taxonID	type	description	language	source
03A8A444FF8FC468FF08FF45FD5DFB9B.taxon	description	Section Petermannia	en	Sang, Julia, Kiew, Ruth (2016): Begonia (Begoniaceae) from Batang Ai National Park and vicinity, Sarawak, Borneo, including six new species. Phytotaxa 252 (1): 17-30, DOI: 10.11646/phytotaxa.252.1.2, URL: http://dx.doi.org/10.11646/phytotaxa.252.1.2
03A8A444FF8FC468FF08FF45FD5DFB9B.taxon	diagnosis	Diagnosis: — Similar to Begonia amidalae C. W. Lin & C. - I Peng in its habit and leaf shape but it is different in being a smaller plant with stem to 20 cm tall (vs. stem 30 – 100 cm tall in B. amidalae), lamina plain olive green, margin serrate (vs. lamina dark green with silver patches, margin entire to inconspicuously dentate), inflorescences with fewer male flowers (vs. inflorescences much-branched with more than 30 flowers), male flowers with small tepals ca. 3 × 3 mm and ca. 18 stamens (vs. male tepals 5 – 6 × 4 – 5 mm and 7 – 11 stamens).	en	Sang, Julia, Kiew, Ruth (2016): Begonia (Begoniaceae) from Batang Ai National Park and vicinity, Sarawak, Borneo, including six new species. Phytotaxa 252 (1): 17-30, DOI: 10.11646/phytotaxa.252.1.2, URL: http://dx.doi.org/10.11646/phytotaxa.252.1.2
03A8A444FF8FC468FF08FF45FD5DFB9B.taxon	materials_examined	Type: — MALAYSIA. Borneo. Sarawak. Lubok Antu District: Batang Ai National Park, Lubang Baya, Bebiyong Mit trail, adjacent to the Ranger Post, 1 ° 18 ’ 00 ” N and 112 ° 04 ’ 10 ” E, 200 m, 2 August 2015, Julia et al. SFC 2762 (holotype SAR!; isotypes KEP!, SNP!). Erect begonia to 20 cm tall, flowering at ca. 4 cm tall, whole plant glabrous. Stems dark brown, little-branched, woody, 4 – 5 mm thick, internodes 1.5 – 3.5 cm long, thicker at nodes. Stipules pale green, lanceolate, 5 – 8 × 2 mm, margin entire, keeled, apex acuminate, acumen 1 – 2 mm long, caducous. Leaves alternate, distant, not oblique, slightly asymmetrical, held more or less horizontally; petioles dark brown, 0.5 – 1.2 cm long, slightly grooved above; lamina plain olive green above, paler beneath, in life slightly succulent, matt, slightly asymmetric, narrowly ovate to almost elliptic, slightly falcate, 9.5 – 14 × 3.3 – 5 cm, broad side 1.7 – 3.5 cm wide, base slightly rounded on the broader side, cuneate on the narrower side, sometimes unequal, basal lobes scarcely developed, margin serrate, apex acuminate, acumen 1.2 – 1.5 cm long; venation pinnate, veins greenish above, dark brown near the leaf base beneath, 3 – 5 veins on either side of the midrib, prominent on both surfaces. Inflorescences protogynous, axillary in the upper leaf axils, racemose, rachis scarcely branched, 5 – 12.5 cm long with 1 – 2 female flowers below and 2 – 4 cymules of male flowers at the apex. Bracts pale green, linear, ca. 11 × 2 mm, margin entire, caducous; bracteoles similar to bracts but smaller, 3 – 4 × 1 – 2 mm, caducous. Male flowers with greenish or yellowish green pedicels, 3 – 4 mm long; tepals 2, yellowish green, oval, ca. 3 × 3 mm, margin entire, apex rounded; stamens ca. 18, cluster conical, subsessile; filaments pale yellow, 0.6 – 1 mm long; anthers pale yellow, obovate, ca. 0.8 × 0.4 mm, apex emarginate. Female flowers with pale green or yellowish green pedicels, 7 – 9 mm long, glabrous; ovary pale green, ovoid, 6 – 7 × 8 – 12 mm, glabrous, wings 3, subequal, locules 3, placentas 2 per locule; tepals 5, pale green or yellowish green, glabrous, outer 4 tepals elliptic, 6 – 7 × 3 – 4 mm, margin entire, apex acute, inner tepal narrowly elliptic, ca. 5 × 2 mm, margin entire, apex acute; styles 3, pale yellow, 2 – 3 mm long, divided to base, shallowly Y-shaped; stigma pale yellow, papillose forming a continuous twisted band. Capsules single or in a pair, pale green, ovate, 8 – 10 × 10 – 18 mm, glabrous, locules 3, wings 3, equal, almost rounded proximally, truncate with rounded wings tips distally, 3 – 6 mm wide, thinly fibrous, dehiscing between locule and wing; pedicel fine and thread-like, pendent, 7 – 13 mm long.	en	Sang, Julia, Kiew, Ruth (2016): Begonia (Begoniaceae) from Batang Ai National Park and vicinity, Sarawak, Borneo, including six new species. Phytotaxa 252 (1): 17-30, DOI: 10.11646/phytotaxa.252.1.2, URL: http://dx.doi.org/10.11646/phytotaxa.252.1.2
