identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03A9E573FFAF165D057FF9157062FE05.text	03A9E573FFAF165D057FF9157062FE05.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cnemaspis spinicollis (Müller 1907) Muller 1907	<div><p>Cnemaspis spinicollis (Müller, 1907)</p><p>Perret (1985, 1986) considered this species to have a broad range from Côte d’Ivoire to Cameroon, but did not include Bénin in his summary of distribution. In Cameroon, where it is most well­documented, it has been recorded chiefly from forested habitats (Perret 1986; Lawson 1993) and from montane savanna (Böhme 1975). Dunger (1968) reported several localities in southwestern Nigeria (Akure, Yemoji, Ibadan) and Joger (1981) provided two localities in Togo. Although no material from Bénin has yet been identified, it is a virtual certainty that C. spinicollis is present.</p></div>	https://treatment.plazi.org/id/03A9E573FFAF165D057FF9157062FE05	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bauer, Aaron M.;Tchibozo, Sévérin;Pauwels, Olivier S. G.;Lenglet, Georges	Bauer, Aaron M., Tchibozo, Sévérin, Pauwels, Olivier S. G., Lenglet, Georges (2006): A review of the gekkotan lizards of Bénin, with the description of a new species of Hemidactylus (Squamata: Gekkonidae). Zootaxa 1242: 1-20, DOI: 10.5281/zenodo.273464
03A9E573FFAF165A057FFC5A72E2F9B5.text	03A9E573FFAF165A057FFC5A72E2F9B5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hemitheconyx caudicinctus (Duméril 1851) Dumeril 1851	<div><p>Hemitheconyx caudicinctus (Duméril, 1851)</p><p>This is the only eublepharid gecko occurring in the region. It is widespread, chiefly in savanna areas from Senegal to Nigeria (Loveridge 1947). Dunger (1968), who provided numerous localities for H. caudicinctus in Nigeria, considered it widespread across the Guinea Savanna. Both Loveridge (1947) and Grandison (1956) explicitly stated that the range of the species included Bénin, but apparently only on the basis of localities originally reported by Chabanaud (1917). This species is common in the pet trade and commercially purchased specimens from Bénin have also been used in laboratory studies, accounting for numerous museum specimens without specific locality data.</p><p>Localities</p><p>Kétou, Département du Plateau: CAS 165588; Diho (=Idiho), Département des Collines (8°05’N, 2°31’E): USNM 199564­65; Agougon (=Agouagon), Département des Collines: Chabanaud (1917); Haute­Dahomey: Chabanaud (1917); no specific locality: FMNH 170554 ­55; MCZ R 104068­69; YPM 8126 (3 specimens), 8129, 8171 (12 specimens), 8416.</p></div>	https://treatment.plazi.org/id/03A9E573FFAF165A057FFC5A72E2F9B5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bauer, Aaron M.;Tchibozo, Sévérin;Pauwels, Olivier S. G.;Lenglet, Georges	Bauer, Aaron M., Tchibozo, Sévérin, Pauwels, Olivier S. G., Lenglet, Georges (2006): A review of the gekkotan lizards of Bénin, with the description of a new species of Hemidactylus (Squamata: Gekkonidae). Zootaxa 1242: 1-20, DOI: 10.5281/zenodo.273464
03A9E573FFA8165D057FFDBF70EAFB10.text	03A9E573FFA8165D057FFDBF70EAFB10.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lygodactylus conraui Tornier 1902	<div><p>Lygodactylus conraui Tornier, 1902</p><p>This small, diurnal gecko is extremely widespread in West Africa from Sierra Leone in the west to Cameroon and possibly Equatorial Guinea in the east and south (Loveridge 1947; Pasteur 1965; Dunger 1968). Pasteur (1965) noted that no specimens had been recorded from either Togo or Bénin. The specimens reported here thus represent the first confirmed record for the country. In addition, we also report a specimen (IRSNB 17158) of this species from the Forêt de Dzogbegan (07°04’15”N, 00°38’15”E), Togo (Fig. 2), constituting the first record for this country as well. Variation within this species throughout its range is high (Perret 1963; Pasteur 1965; van Eijsden 1978). The specimens from Togo and Bénin both definitively key out to L. conraui, but they differ substantially from one another. Whether this reflects intraspecific variation or the presence of multiple taxa within L. conraui as presently construed requires further investigation. The Bénin specimens were collected in swamp forest.</p><p>Locality</p><p>Lokoli, Département du Zou: IRSNB 17161­1, 17161­2.</p></div>	https://treatment.plazi.org/id/03A9E573FFA8165D057FFDBF70EAFB10	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bauer, Aaron M.;Tchibozo, Sévérin;Pauwels, Olivier S. G.;Lenglet, Georges	Bauer, Aaron M., Tchibozo, Sévérin, Pauwels, Olivier S. G., Lenglet, Georges (2006): A review of the gekkotan lizards of Bénin, with the description of a new species of Hemidactylus (Squamata: Gekkonidae). Zootaxa 1242: 1-20, DOI: 10.5281/zenodo.273464
