taxonID	type	description	language	source
03A88784FFF6937159C6132D9A76F92F.taxon	diagnosis	Diagnosis. Pronotum distinctly longer than head. Medial pronotal collar very narrow, about one-fifth the length of inner area. Forewing cubital cell about one-third as thick as medial cell. Male operculum wider than long, small and short, failing to cover timbal cover in ventral view; opercula widely separated. Male abdomen more or less translucent ventrally; about as long as or slightly shorter than distance from head to cruciform elevation. Timbal cover globose. Timbal concealed by timbal cover in dorsal view. Aedeagus very thin. Uncal lobes bifurcate, widely separated at base.	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFF6937159C6132D9A76F92F.taxon	discussion	Remarks. Lee (2010) placed four species in Oncotympana, providing a key to the species: O. pallidiventris Stål, 1870, O. viridicincta Stål, 1870, O. nigristigma (Walker, 1850) and an undescribed species but misidentified as O. viridicincta by Endo & Hayashi (1979), all from the Philippines. Four additional species of this genus are described here as new species from Luzon, bringing the number of species in the genus to eight. A new key to the species of Oncotympana is provided here to incorporate all eight species.	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFF6937159C614609C28FC79.taxon	discussion	Remarks. The genus Oncotympana Stål, 1870 has been placed in the tribe Oncotympanini (e. g., Ishihara 1961; Lee 2010), but this genus seems allied to the genus Cicada Linnaeus, the type genus of the subtribe Cicadina of the tribe Cicadini, considering the hyaline wings, the normality of the wing venation, the smallsized male opercula and the semi-opaque male abdominal sternites. Detailed phylogenetic studies are needed to confirm the correct tribal placement of Oncotympana.	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFF7937659C614FF9E87FB71.taxon	description	(Fig. 1)	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFF7937659C614FF9E87FB71.taxon	materials_examined	Type material. Holotype: male (Fig. 1), N. Luzon, CAR border Abra / Kalinga, E. of Malibcong Basiwag, 1,660 m, 17 ° 30.191 ’ N 120 ° 58.942 ’ E, 3 – 4 IV 2008, J. H. Lourens (IRSNB).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFF7937659C614FF9E87FB71.taxon	etymology	Etymology. The specific name is from the Latin grandis meaning “ large ” in reference to the large body (28.7 mm long) of this new species, which is larger than its known congeners (25.3 mm at longest).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFF7937659C614FF9E87FB71.taxon	description	Measurements of types (in mm, 1 male). Length of body: 28.7; length of forewing: 41.1; width of forewing: 14.3; width of head including eyes: 8.2; width of pronotum: 11.9; wing span: 90.4.	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFF7937659C614FF9E87FB71.taxon	diagnosis	Diagnosis. This species is closely allied to O. pallidiventris as it has a short, wide and obconical male abdomen, but it differs as follows: body much larger (28.7 mm long versus 18.0 mm long in pallidiventris); forewing basal vein of apical cell 1 about one-third (more than half in pallidiventris) as long as longitudinal vein of apical cell 1; forewing with infuscation at bases of apical cells 1 – 7 and on cubitus anterior 2 (CuA 2) (only at bases of apical cells 2, 3, 5 and 7 in pallidiventris); forewing with distinct spot appearing on each hind margin of veins radius anterior 2 (RA 2), radius posterior (RP), median veins 1 – 4 (M 1 – 4) and cubitus anterior 1 (CuA 1), forming a series of spots on subapical margin (these spots indistinct in pallidiventris); timbal cover with broad fuscous fascia along posterior margin (without any marks in pallidiventris).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFF7937659C614FF9E87FB71.taxon	description	Description of male (Fig. 1). Head green with following black to fuscous marks: median spot enclosing ocelli, with its anterior end not reaching frontoclypeal suture and posterior end reaching posterior margin of head; pair of rather indistinct spots on supra-antennal plates; pair of small longitudinal fasciae between the median spot and compound eyes; pair of small spots between the indistinct spots and the longitudinal fasciae. Supra-antennal plates developed and produced anteriad. Distance between lateral ocelli and compound eyes slightly less than twice the distance between lateral ocelli. Postclypeus slightly swollen. Antenna dark brown to fuscous. Postclypeus green with medial longitudinal black fascia on about posterior half, small reddish ochraceous spot on top of the medial fascia and fuscous fasciae along transverse grooves, which are disconnected before median part. Anteclypeus light green with broad central fuscous patch. Rostrum ochraceous but black apically; slightly passing posterior margin of hind coxae. Pronotum green. Inner area of pronotum with following black marks: pair of medial longitudinal fasciae broadened near posterior ends but not reaching anterior margin of pronotum; irregular-shaped marks between paramedian fissures and lateral fissures; pair of fasciae along lateral fissures, not complete; pair of curved fasciae along lateral margins of inner area. Pronotal collar margined with black anterolaterally and with pair of brown spots at lateral corners; with posterior marginal area very narrow. Anterior margin of pronotum wider than head including compound eyes. Anterolateral pronotal collar barely dentate. Mesonotum green to greenish ochraceous with following black to fuscous marks: medial longitudinal fascia suddenly broadened from about middle posteriorly to reach anterior margin of cruciform elevation; pair of small roundish spots enclosing scutal depressions; pair of large obconical paramedian marks falling on submedian sigilla, which are not complete but divided into many small pieces; pair of long obconical sublateral marks falling on lateral sigilla, which are not complete but divided into many small pieces; pair of short fascia along lateral margins; pair of small spots on posterolateral corners. Cruciform elevation brown centrally but green laterally with black anterior subapical parts. Thoracic sternites ochraceous to greenish ochraceous. Legs greenish ochraceous to ochraceous with some brown parts and black to fuscous marks. Fore-femur with minute subapical spine as well as primary and secondary spines, all black. Wings hyaline. Forewing tinged with yellowish green especially in basal part. Vein R + Sc mostly greenish brown to brown. Basal vein of apical cell 1 about one-third as long as longitudinal vein of apical cell 1. Infuscation present at bases of apical cells 1 – 7 and on CuA 2. Distinct spot appearing on each hind margin of RA 2, RP, M 1 – 4 and CuA 1, forming a series of spots on subapical