taxonID	type	description	language	source
03A887D05F4AFF83FF3EEDF5FDAA4C9B.taxon	description	Figs 1 A – E, 46 A, 49, 65 A – B	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F4AFF83FF3EEDF5FDAA4C9B.taxon	diagnosis	Diagnosis: This species can be recognized by the distinctive thorn-like projection on the hind trochanter (Fig. 46 A); surstyli slender and curved towards each other in dorsal view (Fig. 1 A), with a small projection at the base in lateral view (Fig. 1 D – E three small ejaculatory ducts, one with some teeth in basal half (Fig. 1 D – E); phallic guide with a downward projection before apex in lateral view (Fig. 1 D – E). Based on the shape of the surstyli and phallus, this species is related to T. sylvatica (Meigen, 1824) (Földvári & De Meyer 1999: fig. 3 A – D), T. subsylvatica Kazerani, Khaghaninia & Kehlmaier, 2017 (Kazerani et al. 2017: fig. 3 A – D) and T. littoralis (Becker, 1897) (Földvári & De Meyer 1999: fig. 2 A – D). It differs by the distinct thorn-like projection (not spines) of the hind trochanter (Fig. 46 A). Tomosvaryella sylvatica, T. subsylvatica and T. littoralis (Fig. 46 B) have several spines on the hind trochanters.	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F4AFF83FF3EEDF5FDAA4C9B.taxon	description	Description: MALE: Body length (excluding antennae): 2.85 mm. Head. Scape, pedicel dark. Flagellum brown. Eyes meeting for a distance of 6 – 7 facets. Frons silver-gray pollinose. Vertex black, lacking pollinosity. Occiput dark with scattered short black setae. Thorax. Postpronotal lobe light yellow, gray pollinose without setae along upper margin. Prescutum and scutum black, prescutum slightly shining, scutum with a row of long supra-alar setae (up to 0.11 mm) and uniseriate rows of long intra-alar setae (up to 0.09 mm). Scutellum black, light-brown pollinose, with up to 18 thin setae along posterior margin (up to 0.11 mm). Subscutellum and pleura dark brown, silver-gray pollinose. Wing. Length: 3.5 mm. LW: MWW = 3.5. Wing almost entirely covered with microtrichia. Cross-vein r-m reaching cell dm at the middle. M 1 moderately undulating in middle. Halter length: 0.37 mm. Whitish, dark at base. Legs. Dark but narrowly yellow at apex of femora and base of tibiae, gray pollinose. Mid coxa with 3 – 4 anterior setae. Hind trochanter with a distinct thorn-like projection (not spine) (Fig. 46 A). Hind femur with two rows of long setae on anterior and two rows on posterior side. Hind femur with 1 – 2 wrinkled indentations in basal quarter. Tibiae with two rows of short setae on anterior and three rows on posterior side. Hind tibia with 1 – 2 wrinkled indentations in the middle. Tarsi light brown, gray pollinose. Claws white in basal half. Abdomen. Ground color dark brown. Tergite 1 entirely gray pollinose with 8 – 9 brown long lateral setae (up to 0.16 mm). Tergites 2 – 5 with scattered yellowish setae (up to 0.08 mm). Sternites light brown laterally and dark brown centrally, gray pollinose. Syntergosternite 8 with scattered long dark setae. Genitalia. Genital capsule in dorsal view: epandrium and surstyli brown, gray pollinose. Epandrium wider than long (MLE: MWE = 0.72). Surstyli rather symmetrical, elongated and slender, slightly curved to each other at apical third (Fig. 1 A). Genital capsule in ventral view: gonopods small and wide, equal in height, hypandrium broad, subepandrial sclerite wide basically (Fig. 1 B). Genital capsule in lateral view: both surstyli gently curved to sternite, with a small projection in basal towards hypandrium (Figs 1 D – E). Phallic guide long, pointed apically with a small projection pointed towards hypandrium (Fig. 1 D – E); phallus with three small ejaculatory ducts, one bearing some saw-like teeth in basal half (Fig. 1 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 1 C). FEMALE: unknown.	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F4AFF83FF3EEDF5FDAA4C9B.taxon	etymology	Etymology: The specific name is derived from the Latin word ‘ acantha’, meaning thorn, referring to thorn-like projection on its hind trochanter. Specimen examined: HOLOTYPE: ISRAEL: Har Hermon, 33 ° 18 ’ N, 35 ° 46 ’ E, 1500 – 1600 m a. s. l., 6. vi. 2002, L. Friedman, JSS 51689 (1 ♂, TAU).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F4AFF83FF3EEDF5FDAA4C9B.taxon	distribution	Distribution: Israel (Fig. 49). Notes: Based on the DNA barcode, this species is genetically most similar to T. sylvatica (4.6 % pairwise divergence) (Supplementary Table 1).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F48FF85FF3EE8BBFBC54D50.taxon	description	Figs 2 A – E, 27 A, 49, 65 C – D	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F48FF85FF3EE8BBFBC54D50.taxon	diagnosis	Diagnosis: This species can be recognized by the shape of surstyli, which are rectangular-shaped and flatted apically in dorsal view (Fig. 2 A), and almost straight in lateral view (Fig. 2 D – E); right gonopod with a long projection in ventral view (Fig. 2 B); wide subepandrial sclerite basally in ventral view (Fig. 2 B); wide gonopods in ventral view (Fig. 2 B); three small ejaculatory ducts, one with some teeth in basal half (Fig. 2 D). Based on genitalia characters, this species is close to T. susa sp. nov. and T. helwanensis (Collin, 1949). It differs from T. susa (Fig. 40 A ‒ D) in the different shape of the surstyli at the apex in dorsal view, flattened (not rounded) and from T. helwanensis (Kehlmaier et al. 2019: Fig. 8) in the different shape of the surstyli in dorsal view, rectangular shaped (Fig. 2 A); with a broad and short phallic guide in lateral view (Figs 2 D ‒ E).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F48FF85FF3EE8BBFBC54D50.taxon	description	Description: MALE: Body length (excluding antennae): 2.8 mm. Head. Scape dark, pedicel dark brown, flagellum light brown; pedicel with three short upper setae; flagellum tapering (LF: WF = 4.0); arista brown with a dark spot near the base. Eyes meeting for a distance of 5 – 7 facets. Frons silver-gray pollinose. Vertex black, lacking pollinosity. Occiput dark and gray pollinose. Thorax. Postpronotal lobe light yellow, gray pollinose with 2 – 4 small setae along the upper margin (up to 0.04 mm). Prescutum and scutum black, with a row of supra-alar setae and uniseriate rows of intra-alar setae. Scutellum black, silver-gray pollinose, with about 14 – 18 thin setae along posterior margin (up to 0.08 mm). Subscutellum and pleura dark brown. Wing. Length: 2.1 – 2.4 mm. LW: MWW = 2.5. Wing almost entirely covered with microtrichia. Cross-vein r-m reaching cell dm at the middle. M 1 strongly undulating in middle. Halter length: 0.4 mm. Whitish, narrowly light brown at base. Legs. Dark brown but narrowly yellow at apex of femora, tibiae and tarsi. Trochanters and Femora partly gray pollinose. Trochanters smooth. Mid and hind femora bearing two rows of dark, peg-like anteroventral spines in apical one third. Fore, mid and hind femora with 1 – 2 wrinkled indentations in basal quarter. Tibiae with two rows of short setae on anterior and three rows on posterior side. Hind tibia with 1 – 2 wrinkled indentations in the middle. Tarsi yellowish, gray pollinose. Pulvilli shorter than distitarsi. Claws white with black tips. Abdomen. Ground color dark brown. Tergite 1 entirely gray pollinose with 12 – 16 yellowish long lateral setae (up to 0.2 mm). Tergites 1 – 5 with scattered yellowish setae (up to 0.08 mm). Sternites light brown laterally and dark brown centrally, gray pollinose. Syntergosternite 8 dark, gray pollinose. Viewed laterally, higher than long (LS 8: HS 8 = 0.8). Membranous area absent. Genitalia. Genital capsule in dorsal view: epandrium and surstyli brown, gray pollinose. Epandrium wider than long (MLE: MWE = 0.8). Surstyli rather symmetrical, rectangular-shaped, slightly curved to each other at apex, rather flatted apically (Fig. 2 A). Genital capsule in ventral view: gonopods wide, large and slightly equal in height, right wider than left one with elongated projection, subepandrial sclerite wide basally, phallic guide strong, slightly straight (Fig. 2 B). Genital capsule in lateral view: surstyli longer than wide, slightly straight, rounded at apex (Figs 2 D – E). Phallic guide strong, pointed apically (Fig. 2 D); phallus with three small ejaculatory ducts, one bearing some saw-like teeth in basal half (Fig. 2 D). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 2 C). FEMALE: Body length (excluding antennae): 3.2 mm. Head. Eyes separated; with enlarged frontal facets. Frons gray pollinose in lower half. Occiput gray pollinose with scattered light brown setae near ocelli. Thorax. As in male. Wing. Length: 2.75 mm. LW: MWW = 2.08. Legs. As in males except for longer setae on distitarsi. Abdomen. As in males. Ovipositor. Base of ovipositor dark brown with some gray pollinosity. Viewed laterally (Fig. 27 A), piercer straight. LP: LB = 1.6. LDP: LPP = 1.9.	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F48FF85FF3EE8BBFBC54D50.taxon	etymology	Etymology: The specific epithet ampliasa is derived from the Latin “ amplias ”, which means widened or extended, referring to the extended gonopods. Specimens examined: HOLOTYPE: ISRAEL: Hofit, 32 ° 23 ’ N, 34 ° 52 ’ E, 21. ix. 1994, F. Kaplan & A. Freidberg, JSS 51800 (1 ♂, TAU). PARATYPE: ISRAEL: Park haYarden, 32 ° 54 ’ N, 35 ° 37 ’ E, 14. vi. 1996, A. Freidberg, JSS 43959 (1 ♀, TAU).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F48FF85FF3EE8BBFBC54D50.taxon	distribution	Distribution: Israel (Fig. 49) Note: DNA barcodes of this species are closest to T. susa sp. nov., differing by 0.8 % pairwise divergence. Male and female were both sequenced and are genetically identical (Supplementary Table 1).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F4EFF85FF3EE9F0FDAA48A4.taxon	description	Figs 3 A ‒ D, 49, 65 E – F	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F4EFF85FF3EE9F0FDAA48A4.taxon	diagnosis	Diagnosis: This species can be recognized by the shape of surstyli, triangular-shaped in dorsal view (Fig. 3 A), broadened at the middle in lateral view (Fig. 3 C ‒ D); right gonopod with a small finger-like projection in ventral view (Fig. 3 B); three long ejaculatory ducts, two with small teeth in basal third in lateral view (Fig. 3 C ‒ D) (similar to those of T. trichotibialis De Meyer, 1995).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F4EFF85FF3EE9F0FDAA48A4.taxon	description	Description: MALE: Body length (excluding antennae): 4.1 mm. Head. Scape, pedicel and arista brown. Scape and pedicel without setae; flagellum light brown, gray pollinose and long tapering, distinctly paler than pedicel (LF: WF = 5.0). Eyes meeting for a distance of 14 ‒ 15 facets. Frons silver-gray pollinose; vertex shining; occiput lacking pollinosity in upper third with scattered light brown setae. Thorax. Postpronotal lobe light brown with pale setae along the upper margin (up to 0.03 mm). Prescutum and scutum black, silver-gray pollinose; supra-alar area dark brown and shining, scutum with scattered setae at anterior supra-alar area. Scutellum black, silver-gray pollinose, without setae along posterior margin. Subscutellum black, pleura dark brown. Wing. Wing length: 3.7 mm. LW: MWW = 7.5. Wing almost entirely covered with microtrichia. M 1 strongly undulating in middle. Halter length: 0.42 mm. Brown to yellowish, narrowly black at base. Legs. Coxae and trochanters black, gray pollinose, hind trochanter with scattered short pale setae. Femora dark brown with narrow yellow apices, silver pollinose and 5 ‒ 8 ventral spines toward apex. All femora with one wrinkled indentation in basal. Tibiae light brown at base, gray pollinose, with two rows of short black setae on anterior and three rows on posterior side. Hind tibia with two wrinkled indentations in middle. Tibiae without distal spines. Tarsi dark. Abdomen. Ground color dark brown, tergite 1 silver pollinose, tergite 2 with weak silver pollinose on anterior corner, tergites 2 ‒ 5 with scattered brown setae. Sternites white-yellow laterally with dark mid-line centrally, gray pollinose. Genitalia. Genital capsule in dorsal view: epandrium and surstyli light brown, epandrium longer than wide (MLE: MWE = 0.5). Surstyli almost symmetrical, both triangular-shaped and longer than wide (Fig. 3 A). Genital capsule in ventral view: gonopods equal in height, right with a small finger-like projection; subepandrial sclerite rectangular-shaped (Fig. 3 B). Genital capsule in lateral view: both surstyli broadened medially, slightly straight ventrally (Fig. 3 C ‒ D); phallus trifid, with three long straight ejaculatory ducts, two with small teeth in basal third (Fig. 3 C ‒ D); phallic guide small, pointed, slightly bent toward surstyli (Fig. 3 C ‒ D). FEMALE: unknown.	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F4EFF85FF3EE9F0FDAA48A4.taxon	etymology	Etymology: The specific epithet refers to Anahita, the old Persian form of the name of an Iranian goddess who rules water, crops and fertility. Specimen examined: HOLOTYPE: IRAN: Kermanshah: Sarpolezahab, 34 ° 28 ’ N, 45 ° 49 ’ E, 546 m a. s. l., 1. vii. 2015, M. Zardouei, Malaise trap, JSS 51946 (1 ♂, CNC).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F4EFF85FF3EE9F0FDAA48A4.taxon	distribution	Distribution: Iran (Fig. 49). Note: Based on DNA barcoding, this species is closest to T. hamata, differing by 2.5 ‒ 3.1 % (pairwise diver- gence) (Supplementary Table 1).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F4FFF86FF3EEDB5FAA64E55.taxon	description	Figs 4 A ‒ E, 50	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F4FFF86FF3EEDB5FAA64E55.taxon	diagnosis	Diagnosis: This species can be recognized by the shape of surstyli, both apically knobbed in dorsal view (Fig. 4 A) and bent strictly towards the abdominal sternites in lateral view (Fig. 4 D ‒ E); phallus trifid and unequal, one with tooth-like projections in basal half (Fig. 4 D ‒ E); phallic guide short and broad (Fig. 4 D ‒ E); gonopods slightly unequal in height, left is slightly higher than right (Fig. 4 B). The female is described and illustrated in Majnon-Jahromi et al. (2017 b). Specimens examined: IRAN: Sistan & Baluchestan: Saravan, 27 ° 25 ’ N, 62 ° 17 ’ E, 12. v. 2013, F. Hamzavi, Malaise trap, JSS 51917 (1 ♂, CNC); Kermanshah: Sarpolezahab, 34 ° 28 ’ N, 45 ° 49 ’ E, 456 m a. s. l., 14. v. 2016, M. Zardouei, Malaise trap, JSS 52190 (1 ♂, CNC); Khuzestan: Shush, 32 ° 06 ’ N, 48 ° 26 ’ E, 68 m a. s. l., 11. iii – 10. v. 2015, E. Gilasian, Malaise trap, JSS 52192 (1 ♀, CNC); UNITED ARAB EMIRATES: Barqa Al Suqoor Protected Area, 23 ° 59 ’ N, 52 ° 40 ’ E, 29. iii. 2018, A. Saji & A. van Harten, Malaise trap, JSS 52275 (1 ♂, CNC).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F4FFF86FF3EEDB5FAA64E55.taxon	distribution	Distribution: Iran, United Arab Emirates (Skevington 2020) (Fig. 50). Note: Based on DNA barcoding, this species is most similar to T. pseudophanes (Perkins, 1905), differing by 8.3 % (pairwise divergence). Intraspecific variation ranges from 0.1 to 3.2 % (Supplementary Table 1).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F4DFF86FF3EEAF4FC844C6D.taxon	description	Fig. 50	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F4DFF86FF3EEAF4FC844C6D.taxon	diagnosis	Diagnosis: Following De Meyer (1995), this species can be recognized by the shape of surstyli, broadened at base and gradually tapering toward apices in dorsal view, both surstyli slightly curved towards abdominal sternites in lateral view (De Meyer, 1995: fig. 9 a); epandrium longer than wide, phallic guide slender and long in lateral view, hind femur ventrally with long pale setae (De Meyer, 1995: fig. 9 b). Based on the shape of surstyli in dorsal view, this species is similar to T. pistacia Majnon-Jahromi & Kehlmaier in Majnon-Jahromi et al. 2017 b (Fig. 32 A) and T. argyratoides De Meyer, 1995 (Fig. 5 A). It differs from both species by having long pale setae on the hind femur. Moreover, Tomosvaryella pistacia has wider epandrium and its surstyli are more angulate in lateral view (Fig. 32 A, D, E).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F4DFF86FF3EEAF4FC844C6D.taxon	distribution	Distribution: Iran, Israel (Skevington 2020) (Fig. 50).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F4DFF86FF3EE8E1FD024912.taxon	description	Figs 5 A – D, 50	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F4DFF86FF3EE8E1FD024912.taxon	diagnosis	Diagnosis: This species can be recognized by the shape of surstyli, broadened at the base and gradually tapering toward apices in dorsal view, both surstyli not curved to the abdominal sternites in lateral view (Fig. 5 A); epandrium longer than wide (Fig. 5 A); gonopods equal, triangular-shaped with a small projection bent towards phallic guide in ventral view (Fig. 5 B). This species is closely related to T. argyrata and T. pistacia. See diagnosis of T. argyrata for more details. Specimens examined: IRAN: South Khorasan: Mohammadieh, 32 ° 52 ’ N, 59 ° 01 ’ E, 1419 m a. s. l., 5 – 23. ix. 2016, B. Motamedinia, Malaise trap, JSS 52026 (1 ♂, CNC); ISRAEL: Arava Valley, nr Hazeva, Shizaf Nature Res. side channel of Waddi Shahak, 30 ° 45 ’ N, 35 ° 15 ’ E, - 116 m a. s. l., 16. iv. 1995, M. E. Irwin, Malaise trap, JSS 52059 (1 ♂, TAU); Har Horesha, 30 ° 34 ’ N, 34 ° 31 ’ E, 6. iv. 2005, A. Freidberg, JSS 51799 (1 ♂, TAU); Jeruham, 30 ° 59 ’ N, 34 ° 54 ’ E, 26. iv. 1973, A. Freidberg, JSS 51750 (1 ♂, TAU); Nahal Oren, 32 ° 43 ’ N, 35 ° 01 ’ E, 2. vi. 2003, A. Freidberg, JSS 51785 (1 ♂, TAU); Sede Boqer, 30 ° 52 ’ N, 34 ° 47 ’ E, 30. iii. 2004, L. Friedman, JSS 51796, JSS 51797 (2 ♂, CNC, TAU); Se- dom, 31 ° 05 ’ N, 35 ° 22 ’ E, 13. iii. 1994, A. Freidberg, JSS 51702 (1 ♂, TAU).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F4DFF86FF3EE8E1FD024912.taxon	distribution	Distribution: Iran, Israel, Spain (Skevington 2020) (Fig. 50). Note: Based on DNA barcodes, this species is most similar to T. gazliensis Kuznetzov, 1994, differing by 5.05 % (pairwise divergence) (Supplementary Table 1).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F4DFF9BFF3EED31FBE34C09.taxon	description	Figs 6 A – F, 51, 66 A – B	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F4DFF9BFF3EED31FBE34C09.taxon	diagnosis	Diagnosis: This species can be recognized by the shape of surstyli in dorsal view, broad at base and elongated; both surstyli bent toward sternites in lateral view (Fig. 6 A); phallus with three ejaculatory ducts, one with three spines in lateral view (Fig. 6 F). Other species that have such distinct spines are T. israelensis De Meyer, 1995 (Fig. 20 D ‒ E), T. jubata De Meyer, 1995 (Fig. 221 D), T. minuscula (Collin, 1956) (Földvári & De Meyer, 1999: fig. 13 E – F) and T. sedomensis De Meyer, 1995 (De Meyer, 1995: fig. 21 f). On the basis of genitalia characters, this species is close to Tomosvaryella cilifemorata (Becker, 1907) (Földvári & De Meyer, 1999: fig. 11 A – E; Kehlmaier et al. 2019: fig. 6 A) and Tomosvaryella mutata (Földvári & De Meyer, 1999: fig. 15 A – D). It differs by the shape of surstyli in lateral view and having three distinct spines on one of ejaculatory ducts (Figs 6 F).