taxonID	type	description	language	source
03A89857A954FF896A9192BEFBAEFC14.taxon	description	(Figs 1, 4 – 15)	en	Shuai, Qi, Nozaki, Tsubasa, Tang, Liang (2021): Notes on the genus Diochus Erichson, with descriptions of two new species from China (Coleoptera, Staphylinidae). Zootaxa 4908 (2): 276-282, DOI: 10.11646/zootaxa.4908.2.8
03A89857A954FF896A9192BEFBAEFC14.taxon	materials_examined	Material examined. Japan: Kumamoto Pref.: 1 ♁ 1 ♀, Hitoyoshi-shi, 15. X. 1977, Ôishi leg. (KUM); China: Zhejiang: 1 ♀, Lin’an City, West Tianmushan, 15 – 16. VII. 2000, Li-Zhen Li leg. (SHNU); 3 ♁♁ 2 ♀, Longwangshan, 300 – 500 m, 24 – 26. IV. 2004, Hu, Li, Tang & Zhu leg. (SHNU); 2 ♁♁ 1 ♀, Baishanzu Conv., 1050 – 1700 m, 5 – 7. V. 2004, Hu, Tang & Zhu leg. (SHNU); 2 ♀♀, Gutianshan, 5 – 7. V. 2005, Li & Zhu leg. (SHNU); 2 ♁♁ 1 ♀, Qingliangfeng, 1050 – 1080 m, 9 – 10. V. 2005, Li & Zhu leg. (SHNU); 1 ♁, Longwangshan, Anji County, 350 – 550 m, 24. IV. 2006, Shan-Jia Shen leg. (SHNU); 3 ♁♁ 1 ♀, Xianju County, Danzhu, 150 – 600 m, 31. V – 2. VI. 2006, Li & Shen leg. (SHNU); 1 ♀, Lin’an City, West Tianmushan, 300 – 400 m, 11 – 15. VI. 2006, Hu & Tang leg. (SHNU); 1 ♁ 3 ♀♀, Suichang County, Jiulongshan N. R., 500 – 800 m, 29 – 30. VII. 2006, Li & Shen leg. (SHNU); 3 ♁♁, Lin’an City, East Tianmushan, 650 m, 25. IV. 2008, He & Tang leg. (SHNU); 1 ♀, Lin’an City, Longwangshan, 600 – 1000 m, 19. VII. 2009, Feng, Yin & Yuan leg. (SHNU); 3 ♁♁ 4 ♀♀, Lin’an City, West Tianmushan, 300 – 400 m, 29. V. 2010, Wang, Xu & Zhu leg. (SHNU); 1 ♀, Lin’an City, Tianmushan, Haoling, 830 – 900 m, 31. V. 2010, Wang, Xu & Zhu leg. (SHNU); 1 ♁, Lin’an City, East Tianmushan, 1050 – 1150 m, 13. IV. 2011, Peng & Zhu leg. (SHNU); 1 ♀, Lin’an City, West Tianmushan, 300 – 1000 m, 27. VII. 2011, Yu-Hong Pan leg. (SHNU); 1 ♀, Anji City, Qianmutian, 700 – 1325 m, 28. VII. 2011, Yu-Hong Pan leg. (SHNU); 8 ♁♁ 13 ♀♀, Ningbo City, Yinjiang Town, Xishan, 200 m, 18 – 22. VIII. 2011, Jian-Qing Zhu leg. (SHNU); 1 ♁ 3 ♀♀, Kaihua County, Gutianshan, 29 ° 14 ’ N, 118 ° 08 ’ E, 400 – 500 m, 19. VI. 2013, Li, Lv, Xie & Zheng leg. (SHNU); 6 ♁♁, Longquan, Fengyangshan, Datianping, 27 ° 54 ’ 36 ’’ N, 119 ° 10 ’ 20 ’’ E, 1320 m, bush leaf, moss, ferns, sifted, beating, 27. IV. 2014, Peng, Song, Yan, Yin & Yu leg. (SHNU); 4 ♁♁ 3 ♀♀, Qingyuan, Baishanzu N. R., path nr. hotel, 27 ° 44 ’ 42 ’’ N, 119 ° 10 ’ 19 ’’ E, 1060 – 1130 m, leaf litter, sifted, 30. IV. 2014, Peng, Song, Yan, Yin & Yu leg. (SHNU); 5 ♁♁ 1 ♀, Lin’an City, West Tianmushan, Mountain Grand Canyon, 30 ° 22 ’ 54.68 ’’ N, 119 ° 27 ’ 49.71 ’’ E, 749 m, sifted, 6. V. 2018, Cheng & Shen leg. (SHNU); 2 ♁♁ 3 ♀♀, Quzhou City, Kaihua County, Gutianshan N. R., 28 ° 19 ’ 39.60 ’’ N, 118 ° 40 ’ 36.93 ’’ E, 461 m, mixed leaf litter, sifted, 14. VIII. 2018, Cheng & Miao leg. (SHNU); Fujian: 5 ♀♀, Wuyishan, 27 – 31. V. 2002, Li-Zhen Li leg. (SHNU); 1 ♁ 2 ♀♀, Wuyishan City, Guadun Vill., 27 ° 44 ’ N, 117 ° 38 ’ E, 1100 – 1300 m, 30. V. 2012, Dai & Peng leg. (SHNU); Anhui: 1 ♀, Guniujiang, 300 m, 27. IV. 2005, Hu & Tang leg. (SHNU); 2 ♁♁, Huangshan, Tangkou Town, Hougu, 30 ° 5 ’ 3.48 ’’ N, 118 ° 8 ’ 45.96 ’’ E, 569 – 688 m, 29. VI – 7. VII. 2020, sifted, Tang, Li & Zhou leg. (SHNU).	