taxonID	type	description	language	source
03A887BAFFC4FFE72DD9FC0CC87F6060.taxon	type_taxon	Type species: Hapsidopteris diastenus Opitz, 1997: 63. By original designation.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFC4FFE72DD9FC0CC87F6060.taxon	diagnosis	Diagnosis: Specimens of this monotypic genus have an antennal club that appears to be comprised of four antennomeres (fig. 2).	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFC4FFE72DD9FC0CC87F6060.taxon	description	Description: Size: Length 5.0 mm; width 2.0 mm. Form: Oblong rectangulate; elytra about three times longer than wide, posterolateral margin gradually curved towards elytral apex. Vestiture: Body profusely vested with short setae, elytral primary (1 º) setae short, secondary (2 º) setae reclinate, usually pale and very densely distributed. Head: Cranium coarsely punctate; eyes small, moderately bulgy, finely faceted, ocular notch more angular than rounded; antenna (fig. 2) inserted at lower angle of eye incision, comprised of 10 antennomeres, loosely clubbed, scape short and robust, funicular antennomeres variously developed, 3 rd large triangular, 4 th transverse rectangulate, 5 th small transverse, 6 th very large quadrate, 7 th very small transverse, 8 th subrectangulate, 9 th triangular, and 10 th oval; labrum deeply incised; mandible robust, dentes well developed; maxillary and labial terminal palpomeres digitiform; frons wide; gular sutures strongly converging; gular pads small. Thorax: Pronotum (fig. 3) quadrate, lateral tubercle small, disc coarsely punctate, pronotal arch not evident, anterior margin slightly convex, posterior margin transverse, anterior transverse depression not evident, discal trichobothria set in depressions; lateral carina not extended to anterior margin of pronotum and posteriorly confluent with pronotal bead; procoxal cavities open; interprocoxal process linear, not laterally expanded distally; metendosternite without furcal lamina; elytra oblong rectangulate, side margins strongly deflexed, epipleuron extended to elytral apex, punctations small, grouped into distinct rows; metathoracic wing as in figure 4; mesoscutellum quadrate; tibial spur formula 0 - 1 - 1, tarsal pulvilli formula 3 - 3 - 1, anterior margin of tibia spinous; tarsal claws with large basal denticle. Abdomen: Narrows to posterior, six visible sterna; pygidium transverse, sixth visible sternum incised distally in males. Male Genitalia: Aedeagus tubular; phallobasic rod present, linear, phallic plates broad.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFC4FFE72DD9FC0CC87F6060.taxon	distribution	Distribution: This monotypic genus is known from southern México. Evolutionary relationships: It is postulated that Hapsidopteris Opitz has a sister group relationship with Opitzia Nemésio. The synapotypies that bind these genera are: eyes reduced in size, anterior third of elytral sides strongly deflected, and hind pronotal angles acute.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFC5FFE52DD9F929CF7A6721.taxon	description	Figures 2 – 4; map 2.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFC5FFE52DD9F929CF7A6721.taxon	diagnosis	Diagnosis: The extraordinarily large 6 th antennomere will distinguish these epiphloeines.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFC5FFE52DD9F929CF7A6721.taxon	description	Description: Length 5.0 mm; width 2.0 mm. Form: Oblong rectangulate. Integument: Antenna and mouthparts dark brown, frons at sides and postgena behind eyes black, frons and vertex narrowly yellow; pronotum lower sides black, pronotal disc with v-shaped black marking, legs and remainder of venter black, elytral disc yellow-brown, infuscated near epipleural margins. Head: Eyes small, moderately convex; vertex slightly wider than width of eye; coarsely punctate. Thorax: Side margins very sinuous, anterior transverse depression not evident, punctations small, but render disc coarsely sculptured; elytral side margins very deflexed, disc punctations small and organized into 12 rows. Abdomen: Aedeagus tubular. Variation: One specimen examined. Natural history: No information available.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFC5FFE52DD9F929CF7A6721.taxon	distribution	Distribution (map 2): This species is known only from southern México	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFC7FFEB2DD9F977C8E66308.taxon	type_taxon	Type species: Enoplium humerale Klug, 1842: 373. By original designation.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFC7FFEB2DD9F977C8E66308.taxon	diagnosis	Diagnosis: There are three known epiphloeine species in which specimens have 10 antennomeres, two metatarsal pulvilli, elytral punctations that are very small and profusely distributed, and the elytral disc densely vested with very short decumbent setae. Two of these species belong to Iontoclerus, the third species belongs to the exclusively West Indian genus Pericales, whose specimens may be conveniently distinguished from those of Iontoclerus by having the basal segment of the antennal club much shorter than the combined length of the funicular antennomeres (compare figures 11 and 100). MAP 2. Distribution of Katamyurus albopaniculus, Megatrachys bibara, Hapsidopteris diastenus, Opitzia chiapas, M. paniculus, K. paxillus. and M. truncatia.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFC7FFEB2DD9F977C8E66308.taxon	description	Description: Size: Length 6.0 – 9.0. mm; width 2.0 – 3.0 mm. Form: Oblong subrectangulate, elytral outer margin slightly widened at middle; elytra about three times longer than wide, posterolateral margin gradually curved towards elytral apex. Vestiture: Body profusely vested with short setae, elytral primary (1 º) setae very short, secondary (2 º) setae usually pale and very densely distributed. Head (figs. 5, 6): Cranium finely punctate; gular sutures converging, gular pads well developed (fig. 6); eyes (fig. 28) very bulgy, finely faceted, ocular notch more angular than rounded; antenna (fig. 11) inserted at lower angle of eye incision, comprised of 10 antennomeres, loosely clubbed, scape as long as combined length of pedicel and funicular antennomeres, scape curvate, funicular antennomeres (figs. 11, 27) becoming increasingly more serrate from antennomere three to seven, 8 th and 9 th antennomeres triangular, 10 th antennomere oval; labrum (fig. 13) deeply incised; tormal processes horizontal and not welded; mandible (fig. 12) robust, dentes well developed, mandibular penicillus well developed; maxillary (fig. 14) and labial (fig. 20) terminal palpomeres digitiform, laterolacinia present; frons not very wide; gular sutures strongly converging; gular pads very small. Thorax: Pronotum (figs. 7, 8) transverse, lateral tubercle not evident, disc finely punctate, pronotal arch not prominent, anterior margin slightly sinuous, posterior margin transverse, anterior transverse depression not well developed, disc of pronotum proper with shallow elevations and shallow depressions with discal trichobothria (fig. 30); trich of lateral pronotal trichobothrium particularly long (fig. 29); pronotal projection (fig. 7) slightly extended to middle; lateral carina not extended to anterior margin of pronotum and posteriorly confluent with pronotal bead; procoxal cavities open; interprocoxal process linear (fig. 7), not laterally expanded distally; metendosternite (fig. 19) without furcal lamina; elytra oblong subrectangulate, epipleural margins slightly expanded at middle, then gradually converging towards sutural margin, epipleuron extended to elytral distal four-fifths, 2 º setae profusely distributed on elytral disc (fig. 32), 1 º setae sparsely distributed on elytral disc, punctations very small and profusely distributed on elytral disc; metathoracic wing as in figure 21; mesoscutellum (fig. 25) oval; tibial spur formula 0 - 1 - 1, tarsal pulvilli formula 3 - 3 - 2, anterior margin of tibia spinous (fig. 31); tarsal claws with large basal denticle. Abdomen: Narrows to posterior, six visible sterna; pygidium transverse, sixth visible sternum incised distally in males, not incised in females. Male Genitalia: Aedeagus (fig. 16) tubular; phallobasic rod present, linear, phallic plates broad; spicular fork as in figure 15. Alimentary Canal (fig. 26): Proventriculus well developed; ventricular crypts poorly developed; four cryptonephridial malpighian tubules. Male Mesodermal Reproductive Organs (fig. 9): Two pairs of accessory glands; testis comprised of about 40 – 50 follicles. Female Mesodermal Reproductive Organs (fig. 10): Spermathecal capsule not heavily sclerotized; spermathecal gland attached to subapex of spermatheca capsule; bursa copulatrix saccular.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFC7FFEB2DD9F977C8E66308.taxon	distribution	Distribution: This South American genus ranges from French Guiana to northern Argentina. Evolutionary relationships: The following character states establish a sister group relationship between Iontoclerus Opitz and the West Indian genus Pericales, n. gen.: funicular antennomeres serrate, elytral punctations minute, and elytral disc very densely vested with short 2 º setae.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFC9FFE92DD9FD11C9F56540.taxon	description	Figures 5 – 21, 25 – 32; map 3	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFC9FFE92DD9FD11C9F56540.taxon	diagnosis	Diagnosis: These beetles may be easily distinguished from their congeners by the yellow vitta that extends posteriorly from a yellow humerus.