margin of forewing. Base of ulnar cell 1 with black node. Basal membrane and base of hind wing jugum vivid jade green. Operculum light green; much wider than long, with rounded posterior apex barely reaching posterior margin of sternite II and with posterolateral corner truncate and almost straight. Opercula widely separated from each other. Abdomen obconical, shorter than distance from head to cruciform elevation. Tergite 2 brown centrally but black laterally. Tergites 3 – 8 black each with brown caudal margin and medial patch. Tergites 3 – 7 with pair of sublateral brown patches, which are densely covered with silvery hairs. Posterior margin of tergite 3 much wider than anterior margin of mesonotum. Timbal cover greenish ochraceous with broad fuscous fascia along posterior margin; globose, about 1.7 times wider than long, with anterior inner corner not angled and with posterior inner corner being close to posterior margin of tergite 2. Timbal completely concealed by timbal cover in dorsal view. Abdominal sternites mostly black with narrow ochraceous posterior margin on each of sternites IV – VI. Genitalia (Fig. 1 C, D): Pygofer barrel-shaped in ventral view. Uncus bifurcate. Uncal lobe hooked outward with apex directed laterad in ventral view, with its inner margin convex at about middle. Dorsal beak narrow triangular, about as long as anal styles. Basal lobe of pygofer absent.	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFF7937659C614FF9E87FB71.taxon	distribution	Distribution. Philippines (N. Luzon).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFF7937659C614FF9E87FB71.taxon	discussion	Remarks. This species is also similar to the undescribed species from Mindanao (Endo & Hayashi 1979) as it has a short, wide and obconical male abdomen and similar forewing venation and infuscation, but it differs as follows: mesonotum with greenish marks less developed than black marks (with much developed greenish marks and minimal black marks in the undescribed species); male operculum rather semicircular (rectangular in the undescribed species); timbal cover comparatively small, with its lateral margin slightly protruding (much protruding in the undescribed species) beyond lateral margin of abdominal segment 3 and with its posterior margin not reaching (extending beyond in the undescribed species) median center of abdominal tergite 3.	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFF1937459C6122A9E9EF8EE.taxon	description	(Fig. 2)	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFF1937459C6122A9E9EF8EE.taxon	materials_examined	Type material. Holotype: male (Fig. 2), E. Luzon, Aurora, 8 km W. of Baler, Sierra Madre, Mingan Mts., 471 m, 16 ° 41.463 ’ N 121 ° 23.86 ’ E, 18 VI 2007, J. H. Lourens (IRSNB). Paratype: 1 male, E. Luzon, Aurora, Sierra Madre, 15 km W. of Dibulo, 650 m, 16 ° 33.176 ’ N 122 ° 13.386 ’ E, 26 – 27 IX 2008, J. H. Lourens (IRSNB).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFF1937459C6122A9E9EF8EE.taxon	etymology	Etymology. The species is named for the Professor Chris Simon at the University of Connecticut, who supported the author’s works.	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFF1937459C6122A9E9EF8EE.taxon	description	Measurements of types (in mm, 2 males). Length of body: 23.4 (23.3 – 23.5); length of forewing: 32.0 (32.0 – 32.0); width of forewing: 10.9 (10.8 – 10.9); width of head including eyes: 6.9 (6.9 – 6.9); width of pronotum: 9.0 (9.0 – 9.0); wing span: 70.8 (69.6 – 71.9).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFF1937459C6122A9E9EF8EE.taxon	diagnosis	Diagnosis. This species is closely allied to O. viridicincta but distinguished by the following characters: body larger (about 23.4 mm long versus 16.0 – 21.0 mm long in viridicincta); pronotum with pair of medial longitudinal black fasciae (with just one, fused, medial longitudinal black fascia in viridicincta); anterior margin of pronotum wider (narrower in viridicincta) than head including compound eyes; forewing costal vein slightly enlarged near node (distinctly enlarged to form protuberance in viridicincta); basal vein of apical cell 1 comparatively shorter, less than one-third (more than one-third in viridicincta) as long as longitudinal vein of apical cell 1; timbal cover without any marks (margined with fuscous in viridicincta).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFF1937459C6122A9E9EF8EE.taxon	description	Description of male (Fig. 2). Head green with following black marks: median spot enclosing ocelli, with its anterior end reaching frontoclypeal suture and posterior end reaching posterior margin of head; pair of spots on supra-antennal plates; three pairs of small spots between the median spot and compound eyes. Supraantennal plates developed and produced anteriad. Distance between lateral ocelli and compound eyes slightly less than or about as long as twice the distance between lateral ocelli. Postclypeus barely swollen. Antenna ochraceous to dark brown. Ventral part of head light green to greenish ochraceous with black marks. Postclypeus laterally margined and with medial longitudinal fascia on about posterior half, small V-shaped mark on about anterior one-third and fasciae along anterior three to four transverse grooves. Anteclypeus with central spot. Rostrum black apically; passing posterior margin of hind coxae. Pronotum green. Inner area of pronotum with following black marks: pair of medial longitudinal fasciae broadened at posterior ends; pair of spots or longitudinal small fasciae between median parts of paramedian fissures and posterior ends of lateral fissures; pair of fasciae along lateral fissures, often not complete; pair of curved fasciae along lateral margins of inner area; other irregular marks. Pronotal collar with pair of spots at lateral corners (indistinct in holotype); with posterior marginal area very narrow. Anterior margin of pronotum wider than head including compound eyes. Anterolateral pronotal collar barely dentate. Mesonotum green to greenish ochraceous with following black to fuscous marks: medial longitudinal fascia suddenly broadened from about middle posteriorly to reach anterior margin of cruciform elevation, broader than median breadth of sublateral marks on lateral sigilla; pair of small roundish spots just enclosing scutal depressions; pair of large obconical paramedian marks falling on submedian sigilla; pair of long obconical, but partly fading, sublateral marks falling on lateral sigilla; pair of small spots on posterolateral corners. Cruciform elevation green to greenish ochraceous with fuscous anterior subapical parts and posterior margin. Thoracic sternites greenish ochraceous. Legs greenish ochraceous to ochraceous with black to fuscous marks. Fore-femur with one (paratype) or two (holotype) minute subapical spines as well as primary and secondary spines, all