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F4DFF9BFF3EED31FBE34C09.taxon	description	Description: MALE: Body length (excluding antennae): 3.6 mm (n = 5). Head. Scape, pedicel, arista dark. Pedicel with a pair of dorsal setae; flagellum light brown, gray pollinose and tapering, distinctly paler than pedicel (LF: WF = 3.0). Eyes meeting for a distance of seven facets. Frons silver-gray pollinose. Occiput dark, gray pollinose. Thorax. Postpronotal lobe light brown with 2 – 3 pale setae along the upper margin (up to 0.03 mm). Prescutum and scutum black, prescutum silver-gray pollinose and scutum light brown pollinose, with scattered setae at anterior supra-alar area and uniseriate rows of intra-alar setae. Scutellum black, silver-gray pollinose, with 1 – 2 thin short setae along posterior margin (up to 0.03 mm). Subscutellum and pleura black. Wing. Wing length: 2.5 mm., LW: MWW = 2.7. Wing almost entirely covered with microtrichia. M 1 strongly undulating in middle. Halter length: 0.33 mm. Brown to yellowish, narrowly black at base. Legs. Coxae black, trochanters dark brown, smooth. Femora dark brown with narrow yellow apices, silver pollinose. Mid and hind femora with one or two wrinkled indentations in basal. Tibiae brown at base, gray pollinose, with two rows of short brown setae on anterior and three rows on posterior side. Tibiae without distal spines, mid and hind tibiae with one or two wrinkled indentations in middle. Tarsi light brown, distitarsi brown. Abdomen. Ground color dark, tergite 1 silver gray pollinose, tergites 2 ‒ 5 brown pollinose with scattered short dark setae. Tergite 5 asymmetrical, T 5 R: T 5 L = 1.2. Membranous area present. Sternites brown laterally and dark centrally, gray pollinose. Genitalia. Genital capsule in dorsal view: epandrium and surstyli brown, gray pollinose. Surstyli elongated, almost symmetrical, with a broad base and inward bent tips, left surstylus slightly longer (Fig. 6 A). Genital capsule in ventral view: gonopods equal in height, right slightly broader than left one (Fig. 6 B); genital capsule in lateral view: both surstyli broad at base and bent toward sternites (Fig. 6 D ‒ E); phallus with three ejaculatory ducts, one with three long spines towards phallic guide (Fig. 6 F); ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 6 C). FEMALE: unknown.	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F4DFF9BFF3EED31FBE34C09.taxon	etymology	Etymology: The specific epithet refers to Bistoun, an archaeological site in Kermanshah from where the specimens have been collected. Specimens examined: HOLOTYPE: IRAN: Kermanshah: Ghazanchi, 34 ° 26 ’ N, 47 ° 00 ’ E, 1304 m a. s. l., 15. vi. 2015, M. Zardouei, Malaise trap, JSS 51939 (1 ♂, CNC). PARATYPES: IRAN: Kermanshah: Dodan, 35 ° 00 ’ N, 46 ° 12 ’ E, 1011 m a. s. l., Garden, 22. vii. 2016, M. Zardouei, Malaise trap, JSS 52194 (1 ♂, CNC); Ghazanchi, 34 ° 26 ’ N, 47 ° 00 ’ E, 1304 m a. s. l., 15. vi. 2015, M. Zardouei, Malaise trap, JSS 51935, JSS 51940 (2 ♂, TAU, CNC); Gheshlagh, 34 ° 56 ’ N, 46 ° 27 ’ E, 1533 m a. s. l., 1. vi. 2015, M. Zardouei, Malaise trap, JSS 51950 (1 ♂, CNC).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F4DFF9BFF3EED31FBE34C09.taxon	distribution	Distribution: Iran (Fig. 51). Notes: Based on DNA barcodes, this species is closest to T. minuscula (Collin, 1956), differing by 0.9 % (pairwise divergence) (Supplementary Table 1). It is possible that these are a single species with polymorphic genitalia, so future genetic and ecological work are required to confirm their taxonomic status.	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F50FF9BFF3EE83DFB154B8A.taxon	description	Fig. 49	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F50FF9BFF3EE83DFB154B8A.taxon	diagnosis	Diagnosis: Following Földvári & De Meyer (1999), this species can be recognized by the elongated surstyli in dorsal view, broadened at base, both surstyli normally curved in lateral view; epandrium longer than wide; phallus with three ejaculatory ducts, one with saw-like teeth; long hair on hind femur; black halter (Földvári & De Meyer, 1999: fig. 11 A – E; Kehlmaier et al. 2019: fig. 6 A).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F50FF9BFF3EE83DFB154B8A.taxon	distribution	Distribution: Austria, Bulgaria, Cyprus, France, Germany, Greece, Hungary, Italy, Morocco, Spain, Switzerland, Tunisia (Földvári & De Meyer, 1999; Kehlmaier et al. 2019; Skevington 2020) (Fig. 49).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F50FF9BFF3EEE49FD9148C1.taxon	description	Figs 7 A – E, 52	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F50FF9BFF3EEE49FD9148C1.taxon	diagnosis	Diagnosis: This species can be recognized by the shape of epandrium in dorsal view, divided into two parts by an epandrial groove (Fig. 7 A); broad subepandrial sclerite in ventral view (Fig. 7 B); phallus trifid, with three wavy ejaculatory ducts (Fig. 7 D – E); small and slender phallic guide in lateral view (Fig. 7 E); gonopods equal in height (Fig. 7 B). Specimens examined: IRAN: Kermanshah: Ghazanchi, 34 ° 26 ’ N, 47 ° 00 ’ E, 1304 m a. s. l., 1. vii. 2015, M. Zardouei, Malaise trap, JSS 51954 (1 ♂, CNC); North Khorasan: Gelian, 37 ° 15 ’ N, 57 ° 55 ’ E, 1283 m a. s. l., 8 – 22. vii. 2016, B. Motamedinia, Malaise trap, JSS 51977 – 82, JSS 51984 – 85, JSS 51988 (9 ♂, CNC); Darkesh, 37 ° 26 ’ N, 56 ° 44 ’ E, 23. vii. 2016, B. Motamedinia, sweep net, JSS 52036 (1 ♀, CNC); Razavi Khorasan: Sheikhha, 36 ° 36 ’ N, 59 ° 11 ’ E, 1169 m a. s. l., 22. v – 6. vi. 2016, B. Motamedinia, Malaise trap, JSS 51864 (1 ♂, CNC); Torqabeh, 36 ° 18 ’ N, 59 ° 22 ’ E, 1296 m a. s. l., 25. vi – 9. vii. 2016, B. Motamedinia, Malaise trap, JSS 51850, JSS 51858 – 9 (3 ♂, CNC, TAU, USNM); Sistan & Baluchestan: Zabol, Nimrooz, 31 ° 12 ’ N, 61 ° 20 ’ E, 5. vi. 2016, H. Derafshan, sweep net, JSS 51844 (1 ♂, CNC).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F50FF9BFF3EEE49FD9148C1.taxon	distribution	Distribution: Bulgaria, Canada, Czech Republic, Egypt, Finland, France, Greece, Hungary, Iran, Latvia, Portugal, Romania, Slovakia, Spain, Switzerland, Ukraine, United States (Skevington 2020) (Fig. 52). Note: DNA barcodes of this species are genetically most similar to T. parakuthyi, differing by 10.3 % (pairwise divergence) (Supplementary Table 1).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F51FF9CFF3EEC76FD844B69.taxon	description	Figs 8 A – E, 50, 66 C – D	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F51FF9CFF3EEC76FD844B69.taxon	diagnosis	Diagnosis: This species can be recognized by the shape of surstyli in lateral view, elongated and straight (Fig. 8 D – E); epandrium longer than wide in dorsal view (Fig. 8 A); left gonopod higher than right one in ventral view (Fig. 8 B). Based on the shape of surstyli in dorsal view, this species resembles species in the kuthyi group (Tomosvaryella freidbergi De Meyer, 1995, T. hispanica De Meyer, 1997, T. israelensis De Meyer, 1995, T. kuthyi Aczél, 1944, T. parakuthyi De Meyer, 1995). It differs by the straight surstyli in lateral view (Fig. 8 D – E).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F51FF9CFF3EEC76FD844B69.taxon	description	Description: MALE: Body length (excluding antennae): 2.5 mm. Head. Scape, pedicel and arista dark brown, flagellum light brown. Pedicel with three short, pale upper setae; flagellum tapering and gray white pollinose (LF: WF = 2.6). Eyes meeting for a distance of 5 – 6 facets. Frons dark silver-gray pollinose. Vertex black, lacking pollinosity. Occiput dark and gray pollinose. Thorax. Postpronotal lobe light yellow, gray pollinose with 6 – 7 yellowish setae along the upper margin (up to 0.08 mm). Prescutum, scutum, scutellum black. Scutum with two uniseriate rows of conspicuous intra-alar setae and two uniseriate rows of dorsocentral setae. Postalar callus with two dark setae. Scutellum gray pollinose without setae along lateral margin. Subscutellum black, gray pollinose. Pleura dark, gray pollinose. Wing. Length: 2.2 mm. LW: MWW = 3.05. Wing almost entirely covered with microtrichia. Crossvein r-m reaches dm at the middle. M 1 moderately undulating in middle. Halter length: 0.37 mm. Whitish, brown at base. Legs. Coxae dark, gray pollinose. Mid coxa with three dark anterior setae. Trochanters dark, gray pollinose, smooth. Femora dark brown, gray pollinose. All femora with 1 – 2 wrinkled indentations at base. Tibiae brown but light brown in basal half, with two ventral rows of short setae on anterior and three rows on posterior side. Hind tibia with two wrinkled indentations in middle. Tarsi brown with scattered brown setae at anterior margin. Pulvilli yellow. Claws light brown with black tips. Abdomen. Ground color dark brown. Tergite 1 densely gray pollinose, with 6 – 8 brown lateral setae (up to 0.13 mm). Tergites 2 densely gray pollinose in anterior margin. Genitalia. Genital capsule in dorsal view: epandrium and surstyli light brown, gray pollinose. Epandrium longer than wide (MLE: MWE = 1.33). Surstyli rather symmetrical. Both surstyli elongated, left longer than right one (Fig. 8 A). Both surstyli bent towards each other apically (Fig. 8 A). Genital capsule in ventral view: gonopods unequal in height, left is higher than right one (Fig. 8 B). Subepandrial sclerite elongated (Fig. 8 B). Genital capsule in lateral view: both surstyli long and straight, slightly broader at base (Fig. 8 D – E). Phallic guide strong with distinct patch of setae dorsally before apex (Fig. 8 D – E), phallus with three small ejaculatory ducts, one with some small teeth in basal half (Fig. 8 B). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 8 C). FEMALE: unknown.	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F51FF9CFF3EEC76FD844B69.taxon	etymology	Etymology: The specific epithet refers to Cyprus, from where the specimens have been collected. Specimens examined: HOLOTYPE: CYPRUS: Kyrenia, 35 ° 21 ’ N, 33 ° 09 ’ E, 30. vii – 13. viii. 2017, O. Ozden, Malaise trap, JSS 52302 (1 ♂, CNC). PARATYPE: CYPRUS: Kyrenia, 35 ° 21 ’ N, 33 ° 09 ’ E, 10 – 17. ix. 2017, O. Ozden, Malaise trap, JSS 52349 (1 ♂, CNC).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F51FF9CFF3EEC76FD844B69.taxon	distribution	Distribution: Cyprus (Fig. 50).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F57FF9CFF3EEFE8FD9149A8.taxon	description	Figs 9 A – D, 48 A, 52	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F57FF9CFF3EEFE8FD9149A8.taxon	diagnosis	Diagnosis: This species can be recognized by the horn-like projection in middle of frons (Fig. 48 A); epandrium wider than long in dorsal view (Fig. 9 A); surstyli broadened in the middle and gradually tapering toward apices in dorsal view (Fig. 9 A); both surstyli slightly curved in lateral view (Fig. 9 C – D); phallus trifid, with three wavy ejaculatory ducts, one with teeth-like projections in basal half in lateral view (Fig. 9 C – D); small and broad phallic guide (Fig. 9 C); gonopods equal in height in ventral view (Fig. 9 B). Specimens examined: ISRAEL [PALESTINE on the label]: Beth Hakerem, Jerusalem, 31 ° 46 ’ N, 35 ° 12 ’ E, 22. v. 1980, O. Theodor, JSS 51736 (1 ♂, TAU); Har Horesha, 30 ° 34 ’ N, 34 ° 31 ’ E, 900 – 1000 m a. s. l., 18. iv. 1998, JSS 51716 (1 ♂, CNC).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F57FF9CFF3EEFE8FD9149A8.taxon	distribution	Distribution: Israel, Morocco, Tajikistan (Skevington 2020) (Fig. 52). Note: Based on DNA barcodes, this species is closest to T. ellipiensis sp. nov., differing by 6.4 % (pairwise divergence) (Supplementary Table 1).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F57FF9FFF3EEDA8FF6B4E2D.taxon	description	Fig. 49	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F57FF9FFF3EEDA8FF6B4E2D.taxon	diagnosis	Diagnosis: Following Kuznetzov (1993), this species can be recognized by the distinct ventrobasal protuberance of hind femur, bearing some distinct spines (Kuznetzov, 1993: fig. 13; Kehlmaier & Majnon-Jahromi, 2014: fig. 6); syntergosternite 8 with longitudinal suture in dorsal view (Kuznetzov, 1993: fig. 9); epandrium longer than wide (Kuznetzov, 1993: Fig. 18; Kehlmaier & Majnon-Jahromi, 2014: fig. 4); surstyli short, broadened in the middle and gradually tapering toward apices in dorsal view (Kuznetzov, 1993: fig. 18; Kehlmaier & Majnon-Jahromi, 2015: fig. 4); hypandrium with a distinct constriction after basal quarter (Kehlmaier & Majnon-Jahromi, 2015: fig. 3).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F57FF9FFF3EEDA8FF6B4E2D.taxon	distribution	Distribution: Egypt, Iran (Kuznetzov, 1993; Kehlmaier & Majnon Jahromi, 2015; Skevington 2020) (Fig. 49).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F55FF91FF3EEBD5FB9D4EB9.taxon	description	Figs 10 A – E, 49	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F55FF91FF3EEBD5FB9D4EB9.taxon	diagnosis	Diagnosis: This species can be recognized by the asymmetrical shape of surstyli in dorsal view, left surstylus broadened apically and constricted medially, right surstylus broader than left one basally (Fig. 10 A); epandrium wider than long in dorsal view (Fig. 10 A); gonopods unequal in height, left is higher than right one in ventral view (Fig. 10 B); phallus trifid, with three wavy ejaculatory ducts, one with tooth-like projections in basal half in lateral view (Fig. 10 D – E). Based on the shape of the surstyli in dorsal view, it is closely related to T. angolensis De Meyer, 1993 and T. forchhammeri De Meyer, 1993. It differs from these species by the shape of the left surstylus in dorsal view, constricted in the middle (Fig. 10 A). Specimens examined: UNITED ARAB EMIRATES: Bithnah, 25 ° 06 ’ N, 56 ° 08 ’ E, 12. viii – 9. ix. 2006, A. van Harten, Malaise trap, CNCD 160279 (1 ♂, CNC); Sharjah Desert Park, 25 ° 10 ’ N, 55 ° 25 ’ E, 4 – 8. xii. 2004, A. van Harten, pan trap, CNCD 4277 (1 ♂, CNC); Um al-Quwain, 25 ° 19 ’ N, 55 ° 19 ’ E, 23 – 26. vi. 2008, A. van Harten, pan trap, JSS 52251 (1 ♂, CNC); Wadi Bih dam, 25 ° 48 ’ N, 56 ° 04 ’ E, 21 – 30. iv. 2008, A. van Harten, light trap, JSS 52243 (1 ♂, CNC); Wadi Bih, 25 ° 29 ’ N, 56 ° 02 ’ E, 29. iii. 2007, F. Menzel, sweep net, CNCD 159626 (1 ♂, TAU); Wadi Maidaq, 25 ° 11 ’ N, 56 ° 04 ’ E, 29. vii – 26. viii. 2006, A. van Harten, Malaise trap, CNCD 159901 (1 ♂, EAD).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F55FF91FF3EEBD5FB9D4EB9.taxon	distribution	Distribution: Egypt, Iran, United Arab Emirates (Skevington 2020) (Fig. 49).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F5AFF91FF3EEA98FD914819.taxon	description	Figs 11 A – F, 52	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F5AFF91FF3EEA98FD914819.taxon	diagnosis	Diagnosis: This species can be recognized by the shape of surstyli in dorsal view, both broadened in basal half; right surstylus narrowed moderately in in apical half, left surstylus narrowed abruptly in apical half (Fig. 11 A); epandrium longer than wide (Fig. 11 A); subepandrial sclerite wide at the base in ventral view (Fig. 11 B); gonopods with an unequal projection, right is farther than left one (Fig. 11 B); phallus trifid, one with some small teeth-like projections in lateral view (Fig. 11 F). Specimens examined: IRAN: Khuzestan: Shush, 32 ° 06 ’ N, 48 ° 26 ’ E, 55 m a. s. l., 29 – 31. viii. 2013, M. Parchami-Araghi, Malaise trap, JSS 52189 (1 ♀, CNC); Sistan & Baluchestan: Daman, 27 ° 24 ’ N, 60 ° 49 ’ E, 2 – 12. vii. 2016, H. Davari, Malaise trap, JSS 51822 (1 ♂, CNC); Saravan, 27 ° 25 ’ N, 62 ° 17 ’ E, 17. x. 2016, F. Hamzavi, sweep net, JSS 51827 (1 ♀, CNC); Zabol, Dostmohammad, 31 ° 09 ’ N, 61 ° 43 ’ E, 470 m a. s. l., 29. vi. 2016, M. Enaiatnia, Mal- aise trap, JSS 51933 (1 ♀, CNC); Zabol, 31 ° 0 3 ’ N, 61 ° 22 ’ E, 480 m a. s. l., 10. vii. 2016, M. Enaiatnia, Malaise trap, JSS 51860 (1 ♂, CNC); 31 ° 02 ’ N, 61 ° 32 ’ E, 470 m a. s. l., 2. ix. 2016, M. Enaiatnia, Malaise trap, JSS 51907 (1 ♀, CNC); Kerman: Fariab, Horesfandi, 28 ° 08 ’ N, 57 ° 16 ’ E, 5. viii. 2016, M. Arabzadeh, Malaise trap, JSS 51922 (1 ♂, CNC); ISRAEL: Hazeva Field School, 30 ° 43 ’ N, 35 ° 15 ’ E, 2. x. 1993, S. Plotkin, Malaise trap, JSS 51704 (1 ♂, TAU); 5. x. 1997, S. Plotkin, Malaise trap, JSS 52080 (1 ♂, TAU); Hazeva, Field School, 30 ° 43 ’ N, 35 ° 15 ’ E, 11. x. 1997, A. Maklakov, Malaise trap, JSS 51769 – 70 (2 ♂, TAU); CYPRUS: Kyrenia, 35 ° 21 ’ N, 33 ° 09 ’ E, 10 – 17. ix. 2017, O. Ozden, Malaise trap, JSS 52344 (1 ♂, CNC); 16 – 30. vii. 2017, O. Ozden, Malaise trap, JSS 52325, JSS 52355 (2 ♂, CNC); 24. ix – 1. x. 2017, O. Ozden, Malaise trap, JSS 52354 (1 ♂, CNC); 30. vii – 13. viii. 2017, O. Ozden, Malaise trap, JSS 52366 (1 ♂, CNC); 9 – 16. vii. 2017, O. Ozden, Malaise trap, JSS 52330 (1 ♂, CNC); UNITED ARAB EMIR- ATES: Abu Dhabi: Al Wathba Wetland Reserve, 24 ° 15 ’ N, 54 ° 37 ’ E, i. 2015, A. Saji & A. van Harten, Malaise trap, CNC 470746 (1 ♂, CNC); ii. 2015, A. Saji & A. van Harten, Malaise trap, CNC 470512, CNC 470572 (2 ♂, USNM); iii. 2015, A. Saji & A. van Harten, Malaise trap, CNC 470645, CNC 470654, CNC 470679 – 80 (4 ♂, CNC); iv. 2013, A. Saji & A. van Harten, Malaise trap, CNC 470729 (1 ♂, CNC); iv. 2015, A. Saji & A. van Harten, Mal- aise trap, CNC 470808 (1 ♂, CNC); ix. 2013, A. Saji & A. van Harten, Malaise trap, CNC 465503, CNC 470708 – 10, CNC 470713, CNC 470716 (6 ♂, CNC); xi. 2013, A. Saji & A. van Harten, Malaise trap, CNC 470721, CNC 470725, CNC 470728 (3 ♂, CNC); Al Bida’a Protected Area, 23 ° 36 ’ N, 53 ° 58 ’ E, 29. x. 2018, A. Saji & A. van Harten, Malaise trap, JSS 52267 – 70 (4 ♂, CNC); Al Tawi Protected Area, 23 ° 50 ’ N, 53 ° 10 ’ E, 1. xi. 2018, A. Saji & A. van Harten, Malaise trap, JSS 52295 – 98 (4 ♂, CNC); Barqa Al Suqoor Protected Area, 23 ° 60 ’ N, 52 ° 41 ’ E, 20. iii. 2018, A. Saji & A. van Harten, Malaise trap, JSS 52260 (1 ♂, CNC); 29. x. 2018, A. Saji & A. van Harten, Malaise trap, JSS 52282 (1 ♂, CNC); Wadi Wurayah farm, 25 ° 14 ’ N, 56 ° 11 ’ E, 17 – 24. iii. 2009, A. van Harten, Malaise trap, CNCD 137123 (1 ♂, CNC); 19. iv – 19. v. 2009, A. van Harten, Malaise trap, CNCD 137544 (1 ♂, CNC); 31. v – 14. vi. 2009, A. van Harten, Malaise trap, CNCD 137579, CNCD 137582 (2 ♂, EAD); Wadi Wurayah, 25 ° 14 ’ N, 56 ° 11 ’ E, 19 – 26. iv. 2009, A. van Harten, Malaise trap, CNCD 160581 (1 ♂, EAD).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F5AFF91FF3EEA98FD914819.taxon	distribution	Distribution: Cyprus, Egypt, Iran, Israel, Spain, United Arab Emirates (Skevington 2020) (Fig. 52). Note: DNA barcodes of this species are genetically most similar to T. gazliensis, differing by 7.4 % (pairwise divergence) (Supplementary Table 1).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F58FF92FF3EEBD5FE8C4B4D.taxon	description	Figs 12 A – D, 53, 66 E – F	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F58FF92FF3EEBD5FE8C4B4D.taxon	diagnosis	Diagnosis: This species can be recognized by the shape of epandrium in dorsal view, expanded towards the left side (Fig. 12 A); small surstyli with a ventromedial projection in lateral view (Fig. 12 C – D); phallic guide curved towards surstyli, with a pair of dorsolateral spines in lateral view (Fig. 12 C).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F58FF92FF3EEBD5FE8C4B4D.taxon	description	Description: MALE: Body length (excluding antennae): 2.6 mm. Head. Scape, pedicel, arista dark brown, flagellum brown; scape and pedicel without setae; flagellum tapering (LF: WF = 3.0). Eyes meeting for a distance of 7 – 8 facets. Frons silver-gray pollinose. Vertex black, lacking pollinosity. Occiput dark and gray pollinose. Thorax. Postpronotal lobe light yellow, gray pollinose with 2 – 3 small pale setae along the upper margin (up to 0.02 mm). Prescutum and scutum black, with uniseriate rows of intra-alar setae. Scutellum black, silver-gray pollinose, without bristle. Subscutellum and pleura dark brown, gray pollinose. Wing. Length: 2.5 mm. LW: MWW = 2.6. Wing almost entirely covered with microtrichia. Cross-vein r-m reaching cell dm at the middle. M 1 strongly undulating in middle. Halter length: 0.28 mm. Whitish, narrowly light brown at base. Legs. Dark brown but narrowly yellow at base of tibiae and all tarsal segments except distitarsi. Trochanters smooth. Femora partly gray pollinose, hind femur shining in posterior. Hind femur bearing two rows of dark, peg-like anteroventral spines in apical one third. Fore, mid and hind femora with 1 – 2 wrinkled indentations in basal quarter. Tibiae with two rows of short setae on anterior and three rows on posterior side. Hind tibia with 1 – 2 wrinkled indentations in the middle. Tarsi light brown, gray pollinose. Pulvilli shorter than distitarsi. Claws white with black tips. Abdomen. Ground color dark brown. Tergite 1 entirely gray pollinose with 3 – 4 yellowish short lateral setae (up to 0.05 mm). Tergites 1 – 5 with scattered yellowish setae (up to 0.05 mm). Sternites brown laterally and dark centrally, gray pollinose. Genitalia. Genital capsule in dorsal view: epandrium and surstyli brown, gray pollinose. Epandrium wider than long (MLE: MWE = 0.5), expanded towards left side (Fig. 12 A). Surstyli small, rather symmetrical, triangular-shaped, slightly curved to each other at apex (Fig. 12 A). Genital capsule in ventral view: gonopods large and slightly equal in height, subepandrial sclerite wide basically (Fig. 12 B). Genital capsule in lateral view: both surstyli small with ventromedial projection (12 C – D); phallic guide broad at base, curved towards surstyli, with two small dorsolateral spines at basal half (Fig. 12 C); phallus with three small ejaculatory ducts, one broader than other (Fig. 12 C). FEMALE: unknown.	