identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03AB9F22FF83FFF61D5AFEF1FB5AFBE7.text	03AB9F22FF83FFF61D5AFEF1FB5AFBE7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Periclimenaeus karantina Park & De Grave 2021	<div><p>Periclimenaeus karantina sp. nov. Park and De</p><p>Grave (Figs. 2–9) urn:lsid:zoobank.org:act: 5157B8FA-51BE-476E-8B98- 100D6ABDF592</p><p>Material examined: Holotype. 1 male (pocl 3.3); Oct. 22, 2019; Munseom Islet, Jejudo <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=126.570274&amp;materialsCitation.latitude=33.225002" title="Search Plazi for locations around (long 126.570274/lat 33.225002)">Island</a> (33°13'30"N 126°34'13"E), 21 m, leg. JH Park (NIBRIV0000862971) . Paratypes. 1 female (pocl 3.5); Aug. 08, 2016; same location, 20 m, leg. JH Park (NIBRIV 0000862966, transferred from SNU KR_JH 474); 1 ovig. female, 1 male (pocl 4.1, 3.5); Aug. 16, 2019; Seopseom Islet, Jejudo Island (33°13'44.37"N 126°35'43.74"E), 38 m, leg. JH Park (NIBRIV 0000862967-68); 1 female, 1 male (pocl 3.8, 3.5); Aug. 16, 2019; same location, leg. JH Park (OUMNH. ZC.2018 -03-027, OUMNH. ZC.2018 - 03-028); 1 ovig. female, 1 female, 2 males (pocl 4.5, 3.2, 3.4, 2.8); Oct. 21, 2019; same location, depth, leg. JH Park (NIBRIV 0000862972-862974, NIBRIV 0000877272); 1 juv., 1 ovig. female, 1 male (pocl 1.9, 3.4, 3.1); same data as holotype, leg. JH Park (NIBRIV 0000862969-862970, NIBRIV 0000877273); 1 ovig. female, 1 female, 2 males (pocl 4.6, 3.5, 3.8, 3.3); Jan. 13, 2020; Munseom Islet Jejudo Island, 32m, leg. JH Park (NIBRIV 0000877274-877277); 1 ovig. female, 1 female, 2 males (pocl 4.1, 3.4, 3.7, 2.9); Jan. 14, 2020; same location, 24 m, leg. JH Park (NIBRIV 0000877278-877281); 1 ovig. female, 1 male (pocl 4.4, 3.5); Jan. 14, 2020; same location, 24 m, leg. JH Park (NIBRIV 0000877282-877283); 1 male (pocl 3.1); Jan. 15, 2020; same location, 21 m, leg. JH Park (JH 1108). All collected from a colonial didemnid ascidian Leptoclinides sp. ( Ascidiacea: Aplousobranchia: Didemnidae) (Fig. 9). GenBank accession numbers for DNA sequences data are presented as table 1.</p><p>© 2021 Academia Sinica, Taiwan</p><p>Description: Body medium-sized, subcylindrical form (Fig. 2).</p><p>Rostrum (Figs. 2, 3 A) straight, trending</p><p>Used symbols: N/A – not available; * Leptoclinides sp. – tissue sample of the host colonial ascidian of Periclimenaeus karantina sp. nov. (NIBRIV0000862967).</p><p>© 2021 Academia Sinica, Taiwan</p><p>downwards, about 0.4 of pocl, reaching slightly beyond distal margin of basal segment of antennular peduncle, with 4 equally spaced dorsal teeth, without ventral tooth.</p><p>Carapace (Fig. 2) smooth, glabrous, without supraorbital tooth or tubercles, with feeble supraorbital ridges; antennal tooth acute; inferior orbital angle with round blunt process (Fig. 3C); pterygostomial angle produced.</p><p>Abdomen (Fig. 2) with smooth pleon, first segment without anteromedian dorsal lobe; pleura broadly rounded, sixth segment about 1.1 times length of fifth, about 0.5 of telson length, posterolateral angle rounded, posteroventral angle acute (Fig. 3D, E).</p><p>Telson (Fig. 2) about 0.7 of pocl, about 2 times longer than maximal width (Fig. 3E); two pairs of dorsal spiniform setae, subequal in size, at about 0.3 and 0.8 of telson length respectively, posterior margin with three pairs of spiniform setae, lateral posterior spiniform setae short, about 0.5 of length of intermediate pair, intermediate pair long and stout, submedian pair about 0.8 of intermediate pair length, distally setulose (Fig. 3F).</p><p>Eye (Fig. 4A, B) with hemispherical cornea, about 1.5 times longer than maximum dorsal width, about 1.2 times longer than maximum lateral width, nebenauge absent.</p><p>Antennule (Fig. 4C) with proximal segment of peduncle bearing acute distolateral tooth, with acute tooth at ventromedial margin (Fig. 4D); stylocerite broad, bearing sharp point, reaching to about 0.5 of proximal segment; intermediate segment short, about 0.3 times of proximal segment length, subequal to distal segment length; upper flagellum biramous, proximal five segments fused, short free ramus with two segments, longer free ramus with seven segment; lower flagellum with fourteen segments, filiform.</p><p>Antenna (Fig. 4E) with rounded boss proximally on coxa; basicerite without distoventral tooth, ischiocerite and merocerite unarmed; carpocerite exceeding scaphocerite; scaphocerite about 2.4 times as long as maximal width, distal lamella rounded, exceeding acute distolateral tooth situated at about 0.8 of scaphocerite length.</p><p>Mouthparts not dissected. Second maxilliped with normal endopod, exopod, oval epipod without podobranch. Third maxilliped (Fig. 5A) with ultimate segment about 0.5 times as long as antepenultimate segment, tapering distally, with dense tufts of long setae; penultimate segment about 0.6 times antepenultimate segment length, with ventromedial row of long setae; antepenultimate segment with long setae on ventromedial margin; exopod reaching middle of penultimate segment, distally with six plumose setae; coxa with rounded lateral plate, without arthrobranch.