black except their brown tips. Wings hyaline. Forewing tinged with pale brown. Vein R + Sc mostly reddish ochraceous but partly mixed with green. Infuscation present at bases of apical cells 1 – 7 and on CuA 2. Spot appearing on each hind margin of RA 2, RP, M 1 – 4 and CuA 1, forming a series of spots on subapical margin of forewing. Last spot (on CuA 1) largest among marginal spots. Base of ulnar cell 1 with black node. Basal membrane and base of hind wing jugum slightly grayish jade green. Operculum light green to greenish ochraceous; much wider than long, with narrowly rounded posterior apex barely reaching posterior margin of sternite II and with posterolateral corner truncate and slightly concave. Opercula widely separated from each other. Abdomen about as long as or slightly longer than distance from head to cruciform elevation. Tergite 2 mostly brown without medial mark. Tergites 3 – 8 black each with green caudal margin and brown medial patch. Tergites 3 – 7 with pair of sublateral greenish ochraceous patches, which are densely covered with silvery hairs. Posterior margin of tergite 3 slightly wider than anterior margin of mesonotum. Timbal cover light green to greenish ochraceous without marks; globose, about 1.2 times wider than long, with anterior inner corner roundly angled and with posterior inner corner being close to posterior margin of tergite 2. Timbal completely concealed by timbal cover in dorsal view. Abdominal sternites mostly fuscous. Genitalia (Fig. 2 C, D): Pygofer barrel-shaped in ventral view. Uncus bifurcate. Uncal lobe roundly curved outward, with spine on its lateral margin and with apex directed toward center in ventral view. Both uncal lobes forming heart shape with their inner margins, which is wider than long. Dorsal beak triangular, about as long as or slightly shorter than anal styles. Basal lobe of pygofer absent.	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFF1937459C6122A9E9EF8EE.taxon	distribution	Distribution. Philippines (E. Luzon).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFF1937459C6122A9E9EF8EE.taxon	discussion	Remarks. This species is also similar to O. nigristigma (cf. Lee 2010), but the latter is distinguished from all its congeners by the forewing with the costal margin distinctly bent and convex after node, more slender male abdomen, timbal covers semi-quadrangular, very large and slightly longer than wide and timbal covers close together with very narrow space between them. From the next two new species, O. simonae sp. nov. is distinguished by the following characters: mesonotum with medial longitudinal fascia being broad, broader than median breadth of sublateral marks on lateral sigilla; forewing distinctly tinged with pale brown; forewing with infuscation much more developed; forewing with basal membrane being broad and jade green; timbal cover with anterior inner corner roundly angled; timbal cover with posterior inner corner being close to posterior margin of tergite 2; both uncal lobes forming heart shape with their inner margins, which is wider than long; uncal lobes with apex directed toward center in ventral view.	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFFC937959C617F59D7AFF7F.taxon	description	(Fig. 3)	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFFC937959C617F59D7AFF7F.taxon	materials_examined	Type material. Holotype: male (Fig. 3), E. Luzon, Aurora, Sierra Madre, 14 km S. of Dilalongan, Dapalan River, 50 m, 16 ° 02.709 ’ N 121 ° 42.667 ’ E, 11 – 12 II 2008, J. H. Lourens and K. Knoblich (IRSNB).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFFC937959C617F59D7AFF7F.taxon	etymology	Etymology. The specific name is from the Latin brevis meaning “ short ” in reference to the comparatively short wings of this new species.	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFFC937959C617F59D7AFF7F.taxon	description	Measurements of types (in mm, 1 male). Length of body: 23.9; length of forewing: 29.7; width of forewing: 10.5; width of head including eyes: 6.7; width of pronotum: 8.5; wing span: 66.1.	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFFC937959C617F59D7AFF7F.taxon	diagnosis	Diagnosis. This and the next new species are closely allied to O. simonae but distinguished by the following characters: mesonotum with medial longitudinal fascia being narrow, narrower than sublateral marks on lateral sigilla; forewing barely tinged; forewing with infuscation much reduced; forewing with basal membrane being narrower and gray; timbal cover with anterior inner corner not angled; timbal cover with posterior inner corner being widely separated from posterior margin of tergite 2; both uncal lobes forming heart shape with their inner margins, which is longer than wide; uncal lobes with apex directed downward (anteriad) or laterad in ventral view.	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFFC937959C617F59D7AFF7F.taxon	description	Description of male (Fig. 3). Head ochraceous with following black marks: median spot enclosing ocelli, with its anterior end reaching frontoclypeal suture and posterior end reaching posterior margin of head; pair of spots on supra-antennal plates; three pairs of small spots between the median spot and compound eyes; pair of spots on anterolateral corners. Supra-antennal plates developed and produced anteriad. Distance between lateral ocelli and compound eyes slightly less than or about as long as twice the distance between lateral ocelli. Postclypeus barely swollen. Antenna ochraceous to dark brown. Ventral part of head light green to greenish ochraceous with black marks. Postclypeus laterally margined and with medial longitudinal fascia on about posterior half, small V-shaped mark on about anterior one-third and fasciae along anterior three to four transverse grooves. Anteclypeus with central spot. Rostrum black apically; passing posterior margin of hind coxae. Pronotum ochraceous. Inner area of pronotum with following black marks: pair of medial longitudinal fasciae broadened at posterior ends; pair of spots or longitudinal small fasciae between median parts of paramedian fissures and posterior ends of lateral fissures; pair of fasciae along lateral fissures, often not complete; pair of curved fasciae along lateral margins of inner area; other irregular marks. Pronotal collar with pair of spots at lateral corners; with posterior marginal area very narrow. Anterior margin of pronotum wider than head including compound eyes. Anterolateral pronotal collar with two or three indistinct teeth. Mesonotum greenish ochraceous with following black to fuscous marks: medial longitudinal fascia suddenly broadened from about middle posteriorly to reach anterior margin of cruciform elevation, narrower than median breadth of obconical sublateral marks on lateral sigilla; pair of small roundish spots just enclosing scutal