A22BE41FFCB87AB0F9F2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5052059/files/figure.png	https://doi.org/10.5281/zenodo.5052059	FIGURE 16. Opercula of Nanocochlea species and Pseudotricula progenitor. A ­ C, Nanocochlea exigua, C.203677, paratypes, Cane Toad Abuse Streamway, Damper Cave; D ­ F, Nanocochlea damperensis; C–E, C.203685, paratypes, first crossing, Damper Creek; F, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; G, H, Pseudotricula progenitor, C.201821, main streamway, Bauhaus; A, D, G, outer side; B, C, E, F, H, inner side; Scales: A, B, D, 200 m, C, E–H, 100 m.	FIGURE 16. Opercula of Nanocochlea species and Pseudotricula progenitor. A ­ C, Nanocochlea exigua, C.203677, paratypes, Cane Toad Abuse Streamway, Damper Cave; D ­ F, Nanocochlea damperensis; C–E, C.203685, paratypes, first crossing, Damper Creek; F, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; G, H, Pseudotricula progenitor, C.201821, main streamway, Bauhaus; A, D, G, outer side; B, C, E, F, H, inner side; Scales: A, B, D, 200 m, C, E–H, 100 m.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D23A22BE41FFCB87AB0F9F2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050829/files/figure.png	https://doi.org/10.5281/zenodo.5050829	FIGURE 17. Radulae of Nanocochlea species Pseudotricula progenitor. A ­ C, Nanocochlea exigua, A, C.201475, Honey and Cream Streamway, Damper Cave, B, D, C.201465, main streamway, Damper Cave; D– E, Nanocochlea stylesae, C.201476, paratypes, Malani Creek; G–K, Nanocochlea damperensis, G, H, C.203685, paratypes, first crossing, Damper Creek, I–K, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; L–O, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. A, E, central and lateral teeth; B, F, I, M, central, lateral, inner and outer marginal teeth; C, D, J, N central teeth; G, L, lateral teeth; H, central lateral inner and outer marginal teeth; K, outer marginal teeth. Scales: A, G–H, K, L, 10 m; B, I, M, 20 m; C, D, J, N, 5 m.	FIGURE 17. Radulae of Nanocochlea species Pseudotricula progenitor. A ­ C, Nanocochlea exigua, A, C.201475, Honey and Cream Streamway, Damper Cave, B, D, C.201465, main streamway, Damper Cave; D– E, Nanocochlea stylesae, C.201476, paratypes, Malani Creek; G–K, Nanocochlea damperensis, G, H, C.203685, paratypes, first crossing, Damper Creek, I–K, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; L–O, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. A, E, central and lateral teeth; B, F, I, M, central, lateral, inner and outer marginal teeth; C, D, J, N central teeth; G, L, lateral teeth; H, central lateral inner and outer marginal teeth; K, outer marginal teeth. Scales: A, G–H, K, L, 10 m; B, I, M, 20 m; C, D, J, N, 5 m.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D23A22BE41FFCB87AB0F9F2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050831/files/figure.png	https://doi.org/10.5281/zenodo.5050831	FIGURE 18. Female reproductive system of Nanocochlea species and Pseudotricula progenitor. A, Nanocochlea exigua, C.203677, paratype, Cane Toad Abuse Streamway, Damper Cave; B, Nanocochlea damperensis, C.203685, paratype, first crossing, Damper Creek; C, D, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. Scales: 500 m. Abbreviations: ag, albumen gland; bc, bursa copulatrix; bd, bursal duct; cg, capsule gland; co, coiled oviduct; po, genital opening; pw, pallial wall; sr, seminal receptacle; vc, ventral channel.	FIGURE 18. Female reproductive system of Nanocochlea species and Pseudotricula progenitor. A, Nanocochlea exigua, C.203677, paratype, Cane Toad Abuse Streamway, Damper Cave; B, Nanocochlea damperensis, C.203685, paratype, first crossing, Damper Creek; C, D, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. Scales: 500 m. Abbreviations: ag, albumen gland; bc, bursa copulatrix; bd, bursal duct; cg, capsule gland; co, coiled oviduct; po, genital opening; pw, pallial wall; sr, seminal receptacle; vc, ventral channel.