en	Shuai, Qi, Nozaki, Tsubasa, Tang, Liang (2021): Notes on the genus Diochus Erichson, with descriptions of two new species from China (Coleoptera, Staphylinidae). Zootaxa 4908 (2): 276-282, DOI: 10.11646/zootaxa.4908.2.8
03A89857A954FF896A9192BEFBAEFC14.taxon	description	Measurements. BL: 4.3 – 5.9 mm, FL: 2.4 – 2.8 mm. HL: 0.70 – 0.80 mm, HW: 0.54 – 0.62 mm, EyL: 0.18 – 0.23 mm, TL: 0.40 – 0.46 mm, PL: 0.91 – 1.05 mm, PW: 0.72 – 0.85 mm, EL: 0.91 – 1.02 mm, EW: 0.92 – 1.09 mm, SL: 0.63 – 0.72 mm. Head 1.22 – 1.41 times as long as wide, eye 0.40 – 0.55 times as long as tempora, pronotum 1.20 – 1.32 times as long as wide, elytra 0.90 – 1.07 times as long as wide.	en	Shuai, Qi, Nozaki, Tsubasa, Tang, Liang (2021): Notes on the genus Diochus Erichson, with descriptions of two new species from China (Coleoptera, Staphylinidae). Zootaxa 4908 (2): 276-282, DOI: 10.11646/zootaxa.4908.2.8
03A89857A954FF896A9192BEFBAEFC14.taxon	distribution	Distribution. China (Fujian, Hainan?, Zhejiang, Guangxi, Yunnan, Anhui, Jilin?, Liaoning?), Japan. New to Anhui.	en	Shuai, Qi, Nozaki, Tsubasa, Tang, Liang (2021): Notes on the genus Diochus Erichson, with descriptions of two new species from China (Coleoptera, Staphylinidae). Zootaxa 4908 (2): 276-282, DOI: 10.11646/zootaxa.4908.2.8
03A89857A954FF896A9192BEFBAEFC14.taxon	discussion	Remarks. Diochus bicornutus was described by Zhou & Zhou (2016) in a review of the Diochus species of China. The species is the most widely distributed Diochus species in China and was reported from Hainan, Fujian, Zhejiang, Guangxi and Yunnan. Diochus bicornutus can be recognized by the distinct secondary sexual characters of the male: head with an elongate elevation between eyes and sternite VIII with coarse setae along the posterior margin. However, these two characters are also shared with D. japonicus and D. conicollis. Before discussing the relationships of these three species mentioned above, a few characters of D. bicornutus mentioned in the original description must be fixed here: 1. “ Right mandible with a smaller and not very sharp tooth (Fig. 4 J in Zhou & Zhou 2016) ”. This character, one small inner tooth of right mandible, is unusual within Diochus. It seems that unfortunately, the right mandible illustrated in Zhou & Zhou, 2016 is damaged. We checked many specimens of D. bicornutus in our collections and in all specimens the right mandible (Fig. 7) was bearing two inner teeth: a small one and a large one. 2. The body length, stated as “ BL = (5.1 – 5.2) mm ”, seems to be measured on a limited number of specimens. According to the measurements of specimens listed in this paper, the range of the body length should be expanded to 4.3 – 5.9 mm. 3. The shape of the head in the original description, stated as “ HL to HW ratio 1.41 ”, is incorrectly measured or calculated. The measurements of head in the original description were given as “ HL = (0.6 – 0.7) mm, HW = (0.5 – 0.6) mm ”, and therefore it is impossible arrive at such a ratio. The ratio be 1.22 – 1.41. 