03A8A444FF8FC468FF08FF45FD5DFB9B.taxon	etymology	Etymology: — Latin, bayae = of the crocodile; name derived from the type locality ‘ Lubang Baya’ in local Iban language which means ‘ crocodile hole’.	en	Sang, Julia, Kiew, Ruth (2016): Begonia (Begoniaceae) from Batang Ai National Park and vicinity, Sarawak, Borneo, including six new species. Phytotaxa 252 (1): 17-30, DOI: 10.11646/phytotaxa.252.1.2, URL: http://dx.doi.org/10.11646/phytotaxa.252.1.2
03A8A444FF8FC468FF08FF45FD5DFB9B.taxon	distribution	Distribution: — MALAYSIA. Borneo. Sarawak. Endemic in Sarawak. So far only known from Lubang Baya in Batang Ai National Park. Habitat: — Lowland mixed dipterocarp forest below 200 m elevation, on slopes above the river bank. Proposed conservation status: — Least Concern as the species occurs within a Totally Protected Area and their habitat is not threatened by any local disturbance.	en	Sang, Julia, Kiew, Ruth (2016): Begonia (Begoniaceae) from Batang Ai National Park and vicinity, Sarawak, Borneo, including six new species. Phytotaxa 252 (1): 17-30, DOI: 10.11646/phytotaxa.252.1.2, URL: http://dx.doi.org/10.11646/phytotaxa.252.1.2
03A8A444FF8CC466FF08FBB0FAF7FD87.taxon	description	Section Petermannia	en	Sang, Julia, Kiew, Ruth (2016): Begonia (Begoniaceae) from Batang Ai National Park and vicinity, Sarawak, Borneo, including six new species. Phytotaxa 252 (1): 17-30, DOI: 10.11646/phytotaxa.252.1.2, URL: http://dx.doi.org/10.11646/phytotaxa.252.1.2
03A8A444FF8CC466FF08FBB0FAF7FD87.taxon	diagnosis	Diagnosis: — Similar to Begonia vulgare S. Julia & Kiew (2013: 26) in habit, leaf shape and inflorescence that has a basal pair of female flowers and male flowers in a dense cluster above but it is different in its shorter stem to 60 cm tall (vs. to 2 m tall in B. vulgare), longer petioles 4 – 13 cm long (vs. 0.5 – 3.5 cm), shorter inflorescences, 1.5 – 5 cm (vs. 9 – 19 cm) and the shorter fruit, 15 – 25 × 12 – 24 mm (vs. the more oblong fruit, 35 – 45 × 17 – 27 mm in B. vulgare).	en	Sang, Julia, Kiew, Ruth (2016): Begonia (Begoniaceae) from Batang Ai National Park and vicinity, Sarawak, Borneo, including six new species. Phytotaxa 252 (1): 17-30, DOI: 10.11646/phytotaxa.252.1.2, URL: http://dx.doi.org/10.11646/phytotaxa.252.1.2
03A8A444FF8CC466FF08FBB0FAF7FD87.taxon	materials_examined	Type: — MALAYSIA. Borneo. Sarawak. Lubok Antu District: Batang Ai, Sungai Jengin, 1 ° 15 ’ 00 ” N and 112 ° 06 ’ 00 ” E, 127 m, 1 August 2015, Julia et al. SFC 2751 (holotype SAR!; isotypes KEP!, SNP!). Cane-like begonia 15 – 60 cm tall. Stems erect, reddish, 12 – 14 mm thick, little-branched, succulent, minutely pubescent; internodes 3 – 8 (– 16) cm long, thicker at nodes. Stipules brownish, lanceolate, 11 – 19 (– 28) × 5 – 8 (– 12) mm, margin entire, keeled, apex acuminate, caducous. Leaves alternate, distant, oblique, held horizontally; petioles reddish, (4 –) 8 – 13 cm long, grooved, minutely pubescent; lamina plain green on both surfaces, in life succulent, drying papery, matt, ovate to obovate, 15 – 22 × 8 – 17 cm, broad side 5.5 – 15 cm wide, often markedly concave on the narrower side, base slightly cordate, sometimes slightly overlapping, basal lobes 0.75 – 1.1 cm, margin shallowly dentate, apex sharply acuminate, 1.5 – 2 cm long; venation