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D2BA220E41FF9887F43FCBD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050797/files/figure.png	https://doi.org/10.5281/zenodo.5050797	FIGURE 6. Protoconchs and teleoconch microsculpture of Pseudotricula and Nanocochlea species. A–D, Pseudotricula expandolabra, A, B, D, C.201493, paratype, main streamway, Bauhaus, C, C.201495, Persephone Streamway, Bauhaus; E–G, Pseudotricula conica, E, F, C.201464, Cane Toad Abuse Streamway, Damper Cave, G, Persephone Pot, Bauhaus; Honey and Cream Streamway, Damper Cave; H, Nanocochlea exigua, C.201475; I, J, Nanocochlea stylesae, C.201476, paratypes, Malani Creek; K–M, Nanocochlea damperensis, K, L, C.203685, paratypes, first crossing, Damper Creek, M, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; N, O, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. A, B, E, F, H, K–N, protoconch; C, D, G, I, J, O, microsculpture. Scales: A, C–E, J, K, M, O, 200 m; B, F–I, L, N, 100 m.	FIGURE 6. Protoconchs and teleoconch microsculpture of Pseudotricula and Nanocochlea species. A–D, Pseudotricula expandolabra, A, B, D, C.201493, paratype, main streamway, Bauhaus, C, C.201495, Persephone Streamway, Bauhaus; E–G, Pseudotricula conica, E, F, C.201464, Cane Toad Abuse Streamway, Damper Cave, G, Persephone Pot, Bauhaus; Honey and Cream Streamway, Damper Cave; H, Nanocochlea exigua, C.201475; I, J, Nanocochlea stylesae, C.201476, paratypes, Malani Creek; K–M, Nanocochlea damperensis, K, L, C.203685, paratypes, first crossing, Damper Creek, M, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; N, O, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. A, B, E, F, H, K–N, protoconch; C, D, G, I, J, O, microsculpture. Scales: A, C–E, J, K, M, O, 200 m; B, F–I, L, N, 100 m.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D2BA220E41FF9887F43FCBD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050803/files/figure.png	https://doi.org/10.5281/zenodo.5050803	FIGURE 8. Pallial cavities of male specimens of Pseudotricula and Nanocochlea species. A, Pseudotricula expandolabra, C.201493, paratype, Persephone Streamway, Bauhaus; B, Pseudotricula arthurclarkei, C.203671, paratype, Quetzalcoatl Conduit; C, Pseudotricula auriforma, C.201496, paratype, main streamway, Damper Cave; D, E, Pseudotricula conica, D, C.203680, main streamway near entrance, Damper Cave, E, Cane Toad Abuse Streamway, Damper Cave; F, Nanocochlea exigua, C.203677, paratype, Cane Toad Abuse Streamway, Damper Cave; G, Nanocochlea damperensis, C.203685, paratype, first crossing, Damper Creek; H, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. Scales: A–D, F–H, 500 m, E, 1 mm. Abbreviations: a, anus; ct, ctenidium; ev, efferent vein; os, osphradium; pc, pericardium; pe, pallial edge; ppw, posterior pallial wall; r, rectum; rg, renal gland. The stippled area represents the hypobranchial gland.	FIGURE 8. Pallial cavities of male specimens of Pseudotricula and Nanocochlea species. A, Pseudotricula expandolabra, C.201493, paratype, Persephone Streamway, Bauhaus; B, Pseudotricula arthurclarkei, C.203671, paratype, Quetzalcoatl Conduit; C, Pseudotricula auriforma, C.201496, paratype, main streamway, Damper Cave; D, E, Pseudotricula conica, D, C.203680, main streamway near entrance, Damper Cave, E, Cane Toad Abuse Streamway, Damper Cave; F, Nanocochlea exigua, C.203677, paratype, Cane Toad Abuse Streamway, Damper Cave; G, Nanocochlea damperensis, C.203685, paratype, first crossing, Damper Creek; H, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. Scales: A–D, F–H, 500 m, E, 1 mm. Abbreviations: a, anus; ct, ctenidium; ev, efferent vein; os, osphradium; pc, pericardium; pe, pallial edge; ppw, posterior pallial wall; r, rectum; rg, renal gland. The stippled area represents the hypobranchial gland.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D2BA220E41FF9887F43FCBD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050821/files/figure.png	https://doi.org/10.5281/zenodo.5050821	FIGURE 14. Shells of Nanocochlea species and Pseudotricula progenitor. A ­ C, Nanocochlea exigua, A, holotype, C.439393, Cane Toad Abuse Streamway, Damper Cave; B, C, C.201465, main streamway, Damper Cave; D, Nanocochlea stylesae, holotype, C.439398, Malani Creek; E–G, Nanocochlea damperensis, E, C.203685, paratype, first crossing, Damper Creek; F, holotype, C.439399, first crossing, Damper Creek, F, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; H, I, Pseudotricula progenitor, H, paratype, C.439397, Persephone Streamway, Bauhaus; I, holotype, C.439396, Persephone Streamway, Bauhaus. Scales: 500 m.	