</p><p>First pereiopod (Figs. 2, 6A) with coxa and basis without special features; ischium about 0.6 of merus length, unarmed; merus subequal to carpus length, unarmed; carpus about 1.2 times as long as chela, tapering proximally; carpo-propodal cleaning brush developed; chela about 0.4 times as long as pocl, about 0.7 times as long as merus length; palm subcylindrical, smooth, non-tuberculate; fingers about 0.9 of palm length, subspatulate, with subterminal group of setae (Fig. 6B), with three terminal teeth, pair of subterminal teeth short, about 0.6 of median tooth length (Fig. 6C).</p><p>Second pereiopods (Figs. 2, 7) robust, dissimilar in shape, unequal in size.</p><p>Major second pereiopod (Fig. 2) with coxa and basis without special features; ischium about 0.8 of merus length, tapering proximally, unarmed; merus about 0.3 of palm length, with minute tubercles on ventral margin (Fig. 8E, F); carpus about 0.3 of palm length, tapering proximally, unarmed; chela about 1.7 to 2.6 times as long as pocl, about 4.3 times as long as merus length (Fig. 7A); palm subcylindrical, smooth, non-tuberculate; fingers (Fig. 7B) about 0.4 of palm length, distally curved mesially (Fig. 7C), with subterminal group of setae; fixed finger with strong subacute tip, distal cutting edge concave, entire, proximal cutting edge with deep oval fossa, mesial margin with triangular process (Fig. 7B), lateral margin with lower rounded process (Fig. 7A); dactylus with strong subacute tip, distal cutting edge with about 50 small acute teeth (Fig. 7D), proximal cutting edge with large molar process.</p><p>Minor second pereiopod (Fig. 2) with coxa and basis without special features; ischium subequal to merus length, tapering proximally, unarmed; merus about 0.4 of palm length, with minute tubercles on ventral margin (Figs. 7E, 8D); chela about 1.2 times as long as pocl, about 0.5 of major chela length, about 3 times as long as merus, with fingers unequal in size; palm subcylindrical, smooth, non-tuberculate; carpus about 0.3 of palm length, tapering proximally, unarmed; fixed finger about 0.7 of dactylus length, with strong subacute tip, cutting edge with long groove extending to 0.9 of fixed finger from apex (Fig. 8A), lateral cutting edge slightly convex, proximal margin with lower rounded process, with denticulate mesial cutting edge (Fig. 8A, C), proximal margin with triangular tuberculate process (Fig. 8A, C); dactylus (Fig. 7F) about one third of palm length, about 2.0 times as long as maximal depth in midlength, exceeding fixed finger, with broadly rounded dorsal margin, tip with blunt tooth continuous with sinuous cutting edge, distally concave, proximally convex, with about 40 small acute teeth, decreasing in size proximally (Fig. 8 A, B), proximal cutting edge with right-angular obtuse process fitting to proximal end of occlusal groove on fixed finger (Fig. 8B).</p><p>© 2021 Academia Sinica, Taiwan</p><p>© 2021 Academia Sinica, Taiwan</p><p>Ambulatory pereiopods (Fig. 2) subequal in shape, third pereiopod strongest, fourth and fifth gradually slightly slender.</p><p>Third pereiopod (Fig. 6D) robust, sparsely setose; coxa and basis without special features; ischium about 0.7 of merus length, unarmed; merus about 1.2 of carpus length, unarmed; carpus about 0.9 of propodus length, tapering proximally, unarmed; propodus about 2.9 times longer than maximal depth, tapering distally, with two or three stout distoventral spiniform setae (Figs. 6E, 8G); dactylus about 0.2 of propodus length, unguis distinctly demarcated, simple, curved, about 0.3 of dorsal corpus length, corpus without distal accessory tooth, slightly convex distally, with minute acute proximal tooth (Figs. 6E, 8G), with sensory setae distolaterally.</p><p>© 2021 Academia Sinica, Taiwan</p><p>Fourth pereiopod (Fig. 6F) robust, sparsely setose; coxa and basis without special features; ischium about 0.8 of merus length, unarmed; merus about 1.2 of carpus length, unarmed; carpus about 0.9 of propodus length, tapering proximally, unarmed; propodus about 3.3 times longer than maximal depth, tapering distally, with pair of stout distoventral spiniform setae (Fig. 6G); dactylus about 0.16 of propodus length, unguis distinctly demarcated, simple, curved, about 0.4 of dorsal corpus length, corpus without distal accessory tooth, slightly convex distally, with minute acute proximal tooth (Fig. 6G), with sensory setae distolaterally.