4. The configuration of the gular sutures was given as “ Gular sutures deep, confluent at basal 1 / 4, then extended to base ” in the original description of D. bicornutus, versus “ Gular sutures deep, confluent at near base ” in the redescription of D. conicollis. However, the gular sutures are not always stable in D. bicornutus. Forty-eight male specimens of D. bicornutus listed here were checked: 42 males with gular sutures confluent at basal 1 / 4, 6 males with gular sutures confluent at base or at a point closer to base than basal 1 / 4. Therefore, the external differences among D. bicornutus, D. japonicus and D. conicollis mentioned in Zhou & Zhou’s paper no longer exist. Additionally, the female holotype of D. japonicus is the only specimen examined by Zhou & Zhou (2016). By comparing the aedeagi and spermathecae of D. japonicus and D. bicornutus in the present study, no striking differences were found and so Diochus bicornutus Zhou & Zhou, 2016 syn. nov. is here synonymized with D. japonicus Cameron, 1930. The status of D. japonicus Cameron, 1930 is also doubtful, since D. japonicus shares nearly all characters with D. conicollis (Motschulsky, 1858). A male lectotype of D. conicollis was studied by Zhou & Zhou (2016) but the internal structures of the aedeagus were impossible for us interpret as illustrated. Thus, it will be necessary to get more specimens from or near the type locality of D. conicollis to clarify the status of D. conicollis and D. japonicus. Because of the variability observed in the structure of the mandibles and male sternite VIII for D. japonicus, D. conicollis, at present can be only reliably distinguished from D. japonicus by the reported smaller length of forebody (2.0 – 2.2 mm), given by Zhou & Zhou (2016). Due to some slight differences in the aedeagus between specimens of D. japonicus from Hainan and those from other populations, we only tentatively treat Hainan as part of this species’ distribution.	en	Shuai, Qi, Nozaki, Tsubasa, Tang, Liang (2021): Notes on the genus Diochus Erichson, with descriptions of two new species from China (Coleoptera, Staphylinidae). Zootaxa 4908 (2): 276-282, DOI: 10.11646/zootaxa.4908.2.8
03A89857A952FF8E6A9192F3FB87F9B8.taxon	description	(Figs 2, 16 – 22)	en	Shuai, Qi, Nozaki, Tsubasa, Tang, Liang (2021): Notes on the genus Diochus Erichson, with descriptions of two new species from China (Coleoptera, Staphylinidae). Zootaxa 4908 (2): 276-282, DOI: 10.11646/zootaxa.4908.2.8