FIGURE 14. Shells of Nanocochlea species and Pseudotricula progenitor. A ­ C, Nanocochlea exigua, A, holotype, C.439393, Cane Toad Abuse Streamway, Damper Cave; B, C, C.201465, main streamway, Damper Cave; D, Nanocochlea stylesae, holotype, C.439398, Malani Creek; E–G, Nanocochlea damperensis, E, C.203685, paratype, first crossing, Damper Creek; F, holotype, C.439399, first crossing, Damper Creek, F, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; H, I, Pseudotricula progenitor, H, paratype, C.439397, Persephone Streamway, Bauhaus; I, holotype, C.439396, Persephone Streamway, Bauhaus. Scales: 500 m.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D2BA220E41FF9887F43FCBD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050823/files/figure.png	https://doi.org/10.5281/zenodo.5050823	FIGURE 15. Shells of Nanocochlea species and Pseudotricula progenitor. A ­ C, Nanocochlea exigua, A, C.203674, outside Damper Cave, Damper Creek B, C.166855, Persephone Pot, Bauhaus, C, C.201465, main streamway, Damper Cave; D–F, Nanocochlea stylesae, D, C.201476, paratype, Malani Creek, E, F, C.203686, first crossing, Damper Creek; G­I, Nanocochlea damperensis, G,H, C.303685, seepages at Bauhaus entrance, I, C.203685, paratype, first crossing, Damper Creek; J–L, Pseudotricula progenitor, J, L, C.201463, paratype, Persephone streamway, Bauhaus, K, C.201454, Persephone, Bauhaus. Scales: 1 mm.	FIGURE 15. Shells of Nanocochlea species and Pseudotricula progenitor. A ­ C, Nanocochlea exigua, A, C.203674, outside Damper Cave, Damper Creek B, C.166855, Persephone Pot, Bauhaus, C, C.201465, main streamway, Damper Cave; D–F, Nanocochlea stylesae, D, C.201476, paratype, Malani Creek, E, F, C.203686, first crossing, Damper Creek; G­I, Nanocochlea damperensis, G,H, C.303685, seepages at Bauhaus entrance, I, C.203685, paratype, first crossing, Damper Creek; J–L, Pseudotricula progenitor, J, L, C.201463, paratype, Persephone streamway, Bauhaus, K, C.201454, Persephone, Bauhaus. Scales: 1 mm.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D2BA220E41FF9887F43FCBD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5052059/files/figure.png	https://doi.org/10.5281/zenodo.5052059	FIGURE 16. Opercula of Nanocochlea species and Pseudotricula progenitor. A ­ C, Nanocochlea exigua, C.203677, paratypes, Cane Toad Abuse Streamway, Damper Cave; D ­ F, Nanocochlea damperensis; C–E, C.203685, paratypes, first crossing, Damper Creek; F, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; G, H, Pseudotricula progenitor, C.201821, main streamway, Bauhaus; A, D, G, outer side; B, C, E, F, H, inner side; Scales: A, B, D, 200 m, C, E–H, 100 m.	FIGURE 16. Opercula of Nanocochlea species and Pseudotricula progenitor. A ­ C, Nanocochlea exigua, C.203677, paratypes, Cane Toad Abuse Streamway, Damper Cave; D ­ F, Nanocochlea damperensis; C–E, C.203685, paratypes, first crossing, Damper Creek; F, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; G, H, Pseudotricula progenitor, C.201821, main streamway, Bauhaus; A, D, G, outer side; B, C, E, F, H, inner side; Scales: A, B, D, 200 m, C, E–H, 100 m.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D2BA220E41FF9887F43FCBD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050829/files/figure.png	https://doi.org/10.5281/zenodo.5050829	FIGURE 17. Radulae of Nanocochlea species Pseudotricula progenitor. A ­ C, Nanocochlea exigua, A, C.201475, Honey and Cream Streamway, Damper Cave, B, D, C.201465, main streamway, Damper Cave; D– E, Nanocochlea stylesae, C.201476, paratypes, Malani Creek; G–K, Nanocochlea damperensis, G, H, C.203685, paratypes, first crossing, Damper Creek, I–K, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; L–O, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. A, E, central and lateral teeth; B, F, I, M, central, lateral, inner and outer marginal teeth; C, D, J, N central teeth; G, L, lateral teeth; H, central lateral inner and outer marginal teeth; K, outer marginal teeth. Scales: A, G–H, K, L, 10 m; B, I, M, 20 m; C, D, J, N, 5 m.	