</p><p>Fifth pereiopod (Fig. 6H) robust, sparsely setose; coxa and basis without special features; ischium about 0.7 of merus length, unarmed; merus about 1.4 of carpus length, unarmed; carpus about 0.7 of propodus length, tapering proximally, unarmed; propodus about 4.7 times longer than maximal depth, tapering distally, with single stout distoventral spiniform setae, with distoventral setulose setae, with single spiniform setae distomedially (Fig 6I); dactylus about 0.13 of propodus length, unguis distinctly demarcated, simple, curved, about 0.3 of dorsal corpus length, corpus without distal accessory tooth, slightly convex distally, with minute acute proximal tooth (Fig 6I), with sensory setae distolaterally.</p><p>Second pleopod of male (Fig. 5B) with endopod with short appendix masculina in relation to appendix interna, with two long terminal setulose setae; appendix interna about 3.0 times as long as appendix masculina (Fig. 5C).</p><p>Second pleopod of female with protopod medially with two to four ovigerous setae proximally, two to three distally, proximo-laterally with single seta (Fig. 5D, E).</p><p>Uropod (Fig. 3E) reaching to telson tip; exopod slightly shorter than endopod, outer margin entire, with single spiniform setae, curved inward, about 2.3 times longer than acute distolateral tooth.</p><p>Variation: All intact 21 adult specimens have the same rostral formula (4/0) except for a single ovigerous specimen (pocl 4.6 mm, 3/0). The general morphology of both sexes is very similar, although the major second chela is distinctly larger in the relatively smaller male specimens. For example, the major second chela is about 2.6 times as long as the pocl in the holotype male (pocl 3.3 mm) and about 1.7 times in the largest ovigerous female paratype (pocl 4.6 mm). The dentition of the cutting edge of the fixed finger in the minor second chelae varies considerably (Fig. 8A, C) in the number of small teeth, ranging from none to 30 (holotype).</p><p>© 2021 Academia Sinica, Taiwan</p><p>© 2021 Academia Sinica, Taiwan</p><p>Colouration: Whole body and appendages semitransparent with a pale orange-cream background colour when alive (Fig. 9A); bright white and orange chromatophores scattered all over body and appendages (Fig. 9A, B).</p><p>© 2021 Academia Sinica, Taiwan</p><p>© 2021 Academia Sinica, Taiwan</p><p>Etymology: From the Greek karantina (καραντίνα, quarantine), referring to the lifestyle of the new species within the host ascidian species (Fig. 9B). It also alludes to the quarantine of human society due to the coronavirus pandemic (COVID-19), during which time this paper was written. Used as a noun in apposition.</p><p>© 2021 Academia Sinica, Taiwan</p><p>Host: The shrimp specimens were found inside the common cloacal system of an colonial didemnid ascidian Leptoclinides sp. ( Ascidiacea: Aplousobranchia: Didemnidae) (Fig. 9B, C).</p><p>Distribution: Presently only known from the type locality in Jejudo Island, Korea (Fig. 1A, B).</p><p>Remarks: The species is considered to be typical for ascidian associates due to the presence of a denticulate dactylus on the minor second pereiopod. Periclimenaeus karantina sp. nov. appears morphologically close to seven species: P. colemani Bruce, 2014, P. dactylodon Bruce, 2012a, P. devaneyi Bruce, 2010, P. diplosomatis Bruce, 1980, P. kottae Bruce, 2005a, P. myora Bruce, 1998, P. orbitocarinatus Fransen, 2006; and possibly P. zarenkovi Ďuriš, 1990 . All these species share the following characters: 1) first abdominal tergite without anterodorsal medial lobe; 2) major and minor second chela with denticulate cutting edges on dactylus; and 3) the dactyli of the ambulatory pereiopods having a proximal ventral tooth, but no distoventral accessory tooth.</p><p>The new species shares with P. devaneyi, P. dactylodon, P. kottae and P. orbitocarinatus the presence of median tubercles on the ventral margin on the merus of both second pereiopods. However, P. devaneyi can be readily distinguished from the new species by the presence of a large spine on the distoventral angle of the carpus of the ambulatory pereiopods (vs. absent in P. karantina sp. nov.). Periclimenaeus dactylodon and P. kottae can also readily be distinguished from the new species by the presence of the proximal process on the corpus of the dactylus of the third pereiopod (with large rounded boss in P. kottae or large triangular basal tooth in P. dactylodon vs. with minute proximal tooth in P. karantina sp. nov.). Periclimenaeus orbitocarinatus clearly differs from the new species by the strongly pronounced postorbital ridge along the anterior margin of the carapace and the antennal carpocerite not overreaching the scaphocerite (vs. with feebly pronounced postorbital ridge and carpocerite overreaching the scaphocerite in P. karantina sp. nov.).</p><p>Periclimenaeus karantina sp. nov. can also be distinguished from P. diplosomatis, P. myora and P. zarenkovi, on the basis of the dactylus of the minor second pereiopod not exceeding the fixed finger and the ventral margin of the merus of the second pereiopods not being tuberculate (vs. dactylus exceeding fixed finger and tuberculated ventral margin in P. karantina sp. nov.). Periclimenaeus diplosomatis can be distinguished by the absence of an epipod on the second maxilliped and distolateral tooth of the scaphocerite exceeding anterior margin of the lamella (vs. with epipod and not exceeding in P. karantina sp. nov.). The new species also differs from P. myora and P. zarenkovi in the presence of a long carpocerite which exceeds the anterior margin of the scaphocerite (vs. not exceeding in P. myora and P. zarenkovi).