depressions; pair of large obconical paramedian marks falling on submedian sigilla; pair of long obconical, but partly fading, sublateral marks falling on lateral sigilla; pair of small spots on posterolateral corners. Cruciform elevation green to greenish ochraceous with fuscous anterior subapical parts and posterior margin. Thoracic sternites ochraceous. Legs greenish ochraceous to ochraceous with black to fuscous marks. Fore-femur with minute subapical spine as well as primary and secondary spines, all black except their brown tips. Wings hyaline. Forewing barely tinged; comparatively short and broad. Vein R + Sc mostly reddish ochraceous. Infuscation present at bases of apical cells 1, 2, 3, 5 and 7 and on CuA 2. Minute spot appearing on each hind margin of RA 2, RP, M 1 – 4 and CuA 1, forming a series of spots on subapical margin of forewing. Base of ulnar cell 1 with black node. Basal membrane reduced and gray. Hind wing short, its apex barely reaching the junction of m-cu and M 4. Base of hind wing jugum gray. Operculum greenish ochraceous; much wider than long, with widely rounded posterior apex barely reaching posterior margin of sternite II and with posterolateral corner slightly concave. Opercula widely separated from each other. Abdomen about as long as or slightly longer than distance from head to cruciform elevation. Tergite 2 brown with greenish posterior margin and medial longitudinal fuscous fascia. Tergites 3 – 7 black each with blue caudal margin and brown medial patch. Tergites 3 – 7 with pair of sublateral greenish ochraceous patches, which are densely covered with silvery hairs. Posterior margin of tergite 3 slightly wider than anterior margin of mesonotum. Timbal cover brownish gray without marks; globose, about 1.2 times wider than long, with anterior inner margin roundly convex widely, not angled, and with posterior inner corner being widely separated from posterior margin of tergite 2. Timbal completely concealed by timbal cover in dorsal view. Abdominal sternites mostly fuscous with bluish posterior margin on each of sternites IV – VI. Genitalia (Fig. 3 C, D): Pygofer barrel-shaped in ventral view. Uncus bifurcate. Uncal lobe curved outward, with spine on its lateral margin and with truncate and laterally acute apex directed downward in ventral view. Both uncal lobes forming heart shape with their inner margins, which is longer than wide. Dorsal beak triangular, shorter than anal styles. Basal lobe of pygofer absent.	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFFC937959C617F59D7AFF7F.taxon	distribution	Distribution. Philippines (E. Luzon).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFFE937F59C616309D7AFEC4.taxon	description	(Fig. 4)	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFFE937F59C616309D7AFEC4.taxon	materials_examined	Type material. Holotype: male (Fig. 4), E. Luzon, Quirino, Sierra Madre, 35 km E. of Naptipunan, S. Pul. Lupa, 15 ° 58 ’ N 121 ° 29 ’ E, 11 – 12 III 2005, J. H. Lourens (IRSNB).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFFE937F59C616309D7AFEC4.taxon	etymology	Etymology. The specific name is from the Latin undatus meaning “ wavy ” in reference to the long and wavy uncal lobes of this new species.	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFFE937F59C616309D7AFEC4.taxon	description	Measurements of types (in mm, 1 male). Length of body: 25.3; length of forewing: 34.3; width of forewing: 11.4; width of head including eyes: 7.3; width of pronotum: 9.6; wing span: 74.7.	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFFE937F59C616309D7AFEC4.taxon	diagnosis	Diagnosis. This species is distinguished from O. brevis by the following characters: forewing comparatively long and slender (ratio of forewing length to forewing width approximately 3.01 versus 2.83 in O. brevis); hind wing long, its apex passing (barely reaching in O. brevis) the junction of mediocubital crossvein (m-cu) and M 4; timbal cover with anterior inner margin truncate and slightly concave (roundly convex in O. brevis); uncal lobes with triangular apex directed laterad in ventral view (in O. brevis, uncal lobes with truncate and laterally acute apex directed downward in ventral view).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFFE937F59C616309D7AFEC4.taxon	description	Description of male (Fig. 4). Head reddish ochraceous with following black marks: median spot enclosing ocelli, with its anterior end reaching frontoclypeal suture and posterior end reaching posterior margin of head; pair of spots on supra-antennal plates; three pairs of small spots between the median spot and compound eyes; pair of spots on anterolateral corners. Supra-antennal plates developed and produced anteriad. Distance between lateral ocelli and compound eyes slightly less than or about as long as twice the distance between lateral ocelli. Postclypeus barely swollen. Antenna ochraceous to dark brown. Ventral part of head light green to greenish ochraceous with black marks. Postclypeus laterally margined and with medial longitudinal fascia on about posterior half, small V-shaped mark on about anterior one-third and fasciae along anterior three to four transverse grooves. Anteclypeus with central spot. Rostrum black apically; passing posterior margin of hind coxae. Pronotum reddish ochraceous. Inner area of pronotum with following black marks: pair of medial longitudinal fasciae broadened at posterior ends; pair of spots or longitudinal small fasciae between median parts of paramedian fissures and posterior ends of lateral fissures; pair of fasciae along lateral fissures, often not complete; pair of curved fasciae along lateral margins of inner area; other irregular marks. Pronotal collar with pair of spots at lateral corners; with posterior marginal area very narrow. Anterior margin of pronotum wider than head including compound eyes. Anterolateral pronotal collar with one to three indistinct teeth. Mesonotum slightly greenish ochraceous with following black to fuscous marks: medial longitudinal fascia suddenly broadened from about middle posteriorly to reach anterior margin of cruciform elevation, broader than median breadth of sublateral marks on lateral sigilla; pair of small roundish spots just enclosing scutal depressions; pair of rather indistinct obconical paramedian marks falling on submedian sigilla; pair of long obconical, but rather indistinct, sublateral marks falling on lateral sigilla; pair of small spots on posterolateral corners. Cruciform elevation green to greenish ochraceous with fuscous anterior subapical parts and posterior margin. Thoracic sternites greenish ochraceous. Legs ochraceous with black to fuscous marks. Fore-femur with minute subapical spine as well as primary and secondary spines, all black except their brown tips. Wings hyaline, comparatively long and slender. Forewing barely tinged; comparatively