03A9E573FFA8165D057FFADD724BF955.text	03A9E573FFA8165D057FFADD724BF955.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lygodactylus gutturalis (Bocage 1873) Bocage 1873	<div><p>Lygodactylus gutturalis (Bocage, 1873)</p><p>Loveridge (1947) listed numerous localities for this species (as L. picturatus gutturalis) elsewhere in both East and West Africa, but none in countries bordering the eastern Gulf of Guinea. Pasteur (1965) did not specify the localities of material he examined, but depicted the species’ range as extending from Senegal and southernmost Mauritania to the Horn of Africa, and incorporating all but the southernmost parts of Bénin. We were unable to confirm vouchered records of this species for Bénin, but Dunger (1968) reported material from Kadur in western Nigeria and it is clear that this species must also be present in Bénin.</p></div>	https://treatment.plazi.org/id/03A9E573FFA8165D057FFADD724BF955	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bauer, Aaron M.;Tchibozo, Sévérin;Pauwels, Olivier S. G.;Lenglet, Georges	Bauer, Aaron M., Tchibozo, Sévérin, Pauwels, Olivier S. G., Lenglet, Georges (2006): A review of the gekkotan lizards of Bénin, with the description of a new species of Hemidactylus (Squamata: Gekkonidae). Zootaxa 1242: 1-20, DOI: 10.5281/zenodo.273464
03A9E573FFA8165C057FF8E27788FBAD.text	03A9E573FFA8165C057FF8E27788FBAD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ptyodactylus ragazzi Anderson 1898	<div><p>Ptyodactylus ragazzi Anderson, 1898</p><p>Loveridge (1947) regarded Ptyodactylus from northern Togo as referable to P.</p><p>hasselquistii togoensis Tornier, 1901 and referred P. h. ragazzi, with its type from Eritrea, to the synonymy of P. h. hasselquistii Donndorff. However, Heimes (1987) demonstrated that P.h. ragazzi and P.h. togoensis were synonyms and resurrected the older name for the subspecies ranging across the central Sahara and the Sahel. He considered the range in West Africa to span from Côte d’Ivoire to Cameroon, but did not examine any material from Bénin. Schleich et al. (1996) subsequently accorded full species status to P. ragazzi, but without explicit justification. Baha El Din (1999) provided evidence of sympatry between P. hasselquistii and P. ragazzi in southeastern Egypt, demonstrating that they should indeed be considered distinct species. The species is widespread in northern areas of Nigeria (Dunger 1968; Butler 1986) and Cameroon (LeBreton 1999). Dunger (1968) considered P. ragazzi to be a rupicolous or crevice­dwelling savanna species, although Butler (1986) collected it at the Jebba crossing of the Niger River in western Nigeria in a small shed. There appear to be no prior published records of this species from Bénin. Our material was collected on boulders (Fig. 3) or abandoned buildings adjacent to boulders, confirming earlier ecological observations from Nigeria. The largest individual collected (IRSNB 17151­3) measured 77.7 mm SVL.</p><p>Localities</p><p>Chutes de Koudou, Parc National du W du Bénin, Département de l’Alibori (11°38’N, 02°18’E): IRSNB 17151­1, 17151­2, 17151­3; Parc National du W du Bénin, Département de l’Alibori: IRSNB 17154; without specific locality: IRSNB 17171­1, 17171­4.</p></div>	https://treatment.plazi.org/id/03A9E573FFA8165C057FF8E27788FBAD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bauer, Aaron M.;Tchibozo, Sévérin;Pauwels, Olivier S. G.;Lenglet, Georges	Bauer, Aaron M., Tchibozo, Sévérin, Pauwels, Olivier S. G., Lenglet, Georges (2006): A review of the gekkotan lizards of Bénin, with the description of a new species of Hemidactylus (Squamata: Gekkonidae). Zootaxa 1242: 1-20, DOI: 10.5281/zenodo.273464