</p><p>Periclimenaeus karantina sp. nov. most closely resembles P. colemani, with which it shares the carpocerite of the antenna exceeding the distal margin of the scaphocerite, and the scaphocerite with the lamella exceeding the distolateral tooth. The new species can be distinguished from P. colemani on the basis of the combination of the following characters: 1) antennule with rounded distolateral tooth (vs. sharp distolateral tooth in P. karantina sp. nov.); 2) dactylus of the minor chela being about 2.8 times greater than the maximal depth (vs. about 2 times in P. karantina sp. nov.); 3) dactylus of minor second pereiopod not exceeding fixed finger (vs. exceeding in P. karantina sp. nov.); 4) major and minor second pereiopods with cutting edge of dactyli with about 20 and 25 teeth, respectively (vs. with about 50 and 40 teeth in P. karantina sp. nov.); 5) fifth pereiopod propodus without distoventral spiniform setae (vs. single stout distoventral and single subdistal spiniform seta in P. karantina sp. nov.); 6) non-tuberculate ventral margin of merus of second pereiopods (vs. tuberculate ventral margin in P. karantina sp. nov.); 7) sixth pleuron with acutely produced posterolaterally (vs. posterolateral angle rounded in P. karantina sp. nov.).</p></div>	https://treatment.plazi.org/id/03AB9F22FF83FFF61D5AFEF1FB5AFBE7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, Jin-Ho;De Grave, Sammy	Park, Jin-Ho, De Grave, Sammy (2021): Two New Species and a Further Country Record of the Caridean Shrimp Genus Periclimenaeus Borradaile, 1915 from Korea (Decapoda: Palaemonidae). Zoological Studies 60 (1): 1-27, DOI: 10.6620/ZS.2021.60-01, URL: http://dx.doi.org/10.5281/zenodo.15500962
03AB9F22FF8DFFEE1D63FBB1FCEDFDA7.text	03AB9F22FF8DFFEE1D63FBB1FCEDFDA7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Periclimenaeus apomonosi Park & De Grave 2021	<div><p>Periclimenaeus apomonosi sp. nov. Park and</p><p>De Grave (Figs. 10–15) urn:lsid:zoobank.org:act: BDC1C00D-392A-421F-8E7B-2933E1B31FF4</p><p>Material examined: Holotype. 1 female (pocl 3.4); Jun. 20, 2018; Munseom Islet, Jejudo <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=126.568886&amp;materialsCitation.latitude=33.226944" title="Search Plazi for locations around (long 126.568886/lat 33.226944)">Island</a> (33°13'37"N 126°34'8"E), 45m, inside of unidentified sponge host, leg. JH Park (NIBRIV0000862976) . Paratypes. 1 female (pocl 2.4); Mar. 31, 2018; same location, 45 m, leg. JH Park (OUMNH. ZC.2018-03- 029), 1 male (pocl 2.6); Mar. 31, 2018; same location, 30 m, inside of same unidentified sponge host, leg. JH Park (NIBRIV 0000862975). GenBank accession numbers for DNA sequences data are presented in table 1.</p><p>Description: Body medium-sized, subcylindrical form (Figs. 10, 15A). Rostrum (Fig. 11B) straight, slightly inclined downwards, about 0.4 of pocl, overreaching end of basal segment of antennular peduncle, 7–8 dorsal teeth along entire length, without ventral tooth.</p><p>Carapace (Fig. 11A, B) smooth, glabrous, with small acute supraorbital tooth, with feeble supraorbital ridge; antennal tooth acute; inferior orbital angle with pointed process (Fig. 11C); pterygostomial angle rounded, not produced anteriorly.</p><p>© 2021 Academia Sinica, Taiwan</p><p>Abdomen smooth (Fig. 11D), first segment with anteromedian dorsal lobe; pleura broadly rounded, sixth segment about 1.2 times length of fifth, about 0.5 of telson length, posterolateral angle sharply pointed, posteroventral angle acute.</p><p>Telson (Fig. 11E) about 0.6 of pocl, about 2.2 times longer than maximum width; two pairs of dorsal subequal spiniform setae at about 0.1 and 0.6 of telson length respectively, posterior margin (Fig. 11F) with three pairs of spiniform setae, lateral posterior spiniform setae short, about one third of length of intermediate pair, intermediate pair long and stout, submedian pair about 0.8 of intermediate pair length, setulose.</p><p>Eye (Fig. 12A) with hemispherical cornea, about 1.3 times longer than maximum dorsal and lateral width, without nebenauge.</p><p>Antennule (Figs. 11A, 12B) with proximal segment of peduncle bearing acute distolateral tooth, with small acute tooth at ventromedial margin; stylocerite bearing sharp point, reaching to about 0.5 of proximal segment; intermediate segment short, about 0.25 times of proximal segment length, subequal in length to distal segment; upper flagellum biramous, proximal five segments fused, short free ramus with three segments, longer free ramus with eleven segments; lower flagellum filiform.