FIGURE 17. Radulae of Nanocochlea species Pseudotricula progenitor. A ­ C, Nanocochlea exigua, A, C.201475, Honey and Cream Streamway, Damper Cave, B, D, C.201465, main streamway, Damper Cave; D– E, Nanocochlea stylesae, C.201476, paratypes, Malani Creek; G–K, Nanocochlea damperensis, G, H, C.203685, paratypes, first crossing, Damper Creek, I–K, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; L–O, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. A, E, central and lateral teeth; B, F, I, M, central, lateral, inner and outer marginal teeth; C, D, J, N central teeth; G, L, lateral teeth; H, central lateral inner and outer marginal teeth; K, outer marginal teeth. Scales: A, G–H, K, L, 10 m; B, I, M, 20 m; C, D, J, N, 5 m.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D2BA220E41FF9887F43FCBD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050831/files/figure.png	https://doi.org/10.5281/zenodo.5050831	FIGURE 18. Female reproductive system of Nanocochlea species and Pseudotricula progenitor. A, Nanocochlea exigua, C.203677, paratype, Cane Toad Abuse Streamway, Damper Cave; B, Nanocochlea damperensis, C.203685, paratype, first crossing, Damper Creek; C, D, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. Scales: 500 m. Abbreviations: ag, albumen gland; bc, bursa copulatrix; bd, bursal duct; cg, capsule gland; co, coiled oviduct; po, genital opening; pw, pallial wall; sr, seminal receptacle; vc, ventral channel.	FIGURE 18. Female reproductive system of Nanocochlea species and Pseudotricula progenitor. A, Nanocochlea exigua, C.203677, paratype, Cane Toad Abuse Streamway, Damper Cave; B, Nanocochlea damperensis, C.203685, paratype, first crossing, Damper Creek; C, D, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. Scales: 500 m. Abbreviations: ag, albumen gland; bc, bursa copulatrix; bd, bursal duct; cg, capsule gland; co, coiled oviduct; po, genital opening; pw, pallial wall; sr, seminal receptacle; vc, ventral channel.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D2EA222E41FFC507D25FDCD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050791/files/figure.png	https://doi.org/10.5281/zenodo.5050791	FIGURE 4. Dorsal views of penes of Pseudotricula and Nanocochlea species. A,. Pseudotricula eberhardi; B, Pseudotricula expandolabra; C, Nanocochlea stylesae; D, Nanocochlea damperensis; E, Pseudotricula conica. Scales A, B, E, 0.2 mm; C, D, 0.1 mm. Abbreviations: ds, distal swelling; pd, penial duct.	FIGURE 4. Dorsal views of penes of Pseudotricula and Nanocochlea species. A,. Pseudotricula eberhardi; B, Pseudotricula expandolabra; C, Nanocochlea stylesae; D, Nanocochlea damperensis; E, Pseudotricula conica. Scales A, B, E, 0.2 mm; C, D, 0.1 mm. Abbreviations: ds, distal swelling; pd, penial duct.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D2EA222E41FFC507D25FDCD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050797/files/figure.png	https://doi.org/10.5281/zenodo.5050797	FIGURE 6. Protoconchs and teleoconch microsculpture of Pseudotricula and Nanocochlea species. A–D, Pseudotricula expandolabra, A, B, D, C.201493, paratype, main streamway, Bauhaus, C, C.201495, Persephone Streamway, Bauhaus; E–G, Pseudotricula conica, E, F, C.201464, Cane Toad Abuse Streamway, Damper Cave, G, Persephone Pot, Bauhaus; Honey and Cream Streamway, Damper Cave; H, Nanocochlea exigua, C.201475; I, J, Nanocochlea stylesae, C.201476, paratypes, Malani Creek; K–M, Nanocochlea damperensis, K, L, C.203685, paratypes, first crossing, Damper Creek, M, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; N, O, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. A, B, E, F, H, K–N, protoconch; C, D, G, I, J, O, microsculpture. Scales: A, C–E, J, K, M, O, 200 m; B, F–I, L, N, 100 m.	