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFC9FFE92DD9FD11C9F56540.taxon	description	Description: Length 8.0 mm; width 2.8 mm. Form: Oblong subrectangulate. Integument: Brown and yellow, cranium yellow-brown, pronotum predominantly yellow-brown on each side, femora mostly yellow, infuscated, elytra mostly brown, humeral region yellow, yellow region may extend posteriorly into slender discal line. Head: Eyes large, very convex; vertex between eyes narrow. Thorax: Side margins slightly convex, anterior transverse depression not prominent, disc with slightly elevated and depressed regions, punctations very small; elytral side margins slightly broadened at middle, then broadly rounded to apex, disc punctations very small and profusely distributed. Abdomen: Fifth visible abdominal sternite sex dimorphic (compare figures 17, 18). Aedeagus as in figure 16. Alimentary Canal (fig. 26): Proventriculus well developed; ventricular crypts poorly developed; four cryptonephridial malpighian tubules. Male Mesodermal Reproductive Organs (fig. 9): Two pairs of accessory glands, medial gland considerably shorter than lateral gland; testis comprised of about 50 follicles. Female Mesodermal Reproductive Organs (fig. 10): Spermathecal capsule not heavily sclerotized; spermathecal gland attached to subapex of spermatheca capsule; bursa copulatrix saccular. Variation: Length 7.5 – 9.0 mm; width 2.0 – 3.0 mm. The amount of yellow on the humerus and the extent of yellow streaking posteriorly from the yellow humerus varies. Natural history: Specimens have been collected throughout the year, at altitudes ranging from 7 to 330 m; some on bark, others in a Malaise trap.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFC9FFE92DD9FD11C9F56540.taxon	distribution	Distribution (map 3): The composite range for this species extends from Colombia to Argentina. I have examined 180 specimens. Colombia: Putumayo: La Paya, Cabaña La Paya, 0 ° 2 ' S, 75 ° 12 ' W, Malaise trap, 25 - X- 2001, 330 m, E. Lozano. Venezuela: 40 km E Canton Barinas, 8 - III- 1970, 7 m, S. L. Wood: Amazonas: Alto Rio Mavaca, 02 ° 02 ' 10? N 65 ° 06 ' 50? W, 5 - II- 1989, 150 m, J. Lyttke: Cojedes, El Pao Pilancones, 9 ° 43 ' 54? N 68 ° 8 ' 31? W, 12 - 16 - I- 1995, 188 m, A. Alemá. Guyana: Rupununi: Region 8 Iwokrama Forest, Pakatau Hills, 4 ° 44 ' 54? N 59 ° 1 ' 36? W, 27 - X- 2001, 70 m, on bark, downed tree, R. Brooks, Z. Falin; Iwokrama Forest, Turtle Mt. base camp, 4 ° 43 ' 5? N, 58 ° 43 ' 5? W, 1 - XI- 2001, 50 m, on bark, downed tree, R. Brooks, Z. Falin; Iwokrama Forest, 1 km W Kurupukari Iwokrama Field Stn., 4 ° 40 ' 19? N, 58 ° 41 ' 14? W, 2 - XI- 2001, 60 m, on bark, downed trees, R. Brooks, Z. Falin. Ecuador: Napo: Yasuni Res. Sta., 19 -- 30 - X- 1998, 250 m, W. J. Hanson; idem, 30 - IX- - 11 - X- 2002, 0 ° 41 ' S, 76 ° 24 ' W, 250 m, C. Brammer; idem, Garzacocha, 68 air km E. Coca, 16 - III- 1986, 210 m, McKamey. Brazil: Rio Grande do Sol: Vila Oliva, 18 - II- 1959, collector not noted: Mato Grosso: Jacare, P. N. Xingu, collection day not noted-XI- 1961, Alvarenga: Paraná: R. Branco Acre, 15 -- 20 - XI- 1961, F. M. Oliveira; Foz Do Iguacu, 10 - XII- 1966, collector not noted; Coordeuadas Sinop, collection day not noted-X- 1974, M. Alvarenga; idem, 350 m, collection day not noted-X- 1974, M. Alvarenga; idem, collection day not noted-X- 1975; idem, collection day not noted-X- 1975, Malaise trap, M. Alvarenga; idem, collection day not noted-X- 1976, M. Alvarenga: Rondonía: Ron Dovia, Br- 368, 48 km, 27 - IX- 1979, C. Biancardi; idem, 26 -- 27 - IX- 1979, Sucam; idem, 27 - IX- 1979, J. Arias; Porto Velho, Br 364, 48 km, 27 - IX- 1979, J. Arias; idem, 3 - X- 1979, J. Arias; idem, 4 - X- 1979, J. Arias; Br 364, km 48, 28 - IX- 1979, Sucam; Porto Veho, Br- 364, 48 km, 3 - X- 1979, J. Arias: Amazonas: Teffé, M de Mathan, 2 - trimestre- 1879, R. Oberthur; Villanova, Mahnel, collection day not noted-collection month not noted- 1860, collector not noted; Itapiranga, collection day not noted-II- 1934, collector not noted; idem, collection day not noted-collection month not noted- 1861, Bates; Amazonas Paraque, 30 km E. Manaus, Amazon R., 17 - II- 1981, 34 m, Chenwen Young; vic. Caucalandia, 10 ° 32 ' S, 62 ° 48 ' W, 31 - X- 1991, 160 - 350 m, J. MacDonald; idem, 28 - X- 1991, J. R. MacDonald; Ariquemes, RO Fazenda Rancho Grande, 62 km SE Ariquemes, 2 - XII- 1991, S. L. Heydon; idem, 62 km SE Ariquemes, Rancho Grande, 2 - XII- 1991, vapor & headlamp, S. L. Heydon; 13 -- 25 - IX- 1992, W. J. Hanson; idem, Fzda. Rancho Grande, 7 - X- 1993, C. W. & L. O’Brien; idem, 16 - XI- 1994, C. W. & L. O’Brien; idem, 8 -- 20 - XI- 1994, W. J. Hanson; idem, 7 -- 18 - XI- 1995, W. J. Hanson; idem, 5 -- 16 - XI- 1996, W. J. Hanson; idem, 1 -- 17 - IX- 1997, B. K. Dozier; idem, 22 -- 31 - X- 1997, W. J. Hanson; idem, 1 -- 14 - XI- 1997, B. Dozier; Pará: Ourém, Patauateua, 23 - VIII- 1992, Armadilha 1.8 m, B. Mascarenhas; Santarém, collection date not noted, collector not noted. Perú: Madre de Dios: Rio Tambopata Reserve, 20 air km SW of Puerto Maldonado, 1 - 26 - XI- 1982, 290 m, Edward S. Ross; Chambireyacú: Yurimaguas, collection day not notedcollection month not noted- 1885, Juin Aout. Bolivia: Santa Cruz: 4 - 6 k SSE Buena Vista F & F Hotel, 2 - 12 - II- 2000, J. E. Wappes. Argentina: Misiones: Sta. María, collection day not noted-X- 1944, M. J. Viana. Specimens are deposited in AMNH, BMNH, CASC, CMNH, DZUP, EMUS, FSCA, IAVH, JNRC, JEWC, MIZA, OXUM, SEMC, UCDC, WFBC, and WOPC.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFCBFFEC2DD9FBA9C9A765F8.taxon	description	Figures 22 – 24; map 3.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFCBFFEC2DD9FBA9C9A765F8.taxon	diagnosis	Diagnosis: From superficially similar specimens of I. humeralis, beetles of I. sericeus may be distinguished by the concolorous (yellow) condition of the pronotum and concolorous (dark brown) elytra.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFCBFFEC2DD9FBA9C9A765F8.taxon	description	Description: Length 7. 0 mm; width 2.2 mm. Form: Oblong subrectangulate. Integument: Cranium and pronotum yellow, elytra and legs dark brown. Head: Eyes large, very convex; vertex between eyes narrow. Thorax: Side margins slightly convex, anterior transverse depression not prominent, disc with slightly elevated and depressed regions, punctations very small; elytral side margins slightly broadened at middle, then broadly rounded to apex, disc punctations very small and profusely distributed; Abdomen: Aedeagus as in figure 24. Male Mesodermal Reproductive Organs (fig. 23): Two pairs of accessory glands; testis comprised of about 40 follicles. Female Mesodermal Reproductive Organs (fig. 22): Spermathecal capsule not heavily sclerotized; spermathecal gland attached to subapex of spermatheca capsule; bursa copulatrix saccular. Variation: Length 7.0 – 8.0 mm; width 1.8 – 2.2 mm. The amount of yellow on the humerus and the extent of yellow streaking posteriorly from the yellow humerus varies. Natural history: Specimens have been collected throughout the year at altitudes ranging from 160 – 450 m; some in a Malaise trap and at night on tree bark.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFCBFFEC2DD9FBA9C9A765F8.taxon	distribution	Distribution (map 3): The composite range for this species extends from Columbia to Bolivia. I have examined 358 specimens. Columbia: Putumayo: PNN La Paya, Cabaña La Paya, 0 ° 2 ' S 75 ° 12 ' W, 5 - 25 - XII- 2001, Malaise trap, 330 m, E. Lozano. French Guiana: Cayenne: collection date not noted, collector not noted: Ret De L’Amse: Sinnamary, 31 - VII- 1978, Degallier. Brazil: Mato Grosso: Sinop, Coordenadas, collection day not noted-X- 1974, M. Alvarenga; idem, 350 m, M. Alvarenga; idem, collection day not noted-X- 1976, M. Alvarenga: Humboldt, 10 ° 25 ' S 59 ° 28 ' W, 20 - III- 1977, 200 m, Stockwell; Diamantino, 28 - IX- 1981, MAP 3. Distribution of Opitzia apicula, Iontoclerus humeralis, I. sericeus, Turbophloeus simplex, and Pteroferus zolnerowichi. E. Furtado; idem, Fac. Sao Joao, 22 - XI- 1981, E. Furtado; idem, 28 - IX- 1981, E. Furtado; idem, collection day not noted-XI- 1981, E. Furtado; idem, collection day not noted-XII- 1981, E. Furtado; idem, Fazenda Rio Ari- nas, collection day not noted-VI- 1983, E. Furtado: Pará: Itaituba, collection date not noted, collector not noted; Tucurui, collection day not noted-I- 1979, M. Alvarenga: Amazonas: Manaus, 1 km W Taruma Falls, 19 - I- 1981, G. Ekis: Rondônia: BR- 368 kKM- 48, 27 - IX- 1979, Malaise trap, C. Biancardi; 62 km SE Ariquemes, 17 - 24 - III- 1989, 180 m, W. J. Hanson; idem, 15 - 22 - III- 1991, W. Hanson, G. Bohart; vic. Caucalandia, 10 ° 32 ' S 62 ° 48 ' W, 28 - X- 1991, 160 - 350 m, John R. MacDonald; idem, 29 - X- 1991, J. MacDonald; idem, 13 - 25 - IX- 1992, W. J. Hanson; idem, 8 - 20 - XI- 1994, W. J. Hanson; idem, 7 - 18 - XI- 1995, W. J. Hanson; idem, 5 -- 16 - XI- 1996, W. J. Hanson; idem, 22 - 31 - X- 1997, W. J. Hanson; idem, 1 -- 14 - XI- 1997, B. Dozier; idem, 1 - 17 - XI, B. K. Dozier. Ecuador: Napo: Limoncocha, 9 - VI- 1977, W. E. Steiner: Sucumbíos: E Nueva Loja, 5 km W Parahuacu Oil Station, 31 - XII- 1989, MacRae; Sacha Lodge, 5 ° S 77 ° W, 14 - 24 - III- 1994, Malaise trap, 270 m, Hibbs; 24 km F. Atahualpa, 13 - 14 - IV- 1997, 450 m, E. Giesbert, F. Hovore; Yasuni Res. Sta., 6 ° 36 ' W 0 ° 38 ' S, 19 - 30 - X- 1998, 250 m, W. J. Hanson; 28 km NE Campo Cocham, 31 - III- 2001, F. Hovore; idem, 0 ° 41 ' S, 76 ° 24 ' W, 30 - IX- 11 - X- 2002, 250 m, Malaise trap, C. Brammer. Perú: Huallaga: Chambireyacú, Yurimaguas, collection day not noted-collection month not noted- 1885 M. Mathan; Tarapoto, collection day not noted-collection month not noted- 1886, M. Mathan: Huánuco: Tingo Maria, Monzon Valley, 10 - XI- 1954, collector not noted: Madre de Dios: Tambopata, 15 km NE Pto. Maldonado, 1 - VII- 1989, 200 m, under bark, J. Ashe, R. Leschen; idem, Madre de Dios Dpto. 15 km NE Peurto Maldonado Reserva, Cuzco Amazónico, 12 ° 33 ' S 69 ° 03 ' W, 17 - VII- 1989, camp, caught by hand net, 200 m, J. S. Ashe, R. A. Leschen; idem, 19 - VII- 1989, 200 m, at night on logs, J. Ashe, R. Leschen; Mt. Alogre, Rio Pachitoa, collection date not noted, G. Tessmann. Bolivia: Cochabamba: collection day not noted-collection month not noted- 1906, collector not noted; Limbo, collection day not noted-collection month not noted- 1953, Martinez; Chaparé, 400 m, 17 - II- 1948, Zischka; idem, 25 - IX- 1949, 400 m, R. Zischka; idem, 22 - X- 1949, 400 m, R. Zischka, idem, 22 - X- 1949, 400 m, R. Zischka; idem, 5 - XI- 1949, 400 m, Zischka; idem, 15 - IX- 1950, 400 m, R. Zischka; R. Japacani, collection date not noted, J. Steinbach; Rio Madeira, collection date not noted, Mann & Baker: La Paz: Tumupasa, no other locality noted, collection day not noted-XII- 1921 - 1922, W. M. Mann; Specimens are deposited in AMNH, BMNH, CASC, CMNH, DZUP, EMUS, FSCA, IAVH, INSB, JNRC, MNHN, OXUM, SEMC, STRI, USNM, WFBC, and WOPC.