03A89857A952FF8E6A9192F3FB87F9B8.taxon	materials_examined	Type material. Holotype. China: Guizhou: ♁, glued on a card with labels as follows: “ China: Guizhou Province, Libo county, Maolan N. R., Bizuo, 25 ° 16 ’ 59 ’’ N, 108 ° 03 ’ 18 ’’ E, 587 m, mixed leaf litter, sifted, 28. IV. 2017, Jiang, Jiang, Hu, Liu & Zhang leg. ” “ Holotype / Diochus guizhouensis / Shuai, Nozaki & Tang ” [red handwritten label] (SHNU). Paratypes. 1 ♁, same data as for the holotype (SHNU); 1 ♀, Libo county, Maolan N. R., Bizuo, 25 ° 17 ’ 16 ’’ N, 108 ° 04 ’ 18 ’’ E, 478 – 507 m, mixed leaf litter, sifted, 26. IV. 2017, Jiang, Jiang, Hu, Liu & Zhang leg. (SHNU).	en	Shuai, Qi, Nozaki, Tsubasa, Tang, Liang (2021): Notes on the genus Diochus Erichson, with descriptions of two new species from China (Coleoptera, Staphylinidae). Zootaxa 4908 (2): 276-282, DOI: 10.11646/zootaxa.4908.2.8
03A89857A952FF8E6A9192F3FB87F9B8.taxon	description	Description. Body dark brown, head and elytra slightly darker, portions along the posterior margins of abdominal segments lighter, appendages reddish yellow with antennal segments 4 – 8 slightly darker. BL: 5.9 – 6.1 mm, FL: 2.9 – 3.0 mm. HL: 0.78 – 0.81 mm, HW: 0.60 – 0.62 mm, EyL: 0.16 – 0.22 mm, TL: 0.46 – 0.48 mm, PL: 0.99 – 1.07 mm, PW: 0.83 – 0.87 mm, EL: 1.01 – 1.03 mm, EW: 1.07 – 1.09 mm, SL: 0.68 – 0.69 mm. Head 1.27 – 1.31 times as long as wide, tempora slightly widened posterad, posterior angles rounded, eyes 0.38 – 0.50 times as long as tempora. Dorsal surface shiny, microsculpture distinct and transverse, punctation coarse and sparse. Mandibles (Fig. 19) falciform, each with a large tooth and a small tooth on inner margin. Antennae slender, antennomere 3 distinctly longer than 2 (ratio: 1.18 – 1.25). Gular sutures confluent at about basal 1 / 4, then extended to base. Pronotum 1.20 – 1.23 times as long as wide, wider than head, lateral margins widened posteriad to basal 1 / 3, then convergent backwards, anterior and posterior angles broadly rounded. Dorsal surface shiny without microsculpture. Disc with one row of punctures composing of 3 or 4 large punctures on each side of the midline and 1 or 2 additional punctures outside of each row of punctures. Elytra 0.93 – 0.95 times as long as wide, shorter than pronotum at suture, lateral margins divergent posteriad, dorsal surface shiny without microsculpture; each elytron with a row of 5 punctures near suture and a row of 5 punctures in median row. First four segments of protarsi moderately dilated in both sexes. Abdomen with dense and fine punctation and with distinct microsculpture, posterior margin of tergite VII with distinct palisade fringe. Male. Head with a small round elevation between the eyes; sternite VIII (Fig. 21) with posterior margin emarginated at middle; sternite IX (Fig. 20) symmetrical, widest near the middle; aedeagus (Figs 16 – 18) with parameres as long as 1 / 3 length of the median lobe, internal structures sclerotized, consisting of paired apical sclerites and a single Y-shaped basal plate. Female. Head without a small spot elevation between eyes; sternite VIII with posterior margin rounded; spermatheca (Fig. 22) sclerotized, very long, with three bends.	en	Shuai, Qi, Nozaki, Tsubasa, Tang, Liang (2021): Notes on the genus Diochus Erichson, with descriptions of two new species from China (Coleoptera, Staphylinidae). Zootaxa 4908 (2): 276-282, DOI: 10.11646/zootaxa.4908.2.8
