taxonID	type	description	language	source
03B687F62B2BFF8EFCC4C9DD59FB5A2E.taxon	description	(Figs 1 - 5)	en	Ober, Stefan V., Staniczek, Arnold H. (2009): A new genus and species of coenagrionid damselflies (Insecta, Odonata, Zygoptera, Coenagrionidae) from Vanuatu. Zoosystema 31 (3): 485-497, DOI: 10.5252/z2009n3a6, URL: http://www.bioone.org/doi/abs/10.5252/z2009n3a6
03B687F62B2BFF8EFCC4C9DD59FB5A2E.taxon	materials_examined	TYPE SPECIES. — Nesobasis malekulana Kimmins, 1936, by present designation.	en	Ober, Stefan V., Staniczek, Arnold H. (2009): A new genus and species of coenagrionid damselflies (Insecta, Odonata, Zygoptera, Coenagrionidae) from Vanuatu. Zoosystema 31 (3): 485-497, DOI: 10.5252/z2009n3a6, URL: http://www.bioone.org/doi/abs/10.5252/z2009n3a6
03B687F62B2BFF8EFCC4C9DD59FB5A2E.taxon	etymology	ETYMOLOGY. — The name of the related genus Nesobasis refers to the Greek νησΟς (nesos = island) and βάση (basi = basis). The suffix “ basis ” is common in many A new genus and species of Coenagrionidae (Insecta, Odonata) from Vanuatu zygopteran genera. Accordingly, the name of the new genus is composed of “ Vanuatu ”, which refers to its presently known distribution, and the suffix “ basis ”.	en	Ober, Stefan V., Staniczek, Arnold H. (2009): A new genus and species of coenagrionid damselflies (Insecta, Odonata, Zygoptera, Coenagrionidae) from Vanuatu. Zoosystema 31 (3): 485-497, DOI: 10.5252/z2009n3a6, URL: http://www.bioone.org/doi/abs/10.5252/z2009n3a6
03B687F62B2BFF8EFCC4C9DD59FB5A2E.taxon	distribution	DISTRIBUTION. — Currently, the genus is only known from the islands of Aneityum, Espiritu Santo, Maewo, and Malekula, Vanuatu.	en	Ober, Stefan V., Staniczek, Arnold H. (2009): A new genus and species of coenagrionid damselflies (Insecta, Odonata, Zygoptera, Coenagrionidae) from Vanuatu. Zoosystema 31 (3): 485-497, DOI: 10.5252/z2009n3a6, URL: http://www.bioone.org/doi/abs/10.5252/z2009n3a6
03B687F62B2BFF8EFCC4C9DD59FB5A2E.taxon	diagnosis	DIAGNOSIS. — Vanuatubasis n. gen. resembles Nesobasis, but differs from the latter in the following characters: 1) superior anal appendages of males broad and short (Fig. 4); 2) inferior anal appendages significantly longer than superior appendages, forceps-like, apically curved inwards, each ending in a black tip (Fig. 4); and 3) pronotal hind lobe raised and medially protruding, tapering to a rounded or acute apex (Fig. 5).	en	Ober, Stefan V., Staniczek, Arnold H. (2009): A new genus and species of coenagrionid damselflies (Insecta, Odonata, Zygoptera, Coenagrionidae) from Vanuatu. Zoosystema 31 (3): 485-497, DOI: 10.5252/z2009n3a6, URL: http://www.bioone.org/doi/abs/10.5252/z2009n3a6
03B687F62B2BFF8EFCC4C9DD59FB5A2E.taxon	discussion	REMARK Vanuatubasis n. gen. currently contains three species, one of which is described as new in this contribution.	en	Ober, Stefan V., Staniczek, Arnold H. (2009): A new genus and species of coenagrionid damselflies (Insecta, Odonata, Zygoptera, Coenagrionidae) from Vanuatu. Zoosystema 31 (3): 485-497, DOI: 10.5252/z2009n3a6, URL: http://www.bioone.org/doi/abs/10.5252/z2009n3a6
03B687F62B28FF89FF05C93A5D035AC0.taxon	description	(Figs 1 - 3; 4 C, F, I, L; 5 B)	en	Ober, Stefan V., Staniczek, Arnold H. (2009): A new genus and species of coenagrionid damselflies (Insecta, Odonata, Zygoptera, Coenagrionidae) from Vanuatu. Zoosystema 31 (3): 485-497, DOI: 10.5252/z2009n3a6, URL: http://www.bioone.org/doi/abs/10.5252/z2009n3a6
03B687F62B28FF89FF05C93A5D035AC0.taxon	materials_examined	HOLOTYPE. — Vanuatu. Sanma Province, Espiritu Santo, surroundings of Penaoru, Penaoru River, 14.96105 ° S, 166.63316 ° E, 90 m, 13. XI. 2006, leg. A. H. Staniczek & M. Pallmann, ♂ (SMNS: ODO 000242 K). PARATYPES. — Vanuatu. Sanma Province, Espiritu Santo, surroundings of Tasmate, Mamasa River, 15.20976 ° S, 166.67705 ° E, 20 m, 9. XI. 2006, leg. A. H. Staniczek & M. Pallmann, 2 ♂♂ (MNHN); 2 ♂♂ (SMNS: ODO 000243 K, ODO 000244 K). — Sanma Province, Espiritu Santo, surroundings of Tasmate, Paé River, 15.21751 ° S, 166.68706 ° E, 139 m, 11. XI. 2006, leg. A. H. Staniczek & M. Pallmann, 1 ♂ (SMNS: ODO 000245 K). — Sanma Province, Espiritu Santo, surroundings of Penaoru, Penaoru River, 14.96105 ° S, 166.63316 ° E, 90 m, 13. XI. 2006, leg. A. H. Staniczek & M. Pallmann, 2 ♂♂ (SMNS: ODO 000246 K, ODO 000247 K). — Penaoru, Camp de base, 17. XI. 2006, leg. F. Durand & E. Boitier, 2 ♂♂ (MNHN).	en	Ober, Stefan V., Staniczek, Arnold H. (2009): A new genus and species of coenagrionid damselflies (Insecta, Odonata, Zygoptera, Coenagrionidae) from Vanuatu. Zoosystema 31 (3): 485-497, DOI: 10.5252/z2009n3a6, URL: http://www.bioone.org/doi/abs/10.5252/z2009n3a6