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFCEFFED2DD9FB21C9906238.taxon	type_taxon	Type species: Katamyurus paxillus Opitz, 1997: 60. By original designation.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFCEFFED2DD9FB21C9906238.taxon	diagnosis	Diagnosis: Epiphloeine specimens belong to this genus if they show a narrow-oblong body form and very roughened elytral disc. The spring-like modification of the phallic apex is unique within the subfamily (figs. 53, 54).	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFCEFFED2DD9FB21C9906238.taxon	description	Description: Size: Length 4.8 – 6.3 mm; width 1.2 – 1.8 mm. Form: Oblong narrow rectangulate; elytra a little more than three times longer than wide, posterolateral margin sharply curved towards elytral apex. Vestiture: Head, pronotum, and elytral disc vested profusely with dark setae, elytral disc with a few tufts of white setae, disc vested with primary (1 º) setae and secondary (2 º) setae, 1 º setae stout, erect, and dark, 2 º setae short, decumbent, and brown or white, latter may be matted into small tufts. Head (figs. 33 – 35): Cranium rugosely punctate; eyes bulgy (fig. 33), finely faceted, ocular notch (fig. 33) angle rounded at innermost point; antenna (figs. 38, 49) inserted at lower angle of eye incision, comprised of 10 antennomeres, loosely clubbed, scape as long as combined length of pedicel and antennomeres 3 – 4, scape curvate, funicular antennomeres subfiliform, 8 th and 9 th antennomeres triangular, 10 th antennomere oval; labrum (fig. 36) deeply incised; tormal processes horizontal and not welded; mandible (fig. 39) robust, dentes well developed, mandibular penicillus very small; maxillary (fig. 42) and labial (fig. 57) terminal palpomeres digitiform, laterolacinia present; frons not very wide; gular sutures strongly converging; gular pads (fig. 34) very small. Thorax: Pronotum (figs. 40, 41, 44, 45, 48) oblong, lateral tubercle moderately developed, disc finely punctate, pronotal arch roughly punctate, anterior margin convex, posterior margin transverse, anterior transverse depression defined only at pronotal sides, disc of pronotal proper with elevations and depressions, smooth paralateral tumescences usually prominent, discal (fig. 51) and lateral (fig. 50) trichobothria prominent; pronotal projection (fig. 44) slightly extended to middle; lateral carina very brief, not extended to anterior margin of pronotum and posteriorly confluent with pronotal bead; procoxal cavities open; interprocoxal process linear (fig. 44), not laterally expanded distally; metendosternite (fig. 46) without furcal lamina; elytra narrow oblong rectangulate, epipleural margins subparallel then sharply becoming more rounded, then gradually converging towards sutural margin, epipleuron extended to elytral apex, punctations large, deeply set into disc and rendering disc subscabrous; metathoracic wing as in figures; mesoscutellum (fig. 37) quadrate; tibial spur formula 0 - 1 - 1, tarsal pulvilli formula 3 - 3 - 2, anterior margin of tibia spinous; tarsal claws with large basal denticle. Abdomen: Narrows to posterior, six visible sterna; pygidium transverse, sixth visible sternum incised distally in males, not incised in females. Male Genitalia (figs. 53 - 55): Aedeagus long and narrow; phallobasic rod present, linear, phallic plates narrow, modified at apex into spring-like extensions; spicular fork as in figure 47.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFCEFFED2DD9FB21C9906238.taxon	distribution	Distribution: The known distribution of this genus is from Guatemala to Colombia. Evolutionary relationships: Katamyurus Opitz and Ellipotoma Spinola are believed to be sister taxa on the basis of the following shared synapotypic characteristics: eyes very large, cranium reduced, body form narrow oblong, and pronotum longer than wide.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFCFFFED2DD9FC61C870678E.taxon	description	Figures 41, 48, 49, 54, 55; map 2	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFCFFFED2DD9FC61C870678E.taxon	materials_examined	Holotype: ♀. MEX. Sinaloa 14 km NE La Cap. De Taxte, 30 Sept. 1990, J. E. Wappes (FSCA) (Specimen point mounted, gender symbol affixed to paper point; support card; locality label; acronymic repository label; holotype label.). Paratypes: Fourteen specimens: México: Durango: 18.5 km SW El Salto, 3 - VIII- 1964, H. F. Howden (CNCI, 4; WOPC, 6): Sinaloa: 10 km W El Palmito, 7 - VIII- 1964, 1616 m, H. F. Howden (CNCI, 2; WOPC, 1). México: “ Colonia ” (ZMHB, 1)	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFCFFFED2DD9FC61C870678E.taxon	diagnosis	Diagnosis: The members of this species can easily be distinguished from congeners by their significantly more slender forebody. Also, K. albopaniculus specimens are black except for the white tufts of setae on the elytral disc.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFCFFFED2DD9FC61C870678E.taxon	description	Description: Length 6.7 mm; width 1.4 mm. Form: Very slender, particularly forebody (fig. 41). Integument: Black, except six small tufts of white setae on the elytral disc. Head: Eyes large and moderately convex; vertex between eyes narrow Thorax: Side margins subparallel, coarsely punctate; anterior transverse depression absent; elytra, punctations arranged into 10 rows, with six small tufts of white setae. Abdomen: Aedeagus (figs. 54, 55) tubular; phallic plates broad, apex modified into spring-like projections. Variation: Length 5.0 – 9.0 mm; width 1.0 – 2.0 mm. Natural history: Specimens were collected by beating branches of oak (Quercus), in August, at 1616 m.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFCFFFED2DD9FC61C870678E.taxon	distribution	Distribution (map 2): Known only from western México in the states of Durango and Sinaloa.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFCFFFED2DD9FC61C870678E.taxon	etymology	Etymology: The name is a Latin compound name that stems from paniculus (= tuft) and albus (= whitish). I refer to the tufts of white setae on the elytral disc.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFCFFFF12DD9F89CC8A561B8.taxon	description	Figures 33 – 40, 42 – 47, 50 – 53; map 2	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFCFFFF12DD9F89CC8A561B8.taxon	diagnosis	Diagnosis: These beetles are distinguishable from those of K. albopaniculus by the less elongate pronotum (compare figures 40, 41), and the predominantely yellow legs. In K. albopaniculus the legs are entirely black. Moreover, in K. paxillus specimens the base antennomere of the antennal club is yellow, it is black in specimens of K. albopaniculus.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFCFFFF12DD9F89CC8A561B8.taxon	description	Description: Length 5.0 mm; width 1.2 mm. Form: Slender. Integument: Mostly dark brown, scape and basal antennomere of antennal club yellow; legs yellow but femora and tibiae infuscated; elytral humeral angle yellow; elytral disc with a few tufts of yellow setae. Head: Eyes convex; vertex between eyes not particularly narrow. Thorax: Side margins not subparallel, lateral tubercle prominent, disc coarsely punctate; anterior transverse depression absent; elytra, punctations somewhat rowed, with small tufts of yellow setae. Abdomen: Aedeagus (fig. 53) tubular; phallic plates broad, apex modified into spring-like projections. Variation: Length 4.3 – 6.0 mm; width 1.3 – 1.5 mm. In one specimen, from Honduras, the elytral apex is slightly elevated and setose. The humeral yellow spot varies in expression. Natural history: Specimens were collected during March, May, June, and November. The two types were collected at 1400 m, whereas the specimen from Guatemala was collected at 950 m.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFCFFFF12DD9F89CC8A561B8.taxon	distribution	Distribution (Map 2): In addition to the type specimens I have examined five specimens from: Guatemala: Huehuetenango: 15 km N Nentón, 5 - VI- 1997, E. Giesbert & J. Monzón. Honduras: Olancho: Parque Nacional La Muralla, 24 - 27 - V- 1995, J, Huether; idem, 29 - XI- 1995, R. Turnbow; idem, 15 º 07 ' N 86 º 45 ' W, 22 - III- 1995, R. Cave. Nicaragua: Matagalpa: 10 km NW Selva Negra, 12 º 59 ' N 85 º 54 ' W, 18 - X- 2005, 1280 m, beating, W. Opitz. Specimens are deposited in FSCA, RHTC, and WOPC.