short and broad. Vein R + Sc mostly reddish ochraceous. Infuscation present at bases of apical cells 1, 2, 3, 5 and 7 and on CuA 2. Minute spot appearing on each hind margin of RA 2, RP, M 1 – 4 and CuA 1, forming a series of spots on subapical margin of forewing. Base of ulnar cell 1 with black node. Basal membrane reduced and gray. Hind wing long, its apex passing the junction of m-cu and M 4. Base of hind wing jugum gray. Operculum greenish ochraceous; much wider than long, with widely rounded posterior apex barely reaching posterior margin of sternite II and with posterolateral corner slightly concave. Opercula widely separated from each other. Abdomen about as long as or slightly longer than distance from head to cruciform elevation .. Tergite 2 ochraceous to brown (maybe due to discoloration) with medial darker patch. Tergites 3 – 7 mostly fuscous each with ochraceous caudal margin and pair of sublateral ochraceous patches, the latter being densely covered with silvery hairs. Posterior margin of tergite 3 distinctly wider than anterior margin of mesonotum. Timbal cover ochraceous inwardly and greenish outwardly, without marks; globose, about 1.3 times wider than long, with anterior inner margin truncate and slightly concave, not angled, and with posterior inner corner being widely separated from posterior margin of tergite 2. Timbal slightly revealed anteriorly in dorsal view. Abdominal sternites mostly fuscous with bluish posterior margin on each of sternites III-V. Genitalia (Fig. 4 C, D): Pygofer barrel-shaped in ventral view. Uncus bifurcate. Uncal lobe curved outward, with spine on its lateral margin and with triangular apex directed laterad in ventral view. Both uncal lobes forming heart shape with their basal inner margins, which is longer than wide. Dorsal beak triangular, about as long as anal styles. Basal lobe of pygofer absent.	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFFE937F59C616309D7AFEC4.taxon	distribution	Distribution. Philippines (E. Luzon).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFF8937F59C615B89E05F931.taxon	materials_examined	Material examined. 1 male, E. Luzon, Sierra Madre Mts., 10 km W. of Infanta, along Pisa River, Brgy Magsaysay, 100 m, 1 IV 2004, J. H. Lourens (IRSNB); 1 female, E. Luzon, N. of Infanta, Sierra Madre Mts., 4 km W. of Gen. Nakar, lowland forest, 120 m, 10 VIII 2004, J. H. Lourens (IRSNB); 1 male, 1 female, E. Luzon, N. Ec. Bongabong Bgy Laby, Sierra Madre, Mingan Mts., 500 m, 15 ° 38 ’ N 121 ° 15 ’ E, 8 VI 2005, J. H. Lourens (IRSNB); 1 female, N. W. Luzon, Apayao Pagudpud Adams, Bgy Pantiang, Managa River, 340 m, 18 ° 31.278 ’ N 120 ° 54.642 ’ E, 21 IV 2007, J. H. Lourens (IRSNB); 2 females, E. Luzon, Aurora, Sierra Madre, 13 km W. of Diatagon, 585 m, 16 ° 32.886 ’ N 122 ° 14.134 ’ E, 14 – 15 VI 2007, J. H. Lourens (IRSNB); 1 male, 2 female, E. Luzon, Aurora, 8 km W. of Baler, Sierra Madre, Mingan Mts., 471 m, 16 ° 41.463 ’ N 121 ° 23.86 ’ E, 18 VI 2007, J. H. Lourens (IRSNB); 1 male, 11 females, E. Luzon, Aurora, Sierra Madre, S. of Dilalongan, Dapalan River, 50 m, 16 ° 02.707 ’ N 121 ° 42.673 ’ E, 4 IX 2007, J. H. Lourens (IRSNB); 3 males, 8 females, E. Luzon, Aurora, Sierra Madre, 15 km W. of Dibulo, 50 m, 16 ° 32.866 ’ N 122 ° 14.134 ’ E, 5 – 6 IX 2007, J. H. Lourens (IRSNB); 3 males, 7 females, E. Luzon, Aurora, Sierra Madre, 14 km S. of Dilalongan, Dapalan River, 50 m, 16 ° 02.709 ’ N 121 ° 42.667 ’ E, 11 – 12 II 2008, J. H. Lourens and K. Knoblich (IRSNB); 1 female, N. Luzon, 5 km N. of Adams, Managa River, 140 m, 18 ° 31.359 ’ N 120 ° 54.674 ’ E, 15 II 2008, J. H. Lourens and K. Knoblich (IRSNB); 1 female, N. Luzon, 5 km S. of Adams, 350 m, 18 ° 31.338 ’ N 120 ° 55.690 ’ E, 6 – 7 IV 2008, J. H. Lourens (IRSNB); 2 males, E. Luzon, Aurora, Sierra Madre, 13 km W. of Dibulo, 585 m, 16 ° 32.886 ’ N 122 ° 14.134 ’ E, 24 – 25 IX 2008, J. H. Lourens (IRSNB); 2 females, E. Luzon, Aurora, Sierra Madre, 15 km W. of Dibulo, 650 m, 16 ° 33.176 ’ N 122 ° 13.386 ’ E, 26 – 27 IX 2008, J. H. Lourens (IRSNB); 2 females, Marinduque, Sihujao, Mt. Masimot, primary hill forest edge, 560 m, 13 ° 20.830 ’ N 122 ° 00.219 ’ E, 21 – 23 V 2009, J. H. Lourens (IRSNB).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFF8937F59C615B89E05F931.taxon	distribution	Distribution. Philippines (Luzon, Mindoro, Homonhon, Samar, Panay, Leyte, Negros, Mindanao and Basilan).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFF9937E59C6136F9CB7FB37.taxon	discussion	Remarks. This genus was resurrected and redefined by Lee (2010).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFF9937E59C612689CD2F979.taxon	materials_examined	Material examined. 5 females, E. Luzon, Aurora, Sierra Madre, 13 km W. of Diatagon, 585 m, 16 ° 32.886 ’ N 122 ° 14.134 ’ E, 14 – 15 VI 2007, J. H. Lourens (IRSNB); 2 males, 2 females, E. Luzon, Aurora, 8 km W. of Baler, Sierra Madre, Mingan Mts., 471 m, 16 ° 41.463 ’ N 121 ° 23.86 ’ E, 18 VI 2007, J. H. Lourens (IRSNB); 1 male, 2 females, E. Luzon, Aurora, Sierra Madre, 15 km W. of Dibulo, 50 m, 16 ° 32.866 ’ N 122 ° 14.134 ’ E, 5 – 6 IX 2007, J. H. Lourens (IRSNB); 2 male, N. Luzon, 5 km S. of Adams, 350 m, 6 – 7 IV 2008, J. H. Lourens (IRSNB); 1 female, Marinduque, Sihujao, Mt. Masimot, primary hill forest edge, 560 m, 13 ° 20.830 ’ N 122 ° 00.219 ’ E, 21 – 23 V 2009, J. H. Lourens (IRSNB).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFF9937E59C612689CD2F979.taxon	distribution	Distribution. Philippines (Luzon, Sibuyan, Samar and Mindanao).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFF9937E59C617F59D41FC34.taxon	materials_examined	Material examined. 3 males, 1 female, E. Luzon, Sierra Madre Mts., 10 km W. of Infanta, along Pisa River, Brgy Magsaysay, 100 m, 1 IV 2004, J. H. Lourens (IRSNB); 2 females, S. E. Luzon, southern Sierra Madre Mts., 12 km N. of Mauban, primary lowland forest, 120 m, 16 VI 2004, J. H. Lourens (IRSNB); 1 female, same data except 23 VI 2004 (IRSNB); 1 female, E. Luzon, N. of Infanta, Sierra Madre Mts., 4 km W. of Gen. Nakar, lowland forest, 120 m, 10 VIII 2004, J. H. Lourens (IRSNB); 1 male, 1 female, E. Luzon, Quirino, Sierra Madre, 35 km E. of Naptipunan, S. Pul. Lupa, 15 ° 58 ’ N 121 ° 29 ’ E, 11 – 12 III 2005, J. H. Lourens (IRSNB); 1 female, E. Luzon, Nueva Ecija Sierra Madre, Mingan Mts., Brgy Laby, 15 ° 40.432 ’ N 121 ° 17.906 ’ E, 650 m, 3 IX 2007, J. H. Lourens (IRSNB); 4 females, E. Luzon, Aurora, Sierra Madre, 15 km W. of Dibulo, 50 m, 16 ° 32.866 ’ N 122 ° 14.134 ’ E, 5 – 6 IX 2007, J. H. Lourens (IRSNB); 1 male, E. Luzon, Aurora, Sierra Madre, 14 km S. of Dilalongan, Dapalan River, 50 m, 16 ° 02.709 ’ N 121 ° 42.667 ’ E, 11 – 12 II 2008, J. H. Lourens and K. Knoblich (IRSNB); 1 male, Luzon, Quezon Prov., Quezon National Park, Pinagbanderahan, III 2008, J. Bresseel (IRSNB); 1 male, N. Luzon, CAR border Abra / Kalinga, E. of Malibcong Basiwag, 1,660 m, 17 ° 30.191 ’ N 120 ° 58.942 ’ E, 3 – 4 IV 2008, J. H. Lourens (IRSNB); 1 male, 3 females, Marinduque, Sihujao, Mt. Masimot, primary hill forest edge, 560 m, 13 ° 20.830 ’ N 122 ° 00.219 ’ E, 21 – 23 V 2009, J. H. Lourens (IRSNB).