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F58FF92FF3EEBD5FE8C4B4D.taxon	etymology	Etymology: The specific epithet refers to Ellipi, the historical name of Kermanshah, from where the holotype has been collected. Specimen examined: HOLOTYPE: IRAN: Kermanshah: Gheshlagh, 34 ° 56 ’ N, 46 ° 27 ’ E, 1533 m a. s. l., 1. vii. 2015, M. Zardouei, Malaise trap, JSS 51945 (1 ♂, CNC).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F58FF92FF3EEBD5FE8C4B4D.taxon	distribution	Distribution: Iran (Fig. 53). Note: Based on DNA barcodes, T. ellipiensis sp. nov. is genetically most similar to T. teligera (5.6 % pairwise divergence).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F59FF94FF3EEF81FA374DFD.taxon	description	Figs 13 A ‒ D, 27 E, 51, 67 A – B	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F59FF94FF3EEF81FA374DFD.taxon	diagnosis	Diagnosis: This species can be recognized by the rectangular shape of both surstyli in dorsal view (Fig. 13 A); long gonopods; sclerotized hypandrium with long hypandrial apodeme and a pair of membranous sheaths in the middle in ventral view (Fig. 13 D); straight phallic guide with a few distinct dorsolateral spines in lateral and ventral view (Fig. 13 B ‒ D). Based on the shape of the surstyli, this species is closely related to kuthyi group (Tomosvaryella cyprusensis sp. nov., T. freidbergi, T. hispanica, T. israelensis, T. kuthyi, T. parakuthyi). It differs by the round syntergosternite 8 in dorsal view (Fig. 13 A), long gonopods in ventral view (Fig. 13 D) and a few dorsolateral spines on phallic guide in lateral and ventral view (Fig. 13 B ‒ D).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F59FF94FF3EEF81FA374DFD.taxon	description	Description. MALE: Body length: 2.2 ‒ 2.9 mm. Head. Frons silver-gray pollinose. Eyes meeting for a distance of four facets. Pedicel with a pair of dorsal setae; flagellum tapering (LF: WF = 1.9 ‒ 2.1), distinctly paler. Thorax. Postpronotal lobe pale, gray pollinose with about 2 ‒ 3 postpronotal gray setae along upper margin. Prescutum and scutum black, silver-gray pollinose, with uniseriate dorsocentral row of brown setae and some distinct patches of long supra-alar hairs behind postpronotal lobes. Scutellum black, silver-gray pollinose with 8 ‒ 10 dark brown se- tae along the upper margin (up to 0.02 mm). Subscutellum black and pleura brown. Wings. Wing length: 2.3 ‒ 2.5 mm. LW: MWW = 2.6 ‒ 3.0. Wing almost entirely covered with microtrichia. Cross-vein r-m reaching cell dm at the middle. Halter length: 0.3 mm. Yellow, narrowly brown at base. Legs. Legs dark brown, narrowly yellow at apex of femora, base of tibiae and tarsi. Hind trochanter with a patch of dark brown bristle in the middle. Femora with posteroventral row of black spines and anteroventrally with a row of spines on apical third. Tibiae with two rows of short brown setae on anterior and three rows on posterior side. Pulvilli shorter than distitarsi. Claws white with black tips. Abdomen. Ground color of abdomen dark, gray pollinose; tergite 1 with 3 ‒ 4 dark lateral setae in both sides, tergites normally covered with distinct bristly brown setae. Sternite brown, lighter than tergites, gray pollinose; syntergosternite 8 dark brown with scattered brown setae, membranous area large, enlarged towards tip of abdomen and occupying more than half the width of syntergosternite 8. Epandrium darker than surstyli, higher than long (LS 8: HS 8 = 0.7 – 0.8); Genitalia. Genital capsule in dorsal view: surstyli rather symmetrical, rectangular-shaped. Left surstylus slightly larger and wider than right one (Fig. 13 A). Genital capsule in ventral view: gonopods unequal and long, left slightly higher than right one; lobes of hypandrium sclerotized, with a pair of membranous sheaths covered by small hairs in the middle of hypandrium and with long hypandrial apodeme (Fig. 13 D). Genital capsule in lateral view: left surstylus bent towards sternite in apical third, right surstylus gently curved toward sternite (Fig. 13 B – C); phallus straight and trifid. Phallic guide straight with few (2 – 3) short spines laterally (Fig. 13 B – C). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 13 D). FEMALE: Body length: 2.0 mm. Frons dark, widened in middle. Occiput gray pollinose. Eyes separated, with enlarged frontal facets. Long tapering flagellum. Scutum gray pollinose with two row dorsocentral setae and uniseriate rows of supra-alar setae. Wing length: 1.9 ‒ 2.0 mm. LW: MWW = 2.1 ‒ 2.2. Femora with posteroventral row of black spines and anteroventrally with a row of spines on apical third. Pulvilli smaller than distitarsi. Tergites 1 – 6 with scattered brown and black setae. Tergite 8 brown, rounded, gray pollinose, distal part of piercer straight and long (Fig. 27 E). LP: LB = 1.9 ‒ 2.1. LDP: LPP = 1.8 ‒ 2.0.	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F59FF94FF3EEF81FA374DFD.taxon	etymology	Etymology: The specific epithet refers to the Emirates, from where the specimens have been collected. Specimens examined: HOLOTYPE: UNITED ARAB EMIRATES: Wadi Wurayah farm, 25 ° 23 ’ N, 56 ° 19 ’ E, 31. v. ‒ 14. vi. 2009, Malaise trap, A. van Harten, CNCD 137568 (1 ♂, CNC); PARATYPES: UNITED ARAB EMIR- ATES: 19. iv ‒ 19. v. 2009, same data as holotype, CNCD 137555 (1 ♂, EAD); Fujairah, 25 ° 04 ’ N, 56 ° 12 ’ E, 28. ii. ‒ 1. iv. 2006, light trap, A. van Harten, CNCD 8954 (1 ♂, CNC), CNCD 8955 (1 ♂, USNM); Wadi Safad, 25 ° 07 ’ N, 56 ° 11 ’ E, 20. xii. 2005 ‒ 2. i. 2006, light trap, A. van Harten, CNCD 8976 (1 ♂, CNC); 19. iv ‒ 18. v. 2009, same data as holotype, CNCD 137556 (1 ♀, CNC).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F59FF94FF3EEF81FA374DFD.taxon	distribution	Distribution: United Arab Emirates (Fig. 51). Note: Based on DNA barcodes, Tomosvaryella emaratensis sp. nov. is genetically most similar to T. minuscula, differing by 2.2 % (pairwise divergence). The male and female sequenced differ by 0.5 % (Supplementary Table 1).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F5FFF94FF3EE85CFE6E49A9.taxon	description	Figs 14 A ‒ E, 54	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F5FFF94FF3EE85CFE6E49A9.taxon	diagnosis	Diagnosis: This species can be recognized by the slender shape of surstyli in dorsal view (Fig. 14 A); both surstyli straight in basal half, gently bent in apical half in lateral view (Fig. 14 D ‒ E); epandrium longer than wide (Fig. 14 A); gonopods equal in height (Fig. 14 B). Based on the shape of genitalia, this species is part of the kuthyi species group (Tomosvaryella emaratensis sp. nov., T. cyprusensis sp. nov., T. hispanica, T. israelensis, T. kuthyi and T. parakuthyi). It differs from these species by the shape of surstyli in lateral view, slender, slightly straight in basal half and being bent in apical half (De Meyer, 1995: fig. 13 a – c; Földvári & De Meyer, 1999: fig. 10 A – D). See diagnosis of T. emaratensis sp. nov. and T. cyprusensis sp. nov. for more details. Specimens examined: HOLOTYPE: ISRAEL: Mt. Hermon, 33 ° 25 ’ N, 35 ° 51 ’ E, 2000 m a. s. l., 1. vii. 1986, A. Freidberg, JSS 52375 (1 ♂, TAU); PARATYPE: ISRAEL: Meron, 32 ° 59 ’ N, 35 ° 26 ’ E, 11. iv. 1974, F. Nachbar, JSS 52376 (1 ♂, TAU). Other specimens examined: CYPRUS: Kyrenia, 35 ° 21 ’ N, 33 ° 09 ’ E, 16 ‒ 30. vii. 2017, O. Ozden, Malaise trap, JSS 52356 (1 ♂, CNC); IRAN: North Khorasan: Biar, 37 ° 53 ’ N, 57 ° 35 ’ E, 1008 m a. s. l., 10 ‒ 24. vi. 2016, B. Motamedinia, Malaise trap; JSS 52018 (1 ♂, CNC); Chenaran village, 37 ° 27 ’ N, 57 ° 36 ’ E, 1300 m a. s. l., 12. vi. 2016, B. Motamedinia, sweep net, JSS 51963 (1 ♂, CNC); South Khorasan: Ghiok, 32 ° 47 ’ N, 59 ° 07 ’ E, 1917 m a. s. l., 9. vi. 2016, B. Motamedinia, sweep net, JSS 52000 (1 ♂, CNC); Kermanshah: Dodan, 35 ° 00 ’ N, 46 ° 12 ’ E, 1011 m a. s. l., Garden, 20. vi. 2016, M. Zardouei, Malaise trap, JSS 52177 (1 ♂, CNC); ISRAEL: Har Hermon, 33 ° 18 ’ N, 35 ° 46 ’ E, 2000 m a. s. l., 3. viii. 1995, A. Freidberg, JSS 52076 (1 ♂, TAU).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F5FFF94FF3EE85CFE6E49A9.taxon	distribution	Distribution: Czech Republic Egypt, France, Greece, Hungary, Iran, Israel, Italy, Kyrgyz Republic, Portugal, Spain, Syria, Turkey (Skevington 2020) (Fig. 54). Notes: DNA barcodes of T. freidbergi are identical to those of T. kuthyi (0.0 ‒ 1.5 % pairwise divergence). The terminalia of these species differ a little by the shape of left surstylus, so this is likely a case of recently-diverged species or ancestral hybridisation. It is possible that these are a single species with polymorphic genitalia, so future genetic work is warranted.	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F5FFF97FF3EED9DFEA74FE5.taxon	description	Fig. 51	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F5FFF97FF3EED9DFEA74FE5.taxon	diagnosis	Diagnosis. Following Hardy (1967), this species can be recognized by syntergosternite 8, wider than long, without membranous area; epandrium wider than long; symmetrical surstyli, slightly rectangular-shaped, almost twice as wide as long, blunt at apices (Hardy, 1967: fig. 6).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F5FFF97FF3EED9DFEA74FE5.taxon	distribution	Distribution: Egypt, France, Israel, Italy, Malta, Morocco, Palestine, Romania, Spain, Tunisia (Skevington 2020) (Fig. 51). Note: This species has been reared from Tamarix leafhopper, Opsius stactogalus Fieber, 1866, in France (Hardy, 1967).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F5CFF96FF3EEC28FB054D29.taxon	description	Figs 15 A ‒ D, 51	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F5CFF96FF3EEC28FB054D29.taxon	diagnosis	Diagnosis: This species can be recognized by the shape of surstyli in lateral view, rounded apically, with a distinct posteroventral projection before its apex (Fig. 15 B ‒ C); epandrium wider than long in dorsal view (Fig. 15 A); gono- pods equally in height in ventral view (Fig. 15 D); subepandrial sclerite small, basically wide in ventral view (Fig. 15 D). Based on the shape of genitalia, this species resembles T. urdaensis Kuznetzov, 1994. It differs by the shape of surstyli in dorsal view. Base of surstyli in T. urdaensis is broader and more angulated (Majnon-Jahromi et al. 2017 b: fig. 19). Specimens examined: IRAN: Sistan & Baluchestan: Saravan, 27 ° 25 ’ N, 62 ° 17 ’ E, 8. xi. 2016, F. Hamzavi, pan trap, JSS 51918 (1 ♂, CNC).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F5CFF96FF3EEC28FB054D29.taxon	distribution	Distribution: Iran, Uzbekistan (Skevington 2020) (Fig. 51). Note: Tomosvaryella gazliensis and T. urdaensis Kuznetzov, 1994, share identical COI barcode (0.0 % pairwise divergence) (Supplementary Table 1). In addition, the terminalia of these species are rather similar so future work is required to clarify if they are a single species with polymorphic genitalia or different species.	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F5DFFA9FF3EEE87FDAA4EE1.taxon	description	Figs 16 A ‒ E, 53	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F5DFFA9FF3EEE87FDAA4EE1.taxon	diagnosis	Diagnosis: This species can be recognized by the shape of surstyli in dorsal view, both surstyli slightly rectangular shaped, broader at the base and tips (Fig. 16 A), tip of both surstyli flattened apically in lateral view (Fig. 16 D – E); epandrium wider than long (Fig. 16 A); phallus trifid, one of ejaculatory ducts bearing some small spines in lateral view (Fig. 16 E); gonopods equal in height (Fig. 16 B). Specimens examined: IRAN: Kermanshah: Ghazanchi, 34 ° 26 ’ N, 47 ° 00 ’ E, 1304 m a. s. l., 20. vi. 2016, M. Zardouei, funnel Malaise trap, JSS 51824 (1 ♂, CNC); Harsin, 34 ° 19 ’ N, 47 ° 26 ’ E, 1324 m a. s. l., 22. vii. 2016, M. Zardouei, funnel Malaise trap, JSS 51944 (1 ♂, CNC); Sarpolezahab, 34 ° 28 ’ N, 45 ° 49 ’ E, 456 m a. s. l., 14. v. 2016, M. Zardouei, Malaise trap, JSS 52230 (1 ♀, CNC); South Khorasan: Ark, 33 ° 01 ’ N, 58 ° 41 ’ E, 1605 m a. s. l., 30. v – 12. vi. 2016, B. Motamedinia, Malaise trap, JSS 51997 – 8 (1 ♂, 1 ♀, USNM); Ghiok, 32 ° 47 ’ N, 59 ° 07 ’ E, 1917 m a. s. l., 9. vi. 2016, B. Motamedinia, sweep net, JSS 52001 (1 ♂, CNC); Jafarieh, 32 ° 48 ’ N, 59 ° 14 ’ E, 1760 m a. s. l., 21. iv – 5. v. 2016, B. Mo- tamedinia, Malaise trap, JSS 52007 (1 ♀, CNC); Mohammadieh, 32 ° 52 ’ N, 59 ° 01 ’ E, 1419 m a. s. l., 14 – 30. vii. 2016, B. Motamedinia, funnel Malaise trap, JSS 52051 (1 ♂, CNC); 3 – 17. vii. 2016, B. Motamedinia, Malaise trap; funnel Malaise trap, JSS 52041 – 2, JSS 52045 – 6, JSS 52048, JSS 51966 – 7, JSS 51968 – 72 (11 ♂, 1 ♀, CNC); 5 – 23. ix. 2016, B. Motamedinia, Malaise trap, JSS 52025 (1 ♂, CNC); ISRAEL: Nahal Eshharim, 30 ° 28 ’ N, 34 ° 35 ’ E, 790 m a. s. l., 2. v. 1995, I. Yarom, Malaise trap, JSS 51791 (1 ♂, TAU).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F5DFFA9FF3EEE87FDAA4EE1.taxon	distribution	Distribution: Austria, Belgium, Bulgaria, Croatia, Cyprus, Czech Republic, Denmark, Finland, France, Greece, Germany, Great Britain, Hungary, Iran Israel, Italy, Latvia, Macedonia, Mongolia, Morocco, Netherlands, Poland, Portugal, Romania, Russia, Slovakia, Spain, Sweden, Switzerland (Skevington 2020) (Fig. 53). Note: This species is genetically most similar to T. emaratensis sp. nov., differing by 9.7 % (pairwise divergence) (Supplementary Table 1).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F63FFA8FF3EEBD5FB9B4EBA.taxon	description	Fig. 51	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F63FFA8FF3EEBD5FB9B4EBA.taxon	diagnosis	Diagnosis: Following Majnon-Jahromi et al. (2017 b), this species can be recognized by the shape of surstyli in lateral view, both cleaver-shaped; epandrium wider than long in dorsal view; phallic guide enlarged apically in lateral view; phallus trifid, with a small hook at the base of ejaculatory ducts (Majnon-Jahromi et al. 2017 b: fig. 1 A – D).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F63FFA8FF3EEBD5FB9B4EBA.taxon	distribution	Distribution: Iran (Majnon-Jahromi et al. 2017 b; Skevington 2020) (Fig. 51).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F63FFABFF3EEA99FDD74E71.taxon	description	Figs 17 A – E, 27 J, 47 B, 51, 67 C – D	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F63FFABFF3EEA99FDD74E71.taxon	diagnosis	Diagnosis: This species can be recognized by the shape of surstyli in dorsal view, symmetrical, broadened apically (Fig 17 A), both surstyli rather rectangular-shaped in lateral view (Fig. 17 D – E); epandrium wider than long (Fig. 17 A); hind trochanter expanded apically with a small projection (Fig. 47 B). Based on the shape of surstyli in dorsal view, this species resembles T. nodosa and T. pruinosa. It differs by having broader surstyli apically and lacking a hypandrial apodeme (Fig 17 B). Additionally, the hind trochanter of T. pruinosa lacks the small projection found in T. hamounensis sp. nov. (Fig. 47 B) and T. nodosa (Fig. 47 C).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F63FFABFF3EEA99FDD74E71.taxon	description	Description: MALE: Body length (excluding antennae): 2.5 – 2.7 mm (n = 11). Head. Scape dark, pedicel and flagellum brown, arista light brown. Flagellum long tapering and gray-white pollinose (LF: WF = 5.0). Eyes meeting for a distance of 10 – 12 facets. Frons dark silver-gray pollinose. Vertex black, lacking pollinosity, bearing an elevated ocellar triangle. Occiput dark and gray pollinose. Thorax. Postpronotal lobe light yellow, gray pollinose with 3 – 5 yellowish setae along the upper margin (up to 0.03 mm). Prescutum, Scutum, Scutellum black. Scutum with two short uniseriate rows of intra-alar setae and two uniseriate rows of dorsocentral setae. Scutellum gray pollinose with 8 – 10 thin short setae along lateral margin (up to 0.01 mm). Subscutellum black, gray pollinose. Pleura brown. Wing. Length: 2.2 – 2.8 mm. LW: MWW = 1.6. Wing almost entirely covered with microtrichia. Cross-vein r-m reaches dm at the middle. M 1 moderately undulating in middle. Halter length: 0.33 mm. Whitish, narrowly brown at base. Legs. Coxae brown, gray pollinose. Mid coxa with two or three short dark anterior setae. Trochanters brown, gray pollinose. Hind trochanter broader apically with small projection with some short setae (Fig. 47 B). Femora brown with light brown apices, gray pollinose. All femora with one wrinkled indentations at base. Tibiae light brown but darker in middle, with two ventral rows of short setae on anterior and three rows on posterior side. Hind tibia with 1 – 2 wrinkled indentations in middle. Tarsi light brown with scattered brown setae at anterior margin. Pulvilli yellow. Claws light brown with black tips. Abdomen. Ground color brown. Tergite 1 densely gray pollinose, with 3 – 5 brown lateral setae (up to 0.08 mm). Tergite 2 – 5 gray pollinose, with scattered brown setae. Genitalia. Genital capsule in dorsal view: epandrium and surstyli light brown. Epandrium wider than long (MLE: MWE = 0.54). Surstyli rather small and symmetrical. Both surstyli broadened at the middle, with an inner small projection at apex (Fig. 17 A). Genital capsule in ventral view: gonopods rather equal in height (Fig. 17 B). Subepandrial sclerite small and triangular-shaped (Fig. 17 B). Genital capsule in lateral view: both surstyli rather rectangular shape, left surstylus with small ventroapical projection and right surstylus with ventrobasal projection (Fig. 17 D – E). Phallic guide strong, phallus with three ejaculatory ducts. Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 17 C). FEMALE: Body length: 2.6 – 2.7 mm (n = 5). Frons gray polllinose, widened in middle. Occiput gray pollinose. Eyes separated. Long tapering flagellum (LF: WF = 3.5). Scutum gray pollinose with two row dorsocentral setae. Pleura dark brown, silver-gray pollinose. Wing length: 1.9 ‒ 2.0 mm. LW: MWW = 2.1 ‒ 2.3. Fore femur bearing two smaller anteroventral spines in basal third, hind femur with 1 – 2 wrinkled indentations in basal. Pulvilli smaller than distitarsi. Tergites 1 – 6 with scattered gray pollinose. Tergite 8 brown, rounded, gray pollinose, distal part of piercer straight (Fig. 27 J). LP: LB = 1.3 ‒ 1.4. LDP: LPP = 2.8 ‒ 2.9.	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F63FFABFF3EEA99FDD74E71.taxon	etymology	Etymology. The specific epithet refers to Hamoun, the region where the specimens have been collected. Specimens examined: HOLOTYPE: IRAN: Sistan & Baluchestan: Zabol, 31 ° 02 ’ N, 61 ° 32 ’ E, 470 m a. s. l., 13. ix. 2016, M. Enaiatnia, Malaise trap, JSS 51893 (1 ♂, CNC); PARATYPES: IRAN: Sistan & Baluchestan: Zabol, 31 ° 02 ’ N, 61 ° 32 ’ E, 13. ix. 2016, M. Enaiatnia, Malaise trap, JSS 51894 – 5 (2 ♂, TAU, USNM); 2. ix. 2016, M. Enaiat- nia, Malaise trap, JSS 51896 (1 ♀, CNC); 31 ° 03 ’ N, 61 ° 22 ’ E, 7. iv. 2016, M. Enaiatnia, Malaise trap, JSS 51816 – 7, JSS 52164, JSS 52169 (4 ♂, CNC); 2 – 13. ix. 2016, M. Enaiatnia, funnel Malaise trap, JSS 52166, JSS 52156, JSS 52178 (3 ♀, CNC); 6. x. 2016, M. Enaiatnia, Malaise trap, JSS 52170 (1 ♀, CNC).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F63FFABFF3EEA99FDD74E71.taxon	distribution	Distribution: Iran (Fig. 51). Note: Based on DNA barcodes, T. hamounensis sp. nov. is most similar to T. pruinosa, differing by 5.7 % (pairwise divergence) (Supplementary Table 1).