03A9E573FFAA165E057FF9AE7115FE55.text	03A9E573FFAA165E057FF9AE7115FE55.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tarentola ephippiata O'Shaughnessy 1875	<div><p>Tarentola ephippiata O’Shaughnessy, 1875</p><p>Grandison (1961) clarified the taxonomic status of this species, recognizing it as distinct from T. annularis Geoffroy Saint Hilaire and indicating a broad range in West Africa, although she did not examine material from Bénin. Joger (1984) considered the species to range from the Côte d’Ivoire to Cameroon and north into Mali, Burkina Faso, Niger and Tchad, but implied that no specimens were actually recorded from Bénin, although he considered their occurrence certain. Our record below is thus the first confirmed record of T. epphipiata for Bénin. Dunger (1968) reported this species from fig trees and in­ and outside human dwellings and other buildings in northern Nigeria. Our specimens were taken in savanna habitat.</p></div>	https://treatment.plazi.org/id/03A9E573FFAA165E057FF9AE7115FE55	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bauer, Aaron M.;Tchibozo, Sévérin;Pauwels, Olivier S. G.;Lenglet, Georges	Bauer, Aaron M., Tchibozo, Sévérin, Pauwels, Olivier S. G., Lenglet, Georges (2006): A review of the gekkotan lizards of Bénin, with the description of a new species of Hemidactylus (Squamata: Gekkonidae). Zootaxa 1242: 1-20, DOI: 10.5281/zenodo.273464
03A9E573FFAB165E057FFD4F7145FABA.text	03A9E573FFAB165E057FFD4F7145FABA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hemidactylus Oken 1817	<div><p>Hemidactylus Oken, 1817</p><p>Hemidactylus is the dominant gekkonid genus in West Africa. Nonetheless, relatively few species are known from the region in comparison with other parts of the continent, most notably the Horn of Africa (Lanza 1983; Bauer 1993). The genus as a whole is nearly pantropical and also occurs in warm temperate regions. It is a hugely successful group, with at least 84 recognized species (Kluge 2001; Bauer and Pauwels 2002; Henle and Böhme 2003; Baha El Din 2003, 2005; Bauer et al., in press), making it the second most speciose of all gekkonid genera. Henle and Böhme (2003) recognized ten species in West Africa: H. mabouia (Moureau de Jonnés), H. brooki (= H. angulatus Hallowell), H. matschiei (Tornier), H. fasciatus Gray, H. ansorgii Boulenger, H. richardsonii (Gray), H. kamdemtohami Bauer &amp; Pauwels, H. echinus O’Shaughnessy, H. muriceus Peters and H. pseudomuriceus Henle &amp; Böhme. An eleventh species has recently been described from northern Cameroon (Bauer et al., in press). We here confirm the presence of five of these from the territory of Bénin and consider another two Hemidactylus as probable. In addition, among our collections are two specimens not referable to any named species, which we describe here as new.</p></div>	https://treatment.plazi.org/id/03A9E573FFAB165E057FFD4F7145FABA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bauer, Aaron M.;Tchibozo, Sévérin;Pauwels, Olivier S. G.;Lenglet, Georges	Bauer, Aaron M., Tchibozo, Sévérin, Pauwels, Olivier S. G., Lenglet, Georges (2006): A review of the gekkotan lizards of Bénin, with the description of a new species of Hemidactylus (Squamata: Gekkonidae). Zootaxa 1242: 1-20, DOI: 10.5281/zenodo.273464
03A9E573FFAB1651057FFA77707FFBAD.text	03A9E573FFAB1651057FFA77707FFBAD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hemidactylus angulatus Hallowell 1852	<div><p>Hemidactylus angulatus Hallowell, 1852</p><p>This species was only recently elevated to specific rank, having been elevated from a subspecies of the Asian H. brookii, to which it is very distantly related (Carranza and Arnold 2006). It is widespread in tropical Africa. Loveridge (1947) recorded localities from Sudan to Angola and in neighboring Nigeria. Dunger (1968) considered it the most common and widespread of all Nigerian gecko species, occupying a diversity of natural and edificarian habitats. This is certainly also the most commonly encountered gecko in Bénin. It is highly variable in color pattern (Loveridge 1947; Dunger 1968) and may sometimes be confused for the broadly sympatric H. mabouia . The biology of this species was recently considered by Gramentz (2000). Hemidactylus angulatus is the only gecko species in Bénin to have been reported from multiple localities by multiple earlier authors (Chabanaud 1917; Loveridge 1947, 1952). Chabanaud (1917) recorded specimens from Agougon (=Agouagon) as both H. brookii and H. stellatus Boulenger. Most of the new material was collected from edificarian habitats, but specimens from Niaouli were found in semi­deciduous forest. The largest Bénin specimen (IRSNB 17168­3) measured 66.2 mm SVL.