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFD4FFF62DD9FF41C8D867F8.taxon	type_taxon	Type species: Megatrachys paniculus Opitz, 1997: 61. By original designation.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFD4FFF62DD9FF41C8D867F8.taxon	diagnosis	Diagnosis: The tufts of dark setae near the apex of the elytra will conveniently distinguish the members of this genus.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFD4FFF62DD9FF41C8D867F8.taxon	description	Description: Size: Length 5.0 – 8.5 mm; width 1.5 – 2.5 mm. Form: Oblong rectangulate; elytra a little more than three times longer than wide, posterolateral margin gradually curved towards elytral apex. Vestiture: Cranium matted with light setae, vertex between eyes always with one or more setal tufts, latter may also be present on frons; pronotum densely matted with light setae that may be arranged into small wisps; elytral disc vested with primary (1 º) setae and secondary (2 º) setae, 2 º setae white, dark brown or yellow, setae matted, setal tufts prominent. Head: Cranium (figs. 59 – 61) rugosely punctate; eyes very bulgy, finely faceted, ocular notch angle rounded at innermost point; antenna (fig. 63) inserted at lower angle of eye incision, comprised of 10 antennomeres, loosely clubbed, scape curvate, as long as combined length of pedicel and antennomeres 3 – 4, scape curvate, funicular antennomeres subfiliform, 7 th and 8 th antennomeres narrow triangular, 10 th antennomere oval; labrum (fig. 59) deeply incised; tormal processes horizontal and not welded; mandible robust, dentes well developed, mandibular penicillus very small; maxillary and labial (fig. 57) terminal palpomeres digitiform, laterolacinia present; frons very wide; gular sutures (fig. 60) strongly converging; gular pads very small. Thorax: Pronotum (figs. 56, 62) transverse, lateral tubercle very prominent, disc highly undulated, coarsely punctate, pronotal arch domed, anterior margin curvate, posterior margin transverse, anterior transverse depression well developed, disc paralateral tumescences usually prominent, depressed obliquely at sides where discal trichobothria are prominent; pronotal projection (fig. 62) slightly extended to middle; lateral carina not extended to anterior margin of pronotum and posteriorly confluent with pronotal bead; procoxal cavities open; interprocoxal process linear (fig. 62), not laterally expanded distally; metendosternite (fig. 70) without furcal lamina; elytra oblong rectangulate, epipleural margins subparallel then gradually becoming more rounded, then gradually converging towards sutural margin, minutely serrated or not, epipleuron extended to elytral distal four-fifth, punctations small and obscured by rugose surface of disc; metathoracic wing as in figure 133; mesoscutellum transverse (fig. 58); tibial spur formula 0 - 1 - 1, tarsal pulvilli formula 3 - 3 - 1, anterior margin of tibia spinous; tarsal claws with large basal denticle. Abdomen: Narrows to posterior, six visible sterna; pygidium transverse, sixth visible sternum incised distally in males, not incised in females. Male Genitalia (fig. 68): Aedeagus tubular; phallobasic rod present, linear, phallic plates very broad; spicular fork as in figure 67. Male Mesodermal Reproductive Organs: Two pairs of accessory glands. Female Mesodermal Reproductive Organs: Spermathecal not heavily sclerotized; spermathecal gland attached to subapex of spermathecal gland; bursa copulatrix saccular.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFD4FFF62DD9FF41C8D867F8.taxon	distribution	Distribution: The known distribution of this genus extends from Southern México to southeastern Guatemala. Evolutionary considerations: This Middle American (Opitz, 2005: 97) genus is most closely related to the North American genus Pennasolis, n. gen. with which it shares the following synapotypic characteristics: elytral disc corrugated and epipleural margin minutely serrated.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFD4FFF42DD9F92EC87B6320.taxon	materials_examined	Holotype. ♀: GUAT., ZACAPA, 2 km San Lorenzo, 5400 ', JUN 3 – 5, 1989, E. Giesbert (FSCA) (Specimen point mounted, antenna and gender symbol affixed to paper point; support card; locality label; FSCA acronymic label; holotype label.)	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFD4FFF42DD9F92EC87B6320.taxon	diagnosis	Diagnosis: These small specimens (about 4.0 mm) are conveniently distinguished from congeners by having two small black setal tufts near the elytral apex.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFD4FFF42DD9F92EC87B6320.taxon	description	Description: Length 4. 2 mm; width 1.3 mm. Form: Short oblong rectangulate. Integument: Mostly brown, cranial vertex with two brown setal tufts, legs yellow and with brown infuscations, elytra admixture of light brown and dark brown regions, elytral setae yellow, dark brown, or black, two black setal tufts near the elytral apex. Head: Eyes large, very convex; vertex between eyes broad. Thorax: Pronotal side margins with slightly-developed tubercle, anterior transverse depression not prominent, disc roughly sculptured and with coarse punctations; elytral side margins parallel to posterior four-fifths, then acutely rounded to apex; disc subcorrugated and coarsely punctate. Abdomen: Female pygidium transverse. Variation: One specimen examined. Natural history: The holotype specimen was collected in June, at 1646 m.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFD4FFF42DD9F92EC87B6320.taxon	distribution	Distribution (Map 2): Known only from the type locality.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFD4FFF42DD9F92EC87B6320.taxon	etymology	Etymology: The specific epithet is a Latin compound name formed from bi (= two) and barba (= beard). It refers to the two black setal tufts near the elytra apex.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFD6FFF42DD9FD49CBAB67B6.taxon	description	Figures 56 – 63, 67, 68, 70, 133; map 2.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFD6FFF42DD9FD49CBAB67B6.taxon	diagnosis	Diagnosis: Readily distinguishable from the superficially similar specimens of M. truncatia by large aggregate of decumbent white setae on the sides of the pronotum, the three brown setal tufts on the cranium, and the gradually rounded deflexion of the elytral preapex.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFD6FFF42DD9FD49CBAB67B6.taxon	description	Description: Length 7. 0 mm; width 1.8 mm. Form: Long oblong rectangulate. Integument: Dark brown, except pronotal side with mat of white setae; legs yellow and with brown infuscations, cranium with three small brown setal tufts, elytra with ten small brown setal tufts, elytral setae gray, brown, or yellow. Head: Cranium coarsely punctate, eyes large, very convex; vertex between eyes broad. Thorax: Pronotal side margins with well-developed tubercle, anterior transverse depression prominent, anterior not projected, disc with shallow and elevated sculpturing and with coarse punctations; elytral side margins parallel to posterior four-fifths, then acutely angled to apex; disc subcorrugated and coarsely punctuate. Abdomen: Aedeagus as in figure 68. Variation: Length 6.8 – 9.0 mm; width 2.0 – 2.5 mm. Natural history: Specimens were collected from southern México from May through July, at elevations ranging from 2287 to 2439 m.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFD6FFF42DD9FD49CBAB67B6.taxon	distribution	Distribution (Map 2): In addition to the type specimens I examined five specimens from México: Chiapas: San Cristóbal de las Casas, vicinity El Chivero, 3 - 5 - VII- 1986, 2439 m, E. Giesbert: Junction highway 199 & 190, 22 - VI- 1990, J. Huether. Specimens are deposited in CNCI, FSCA, JNRC, JPHC, WFBC, and WOPC.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFD6FFF52DD9F8E4CF4C6380.taxon	materials_examined	Holotype. ♀: MEXICO: Chiapas, 47.5 km NW Comitan, 18 Oct. 1988, R. Turnbow (FSCA) (Specimen point mounted, gender symbol affixed to paper point; support card; locality label; FSCA acronymic label; holotype label.)	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFD6FFF52DD9F8E4CF4C6380.taxon	diagnosis	Diagnosis: The abruptly deflexed condition of the elytral apex and the large brown setal tuft on the cranial vertex will distinguish these large (about 8 mm) specimens from congeners.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFD6FFF52DD9F8E4CF4C6380.taxon	description	Description: Length 7. 5 mm; width 1.8 mm. Form: Long oblong rectangulate. Integument: Dark brown; except legs yellow and with brown infuscations, cranial vertex with one large brown setal tufts, elytra with four small brown setal tufts, elytral setae gray, brown, or yellow. Head: Cranium coarsely punctate, eyes large, very convex; vertex between eyes broad. Thorax: Pronotal side margins with tubercle, anterior transverse depression prominent, anterior margin projected, disc roughly sculptured and with coarse punctations; elytral side margins parallel to posterior four-fifths, then acutely angled to apex, elytra subapical region truncated; disc subcorrugated and coarsely punctate; Abdomen: Female pygidium transverse. Variation: One specimen examined. Natural history: The holotype specimen was collected in October.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFD6FFF52DD9F8E4CF4C6380.taxon	distribution	Distribution (Map 2): Known only from southern México.