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F61FFAAFF3EEBD5FBEB4E9E.taxon	description	Fig. 53	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F61FFAAFF3EEBD5FBEB4E9E.taxon	diagnosis	Diagnosis: Following Kehlmaier et al. (2019), this species can be recognized by the symmetrical surstyli in dorsal view, broadened at the base, both surstyli slightly bean-shaped in lateral view; epandrium wider than long; phallus trifid, one of ejaculatory duct with small spines; phallic guide bent towards surstyli in lateral view (Kehlmaier et al. 2019: fig. 8 A – D).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F61FFAAFF3EEBD5FBEB4E9E.taxon	distribution	Distribution: Egypt, Israel (Kehlamier et al. 2019; Skevington 2020) (Fig. 53).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F61FFAAFF3EEABDFEE74DC1.taxon	description	Fig. 49	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F61FFAAFF3EEABDFEE74DC1.taxon	distribution	Distribution: Iran (Skevington 2020) (Fig. 49). Notes. The female of this species was described by Becker (1913) and the male has not been described or illustrated so far. The type specimen is currently on loan to Sergey Kuznetzov and unavailable (Kehlmaier & Majnon- Jahromi, 2014).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F61FFAAFF3EE801FE384AAE.taxon	description	Figs 18 A – E, 55	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F61FFAAFF3EE801FE384AAE.taxon	diagnosis	Diagnosis: This species can be recognized by its small epandrium, wider than long (Fig. 18 A); both surstyli beanshaped, with an inner small projection shortly before apex in dorsal view (Fig. 18 A); phallus with four ejaculatory ducts in lateral view (Fig. 18 C – D); equal gonopods in ventral view (Fig. 18 B). Specimens examined: IRAN: Sistan & Baluchestan: Bandan, 31 ° 24 ’ N, 60 ° 44 ’ E, 12. ix. 2016, N. Khaje, sweep net, JSS 51825 (1 ♂, CNC); ISRAEL: 5 km S Qiryat Gat, Phillip Farm, 31 ° 31 ’ N, 34 ° 47 ’ E, 12. x. 2004, A. Freid- berg, JSS 51801 (1 ♂, TAU); Sappir, 30 ° 36 ’ N, 35 ° 11 ’ E, 1. v. 1997, A. Freidberg, JSS 51703 (1 ♂, TAU); Nizzanim, 31 ° 45 ’ N 34 ° 38 ’ E, 23. ix. 2004, A. Freidberg, JSS 52098 (1 ♂, TAU); UNITED ARAB EMIRATES: Bithnah, 25 ° 06 ’ N, 56 ° 08 ’ E, 12. viii – 9. ix. 2006, A. van Harten, Malaise trap, CNCD 160273 – 8 (5 ♂, CNC); 19. x – 16. xi. 2006, A. van Harten, Malaise trap, CNCD 11521 – 2, CNCD 11524 (3 ♂, CNC); 4. vii – 12. viii. 2006, A. van Harten, CNCD 159939 (1 ♂, USNM); Wadi Bih dam, 25 ° 48 ’ N, 56 ° 04 ’ E, 19 – 25. ii. 2009, A. van Harten, Malaise trap, JSS 52253, JSS 52258 (2 ♂, CNC); Wadi Maidaq, 25 ° 11 ’ N, 56 ° 04 ’ E, 29. vii – 26. viii. 2006, A. van Harten, CNCD 159902 – 3 (2 ♂, CNC); al- Ajban, 24 ° 22 ’ N, 55 ° 01 ’ E, 27. v – 26. vi. 2006, A. van Harten, light trap, JSS 52245 (1 ♂, EAD).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F61FFAAFF3EE801FE384AAE.taxon	distribution	Distribution: Greece, Iran, Israel, Syria, United Arab Emirates (De Meyer, 1995; Kehlmaier et al. 2019; Skevington 2020) (Fig. 55).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F61FFAAFF3EEEADFEF348ED.taxon	description	Figs 19 A – E, 56	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F61FFAAFF3EEEADFEF348ED.taxon	diagnosis	Diagnosis: This species is related to T. oligoseta De Meyer, 1993 from Botswana (South Africa). Both have rather symmetrical surstyli in dorsal view, broadened at base and apically, constricted at middle (Fig. 19 A); epandrium wider than long (Fig. 19 A). It slightly differs by the shape of surstyli in dorsal view, the bristling of the phallus and lateral setae in tergite 1, being completely reduced in T. oligoseta. Specimens examined: ISRAEL: Nahal Oren, 32 ° 43 ’ N, 35 ° 01 ’ E, 25. vii. 2002, A. Freidberg, JSS 52071 (1 ♂, TAU); Nahal Oren, S. facing slope, 32 ° 43 ’ N, 35 ° 01 ’ E, 2. vii. 1998, A. Freidberg, JSS 52081 (1 ♂, TAU); Nizzanim, A. Dunes, 31 ° 43 ’ N, 34 ° 38 ’ E, 6. iv. 2009, A. Freidberg, JSS 52068 – 9 (2 ♂, TAU); UNITED ARAB EMIRATES: Wadi Safad, 25 ° 07 ’ N, 56 ° 11 ’ E; 26. xii. 2005 – 2. i. 2006, A. van Harten, CNCD 8963, CNCD 8969 (2 ♂, CNC).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F61FFAAFF3EEEADFEF348ED.taxon	distribution	Distribution: Egypt, Israel (Skevington 2020) (Fig. 56). Note: T. inopinata is genetically similar to T. dentiterebra, differing by 3.9 % (pairwise divergence) (Supplementary Table 1).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F64FFAFFF3EEBD5FEE64D9A.taxon	description	Figs 20 A – E, 55	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F64FFAFFF3EEBD5FEE64D9A.taxon	diagnosis	Diagnosis: Based on the shape of the surstyli in dorsal view, this species is related to the kuthyi species group (Tomosvaryella emaratensis sp. nov. T. cyprusensis sp. nov., T. freidbergi, T. hispanica, T. kuthyi and T. parakuthyi). It differs from these species by the shape of the surstyli in lateral view, broadened, being normally curved (Fig. 20 D – E). This species also can be recognized by the shape of phallus, with three ejaculatory ducts, one with more than eight small spines in basal two thirds in lateral view (Fig. 20 D – E); epandrium longer than wide (Fig. 20 A), gonopods equal in height (Fig. 20 B). See diagnosis of T. emaratensis sp. nov. and T. cyprusensis sp. nov. for more details. Specimens examined: HOLOTYPE: ISRAEL: Kfar Shamai, 32 ° 57 ’ N, 35 ° 27 ’ E, 30. ix. 1975. A. Freidberg, JSS 52377 (1 ♂, TAU); PARATYPE: ISRAEL: Mt. Hermon, 33 ° 25 ’ N, 35 ° 51 ’ E, 1300 m a. s. l., 22. v. 1973, A. Freidberg, JSS 52378 (1 ♂, CNC).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F64FFAFFF3EEBD5FEE64D9A.taxon	distribution	Distribution: Czech Republic, Greece, Hungary, Iran, Israel, Italy, Romania, Slovakia, Syria (Skevington 2020) (Fig. 55).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F64FFAFFF3EE9B9FDD74B91.taxon	description	Figs 21 A – E, 47 A, 53	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F64FFAFFF3EE9B9FDD74B91.taxon	diagnosis	Diagnosis: This species can be recognized by the small epandrium, wider than long (Fig. 21 A); both surstyli rather straight in lateral view (Fig. 21 D – E); phallus with three ejaculatory ducts, one with some long spines towards surstyli in lateral view (Fig. 21 D); subepandrial sclerite long, both sides with about seven setae apically (Fig. 21 B); gonopods broad, equal in height (Fig. 21 B); hind trochanter with a small projection (Fig. 47 A). Specimens examined: IRAN: Khuzestan: Shush, 32 ° 06 ’ N, 48 ° 26 ’ E, 68 m a. s. l., 11. iii – 10. v. 2015, E. Gilasian, Malaise trap, JSS 52231, JSS 52203 (1 ♂, ♀, CNC).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F64FFAFFF3EE9B9FDD74B91.taxon	distribution	Distribution: Israel, Iran (De Meyer, 1995) (Fig. 53). Note: Based on DNA barcoding, T. jubata is very similar to T. bistounensis sp. nov., differing by 1.9 % (pairwise divergence) (Supplementary Table 1).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F64FFA3FF3EEFB1FDA84E55.taxon	description	Figs 22 A – D, 53, 67 E – F	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F64FFA3FF3EEFB1FDA84E55.taxon	diagnosis	Diagnosis: This species can be recognized by the shape of surstyli in dorsal view, right surstylus rectangular-shaped and left surstylus roughly club-shaped (Fig. 22 A); left surstylus with apicoventral projection in lateral view (Fig. 22 C); long phallic guide, straight dorsally, wavy ventrally in lateral view (Fig. 22 C – D).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F64FFA3FF3EEFB1FDA84E55.taxon	description	Description: MALE: Body length (excluding antennae): 2.6 mm. Head. Face dark, silver-gray pollinose. Scape and pedicel dark; pedicel with a pair of short upper setae and 1 – 2 short lower setae; flagellum light brown, tip paler than base, tapering and gray pollinose (LF: WF = 2.6); arista dark, with thickened base. Eyes converging but not meeting and separated by less than diameter of frontal facets. Frons dark, silver-gray pollinose, vertex dark, lacking pollinosity, bearing an elevated ocellar triangle; vertex dark, lacking pollinosity; occiput dark, gray pollinose. Thorax. Pleura, prescutum, scutum and scutellum dark. Pleura gray pollinose. Postpronotal lobe yellow, weakly gray pollinose and with 2 – 3 light brown postpronotal setae along upper margin. Prescutum and scutum narrowly gray pollinose, with two uniseriate dorsocentral rows of setae and some supra-alar setae. Scutellum gray pollinose, with about 2 – 3 thin short setae along posterior margin (up to 0.03 mm). Subscutellum gray pollinose. Wing. Length: 2.5 mm. LW: MWW = 2.5. Wing almost entirely covered in microtrichia. M 1 gently undulating. Halter length: 0.33 mm. Base and stem brown, knob paler than base. Legs. Coxae dark, gray pollinose. Mid coxa with 2 – 3 light brown setae on apical margin. Trochanters dark, smooth. Femora dark brown, slightly light brown at apex, gray pollinose. Fore femur bearing two smaller anteroventral spines in basal third, Fore and hind femora with 1 – 2 wrinkled indentations in basal. Tibiae dark brown, distinctly light brown in basal third, gray pollinose with one wrinkled indentation in middle, hind tibia with some light brown setae in apical third. Tarsi light brown and paler than tibiae, gray pollinose, with some light brown setae dorsally. Pulvilli yellow. Claws light brown with black tips. Abdomen. Ground color dark, gray pollinose. Tergite 1 with six to eight strong lateral setae. Tergites 1 – 5 with scattered brown setae. Syntergosternite 8 dark, brown pollinose. Genitalia. Genital capsule in dorsal view: epandrium brown, wider than long (MLE: MWE = 0.71). Surstyli brown and asymmetrical, right surstylus rectangular-shaped and left surstylus roughly club-shaped, wider than right one, tip of left surstylus rounded (Fig. 22 A). Genital capsule in ventral view: gonopods equal in height, rather broad and almost symmetrical (Fig. 22 B); phallic guide strong and long, reaches in middle of surstyli, subepandrial sclerite wide, rather rectangular-shaped (Fig. 22 B). Genital capsule in lateral view: both surstyli rectangular-shaped, bent towards sternite (Fig. 22 C – D); left surstylus with apicoventral projection (Fig. 22 C); phallic guide thin and long, straight dorsally, wavy ventrally (Fig. 22 C – D); phallus trifid with three ejaculatory ducts, one bearing small teeth along basal two thirds (Fig. 22 C – D). FEMALE: unknown.	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F64FFA3FF3EEFB1FDA84E55.taxon	etymology	Etymology: The specific epithet refers to “ Kiansi ”, the historical name of a Hamoun wetland, from where the specimens were collected. The name is formed as a feminine noun in the genitive case. Specimens examined: HOLOTYPE: IRAN: Sistan & Baluchestan: Zabol, 31 ° 07 ’ N, 61 ° 28 ’ E, 481 m a. s. l., 3. iv. 2015, H. Derafshan, Malaise trap, JSS 51921 (1 ♂, CNC). PARATYPE: IRAN: Sistan & Baluchestan: Zabol, 31 ° 12 ’ N, 61 ° 20 ’ E, 14. iv. 2014, H. Derafshan, sweep net, JSS 51828 (1 ♂, CNC).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F64FFA3FF3EEFB1FDA84E55.taxon	distribution	Distribution: Iran (Fig. 53).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F68FFA2FF3EEAF4FB594CDD.taxon	description	Figs 23 A – D, 57	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F68FFA2FF3EEAF4FB594CDD.taxon	diagnosis	Diagnosis: This species can be recognized by elongated surstyli and epandrium in dorsal view (Fig. 23 A); long subepandrial sclerite mostly covered by dense hair in ventral view (Fig. 23 B); unequal gonopods, left is higher than right one in ventral view (Fig. 23 B); both surstyli normally curved in lateral view (Fig. 23 C ‒ D). Based on the shape of the surstyli in dorsal view, this species is related to the kuthyi species group, Tomosvaryella cyprusensis sp. nov., T. emaratensis sp. nov., T. freidbergi, T. hispanica, T. israelensis and T. parakuthyi. It differs from these species by the shape of surstyli in lateral view, slightly narrow, being smoothly curved (Fig. 23 C ‒ D). See diagnosis of T. emaratensis sp. nov. and T. cyprusensis sp. nov. for more details. Specimens examined: CYPRUS: Kyrenia, 35 ° 21 ’ N, 33 ° 09 ’ E, 1 ‒ 8. x. 2017, O. Ozden, Malaise trap, JSS 52335 – 6, JSS 52338 – 9 (4 ♂, CNC); 19 ‒ 25. v. 2017, O. Ozden, Malaise trap, JSS 52326 (1 ♂, CNC); 22 ‒ 29. x. 2017, O. Ozden, Malaise trap, JSS 52359 (1 ♂, CNC); 3 ‒ 17. vi. 2017, O. Ozden, Malaise trap, JSS 52331 (1 ♂, USNM); IRAN: Ker- manshah: Dodan, 35 ° 00 ’ N, 46 ° 12 ’ E, 1011 m a. s. l., Garden, 20. v. 2016, M. Zardouei, Malaise trap, JSS 52176 (1 ♂, CNC); 20. vi. 2016, M. Zardouei, Malaise trap, JSS 52183 – 4, JSS 52181 (2 ♂, 1 ♀, CNC); Ghazanchi, 34 ° 26 ’ N, 47 ° 00 ’ E, 1304 m a. s. l., 15. vi. 2015, M. Zardouei, Malaise trap, JSS 51941 (1 ♀, CNC); North Khorasan: Biar, 37 ° 53 ’ N, 57 ° 35 ’ E, 1008 m a. s. l., 10 ‒ 24. vi. 2016, B. Motamedinia, Malaise trap, JSS 52015, JSS 52019 (2 ♂, CNC); Darkesh, 37 ° 26 ’ N, 56 ° 44 ’ E, 23. vii. 2016, B. Motamedinia, sweeping, JSS 52032, JSS 52035 (1 ♂, 1 ♀, CNC); Ge- lian, 37 ° 15 ’ N, 57 ° 55 ’ E, 1283 m a. s. l., 8 ‒ 22. vii. 2016, B. Motamedinia, Malaise trap, JSS 51994, JSS 51986 (2 ♂, CNC, USNM); Sistan & Baluchestan: Zabol, 31 ° 07 ’ N, 61 ° 28 ’ E, 481 m a. s. l., 6. vi. 2016, H. Derafshan, sweep net, JSS 51877 (1 ♀, CNC); South Khorasan: Mohammadieh, 32 ° 52 ’ N, 59 ° 01 ’ E, 1419 m a. s. l., 3 ‒ 17. vii. 2016, B. Motamedinia, funnel Malaise trap, JSS 51974 (1 ♂, CNC); Salmanefarsi, 32 ° 30 ’ N, 59 ° 13 ’ E, 2. vi. 2016, B. Mota- medinia, sweep net, JSS 52029 (1 ♀, CNC); Tehran: Shahriar, 35 ° 39 ’ N, 51 ° 01 ’ E, 20. vii. 2012, E. Gilasian, Malaise trap, JSS 52219 (1 ♀, CNC); ISRAEL: Arava Valley, 0.2 km N. Hazeva Field School, small wadi, 30 ° 46 ’ N, 35 ° 14 ’ E, - 116 m a. s. l., 30. iii. 1995, M. E. Irwin, Malaise trap, JSS 52062 (1 ♂, TAU); Arava Valley, Iddan, wadi running east of date palm orchard, 30 ° 48 ’ N, 35 ° 16 ’ E, - 110 m a. s. l., 23. iii. 1995, M. E. Irwin, Malaise trap, JSS 51814 (1 ♂, TAU); Arava Valley, Iddan, Small wadi, 2.4 km. W. of hwy. 90 at km 149, 30 ° 42 ’ N, 35 ° 11 ’ E, - 60 m a. s. l., 16. iv. 1995, M. E Irwin, hand net, JSS 51727 (1 ♂, TAU); B. S. Hazeva, 13. iii. 1998, S. Alfi, JSS 51795 (1 ♂, TAU); Deir Sha- man 500 m N near Yarden, 32 ° 02 ’ N, 35 ° 30 ’ E, 15. iii. 2005, L. Friedman, JSS 51759 (1 ♂, TAU); En Mor, 30 ° 49 ’ N, 34 ° 46 ’ E, 29. vi. 1994, A. Freidberg, JSS 52087 (1 ♂, TAU); Eshkolot, 31 ° 23 ’ N, 34 ° 45 ’ E, 29. v. 2002, A. Freidberg, JSS 51693 (1 ♂, TAU); Hadera Berekhat Atta, 32 ° 26 ’ N, 34 ° 55 ’ E, 1. v. 1998, A. Freidberg, JSS 52088 (1 ♂, TAU); Herzliyya Hill, 32 ° 09 ’ N, 34 ° 50 ’ E, 11. v. 2007, A. Freidberg, JSS 51798 (1 ♂, TAU); Holon, 16. ii. 1995, A. Freidberg, JSS 51772 (1 ♂, TAU); Isfiya, 32 ° 43 ’ N, 35 ° 04 ’ E, 450 m a. s. l., 21. v. 2010, A. Freidberg, JSS 52118 (1 ♂, TAU); La- trun, 31 ° 49 ’ N, 34 ° 58 ’ E, 10. iii. 2004, L. Friedman, JSS 52119 (1 ♂, TAU); Maagar Bental, 7. v. 2007, L. Friedman, JSS 51794 (1 ♂, TAU); Maalot, 33 ° 00 ’ N, 35 ° 17 ’ E, 580 m a. s. l., 25. v. 2009, A. Freidberg, JSS 51741 (1 ♂, TAU); Nahal Dishon, Rt. 886, 33 ° 03 ’ N, 35 ° 30 ’ E, 22. v. 1998, A. Freidberg, light trap, JSS 51713 – 4 (2 ♂, TAU); Nahal Oren, 32 ° 43 ’ N, 35 ° 02 ’ E, 2. vi. 2003, A. Freidberg, JSS 51784 (1 ♂, TAU); 30. v. 1995, A. Freidberg, JSS 52070 (1 ♂, TAU); Nahal Oren, river bed, 32 ° 43 ’ N, 35 ° 02 ’ E, 14. v. 2002, A. Freidberg, JSS 51712 (1 ♂, TAU); Nahal Qidron, 31 ° 42 ’ N, 35 ° 20 ’ E, 7. vi. 1996, A. freidberg, JSS 52074 – 5 (2 ♂, TAU); Nahal Tavor, south facing slope, 32 ° 39 ’ N, 35 ° 27 ’ E, 26. iii. 2002, L. Friedman, JSS 51706 (1 ♂, TAU); Nahal Teqoa, Maale Rehavam, 31 ° 39 ’ N, 35 ° 15 ’ E, 460 m a. s. l., 31. iii. 2009, A. Freidberg, JSS 51743 – 4 (2 ♂, TAU); Nahal Teqoa, 31 ° 38 ’ N, 35 ° 14 ’ E, 650 m a. s. l., 31. iii. 2009, L. Friedman, JSS 51742 (1 ♂, TAU); Nahal Yeelim, 31 ° 14 ’ N, 35 ° 14 ’ E, 20. iv. 2011, A. Freidberg, JSS 51745 (1 ♂, TAU); Nahal Yitav spill, 31 ° 55 ’ N, 35 ° 31 ’ E, 16. iii. 2005, I. Zonstein, JSS 51778 (1 ♂, TAU); L. Friedman, JSS 51779 – 80 (2 ♂, TAU); Nahal Zedim, Bitronot Ruhama, 31 ° 33 ’ N, 34 ° 41 ’ E, 5. iv. 2005, A. Freidberg, JSS 51787 – 8 (2 ♂, TAU); L. Friedman, JSS 51789 (1 ♂, TAU); Nahal lyyon, HaTamur Waterfall, 33 ° 16 ’ N, 35 ° 34 ’ E, 15. iii. 2011, A. Freidberg, JSS 51760 (1 ♂, TAU); Nizzanim Nature Reserve, 31 ° 43 ’ N, 44 ° 36 ’ E, 25 m a. s. l., sand dune swales, 11. v. 2005, M. E. Irwin, hand net, CNCD 157658 (1 ♂, CNC); Nizzanim, D. Dunes, 31 ° 45 ’ N, 34 ° 38 ’ E, 6. iv. 2009, A. Freidberg, light trap, JSS 52099, JSS 51734 – 5 (3 ♂, TAU); Nizzanim, 31 ° 45 ’ N, 34 ° 38 ’ E, 2. vi. 2011, L. Friedman, light trap, JSS 51733 (1 ♂, TAU); Panyas, 33 ° 14 ’ N, 35 ° 40 ’ E, 14. vi. 1996, A. Freidberg, JSS 52094 (1 ♂, TAU); Park Rosh ha Ayin, 16. iv. 1993, A. Freidberg & F. Kaplan, JSS 51766 (1 ♂, TAU); Qalya, 31 ° 45 ’ N, 35 ° 27 ’ E, 1. vi. 1996, I. Yarom & A. Freidberg, JSS 52092 (1 ♂, TAU); Ramot Naftali, 33 ° 06 ’ N, 35 ° 33 ’ E, 13. v. 1998, A. Freidberg, JSS 51708 – 9, JSS 52083 – 4 (4 ♂, TAU); Tel Maresha, Bet Guvrin, 31 ° 36 ’ N, 34 ° 53 ’ E, 30. iii. 2002, A. Freidberg, JSS 51717 (1 ♂, TAU); Tel Qeshet, 31 ° 32 ’ N, 34 ° 45 ’ E, 163 m, 18. iii. 2010, A. Freidberg, JSS 51747 (1 ♂, TAU); Timna, 29 ° 46 ’ N, 34 ° 58 ’ E, 18. iii. 1995, A. Freidberg, JSS 51777 (1 ♂, TAU); Umm el Ghanam, 32 ° 40 ’ N, 35 ° 24 ’ E, on Polygonum arenastrum, 26. v. 2005, L. Friedman, Malaise trap, JSS 51793 (1 ♂, TAU); Wadi Ahmar near Yarden, 32 ° 01 ’ N, 35 ° 30 ’ E, 15. iii. 2005, L. Friedman, JSS 51783 (1 ♂, TAU); Zafririm, 31 ° 39 ’ N, 34 ° 56 ’ E, 30. iii. 2002, A. Freidberg, JSS 51719 (1 ♂, TAU); Zin Wilderness Nakhal Zin at En Akrabim cane-covered sandy wadi, 30 ° 53 ’ N, 35 ° 09 ’ E, - 61 m a. s. l., 7. iv. 1995, M. E. Irwin, Malaise trap, JSS 51701 (1 ♂, TAU); Zomet Mezada, 31. v. 1998, A. Freidberg, JSS 51691 (1 ♂, TAU); Zomet haEla, 12. iv. 2009, L. Freidman, JSS 51731 (1 ♂, TAU); Zomet, Mash abbe Sade, 23. iv. 2009, A. Freidberg, JSS 51761 (1 ♂, TAU); Panyas, 33 ° 14 ’ N, 35 ° 40 ’ E, 16. vii. 1995, A. Freidberg, JSS 52077 (1 ♂, TAU); UNITED ARAB EMIRATES: Wadi Wurayah, 25 ° 24 ’ N, 56 ° 17 ’ E, 11 ‒ 18. v. 2007, Malaise trap, A. van Harten, CNCD 160299 (1 ♂, CNC); Al Wathba Wetland Reserve, 24 ° 15 ’ N, 54 ° 36 ’ E, i. 2015, Malaise trap, A. Saji & A. van Harten, CNC 470760 (1 ♂, CNC); Fujairah, 25 ° 04 ’ N, 56 ° 12 ’ E, 19. iv. ‒ 02. v. 2005, light trap, A. van Harten, CNCD 4278 (1 ♂, CNC); Jebel Hafeet National Park, 24 ° 04 ’ N, 55 ° 46 ’ E, 25. ii. 2018, Malaise trap, A. Saji & A. van Harten, JSS 52263 (1 ♂, EAD).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F68FFA2FF3EEAF4FB594CDD.taxon	distribution	Distribution: Cyprus, Greece, Iran, Israel, Malta, Morocco, Turkey, United Arab Emirates, West and Central Europe (Földvári & De Meyer, 1999; Kehlmaier et al. 2019; Skevington 2020) (Fig. 57). Note: DNA barcodes of T. kuthyi and T. freidbergi are similar. See the notes under T. freidbergi.	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F69FFA4FF3EEF7CFEF3484A.taxon	description	Figs 24 A ‒ D, 58	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F69FFA4FF3EEF7CFEF3484A.taxon	diagnosis	Diagnosis: This species can be recognized by the shape of surstyli in dorsal view, constricted in middle (Fig. 24 A), broadened in basal third in lateral view (Fig. 24 C ‒ D); epandrium longer than wide in dorsal view (Fig. 24 A); sub- epandrial sclerite long and broad in ventral view (Fig. 24 B); gonopods equal in height (Fig. 24 B); phallic guide long, curved to abdominal sternite in lateral view (Fig. 24 C ‒ D). Specimens examined: IRAN: North Khorasan: Darkesh, 37 ° 26 ’ N, 56 ° 44 ’ E, 23. vii. 2016, B. Motamedinia, sweep net, JSS 52031, JSS 52033 – 4 (3 ♂, CNC); Gelian, 37 ° 15 ’ N, 57 ° 55 ’ E, 1283 m a. s. l., 8 – 22. vii. 2016, B. Mota- medinia, Malaise trap, JSS 51983; JSS 51987 (2 ♂, CNC); Ghazi, 37 ° 29 ’ N, 56 ° 44 ’ E, 1200 m a. s. l., 21. vi. 2016, B. Motamedinia, sweep net, JSS 51960 (1 ♂, USNM); South Khorasan: Jafarieh, 32 ° 49 ’ N, 59 ° 14 ’ E, 1760 m a. s. l., 21. iv – 5. v. 2016, B. Motamedinia, Malaise trap, JSS 52012 (1 ♀, CNC); Mohammadieh, 32 ° 52 ’ N, 59 ° 01 ’ E, 1419 m a. s. l., 3 – 17. vii. 2016, B. Motamedinia, Malaise trap; funnel Malaise trap, JSS 52043 – 4; JSS 51973 (3 ♂, CNC); 9. vi. 2015, B. Motamedinia, Malaise trap, JSS 52002 (1 ♀, CNC); Zabol, 31 ° 03 ’ N, 61 ° 22 ’ E, 22. vii. 2016, M. Enaiatnia, Mal- aise trap, JSS 52145 (1 ♂, CNC); ISRAEL: Be’er Sheva, 31 ° 15 ’ N, 34 ° 47 ’ E, 9. vii. 2001, A. Freidberg, JSS 52079 (1 ♂, TAU); Haluza Dunes, 31 ° 04 ’ N, 34 ° 27 ’ E, 165 m a. s. l., 3. iv. 2009, A. Freidberg, JSS 52120 (1 ♂, TAU); Nahal Lavan Rt. 10, 30 ° 51 ’ N, 34 ° 36 ’ E, 17. iv. 1998, A. Freidberg, JSS 51723, JSS 52089, JSS 52091 (3 ♂, TAU); Negev, 1 km South of Midreshet Ben Gurion, Nakhal Khavarim, 30 ° 51 ’ N, 34 ° 47 ’ E, 380 m a. s. l., 7 – 8. vi. 2004, M. E. Irwin, Malaise trap, CNCD 135039 (1 ♂, CNC).