palmate-pinnate, veins pale green on both surfaces, minutely pubescent, 3 – 5 veins on either side of the midrib, 3 – 5 veins in the basal lobe, prominent on both surfaces. Inflorescences protogynous, axillary in the upper leaf axils, rachis 1.5 – 5 cm long, with a pair of female flowers below and a dense cluster of male flowers above, each cymule enveloped in upto 2 pairs of bracteoles. Bracts reddish, lanceolate, 12 – 20 × 8 – 12 mm, margin entire, caducous; bracteoles broadly lanceolate, setose at apex, increasing in size to 11 – 13 × 8 – 10 mm, then abruptly diminishing in size towards the developing apex. Male flowers with pale pink pedicels, 4 – 8 mm long; tepals 2, pale yellowish green, oval, 5 – 6 × 3 – 5 mm, margin entire, apex rounded; stamens 29 – 32, cluster globose, subsessile; filaments pale yellow, 0.8 – 1 mm long; anthers pale yellow, obovate, ca. 1.5 × 0.8 mm, apex emarginate. Female flowers with greenish pedicels, 2 – 4 mm long; ovary pale green, wings tips pale green, obovoid, 9 – 10 × 8 – 12 mm, wings 3, equal, locules 3, placentas 2 per locule; tepals 5, pale green, outer 4 tepals ovate, 7 – 8 × 5 – 6 mm, margin entire, apex broadly acute, inner tepals ovate, ca. 6 × 5 mm, margin entire, apex rounded; styles 3, pale yellow, 3 – 4 mm long, divided to base, deeply Y-shaped; stigma pale yellow with reddish stripe, papillose forming a continuous twisted band. Capsules single or a pair, pale green, triangular or sometimes to slightly oblong-triangular in outline, 15 – 25 × 12 – 24 mm, glabrous, locules 3, wings 3, equal, rounded proximally and expanded and pointed distally, wings 6 – 7 mm wide, thinly fibrous, dehiscing between locule and wing; pedicel pendent, 7 – 11 mm long, reddish towards the base.	en	Sang, Julia, Kiew, Ruth (2016): Begonia (Begoniaceae) from Batang Ai National Park and vicinity, Sarawak, Borneo, including six new species. Phytotaxa 252 (1): 17-30, DOI: 10.11646/phytotaxa.252.1.2, URL: http://dx.doi.org/10.11646/phytotaxa.252.1.2
03A8A444FF8CC466FF08FBB0FAF7FD87.taxon	etymology	Etymology: — Latin, compacta = close-packed, referring to the compact male inflorescence subtended by many pairs of bracteoles.	en	Sang, Julia, Kiew, Ruth (2016): Begonia (Begoniaceae) from Batang Ai National Park and vicinity, Sarawak, Borneo, including six new species. Phytotaxa 252 (1): 17-30, DOI: 10.11646/phytotaxa.252.1.2, URL: http://dx.doi.org/10.11646/phytotaxa.252.1.2
03A8A444FF8CC466FF08FBB0FAF7FD87.taxon	distribution	Distribution: — MALAYSIA. Borneo. Sarawak. Endemic in Sarawak, known only from the Batang Ai. Habitat: — Riparian forest to 127 m elevation on steep earth or rocky slopes above the river bank in light shade. Additional specimen examined: — MALAYSIA. Borneo. Sarawak. Lubok Antu District: Batang Ai National Park, Sungai Bebiyong Besai, 2 August 2015, Julia et al. SFC 2760 (KEP, SAR, SNP). Proposed conservation status: — Least Concern as the species occurs within a Totally Protected Area and their habitat is not threatened by any local disturbance. 6. Begonia darthvaderiana C. W. Lin & C. - I Peng (2014: 132). Type: — MALAYSIA. Borneo, Sarawak. Lubok Antu District: Batang Ai, 5 July 2013, C. W. Lin 549 (holotype SAR n. v.; isotypes E, HAST, KEP, TAIF). Distribution: — Endemic in Sarawak; known only from Batang Ai in Lubok Antu District. Habitat: — In lowland mixed dipterocarp forest at 130 – 200 m elevation in valleys, on lower to upper soil slopes or soil at the cliff base in deep shade. Notes: — The species was collected from forest areas outside the Batang Ai National Park boundary (Lin, C. W. pers. comm.). So far no record of this species has been recorded within the Park boundary. Proposed conservation status: — Data Deficient. The species was previously recorded from forest area outside the park boundary and the species occurrence within the park boundary is yet to be ascertained. It may also be under pressure from commercial collecting because it is currently the most sought after begonia from Sarawak.	en	Sang, Julia, Kiew, Ruth (2016): Begonia (Begoniaceae) from Batang Ai National Park and vicinity, Sarawak, Borneo, including six new species. Phytotaxa 252 (1): 17-30, DOI: 10.11646/phytotaxa.252.1.2, URL: http://dx.doi.org/10.11646/phytotaxa.252.1.2