03A89857A952FF8E6A9192F3FB87F9B8.taxon	distribution	Distribution. China (Guizhou).	en	Shuai, Qi, Nozaki, Tsubasa, Tang, Liang (2021): Notes on the genus Diochus Erichson, with descriptions of two new species from China (Coleoptera, Staphylinidae). Zootaxa 4908 (2): 276-282, DOI: 10.11646/zootaxa.4908.2.8
03A89857A952FF8E6A9192F3FB87F9B8.taxon	discussion	Remarks. The new species is similar to D. japonicus Cameron, 1930 in appearance, but can be distinguished from the latter by male frons with a small round elevation between the eyes, male sternite VIII without coarse setae and different morphology of aedeagus and spermatheca.	en	Shuai, Qi, Nozaki, Tsubasa, Tang, Liang (2021): Notes on the genus Diochus Erichson, with descriptions of two new species from China (Coleoptera, Staphylinidae). Zootaxa 4908 (2): 276-282, DOI: 10.11646/zootaxa.4908.2.8
03A89857A952FF8E6A9192F3FB87F9B8.taxon	etymology	Etymology. The specific name is derived from the locality, Guizhou, of the species.	en	Shuai, Qi, Nozaki, Tsubasa, Tang, Liang (2021): Notes on the genus Diochus Erichson, with descriptions of two new species from China (Coleoptera, Staphylinidae). Zootaxa 4908 (2): 276-282, DOI: 10.11646/zootaxa.4908.2.8
03A89857A952FF8C6A91943FFF3CFEAC.taxon	description	(Figs 3, 23 – 29)	en	Shuai, Qi, Nozaki, Tsubasa, Tang, Liang (2021): Notes on the genus Diochus Erichson, with descriptions of two new species from China (Coleoptera, Staphylinidae). Zootaxa 4908 (2): 276-282, DOI: 10.11646/zootaxa.4908.2.8
03A89857A952FF8C6A91943FFF3CFEAC.taxon	materials_examined	Type material. Holotype. China: Yunnan: ♁, glued on a card with labels as follows: “ China: Yunnan Province, Dehongmang, Yingjiang County, Nabang Town, 24 ° 42 ’ 59.10 ’’ N, 97 ° 35 ’ 04.54 ’’ E, 356 m, mixed leaf litter, sifted, 23. VII. 2019, Cheng & Shen leg. ” “ Holotype / Diochus caudapiscis / Shuai, Nozaki & Tang ” [red handwritten label] (SHNU). Paratypes. 1 ♁ 2 ♀♀, same data as for the holotype (SHNU); 1 ♁ 2 ♀♀, Dehongmang, Yingjiang County, Tongbiguan Town, 24 ° 36 ’ 13.78 ’’ N, 97 ° 39 ’ 50.41 ’’ E, 1407 m, mixed leaf litter, sifted, 20. VII. 2019, Cheng & Shen leg. (SHNU); 1 ♁, Dehongmang, Yingjiang County, Nabang Town, 24 ° 42 ’ 05.46 ’’ N, 97 ° 34 ’ 25.18 ’’ E, 369 m, mixed leaf litter, sifted, 24. VII. 2019, Cheng & Shen leg. (SHNU); 1 ♀, Dehongmang, Yingjiang County, Taiping Town, Near Mangna Road, 24 ° 37 ’ 42.99 ’’ N, 97 ° 45 ’ 34 ’’ E, 1471 m, mixed leaf litter, sifted, 28. VII. 2019, Cheng & Shen leg. (SHNU); 4 ♀♀, Dehongmang, Yingjiang County, Tongbiguan Town, near Mangna Road, 24 ° 36 ’ 59.75 ’’ N, 97 ° 44 ’ 30.89 ’’ E, 1550 m, mixed leaf litter, sifted, 29. VII. 2019, Cheng & Shen leg. (SHNU); 1 ♁, Lushui County, Liuku, Qingshan Park, 7. VII. 2010, Liang Tang leg. (SHNU).	en	Shuai, Qi, Nozaki, Tsubasa, Tang, Liang (2021): Notes on the genus Diochus Erichson, with descriptions of two new species from China (Coleoptera, Staphylinidae). Zootaxa 4908 (2): 276-282, DOI: 10.11646/zootaxa.4908.2.8