</p><p>Antenna (Fig. 12C, D) with rounded medial boss proximally; basicerite with distinct teeth distolaterally an d d is to med ially (F ig. 1 2 C), is ch io cer ite a n d merocerite unarmed; carpocerite reaching about 0.6 of scaphocerite; scaphocerite about 2.0 times as long as maximum width, rounded distal end of lamella almost in line with distolateral tooth.</p><p>Mouthparts not dissected, typical for genus in external observation. Third maxilliped (Fig. 12E) with ultimate segment length about 0.4 times antepenultimate segment, tapering distally, with dense tufts of long setae; penultimate segment length about 0.6 times antepenultimate segment, with ventral row of long setae; antepenultimate segment with long setae on ventromedial margin; exopod reaching 0.9 of antepenultimate segment, with four plumose distal setae; coxa with rounded lateral plate. Arthrobranch absent.</p><p>First pereiopod (Figs. 10, 13A) with coxa and basis without special features; ischium about 0.7 of merus length, unarmed; merus about 1.2 times as long as carpus length, unarmed; carpus about 1.1 times as long as chela length; carpo-propodal cleaning setae developed; chela about 0.3 times as long as pocl; palm subcylindrical, smooth, non-tuberculate; fingers (Fig. 13B) similar in shape, about 0.4 of palm length, with group of setae, subspatulate, with elongate apices of fingers, cutting edges medial, entire.</p><p>© 2021 Academia Sinica, Taiwan</p><p>Second pereiopods (Figs. 10, 13C, E) robust, dissimilar in shape, unequal in size.</p><p>Major second pereiopod (Figs. 10, 13C) with coxa and basis without special features; ischium subequal to merus length, tuberculate ventrally; merus about 0.4 of palm length, tuberculate ventrally; carpus one third of palm length, tapering proximally, unarmed; chela (Fig. 13D) about 1.4 times as long as pocl, about 3.4 times as long as merus length; palm subcylindrical, tuberculate; fingers unequal in size, slightly curved mesially; fixed finger with strong subacute tip, distal cutting edge entire, proximal cutting edge with deep oval fossa, mesial margin with triangular process; dactylus slightly exceeding fixed finger, about 0.4 of palm length, with strong subacute tip, distal cutting edge entire, proximal cutting edge with large molar process.</p><p>© 2021 Academia Sinica, Taiwan</p><p>© 2021 Academia Sinica, Taiwan</p><p>Minor second pereiopod (Figs. 10, 13E) with coxa and basis without special features; ischium subequal to merus length, tuberculate ventrally; merus about 0.5 of palm length, tuberculate ventrally; carpus about 0.3 of palm length, tapering proximally, unarmed; chela (Fig. 13F) about 0.85 times as long as pocl, about 0.6 of major chela length, about 2.6 times as long as merus; palm subcylindrical, tuberculate; fingers unequal in size; fixed finger about 0.85 of dactylus length, with strong subacute tip, cutting edge with groove extending to 0.9 of fixed finger, cutting edge slightly concave, proximal margin with lower triangular process; dactylus about one third of palm length, about 1.7 times as long as maximal depth, slightly exceeding fixed finger, with broadly rounded dorsal margin, tip with large tooth, cutting edge entire, convex, proximal cutting edge angular.</p><p>© 2021 Academia Sinica, Taiwan</p><p>Ambulatory pereiopods subequal (Figs. 10, 14) in shape, fifth pereiopod slightly more slender than third and fourth.</p><p>Third pereiopod (Fig. 14A, B) with coxa and basis without special features; ischium about 0.8 of merus length, unarmed; merus about 1.5 of carpus length, unarmed; carpus about 0.7 of propodus length, unarmed; propodus about 4.6 times as long as maximal depth, sparsely setose, with pair of stout distoventral spiniform setae, five or six spiniform setae along whole ventral border; dactylus about 0.2 of propodus length, biunguiculate; unguis demarcated, curved, corpus compressed, tapering distally, dorsal margin slightly convex, ventral margin slightly concave with distinct acute distal accessory tooth about 0.4 of unguis length, distal two thirds with five acute denticles, sensory setae distolaterally.</p><p>Fourth pereiopod (Fig. 14C, D) with coxa and basis without special features; ischium about 0.7 of merus length, unarmed; merus about 1.5 of carpus length, unarmed; carpus about 0.7 of propodus length, unarmed; propodus about 5.5 times as long as maximal depth, sparsely setose, with pair of stout distoventral spiniform setae, five spiniform setae on ventral border; dactylus about 0.2 of propodus length, biunguiculate; unguis demarcated, curved, corpus compressed, tapering distally, dorsal margin slightly convex, ventral margin slightly concave with distinct acute distal accessory tooth, distal two thirds with four acute denticles, sensory setae distolaterally.