03A8A444FF82C467FF08FD8BFD5DF8A5.taxon	description	Section Petermannia	en	Sang, Julia, Kiew, Ruth (2016): Begonia (Begoniaceae) from Batang Ai National Park and vicinity, Sarawak, Borneo, including six new species. Phytotaxa 252 (1): 17-30, DOI: 10.11646/phytotaxa.252.1.2, URL: http://dx.doi.org/10.11646/phytotaxa.252.1.2
03A8A444FF82C467FF08FD8BFD5DF8A5.taxon	diagnosis	Diagnosis: — It most resembles Begonia matangensis S. Julia & Kiew (2015: 95) in its habit, its dense hispid indumentums and its subopposite velvety leaves but it is different in its broader leaves about twice as long as wide (vs. leaves about three times longer than wide in B. matangensis), its female flowers with dimorphic tepals with acute apex (vs. isomorphic tepals with pointed apex), style 3 – 4 mm long, more than half of the length of the tepals with only a shallow Y-shaped stigma (vs. style ca. 2 mm long, less than half of the tepal length and a with wide Y-shaped stigma) and a broader capsule, triangular in outline and 10 – 15 × 11 – 14 mm with wider wings, 4 – 6 mm wide (vs. an oblong capsule ca. 10 × 8 mm with narrower wings, 2 – 3 mm wide).	en	Sang, Julia, Kiew, Ruth (2016): Begonia (Begoniaceae) from Batang Ai National Park and vicinity, Sarawak, Borneo, including six new species. Phytotaxa 252 (1): 17-30, DOI: 10.11646/phytotaxa.252.1.2, URL: http://dx.doi.org/10.11646/phytotaxa.252.1.2
03A8A444FF82C467FF08FD8BFD5DF8A5.taxon	materials_examined	Type: — MALAYSIA. Borneo, Sarawak. Sarawak. Lubok Antu District: Batang Ai National Park, Sungai Bebiyong Besai, 1 ° 19 ’ 00 ” N and 112 ° 04 ’ 00 ” E, 222 m, 2 August 2015, Julia et al. SFC 2758 (holotype SAR!; isotypes BRUN!, KEP!, SNP!). Erect begonia 26 – 45 cm tall. Indumentum of stem, stipules and petioles densely hispid, hairs white, 5 – 6 mm long. Stem branched, dark brown, succulent, slender, 3 – 4 mm thick, internodes (0.7 –) 3 – 8 cm long, thicker at nodes. Stipules reddish brown, 6 – 12 × 4 – 5 mm, margin entire, keeled, apex acuminate, seta to 6 mm long, caducous. Leaves alternate below, upper leaves sub-opposite, pairs distant, held horizontally, not oblique; petiole brownish, 0.3 – 0.5 cm long, slightly grooved above; lamina plain dark green above, paler beneath, velvety, densely covered in appressed hairs ca. 2 mm long, less dense beneath, in life papery and soft (not fleshy), matt, slightly asymmetric, obovate, slightly falcate, 6 – 14 × 2.5 – 6.5 cm, broad side 1.5 – 4 cm wide, base unequal, slightly lobed on narrow side, rounded on broad side, basal lobe rounded, 0.2 – 0.8 cm long, margin ciliate, minutely dentate in the lower half, becoming more deeply dentate towards the apex and appearing scalloped, apex acuminate, acumen 1 – 1.5 cm long; venation pinnate, 4 – 6 veins on either side of the midrib, concolorous on both surfaces, densely pilose, 1 – 2 veins in basal lobe, prominent on both surfaces, more so beneath. Inflorescences on short shoots below the subopposite leaf, protogynous, densely pilose, female flower single or occasionally in a pair, male flowers in umbellate clusters with individual flowers subtended by a bract. Bracts and bracteoles similar to stipules, persistent. Male flowers with white pedicels, sometimes dull red, 2 – 5 mm long; tepals 