</p><p>Localities</p><p>Ajuda (Ouidah), Département de l’Atlantique: Loveridge (1947); Zomai (Ouidah), Département de l’Atlantique: Loveridge (1947); Godomey (=Godomé), Département de l’Atlantique: Loveridge (1947); Attogon, Département de l’Atlantique (06°43’N, 02°09’E): IRSNB 17168­1, 17168­2, 17168­3, 17168­4; Niaouli, Département de l’Atlantique (06°44’N, 02°08’E): IRSNB 17173­2; Amou Oblo, Togo, IRSNB 17166­1, 17166­2; Grand Popo, Département du Mono: Loveridge (1947); Abomey, Département du Zou: Loveridge (1952); Porto Novo, Département de l’Ouémé: Loveridge (1947); Agougon (=Agouagon), Département des Collines: Chabanaud (1917); Collines de Dassa­Zoumè, Département des Collines (07°45’N, 02°10’E): IRSNB 17150­1; Bassila, Département de la Donga: Loveridge (1952); Manigri, Département de la Donga: IRSNB 17245­1–17245­ 9; Koussokoinga (=Koussoukoingou), Département de l’Atacora: Loveridge (1952), MCZ R 51759­60; Parc National de la Pendjari, Département de l’Atacora (11°23’N, 01°31’E); IRSNB 17170­3; without specific locality: IRSNB 17171­2, 17171­3; MCZ R 159974; YPM 8133 (39 specimens), 8422 (3 specimens).</p></div>	https://treatment.plazi.org/id/03A9E573FFAB1651057FFA77707FFBAD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bauer, Aaron M.;Tchibozo, Sévérin;Pauwels, Olivier S. G.;Lenglet, Georges	Bauer, Aaron M., Tchibozo, Sévérin, Pauwels, Olivier S. G., Lenglet, Georges (2006): A review of the gekkotan lizards of Bénin, with the description of a new species of Hemidactylus (Squamata: Gekkonidae). Zootaxa 1242: 1-20, DOI: 10.5281/zenodo.273464
03A9E573FFA41651057FFB577141F9CA.text	03A9E573FFA41651057FFB577141F9CA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hemidactylus ansorgii Boulenger 1901	<div><p>Hemidactylus ansorgii Boulenger, 1901</p><p>Hemidactylus ansorgii is widespread in West Africa from Liberia to Cameroon (Loveridge 1947; Henle &amp; Böhme 2003), although as yet there are no confirmed records from Bénin. Henle and Böhme (2003) clarified the status of this species, which has been frequently misidentified in the past, thus calling in to question the validity of literature records (Dunger 1968; van Eijsden 1978) that have not been verified in this new light. Perret (1975) considered H. ansorgii as a junior synonym of H. intestinalis Werner, 1897, but Henle and Böhme (2003) have presented compelling evidence that the latter is, in reality, a junior synonym of H. muriceus .</p></div>	https://treatment.plazi.org/id/03A9E573FFA41651057FFB577141F9CA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bauer, Aaron M.;Tchibozo, Sévérin;Pauwels, Olivier S. G.;Lenglet, Georges	Bauer, Aaron M., Tchibozo, Sévérin, Pauwels, Olivier S. G., Lenglet, Georges (2006): A review of the gekkotan lizards of Bénin, with the description of a new species of Hemidactylus (Squamata: Gekkonidae). Zootaxa 1242: 1-20, DOI: 10.5281/zenodo.273464
03A9E573FFA41650057FF96D7240FE2D.text	03A9E573FFA41650057FF96D7240FE2D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hemidactylus fasciatus Gray 1842	<div><p>Hemidactylus fasciatus Gray, 1842</p><p>This large (to at least 95 mm SVL elsewhere in its range; Schmidt 1919; Burger et al. 2004) and distinctive species is broadly distributed from Liberia to the eastern Democratic Republic of Congo. It has been recorded from both forest (Loveridge 1947; Perret 1963; Lawson 1993; Bauer &amp; Pauwels 2002; Pauwels et al. 2004) and edificarian habitats (Romer 1953; Joger 1982; Ota et al. 1987; Lawson 1993) elsewhere in its range. There are numerous records from Togo (Matschie 1893b; Tornier 1901; Loveridge 1947; Joger 1981) and we also have material from the Forêt de Missahoé (IRSNB 17149, 17159, 17167), but the specimens noted here constitute the first published record of H. fasciatus for Bénin.</p></div>	https://treatment.plazi.org/id/03A9E573FFA41650057FF96D7240FE2D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bauer, Aaron M.;Tchibozo, Sévérin;Pauwels, Olivier S. G.;Lenglet, Georges	Bauer, Aaron M., Tchibozo, Sévérin, Pauwels, Olivier S. G., Lenglet, Georges (2006): A review of the gekkotan lizards of Bénin, with the description of a new species of Hemidactylus (Squamata: Gekkonidae). Zootaxa 1242: 1-20, DOI: 10.5281/zenodo.273464