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFD6FFF52DD9F8E4CF4C6380.taxon	etymology	Etymology: The trivial name is a Latin compound name formed from truncus (= cut off) and antia (= forlock). I refer to the highly setose and truncated condition of the elytral apex.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFD7FFFA2DD9FCE9C8696060.taxon	type_taxon	Type species: Arenaria chiapas Opitz, 1997: 57. By original designation.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFD7FFFA2DD9FCE9C8696060.taxon	diagnosis	Diagnosis: Epiphloeine specimens belong to this genus if the seventh antennomere is acuminate (fig. 73) and the elytral sides are strongly deflexed.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFD7FFFA2DD9FCE9C8696060.taxon	description	Description: Size: Length 7.0 – 10.0 mm; width 1.8 – 3.0 mm. Form: Oblong subrectangulate; elytra about three times longer than wide, posterolateral margin gradually curved towards elytral apex. Vestiture: Body profusely vested with short setae, elytral primary (1 º) setae very short, secondary (2 º) setae usually pale and very densely distributed. Head (figs. 64 – 66): Cranium coarsely punctate; eyes moderately bulgy, finely faceted, ocular notch more angular than rounded; antenna (fig. 73) inserted at lower angle of eye incision, comprised of 10 antennomeres, loosely clubbed, scape as long as combined length of pedicel and funicular antennomeres, scape short and stout, funicular antennomeres very transverse, 5 th and 7 th antennomeres particularly acuminate, 8 th and 9 th antennomeres rectangular, 10 th antennomere oval; labrum (fig. 79) deeply incised; tormal processes horizontal and not welded; mandible (fig. 74) robust, dentes well developed, mandibular penicillus well developed; maxillary (fig. 76) and labial (fig. 75) terminal palpomeres digitiform, laterolacinia present; frons wide; gular sutures strongly converging; gular pads small. Thorax: Pronotum (figs. 71, 72) quadrate or slightly oblong, lateral tubercle prominent, disc coarsely punctate, pronotal arch not evident, anterior margin slightly convex, posterior margin transverse, anterior transverse depression not evident, discal trichobothria set in depressions; lateral carina not extended to anterior margin of pronotum and posteriorly confluent with pronotal bead; procoxal cavities open; interprocoxal process linear, not laterally expanded distally; metendosternite (fig. 80) without furcal lamina; elytra oblong rectangulate, side margins strongly deflexed, epipleuron extended to elytral apex, punctations small, grouped into distinct rows or somewhat diffusely distributed and profusely distributed; metathoracic wing not examined; mesoscutellum quadrate; tibial spur formula 0 - 1 - 1, tarsal pulvilli formula 3 - 3 - 1, anterior margin of tibia spinous; tarsal claws with large basal denticle. Abdomen: Narrows to posterior, six visible sterna; pygidium transverse, sixth visible sternum incised distally in males, not incised in females. Male Genitalia (fig. 69): Aedeagus tubular; phallobasic rod present, linear, phallic plates broad; spicular fork (fig. 78) comprised of completely separated spicular plates. Male Mesodermal Reproductive Organs (fig. 77): Two pairs of accessory glands; testis comprised of about 12 follicles.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFD7FFFA2DD9FCE9C8696060.taxon	distribution	Distribution: One species is known from southern México, the second from eastern Bolivia. Evolutionary relationship: Within Epiphloeinae Opitzia Nemésio is most closely related to Hapsidopteris Opitz as discussed under the latter mentioned genus.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFD8FFFA2DD9FE89C9E565AE.taxon	materials_examined	Holotype: ♀. BOLIVIA, Santa Cruz: Amboro Rd. above Achira Campo 5 - 5, 800 ' 9 - 11 - Oct. 2004 Wappes & Morris. (UASC) (Specimen point mounted, antenna and gender symbol affixed to paper point; locality label; holotype label.). Paratypes: None.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFD8FFFA2DD9FE89C9E565AE.taxon	diagnosis	Diagnosis: Distinguishable from congeners by the black discal vittae of the pronotum, black streak along the elytral sutural margin, black coloration of the distal third portion of the elytral disc, and yellow antennal apex.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFD8FFFA2DD9FE89C9E565AE.taxon	description	Description: Length 7.3 mm; width 1.8 mm. Form: Oblong narrow rectangulate. Integument: Last antennomere mostly yellow, remainder of antenna, mouthparts, clypeus, lower frons, and postgena black, frons and vertex narrowly yellow; pronotum with three longitudinal bands of yellow and four of black, legs and remain- der of venter black, except femora light brown, elytra black along basal half of sutural margin, distal half of elytral disc, and along basal half of epipleural margin. Head: Eyes small, moderately convex; vertex about as wide as width of eye; coarsely punctate. Thorax: Side margins very sinuous, anterior transverse depression not evident, punctations small, but render disc coarsely sculptured; elytral side margins very deflexed, disc punctations small and organized into 13 rows that are not clearly defined, punctations seem diffusely distributed. Abdomen: Female pygidium transverse. Variation: One specimen examined. Natural history: The holotype was collected during October at 244 m.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFD8FFFA2DD9FE89C9E565AE.taxon	distribution	Distribution (map 3): This species is known only from eastern Bolivia.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFD8FFFA2DD9FE89C9E565AE.taxon	etymology	Etymology: The trivial name apicula stems from the Latin diminutive apiculus, referring to the yellow coloration on the apex of the last antennomere.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFD8FFF82DD9FAFCCBC66030.taxon	description	Figures 64 – 66, 69, 71 – 80; map 2.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFD8FFF82DD9FAFCCBC66030.taxon	diagnosis	Diagnosis: These beetles may be easily distinguished from members of its congener species by the concolorous (reddish yellow) condition of the elytral disc.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFD8FFF82DD9FAFCCBC66030.taxon	description	Description: Length 8.5 mm; width 2.0 mm. Form: Oblong rectangulate. Integument: Cranium mostly yellow, brown near antennal carina and behind eyes; pronotum dark brown along lower sides, narrowly yellow along upper sides, disc light brown, legs and abdomen brown, elytra reddish yellow, infuscated along elytral deflexion. Head: Eyes small, moderately convex; vertex between eyes broad. Thorax: Side margins very sinuous, anterior transverse depression not evident, punctations small; elytral side margins very deflexed, disc punctations small and organized into 14 rows. Abdomen: Aedeagus as in figure 69. Variation: Length 6.0 – 8.0 mm; width 1.5 – 2.5 mm. Except for size, the available specimens are quite homogeneous. Natural history: Specimens have been collected from April through July, from elevations that range from 694 to 2165 m.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFD8FFF82DD9FAFCCBC66030.taxon	distribution	Distribution (map 2): In addition to the holotype I have examined four additional specimens from: Guatemala: El Progresso, Sierra de las Minas, 19 km N Estancia de La Virgen, 18 - 29 - IV- 1990, 1829 m, E. Giesbert; Zacapa: Cerro de Mono, 8 - VI- 1997, 2275 m, E. Giesbert, J. Monzon. Honduras: Olancho: PN La Muralla, Sendero Pizote, 10 - VI- 2003, R. Turnbow. Specimens are deposited in FSCA, RHTC, and WOPC..	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFDAFFF82DD9FE51CADB66B6.taxon	type_taxon	Type species: Phyllobaenus merkeli Horn, 1896: 374. By present designation.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFDAFFF82DD9FE51CADB66B6.taxon	diagnosis	Diagnosis: The paralateral-elongated depressions on the sides of the elytral disc will distinguish the members of this genus. Additional diagnostic characteristics involve the pronotal setae, which are set on papillose elevations, the subequal antennal club antennomeres are particularly small, the head is extraordinarily transverse, and the phallobasic apodeme is very short.