2, white or greenish white at the centre, sometimes dull red, densely pilose outside, hairs white, glabrous inside, oval, 4 – 5 × 3 mm, margin entire, apex acute; stamens 14 – 16, cluster conical, stalked to 0.5 mm long; filaments whitish, ca. 1 mm long; anthers pale yellow, obovate, 1 – 1.2 × 0.5 mm, apex emarginate. Female flowers with greenish white pedicels 2 – 6 mm long, sometimes reddish, sparsely hairy; ovary greenish yellow, sometimes reddish, sparsely hairy, oblong, 11 – 15 × 12 – 13 mm, wings 3, equal, locules 3, placentas 2 per locule; tepals 5, pale greenish yellow to white tinged pink or reddish, sparsely pilose outside, glabrous inside, lanceolate; outer 2 tepals 6 – 9 × 2 – 3 mm, margin entire, apex acute; inner 3 tepals 6 – 8 × 1.5 – 3 mm, styles 3, pale green, anchor-shaped, 3 – 4 mm long, divided to base, papillose forming a continuous twisted band. Capsules single, axillary, hispid, 10 – 15 × 11 – 14 mm, triangular in outline, narrowed proximally, expanded distally to a sharp point, locules 3, wings 3, equal, wings 4 – 6 mm wide, papery, dehiscing between the locule and wing; pedicel 2 – 5 mm long.	en	Sang, Julia, Kiew, Ruth (2016): Begonia (Begoniaceae) from Batang Ai National Park and vicinity, Sarawak, Borneo, including six new species. Phytotaxa 252 (1): 17-30, DOI: 10.11646/phytotaxa.252.1.2, URL: http://dx.doi.org/10.11646/phytotaxa.252.1.2
03A8A444FF82C467FF08FD8BFD5DF8A5.taxon	etymology	Etymology: — Named after Edgar Irmscher (1887 – 1968), a German botanist expert in Begonia, who in 1953 described 11 Begonia species collected by Odoardo Beccari from Sarawak (Irmscher, 1953) and who explained the growth pattern of begonias with subopposite leaves.	en	Sang, Julia, Kiew, Ruth (2016): Begonia (Begoniaceae) from Batang Ai National Park and vicinity, Sarawak, Borneo, including six new species. Phytotaxa 252 (1): 17-30, DOI: 10.11646/phytotaxa.252.1.2, URL: http://dx.doi.org/10.11646/phytotaxa.252.1.2
03A8A444FF82C467FF08FD8BFD5DF8A5.taxon	distribution	Distribution: — MALAYSIA. Borneo. Sarawak. Endemic in Sarawak, known only from the Batang Ai National Park. Habitat: — Lowland mixed dipterocarp forest, to 222 m elevation on steep or gentle slopes in deep shade. Notes: — This species is common where it occurs. Two colour forms of leaves and flowers were observed in the field: (i) plants with green or bronzy-dark green leaves and (ii) male flowers usually white or occasionally dull red and the female flowers usually pale green or white tinged pink, occasionally dull red. However, the leaf colour is not associated with flower colour, thus plants with bronzy leaves may have white flowers and those with green leaves have red flowers. The species is monoecious but because the male phase does not overlap with the female, individual plants tend to have either female or male flowers. Proposed conservation status: — Least Concern as the species occurs within a Totally Protected Area and their habitat is not threatened by any local disturbance.	en	Sang, Julia, Kiew, Ruth (2016): Begonia (Begoniaceae) from Batang Ai National Park and vicinity, Sarawak, Borneo, including six new species. Phytotaxa 252 (1): 17-30, DOI: 10.11646/phytotaxa.252.1.2, URL: http://dx.doi.org/10.11646/phytotaxa.252.1.2
03A8A444FF83C465FF08F8E9FD5DFB0F.taxon	description	Section Petermannia	en	Sang, Julia, Kiew, Ruth (2016): Begonia (Begoniaceae) from Batang Ai National Park and vicinity, Sarawak, Borneo, including six new species. Phytotaxa 252 (1): 17-30, DOI: 10.11646/phytotaxa.252.1.2, URL: http://dx.doi.org/10.11646/phytotaxa.252.1.2