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F69FFA4FF3EEF7CFEF3484A.taxon	distribution	Distribution: Belgium, Czech Republic, Denmark, France, Germany, Great Britain, Hungary, Iran, Israel, Italy, Latvia, Mongolia, Morocco, Netherlands, Poland, Romania, Russia, Slovakia, Spain, Sweden, Switzerland (Skevington 2020) (Fig. 58). Note: Based on DNA barcodes, T. minima is most similar to T. minuscula (3.6 % pairwise divergence) (Supplementary Table 1).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F6CFFA7FF3EEBD5FD3E4D3F.taxon	description	Fig. 56	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F6CFFA7FF3EEBD5FD3E4D3F.taxon	diagnosis	Diagnosis: Following Földvári & De Meyer (1999), this species can be recognized by the apically divided phallic guide in lateral view; slightly broadened and elongated surstyli; phallus with three ejaculatory ducts, one bearing some long spines in lateral view; epandrium wider than long (Földvári & De Meyer, 1999: fig. 13 A – F).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F6CFFA7FF3EEBD5FD3E4D3F.taxon	distribution	Distribution: Czech Republic, Finland, Germany, Hungary, Iran, Slovakia, Sweden, Switzerland (Majnon-Jahromi et al. 2018; Skevington 2020) (Fig. 56).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F6CFFA7FF3EE922FC164CD2.taxon	description	Fig. 58	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F6CFFA7FF3EE922FC164CD2.taxon	diagnosis	Diagnosis: Following Földvári & De Meyer (1999), this species can be recognized by the shape of surstyli in dorsal view, broadened at base, constricted before apex; both surstyli bent in middle in lateral view; epandrium wider than long in dorsal view; short phallic guide, broadened at base; phallus with three ejaculatory ducts, one bearing some small teeth in basal half (Földvári & De Meyer, 1999: fig. 15 A – D).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F6CFFA7FF3EE922FC164CD2.taxon	distribution	Distribution: Egypt, France, Hungary, Israel, Japan, Morocco, Romania, Spain, Syria (Skevington 2020) (Fig. 58). Note: Földvári & De Meyer (1999) mentioned that there is some variation in the male genitalia features of this species from Israel (and Hungary) and this might be a species complex.	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F6CFFA7FF3EEF76FCBA4A17.taxon	description	Fig. 53	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F6CFFA7FF3EEF76FCBA4A17.taxon	distribution	Distribution: Croatia, Israel (De Meyer, 1995; Skevington 2020) (Fig. 53). Notes: This species was described based on a female, and the male sex is unknown. The female can be identified by the black halter and long piercer reaching the first abdominal sternite (De Meyer, 1995). It may be synonymous with T. cilifemorata (Földvári & De Meyer (1999).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F6CFFB9FF3EEE3AFF1A4DB5.taxon	description	Figs 25 A – F, 56, 68 A – B	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F6CFFB9FF3EEE3AFF1A4DB5.taxon	diagnosis	Diagnosis: This species can be recognized by the asymmetrical surstyli in dorsal view, broadened in basal, left surstylus knobbed apically, right surstylus narrowed in apical half (Fig. 25 A); gonopods equal in height in ventral view (Fig. 25 B); small phallic guide in lateral view (Fig. 25 D).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F6CFFB9FF3EEE3AFF1A4DB5.taxon	description	Description: MALE: Body length (excluding antennae): 3.9 mm. Head. Scape, pedicel, flagellum, arista dark brown. Pedicel gray pollinose in apical half, with three upper setae and 1 - 2 lower setae. Flagellum gray pollinose and tapering (LF: WF = 3.0); arista dark, with thickened base. Eyes meeting for a distance of 6 ‒ 8 facets. Frons dark silver-gray pollinose. Vertex black, lacking pollinosity. Occiput dark and gray pollinose with scattered short black setae. Thorax. Postpronotal lobe light yellow, gray pollinose with 4 - 6 yellowish setae along the upper margin (up to 0.02 mm). Prescutum, scutum and scutellum black. Scutum with two uniseriate rows of intra-alar setae. Scutellum with 14 – 16 thin short setae along lateral margin (up to 0.02 mm). Subscutellum black, gray pollinose. Pleura black but dark brown on pteropleuron, hypopleuron and sternopleuron. Wing. Length: 3.85 mm. LW: MWW = 5.13. Wing almost entirely covered with microtrichia. Cross-vein r-m reaches dm at the middle. M 1 strongly undulating in middle. Halter length: 0.4 mm. Whitish, narrowly brown at base. Legs. Coxae dark brown, gray pollinose. Mid coxa with four dark anterior setae. Trochanters brown, smooth. Femora dark with light brown apices, gray pollinose. Mid and hind femora bearing two rows of small spines in apical half and with one wrinkled indentation at the base. Tibiae brown but dark at middle, with two ventral rows of short setae on anterior and three rows on posterior side. Hind tibia with 1 – 2 wrinkled indentations in middle. Tarsi light brown with scattered brown setae at anterior margin, basitarsi longer than other tarsomeres. Pulvilli yellow. Claws light brown with black tips. Abdomen. Ground color dark. Tergite 1 densely gray pollinose, with 8 – 10 dark lateral setae (up to 0.11 mm). Tergites 2 – 5 gray pollinose, with scattered brown setae. Sternites light brown laterally and dark brown centrally, gray pollinose. Genitalia. Genital capsule in dorsal view: epandrium and surstyli brown, gray pollinose. Epandrium wider than long (MLE: MWE = 0.5). Surstyli asymmetrical, elongated (Fig. 25 A). Both surstyli broad at base, base of left surstylus triangular-shaped, base of right surstylus square-shaped, left surstylus entirely constricted at middle and knobbed apically, right surstylus narrowed in apical half, left surstylus higher than right one (Fig. 25 A). Genital capsule in ventral view: gonopods wide, equal in height (Fig. 25 B). Subepandrial sclerite small (Fig. 25 B). Genital capsule in lateral view: both surstyli bent in middle towards sternite (Fig. 25 E – F). Phallic guide short, phallus with three ejaculatory ducts (Fig. 25 D). Ejaculatory apodeme rather spade-shaped (Fig. 25 C). FEMALE: unknown.	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F6CFFB9FF3EEE3AFF1A4DB5.taxon	etymology	Etymology: The specific epithet refers to Nimrooz, from where the holotype was collected. Specimen examined: HOLOTYPE: IRAN: Sistan & Baluchestan: Zabol, 31 ° 03 ’ N, 61 ° 22 ’ E, 7. iv. 2016, M. Enaiatnia, Malaise trap, JSS 51819 (1 ♂, CNC).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F6CFFB9FF3EEE3AFF1A4DB5.taxon	distribution	Distribution: Iran (Fig. 56). Notes: T. nimroozensis sp. nov. is not genetically similar to any other known Tomosvaryella species. Tomosvaryella pilosiventris is closest to T. nimroozensis sp. nov. with 12.2 % pairwise difference (Supplementary Table 1).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F72FFBBFF3EE994FE1C4FC9.taxon	description	Figs 26 A – D, 27 I, 47 C, 59	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F72FFBBFF3EE994FE1C4FC9.taxon	diagnosis	Diagnosis: This species can be recognized by a small projection in hind trochanter (Fig. 47 C); epandrium wider than long in dorsal view (Fig. 26 A); syntergosternite 8 slightly rounded in dorsal view (Fig. 26 A); ejaculatory ducts coiled (Fig. 26 C – D); gonopods equal in height (Fig. 26 B); hypandrium slightly rounded, wider than gonopods in ventral view (Fig. 26 B). Based on the shape of surstyli in dorsal and lateral view, this species resembles T. pruinosa Kozánek, 1992. Tomosvaryella pruinosa lacks the small projection in hind trochanter. In addition, there is a gap between the surstyli and cerci of T. nodosa in lateral view (Fig. 26 C – D). Meanwhile, It is similar to T. hamounensis sp. nov. See the diagnosis notes under T. hamounensis sp. nov. for more details. Specimens examined: IRAN: South Khorasan: Gholonabad, 33 ° 10 ’ N, 59 ° 22 ’ E, 1914 m a. s. l., 11. v. 2015, B. Motamedinia, Malaise trap, JSS 51962 (1 ♂, CNC); ISRAEL: Arava Valley, 0.2 km N. Hazeva Field School, small wadi, 30 ° 46 ’ N, 35 ° 14 ’ E, - 116 m a. s. l., 30. iii. 1995, M. E. Irwin, Malaise trap, JSS 52061 (1 ♂, TAU); Arava Valley, Iddan, wadi running east of date palm orchard, 30 ° 48 ’ N, 35 ° 16 ’ E, - 110 m a. s. l., 16. iv. 1995, M. E. Irwin, Malaise trap, JSS 52052 (1 ♂, TAU); 17. iii. 1995, M. E. Irwin, Malaise trap, JSS 52054 (1 ♂, TAU); 24. iii. 1995, M. E. Irwin, Malaise trap, JSS 51699 (1 ♂, TAU); 31. iii. 1995, M. E. Irwin, Malaise trap, JSS 52053 (1 ♂, TAU); Arava Valley, Moshaav Hazeva, Wadi Shahak between agricultural fields, 30 ° 46 ’ N, 35 ° 16 ’ E, - 110 m a. s. l., 12. iii. 1995, E. Irwin, Malaise trap, JSS 51622 (1 ♂, TAU); 15. iii. 1995, M. E. Irwin, Malaise trap, JSS 52060 (1 ♂, TAU); 19. iii. 1995, M. E Irwin, Malaise trap, JSS 51730 (1 ♂, TAU); Arava Valley, Shizaf Nature Reserve, Hazeva, north of water treatment plant low sandy hummocks in small wadi, 30 ° 46 ’ N, 35 ° 15 ’ E, - 80 m a. s. l., 16. iv. 1995, M. E. Irwin, JSS 51698 (1 ♂, TAU); Arava Valley, small wadi, 2.4 km W. of hwy. 90 at km. 149, 30 ° 42 ’ N, 35 ° 11 ’ E, - 60 m a. s. l., 16. iv. 1995, M. E. Irwin, hand net, JSS 52058 (1 ♂, TAU); Arava Valley, Iddan Springs, in damp seep, east of spring, 30 ° 49 ’ N, 35 ° 16 ’ E, - 116 m a. s. l., 21. iii. 1995, M. E. Irwin, JSS 51811 (1 ♂, TAU); 31. iii. 1995, M. E. Irwin, JSS 51812 (1 ♂, TAU); Haze- va Field School, 30 ° 43 ’ N, 35 ° 15 ’ E, 2. xi. 1997, S. Plotkin, Malaise trap, JSS 52096 (1 ♂, TAU); Nahal Eshharim, 30 ° 28 ’ N, 34 ° 35 ’ E, 790 m a. s. l., 2. v. 1995, I. Yarom, Malaise trap, JSS 51790 (1 ♂, TAU); Nahal Mishmar, 31 ° 24 ’ N, 35 ° 20 ’ E, 11. iv. 1994, A. Freidberg & F. Kaplan, JSS 51746 (1 ♂, TAU); B. S. Hazeva, 13. iii. 1998, S. Alfi, JSS 51771 (1 ♂, TAU); Sniai Ofira, 27 ° 52 ’ N, 34 ° 17 ’ E, 22. iii. 1981, A. Freidberg, JSS 51721 (1 ♂, TAU); Taba, 29 ° 30 ’ N, 34 ° 53 ’ E, 14. v. 2010, A. Freidberg, JSS 51764 (1 ♂, TAU); Zin Wilderness Nakhal Zin at En Akrabim cane-covered sandy wadi, 30 ° 53 ’ N, 35 ° 09 ’ E, - 61 m a. s. l., 7. iv. 1995, M. E. Irwin, Malaise trap, JSS 51700 (1 ♂, TAU); UNITED ARAB EMIR- ATES: Abu Dhabi: Al Wathba Wetland Reserve, 24 ° 15 ’ N, 54 ° 37 ’ E, i. 2015, A. Saji & A. van Harten, Malaise trap, CNC 470786, CNC 470791, CNC 470801 (3 ♂, CNC); ii. 2015, A. Saji & A. van Harten, Malaise trap, CNC 470524, CNC 470585, CNC 470508, CNC 470473, CNC 470465, CNC 470545, CNC 470448 – 9 CNC 470498 (4 ♂, 5 ♀, CNC); iii. 2015, A. Saji & A. van Harten, Malaise trap, CNC 470669, CNC 470688, CNC 470648 (2 ♂, 1 ♀, CNC); ix. 2013, A. Saji & A. van Harten, Malaise trap, CNC 470711 – 2 (2 ♂, CNC); al-Ajban, 24 ° 22 ’ N, 55 ° 01 ’ E, 5 – 12. vi. 2006, A. van Harten, Malaise trap, CNCD 159561 (1 ♂, USNM); Al Bida’a Protected Area, 23 ° 36 ’ N, 53 ° 58 ’ E, 29. x. 2018, A. Saji & A. van Harten, Malaise trap, JSS 52291 – 4 (4 ♂, CNC); Barqa Al Suqoor Protected Area, 23 ° 60 ’ N, 52 ° 41 ’ E, 29. iii. 2018, A. Saji & A. van Harten, Malaise trap, JSS 52280 – 1 (1 ♂, 1 ♀, CNC); 29. x. 2018, A. Saji & A. van Harten, Malaise trap, JSS 52283 (1 ♂, USNM); Wadi Shawkah, 25 ° 04 ’ N, 56 ° 01 ’ E, 5 – 12. v. 2007, A. van Harten, pan trap, JSS 52125 (1 ♂, EAD).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F72FFBBFF3EE994FE1C4FC9.taxon	distribution	Distribution: Egypt, Iran, Israel, United Arab Emirates (Skevington 2020) (Fig. 59). Note: Based on DNA barcodes, T. nodosa is similar to T. pruinosa, differing by 4.2 %. (pairwise divergence) (Supplementary Table 1).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F70FFBBFF3EEA08FB594889.taxon	description	Figs 27 G, 28 A – E, 59, 68 C – D	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F70FFBBFF3EEA08FB594889.taxon	diagnosis	Diagnosis: The male of this species can be recognized by the shape of surstyli in dorsal view, broad basically, constricted in apical third (Fig. 28 A); large gonopods in ventral view (Fig. 28 B); broad phallic guide; ejaculatory ducts strictly bent into phallic guide in lateral view (Fig. 28 C). The female can be recognized by long ovipositor, curved and reaching the mid coxa (Fig. 27 G).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F70FFBBFF3EEA08FB594889.taxon	description	Description: MALE: Body length (excluding antennae): 4.5 mm. Head. Scape, pedicel, flagellum and arista dark. Pedicel with three upper dark setae; flagellum light brown, gray pollinose and tapering (LF: WF = 1.7). Eyes meeting for a distance of 13 ‒ 14 facets. Frons silver-gray pollinose; vertex shining; occiput gray pollinose with scattered light brown setae. Thorax. Postpronotal lobe light brown with pale setae along the upper margin (up to 0.01 mm). Prescutum and scutum black, light brown pollinose; Scutum with two dark uniseriate rows of intra-alar setae. Scutellum black, silver-gray pollinose, with 12 setae along posterior margin. Subscutellum black, pleura dark brown. Wing. Wing length: 2.4 mm. LW: MWW = 3.4. Wing almost entirely covered with microtrichia. M 1 strongly undulating in middle. Halter length: 0.5 mm. Brown to yellowish, narrowly black at base. Legs. Coxae and trochanters black, gray pollinose, mid coxa with three black anterior setae; hind trochanter with scattered short pale bristle. Femora dark brown with narrow yellow apices, silver pollinose and 5 ‒ 8 ventral spines towards apex. All femora with one wrinkled indentation in basal. Tibiae light brown at base, gray pollinose, with two rows of short black setae on anterior and three rows on posterior side. Hind tibia with one wrinkled indentation in middle. Tibiae without distal spines. Tarsi light brown, distitarsi dark, claws light brown with black tips. Abdomen. Ground color brown, tergite 1 covered by densely silver-gray pollinose, with eight short dark lateral setae, tergite 2 ‒ 5 with weak silvergray pollinose and with scattered brown setae. Sternites white-yellow laterally with dark mid-line centrally, gray pollinose. Genitalia. Genital capsule in dorsal view: surstyli asymmetrical, both with broad base and constricted in apical third but base of the right boarder than left one, left surstyli knobbed apically, right surstyli curved toward left one apically (Fig. 28 A), epandrium wider than long. Genital capsule in ventral view: gonopods large, broadened, equal in height (Fig. 28 B). Genital capsule in lateral view: both surstyli broadened at base, strictly bent in apical third towards sternite (Fig. 28 D ‒ E); phallic guide broad, pointed; phallus wavy, bent into phallic guide (Fig. 28 C). FEMALE: Body length (excluding antennae): 3.6 mm. Head. Eyes separated. Frons silver-gray pollinose, concaved shortly after ocelli to lower third of frons. Occiput gray pollinose. Pedicel with four upper dark setae and one lower setae. Thorax. Postpronotal lobe light yellow, gray pollinose, with 4 – 5 light brown setae along the upper margin (up to 0.01 mm). Prescutum and scutum, scutellum black, light brown pollinose in dorsocentral area. Wing. Length: 3.5 mm. LW: MWW = 3.8. Legs. Hind trochanter with 6 – 8 light brown setae. All femora bearing one wrinkled indentation in basal. Fore femur more widened. Hind trochanter with two wrinkled indentations in middle. Distitarsi with long brown setae at anterior margin. Pulvilli broadened and leaf-shaped; claws long, twice the length of distitarsi. Abdomen. Tergites 1 – 6 gray pollinose with scattered short light brown setae. Ovipositor. Base of ovipositor dark brown, gray pollinose dorsally, with some scattered short pale setae. Viewed laterally (Fig. 27 G), base of piercer curved, piercer long and angled between proximal and distal part and reached mid coxa. LP: LB = 2.5. LDP: LPP = 4.0.	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F70FFBBFF3EEA08FB594889.taxon	etymology	Etymology: The specific epithet refers to Oshida, the old name of Khaje mountain and a female name in mythology, from where the holotype specimen was collected. Specimens examined: HOLOTYPE: IRAN: Sistan & Baluchestan: Zabol, 31 ° 07 ’ N, 61 ° 28 ’ E, 481 m a. s. l., 6. vi. 2016, H. Derafshan, sweep net, JSS 51818 (1 ♂, CNC); PARATYPE: IRAN: South Khorasan: Mohammadieh, 32 ° 52 ’ N, 59 ° 01 ’ E, 1419 m a. s. l., 3 – 17. vii. 2016, B. Motamedinia, funnel Malaise trap, JSS 51976 (1 ♀, CNC).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F70FFBBFF3EEA08FB594889.taxon	distribution	Distribution: Iran (Fig. 59). Note: Based on DNA barcodes, T. oshidae sp. nov. is genetically most similar to T. pseudophanes, with 11.1 % pairwise difference. The male and female sequenced differ by only 0.17 % (Supplementary Table 1).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F77FFBFFF3EEBD5FDAA4B21.taxon	description	Figs 29 A ‒ E, 58, 68 E – F	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F77FFBFFF3EEBD5FDAA4B21.taxon	diagnosis	Diagnosis: This species can be recognized by the asymmetrical shape of surstyli in dorsal view, both long, left surstylus bone-shaped and longer than right one (Fig. 29 A); unequal gonopods, right one with a projection towards phallic guide (Fig. 29 B); strong and straight phallus, three short ejaculatory ducts, one bearing some teeth; membranous sheaths bearing saw-like teeth (Fig. 29 B, D ‒ E).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F77FFBFFF3EEBD5FDAA4B21.taxon	description	Description: MALE: Body length: 2.1 mm. Head. Frons silver-gray pollinose. Eyes meeting for a distance of five facets. Pedicel with 1 ‒ 2 small dorsal setae; flagellum tapering, distinctly paler than pedicel (LF: WF = 2.3). Postpronotal lobe pale, gray pollinose. Prescutum and scutum brown, silver-gray pollinose, with some patches of small supra-alar setae behind postpronotal lobes. Scutellum brown, silver-gray pollinose. Subscutellum darker than pleura. Wing. Wing length: 2.6 mm. LW: MWW = 2.4. Wing almost entirely covered with microtrichia except at wing base. Cross-vein r-m reaching cell dm at the middle. Halter length 0.4 mm., yellow, narrowly brown at base. Legs. Light brown. Trochanters black, smooth. Femora with posteroventral and anteroventral row of 3 ‒ 8 black spines on apical half. Tibiae with two rows of short brown setae on anterior and three rows on posterior side. Distitarsi darker that tarsi; pulvilli shorter than distitarsi. Claws white with black tips. Abdomen. Ground color brown, gray pollinose; tergite 1 with four brown lateral setae, tergites normally covered with short but scattered brown hairs. Syntergosternite 8 brown. Genitalia. Genital capsule in dorsal view: epandrium and surstyli dark brown, epandrium wider than long (MLE: MWE = 0.8). Surstyli long and asymmetrical. Left surstylus slightly larger than right surstylus, bone-shaped. Right surstylus broadened at base and narrowed at apex, curved toward left surstylus in apical third (Fig. 29 A). Genital capsule in ventral view: subepandrial sclerite wide, gonopods unequal; right with a projection curved towards phallic guide, lobes of hypandrium sclerotized, giving it the shape of an inverted triangle (Fig. 29 B). Phallus strong and straight, with three short ejaculatory ducts, one bearing some saw-like teeth along its side (Fig. 29 B). Genital capsule in lateral view: membranous sheath bearing saw-like teeth (Fig. 29 D ‒ E). Phallic guide thickened at base. Ejaculatory apodeme tube-like with a bulb in its middle (Fig. 29 C). FEMALE: unknown.	