</p><p>Fifth pereiopod (Fig. 14E, F) with coxa and basis without special features; ischium about 0.6 of merus length, unarmed; merus about 1.3 of carpus length, unarmed; carpus about 0.7 of propodus length, unarmed; propodus about 7.6 times as long as maximal depth, with two rows of distolateral setae, with single stout distoventral spiniform setae, two spiniform setae distomedially; dactylus about 0.2 of propodus length, biunguiculate; unguis demarcated, curved, corpus compressed, tapering distally, dorsal margin slightly convex, ventral margin slightly concave with distinct acute distal accessory tooth, with five acute denticles, sensory setae distolaterally.</p><p>Second pleopod of male (Fig. 12G) with endopod with long appendix masculina in relation to appendix interna, with two long terminal setulose setae; appendix interna about 1.3 times longer than appendix masculina (Fig. 12H).</p><p>Second pleopod of female (Fig. 12F) with endopod with appendix interna, protopod medially two ovigerous setae proximally, one distally, laterally with single seta proximally.</p><p>Uropodal exopod (Fig. 11E) shorter than endopod, outer margin entire, slightly convex, with single spiniform setae, curved inward, about 2.0 times as long as acute distolateral tooth.</p><p>Variation: The general morphology is very similar between both sexes, even in the relative sizes of the major and minor second chela.</p><p>Host: The specimens were collected from an unidentified sponge (Fig. 15B).</p><p>Etymology: From the Greek apomonosi (απομόνωση, isolation and seclusion), referring to the lifestyle of the new species within the host sponge species. It also alludes to the seclusion of human society due to the coronavirus pandemic (COVID-19), during which time this paper was written. Used as a noun in apposition.</p><p>Distribution: Presently only known from the type locality in Jejudo Island, Korea (Fig. 1A, B).</p><p>Remarks: Periclimenaeus apomonosi sp. nov. appears morphologically close to P. hebedactylus Bruce, 1970 and P. nufu Ďuriš, Horká &amp; Hoc, 2009, sharing with those species the presence of a supraorbital tooth or ridge, the non-denticulate cutting edges of the dactyli of the first and second pereiopods, and a distal accessory tooth as well as the corpus with ventral denticles on the ambulatory dactyli. Periclimenaeus tuamotae Bruce, 1969 and P. bidentatus Bruce, 1970 also resemble the species in having a supraorbital tubercle and non-tuberculate cutting edges. However, the presence of the anterodorsal medial lobe of the first abdominal tergite is not clear in the type (and only) descriptions (Bruce 1969 1970).</p><p>Periclimenaeus hebedactylus is readily distinguished from P. apomonosi by the position of the dorsal spiniform setae on the telson (both pairs at 0.2 of telson length in P. hebedactylus vs. proximal pair at about 0.2 and distal pair at 0.6 in P. apomonosi sp. nov.). Periclimenaeus nufu is also readily separated from the new species by the presence of serrations on the uropodal exopod (vs. non-serrated in P. apomonosi sp. nov.). Periclimenaeus bidentatus clearly differs from the new species by the bidentate distal dactylus of the minor second pereiopod (vs. single tooth in P. apomonosi sp. nov.).</p><p>© 2021 Academia Sinica, Taiwan</p><p>Bruce (1969) provided a detailed, but unillustrated description of P. tuamotae . Based on his description, P. tuamotae and the new species share several characters: 1) carpocerite not exceeding the distal margin of the scaphocerite, 2) a finely tuberculate ventral margin of the second pereiopod chelae, and 3) the presence of tubercles along the ventral margin on the merus and ischium of both second pereiopods. However, P. tuamotae differs from P. apomonosi as follows: 1)</p><p>© 2021 Academia Sinica, Taiwan</p><p>inferior orbital angle obsolete (vs. with pointed process in P. apomonosi sp. nov.), 2) basicerite of antenna unarmed (vs. with distinct teeth distolaterally and distomedially on dorsal margin in P. apomonosi sp. nov.), 3) scaphocerite not exceeding second segment of antennular peduncle (vs. exceeding distal margin of antennular peduncle in P. apomonosi sp. nov.), 4) distolateral tooth of scaphocerite extending beyond the lamella (vs. slightly exceeding or almost in line in P. apomonosi sp. nov.), 5) eye exceeding the rostrum (vs. not in P. apomonosi sp. nov.), 6) fingers of first pereiopod 0.6 of length of palm (vs. 0.4 in P. apomonosi sp. nov.), 7) carpus of first pereiopod about 1.5 times the length of the chela (vs. about 1.1 times in P. apomonosi sp. nov.), 8) two to three spiniform setae along ventral border of fourth pereiopod and none on fifth pereiopod (vs. five and two respectively in P. apomonosi sp. nov.).</p></div>	https://treatment.plazi.org/id/03AB9F22FF8DFFEE1D63FBB1FCEDFDA7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, Jin-Ho;De Grave, Sammy	Park, Jin-Ho, De Grave, Sammy (2021): Two New Species and a Further Country Record of the Caridean Shrimp Genus Periclimenaeus Borradaile, 1915 from Korea (Decapoda: Palaemonidae). Zoological Studies 60 (1): 1-27, DOI: 10.6620/ZS.2021.60-01, URL: http://dx.doi.org/10.5281/zenodo.15500962