03A8A444FF83C465FF08F8E9FD5DFB0F.taxon	diagnosis	Diagnosis: — Similar to Begonia awongii Sands (1997: 432) in its habit, hispid inflorescence, leaf shape and racemose inflorescence but differs in the shallowly cordate base with a basal lobe 1 – 2 cm long (vs. deeply cordate with a basal lobe 5.5 – 8.5 cm long in B. awongii), male flowers with pale pink male tepals and fewer (28 – 30) but larger stamens with filaments ca. 2 mm long and anthers ca. 1.5 mm long (vs. white tepals and ca. 46 subsessile stamens with filaments ca. 0.5 mm long and anthers ca. 0.25 mm long), smaller tepals in the female flower, 6 – 7 × 4 – 6 mm (vs. 12 – 14 × 7 – 9 mm) and larger glabrous capsules, 12 – 14 × 14 – 24 mm (vs. densely hispid and 7 – 12 × 16 – 18 mm).	en	Sang, Julia, Kiew, Ruth (2016): Begonia (Begoniaceae) from Batang Ai National Park and vicinity, Sarawak, Borneo, including six new species. Phytotaxa 252 (1): 17-30, DOI: 10.11646/phytotaxa.252.1.2, URL: http://dx.doi.org/10.11646/phytotaxa.252.1.2
03A8A444FF83C465FF08F8E9FD5DFB0F.taxon	materials_examined	Type: — MALAYSIA. Borneo. Sarawak. Lubok Antu District: Batang Ai, Sungai Jengin, 1 ° 15 ’ 00 ” N and 112 ° 06 ’ 00 ” E, 194 m, 1 August 2015, Julia et al. SFC 2750 (holotype SAR!; isotypes BRUN!, KEP!, SNP!). Cane-like begonia to 1.35 m tall. Stems reddish brown, pilose, hairs brownish, terete and finely ridged, 6 – 8 mm thick, internodes 4 – 6 (– 10) cm, swollen at the nodes. Stipules narrowly or broadly lanceolate, 12 – 18 × 4 – 11 mm, keeled terminating in a seta 1 – 2 mm long, hispid outside. Leaves alternate, very oblique, held horizontally or sometimes slightly vertical; petioles reddish brown, (3 –) 5 – 10 (– 13.5) cm long, grooved, hispid; lamina plain green above, paler beneath, above sparsely hispid between veins, veins densely hispid, glabrous beneath except on veins, in life succulent, membranous when dry, matt, broadly ovate, 13.5 – 25 × 11.5 – 20.5 cm, broad side 9 – 14.5 cm wide, base shallowly cordate, basal lobes 1 – 2 cm, margin minutely dentate, apex acuminate, 1.5 – 2 cm long; venation palmate, prominent on both surfaces, 4 – 6 veins radiating from the base, in life veins pale green on both surfaces, reddish brown towards the base, drying reddish, 3 – 4 veins along the midrib on the broad side, 1 – 2 veins in the basal lobe. Inflorescences protogynous, axillary in the upper leaf axils, racemose, rachis unbranched, (2.5 –) 10 – 15 cm long. Bracts greenish, lanceolate, 12 – 17 × 4 – 5 mm, margin entire, apex setose to 4 mm long, caducous; bracteoles similar to bracts but smaller, 4 – 10 × 4 mm, caducous. Male flowers with pale pink pedicels, 5 – 10 mm long; tepals 4, pale pink, reddish towards base, oval, outer 2 tepals 8 – 10 × 5 – 8 mm, inner 2 tepals 5 – 8 × 2 – 4 mm, margin entire, apex rounded; stamens 28 – 30, cluster globose, subsessile; filaments pale yellow, ca. 2 mm long; anthers pale yellow, obovate, ca. 1.5 × 1 mm, apex emarginate. Female flowers with greenish pedicels, 4 – 8 mm long, glabrous; ovary pale green, wing tips pale green, ovate, 7 – 8 × 8 – 11 mm, wings 3, equal, locules 3, placentas 1 per locule; tepals 5, ovate, 6 – 7 × 4 – 6 mm, margin entire, apex rounded; styles 3, pale yellow, 2 – 3 mm long, divided to base, widely Y-shaped; stigma reddish, papillose cells large and pale yellow forming a continuous twisted band. Capsules 2 – 4 pairs along the rachis, greenish yellow, glabrous, 12 – 14 × 14 – 24 mm, glabrous, locules 3, wings 3, equal, rounded proximally and distally or truncate distally, 3 – 6 mm wide, thinly fibrous, dehiscing between locule and wing; pedicel pendent, 6 – 15 mm long, glabrous, reddish towards the base.	en	Sang, Julia, Kiew, Ruth (2016): Begonia (Begoniaceae) from Batang Ai National Park and vicinity, Sarawak, Borneo, including six new species. Phytotaxa 252 (1): 17-30, DOI: 10.11646/phytotaxa.252.1.2, URL: http://dx.doi.org/10.11646/phytotaxa.252.1.2