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F77FFBFFF3EEBD5FDAA4B21.taxon	etymology	Etymology: The specific epithet derived from the Greek “ osteodes ” (= bone like), referring to the shape of left surstylus. Specimen examined: HOLOTYPE: UNITED ARAB EMIRATES: Wadi Wurayah, 25 ° 24 ’ N, 56 ° 17 ’ E, 25. ii. 2007, sweep net, F. Menzel & A. Stark, CNCD 175298 (1 ♂, CNC).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F77FFBFFF3EEBD5FDAA4B21.taxon	distribution	Distribution: United Arab Emirates (Fig. 58). Note: Based on DNA barcodes, this species is most similar to T. sylvatica, differing by 8.5 % (pairwise divergence) (Supplementary Table 1).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F74FFB1FF3EEF20FE1C498D.taxon	description	Figs 27 F, 30 A ‒ D, 60	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F74FFB1FF3EEF20FE1C498D.taxon	diagnosis	Diagnosis: This species can be recognized by elongated surstyli and epandrium in dorsal view (Fig. 30 A); unequal gonopods, left is higher than right one in ventral view (Fig. 30 B); both surstyli slightly straight, bent towards sternite in apical third in lateral view (Fig. 30 C – D). Based on the shape of surstyli in dorsal view, this species resembles species in the kuthyi group (Tomosvaryella cyprusensis sp. nov., T. emaratensis sp. nov., T. freidbergi, T. hispanica, T. israelensis, T. kuthyi). It differs by the shape of surstyli in lateral view, bent towards sternite in apical third (Fig. 30 C – D). See diagnosis of T. emaratensis sp. nov. and T. cyprusensis sp. nov. for more details. Specimens examined: HOLOTYPE: ISRAEL: Sinai Ofira, 22 iii. 1981, A. Freidberg, JSS 52379 (1 ♂, TAU); PARATYPE: ISRAEL: Sinai, 20 Km N. Dahab, 12. iii. 1982, A. Freidberg, JSS 52380 (1 ♂, TAU); Other specimens examined: IRAN: Kermanshah: Dodan, 35 ° 00 ’ N, 46 ° 12 ’ E, 1011 m a. s. l., Garden, 20. vi. 2016, M. Zardouei, Malaise trap, JSS 52186 (1 ♂, CNC); 5. vii. 2016, M. Zardouei, funnel Malaise trap, JSS 51951 (1 ♂, CNC); North Khorasan: Kohnekand, 37 ° 51 ’ N, 57 ° 31 ’ E, 1200 m a. s. l., 13 – 27. vi. 2016, B. Motamedinia, Malaise trap, JSS 51823 (1 ♂, CNC); Biar, 37 ° 53 ’ N, 57 ° 35 ’ E, 1008 m a. s. l., 10 ‒ 24. vi. 2016, B. Motamedinia, Malaise trap, JSS 52016 (1 ♂, CNC); Sistan & Baluchestan: Rask, 26 ° 16 ’ N, 61 ° 25 ’ E, 139 m a. s. l., 12. vi – 14. vii. 2016, M. Ghaforimoghadam, Malaise trap, JSS 52146 (1 ♂, CNC); ISRAEL: Arava Valley, Iddan, wadi running east of date palm orchard, 30 ° 48 ’ N, 35 ° 16 ’ E, - 110 m a. s. l., 21. iii. 1995, M. E. Irwin, Malaise trap, JSS 51813 (1 ♂, TAU); 24. iii. 1995, M. E. Irwin, Malaise trap, JSS 52055 (1 ♂, TAU); 31. iii. 1995, M. E. Irwin, Malaise trap, JSS 51815 (1 ♂, TAU); Arava Valley, Shizaf Nature Reserve, Hazeva, north of water treatment plant low sandy hummocks in small wadi, 30 ° 46 ’ N, 35 ° 15 ’ E, - 80 m a. s. l., 16. iv. 1995, M. E. Irwin, JSS 51697 (1 ♂, TAU); Arava Valley, Shizaf Nature Reserve, Hazeva, north of water treatment plant, low sandy hummocks in small wadi, 30 ° 46 ’ N, 35 ° 15 ’ E, - 80 m a. s. l., 3. iv. 1995, M. E. Irwin, JSS 52057 (1 ♂, TAU); Arava Valley, small wadi, 2.4 km. W. of hwy. 90 at km 149, 30 ° 42 ’ N, 35 ° 11 ’ E, - 60 m a. s. l., 16. iv. 1995, M. E. Irwin, hand net, JSS 51696 (1 ♂, TAU); Arava Valley, En Yahav Makhteshim Res., En Shahak, 2.5 km. w. hwy 90 at km 149, 30 ° 42 ’ N, 35 ° 11 ’ E, - 60 m a. s. l., 16. iv. 1995, M. E Irwin, hand net, JSS 51729 (1 ♂, TAU); 23. iii. 1995, M. E Irwin, Malaise trap, JSS 51728 (1 ♂, TAU); Arava Valley, Iddan, in damp seep, east of spring, 30 ° 49 ’ N, 35 ° 16 ’ E, - 116 m a. s. l., 21. iii. 1995, M. E. Irwin, JSS 52106 (1 ♂, TAU); Arava Valley, Iddan, wadi running east to date palm orchard, 30 ° 48 ’ N, 35 ° 16 ’ E, - 110 m a. s. l., 17. iii. 1995, M. E Irwin, Malaise trap, JSS 51726 (1 ♂, TAU); Avenat, Rt. 90, 31 ° 41 ’ N, 35 ° 26 ’ E, 17. iii. 2004, A. Freidberg, JSS 51768 (1 ♂, TAU); Beer Sheva, 31 ° 13 ’ N, 34 ° 48 ’ E, 1. x. 2001, A. Freidberg, JSS 51718 (1 ♂, TAU); En Gev, 32 ° 46 ’ N, 35 ° 38 ’ E, 7. v. 1997, A. Freidberg, JSS 51715 (1 ♂, TAU); En Zn, 28. ix. 1999, A. Freidberg, JSS 51782 (1 ♂, TAU); Har Hermon, 33 ° 18 ’ N, 35 ° 46 ’ E, 1600 m a. s. l., 18. v. 2009, A. Freidberg, JSS 51763 (1 ♂, TAU); 3. ix. 2008, A. Freidberg, JSS 51762 (1 ♂, TAU); Maalot, 33 ° 00 ’ N, 35 ° 17 ’ E, 580 m a. s. l., 25. v. 2009, A. Freidberg, JSS 51740 (1 ♂, TAU); Maoz Hayim, 32 ° 29 ’ N, 35 ° 32 ’ E, 23. x. 1978, A. Freidberg, JSS 52105 (1 ♂, TAU); Nahal Oren, 32 ° 43 ’ N, 35 ° 01 ’ E, 30. v. 1998, A. Freidberg, JSS 51711 (1 ♂, TAU); Nahal Yitav spill, 31 ° 55 ’ N, 35 ° 31 ’ E, 16. iii. 2005, L. Friedman, JSS 52086 (1 ♂, TAU); Paran, 30 ° 21 ’ N, 35 ° 09 ’ E, 9. iv. 2002, A. Freidberg, JSS 52078 (1 ♂, TAU); Park Timna, Gate, 29 ° 47 ’ N, 34 ° 59 ’ E, 145 m a. s. l., 13. v. 2010, A. Freidberg, JSS 52066 (1 ♂, TAU); Sinai Mts. Wadi Geragenia, 2000 m a. s. l., 16. vii. 1974, A. Freidberg, JSS 52116 (1 ♂, TAU); Zin Wilderness Nakhal Zin at En Akrabim cane covered sandy wadi, 30 ° 53 ’ N, 35 ° 09 ’ E, - 61 m a. s. l., 21. iii. 1995, M. E. Irwin, Malaise trap, JSS 52063 (1 ♂, TAU); Nahal Eshharim, 30 ° 28 ’ N 34 ° 35 ’ E, 790 m a. s. l., 2. v. 1995, I. Yarom, Malaise trap, JSS 51792 (1 ♂, TAU); Nahal Parsa, 31 ° 13 ’ N 35 ° 20 ’ E, 13. iv. 2009, A. Freidberg, JSS 52082 (1 ♂, TAU); UNITED ARAB EMIRATES: Abu Dhabi: Al Wathba Wetland Reserve, 24 ° 15 ’ N, 54 ° 37 ’ E, i. 2015, A. Saji & A. van Harten, Malaise trap, CNC 465506, CNC 470736 ‒ 37, CNC 470740 ‒ 41, CNC 470748 ‒ 9, CNC 470753 ‒ 4, CNC 470769, CNC 470771, CNC 470774, CNC 470779 ‒ 80, CNC 470782 ‒ 3, CNC 470788, CNC 470792, CNC 470797 ‒ 8, CNC 470806 (21 ♂, CNC); ii. 2015, A. Saji & A. van Harten, Malaise trap, CNC 465507, CNC 470450, CNC 470452, CNC 470454, CNC 470457 ‒ 9, CNC 470461 ‒ 2, CNC 470464, CNC 470466 ‒ 8, CNC 470474, CNC 470478, CNC 470481, CNC 470484 ‒ 7, CNC 470490, CNC 470495, CNC 470496, CNC 470499, CNC 470501, CNC 470506, CNC 470509, CNC 470511, CNC 470513 ‒ 5, CNC 470519 ‒ 20, CNC 470522, CNC 470527 ‒ 30, CNC 470533, CNC 470535, CNC 470541, CNC 470547, CNC 470553 ‒ 6, CNC 470560, CNC 470562, CNC 470566, CNC 470568 ‒ 9, CNC 470571, CNC 470573 ‒ 5, CNC 470577, CNC 470580, CNC 470584, CNC 470590, CNC 470592, CNC 470598, CNC 470601 ‒ 2 (62 ♂, 1 ♀, CNC); iii. 2015, A. Saji & A. van Harten, Mal- aise trap, CNC 470623 ‒ 4, CNC 470636 ‒ 8, CNC 470640, CNC 470653, CNC 470666, CNC 470672, CNC 470693 ‒ 4, CNC 470698, CNC 470634, CNC 470621, CNC 470701 (14 ♂, 1 ♀, CNC); iv. 2013, A. Saji & A. van Harten, Malaise trap, CNC 470731, CNC 470733 (2 ♂, USNM); iv. 2015, A. Saji & A. van Harten, Malaise trap, CNC 470833 ‒ 4, CNC 470842 (3 ♂, CNC); ix. 2013, A. Saji & A. van Harten, Malaise trap, CNC 470714, CNC 470717 (2 ♂, USNM); Wadi Wurayah farm, 25 ° 14 ’ N, 56 ° 11 ’ E, 19. iv – 19. v. 2009, A. van Harten, Malaise trap, CNCD 137557 (1 ♂, CNC); 31. v – 14. vi. 2009, A. van Harten, Malaise trap, CNCD 137569, CNCD 137571 (2 ♂, CNC); Wadi Wurayah, 25 ° 14 ’ N, 56 ° 10 ’ E, 11 – 18. v. 2007, A. van Harten, Malaise trap, CNCD 160298 (1 ♂, CNC); near ad-Dhaid, 25 ° 13 ’ N, 55 ° 35 ’ E, 16 – 19. iii. 2007, J. Btelka, pan trap, CNCD 159475 (1 ♂, CNC); Fujairah, 25 ° 05 ’ N, 56 ° 13 ’ E, 20 – 27. v. 2006, A. van Harten, light trap, CNCD 160289 (1 ♂, CNC); 14 – 25. i. 2006, A. van Harten, white & yellow pan trap, CNCD 8961 ‒ 2 (2 ♂, CNC); Wadi Bih dam, 25 ° 48 ’ N, 56 ° 04 ’ E, 13 ‒ 21. iv. 2008, light trap, A. van Harten, JSS 52254 (1 ♂, EAD); Houbara Protected Area, 24 ° 05 ’ N, 52 ° 58 ’ E, 15. iii. 2017, Malaise trap, A. Saji & A. van Harten, JSS 52272 (1 ♂, EAD); Wadi Maidaq, 24 ° 10 ’ N, 56 ° 04 ’ E, 3 ‒ 17. ii. 2008, A. van Harten, pan trap, JSS 52256 (1 ♂, CNC); Al-Ajban, 24 ° 21 ’ N, 55 ° 00 ’ E, A. van Harten, Malaise trap, CNCD 159295 (1 ♀, CNC); YEMEN: 12 km NW of Manakhah, 15 ° 04 ’ N 43 ° 44 ’ E, 24. vi – 4. viii. 2003, Malaise trap, CNCD 6824 (1 ♂, CNC).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F74FFB1FF3EEF20FE1C498D.taxon	distribution	Distribution: Canary Island, Cyprus, Egypt, Iran, Israel, North Africa, United Arab Emirates, Yemen (Kehlmaier & Majnon Jahromi, 2015; De Meyer, 1995; De Meyer et al. 2000) (Fig. 60). Note: DNA barcodes of T. parakuthyi are most similar to T. kuthyi, differing by 2.3 % (pairwise divergence) (Supplementary Table 1).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F7AFFB3FF3EEC4CFA2F4CF9.taxon	description	Figs 27 B, 31 A – E, 47 E, 61	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F7AFFB3FF3EEC4CFA2F4CF9.taxon	diagnosis	Diagnosis: This species can be recognized by the asymmetrical and irregular-shaped surstyli in dorsal view (Fig. 31 A); epandrium wider than long in dorsal view (Fig. 31 A); phallus trifid (Fig. 31 D – E). The surstyli shape is similar to T. subvirescens (Loew, 1872). It differs from this species by the shape of left surstylus in dorsal view, wrinkled and wavy-shaped (Fig. 31 A) and triangular-shaped projections on the hind trochanter (Fig. 47 E). Specimens examined: IRAN: Kermanshah: Ghazanchi, 34 ° 26 ’ N, 47 ° 00 ’ E, 1304 m a. s. l., 1. vii. 2015, M. Zardouei, Malaise trap, JSS 51953 (1 ♂, CNC); North Khorasan: Ghazi, 37 ° 29 ’ N, 56 ° 44 ’ E, 1200 m a. s. l., 21. vi. 2016, B. Motamedinia, sweep net, JSS 51961 (1 ♀, CNC); Sistan & Baluchestan: Iranshahr, 27 ° 24 ’ N, 60 ° 50 ’ E, 769 m a. s. l., 2 – 12. v. 2016, M. Ghaforimoghadam, Malaise trap, JSS 52165 (1 ♀, CNC); Zabol, 31 ° 07 ’ N, 61 ° 28 ’ E, 481 m a. s. l., 6. vi. 2016, H. Derafshan, sweep net, JSS 51875 ‒ 6 (2 ♀, CNC); South Khorasan: Mohammadieh, 32 ° 52 ’ N, 59 ° 01 ’ E, 1419 m a. s. l., 3 – 17. vii. 2016, B. Motamedinia, funnel Malaise trap, JSS 51975 (1 ♀, CNC); Salmanefar- si, 32 ° 30 ’ N, 59 ° 13 ’ E, 2. vi. 2016, B. Motamedinia, sweep net, JSS 52028 (1 ♂, CNC); ISRAEL: Hof Rotem She- zaf, 32 ° 46 ’ N, 35 ° 38 ’ E, - 200 m, 21. iii. 2010, A. Freidberg, Malaise trap, JSS 52064 ‒ 5 (2 ♂, TAU); Kinneret, Zee- lon, Beach, 32 ° 50 ’ N, 35 ° 39 ’ E, - 190 m a. s. l., 28. vi. 2011, A. Freidberg, JSS 51737 ‒ 9 (3 ♂, TAU); Park haYarden, 32 ° 50 ’ N, 35 ° 39 ’ E, - 192 m a. s. l., 28. vi. 2011, L. Bodner, JSS 43963 (1 ♀, TAU); RT. 90, nr. En Admon, 31 ° 01 ’ N, 35 ° 22 ’ E, 11. iv. 1994, A. Freidberg & F. Kaplan, JSS 51748 ‒ 9 (2 ♂, TAU); UNITED ARAB EMIRATES: Bithnah, 25 ° 06 ’ N, 56 ° 08 ’ E, 16. xi – 26. xii. 2006, A. van Harten, Malaise trap, JSS 52242 (1 ♂, CNC); 19. x – 16. xi. 2006, A. van Harten, Malaise trap, CNCD 11525 (1 ♂, CNC); 4. vii – 12. viii. 2006, A. van Harten, CNCD 159940 (1 ♂, CNC); Wadi Maidaq, 25 ° 11 ’ N, 56 ° 04 ’ E, 26. xii. 2006 – 20. ii. 2007, A. van Harten, Malaise trap, JSS 52252 (1 ♂, CNC); 27. vi – 29. vii. 2006, A. van Harten, Malaise trap, CNCD 159304 (1 ♂, CNC); Wadi Safad, 25 ° 07 ’ N, 56 ° 11 ’ E, 26. xii. 2005 – 2. i. 2006, A. van Harten, yellow & white pan traps, CNCD 8965, CNCD 8971 (2 ♂, CNC); Wadi Shawkah, 25 ° 04 ’ N, 56 ° 01 ’ E, 18. iii. 2007, F. Menzel & A. Stark, sweep net, JSS 52246 (1 ♂, EAD); Wadi Shawwkah, 25 ° 04 ’ N, 56 ° 01 ’ E, 31. x – 27. xi. 2006, A. van Harten, pan trap, CNCD 159615 (1 ♂, CNC); Wadi Wurayah farm, 25 ° 14 ’ N, 56 ° 11 ’ E, 19. iv – 19. v. 2009, A. van Harten, Malaise trap, CNCD 137547 (1 ♀, CNC); near ad-Dhaid, 25 ° 13 ’ N, 55 ° 35 ’ E, 16 – 19. iii. 2007, J. Btelka, pan trap, CNCD 159474 (1 ♂, USNM).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F7AFFB3FF3EEC4CFA2F4CF9.taxon	distribution	Distribution: Egypt, Iran, Israel, Kenya, South Africa, Spain (Canary Islands), Syria, Turkey, United Arab Emirates, West Bank, Zimbabwe (Kehlmaier et al. 2019; Skevington 2020) (Fig. 61). Notes: DNA barcodes of T. pilosiventris overlap with T. subvirescens (0.3 – 1.9 % pairwise divergence). Despite this, the genitalia of these species are rather different. This is thus likely a case of recently diverged species, as seen in many other taxa (e. g. Skevington et al. 2007, Young et al. 2016, Motamedinia et al. 2020). There is always a possibility that it is a single species with polymorphic genitalia, so future genetic work may shed light on this issue.	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F78FFB3FF3EEF58FC2549D1.taxon	description	Figs 32 A – E, 59	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F78FFB3FF3EEF58FC2549D1.taxon	diagnosis	Diagnosis: This species can be recognized by the shape of surstyli, broadened at base and gradually tapering toward apices in dorsal view (Fig. 32 A), both surstyli bent by 90 ˚ towards abdominal sternite in lateral view (Fig. 32 D – E); epandrium small and wider than long in dorsal view (Fig. 32 A); phallic guide short, broadened at base; phallus with three ejaculatory ducts, one bearing some small teeth in basal half (Fig. 32 D); gonopods and hypandrium symmetrical in ventral view (Fig. 32 B). The shape of the surstyli and small epandrium in dorsal view are similar to T. congoana Hardy, 1950 (for illustration see De Meyer, 1993: fig. 10), T. debruyni De Meyer, 1995 (for illustration see De Meyer, 1995: fig. 11) and T. mutata (for illustration see Földvári & De Meyer, 1999: 15 A – D). It differs from T. congoana and T. debruyni by the shape of surstyli in lateral view, which is bent by 90 ˚ towards the abdominal sternites and differs from T. mutata by the shape of the right surstylus in dorsal view. It is also similar to T. angulata Majnon-Jahromi et al. 2017, as the surstyli of both species are bent by 90 ˚ towards the abdominal sternite in lateral view and both have a small epandrium in dorsal view. It differs from T. angulata by the lack of distinct and distally-knobbed surstyli in dorsal view (Fig. 32 B). Moreover, this species is closely related to T. argyrata and T. argyratoides. See the diagnosis notes under T. argyrata for more details. Specimens examined: CYPRUS: Kyrenia, 35 ° 21 ’ N, 33 ° 09 ’ E, 1 – 8. x. 2017, O. Ozden, Malaise trap, JSS 52304 (1 ♂, CNC); ISRAEL: Har Hermon, 33 ° 18 ’ N, 35 ° 46 ’ E, 2000 m a. s. l., 12. vi. 1996, A. Freidberg, JSS 52095 (1 ♂, TAU).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F78FFB3FF3EEF58FC2549D1.taxon	distribution	Distribution: Cyprus, Iran, Israel (Skevington 2020) (Fig. 59).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F78FFB2FF3EEC70FD904FC9.taxon	description	Fig. 60	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F78FFB2FF3EEC70FD904FC9.taxon	distribution	Distribution: Iran (Skevington 2020) (Fig. 60). Notes: This species has been recorded by Becker (1913) from Iran (Sistan & Baluchestan) but has not been completely illustrated yet. Type material (one male and one female) are currently on loan to Sergey Kuznetzov and unavailable. Majnon-Jahromi et al. (2018) mentioned its distribution in Alborz, Iran but it has been reported only in Siatan & Baluchestan province so far.	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F7EFFB4FF3EEBD5FD914DC0.taxon	description	Figs 27 C, 33 A – E, 62	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F7EFFB4FF3EEBD5FD914DC0.taxon	diagnosis	Diagnosis: This species can be recognized by the shape of surstyli in lateral view; both bean-shaped (Fig. 33 D – E); epandrium wider than long in dorsal view (Fig. 33 A); coiled ejaculatory ducts (Fig. 33 D – E); gonopods equal in height (Fig. 33 B); rounded hypandrium with distinct hypandrial apodeme in ventral view (Fig. 33 B). This species is closely related to T. nodosa and T. hamounensis sp. nov. See the diagnosis notes under T. nodosa and T. hamounensis sp. nov. for more details. Specimens examined: IRAN: Kermanshah: Sarpolezahab, 34 ° 28 ’ N, 45 ° 49 ’ E, 456 m a. s. l., 1. vi. 2015, M. Zardouei, Malaise trap, JSS 51831; JSS 51834 (2 ♂, CNC); 456 m a. s. l., 14. v. 2016, M. Zardouei, Malaise trap, JSS 52235 (1 ♀, CNC); Sistan & Baluchestan: Zabol, Dostmohammad, 31 ° 09 ’ N, 61 ° 43 ’ E, 470 m a. s. l., 13. ix. 2016, M. Enaiat- nia, Malaise trap, JSS 51931 (1 ♂, CNC); Zabol, 31 ° 03 ’ N, 61 ° 22 ’ E, 7. x. 2016, M. Enaiatnia, Malaise trap, JSS 52144, JSS 52155, JSS 52174 (3 ♂, CNC); South Khorasan: Mohammadieh, 32 ° 52 ’ N, 59 ° 01 ’ E, 1419 m a. s. l., 15. iv. 2015, B. Motamedinia, Malaise trap, JSS 52050 (1 ♂, CNC); ISRAEL: Arava Valley, Iddan Springs, in damp seep, east of spring., 30 ° 49 ’ N, 35 ° 16 ’ E, - 116 m a. s. l., 16. iv. 1995, M. E. Irwin, JSS 52107 (1 ♂, TAU); Enot Zuqim, north of gate, 30 ° 29 ’ N, 35 ° 09 ’ E, 26. iv. 2006, L. Freidman, JSS 51732 (1 ♂, TAU); Nahal Qumeran RT. 90, 31 ° 44 ’ N, 35 ° 27 ’ E, - 370 m a. s. l., 28. iii. 2007, A. Freidberg, Malaise trap, JSS 52097 (1 ♂, TAU); Zomet Mezada, 31. v. 1998, A. Freid- berg, JSS 51695; JSS 51724 (2 ♂, TAU); UNITED ARAB EMIRATES: Abu Dhabi, Al Wathba Wetland Reserve, 24 ° 15 ’ N, 54 ° 36 ’ E, i. 2015, A. Saji & A. van Harten, Malaise trap, CNC 470750 (1 ♂, EAD); ii. 2015, A. Saji & A. van Harten, Malaise trap, CNC 470517, CNC 470597, CNC 470475, CNC 470591 (2 ♂, 2 ♀, CNC); iii. 2015, A. Saji & A. van Harten, Malaise trap, CNC 470682 (1 ♂, CNC); iv. 2015, A. Saji & A. van Harten, Malaise trap, CNC 470828, CNC 470831 (1 ♂, 1 ♀, EAD); iv. 2015, A. Saji & A. van Harten, Malaise trap, CNC 470837 (1 ♂, USNM).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F7EFFB4FF3EEBD5FD914DC0.taxon	distribution	Distribution: Iran, Israel, Mongolia, United Arab Emirates (Kozánek, 1992, Skevington 2020) (Fig. 62). Note: Based on DNA barcodes, T. pruinosa is genetically closest to T. urdaensis, differing by 0.6 % (pairwise divergence) (Supplementary Table 1).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F7FFFB4FF3EE800FD914938.taxon	description	Figs 34 A – E, 63	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F7FFFB4FF3EE800FD914938.taxon	diagnosis	Diagnosis: This species can be recognized by the shape of surstyli in dorsal view, broadened at base and middle, the middle covered by dense setae dorsomedially (Fig. 34 A); epandrium wider than long in dorsal view (Fig. 34 A); phallic guide strong, straight and tapering apically in lateral view, with one strong spine dorsomedially (Fig. 34 D – E); phallus with three ejaculatory ducts, longer one divided into two branch, smaller one with two or three small spines in lateral view (Fig. 34 D – E); symmetrical gonopods, wider than hypandrium in ventral view (Fig. 34 B). Specimens examined: IRAN: Sistan & Baluchestan: Nikshahr, Pip, 26 ° 37 ’ N, 60 ° 07 ’ E, 17. vi – 19. vii. 2016, M. Ghaforimoghadam, Malaise trap, JSS 51878, JSS 51869 (1 ♂, 1 ♀, CNC); Nikshahr, Pip, 26 ° 37 ’ N, 60 ° 07 ’ E, 17. vi – 19. vii. 2016, M. Ghaforimoghadam, Malaise trap, JSS 51879 ‒ 2 (4 ♂, CNC); Rask, 26 ° 16 ’ N, 61 ° 25 ’ E, 139 m a. s. l., 10. vi – 14. vii. 2016, M. Ghaforimoghadam, Malaise trap, JSS 52163, JSS 52171 (2 ♂, CNC ISRAEL: En Mor, 30 ° 49 ’ N, 34 ° 46 ’ E, 4. vi. 2000, A. Freidberg, JSS 51692, JSS 52073 (2 ♂, TAU); UNITED ARAB EMIRATES: Wadi Bih dam, 25 ° 29 ’ N, 56 ° 02 ’ E, 19. ii – 8. iii. 2009, A. van Harten, Malaise trap, CNCD 175304 (1 ♂, USNM); Wadi Wurayah farm, 25 ° 14 ’ N, 56 ° 11 ’ E, 31. v – 14. vi. 2009, A. van Harten, Malaise trap, CNCD 137566, CNCD 137572 ‒ 3, CNCD 137577, CNCD 137585 ‒ 7, CNCD 137592 (7 ♂, 1 ♀, CNC); near ad-Dhaid, 25 ° 13 ’ N, 55 ° 35 ’ E, 16 – 19. iii. 2007, J. Btelka, pan trap, CNCD 159485 (1 ♂, EAD).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F7FFFB4FF3EE800FD914938.taxon	distribution	Distribution: Egypt, Iran, Israel, United Arab Emirates (Skevington 2020) (Fig. 63). Notes: Based on DNA barcodes, this species is genetically closest to T. jubata, differing by 5.1 % (pairwise divergence) (Supplementary Table 1).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F7FFFC9FF3EED18FD914C95.taxon	description	Figs 35 A – E, 46 C, 54, 69 A – B	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F7FFFC9FF3EED18FD914C95.taxon	diagnosis	Diagnosis: This species can be recognized by one distinct thorn-like projection and two small ones on the hind trochanter (Fig. 46 C); elongated surstyli, broadened in basal two thirds in dorsal view (Fig. 35 A), rather curved in lateral view (Fig. 35 D – E); long phallic guide; phallus with three ejaculatory ducts, one with a downward projection in lateral view (Fig. 35 D – E).