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFF9937E59C617F59D41FC34.taxon	distribution	Distribution. S. China, Philippines (Luzon, Sibuyan, Panay, Palawan and Mindanao), Malay Archipelago, S. Myanmar and India.	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFF9937D59C6102D9E95FDCF.taxon	materials_examined	Material examined. 1 male, E. Luzon, Isabela, Sierra Madre, 22 km N. W. of Dinapique, logged primary forest, 700 m, 16 ° 33 ’ N 122 ° 13 ’ E, 4 IX 2006, J. H. Lourens (IRSNB); 1 male, E. Luzon, N. Ec. Bongabon Bgy Laby, Sierra Madre, Mingan Mts., 950 m, 15 ° 38 ’ N 121 ° 15 ’ E, 6 – 7 IX 2006, J. H. Lourens (IRSNB); 1 male, 1 female, E. Luzon, Aurora, Sierra Madre, 13 km W. of Diatagon, 585 m, 16 ° 32.886 ’ N 122 ° 14.134 ’ E, 14 – 15 VI 2007, J. H. Lourens (IRSNB); 1 male, 5 females, E. Luzon, Aurora, 8 km W. of Baler, Sierra Madre, Mingan Mts., 471 m, 16 ° 41.463 ’ N 121 ° 23.86 ’ E, 18 VI 2007, J. H. Lourens (IRSNB); 1 female, E. Luzon, Nueva Ecija Sierra Madre, Mingan Mts., Brgy Laby, 15 ° 40.432 ’ N 121 ° 17.906 ’ E, 650 m, 3 IX 2007, J. H. Lourens (IRSNB); 1 female, E. Luzon, Aurora, Sierra Madre, 15 km W. of Dibulo, 50 m, 16 ° 32.866 ’ N 122 ° 14.134 ’ E, 5 – 6 IX 2007, J. H. Lourens (IRSNB); 1 male, E. Luzon, Aurora, Sierra Madre, 13 km W. of Dibulo, 585 m, 16 ° 32.886 ’ N 122 ° 14.134 ’ E, 24 – 25 IX 2008, J. H. Lourens (IRSNB).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFF9937D59C6102D9E95FDCF.taxon	distribution	Distribution. Philippines (Luzon (including Camigun Island )), Negros, Samar, Leyte, Mindanao and Basilan) and Borneo.	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFFA937D59C615FF9D5AFCEF.taxon	materials_examined	Material examined. Type material in Lee (2009 a). Distribution. Philippines (Luzon).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFFA937D59C614A09D5AFB11.taxon	materials_examined	Material examined. Type material in Lee (2009 a). Distribution. Philippines (Luzon).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFFA937D59C612A79E6DF95F.taxon	discussion	Remarks. Lee & Hill (2010) placed 12 species in Psithyristria, providing a key to the species, all from Luzon. Two additional species of this genus are described here as new species from Luzon, bringing the number of species in the genus to 14. A new key to the species of Psithyristria is provided here to incorporate all the 14 species.	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFFB937C59C611B59D7AF824.taxon	materials_examined	Material examined. Type material in Lee & Hill (2010). Distribution. Philippines (E. Luzon).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFE4936359C616F59D7AFDE7.taxon	materials_examined	Material examined. Type material in Lee & Hill (2010). Distribution. Philippines (E. Luzon).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFE4936359C614D89D84FB09.taxon	materials_examined	Material examined. Type material in Lee & Hill (2010). Distribution. Philippines (C. Luzon).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFE4936359C615D89D71FCE7.taxon	distribution	Distribution. Philippines (N. W. Luzon).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFE4936359C613BD9D4AFA04.taxon	distribution	Distribution. Philippines (S. Luzon).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFE4936159C612BF9D86F838.taxon	description	(Fig. 5)	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFE4936159C612BF9D86F838.taxon	materials_examined	Type material. Holotype: male (Fig. 5), N. Luzon, CAR border Abra / Kalinga, E. of Malibcong Basiwag, 1,660 m, 17 ° 30.191 ’ N 120 ° 58.942 ’ E, 3 – 4 IV 2008, J. H. Lourens (IRSNB). Paratypes: 2 males, same data as holotype (IRSNB).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFE4936159C612BF9D86F838.taxon	etymology	Etymology. The specific name refers to the morphological similarity of this species to P. tenuinervis.	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFE4936159C612BF9D86F838.taxon	description	Measurements of types (in mm, 3 males). Length of body: 22.2 (21.2 – 23.5); length of forewing: 29.1 (28.7 – 29.3); width of forewing: 9.2 (9.1 – 9.3); width of head including eyes: 5.6 (5.5 – 5.6); width of pronotum: 7.1 (7.0 – 7.1); wing span: 63.1 (61.7 – 64.7).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFE4936159C612BF9D86F838.taxon	diagnosis	Diagnosis. This species is closely allied to P. tenuinervis as it has a very slender forewing and the least developed medial and cubital cells, but it differs as follows: body larger (about 22.2 mm long versus 16.0 – 19.0 mm long in tenuinervis); forewing nodal line indistinct or absent in medial cell (distinct in other Psithyristria species except for P. specularis); M 2 derived from ulnar cell 2 (from ulnar cell 3 in other Psithyristria species except for P. specularis, P. nodinervis, P. crassinervis and P. moderabilis, of which latter two have unstable M 2 divergence); M 1 + 2 and M 3 + 4 becoming distinctly closer to each other distally (not becoming distinctly closer in other species except for P. paraspecularis); M along medial cell short, about half as long as M along ulnar cell 3 (about as long as M along ulnar cell 3 in tenuinervis); CuA 1 about three times as long as CuA 2 (less than twice as long as CuA 2 in tenuinervis); timbal cover well developed, covering most part of timbal (very small, exposing most part of timbal in tenuinervis); uncal lobes widely separated (narrowly separated in tenuinervis) from each other at base; uncal lobe in lateral view thin with its inner margin considerably concave (rather thick with its inner margin slightly convex in tenuinervis).