FIGURE 6. Protoconchs and teleoconch microsculpture of Pseudotricula and Nanocochlea species. A–D, Pseudotricula expandolabra, A, B, D, C.201493, paratype, main streamway, Bauhaus, C, C.201495, Persephone Streamway, Bauhaus; E–G, Pseudotricula conica, E, F, C.201464, Cane Toad Abuse Streamway, Damper Cave, G, Persephone Pot, Bauhaus; Honey and Cream Streamway, Damper Cave; H, Nanocochlea exigua, C.201475; I, J, Nanocochlea stylesae, C.201476, paratypes, Malani Creek; K–M, Nanocochlea damperensis, K, L, C.203685, paratypes, first crossing, Damper Creek, M, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; N, O, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. A, B, E, F, H, K–N, protoconch; C, D, G, I, J, O, microsculpture. Scales: A, C–E, J, K, M, O, 200 m; B, F–I, L, N, 100 m.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D2EA222E41FFC507D25FDCD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050821/files/figure.png	https://doi.org/10.5281/zenodo.5050821	FIGURE 14. Shells of Nanocochlea species and Pseudotricula progenitor. A ­ C, Nanocochlea exigua, A, holotype, C.439393, Cane Toad Abuse Streamway, Damper Cave; B, C, C.201465, main streamway, Damper Cave; D, Nanocochlea stylesae, holotype, C.439398, Malani Creek; E–G, Nanocochlea damperensis, E, C.203685, paratype, first crossing, Damper Creek; F, holotype, C.439399, first crossing, Damper Creek, F, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; H, I, Pseudotricula progenitor, H, paratype, C.439397, Persephone Streamway, Bauhaus; I, holotype, C.439396, Persephone Streamway, Bauhaus. Scales: 500 m.	FIGURE 14. Shells of Nanocochlea species and Pseudotricula progenitor. A ­ C, Nanocochlea exigua, A, holotype, C.439393, Cane Toad Abuse Streamway, Damper Cave; B, C, C.201465, main streamway, Damper Cave; D, Nanocochlea stylesae, holotype, C.439398, Malani Creek; E–G, Nanocochlea damperensis, E, C.203685, paratype, first crossing, Damper Creek; F, holotype, C.439399, first crossing, Damper Creek, F, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; H, I, Pseudotricula progenitor, H, paratype, C.439397, Persephone Streamway, Bauhaus; I, holotype, C.439396, Persephone Streamway, Bauhaus. Scales: 500 m.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D2EA222E41FFC507D25FDCD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050823/files/figure.png	https://doi.org/10.5281/zenodo.5050823	FIGURE 15. Shells of Nanocochlea species and Pseudotricula progenitor. A ­ C, Nanocochlea exigua, A, C.203674, outside Damper Cave, Damper Creek B, C.166855, Persephone Pot, Bauhaus, C, C.201465, main streamway, Damper Cave; D–F, Nanocochlea stylesae, D, C.201476, paratype, Malani Creek, E, F, C.203686, first crossing, Damper Creek; G­I, Nanocochlea damperensis, G,H, C.303685, seepages at Bauhaus entrance, I, C.203685, paratype, first crossing, Damper Creek; J–L, Pseudotricula progenitor, J, L, C.201463, paratype, Persephone streamway, Bauhaus, K, C.201454, Persephone, Bauhaus. Scales: 1 mm.	FIGURE 15. Shells of Nanocochlea species and Pseudotricula progenitor. A ­ C, Nanocochlea exigua, A, C.203674, outside Damper Cave, Damper Creek B, C.166855, Persephone Pot, Bauhaus, C, C.201465, main streamway, Damper Cave; D–F, Nanocochlea stylesae, D, C.201476, paratype, Malani Creek, E, F, C.203686, first crossing, Damper Creek; G­I, Nanocochlea damperensis, G,H, C.303685, seepages at Bauhaus entrance, I, C.203685, paratype, first crossing, Damper Creek; J–L, Pseudotricula progenitor, J, L, C.201463, paratype, Persephone streamway, Bauhaus, K, C.201454, Persephone, Bauhaus. Scales: 1 mm.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D2EA222E41FFC507D25FDCD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050829/files/figure.png	https://doi.org/10.5281/zenodo.5050829	FIGURE 17. Radulae of Nanocochlea species Pseudotricula progenitor. A ­ C, Nanocochlea exigua, A, C.201475, Honey and Cream Streamway, Damper Cave, B, D, C.201465, main streamway, Damper Cave; D– E, Nanocochlea stylesae, C.201476, paratypes, Malani Creek; G–K, Nanocochlea damperensis, G, H, C.203685, paratypes, first crossing, Damper Creek, I–K, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; L–O, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. A, E, central and lateral teeth; B, F, I, M, central, lateral, inner and outer marginal teeth; C, D, J, N central teeth; G, L, lateral teeth; H, central lateral inner and outer marginal teeth; K, outer marginal teeth. Scales: A, G–H, K, L, 10 m; B, I, M, 20 m; C, D, J, N, 5 m.	