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F7FFFC9FF3EED18FD914C95.taxon	description	Description: MALE: Body length (excluding antennae): 2.7 mm. Head. Face dark, light pollinose. Scape and pedicel dark; pedicel with a pair of short upper setae; flagellum light brown, tip paler than base, tapering and gray-light brown pollinose (LF: WF = 2.4); arista dark brown, with thickened base. Eyes meeting for a distance of 6 – 7 facets. Frons dark, light brown pollinose, vertex dark, lacking pollinosity, bearing an elevated ocellar triangle; vertex dark, lacking pollinosity; occiput dark, gray pollinose with scattered short dark setae. Thorax. Pleura, prescutum, scutum and scutellum brown. Pleura gray pollinose. Postpronotal lobe yellow, weakly gray pollinose and with 2 – 3 light brown along upper margin. Prescutum and scutum narrowly light brown pollinose, with some setae along anterior margin of prescutum area and some supra-alar setae. Scutellum light brown pollinose on anterior margin, with about 2 – 3 thin short setae along posterior margin (up to 0.03 mm). Subscutellum gray pollinose. Wing. Length: 2.1 mm. LW: MWW = 3.6. Wing almost entirely covered in microtrichia. M 1 gently undulating. Halter length: 0.25 mm. Base brown, knob paler than base. Legs. Coxae dark brown, gray pollinose. Mid coxa with 2 – 3 light brown setae on apical margin. Trochanters dark, gray pollinose. Hind trochanter with a long (0.08 mm) thorn-like projection on basal margin and two smaller one on apical margin (Fig. 46 C). Femora dark brown, slightly light brown at apex, gray pollinose. Hind femur with one wrinkled indentation in basal. Tibiae light brown, distinctly dark brown in middle, gray pollinose with one wrinkled indentation in middle. Tarsi light brown and paler than tibiae, gray pollinose, with some light brown setae dorsally. Pulvilli yellow. Claws light brown with black tips. Abdomen. Ground color dark brown, light brown pollinose. Tergite 1 with three to four strong lateral setae. Genitalia. Genital capsule in dorsal view: epandrium brown, slightly longer than wide (MLE: MWE = 1.1). Surstyli brown and rather symmetrical, elongated and rather broadened in basal two thirds, curved to each other at apex (Fig. 35 A). Genital capsule in ventral view: gonopods equal in height, right broader than left one (Fig. 35 B); subepandrial sclerite elongated, rather rectangular-shaped (Fig. 35 B). Genital capsule in lateral view: both surstyli weakly concave (Fig. 35 D – E); phallic guide thin and long, (Fig. 35 D – E); phallus with three ejaculatory ducts, one with a downward projection (Fig. 35 D – E). Ejaculatory apodeme cylindrical-shaped (Fig. 35 C). FEMALE: unknown.	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F7FFFC9FF3EED18FD914C95.taxon	etymology	Etymology: The specific epithet refers to Saudi Arabia, from where the specimen was collected. Specimen examined: HOLOTYPE: SAUDI ARABIA: Hada Al Asham Valle, ~ 125 km Northeast of Jeddah, 22 ° 22 ’ N, 40 ° 01 ’ E, viii-ix. 2000, ex. cultivated Alfalfa, CNCD 147709 (1 ♂, CNC).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F7FFFC9FF3EED18FD914C95.taxon	distribution	Distribution: Saudi Arabia (Fig. 54). Note: Based on DNA barcodes, this species is genetically most similar to T. jubata, differing by 4.0 % (pairwise divergence) (Supplementary Table 1).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F02FFC9FF3EE8B4FD484BDA.taxon	description	Fig. 63	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F02FFC9FF3EE8B4FD484BDA.taxon	diagnosis	Diagnosis: Following De Meyer (1995), this species can be recognized by the asymmetrical surstyli in dorsal view, left is higher than right one, left surstylus protruded in ventromedial margin; separated eyes; epandrium as long as wide; phallus with three ejaculatory ducts, one with some small spines (De Meyer, 1995: fig. 21 a – f).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F02FFC9FF3EE8B4FD484BDA.taxon	distribution	Distribution: Israel (De Meyer, 1995) (Fig. 63).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F02FFCBFF3EEE78FB164B05.taxon	description	Figs 27 H, 36 A – E, 56, 69 C – D	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F02FFCBFF3EEE78FB164B05.taxon	diagnosis	Diagnosis: This species can be recognized by its asymmetrical surstyli in dorsal view, left surstylus slightly rectangular-shaped, right surstylus slightly triangular-shaped (Fig. 36 A), left surstylus with ventromedial triangularshaped projection in lateral view (Fig. 36 D); ejaculatory ducts long and different-sized, one bearing saw-like teeth along its side (Fig. 36 D – E); long phallic guide with a distinct membranous sheath basally (Figs 36 D – E).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F02FFCBFF3EEE78FB164B05.taxon	description	Description: MALE: Body length (excluding antennae): 2.7 mm. Head. Scape dark brown, pedicel brown, flagellum light brown. Pedicel with a pair of short upper setae and a single short lower bristle; flagellum tapering and gray-white pollinose (LF: WF = 3.0); arista dark, with thickened base. Eyes converging but not meeting and separat- ed by less than diameter of frontal facets. Frons dark silver-gray pollinose. Vertex black, lacking pollinosity. Occiput dark and gray pollinose. Thorax. Postpronotal lobe light yellow, gray pollinose with 5 – 6 yellowish setae along the upper margin (up to 0.03 mm). Prescutum, scutum, scutellum black. Scutum with two uniseriate rows of intra-alar setae and two uniseriate rows of dorsocentral setae. Scutellum with 1 – 2 thin short setae along lateral margin (up to 0.03 mm). Subscutellum black, gray pollinose. Pleura black but dark brown on pteropleuron, hypopleuron and sternopleuron. Wing. Length: 2.5 mm. LW: MWW = 2.5. Wing almost entirely covered with microtrichia. Crossvein r-m reaches dm at the middle. M 1 strongly undulating in middle. Halter length: 0.3 mm. Whitish, narrowly brown at base. Legs. Coxae dark but mid coxa brown, gray pollinose. Mid coxa with two light brown anterior setae. Trochanters brown, smooth. Femora dark with light brown apices, gray pollinose. Hind femur bearing two rows of small spines in apical half with 1 – 2 wrinkled indentations at base. Tibiae brown but dark at the middle, with two ventral rows of short setae on anterior and three rows on posterior side. Hind tibia with 1 – 2 wrinkled indentations in middle. Tarsi light brown with scattered brown setae at anterior margin, basitarsi longer than other tarsomeres. Pulvilli yellow. Claws light brown with black tips. Abdomen. Ground color dark. Tergite 1 gray pollinose, with 5 – 6 dark lateral setae (up to 0.11 mm). Tergites 2 – 5 with scattered brown setae. Tergite 5 slightly longer than tergite 4 and almost symmetrical in dorsal view (LT 35: WT 5 = 1.3, WT 5: LT 5 = 2.0, T 5 R: T 5 L = 1.0). Sternites light brown laterally and dark brown centrally, gray pollinose. Syntergosternite 8 dark. Membranous area ovate in caudal view. Genitalia. Genital capsule in dorsal view: epandrium and surstyli light brown, gray pollinose. Epandrium wider than long (MLE: MWE = 0.90). Surstyli entirely asymmetrical. Left surstylus slightly rectangular-shaped and rather larger than left one, right surstylus slightly triangular-shaped and curved to left one (Fig. 36 A). Genital capsule in ventral view: gonopods equal in height, right wider than left one, left more sclerotized than right one (Fig. 36 B). Phallus with different-sized ejaculatory ducts, one bearing saw-like teeth along its side (Fig. 36 B). Phallic guide strong, straight with small projection at the base, pointed towards right surstylus (Fig. 36 B). Genital capsule in lateral view: left surstylus with distinct ventromedial projection (Fig. 36 D). Phallic guide strong, elongated, with a distinct membranous sheath basally (Fig. 36 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 36 C). FEMALE: Body length (excluding antennae): 2.75 mm. Head. Eyes separated. Frons silver-gray pollinose except ocellar triangle, widened in middle (MWF: 0.1 mm; MWF: WFA = 2.4). Occiput gray pollinose. Thorax. Postpronotal lobe light yellow, gray pollinose with 2 – 3 light brown setae along the upper margin (up to 0.03 mm). Prescutum and scutum, scutellum black, gray pollinose with scattered setae at anterior supra-alar area. Wing. Length: 2.5 mm. LW: MWW = 3.3. Legs. Mid coxa with 2 – 3 black anterior setae. Mid femur bearing two small ventral rows of dark peg-like spines in the apical third. Distitarsi with long brown setae at anterior margin. Pulvilli smaller than distitarsi. Abdomen. Tergites 1 – 6 gray pollinose with scattered brown setae. Ovipositor. Base of ovipositor dark brown, gray pollinose, dorsally with some scattered brown setae. Viewed laterally (Fig. 27 H), base of piercer curved, piercer straight, as long as the base. LP: LB = 1.0.	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F02FFCBFF3EEE78FB164B05.taxon	etymology	Etymology: The specific epithet refers to the historical name of Khuzestan, from where the specimens were collected. Specimens examined: HOLOTYPE: IRAN: Khuzestan: Shush, 32 ° 06 ’ N, 48 ° 26 ’ E, 68 m a. s. l., 11. iii – 10. v. 2015, E. Gilasian, Malaise trap, JSS 52193 (1 ♂, CNC); PARATYPE: IRAN: Khuzestan: Shush, same data as holotype, JSS 52205 (1 ♀, CNC).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F02FFCBFF3EEE78FB164B05.taxon	distribution	Distribution: Iran (Fig. 56). Note: Based on DNA barcoding, this species is genetically similar to T. spinula sp. nov., differing by 1.8 % (pairwise divergence). The male and female are genetically identical (Supplementary Table 1).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F00FFCAFF3EEFC4FBE0482E.taxon	description	Figs 27 D, 37 A – D, 47 F, 63, 69 E – F	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F00FFCAFF3EEFC4FBE0482E.taxon	diagnosis	Diagnosis: This species can be recognized by the elongated surstyli in dorsal view, constricted in basal third and curved to each other (Fig. 37 A); epandrium longer than wide (Fig. 37 A); gonopods elongated with a small projection (Fig. 37 D); phallus with three short ejaculatory ducts, one with a finger-like projection and a feather-like projection, another one with a small spine in its middle in lateral view (Fig. 37 B – C); hind trochanter with two distinct projections (Fig. 47 F).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F00FFCAFF3EEFC4FBE0482E.taxon	description	Description: MALE: Body length (excluding antennae): 2.5 mm. Head. Scape and arista dark, pedicel and flagellum light brown. Pedicel with one or two short upper pale setae; flagellum tapering and gray-white pollinose (LF: WF = 2.4). Eyes meeting for a distance of 6 – 8 facets. Frons dark, silver-gray pollinose. Vertex black, lacking pollinosity. Occiput dark and gray pollinose with scattered pale setae. Thorax. Postpronotal lobe light yellow, gray pollinose with 3 – 4 short setae along the upper margin (up to 0.02 mm). Prescutum, scutum, scutellum black. Scutum with two uniseriate rows of intra-alar setae and two uniseriate rows of dorsocentral setae. Scutellum with 3 – 4 thin short setae along lateral margin (up to 0.01 mm). Subscutellum black, gray pollinose. Pleura brown. Wing. Length: 2.5 mm. LW: MWW = 3.3. Wing almost entirely covered with microtrichia. Cross-vein r-m reaches dm at the middle. M 1 moderately undulating in middle. Halter length: 0.33 mm. Whitish, narrowly brown at base. Legs. Fore and mid coxae light brown, hind coxa brown, gray pollinose. Mid coxa with three dark anterior setae. Trochanters brown, gray pollinose. Hind trochanter with two distinct projections, longer one (0.08 mm) is thorn-like and located on basal margin with two short setae at its apex and wider one located on apical margin (Fig. 47 F). Femora brown with light brown apices, gray pollinose. All femora with one wrinkled indentations at base. Tibiae brown but light brown in basal third, with two ventral rows of short setae on anterior and three rows on posterior side. Hind tibia with 1 – 2 wrinkled indentations in middle. Tarsi light brown with scattered brown setae at anterior margin, basitarsi lighter than other tarsomeres. Pulvilli yellow. Claws light brown with black tips. Abdomen. Ground color dark brown. Tergite 1 densely gray pollinose, with 4 – 5 light brown lateral setae (up to 0.11 mm). Tergite 2 – 5 gray pollinose, with scattered brown setae. Sternites light brown laterally and dark brown centrally. Genitalia. Genital capsule in dorsal view: epandrium and surstyli brown, gray pollinose. Epandrium longer than wide (MLE: MWE = 1.14). Surstyli rather symmetrical. Both surstyli elongated, restricted in basal third, curved to each other at apex, left is higher than right one (Fig. 37 A). Genital capsule in ventral view: gonopods elongated and equal in height with a small projection (Fig. 37 D). Subepandrial sclerite elongated (Fig. 37 D). Genital capsule in lateral view: both surstyli bent towards sternite, right is broader than left one (Fig. 37 B – C). Phallic guide strong, phallus with three ejaculatory ducts, one with finger-like projection and feather-like projection, another one with a small spine in its middle (Fig. 37 B – C). FEMALE: Body length (excluding antennae): 2.7 mm. Head. Eyes separated, pedicel without short setae. Frons gray pollinose. Occiput gray pollinose with scattered dark setae. Thorax. As in male. Wing. Length: 2.2 mm. LW: MWW = 2.09. Legs. As in males except for smooth hind trochanter. Abdomen. As in males. Ovipositor. Base of ovipositor dark brown with some gray pollinosity. Viewed laterally (Fig. 27 D), piercer short and straight. LP: LB = 1.8. LDP: LPP = 3.0.	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F00FFCAFF3EEFC4FBE0482E.taxon	etymology	Etymology: The specific name is derived from the Latin word spinula, meaning thorn, referring to the thornlike projection on its hind trochanter. Specimens examined: HOLOTYPE: IRAN: Zabol, 31 ° 03 ’ N, 61 ° 22 ’ E, 470 m a. s. l., 4. vii. 2016, M. Enaiatnia, Malaise trap, JSS 51924 (1 ♂, CNC); PARATYPE: IRAN: Zabol, 31 ° 03 ’ N, 61 ° 22 ’ E, 470 m a. s. l., 7. iv. 2016, M. Enaiatnia, Malaise trap, JSS 51820 (1 ♀, CNC). Distribution: Iran (Fig. 63). Note: Based on DNA barcodes, this species is genetically similar to T. soziana sp. nov., differing by 1.8 % (pairwise divergence). The male and female differ by 0.0 % (Supplementary Table 1).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F01FFCDFF3EEC2DFC7E4E71.taxon	description	Fig. 62	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F01FFCDFF3EEC2DFC7E4E71.taxon	diagnosis	Diagnosis: Following Kazerani et al. (2017), this species can be recognized by the elongated surstyli in dorsal view, broadened in basal fourth, narrowed moderately apically; epandrium longer than wide; phallus with three ejaculatory ducts without projection (Kazerani et al. 2017: fig. 3 A – D). Based on the shape of phallus in lateral view and surstyli in dorsal view, this species resembles T. sylvatica. It differs by the shape of surstyli in lateral view, being pointed apically; the hind trochanter with three small spines; the r-m cross-vein reaching cell dm before its middle.	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F01FFCDFF3EEC2DFC7E4E71.taxon	distribution	Distribution: Iran (Kazerani et al. 2017; Skevington 2020) (Fig. 62).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F06FFCDFF3EEAD0FC75498C.taxon	description	Figs 38 A – E, 54, 70 A – B	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F06FFCDFF3EEAD0FC75498C.taxon	diagnosis	Diagnosis: This species can be recognized by the shape of surstyli in dorsal view, elongated and constricted in middle (Fig. 38 A); wide epandrium in dorsal view (Fig. 38 A); asymmetrical gonopod and hypandrium in ventral view (Fig. 38 B); ejaculatory apodeme funnel-shaped (Fig. 38 C). Based on the shape of surstyli, this species resembles T. transvaalensis De Meyer, 1993 (Fig. 42 A – E). The shape of surstyli is slightly different in lateral view, in T. transvaalensis, straight in medial, narrowed in apical third (Fig. 42 D – E) and T. subtransvaalensis has a hypandrial apodeme in ventral view (Fig. 38 B).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F06FFCDFF3EEAD0FC75498C.taxon	description	Description: MALE: Body length (excluding antennae): 4.6 mm. Head. Scape dark, pedicel dark brown, flagellum brown; pedicel with three short upper setae; flagellum tapering (LF: WF = 3.0); arista dark. Eyes meeting for a distance of 8 – 9 facets. Frons silver-gray pollinose. Vertex black, lacking pollinosity. Occiput dark and gray pollinose in lower half. Thorax. Postpronotal lobe light yellow, gray pollinose with 4 – 6 small setae along the upper margin (up to 0.03 mm). Prescutum and scutum black, with a row of supra-alar setae. Scutellum black, silver-gray pollinose, with about 12 – 14 short dark setae along posterior margin (up to 0.03 mm). Subscutellum and pleura dark. Wing. Length: 4.1 mm. LW: MWW = 4.0. Wing almost entirely covered with microtrichia. Cross-vein r-m reaching cell dm at the middle. M 1 strongly undulating in middle. Halter length: 0.37 mm. Yellowish, dark brown at base. Legs. Dark but narrowly yellow at apex of femora, base of tibiae. Trochanters black, smooth. Femora partly gray pollinose. All femora bearing two rows of dark, peg-like anteroventral spines in apical half. Hind femur with one wrinkled indentation in basal quarter. Tibiae with two rows of short setae on anterior and three rows on posterior side. Hind tibia with one wrinkled indentation in the middle. Tarsi brown, gray pollinose. Pulvilli shorter than distitarsi. Claws brown with black tips. Abdomen. Ground color dark. Tergite 1 entirely gray pollinose with 8 – 9 short black lateral setae (up to 0.1 mm). Tergites 2 – 5 with scattered short dark setae. Sternites brown laterally and dark centrally, gray pollinose. Genitalia. Genital capsule in dorsal view: epandrium and surstyli dark brown, gray pollinose. Epandrium wider than long (MLE: MWE = 0.3), left side more extended than right one. Surstyli rather symmetrical, elongated, constricted in middle (Fig. 38 A). Genital capsule in ventral view: gonopods rather large and unequal in height, left slightly higher than right one; hypandrium unequal in height, left is higher than right one (Fig. 38 B). Genital capsule in lateral view: surstyli curved towards sternite (Fig. 38 D – E). Phallic guide short, strong, pointed apically (Fig. 38 D – E); phallus with two long and one short ejaculatory ducts (Fig. 38 D – E). Ejaculatory apodeme funnel-shaped (Fig. 38 C). FEMALE: unknown.	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F06FFCDFF3EEAD0FC75498C.taxon	etymology	Etymology: The specific name is derived from the Latin preposition ‘ sub’ meaning near, referring to this species close relation to T. transvaalensis De Meyer, 1993. Specimens examined: HOLOTYPE: ISRAEL: Rehovot, 31 ° 53 ’ N, 34 ° 48 ’ E, 2. i. 1999, Y. Nussbaum, JSS 51765 (1 ♂, TAU). PARATYPE: ISRAEL: Sappir, 30 ° 36 ’ N, 35 ° 11 ’ E, 1. v. 1997, A. Freidberg, JSS 51690 (1 ♂, CNC).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F06FFCDFF3EEAD0FC75498C.taxon	distribution	Distribution: Israel (Fig. 54). Notes: Based on DNA barcoding, this species is genetically most similar to T. inopinata, differing by 5.3 % (pairwise divergence). Note that T. transvaalensis has not been sequenced.	