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFDAFFF82DD9FE51CADB66B6.taxon	description	Description: Size: Length 4.8 – 6.0 mm; width 1.7 – 2.0 mm. Form: Oblong short rectangulate; elytra little less than three times longer than wide, posterolateral margin gradually curved towards elytral apex. Vestiture: Cranium vested profusely with stout pale setae; pronotum vested with porrect stout setae; elytral disc vested with primary (1 º) setae and secondary (2 º) setae, 1 º setae stout, erect, and dark, particularly conspicuous along epipleural and sutural margins, 2 º setae short, decumbent, white or yellow. Head (figs. 82 – 84): Extraordinarily transverse; cranium finely rugosely punctate; eyes very bulgy, finely faceted, ocular notch angle acute (fig. 82) at innermost point; antenna inserted at lower angle of eye incision, antenna (fig. 86) comprised of 10 antennomeres, loosely clubbed, scape as long as combined length of pedicel and antennomeres 3 - 5, scape curvate, pedicel oval; funicular antennomeres 3 -- 6 subfiliform, 7 globose, club antennomeres, small, subequal; labrum (fig. 89) deeply incised; tormal processes horizontal and not welded; mandible (fig. 87) robust, dentes well developed, mandibular pinicillus well developed; maxillary (fig. 92) and labial (fig. 88) terminal palpomeres digitiform, laterolacinia present; frons very wide; gular sutures arcuate (fig. 83), disc strongly converging; gular pads very small. Thorax: Pronotum (figs. 81, 85) transverse, lateral tubercle not present, disc finely highly undulating, pronotal arch rough, anterior margin slightly pointed medially, posterior margin transverse, side margins minutely serrated, anterior transverse depression notable only at pronotal lateral margins, disc of pronotal proper with elevations and depressions, depressed extensively at sides where discal trichobothria are prominent; pronotal projection (fig. 81) slightly extended to middle; lateral carina extended to anterior margin of pronotum and posteriorly confluent with pronotal bead; procoxal cavities open; interprocoxal process linear (fig. 81), not laterally expanded distally; metendosternite (fig. 90) without furcal lamina; elytra short oblong rectangulate, epipleural margins subparallel then gradually becoming more rounded, then gradually converging towards sutural margin, epipleuron extended to elytral distal four-fifth, punctations profusely distributed on elytral disc; metathoracic wing as in figures; mesoscutellum subglobose; tibial spur formula 0 - 1 - 1, tarsal pulvilli formula 3 - 3 - 1, anterior margin of tibia spinous; tarsal claws with large basal denticle. Abdomen: Narrows to posterior, six visible sterna; pygidium transverse, sixth visible sternum incised distally in males, not incised in females. Male Genitalia (fig. 93): Aedeagus tubular; phallobasic rod present, linear, phallic plates very narrow; spicular fork as in figure 91. Male Internal Reproductive Organs (fig. 94): Two pairs of accessory glands.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFDAFFF82DD9FE51CADB66B6.taxon	distribution	Distribution: Known only from western USA.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFDAFFF82DD9FE51CADB66B6.taxon	etymology	Etymology: The generic name is a Latin compound name that stems from the noun penna (- wing) and the Latin noun sol (= sun), and the Latin suffix – is (= having the nature of). I refer to the light coloration on the elytral disc of the type species. Evolutionary relationships: This North American genus is most closely allied to the Middle American genus Megatrachys.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFDCFFFE2DD9FF41C90665FE.taxon	description	Figure 94 a; map 1. Phyllobaenus californicus Van Dyke, 1923: 47. Holotype: ♀. Type locality: Yosemite Val Cal V 18 1921 (CASC). (Specimen point mounted, gender label affixed to paper point; locality label; Van Dyke Collection label; Alice Riedy collector label, Type label; CASC repository label-type number 3330; Holotype label; identification label; plastic vial with abdomen and ovipositor).	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFDCFFFE2DD9FF41C90665FE.taxon	diagnosis	Diagnosis: The most convenient characteristic that distinguished the members of this species from those of P. merkeli (Horn) is the absence of profusely distributed white 2 º setae on the elytral disc. The absence of the phallobasic rod is also diagnostic for the male members of this species.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFDCFFFE2DD9FF41C90665FE.taxon	description	Description: Length 6.0 mm; width 2.0 mm. Form: Oblong short-rectangulate. Integument: Dark brown, except macula on frons and tibiae and tarsi dark brown, and elytral disc with w few pale setae. Head: Eyes not particularly large, moderately convex; vertex between eyes broad. Thorax: Side margins with slightly developed tubercles, anterior transverse depression not very prominent, disc not smoothly sculptured, with elevated and depressed regions, punctations coarse, but not easily discernable; elytral side margins parallel to posterior three-fourths, then gradually rounded to apex, side margins minutely serrate in posterior fourth, disc with several ridges, punctations small and profusely distributed. Abdomen: Aedeagus (fig. 94 a) with very short phallobasic apodeme; phallobasic rod present, phallic plates broad, apex in form of oblong knob. Variation: Length 4.0 – 5.5 mm; width 1.5 – 1.8 mm. There is variation in the number of white 2 º setae on the elytral disc. Natural history: Specimens were collected in May, June, and September, one on Juniperus occidentalis 1183 – 1220 m.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFDCFFFE2DD9FF41C90665FE.taxon	distribution	Distribution (map 1): In addition to the holotype, I have examined six specimens from: USA: California: Shasta Co., 17 - IX- 1942, on Juniper occidentalis, D. D. Leon: Yosemite National Park, 1 - VI- 1928, 1183 - 1220 m, E. O. Essig. Arizona: Coconino Co., Flagstaff, 5 - V- 1979, R. Baranowski. Specimens are deposited in EMEC, FSCA, UNSM, and WOPC.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFDCFFFD2DD9FB2CCBAB6298.taxon	description	Figures 81 – 94; map 1.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFDCFFFD2DD9FB2CCBAB6298.taxon	diagnosis	Diagnosis: The profuse distribution of white declinate 2 º degree setae on the elytral disc and the faintly visible pale fascia behind the elytral middle will distinguish the members of this species from those of P. californica (Van Dyke).	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFDCFFFD2DD9FB2CCBAB6298.taxon	description	Description: Length 5.5 mm; width 1.9 mm. Form: Oblong short-rectangulate. Integument: Black, except clypeus, mouthparts, first seven antennomeres, macula on frons, anterior corners and briefly posterior margin, legs, and abdomen light brown, elytral disc with many white setae and with pale fascia behind middle. Head: Eyes not particularly large, moderately convex; vertex between eyes broad. Thorax: Side margins with slightly developed tubercles, anterior transverse depression not very prominent, disc not smoothly sculptured, with elevated and depressed regions, punctations coarse, but not easily discernable; elytral side margins parallel to posterior three-fourths, then gradually rounded to apex, side margins minutely serrate in posterior fourth, disc with several ridges, punctations small and profusely distributed. Abdomen: Aedeagus (fig. 93) with very short phallobasic apodeme; phallobasic rod absent, phallic plates broad, apex in form of oblong knob. Variation: Length 4.8 – 6.1 mm; width 1.5 – 2.1 mm. There is variation in the number of white 2 º setae on the elytral disc. Natural history: Specimens were collected from dead wood infested with various lignicolous insects such as the cerambycid Semanotus sp. and buprestids. Some specimens were reared from branches of juniper infested with Phlaeosinus. Other plant associations involve Juniperus pachyphloea and Pinus ponderosa. Other emergence records are from November, January and February. In general, specimens were collected during April, May, June, July, August, and September, one at 1677 m. Champlain (1920: 637) reported that this species is a predator of bark beetles. Specimens were collected on Jeffery pine infested with Ips, on alligator juniper infested with Phlocosinus. Also, specimens were reared from cypress in southwestern USA.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFDCFFFD2DD9FB2CCBAB6298.taxon	distribution	Distribution (map 1): In addition to the holotype I have examined 66 specimens from: USA: Utah: Juab Co., Levan, 20 - IV- 1955; Millard Co., Oak Creek Canyon, 26 - IV- 1958, emerged from juniper, D. E. Johnson; Utah Co. Cedar Fort, emerged 20 - 23 - I- 1992, ex. Juniperus sp., R. Turnbow: New Mexico: Socorro Co. Gallinas Mountains, highway 169 (52), 5 km NW Magdalena, 27 - XII- 1989, ex. Juniperus scopulorum, wood infested with Semanotus sp. (Cerambycidae) and buprestids, L. Beierl; Grant Co. highway 15, 26 km N Silver City, 2 - VI- 1990, G. H. Nelson; Texas: Jeff Davis Co., Davis Mountains, Madera Canyon, emerged 20 - XII- 1982, ex. Juniperus sp., R. Turnbow: Colorado: La Plata Co., Hesperus, 25 - I- 1955, R. W. Dawson; Freemont Co., 11 - VIII- 1997, Juniper infested Phloeosinus, emerged January 1998, D. Leatherman, Temple, Cyn: California; Baca Co. 16 - VIII- 2002, ex. Phloeosinus, D. Leatherman, GR 53 km GR M; Montezuma Co., 10 E Cortez, emerged 26 - XII- 1995, ex. Juniperus sp. R. Turnbow; Arizona: Graham Co. Pinaleno Mountains, Wet Canyon, 5 - II- 1977, under bark & sift leaf litter, D. S. Chandler; Santa Cruz Co. Santa Rita Mountains, Madera Canyon, 17 - IX- 1968, collector not noted; Pima Co, Santa Catalina Mountains, 20 - IX- 1913, Juniperus pachyphloea, collector not noted; Coconino Co., Coconino National Forest, Oak Creek Canyon, 8 - IV- 1987, R. K. J. & Gemmill; Yavapai Co., 11 km E Ash Fork, 31 - V- 1962, on Pinus ponderosa, G. H. Nelson; idem, Williams, 31 - V- 1962, on Pinus ponderosa, G. H. Nelson; Sabino Canyon, J. N. Knull; Cochise County, Chiricahua Mountains, 5 - X- 1968, 1677 m, collector not noted. AMNH, CASC, FMNH, FSCA, JNRC, RHTC, CSUC and WOPC.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFDFFFC22DD9FB81C9536308.taxon	type_taxon	Type species: Pericales albogilvus, new species. By present designation.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFDFFFC22DD9FB81C9536308.taxon	diagnosis	Diagnosis: Within Epiphloeinae, only the members of this genus possess pronotum sculpturing comprised of whorls of shallow ridges.