03A8A444FF83C465FF08F8E9FD5DFB0F.taxon	etymology	Etymology: — Named after the type locality, Jengin River (Sungai = River) where the species was discovered.	en	Sang, Julia, Kiew, Ruth (2016): Begonia (Begoniaceae) from Batang Ai National Park and vicinity, Sarawak, Borneo, including six new species. Phytotaxa 252 (1): 17-30, DOI: 10.11646/phytotaxa.252.1.2, URL: http://dx.doi.org/10.11646/phytotaxa.252.1.2
03A8A444FF83C465FF08F8E9FD5DFB0F.taxon	distribution	Distribution: — MALAYSIA. Borneo. Sarawak. Endemic in Sarawak where the species is known only from Sungai Bebiyong Besai in Batang Ai National Park and Sungai Jengin in Batang Ai. Habitat: — Riparian forest at 194 m elevation, growing on steep earth slopes above the river bank or on boulders with thin layer of soil in light shade. Additional specimen examined: — MALAYSIA. Borneo. Sarawak. Lubok Antu District: Batang Ai National Park, Sungai Bebiyong Besai, Julia et al. SFC 2755 (SAR). Notes: — The species is more common along the bank of Sungai Jengin than along Sungai Bebiyong Besai. Proposed conservation status: — Least Concern as the species occurs within a Totally Protected Area and their habitat is not threatened by any local disturbance.	en	Sang, Julia, Kiew, Ruth (2016): Begonia (Begoniaceae) from Batang Ai National Park and vicinity, Sarawak, Borneo, including six new species. Phytotaxa 252 (1): 17-30, DOI: 10.11646/phytotaxa.252.1.2, URL: http://dx.doi.org/10.11646/phytotaxa.252.1.2
03A8A444FF81C463FF08FB03FD5DFE5B.taxon	description	Section Petermannia	en	Sang, Julia, Kiew, Ruth (2016): Begonia (Begoniaceae) from Batang Ai National Park and vicinity, Sarawak, Borneo, including six new species. Phytotaxa 252 (1): 17-30, DOI: 10.11646/phytotaxa.252.1.2, URL: http://dx.doi.org/10.11646/phytotaxa.252.1.2
03A8A444FF81C463FF08FB03FD5DFE5B.taxon	diagnosis	Diagnosis: — Similar to Begonia johariana S. Julia & C. Y. Ling (2015: 110) in its slender creeping stem, small leaves, racemose inflorescence with large bracteoles and its small fruits wider than long but it is different in its succulent stem 3 – 4 mm thick (vs. wiry and 1 – 3 mm thick in B. johariana), its larger stipules 10 – 15 mm long with an entire margin (vs. 5 – 8 mm long with a serrate margin), longer inflorescences 4 – 8.5 cm long with larger bracts 7 – 17 × 4 – 11 mm (vs. inflorescences to 5 cm long and bracts 5 – 7 × 2 – 4 mm) and outer tepals of female flower less than twice as long as wide (vs. 2 – 3 times as long as wide).	en	Sang, Julia, Kiew, Ruth (2016): Begonia (Begoniaceae) from Batang Ai National Park and vicinity, Sarawak, Borneo, including six new species. Phytotaxa 252 (1): 17-30, DOI: 10.11646/phytotaxa.252.1.2, URL: http://dx.doi.org/10.11646/phytotaxa.252.1.2
03A8A444FF81C463FF08FB03FD5DFE5B.taxon	materials_examined	Type: — MALAYSIA. Borneo. Sarawak. Lubok Antu District: Batang Ai National Park, Lubang Baya, Bebiyong Mit trail, adjacent to the Ranger Post, 1 ° 18 ’ 00 ” N and 112 ° 04 ’ 10 ” E, 200 m, 2 August 2015, Julia et al. SFC 2763 (holotype SAR!; isotypes KEP!, SNP!). Creeping herb. Indumentum on stems and petioles pubescent. Stems dull brown, slender, 3 – 4 mm thick, branched, succulent, internodes 2.5 – 8.5 cm long, slightly thicker at nodes. Stipules pale green, broadly ovate, 10 – 15 × 7 – 10 mm, margin entire and ciliate, apex acuminate, acumen ca. 1 mm long, caducous. Leaves alternate, distant, slightly oblique, held more or less horizontally; petioles dull brown, 1 – 3.5 (– 5) cm long, slightly grooved; lamina plain green above, paler beneath, in life slightly fleshy, glossy, with scattered short hairs, asymmetric, ovate, 5.5 – 8 (– 10.5) × 4.5 – 6 (– 8.5) cm, broad side 2.5 – 4 (– 5.5) cm wide, base slightly cordate, sometimes overlapping, basal