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFE4936159C612BF9D86F838.taxon	description	Description of male (Fig. 5). Head light green with following marks: median large black spot enclosing ocelli, with its anterior end reaching frontoclypeal suture and posterior end reaching pronotum; pair of large oblique dark brown fasciae on sides of the median spot, narrowly connecting with the median spot and their lateral sides nearly reaching compound eyes; pair of dark brown spots on supra-antennal plates; one or two pairs of minute spots posterolaterally. Distance between lateral ocelli and compound eyes about 1.5 times as wide as distance between lateral ocelli. Postclypeus moderately swollen. Antennae mostly fuscous. Ventral part of head light green to greenish ochraceous. Postclypeus with pair of roundish dark brown patch on anterior part neighboring supra-antennal plates and dark brown fasciae along anterior four to five transverse grooves. Anteclypeus without marks. Lorum with black spot on inner margin of posterior part. Gena with minute dark brown to fuscous spot at about middle of lateral margin. Rostrum fuscous apically; slightly passing hind coxae. Pronotum light green. Inner area of pronotum with following dark brown to fuscous marks: pair of central longitudinal fasciae broadened at both anterior and posterior ends; pair of fasciae between anterior parts of paramedian fissures and posterior ends of lateral fissures, often disconnected in the middle; pair of fasciae along lateral fissures; pair of curved fasciae along lateral margins of inner area. Pronotal collar with two pairs of, sometimes indistinct, dark brown spots at lateral inner corner. Anterolateral pronotal collar with or without minute tooth. Mesonotum light green to greenish ochraceous, covered with silvery hairs especially on lateral and posterior parts and with following black to fuscous marks: median longitudinal fascia broadened posteriorly to reach posterior margin of cruciform elevation; pair of inwardly curved fasciae along parapsidal sutures; pair of small roundish spots enclosing scutal depressions; pair of broad longitudinal fasciae on lateral sides of the roundish spots all the way to anterior margin and to posterior margin of mesonotum; pair of longitudinal short fasciae between the inwardly curved fasciae and the broad longitudinal patch, with their anterior ends reaching anterior margin of mesonotum. Cruciform elevation green to greenish ochraceous with median longitudinal fuscous fascia and narrowly fuscous posterior margin. Thoracic sternites light green to greenish ochraceous. Legs greenish ochraceous to ochraceous. Fore-femur primary and secondary spines dark brown to fuscous. Fore-tibia and fore- and mid-tarsi mostly dark brown. Fore-, mid- and hind pretarsal claws apically fuscous. Wings hyaline. Forewing M 2 derived from ulnar cell 2. M 3 derived from ulnar cell 3. M 4 derived from medial cell. Ulnar cell 3 about 1.3 times as large as ulnar cell 2. Medial cell about 1.2 times as large as ulnar cell 3. Medial cell about as large as cubital cell. CuA 1 about three times as long as CuA 2. Infuscation present on bases of apical cells 2 – 5 and on CuA 2. Spot appearing on each hind margin of RA 2, RP, M 1 – 4 and CuA 1, forming a series of spots on subapical margin of forewing. Smoky distal suffusion usually present along apical margins of apical cells 1 – 5. Basal membrane and hind wing jugum reddish orange. Operculum light green or ochraceous, narrowly margined with fuscous laterally; semicircular. Lateral margin of operculum sinuate at base. Opercula widely separated from each other. Abdomen longer than distance from head to cruciform elevation; covered with silvery or fuscous hairs. Tergites 2 – 6 ochraceous with median longitudinal dark brown to brown fascia, with its lateral margins usually darker than medial part. Tergites 7 and 8 dark brown to fuscous. Tergites 3 – 4 with pair of fuscous spots laterally. Tergites 5 – 6 with pair of small fuscous spots sublaterally. Caudal margin of tergite 3 slightly wider than anterior margin of mesonotum. Timbal cover ochraceous or brown; about as long as the median width, semicircular. Timbal mostly covered with timbal cover. Abdominal sternites mostly ochraceous except sternite VII, which is mostly black. Sternite VI brown to dark brown caudally. Sternite VIII entirely ochraceous. Genitalia (Fig. 5 C, D): Pygofer barrel-shaped in ventral view. Uncal lobes separated from each other at base; simple, without additional spine, in ventral view lateral margins curved first convexly then concavely, terminating in single acute apex pointing outward. Dorsal beak long, triangular.	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFE4936159C612BF9D86F838.taxon	distribution	Distribution. Philippines (N. Luzon).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFE7936059C617F59DA6FE31.taxon	distribution	Distribution. Philippines (N. W. Luzon).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFE7936659C615659D7BFDDC.taxon	description	(Figs 6, 7)	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFE7936659C615659D7BFDDC.taxon	materials_examined	Type material. Holotype: male (Fig. 6), S. Luzon, 10 km E. of Irosin, Mt. Bulusan lake, 280 m, 12 ° 44.872 ’ N 124 ° 05.925 ’ E, 12 III 2009, J. H. Lourens & K. Cerny (IRSNB).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFE7936659C615659D7BFDDC.taxon	etymology	Etymology. The specific name refers to the morphological similarity of this species to P. crassinervis.	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFE7936659C615659D7BFDDC.taxon	description	Measurements of types (in mm, 1 male). Length of body: 18.7; length of forewing: 21.0; width of forewing: 8.0; width of head including eyes: 5.2; width of pronotum: 6.2; wing span: 47.5.	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFE7936659C615659D7BFDDC.taxon	diagnosis	Diagnosis. This species has the enlarged and elevated junctions of the forewing M 4 and M 3 + 4, of M 3 and ulnar cell 3 and of M 2 and M 1 + 2 as in P. crassinervis and is nearly indistinguishable from crassinervis in external morphology except for its smaller body (about 18.7 mm long versus 20.2 – 22.1 mm long in crassinervis) and the reduced forewing ulnar cell 2. However, this species differs from all its congeners in having an unusual shape of the uncal lobe, which has a rounded apex (without a lateral spine as in crassinervis) and a pair of narrow, sharply pointed fin-like projections extending downward from the apex and from the paramedian margin of the uncal lobe (Figs 6 C, D, 7).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFE7936659C615659D7BFDDC.taxon	description	Description of male (Figs 6, 7). Head light green with following marks: median black mark enclosing ocelli, with its anterior end reaching frontoclypeal suture and posterior end reaching pronotum; pair of fuscous patches on sides of the median mark, narrowly connecting with the median mark and their lateral sides widely connecting with compound eyes; pair of fuscous patches on supra-antennal plates; pair of fuscous minute spots posterolaterally. Distance between lateral ocelli and compound eyes about twice as wide as distance between lateral ocelli. Postclypeus moderately swollen. Antennae mostly fuscous. Ventral part of head light green to greenish ochraceous. Postclypeus with fuscous fasciae along anterior six to seven transverse grooves laterally. Anteclypeus without distinct marks. Lorum without distinct marks. Gena with small fuscous spot at about middle of lateral margin. Rostrum fuscous apically; reaching posterior margin of hind coxae. Pronotum light green. Inner area of pronotum with following black to fuscous marks: pair of central longitudinal fasciae broadened at both anterior and posterior ends; pair of fasciae between anterior parts