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFDFFFC22DD9FB81C9536308.taxon	description	Description: Size: Length 6.0 mm; width 2.0 mm. Form: Oblong rectangulate; elytra a little less than three times longer than wide, posterolateral margin gradually curved towards elytral apex. Vestiture: Cranium vested profusely with short stout yellow setae; pronotum profusely vested with short porrect yellow setae; elytral disc vested with primary (1 º) setae and secondary (2 º) setae, 1 º setae short, erect, and dark, 2 º setae short, decumbent, dark and profusely distributed. Head (fig. 95): Cranium coarsely punctate; eyes moderately bulgy, finely faceted, ocular notch angle acute at innermost point; antenna inserted at lower angle of eye incision, antenna (fig. 100) comprised of 10 antennomeres, loosely clubbed, scape short as long as combined length of pedicel and antennomeres 3, scape curvate, funicular antennomeres serrate 8 th and 9 th antennomeres triangular, 10 th antennomere oval; labrum (fig. 106) deeply incised; tormal processes horizontal and not welded; mandible (fig. 101) robust, dentes well developed, mandibular pinicillus well developed; maxillary (fig. 990 and labial (fig. 102) terminal palpomeres digitiform, laterolacinia present; frons very wide; gular sutures strongly converging, then parallel; gular pads very small. Thorax: Pronotum (figs. 96, 97) transverse, disc very concave paralaterally, lateral tubercle obscure, disc finely sculptured by shallow whorls of ridges, pronotal arch obscure, anterior margin convex, posterior margin transverse, anterior transverse depression absent, discal trichobothria prominent; pronotal projection (fig 96) slightly extended to middle; lateral carina not extended to anterior margin of pronotum and posteriorly confluent with pronotal bead; procoxal cavities open (fig. 96); interprocoxal process linear, not laterally expanded distally; metendosternite (fig. 105) without furcal lamina; elytra oblong rectangulate, epipleural margins subparallel then gradually becoming more rounded, then gradually converging towards sutural margin, epipleuron extended to elytral apex, punctations small, oval, and profusely distributed on elytral disc; metathoracic wing as in figure 98; mesoscutellum triangular (fig. 107); tibial spur formula 0 - 1 - 1, tarsal pulvilli formula 3 - 3 - 2, anterior margin of tibia spinous; tarsal claws with large basal denticle. Abdomen: Narrows to posterior, six visible sterna; pygidium transverse, sixth visible sternum incised distally in males, not incised in females. Male Genitalia: Aedeagus (fig. 104) tubular; phallobasic rod present, linear, phallic plates not very narrow; spicular fork as in figure 103.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFDFFFC22DD9FB81C9536308.taxon	distribution	Distribution: The known distribution of this genus is Haiti and the Dominican Republic, of Hispaniola.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFDFFFC22DD9FB81C9536308.taxon	etymology	Etymology: The genus name is a Greek compound name that stems from peri (= very) and kallos (= beauty). I refer to the attractiveness of the epiphloeines of this insular genus. Evolutionary relationships: Pericales, n. gen. is most closely related to Iontoclerus Opitz as discussed under the last mentioned genus.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFE0FFC22DD9FD11C93A674E.taxon	description	Figures 95 – 107; map 4.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFE0FFC22DD9FD11C93A674E.taxon	materials_examined	Holotype: ♂. HAITI: Dept. Sud-Oueste, Massif de La Selle, Morne d’ Enfer, 1850 m, 15 - V- 1984 (FSCA). (Specimen point mounted, antenna and gender symbol affixed to paper point; support card, metathoracic wing affixed to mount card; locality label; FSCA acronymic label; holotype label; plastic vial with abdomen and genitalia.). Paratypes: None	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFE0FFC22DD9FD11C93A674E.taxon	diagnosis	Diagnosis: This large epiphloeine beetle may be easily distinguished from any other epiphloeine from Hispaniola by the completely yellow pronotum and the completely dark brown coloration of the elytra. Also, the shallow whorls of ridges on the pronotal disc are diagnostic for the members of this species.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFE0FFC22DD9FD11C93A674E.taxon	description	Description: Length 6.0 mm; width 2.0 mm. Cranium dark brown in front, epicranium behind eyes yellow, antenna dark brown; pronotum yellow, setae yellow; elytra oblong long rectangulate; anterior margin of protibia with 4 spines; antenna as in figure 100. Abdomen: Aedeagus as in figure 104. Variation: Length 5.0 – 6.0 mm; width 1.5 – 2.0 mm. Natural history: Specimens have been collected in May and August, at altitudes ranging from 1250 – 1850 m.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFE0FFC22DD9FD11C93A674E.taxon	distribution	Distribution (map 4): In addition to the holotype I examined another specimen from: Dominican Republic: Pedernales: 35 km N Cabo Rojo, El Aceitillar, Las Abejas, 23 - VIII- 1988, 125 - 1430 m, M. Ivie, Philips & Johnson.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFE0FFC22DD9FD11C93A674E.taxon	etymology	Etymology: The specific epithet is a Latin compound name that stems from the adjective gilvus (= pale yellow) and the adjective albus (= white). I refer to the light coloration of the pronotum and the profusely distributed pale setae on the pronotal disc.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFE0FFC32DD9F95CCED965AA.taxon	type_taxon	Type species: Pteroferus zolnerowichi, new species. By present designation.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFE0FFC32DD9F95CCED965AA.taxon	diagnosis	Diagnosis: Specimens belong to this genus if they are characterized by having 11 antennomeres, have a short body (about 4.4 mm) and somewhat triangular body form, and have corrugations on the elytral disc.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFE0FFC32DD9F95CCED965AA.taxon	description	Description: Size: Length 4.0 – 5.0 mm; width 1.5 – 1.8 mm. Form: Oblong short triangular; elytra a little less than three times longer than wide, posterolateral margin sharply curved towards elytral apex. Vestiture: Cranium and pronotum vested profusely with yellow setae; elytral disc vested with primary (1 º) setae and sec- ondary (2 º) setae, 1 º setae stout, erect, and dark, 2 º setae short, decumbent, and yellow or brown, latter may be matted onto shallow ridges. Head (figs. 108, 109): Cranium rugosely punctate; eyes very bulgy, finely faceted, ocular notch (fig. 108) angle rounded at innermost point; antenna inserted at lower angle of eye incision, antenna (fig. 118) comprised of 11 antennomeres, loosely clubbed, scape as long as combined length of pedicel and antennomeres 3 – 5, scape curvate, funicular antennomeres subfiliform, 9 th and 10 th antennomeres narrow triangular, 11 th antennomere oval; labrum (fig. 116) deeply incised; tormal processes horizontal and not welded; mandible (fig. 114) robust, dentes well developed, mandibular penicillus well developed; maxillary (fig. 113) and labial (fig. 119) terminal palpomeres digitiform, laterolacinia present; frons very wide; gular sutures strongly converging; gular pads very small. Thorax (figs. 123 – 124): Pronotum transverse, disc highly undulating; lateral tubercle prominent, disc punctations not prominent, pronotal arch anterior margin recurved, anterior somewhat projecting, posterior margin transverse, anterior transverse depression well developed, paralateral tumescences usually prominent, depressed obliquely at sides where discal trichobothria are prominent; pronotal projection (fig. 123) slightly extended to middle; lateral carina not extended to anterior margin of pronotum and posteriorly confluent with pronotal bead; procoxal cavities open; interprocoxal process linear (fig. 123), not laterally expanded distally; metendosternite (fig. 117) without furcal lamina; elytra oblong rectangulate, epipleural margins diverging giving elytra a triangular shape, epipleuron margin minutely serrate, extended to elytral distal four-fifth, punctations small, oval, and profusely distributed on elytral disc; metathoracic wing as in figure 110; mesoscutellum triangular; tibial spur formula 0 - 1 - 1, tarsal pulvilli formula 3 - 3 - 2, anterior margin of tibia spinous; tarsal claws with large basal denticle. Abdomen: Narrows to posterior, six visible sterna; pygidium transverse, sixth visible sternum incised distally in males, not incised in females. Male Genitalia: Aedeagus (fig. 115) long and narrow; phallobasic rod present, linear, phallic plates very narrow; spicular fork as in figure 115 a.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFE0FFC32DD9F95CCED965AA.taxon	distribution	Distribution: This genus is known only from Brazil.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFE0FFC32DD9F95CCED965AA.taxon	etymology	Etymology: The generic name is a compound name that stems from the Greek pteron (= wing) and the Latin ferus (= rough). I refer to the rough sculpturing on the elytra. Evolutionary relationships: Pteroferus is most closely allied to those epiphloeine genera with 11 antennomeres that lack the basal portion of the MP 1 vein. The sister taxon of Pteroferus is unknown at this time.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFE1FFC02DD9FAF3C9096030.taxon	description	Figures 108 – 119; map 3.