03A9E573FFA51650057FFD377007FABA.text	03A9E573FFA51650057FFD377007FABA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hemidactylus mabouia	<div><p>Hemidactylus mabouia (Moreau de Jonnés, 1818)</p><p>Although very widespread in sub­Saharan Africa (Loveridge 1947; Kluge 1969; Broadley 1977), Dunger (1968) considered the species uncommon in Nigeria, and regarded populations in port areas as introduced. There appear to be no published records of this species from Bénin, so the specimens cited below constitute a new country record. This species shares with H. angulatus broad transverse subcaudal plates and largely overlapping ranges of precloacal pores and tubercle rows. It is possible that some literature records of H. angulatus may be referable to H. mabouia . The biology of this species was recently considered by Gramentz (2000, 2003). The largest Bénin specimen (IRSNB 17169) measures 67.6 mm SVL.</p><p>Localities</p><p>Godomey (=Godomé), Département de l’Atlantique (06°22’N, 02°21’E): IRSNB 17160, 17246,; Abomey, Département du Zou (07°11’N, 01°59’E): IRSNB 17169; Niaouli (06°44’N, 02°08’E): IRSNB 17173­1, 17173­3.</p></div>	https://treatment.plazi.org/id/03A9E573FFA51650057FFD377007FABA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bauer, Aaron M.;Tchibozo, Sévérin;Pauwels, Olivier S. G.;Lenglet, Georges	Bauer, Aaron M., Tchibozo, Sévérin, Pauwels, Olivier S. G., Lenglet, Georges (2006): A review of the gekkotan lizards of Bénin, with the description of a new species of Hemidactylus (Squamata: Gekkonidae). Zootaxa 1242: 1-20, DOI: 10.5281/zenodo.273464
03A9E573FFA51650057FFA7D762BF958.text	03A9E573FFA51650057FFA7D762BF958.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hemidactylus matschiei (Tornier 1901) Tornier 1901	<div><p>Hemidactylus matschiei (Tornier, 1901)</p><p>This poorly­known, savanna­dwelling species was described from Togo (Tornier 1901) and has subsequently been collected as far east as west central Nigeria at Yankari Game Reserve (Dunger 1968), suggesting that it also occurs in Bénin. It is possible that the apparent rarity of this species is yet another reflection of the long­standing confusion regarding the identity of many of the West African Hemidactylus (Henle &amp; Böhme 2003) .</p></div>	https://treatment.plazi.org/id/03A9E573FFA51650057FFA7D762BF958	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bauer, Aaron M.;Tchibozo, Sévérin;Pauwels, Olivier S. G.;Lenglet, Georges	Bauer, Aaron M., Tchibozo, Sévérin, Pauwels, Olivier S. G., Lenglet, Georges (2006): A review of the gekkotan lizards of Bénin, with the description of a new species of Hemidactylus (Squamata: Gekkonidae). Zootaxa 1242: 1-20, DOI: 10.5281/zenodo.273464
03A9E573FFA51653057FF8E270EFFD3D.text	03A9E573FFA51653057FF8E270EFFD3D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hemidactylus muriceus Peters 1870	<div><p>Hemidactylus muriceus Peters, 1870</p><p>Allocation of literature records to this species are confounded by the long standing confusion surrounding this and several other Hemidactylus that has only recently (Henle &amp; Böhme 2003) been satisfactorily resolved. Loveridge (1947) listed localities from Liberia to the Democratic Republic of Congo. Henle and Böhme (2003) did not examine all available museum material, but confirmed H. muriceus from Côte d’Ivoire to Cameroon. This includes the holotype of H. intestinalis from Missahöhe (Missahoé), Togo (see also Perret 1975). This distribution strongly suggested that H. muriceus is present in Bénin as well, but the specimens cited here constitute the first confirmed records for the country. The biology of this species was recently considered by Gramentz (2000).</p><p>Locality</p><p>vic. of Kétou, Département du Plateau: CAS 165586­87.</p></div>	https://treatment.plazi.org/id/03A9E573FFA51653057FF8E270EFFD3D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bauer, Aaron M.;Tchibozo, Sévérin;Pauwels, Olivier S. G.;Lenglet, Georges	Bauer, Aaron M., Tchibozo, Sévérin, Pauwels, Olivier S. G., Lenglet, Georges (2006): A review of the gekkotan lizards of Bénin, with the description of a new species of Hemidactylus (Squamata: Gekkonidae). Zootaxa 1242: 1-20, DOI: 10.5281/zenodo.273464