of paramedian fissures and posterior ends of lateral fissures, disconnected in the middle; pair of thick fasciae along lateral fissures; pair of curved fasciae along lateral parts of anterior margin and lateral margins of inner area. Pronotal collar with pair of indistinct spots at lateral inner corner and pair of indistinct spots sublaterally. Anterolateral pronotal collar not dentate. Mesonotum light green to greenish brown, covered with silvery hairs especially on lateral and posterior parts and with following marks: median longitudinal fuscous fascia broadened posteriorly to reach anterior margin of cruciform elevation; pair of inwardly curved fuscous fasciae along parapsidal sutures; pair of small roundish fuscous spots on scutal depressions; pair of broad longitudinal dark brown fasciae on lateral sides of the roundish spots all the way to anterior margin and to posterior margin of mesonotum; pair of longitudinal short fasciae between the inwardly curved fasciae and the broad longitudinal fasciae, with their anterior ends reaching anterior margin of mesonotum. Cruciform elevation greenish ochraceous without distinct marks. Thoracic sternites greenish ochraceous without marks. Legs greenish ochraceous to ochraceous. Fore-, mid- and hind tarsi ochraceous to brown. Fore-, mid- and hind pretarsal claws brown, but apically fuscous. Wings hyaline. Forewing M 2 derived from ulnar cell 3, very close to junction of M 2 and M 1 + 2. M 3 derived from ulnar cell 3, closer to junction of M 4 and M 3 + 4. M 4 derived from junction of M 4 and M 3 + 4. Ulnar cell 3 about 1.7 times as large as ulnar cell 2. Medial cell about 1.3 times as large as ulnar cell 3. Medial cell slightly smaller than cubital cell. M 1 + 2 slightly curved anteriorly. CuA 1 about three times as long as CuA 2. Infuscation present on bases of apical cells 2 – 4 and on CuA 1 and CuA 2. Spot appearing on each hind margin of RA 2, RP and M 1 – 4, forming a series of spots on subapical margin of forewing. Smoky distal suffusion usually present inside and along apical margins of apical cells 1 – 6. Basal membrane and hind wing jugum reddish ochraceous. Operculum greenish ochraceous; semicircular. Opercula widely separated from each other. Abdomen much longer than distance from head to cruciform elevation; covered with silvery or fuscous hairs. Tergites 2 – 6 greenish ochraceous with median longitudinal brown fascia, with its lateral parts usually looking darker than medial part because of fuscous hairs. Tergites 7 and 8 dark brown to fuscous. Tergites 3 – 6 with pair of fuscous spots laterally. Caudal margin of tergite 3 about as wide as anterior margin of mesonotum. Timbal cover brown; wider than long in median width, semicircular but inner margin nearly straight. Timbal largely exposed medially. Abdominal sternites mostly greenish ochraceous except sternites VI – VIII. Sternite VI brown caudally. Sternites VII and VIII mostly brown. Genitalia (Figs 6 C, D, 7): Pygofer barrel-shaped in ventral view. Uncal lobes short, separated from each other at base, terminating in single rounded apex and with pair of narrow, slightly curved, sharply pointed finlike projections directed downward from apex and from paramedian margin of uncal lobe, in ventral view, of which the paramedian projection is long but the apical one short, about one-third as long as the paramedian one. Dorsal beak long, triangular.	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFE7936659C615659D7BFDDC.taxon	distribution	Distribution. Philippines (S. Luzon).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFE1936659C611E69D7AF8D2.taxon	materials_examined	Material examined. Type material in Lee & Hill (2010). Distribution. Philippines (E. Luzon).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFE1936659C612039D7AF9AD.taxon	materials_examined	Material examined. Type material in Lee & Hill (2010). Distribution. Philippines (E. Luzon).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFE2936559C617F59C25FD2F.taxon	materials_examined	Material examined. Type material in Lee & Hill (2010). Distribution. Philippines (E. Luzon). 28. Psithyristria nodinervis Stål, 1870	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFE2936559C617F59C25FD2F.taxon	materials_examined	Material examined. 1 male, N. W. Luzon, S. E. of Pagudpud Adams, Masasabug Bgy Bucarat, 400 m, 20 IV 2007, J. H. Lourens (IRSNB); 2 males, E. Luzon, N. Ec. Bongabon Bgy Laby, Sierra Madre, Mingan Mts., 950 m, 16 – 17 VIII 2005, J. H. Lourens (IRSNB); 1 female, same data except 26 – 27 IV 2006 (IRSNB); 1 female, N. Luzon, Kalinga-Apayao Butbut, VI – VII 1988, P. Lays (IRSNB).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFE2936559C617F59C25FD2F.taxon	distribution	Distribution. Philippines (N., N. W. and E. Luzon).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFE2936559C614609D35FC29.taxon	distribution	Distribution. Philippines (Luzon).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFE2936559C612B89B0EF80F.taxon	materials_examined	Material examined. 2 females, E. Luzon, Aurora, Sierra Madre, 15 km W. of Dibulo, 650 m, 16 ° 33.176 ’ N 122 ° 13.386 ’ E, 26 – 27 IX 2008, J. H. Lourens (IRSNB).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFE2936559C612B89B0EF80F.taxon	distribution	Distribution. Philippines (Luzon).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFE2936559C612B89B0EF80F.taxon	discussion	Remarks. This species is believed to be a new species and thus newly recorded from Luzon but could not be described due to unavailable male specimens. With the wing venation and the pigmentation on the pro- and mesonotum and the abdomen similar to those of Purana abdominalis Lee, 2009 and Purana nana Lee, 2009, this species probably belongs to the Purana abdominalis species group, which has been reported only from the Philippines (Lee 2009 b). The female specimens of this species is distinguished from the females of P. abdominalis from Negros and P. nana from Mindanao by the reduced infuscation at bases of apical cells 2 and 3 and the broad smoky membrane suffusion on apical cells of the forewing.	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFE3936459C611D59DB5F807.taxon	distribution	Distribution. Philippines (Luzon and Panay).	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFE3936459C613E09A99FAE9.taxon	distribution	Distribution. Philippines (Luzon, Samar, Palawan and Balabac Island), Malaysia, Indonesia (Java) and India.	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
03A88784FFE3936459C615459CFCFD6C.taxon	distribution	Distribution. China (southern part), Philippines (Luzon) and N. Vietnam.	en	Lee, Young June (2010): Checklist of cicadas (Hemiptera: Cicadidae) of Luzon, Philippines, with six new species and revised keys to the species of Oncotympana Stål and Psithyristria Stål. Zootaxa 2621: 1-26, DOI: 10.5281/zenodo.198106