FIGURE 17. Radulae of Nanocochlea species Pseudotricula progenitor. A ­ C, Nanocochlea exigua, A, C.201475, Honey and Cream Streamway, Damper Cave, B, D, C.201465, main streamway, Damper Cave; D– E, Nanocochlea stylesae, C.201476, paratypes, Malani Creek; G–K, Nanocochlea damperensis, G, H, C.203685, paratypes, first crossing, Damper Creek, I–K, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; L–O, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. A, E, central and lateral teeth; B, F, I, M, central, lateral, inner and outer marginal teeth; C, D, J, N central teeth; G, L, lateral teeth; H, central lateral inner and outer marginal teeth; K, outer marginal teeth. Scales: A, G–H, K, L, 10 m; B, I, M, 20 m; C, D, J, N, 5 m.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D2CA23CE41FFD607A42F905.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050791/files/figure.png	https://doi.org/10.5281/zenodo.5050791	FIGURE 4. Dorsal views of penes of Pseudotricula and Nanocochlea species. A,. Pseudotricula eberhardi; B, Pseudotricula expandolabra; C, Nanocochlea stylesae; D, Nanocochlea damperensis; E, Pseudotricula conica. Scales A, B, E, 0.2 mm; C, D, 0.1 mm. Abbreviations: ds, distal swelling; pd, penial duct.	FIGURE 4. Dorsal views of penes of Pseudotricula and Nanocochlea species. A,. Pseudotricula eberhardi; B, Pseudotricula expandolabra; C, Nanocochlea stylesae; D, Nanocochlea damperensis; E, Pseudotricula conica. Scales A, B, E, 0.2 mm; C, D, 0.1 mm. Abbreviations: ds, distal swelling; pd, penial duct.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D2CA23CE41FFD607A42F905.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050797/files/figure.png	https://doi.org/10.5281/zenodo.5050797	FIGURE 6. Protoconchs and teleoconch microsculpture of Pseudotricula and Nanocochlea species. A–D, Pseudotricula expandolabra, A, B, D, C.201493, paratype, main streamway, Bauhaus, C, C.201495, Persephone Streamway, Bauhaus; E–G, Pseudotricula conica, E, F, C.201464, Cane Toad Abuse Streamway, Damper Cave, G, Persephone Pot, Bauhaus; Honey and Cream Streamway, Damper Cave; H, Nanocochlea exigua, C.201475; I, J, Nanocochlea stylesae, C.201476, paratypes, Malani Creek; K–M, Nanocochlea damperensis, K, L, C.203685, paratypes, first crossing, Damper Creek, M, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; N, O, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. A, B, E, F, H, K–N, protoconch; C, D, G, I, J, O, microsculpture. Scales: A, C–E, J, K, M, O, 200 m; B, F–I, L, N, 100 m.	FIGURE 6. Protoconchs and teleoconch microsculpture of Pseudotricula and Nanocochlea species. A–D, Pseudotricula expandolabra, A, B, D, C.201493, paratype, main streamway, Bauhaus, C, C.201495, Persephone Streamway, Bauhaus; E–G, Pseudotricula conica, E, F, C.201464, Cane Toad Abuse Streamway, Damper Cave, G, Persephone Pot, Bauhaus; Honey and Cream Streamway, Damper Cave; H, Nanocochlea exigua, C.201475; I, J, Nanocochlea stylesae, C.201476, paratypes, Malani Creek; K–M, Nanocochlea damperensis, K, L, C.203685, paratypes, first crossing, Damper Creek, M, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; N, O, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. A, B, E, F, H, K–N, protoconch; C, D, G, I, J, O, microsculpture. Scales: A, C–E, J, K, M, O, 200 m; B, F–I, L, N, 100 m.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D2CA23CE41FFD607A42F905.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050803/files/figure.png	https://doi.org/10.5281/zenodo.5050803	FIGURE 8. Pallial cavities of male specimens of Pseudotricula and Nanocochlea species. A, Pseudotricula expandolabra, C.201493, paratype, Persephone Streamway, Bauhaus; B, Pseudotricula arthurclarkei, C.203671, paratype, Quetzalcoatl Conduit; C, Pseudotricula auriforma, C.201496, paratype, main streamway, Damper Cave; D, E, Pseudotricula conica, D, C.203680, main streamway near entrance, Damper Cave, E, Cane Toad Abuse Streamway, Damper Cave; F, Nanocochlea exigua, C.203677, paratype, Cane Toad Abuse Streamway, Damper Cave; G, Nanocochlea damperensis, C.203685, paratype, first crossing, Damper Creek; H, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. Scales: A–D, F–H, 500 m, E, 1 mm. Abbreviations: a, anus; ct, ctenidium; ev, efferent vein; os, osphradium; pc, pericardium; pe, pallial edge; ppw, posterior pallial wall; r, rectum; rg, renal gland. The stippled area represents the hypobranchial gland.	