03A9E573FFA61653057FFCC7763BFBFA.text	03A9E573FFA61653057FFCC7763BFBFA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hemidactylus pseudomuriceus Henle & Bohme 2003	<div><p>Hemidactylus pseudomuriceus Henle &amp; Böhme, 2003</p><p>This recently described species is known from two localities, one in the Parc National d’Azagny, Côte d’Ivoire and the other at Jingwe (Yingui) in western Cameroon. The species inhabits primary forest near Raphia swamps. This distribution suggests that the species may be found in Bénin, perhaps in habitats such as the swamp forest of Lokoli.</p></div>	https://treatment.plazi.org/id/03A9E573FFA61653057FFCC7763BFBFA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bauer, Aaron M.;Tchibozo, Sévérin;Pauwels, Olivier S. G.;Lenglet, Georges	Bauer, Aaron M., Tchibozo, Sévérin, Pauwels, Olivier S. G., Lenglet, Georges (2006): A review of the gekkotan lizards of Bénin, with the description of a new species of Hemidactylus (Squamata: Gekkonidae). Zootaxa 1242: 1-20, DOI: 10.5281/zenodo.273464
03A9E573FFA61656057FFBBD7109FC9D.text	03A9E573FFA61656057FFBBD7109FC9D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hemidactylus beninensis	<div><p>Hemidactylus beninensis sp. nov.</p><p>Figures 4–5</p><p>Holotype</p><p>Institut Royal des Sciences Naturelles de Belgique (IRSNB) 2617, adult male; Bénin, Département des Collines, Collines de Dassa­Zoumè, (07°45’N, 02°10’E); collected by Sévérin Tchibozo, 13 June 2005. Paratype. IRSNB 2618, subadult male; same data as holotype.</p><p>Diagnosis</p><p>A moderate­sized Hemidactylus, snout­vent length at least 68 mm. One pair of greatly enlarged postmentals, in contact behind mental. 14–16 rows of relatively small tubercles. 49 scale rows across venter between lowest rows of tubercles. Six to seven enlarged paired scansors beneath fourth toes of both manus and pes. A single row of 38 precloacal­femoral pores in adult males. Original tail atuberculate, with median subcaudal scales forming broad transverse plates. Dorsal pattern with a series of four complete, dark crossmarkings between nape and sacrum, alternating with incomplete transverse bands extending dorsally form the flanks but failing to meet along the dorsal midline.</p><p>Hemidactylus beninensis may be distinguished from all other mainland West African congeners on the basis of (sympatric taxa with differing or non­overlapping character states indicated parenthetically): 14–16 rows of dorsal tubercles (0 rows in H. matschiei, 0–4 rows in H. richardsonii, 8–10 in H. ansorgii, 18–24 rows in H. fasciatus, 18–20 rows in H. n. sp. from Cameroon), precloacal­femoral pores in male in single row of 38 (fewer than 12 in H. echinus, H. muriceus, and H. ansorgii, 14–17 in H. pseudomuriceus, 45 in H. n. sp. from Cameroon), median subcaudal scale row greatly enlarged transversely (not enlarged or hexagonally enlarged in H. echinus, H. kamdemtohami, H. pseudomuriceus, H. muriceus, and H. ansorgii). Both H. angularis and H. mabouia share overlapping precloacal­femoral pore counts and transversely enlarged subcaudal scales with H. beninensis; however, both species have conspicuous tubercles on the dorsum of the tail, whereas that in H. beninensis . The new species is most similar in general appearance to a newly described species of Hemidactylus from Cameroon (Bauer et al., in press), with which it shares a mostly smooth dorsal tail surface (atuberculate in H. beninensis, very weakly tuberculate in H. n. sp.), transversely enlarged subcaudal scales, and a similar dorsal color pattern. However, these taxa differ with respect to ( H. beninensis versus H. n. sp.): size (68 versus 100 mm SVL), scansors beneath the fourth toe (6–7 versus 10–11), precloacal­femoral pores (38 versus 45 in the respective holotypes), and rows of dorsal tubercles (14–16 versus 18–20).