03AB9F22FF95FFE21EDAFDF1FEEEFB04.text	03AB9F22FF95FFE21EDAFDF1FEEEFB04.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Periclimenaeus djiboutensis Bruce 1970	<div><p>Periclimenaeus djiboutensis Bruce, 1970</p><p>(Figs. 16–18)</p><p>Periclimenaeus djiboutensis Bruce, 1970: 307 (type locality: Djibouti); 1975: 1568, figs. 3G, H, 8, 9, 13A-D; 1976: 472; 1978: 256, figs. 28, 29; 1981: 11; 1984: 205; Nomura et al., 1996: 10; Marin, 2007: 227, fig. 10; Marin and Caley, 2011, 7.</p><p>Material examined: 1 ovig. female (pocl 3.7); Oct. 15, 2001; Geomundo Island, Dadohaehaesang National Park, Korea (34°2'60"N 127°19'43"E), inside of unidentified sponge host, leg. SNU staff (Laboratory of Systematics and Molecular Evolution) (NIBRIV 0000877271).</p><p>Description of Korean specimen: Body medium-sized, subcylindrical form (Fig. 16). Rostrum (Fig. 17A) straight, downward, about 0.3 of pocl, reaching 0.9 of basal segment of antennular peduncle, 7 dorsal teeth along entire length, without ventral tooth.</p><p>Carapace (Fig. 16) smooth, glabrous, with feeble supraorbital ridge, without supraorbital tooth or tubercle; antennal tooth acute; inferior orbital angle (Fig. 17B) with pointed process; pterygostomial angle rounded, produced anteriorly, with simple setae.</p><p>Abdomen (Fig. 16) smooth, first segment with anteromedian dorsal lobe; pleura broadly rounded, sixth segment as long as fifth length, about 0.6 of telson length, posterolateral angle rounded without tooth, porsteroventral angle rounded.</p><p>Telson (Fig. 17G) about 0.6 of pocl, about 2.5 times maximum width; two pairs of dorsal spiniform subequal setae at about 0.05 and 0.1 of telson length respectively, posterior margin (Fig. 17H) with three pairs of spiniform setae, lateral posterior spiniform setae short, about 0.2 of length of intermediate pair, intermediate pair long and stout, submedian pair about 0.9 of intermediate pair length, reaching distal margin of intermediate pair, setulose.</p><p>© 2021 Academia Sinica, Taiwan</p><p>Eye (Figs. 16, 17C) with hemispherical cornea, about 1.8 times longer than maximum dorsal width, without nebenauge.</p><p>Antennule (Figs. 16, 17C) with proximal segment of peduncle bearing acute distolateral tooth, with acute ventromedial tooth at about 0.5 of proximal segment of peduncle (Fig. 17D); stylocerite bearing sharp point, reaching to about 0.5 of proximal segment; intermediate segment about 0.3 times of proximal segment length; distal segment about 0.7 times of proximal segment length; upper flagellum biramous, lower flagellum filiform.</p><p>Antenna (Fig. 17E, F) with basicerite with distinct rounded lobe laterally, with distomedial tooth; ischiocerite and merocerite unarmed; carpocerite extending to distal end of scaphocerite; scaphocerite about 2.3 times as long as maximum width, rounded distal margin of lamella exceeding acute distolateral tooth.</p><p>Mouthparts not dissected, typical for the genus in external observation.</p><p>First pereiopod (Figs. 16, 18A) with coxa and basis without special features; ischium about 0.6 of merus length, unarmed; merus about 1.1 times as long as carpus length, unarmed; carpus about 1.6 times as long as chela length; chela about 0.3 times of pocl; palm subcylindrical, smooth, non-tuberculate; fingers (Fig. 18B) about 0.2 of palm length, with group of terminal setae, with three terminal teeth.</p><p>Second pereiopods (Figs. 16, 18C, D) robust, dissimilar in shape, unequal in size.</p><p>Major second pereiopod (Figs. 16, 18C) with coxa and basis without special features; ischium about 0.8 of merus length, tuberculate ventrally; merus about 0.4 of palm length, tuberculate ventrally; carpus about 0.3 of palm length, tapering proximally, unarmed; chela about 1.9 times as long as pocl; chela about 3.6 times as long as merus length; palm subcylindrical, tuberculate; fingers unequal in size, slightly curved mesially; fixed finger with strong subacute tip, distal cutting edge entire, proximal cutting edge with deep oval fossa, mesial margin with triangular process, lateral margin with angular process; dactylus slightly exceeding fixed finger, about 0.4 of palm length, with strong subacute tip, distal cutting edge entire, proximal cutting edge with large molar process.</p><p>Minor second pereiopod (Figs. 16, 18D) with coxa and basis without special features; ischium and merus subequal in length, tuberculate ventrally; carpus about 0.3 of palm length, tapering proximally, unarmed; chela about 0.9 times as long as pocl, about 0.5 of major chela length, about 2.7 times as long as merus; palm subcylindrical, tuberculate; fixed finger about 0.4 of palm length, with acute tip, cutting edge with groove, concave, entire; dactylus incomplete, with rounded dorsal margin, cutting edge entire, proximal cutting edge angular.</p><p>Ambulatory pereiopods (Figs. 16, 18E, I, G) subequal in shape, third pereiopod longer and more robust than fourth and fifth; fourth slightly longer and more robust than fifth.