FIGURE 8. Pallial cavities of male specimens of Pseudotricula and Nanocochlea species. A, Pseudotricula expandolabra, C.201493, paratype, Persephone Streamway, Bauhaus; B, Pseudotricula arthurclarkei, C.203671, paratype, Quetzalcoatl Conduit; C, Pseudotricula auriforma, C.201496, paratype, main streamway, Damper Cave; D, E, Pseudotricula conica, D, C.203680, main streamway near entrance, Damper Cave, E, Cane Toad Abuse Streamway, Damper Cave; F, Nanocochlea exigua, C.203677, paratype, Cane Toad Abuse Streamway, Damper Cave; G, Nanocochlea damperensis, C.203685, paratype, first crossing, Damper Creek; H, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. Scales: A–D, F–H, 500 m, E, 1 mm. Abbreviations: a, anus; ct, ctenidium; ev, efferent vein; os, osphradium; pc, pericardium; pe, pallial edge; ppw, posterior pallial wall; r, rectum; rg, renal gland. The stippled area represents the hypobranchial gland.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D2CA23CE41FFD607A42F905.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050821/files/figure.png	https://doi.org/10.5281/zenodo.5050821	FIGURE 14. Shells of Nanocochlea species and Pseudotricula progenitor. A ­ C, Nanocochlea exigua, A, holotype, C.439393, Cane Toad Abuse Streamway, Damper Cave; B, C, C.201465, main streamway, Damper Cave; D, Nanocochlea stylesae, holotype, C.439398, Malani Creek; E–G, Nanocochlea damperensis, E, C.203685, paratype, first crossing, Damper Creek; F, holotype, C.439399, first crossing, Damper Creek, F, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; H, I, Pseudotricula progenitor, H, paratype, C.439397, Persephone Streamway, Bauhaus; I, holotype, C.439396, Persephone Streamway, Bauhaus. Scales: 500 m.	FIGURE 14. Shells of Nanocochlea species and Pseudotricula progenitor. A ­ C, Nanocochlea exigua, A, holotype, C.439393, Cane Toad Abuse Streamway, Damper Cave; B, C, C.201465, main streamway, Damper Cave; D, Nanocochlea stylesae, holotype, C.439398, Malani Creek; E–G, Nanocochlea damperensis, E, C.203685, paratype, first crossing, Damper Creek; F, holotype, C.439399, first crossing, Damper Creek, F, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; H, I, Pseudotricula progenitor, H, paratype, C.439397, Persephone Streamway, Bauhaus; I, holotype, C.439396, Persephone Streamway, Bauhaus. Scales: 500 m.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D2CA23CE41FFD607A42F905.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050823/files/figure.png	https://doi.org/10.5281/zenodo.5050823	FIGURE 15. Shells of Nanocochlea species and Pseudotricula progenitor. A ­ C, Nanocochlea exigua, A, C.203674, outside Damper Cave, Damper Creek B, C.166855, Persephone Pot, Bauhaus, C, C.201465, main streamway, Damper Cave; D–F, Nanocochlea stylesae, D, C.201476, paratype, Malani Creek, E, F, C.203686, first crossing, Damper Creek; G­I, Nanocochlea damperensis, G,H, C.303685, seepages at Bauhaus entrance, I, C.203685, paratype, first crossing, Damper Creek; J–L, Pseudotricula progenitor, J, L, C.201463, paratype, Persephone streamway, Bauhaus, K, C.201454, Persephone, Bauhaus. Scales: 1 mm.	FIGURE 15. Shells of Nanocochlea species and Pseudotricula progenitor. A ­ C, Nanocochlea exigua, A, C.203674, outside Damper Cave, Damper Creek B, C.166855, Persephone Pot, Bauhaus, C, C.201465, main streamway, Damper Cave; D–F, Nanocochlea stylesae, D, C.201476, paratype, Malani Creek, E, F, C.203686, first crossing, Damper Creek; G­I, Nanocochlea damperensis, G,H, C.303685, seepages at Bauhaus entrance, I, C.203685, paratype, first crossing, Damper Creek; J–L, Pseudotricula progenitor, J, L, C.201463, paratype, Persephone streamway, Bauhaus, K, C.201454, Persephone, Bauhaus. Scales: 1 mm.