</p><p>Description (based on the holotype, IRSNB 2617)</p><p>Adult male, SVL 68.3 mm. Head long (HeadL/SVL ratio 0.29), relatively narrow (HeadW/HeadL ratio 0.67), somewhat depressed (HeadH/HL ratio 0.44), strongly distinct from neck. Lores and interorbital region slightly inflated. Snout moderate (SnEye/HeadL ratio 0.41), less than twice eye diameter (OrbD/SnEye ratio 0.57); scales on snout and forehead small, granular; scales on snout much larger than those on occipital region. Eye relatively large (OrbD/HeadL ratio 0.24); pupil vertical with crenelated margins; supraciliaries short, without spines. Ear opening oval, relatively small (EarL/HeadL ratio 0.06); eye to ear distance greater than diameter of eyes (EyeEar/OrbD ratio 1.22). Rostral approximately 1.5 times wider (3.0 mm) than deep (1.9 mm), incompletely divided dorsally by a deeply­incised rostral groove; two enlarged supranasals separated by a single internasal; rostral in contact with supralabial I, supranasals, and internasal; nostrils oval, each surrounded by supranasal, rostral, first supralabial (narrow contact), and three postnasals, the middle one of which smallest, with very narrow contact with nostril rim; each nostril with a recessed valvular scale at posteromedial border; 3–4 rows of scales separate orbit from supralabials. Mental triangular, minimally deeper (3.2 mm) than wide (2.9 mm) and much deeper than infralabials; one pair of greatly enlarged postmentals meeting at a point behind the mental, each postmental bordered anteriorly by first infralabial, medially by mental, and laterally and posteriorly by a series of four chin shields, the lateralmost much enlarged and also bordering first and second infralabials; remaining three chinshields smaller, approximately four times the size of a granular gular scale. Infralabials bordered by a row of enlarged and axially elongated scales, decreasing in size posteriorly. Enlarged supralabials to midpoint of orbit 8 (left)–9 (right); supralabials to angle of jaws 10 (right)–13 (left); infralabials 10 (left)–9 (right); interorbital scale rows across narrowest point of frontal 14, between supraciliaries 36.</p><p>Body moderately slender, trunk relatively long (TrunkL/SVL ratio 0.40), oval in crosssection, with very feebly developed ventrolateral folds without denticulate scales. Dorsal scales heterogeneous, granular; regularly arranged, small (4 times size of granules), weakly conical to slightly keeled, erect to posteriorly­directed tubercles extending from behind occiput to tail base; no tubercles on temporal region; tubercles more or less uniform across dorsum, somewhat more prominent on flanks; tubercles in 14 rows at midbody. Ventral scales larger than dorsal tubercles, subimbricate; somewhat larger on abdomen than on chest, granular and much smaller in gular region; midbody scale rows across belly to lowest row of tubercles 49. Precloacal­femoral pores in a single row of 38, with a single poreless scales in midline separating continuous series of 19 pored scales on each side, extending nearly to knees. Scales on palm and sole smooth, flat, rounded; scales on dorsal aspects of limbs heterogeneous – granular, intermixed with larger domed to conical tubercules, particularly on knees and shanks.</p><p>Fore­ and hindlimbs moderately long, stout; forearm and tibia moderately long (ForeaL/SVL ratio 0.15; CrusL/SVL ratio 0.17); digits relatively short, strongly clawed; all digits of manus and pes webbed proximally only; distal portions of digits curved, arising from distal portion of expanded subdigital pad; scansors beneath each toe divided, except for distalmost and a few, basal scansors (usually 0–1); scansors from proximalmost at least twice diameter of palmar scales to distalmost divided scansor: 5­6­7­7­7 (left manus), 5­7­6­7­7 (right manus), 5­7­8­7­6 (left pes), 4­7­7­6­6 (right pes). Relative length of digits of manus: IV ~ III&gt; V&gt; II&gt; I; of pes: IV&gt; III&gt; V&gt; II&gt; I.</p><p>Original portion of tail depressed; regenerated portion more slender; length of partly regenerated tail approximately equal to snout­vent length (TailL/SVL ratio 0.97); basal part of original portion of tail covered dorsally with small, posteriorly­pointed, subimbricate to imbricate scales, atuberculate; scales forming regular transverse rows, about 8 such rows per distinct caudal segment (corresponding to underlying muscle segments); ventral scales much larger, smooth, imbricate, with a median row of greatly enlarged subcaudal plates extending nearly across the width of tail. Regenerated portion of tail like original but without segmentation. One small, rounded postcloacal spur on each side of tail base.