03A89857A952FF8C6A91943FFF3CFEAC.taxon	description	Description. Body dark brown with pronotum and portions along posterior margins of abdominal segments lighter, appendages reddish yellow with antennal segments 4 – 11 slightly darker. Similar to Diochus guizhouensis sp. nov. except for the measurements and sexual characters. BL: 4.8 – 5.8 mm, FL: 2.7 – 2.9 mm. HL: 0.68 – 0.80 mm, HW: 0.57 – 0.62 mm, EyL: 0.17 – 0.21 mm, TL: 0.43 – 0.47 mm, PL: 1.01 – 1.07 mm, PW: 0.72 – 0.89 mm, EL: 0.98 – 1.09 mm, EW: 1.05 – 1.12 mm, SL: 0.68 – 0.76 mm. Head 1.28 – 1.36 times as long as wide; pronotum 1.17 – 1.23 times as long as wide; elytra 0.88 – 0.99 times as long as wide. Male. Head with a small round elevation between eyes; sternite VIII (Fig. 28) with posterior margin emarginate at middle, lateral corner produced apicolaterad; sternite IX (Fig. 27) symmetrical, widest near the middle; aedeagus (Figs 23 – 25) with parameres as long as 1 / 3 length of the median lobe, internal structures sclerotized, consisting of paired apical sclerites and a long basal plate. Female. Head without a small round elevation between eyes; sternite VIII with posterior margin rounded; spermatheca (Fig. 29) sclerotized, very long, with three bends.	en	Shuai, Qi, Nozaki, Tsubasa, Tang, Liang (2021): Notes on the genus Diochus Erichson, with descriptions of two new species from China (Coleoptera, Staphylinidae). Zootaxa 4908 (2): 276-282, DOI: 10.11646/zootaxa.4908.2.8
03A89857A952FF8C6A91943FFF3CFEAC.taxon	distribution	Distribution. China (Yunnan).	en	Shuai, Qi, Nozaki, Tsubasa, Tang, Liang (2021): Notes on the genus Diochus Erichson, with descriptions of two new species from China (Coleoptera, Staphylinidae). Zootaxa 4908 (2): 276-282, DOI: 10.11646/zootaxa.4908.2.8
03A89857A952FF8C6A91943FFF3CFEAC.taxon	discussion	Remarks. The new species is similar to D. guizhouensis sp. nov. in appearance, but can be distinguished from the latter by produced apical angles of male sternite VIII and different morphology of aedeagus and spermatheca.	en	Shuai, Qi, Nozaki, Tsubasa, Tang, Liang (2021): Notes on the genus Diochus Erichson, with descriptions of two new species from China (Coleoptera, Staphylinidae). Zootaxa 4908 (2): 276-282, DOI: 10.11646/zootaxa.4908.2.8
03A89857A952FF8C6A91943FFF3CFEAC.taxon	etymology	Etymology. The specific name caudapiscis means fish tail, which is derived from the shape of male sternite VIII.	en	Shuai, Qi, Nozaki, Tsubasa, Tang, Liang (2021): Notes on the genus Diochus Erichson, with descriptions of two new species from China (Coleoptera, Staphylinidae). Zootaxa 4908 (2): 276-282, DOI: 10.11646/zootaxa.4908.2.8