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFE1FFC02DD9FAF3C9096030.taxon	materials_examined	Holotype: ♀. BRAZIL: Santa Catarina: Nova Teutonia, XI- 1973, Fritz Plaumann (FSCA) (Specimen point mounted, gender symbol affixed to paper point; support card; locality label; FSCA repository label; holotype label.). Paratypes: Twelve specimens: Brazil: Santa Catarina: Nova Teutonia: collection day not noted-XI- 1973, F. Plaumann (FSCA, 7; WOPC, 1); idem, 27 º 11 ' S 52 º 23 ' W, collection day not noted-X- 1973, 300 – 500 m, F. Plaumann (FSCA, 1); idem, 27 º 11 ' S 52 º 23 ' W, collection day not noted-XI- 1966, 300 – 500 m, F. Plaumann (FSCA, 1; WOPC, 1); idem, 27 º 11 ' S 52 º 23 ' W, collection day not noted-XI- 1973, 300 – 500 m, F. Plaumann (FSCA, 1).	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFE1FFC02DD9FAF3C9096030.taxon	diagnosis	Diagnosis: The short triangular shape of the body in combination with the corrugated elytral disc will easily distinguish the members of this species from other epiphloeines.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFE1FFC02DD9FAF3C9096030.taxon	description	Description: Length 5.0 mm; width 1.8 mm. Cranium red-brown, antenna brown; pronotum periphery light brown, disc black; elytra variegated, brown with yellow streaks, setae intermixture of yellow and brown setae, surface corrugated, some regions shallowly concave and with slightly raised ridges margin; legs yellow, femora and tibiae infuscated; anterior margin of protibia with 4 spines; antenna as in figure 118. Abdomen: Aedeagus tubular; phallic apex rounded knob. Variation: Length 4.0 – 5.0 mm; width 1.5 – 1.8 mm. Also, there is some variation in the corrugated sculpturing of the elytral disc. Natural history: Specimens were collected in southern Brazil during October, November, and February, at an altitude of 300 – 500 m.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFE1FFC02DD9FAF3C9096030.taxon	distribution	Distribution (map 3): This species is known only from southern Brazil.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFE1FFC02DD9FAF3C9096030.taxon	etymology	Etymology: It gives me great pleasure to name this species after my very helpful colleague Gregory Zolnerowich, of Kansas State University.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFE2FFC62DD9FE79C9D9655C.taxon	type_taxon	Type species: Epiphlaeus simplex Schenkling, 1900: 397. By present designation.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFE2FFC62DD9FE79C9D9655C.taxon	diagnosis	Diagnosis: The corrugated-like appearance of the elytral disc, which also exhibits blotches of yellow setae, along with the oblong rectangulate body form will conveniently distinguish the members of this genus from members of other epiphloeine genera. The presence of a furcal lamina on the metendosternite is unique within Epiphloeinae.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFE2FFC62DD9FE79C9D9655C.taxon	description	Description: Size: Length 3.7 – 6.8. mm; width 1.0 – 2.0 mm. Form: Oblong rectangulate; elytra a little more than three times longer than wide, posterolateral margin gradually curved towards elytral apex. Vestiture: Cranium vested profusely with stout yellow setae; pronotum loosely matted with stout yellow setae; elytral disc vested with primary (1 º) setae and secondary (2 º) setae, 1 º setae stout, erect, and dark, particularly conspicuous along epipleural and sutural margins, 2 º setae short, decumbent, and yellow, latter may be matted into ridge-like aggregates; elytral trichobothria prominent near epipleuron. Head (figs. 120 – 122): Cranium rugosely punctate; eyes very bulgy, finely faceted, ocular notch angle acute at innermost point; antenna inserted at lower angle of eye incision, antenna (fig. 127) comprised of 11 antennomeres, loosely clubbed, scape as long as combined length of pedicel and antennomeres 3 – 5, scape curvate, funicular antennomeres 3 – 5 subfiliform, 6 – 8 slightly increasing in width, 9 th and 10 th antennomeres narrow triangular, 10 th antennomere oval; labrum (fig. 128) deeply incised; tormal processes horizontal and not welded; mandible (fig. 126) robust, dentes well developed, mandibular penicillus very small; maxillary (fig. 125) and labial (fig. 129) terminal palpomeres digitiform, laterolacinia present; frons not very wide; gular sutures strongly converging (fig. 121); gular pads very small. Thorax (figs 123 – 124): Pronotum transverse, lateral tubercle prominent, disc finely punctate, pronotal arch roughly punctate, anterior margin curvate, posterior margin transverse, anterior transverse depression well developed, disc of pronotal proper with elevations and depressions, paralateral tumescences usually prominent, depressed obliquely at sides where discal trichobothria are prominent; pronotal projection (fig. 123) slightly extended to middle; lateral carina not extended to anterior margin of pronotum and posteriorly confluent with pronotal bead; procoxal cavities open; interprocoxal process linear (fig. 123), not laterally expanded distally; metendosternite (fig. 131) with furcal lamina; elytra oblong rectangulate, epipleural margins subparallel then gradually becoming more rounded, then gradually converging towards sutural margin, epipleuron extended to elytral distal four-fifth, punctations small, oval, and profusely distributed on elytral disc, become progressively smaller to distal limit of disc, elytral trichobothria present near epipleural margin; metathoracic wing as in figures; mesoscutellum triangular; tibial spur formula 0 - 1 - 1, tarsal pulvilli formula 3 - 3 - 2, anterior margin of tibia spinous; tarsal claws with large basal denticle. Abdomen: Narrows to posterior, six visible sterna; pygidium transverse, fringed with large setae, sixth visible sternum incised distally in males, not incised in females. Male Genitalia (fig. 132): Aedeagus long and narrow; phallobasic rod present, linear, phallic plates very narrow; spicular fork as in figure 130.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFE2FFC62DD9FE79C9D9655C.taxon	distribution	Distribution: The known distribution of this genus is Colombia to Bolivia.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFE2FFC62DD9FE79C9D9655C.taxon	etymology	Etymology: The generic epithet is a compound name that stems from the Latin turba (= disorder) and the Greek phloios (= bark). I refer to the taxonomic enigma that this monotypic genus has presented. This is the only epiphloeine species known to me that has fully developed furcal laminae (fig. 131). Evolutionary considerations: Turbophloeus is most closely allied to those epiphloeine genera with 11 antennomeres that lack the basal portion of the MP 1 vein. It is not possible to predict the sister taxon at this time. MAP 4. Distribution of Pericales albogilvus.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFE4FFC72DD9FB4CC85C668F.taxon	description	Figures 1, 120 – 132; map 3.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFE4FFC72DD9FB4CC85C668F.taxon	diagnosis	Diagnosis: As in the generic section of diagnosis.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFE4FFC72DD9FB4CC85C668F.taxon	description	Description: Length 6. 0 mm; width 2.0 mm. Form: Oblong rectangulate. Integument: Mostly brown, legs yellow and with brown infuscations, elytra with admixture of light brown and dark brown regions, elytral setae yellow or dark brown. Head: Eyes large, very convex; vertex between eyes broad. Thorax: Side margins with well developed tubercles, anterior transverse depression prominent, disc not smoothly sculptured, with elevated and depressed regions, punctations coarse, but not easily discernable; elytral side margins parallel to posterior four-fifths, then sharply rounded to apex, disc subcorrugated and coarsely punctuate. Abdomen: Aedeagus (fig. 132) very long, narrow, tubular; phallobasic rod present, phallic plates very narrow, apex in form of rounded knob. Variation: Length 5.0 – 7.0 mm; width 1.5 – 2.0 mm. The intensity of contrast between the light brown and dark brown regions on the elytral disc varies. Natural history: A large series of this species was collected in a Malaise trap on tree fall by Max Barkley between November and December.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
03A887BAFFE4FFC72DD9FB4CC85C668F.taxon	distribution	Distribution (map 3): Seventy-five specimens examined. Colombia: Caquetá: Mun. San José del Fragua, Vda. La esmeralda, 1 º 20 ' N 7 º 66 ' W, 9 - 13 - IX- 2000, 1250 m, Malaise, E. Gonzáles: Norte de Santander: ANU, Los Estoraques, Qda. La Media, 8 º 13 ' N 73 º 14 ', 1 - 16 - XII- 2003, Malaise, 1480 m, J. Vargas; idem, 8 º 14 ' N 73 º 15 ' W, 29 - X- 18 - XI- 2003, Malaise, 1850 m, J. Vargas & E. Bayona; Risaralda: SFF, Otún Quimbaya Cuchilla, Camino, 04 ° 43 ' N 75 ° 35 ' W, 19 - I- 4 - II- 2003, G. López: Boyacá: SFF., Iguanaque, La Planada, 05 ° 25 ' N 73 ° 27 ' W, 21 - I- 7 - II- 2001, Malaise, 2850 m, P. Reina: SFF Iguaque, Cerro Pan de Azucar, 5 º 25 ' N 73 º 27 ' W, 18 - V- 10 - VI- 2001, 2860 m, R. Reina; idem, 12 - IX- 13 - X- 2001, 3300 m, P. Reina; idem, Quebrd Los Francos, 5 º 25 ' N 28 º 60 ' W, 7 - 24 - II- 2001, Malaise, P. Reina; Cabaña Mamarramos, 5 º 25 ' N 73 º 27 ' W, 2855 m, Malaise, 13 - 30 - VII- 2000, P. Reina. Huila: PNN Cueva de los Guácharos Alto el Mirador, 01 º 38 ' N 76 º 06 ' W, 6 - 20 - 1 - 2002, J. Urbano. Bolivia: Cochabamba: Cochabamba. Santa Cruz: Amboro National Park, Los Volcanes, 18 ° 6 ' S 63 ° 36 ' W, 20 - XI- - 12 - XII- 2004, 1000 m, Malaise trap, M. V. L. Barclay & H. Mendel. Specimens are deposited in BMNH, IAVH, USNM, and WOPC.	en	Opitz, Weston (2008): Classification, natural history, and evolution of Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nemésio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus Opitz, new genus. Zootaxa 1754: 1-40