</p><p>Mensural features (holotype / paratype). SVL 68.3/ 54.3 mm, ForeaL 10.5/ 8.3 mm, CrusL 11.5/ 9.9 mm, TailL 66.5 (31.3 regenerated)/51.3 (5.9 regenerated) mm, TailW 7.3/ 5.1 mm, TrunkL 27.2/21.0 mm, HeadL 19.7/ 16.4 mm, HeadW 13.2/ 10.3 mm, HeadH 8.7/ 6.1 mm, OrbD 4.6/ 4.1 mm, EyeEar 5.6/ 4.3 mm, SnEye 8.1/ 6.4 mm, NarEye 5.5/ 4.2 mm, Interorb 2.4/ 1.7 mm, EarL 1.3/ 1.5 mm, Internar 2.1/ 1.5 mm.</p><p>Osteology</p><p>Parietal bones paired. Stapes imperforate. Phalangeal formulae 2­3­4­5­3 for manus and 2­3­4­5­4 for pes. Presacral vertebrae 26, including 3 anterior cervical (without ribs), 1 lumbar, and 2 sacral vertebrae; 5 pygal and 8.5 post pygal caudal vertebrae to point of regeneration (18.5 in paratype). One pair of slender, crescentic cloacal bones present in both holotype and paratype. Endolymphatic sacs enlarged extracranially, extending to level of 4th–6th vertebrae in both specimens. Radiographs reveal that the holotype is skeletally mature, as is confirmed by its well­developed precloacal­femoral pores, whereas the subadult paratype exhibits incomplete ossification of the carpal and tarsal elements.</p><p>Coloration (in preservative)</p><p>Body grayish with a series of dark grayish­brown crossbands with darker brown borders; one across nape, one behind axilla, one at mid­trunk, and one in lumbar position. These complete dorsal bands alternating with bands extending dorsally from flanks but not meeting in the dorsal midline, first over scapula. Tips of tubercles whitish, particularly on the flanks. Parietal and temporal regions of head with grayish­brown markings without darker margins. A thick white to cream stripe from lateral surface of rostral and anterior of first supralabial to anterodorsal corner of orbit ­ continuing behind orbit and on to occiput as a series of irregular pale blotches. This stripe bordered above by a grayish­brown stripe extending from nostril to top of orbit and below by a similarly colored triangular marking extending posteriorly from the 2nd to 3rd supralabial scales to ventral half of the anterior margin of the orbit. Limbs mottled with alternating brown and pale gray blotches, particularly distal to the knees and elbows, basal limb segments mostly grayish with faint darker irregular markings. Tail with alternating markings as on dorsum; two complete, dark­bordered brown bands on original portion of post­pygal tail; regenerated portion grayish­brown with darker speckles, without distinct pattern. Venter grayish­cream becoming pale gray subcaudally.</p><p>Va r i a t i o n</p><p>The subadult paratype differs from the holotype in the following features: postmentals bordered posteriorly by 3 chin shields; supralabials 11, 9 to midpoint of orbit (L), 8(R); infralabials 9 (L&amp;R); dorsal tubercles in 16 rows at midbody; no precloacal or femoral pores; subdigital lamellae: (left manus) 4­6­7­7­7, (right manus) 4­7­7­7­6, (left pes) 4­7­ 7­7­6, (right pes) 4­7­7­7­6. Paratype similar in color to holotype, but more boldly patterned with four complete dark bands on original portion of post­pygal tail.</p><p>Distribution and natural history</p><p>At present known only from the type locality in the relatively hilly region of the Département des Collines in central Bénin (Fig. 6). The area lies adjacent to the Nigerian border and it is likely that this species occurs in that country as well. A juvenile Hemidactylus angulatus was collected at the same site as the types of H. beninensis . The holotype exhibits damage to the skin of the dorsum of the head (Fig. 1). Such damage is indicative of mechanically weak skin, which is associated with regional integumentary loss, an escape mechanism employed against certain predators by a variety of geckos (Bauer et al. 1989, 1993).</p></div>	https://treatment.plazi.org/id/03A9E573FFA61656057FFBBD7109FC9D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bauer, Aaron M.;Tchibozo, Sévérin;Pauwels, Olivier S. G.;Lenglet, Georges	Bauer, Aaron M., Tchibozo, Sévérin, Pauwels, Olivier S. G., Lenglet, Georges (2006): A review of the gekkotan lizards of Bénin, with the description of a new species of Hemidactylus (Squamata: Gekkonidae). Zootaxa 1242: 1-20, DOI: 10.5281/zenodo.273464