</p><p>Third pereiopod (Fig. 18E, F) with coxa and basis without special features; ischium about 0.7 of merus length, unarmed; merus about 1.5 of carpus length, about 2.0 times as long as maximal depth, unarmed; carpus about 0.9 of propodus length, about 2.6 times as long as maximal depth, unarmed; propodus about 3.2 times as long as maximal depth, sparsely setose, with pair of stout distoventral spiniform setae, three spiniform setae on ventral border; dactylus about 0.2 of propodus length, biunguiculate; unguis demarcated, curved, corpus ventrally with distinct acute distal accessory tooth, distoventral tubercles obsolescent, entire proximally, sinuous, concave distally, convex proximally, sensory setae distolaterally.</p><p>Fourth pereiopod (Fig. 18G, H) with coxa and basis without special features; ischium about 0.8 of merus length, unarmed; merus about 1.5 of carpus length, about 3.1 times as long as maximal depth, unarmed; carpus about 0.8 of propodus length, about 3.3 times as long as maximal depth, unarmed; propodus about 4.9 times as long as maximal depth, sparsely setose, with pair of stout distoventral spiniform setae, two spiniform setae on ventral border; dactylus about 0.16 of propodus length, biunguiculate; unguis demarcated, curved, corpus ventrally with distinct acute distal accessory tooth, entire, sinuous, concave distally, convex proximally, sensory setae distolaterally.</p><p>Fifth pereiopod (Fig. 18I, J) with coxa and basis without special features; ischium about 0.7 of merus length, unarmed; merus about 1.2 of carpus length, about 3.7 times as long as maximal depth, unarmed; carpus about 0.8 of propodus length, about 3.6 times as long as maximal depth, unarmed; propodus about 5.3 times as long as maximal depth, with row of distolateral setae, with single stout distoventral spiniform setae, one spiniform setae distomedially; dactylus about 0.2 of propodus length, biunguiculate; unguis demarcated, curved, corpus ventrally with distinct acute distal accessory tooth, entire, sinuous, concave distally, convex proximally, sensory setae distolaterally.</p><p>Uropodal exopod (Fig. 17G) shorter than endopod, outer margin entire, with single spiniform setae, curved inward, about 2.5 times longer than acute distolateral tooth.</p><p>Host: The specimens were collected from an unidentified sponge.</p><p>Distribution: Periclimenaeus djiboutensis was described from Djibouti (Bruce 1970) and has since been reported from Tanzania (Bruce 1976), Madagascar (Bruce 1978), Vietnam (Marin 2007), Queensland, Australia (Bruce 1981; Marin and Caley 2011), and the Ryukyus, Japan (Nomura et al. 1996) (Fig. 1C). The record from Eilat, Israel (Fishelson 1974) was considered to be erroneous by Ďuriš et al. (2011a). The present record is the first for Korea (Fig. 1).</p><p>© 2021 Academia Sinica, Taiwan</p><p>© 2021 Academia Sinica, Taiwan</p><p>Remark: Although the distal margin of the dactylus of the minor second pereiopod is incomplete, the present specimen agrees well with previous descriptions of Periclimenaeus djboutensis by having a rounded lateral lobe on the antennal basicerite, the shape of the chela of the first pereiopod and the characteristic, dorsal spiniform setae situated on the proximal part of the telson (Bruce 1970 1975 1978). These morphological affinities easily distinguish the species from other sponge associated Periclimenaeus species. However, the Korean specimen exhibits the following differences compared with previous descriptions: 1) inferior orbital angle with dentate process in Korean specimen (Fig. 17B), versus absent in the type description (Bruce 1970), 2) corpus of the ambulatory dactyli without distoventral tubercles in Korean (Fig. 18F, H, J) and Vietnamese specimens (see Fig. 10L in Marin 2007), versus blunt tubercles present on the corpus in Madagascan specimens (see Fig. 29G in Bruce 1978), and 3) the antennal basicerite with distomedial dorsal process in Korean (Fig. 17F) and Djibouti specimens (see Fig. 9B in Bruce 1975), but absent in Vietnamese specimens (see Fig. 10B in Marin, 2007). Further, Bruce (1975 1978) illustrated a strongly hooked bidentate tip on the dactylus of the minor second pereiopod. However, Marin (2007) illustrated only a single acute tip in a Vietnamese specimen. It is presently unclear if these differences are intraspecific morphological variation or potentially could reveal further undescribed species.</p></div>	https://treatment.plazi.org/id/03AB9F22FF95FFE21EDAFDF1FEEEFB04	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Park, Jin-Ho;De Grave, Sammy	Park, Jin-Ho, De Grave, Sammy (2021): Two New Species and a Further Country Record of the Caridean Shrimp Genus Periclimenaeus Borradaile, 1915 from Korea (Decapoda: Palaemonidae). Zoological Studies 60 (1): 1-27, DOI: 10.6620/ZS.2021.60-01, URL: http://dx.doi.org/10.5281/zenodo.15500962