lobes 0.7 – 1.5 cm long, margin minutely serrate, teeth more prominent at the vein endings, apex acuminate, acumen to 1 cm long; venation palmate-pinnate, veins greenish above, dark brown towards the base above and beneath, minutely pubescent above and beneath, 2 pairs at the base and 3 – 4 veins on either side of the midrib, 2 – 3 veins in basal lobe, slightly impressed above, prominent beneath. Inflorescences protogynous, axillary in upper leaf axils, racemose, slender, 4 – 8.5 cm long, sometimes with a single lateral branch 1.5 – 2 cm long with a female flower. Bracts pale green, broadly lanceolate, (7 –) 10 – 12 (– 17) × (4 –) 6 – 7 (– 11) mm, margin ciliate, persistent; bracteoles 3 – 4 pairs, lower bracteoles subtending female flowers oval, pale green, ca. 8 × 4.5 mm, similar to bracts; upper bracteoles subtending male flowers broadly oval, 6 – 7 × 5 – 6 mm, sometimes tinged reddish at the tip, keeled, margin ciliate, persistent. Male flowers with pinkish or white pedicels, 2 – 5 mm long, pubescent; tepals 4, white sometimes tinged red or reddish, glabrous, margin entire, apex rounded, outer 2 tepals broadly elliptic, 6 – 8 × 4 – 6 mm, inner 2 tepals lanceolate, 4 – 5 × 2 – 3 mm; stamens 42 – 46, cluster globose, subsessile; filaments pale yellow, 0.8 – 1.1 mm long; anthers pale yellow, obovate, 0.6 – 1 × 0.2 – 0.4 mm, apex emarginate. Female flowers with pale green pedicels, ca. 1 mm long, glabrous; ovary pale green with reddish wings, broadly ovate, 5 – 6 × 14 – 15 mm, glabrous, wings 3, unequal, locules 3, placentas 2 per locule; tepals 5, reddish or greenish outside, white tinged reddish inside, glabrous, outer 3 tepals elliptic, 5 – 6 × 3 – 5 mm, margin entire, apex acute, inner 2 tepals slightly smaller, ca. 4 × 2 mm; styles 3, pale yellow, ca. 1 mm long, divided to base, shallow Yshaped; stigma pale yellow, papillose forming a continuous twisted band. Capsules broadly obovate, 6 – 7 × 9 – 14 mm, glabrous, locules 3, wings 3, equal, broadly rounded, acute proximally, rounded or truncate distally, 3 – 4 mm wide, thinly fibrous, dehiscing between locule and wing; pedicel slender, pendent, 5 – 8 mm long.	en	Sang, Julia, Kiew, Ruth (2016): Begonia (Begoniaceae) from Batang Ai National Park and vicinity, Sarawak, Borneo, including six new species. Phytotaxa 252 (1): 17-30, DOI: 10.11646/phytotaxa.252.1.2, URL: http://dx.doi.org/10.11646/phytotaxa.252.1.2
03A8A444FF81C463FF08FB03FD5DFE5B.taxon	etymology	Etymology: — From local Iban word tebiang = slope, referring to the habitat where the species grows abundantly.	en	Sang, Julia, Kiew, Ruth (2016): Begonia (Begoniaceae) from Batang Ai National Park and vicinity, Sarawak, Borneo, including six new species. Phytotaxa 252 (1): 17-30, DOI: 10.11646/phytotaxa.252.1.2, URL: http://dx.doi.org/10.11646/phytotaxa.252.1.2
03A8A444FF81C463FF08FB03FD5DFE5B.taxon	distribution	Distribution: — Endemic in Sarawak, as yet known only from Batang Ai National Park. Habitat: — Lowland mixed dipterocarp forest at elevations below 200 m on steep or gentle earth slopes in deep to light shade areas. Additional specimens examined: — MALAYSIA. Borneo. Sarawak. Lubok Antu District: Batang Ai National Park, Nanga Lubang Baya, Bebiyong Mit Trail, Julia et al. SFC 2764 (KEP, SAR), Rantai et al. S 88048 (SAR), Nanga Sungai Lelayang, Rantai et al. S 82793 (SAR). Proposed conservation status: — Least Concern as the species occurs within a Totally Protected Area and their habitat is not threatened by any local disturbance.	en	Sang, Julia, Kiew, Ruth (2016): Begonia (Begoniaceae) from Batang Ai National Park and vicinity, Sarawak, Borneo, including six new species. Phytotaxa 252 (1): 17-30, DOI: 10.11646/phytotaxa.252.1.2, URL: http://dx.doi.org/10.11646/phytotaxa.252.1.2