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F06FFCEFF3EEC4CFACD4D0D.taxon	description	Figs 39 A – E, 46 D, 64	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F06FFCEFF3EEC4CFACD4D0D.taxon	diagnosis	Diagnosis: This species can be recognized by the shape of surstyli in dorsal view, right surstylus quadratic-shaped with small inner projection, left surstylus elongated, rather rectangular-shaped, longer than right one (Fig. 39 A); epandrium wider than long, slightly ovate-shaped (Fig. 39 A); phallus trifid with wavy ejaculatory ducts (Fig. 39 D – E); asymmetrical gonopods in ventral view, right higher than left one (Fig. 39 B); hypandrium ovate-shaped with distinct hypandrial apodeme in ventral view (Fig. 39 B). Specimens examined: IRAN: Sistan & Baluchestan: Saravan, 27 ° 25 ’ N, 62 ° 17 ’ E, 12. v. 2013, F. Hamzavi, pan trap, JSS 51916 (1 ♂, CNC); ISRAEL: Ne’ot haKikkar, 30 ° 56 ’ N, 35 ° 22 ’ E, 1. v. 1997, A. Freidberg, JSS 51694 (1 ♂, TAU); QATAR: Al Shamal distnict, Al Zubarah exclusion zone, Grassy meadow, 25 ° 58 ’ N, 51 ° 02 ’ E, 20. iii. 2012, K. P. Puliafico & A. M. Jensen, yellow & white pan traps, JSS 52318 – 9 (2 ♂, CNC); UNITED ARAB EMIRATES: Bithnah, 25 ° 06 ’ N, 56 ° 08 ’ E, 4 – 26. iii. 2006, A. van Harten, light trap, JSS 52240 (1 ♂, CNC); NARC, near Sweihan, 24 ° 14 ’ N, 55 ° 16 ’ E, 9 – 20. ix. 2005, A. van Harten, light trap, CNCD 174944 – 7 (4 ♂, CNC); Wadi Maidaq, 25 ° 11 ’ N, 56 ° 04 ’ E, 7 – 14. iii. 2006, A. van Harten, pan trap, CNCD 8951 (1 ♂, USNM); Wadi Wurayah farm, 25 ° 14 ’ N, 56 ° 11 ’ E, 17 – 24. iii. 2009, A. van Harten, Malaise trap; CNCD 137124 – 5, CNCD 137128 (3 ♂, CNC); 19. iv – 19. v. 2009, A. van Harten, Malaise trap, CNCD 137532, CNCD 137534, CNCD 137536, CNCD 137538, CNCD 137540, CNCD 137545, CNCD 137552 (7 ♂, CNC); 24 – 30. iii. 2009, A. van Harten, Malaise trap, CNCD 136930, CNCD 136932 (2 ♂, CNC); 31. v – 14. vi. 2009, A. van Harten, Malaise trap, CNCD 137578 (1 ♂, EAD); Wadi Wurayah, 25 ° 14 ’ N, 56 ° 11 ’ E, 19 – 26. iv. 2009, A. van Harten, Malaise trap, CNCD 160583 (1 ♂, CNC); al-Rafa, 25 ° 26 ’ N, 55 ° 31 ’ E, 20 – 23. iv. 2009, A. van Harten, pan trap, CNCD 189979 (1 ♂, CNC); near ad-Dhaid, 25 ° 13 ’ N, 55 ° 35 ’ E, 16 – 19. iii. 2007, J. Btelka, pan trap, CNCD 159476, CNCD 159478 – 9, CNCD 159483 – 4, CNCD 159486, CNCD 159488 (7 ♂, CNC).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F06FFCEFF3EEC4CFACD4D0D.taxon	distribution	Distribution: Bahamas, Belize, Bermuda, Brazil, Canada, Chile, Colombia, Costa Rica, Cuba, Dominica, Egypt, Gabon, Iran, Israel, Jamaica, Mexico, Mozambique, Namibia, Nicaragua, Panama, Peru, Puerto Rico, Qatar, Thailand, the United States, United Arab Emirates (Skevington 2020) (Fig. 64). Note: DNA barcodes of T. subvirescens overlap with T. pilosiventris. See the notes under T. pilosiventris.	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F05FFCEFF3EE9CCFD024841.taxon	description	Figs 40 A – D, 61, 70 C – D	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F05FFCEFF3EE9CCFD024841.taxon	diagnosis	Diagnosis: This species can be recognized by the shape of surstyli in dorsal view, broadened at base and rather triangular-shaped (Fig. 40 A); broadened gonopods, right wider than left one with small projection in ventral view (Fig. 40 B); short ejaculatory ducts, one bearing some saw-like teeth in basal half in lateral view (Fig. 40 C ‒ D).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F05FFCEFF3EE9CCFD024841.taxon	description	Description: MALE: Body length (excluding antennae): 3.1 mm. Head. Scape, pedicel and arista dark. Pedicel with 2 – 3 upper setae and a single lower bristle; flagellum light brown, gray pollinose and long tapering (LF: WF = 3.0). Eyes converging but not meeting and separated by less than diameter of frontal facets. Frons silver-gray pollinose, vertex dark and shine, lacking pollinosity, bearing an elevated ocellar triangle. Occiput dark, gray pollinose with scattered brown setae. Thorax. Postpronotal lobe light brown. Prescutum and scutum black, silver-gray and brown pollinose, with scattered setae at anterior supra-alar area and uniseriate rows of intra-alar setae. Scutellum black, silver-gray pollinose, with about 14 brown setae along posterior margin (up to 0.08 mm). Subscutellum and pleura black; pleuron light brown. Wing. Wing length: 3.1 mm., LW: MWW = 3.0. Wing almost entirely covered with microtrichia. M 1 strongly undulating in middle. Halter length: 0.37 mm. light brown, narrowly black at base. Legs. Coxae black, trochanters light brown, smooth. Femora dark brown with narrow but distinct pale base and apices, silver pollinose. Femora without ventral spines. Mid and hind femora with one wrinkled indentation in basal. Tibiae brown at middle, gray pollinose, with two rows of short brown setae on anterior and three rows on posterior side. Mid and hind tibiae with one or two wrinkled indentations in middle. Tarsi light brown. Abdomen. Ground color dark brown, tergite 1 silver gray pollinose with a patch of light brown setae (15 ‒ 17) laterally extending along posterior margin, tergites 2 ‒ 5 with relatively long brown setae. Sternites white-yellow laterally and brown centrally, gray pollinose. Genitalia. Genital capsule in dorsal view: epandrium and surstyli brown, gray pollinose. Epandrium wider than long (MLE: MWE = 0.74). Surstyli symmetrical, broadened at base, rather triangular-shaped (Fig. 40 A). Genital capsule in ventral view: gonopods broadened and slightly equal in height, right wider than left one with a small projection, subepandrial sclerite wide basically, phallic guide strong, slightly straight (Fig. 40 B). Genital capsule in lateral view: surstyli longer than wide, slightly straight, broadened at apex (Figs 40 C ‒ D). Phallic guide strong, pointed apically (Fig. 40 C); phallus with three ejaculatory ducts, one bearing some saw-like teeth in basal half (Fig. 40 C ‒ D). FEMALE: unknown.	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F05FFCEFF3EE9CCFD024841.taxon	etymology	Etymology: The specific epithet refers to “ Susa ”, the historical name of Shush, from where the specimen was collected. The name is a noun in apposition. Specimen examined: HOLOTYPE: IRAN: Khuzestan: Shush, 32 ° 06 ’ N, 48 ° 26 ’ E, 68 m a. s. l., 11. iii – 10. v. 2015, E. Gilasian, Malaise trap, JSS 52217 (1 ♂, CNC).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F05FFCEFF3EE9CCFD024841.taxon	distribution	Distribution: Iran (Fig. 61). Notes: Tomosvaryella susa sp. nov. is genetically very close to T. ampliasa sp. nov., differing by 0.8 ‒ 1.2 % (pairwise divergence) (Supplementary Table 1).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F0AFFC3FF3EEC33FD024E54.taxon	description	Figs 41 A ‒ E, 48 C, 64	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F0AFFC3FF3EEC33FD024E54.taxon	diagnosis	Diagnosis: This species can be recognized by the distinct frontal prominence (Fig. 48 C); shape of surstyli in dorsal view, both broadened in basal half, moderately narrowed to apical half, curved to each other at apex; left surstylus slightly longer than right one (Fig. 41 A), gonopods slightly equal in height, right with rounded projection towards phallic guide (Fig. 41 B). Specimens examined: IRAN: Kermanshah: Dodan, 35 ° 00 ’ N, 46 ° 12 ’ E, 1011 m a. s. l., Garden, 5. vii. 2016, M. Zardouei, funnel Malaise trap, JSS 51938 (1 ♂, CNC); Sarpolezahab, 34 ° 28 ’ N, 45 ° 49 ’ E, 456 m a. s. l., 14. v. 2015, M. Zardouei, Malaise trap, JSS 51936 ‒ 7 (2 ♂, CNC). Distribution: Iran, Kazakhstan (Kuznetzov 1994; Skevington 2020) (Fig. 64). Note: Based on DNA barcoding, this species is genetically similar to T. ellipiensis sp. nov., differing by 5.7 % (pairwise divergence) (Supplementary Table 1).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F09FFC2FF3EEBD5FEA74DB5.taxon	description	Figs 42 A ‒ E, 64	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F09FFC2FF3EEBD5FEA74DB5.taxon	diagnosis	Diagnosis: This species can be recognized by the shape of surstyli in dorsal view, constricted in middle, knobshaped apically (Fig. 42 A); epandrium wider than long in dorsal view (Fig. 42 A); phallus with three ejaculatory ducts, one longer, coiled, bearing small setae in lateral view (Fig. 42 D ‒ E); gonopods equal in height (Fig. 42 B), hypandrium ovate-shaped with distinct hypandrial apodeme in ventral view (Fig. 42 B). Specimens examined: ISRAEL: Nahal Lavan, 30 ° 51 ’ N, 34 ° 36 ’ E, 30. iv. 1996, I. Yarom, JSS 51786 (1 ♂, TAU); Sede Halamish, 31 ° 00 ’ N, 35 ° 07 ’ E, 30. iv. 1996, A. Freidberg, JSS 52093 (1 ♂, CNC).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F09FFC2FF3EEBD5FEA74DB5.taxon	distribution	Distribution: Israel, South Africa (Fig. 64). Note: This sort of disjunct distribution raises some questions. Could this be a crypic pair of species? Is the gap caused by collecting bias? These questions can only be answered by more field work to try to fill gaps, and molecular work to test existing species concepts. We attempted to sequence these specimens but failed. More work is needed.	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F09FFC2FF3EE98EFC1E4C8F.taxon	description	Fig. 62	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F09FFC2FF3EE98EFC1E4C8F.taxon	diagnosis	Diagnosis: Following De Meyer (1995), this species can be recognized by the rectangular-shaped surstyli in dorsal view, slightly broadened at base; epandrium as long as wide; straight ejaculatory ducts; hind tibia with long setae apically (De Meyer, 1995: fig. 22 a – f).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F09FFC2FF3EE98EFC1E4C8F.taxon	distribution	Distribution: Israel, Morocco, Spain (Skevington 2020) (Fig. 62).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F09FFC4FF3EEF52FD914FC8.taxon	description	Figs 43 A – E, 48 B, 63, 70 E – F	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F09FFC4FF3EEF52FD914FC8.taxon	diagnosis	Diagnosis: This species can be recognized by a distinct horn-like prominence in the middle of frons (Fig. 48 B); shape of surstyli more elongated in dorsal view (Fig. 43 A), rather curved in lateral view (Fig. 43 D – E).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F09FFC4FF3EEF52FD914FC8.taxon	description	Description: MALE: Body length (excluding antennae): 3.5 mm. Head. Scape, pedicel and arista dark brown, flagellum light brown. Pedicel with a pair of short upper pale setae; flagellum tapering and gray-white pollinose (LF: WF = 2.2). Eyes converging but not meeting and separated by less than diameter of frontal facets. Frons dark silver-gray pollinose, with a distinct prominence in middle (Fig. 48 B). Vertex black, lacking pollinosity. Occiput dark and gray pollinose. Thorax. Postpronotal lobe light yellow, gray pollinose with 6 – 7 yellowish setae along the upper margin (up to 0.04 mm). Prescutum, scutum, scutellum black. Scutum with two uniseriate rows of intra-alar setae and two uniseriate rows of dorsocentral setae. Scutellum with 1 – 2 thin short setae along lateral margin (up to 0.01 mm). Subscutellum black, gray pollinose. Pleura brown. Wing. Length: 3.2 mm. LW: MWW = 5.08. Wing almost entirely covered with microtrichia. Cross-vein r-m reaches dm at the middle. M 1 moderately undulating in middle. Halter length: 0.37 mm. Whitish, narrowly brown at base. Legs. Coxae dark brown, gray pollinose. Mid coxa with two dark anterior setae. Trochanters dark brown, smooth. Femora brown with light brown apices, gray pollinose. All femora with 1 – 2 wrinkled indentations at base. Tibiae brown but light brown in basal half, with two ventral rows of short setae on anterior and three rows on posterior side. Hind tibia with 1 – 2 wrinkled indentations in middle. Tarsi light brown with scattered brown setae at anterior margin, basitarsi lighter than other tarsomeres. Pulvilli yellow. Claws light brown with black tips. Abdomen. Ground color dark brown. Tergite 1 densely gray pollinose, with 4 – 5 brown lateral setae (up to 0.11 mm). Tergites 2 densely gray pollinose in anterior margin, Tergite 3 – 5 gray pollinose, with scattered brown setae. Sternites light brown laterally and dark brown centrally, gray pollinose. Genitalia. Genital capsule in dorsal view: epandrium and surstyli light brown, gray pollinose. Epandrium wider than long (MLE: MWE = 0.60). Surstyli symmetrical. Both surstyli more elongated, longer than epandrium (Fig. 43 A). Left surstylus gently bent in basal third (Fig. 43 A). Genital capsule in ventral view: gonopods equal in height (Fig. 43 B). Subepandrial sclerite rectangular-shaped (Fig. 43 B). Genital capsule in lateral view: both surstyli long, weakly concave (Fig. 43 D – E). Phallic guide strong, phallus with three small ejaculatory ducts (Fig. 43 D – E). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 43 C). FEMALE: unknown.	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F09FFC4FF3EEF52FD914FC8.taxon	etymology	Etymology: The specific name is derived from the mythical unicorn, in reference to the horn-like prominence on the frons. Specimen examined: HOLOTYPE: IRAN: Kermanshah: Dagasyave, 35 ° 01 ’ N, 46 ° 07 ’ E, 810 m a. s. l., 7. vi. 2016, M. Zardouei, Malaise trap, JSS 51821 (1 ♂, CNC).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F09FFC4FF3EEF52FD914FC8.taxon	distribution	Distribution: Iran (Fig. 63). Note: Based on DNA barcodes, this species is genetically similar to T. debruyni, differing by 7.4 % (pairwise divergence) (Supplementary Table 1).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F0FFFC4FF3EEA08FCDF4DB5.taxon	description	Fig. 60	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F0FFFC4FF3EEA08FCDF4DB5.taxon	diagnosis	Diagnosis: Following Kuznetzov (1994), this species can be recognized by the shape of surstyli in dorsal view, broadened and angled at base, pointed at apices (Majnon-Jahromi et al. 2017 b: Fig. 19); both surstyli rounded apically, with a distinct posteroventral projection before its apex in lateral view (Kuznetzov, 1994: Figs 302 – 303). Tomosvaryella urdaensis appears to be closely related to T. gazliensis (Fig. 15 A – D). See diagnosis of T. gazliensis for more details.	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F0FFFC4FF3EEA08FCDF4DB5.taxon	distribution	Distribution: Iran, Kazakhstan (Skevington 2020) (Fig. 60).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F0FFFC4FF3EE994FD914A3C.taxon	description	Figs 44 A – F, 64	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F0FFFC4FF3EE994FD914A3C.taxon	diagnosis	Diagnosis: This species can be recognized by the shape of surstyli in lateral view, with extended dorsomedial lobe, lobe of right surstylus larger than left one (Fig. 44 E – F), both surstyli slightly flatted apically in dorsal view (Fig. 44 A – B); phallic guide straight, divided apically with plumose projection shortly before apex in lateral view (Fig. 44 E – F); subepandrial sclerite quadratic-shaped in ventral view (Fig. 44 C); gonopods equal in height, right wider than left one in ventral view (Fig. 44 C). Specimens examined: IRAN: Sistan & Baluchestan: Zabol, 31 ° 02 ’ N, 61 ° 32 ’ E, 470 m a. s. l., 24. v. 2016, M. Enaiatnia, funnel Malaise trap, JSS 51902 (1 ♂, CNC); 31 ° 03 ’ N, 61 ° 22 ’ E, 6. x. 2016, M. Enaiatnia, Malaise trap, JSS 52175 (1 ♂, CNC); ISRAEL: Akko, 32 ° 55 ’ N, 35 ° 05 ’ E, 12. viii. 2002, A. Freidberg, JSS 52067 (1 ♂, TAU); Neot haKikkar, 30 ° 57 ’ N, 35 ° 22 ’ E, 9. iv. 2002, L. Friedman, JSS 51767, JSS 51773 – 5 (4 ♂, TAU); Sappir, 30 ° 36 ’ N, 35 ° 11 ’ E, 1. v. 1997, A. Freidberg, JSS 51720 (1 ♂, TAU).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F0FFFC4FF3EE994FD914A3C.taxon	distribution	Distribution: Congo, Egypt, Iran, Israel, Madagascar (Skevington 2020) (Fig. 64). Note: Based on DNA barcoding, this species is genetically similar to T. hamata, differing by 5.3 % (pairwise divergence) (Supplementary Table 1).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F0FFFDDFF3EEE11FA2048FB.taxon	description	Figs 45 A – E, 47 D, 55, 71 A – B	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F0FFFDDFF3EEE11FA2048FB.taxon	diagnosis	Diagnosis: This species can be recognized by the shape of surstyli in dorsal view, symmetrical, broadened at base and rather rectangular-shaped (Fig. 45 A); epandrium longer than wide (Fig. 45 A); long phallic guide with distinct projection before apex, pointing upwards in lateral view (Fig. 45 D – E); hind tibia with three to four long dark posterodorsal setae at apices (Fig. 47 D).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F0FFFDDFF3EEE11FA2048FB.taxon	description	Description: MALE: Body length (excluding antennae): 3.05 mm. Head. Scape, pedicel and arista dark. Pedicel with three short upper setae and a single lower bristle; flagellum brown, gray pollinose and tapering (LF: WF = 2.4). Eyes meeting for a distance of 12 – 13 facets. Frons silver-gray pollinose, vertex dark and shine, lacking pollinosity, bearing an elevated ocellar triangle. Occiput dark, gray pollinose with scattered dark seate. Thorax. Postpronotal lobe light brown, with 2 – 3 short light brown setae along the upper margin (up to 0.01 mm). Prescutum and scutum black, silver-gray and brown pollinose, with scattered setae at anterior supra-alar area and uniseriate rows of intra-alar setae. Scutellum black, silver-gray pollinose, with about 10 short dark setae along posterior margin (up to 0.04 mm). Subscutellum black and pleura brown. Wing. Wing length: 3.1 mm., LW: MWW = 3.4. Wing almost entirely covered with microtrichia. Halter length: 0.3 mm., light brown, narrowly black at base. Legs. Dark brown, trochanters smooth, femora and tibiae with narrow but distinct pale base and apices, silver pollinose. Fore and mid femora with two rows of dark, peg-like anteroventral spines in apical half. Mid and hind femora with one wrinkled indentation in basal. Hind femur with two wrinkled indentation in basal. Tibiae with two rows of short brown setae on anterior and three rows on posterior side. Hind tibia with one or two wrinkled indentations in middle and with three to four long dark posterodorsal setae (up to 0.16 mm) at apices (Fig. 47 D). Tarsi light brown, distitarsi brown. Abdomen. Ground color dark, tergite 1 silver gray pollinose with a patch of dark setae (5 ‒ 6) laterally, tergites 4 ‒ 5 gray pollinose dorsolaterally, slightly extending dorsally along anterior margin. Sternites light brown laterally and brown centrally, gray pollinose. Genitalia. Genital capsule in dorsal view: epandrium and surstyli brown, gray pollinose. Epandrium longer than wide (MLE: MWE = 1.16). Surstyli symmetrical, slightly broadened at base, rectangular-shaped (Fig. 45 A). Genital capsule in ventral view: gonopods long and equal in height, phallic guide strong, straight, with a finger-like projection in right side before its apex (Fig. 45 B). Genital capsule in lateral view: surstyli slightly straight in basal two thirds, curved towards sternite in apical third (Fig. 45 D – E). Phallic guide long, with a distinct projection pointed upwards towards surstyli (Fig. 45 D – E); phallus with three narrow ejaculatory ducts, twisted apically (Fig. 45 D). FEMALE: unknown.	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F0FFFDDFF3EEE11FA2048FB.taxon	etymology	Etymology: The specific epithet refers to Yemen, from where the type series was collected. Specimens examined: HOLOTYPE: YEMEN: 12 km NW of Manakhah, 15 ° 04 ’ N, 43 ° 44 ’ E, 21. viii – 28. x. 2002, A. van Harten, Malaise trap, CNCD 9082 (1 ♂, CNC); PARATYPES: YEMEN: Same data as holotype, CNCD 9081 (1 ♂, CNC); 12 km NW of Manakhah, 15 ° 04 ’ N, 43 ° 44 ’ E, 6. vii – 21. viii. 2002, A. van Harten, Malaise trap, CNCD 9079 (1 ♂, CNC).	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
03A887D05F0FFFDDFF3EEE11FA2048FB.taxon	distribution	Distribution: Yemen (Fig. 55). Note: Based on DNA barcodes, this species is genetically most similar to T. pilosiventris, differing by 10.3 % (pairwise divergence) (Supplementary Table 1). FIGURE 47. Hind leg of Tomosvaryella species in lateral view A) T. jubata (JSS 52231, CNC), B) T. hamounensis sp. nov., paratype (JSS 51816, CNC), C) T. nodosa (JSS 51746, TAU), D) T. yemenensis sp. nov., paratype (CNCD 9079, CNC), E) T. pilosiventris (JSS 52252, CNC), F) T. spinula sp. nov., holotype (JSS 51924, CNC). ...... continued on the next page ...... continued on the next page ...... continued on the next page	en	Motamedinia, Behnam, Skevington, Jeffrey H., Kelso, Scott (2021): Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species. Zootaxa 5002 (1): 1-103, DOI: 10.11646/zootaxa.5002.1.1