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D2CA23CE41FFD607A42F905.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5052059/files/figure.png	https://doi.org/10.5281/zenodo.5052059	FIGURE 16. Opercula of Nanocochlea species and Pseudotricula progenitor. A ­ C, Nanocochlea exigua, C.203677, paratypes, Cane Toad Abuse Streamway, Damper Cave; D ­ F, Nanocochlea damperensis; C–E, C.203685, paratypes, first crossing, Damper Creek; F, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; G, H, Pseudotricula progenitor, C.201821, main streamway, Bauhaus; A, D, G, outer side; B, C, E, F, H, inner side; Scales: A, B, D, 200 m, C, E–H, 100 m.	FIGURE 16. Opercula of Nanocochlea species and Pseudotricula progenitor. A ­ C, Nanocochlea exigua, C.203677, paratypes, Cane Toad Abuse Streamway, Damper Cave; D ­ F, Nanocochlea damperensis; C–E, C.203685, paratypes, first crossing, Damper Creek; F, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; G, H, Pseudotricula progenitor, C.201821, main streamway, Bauhaus; A, D, G, outer side; B, C, E, F, H, inner side; Scales: A, B, D, 200 m, C, E–H, 100 m.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D2CA23CE41FFD607A42F905.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050829/files/figure.png	https://doi.org/10.5281/zenodo.5050829	FIGURE 17. Radulae of Nanocochlea species Pseudotricula progenitor. A ­ C, Nanocochlea exigua, A, C.201475, Honey and Cream Streamway, Damper Cave, B, D, C.201465, main streamway, Damper Cave; D– E, Nanocochlea stylesae, C.201476, paratypes, Malani Creek; G–K, Nanocochlea damperensis, G, H, C.203685, paratypes, first crossing, Damper Creek, I–K, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; L–O, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. A, E, central and lateral teeth; B, F, I, M, central, lateral, inner and outer marginal teeth; C, D, J, N central teeth; G, L, lateral teeth; H, central lateral inner and outer marginal teeth; K, outer marginal teeth. Scales: A, G–H, K, L, 10 m; B, I, M, 20 m; C, D, J, N, 5 m.	FIGURE 17. Radulae of Nanocochlea species Pseudotricula progenitor. A ­ C, Nanocochlea exigua, A, C.201475, Honey and Cream Streamway, Damper Cave, B, D, C.201465, main streamway, Damper Cave; D– E, Nanocochlea stylesae, C.201476, paratypes, Malani Creek; G–K, Nanocochlea damperensis, G, H, C.203685, paratypes, first crossing, Damper Creek, I–K, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; L–O, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. A, E, central and lateral teeth; B, F, I, M, central, lateral, inner and outer marginal teeth; C, D, J, N central teeth; G, L, lateral teeth; H, central lateral inner and outer marginal teeth; K, outer marginal teeth. Scales: A, G–H, K, L, 10 m; B, I, M, 20 m; C, D, J, N, 5 m.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
03AB87BA2D2CA23CE41FFD607A42F905.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5050831/files/figure.png	https://doi.org/10.5281/zenodo.5050831	FIGURE 18. Female reproductive system of Nanocochlea species and Pseudotricula progenitor. A, Nanocochlea exigua, C.203677, paratype, Cane Toad Abuse Streamway, Damper Cave; B, Nanocochlea damperensis, C.203685, paratype, first crossing, Damper Creek; C, D, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. Scales: 500 m. Abbreviations: ag, albumen gland; bc, bursa copulatrix; bd, bursal duct; cg, capsule gland; co, coiled oviduct; po, genital opening; pw, pallial wall; sr, seminal receptacle; vc, ventral channel.	FIGURE 18. Female reproductive system of Nanocochlea species and Pseudotricula progenitor. A, Nanocochlea exigua, C.203677, paratype, Cane Toad Abuse Streamway, Damper Cave; B, Nanocochlea damperensis, C.203685, paratype, first crossing, Damper Creek; C, D, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. Scales: 500 m. Abbreviations: ag, albumen gland; bc, bursa copulatrix; bd, bursal duct; cg, capsule gland; co, coiled oviduct; po, genital opening; pw, pallial wall; sr, seminal receptacle; vc, ventral channel.	2005-11-01	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.		Zenodo	biologists	Ponder, W. F.;Clark, S. A.;Eberhard, S.;Studdert, J. B.			