03B687F62B28FF89FF05C93A5D035AC0.taxon	etymology	ETYMOLOGY. — The species is named after Santo, the shortened colloquial form of Espiritu Santo. ADDITIONAL MATERIAL. — Vanuatu. Sanma Province, Espiritu Santo, surroundings of Tasmate, Mamasa River, 15.20976 ° S, 166.67705 ° E, 20 m, 9. XI. 2006, leg. A. H. Staniczek & M. Pallmann, 1 ♀ (MNHN); 4 ♂♂, 1 ♀ (SMNS: ODO 000248 K-ODO 000251 K, ODO 000252 A). — Sanma Province, Espiritu Santo, surroundings of Penaoru, Penaoru River, 14.96105 ° S, 166.63316 ° E, 90 m, 13. XI. 2006, leg. A. H. Staniczek & M. Pallmann, 2 ♂♂, 2 ♀♀ (SMNS: ODO 000253 K-ODO 000255 K, ODO 000256 A). — Penaoru, Camp de base, 17. XI. 2006, leg. F. Durand & E. Boitier, 2 ♂♂ (MNHN).	en	Ober, Stefan V., Staniczek, Arnold H. (2009): A new genus and species of coenagrionid damselflies (Insecta, Odonata, Zygoptera, Coenagrionidae) from Vanuatu. Zoosystema 31 (3): 485-497, DOI: 10.5252/z2009n3a6, URL: http://www.bioone.org/doi/abs/10.5252/z2009n3a6
03B687F62B28FF89FF05C93A5D035AC0.taxon	distribution	DISTRIBUTION. — The species is only known from the island of Espiritu Santo.	en	Ober, Stefan V., Staniczek, Arnold H. (2009): A new genus and species of coenagrionid damselflies (Insecta, Odonata, Zygoptera, Coenagrionidae) from Vanuatu. Zoosystema 31 (3): 485-497, DOI: 10.5252/z2009n3a6, URL: http://www.bioone.org/doi/abs/10.5252/z2009n3a6
03B687F62B28FF89FF05C93A5D035AC0.taxon	description	DESCRIPTION OF THE HOLOTYPE Preserved dry, pinned. In good condition. Head Labium light beige; labrum aquamarine, with a very small, central brown spot at the basal margin (colour layer below the cuticula due to preservation broken and partly shifted); genae, anteclypeus, and postclypeus aquamarine; postclypeus medially brownish at base and anteriorly with a blackish triangle, on each side flanked by two brown stripes; frons aquamarine, medially changing to yellowish with slight whitish pruinosity, dorsal part blackish; basal joints of antennae aquamarine with pale end rings, each scapus black with pale end rings, pedicelli brown, paler medially, flagella dark brown; vertex and occiput black with bronze shimmer, except for a small, creamy brown spot in front of the median ocellus and the central hind margin of the occiput, diminutive postocular spots, composed of bluish-white pruinosity; head dorsally brownish, posteriorly creamy-yellowish with slight whitish pruinosity; eyes dark brown. Thorax Prothorax dorsally black with slight greenish metallic shimmer, laterally brownish and slighty covered with bluish-white pruinosity in its lower half; hind lobe of pronotum raised, each side of upper rim slightly curved and forming an acute apex, lower rim broadly V-shaped and slightly protruding posteriorly, surface between both ridges concave (Fig. 5 B). Synthorax: mesanepisternum with green-black metallic median band, reaching the mesopleural suture in its hindmost fourth, but elsewhere separated from it by a pale olive-green band, dorsal carina black; mesostigmal lamina slightly triangular, posterior margin proximally with a rounded protuberance; apex of lamina forming a slightly overhanging process (Fig. 5 B); mesokatepisternum brown to dark brown in the upper two-thirds, pale aquamarine in its lower third, entire mesokatepisternum with slightly bluish-white pruinosity; mesepimeron yellow-brown to pale aquamarine posteriorly, with blackish median stripe; metanepisternum pale aquamarine, brownish yellow anteriorly, posteriorly with small, elongate blackish spots at the intersegmental and metapleural suture; metakatepisternum brownish, covered with a bluish-white pruinosity in its lower half; metepimeron pale aquamarine becoming brownish yellow anteriorly, slightly bluish-white pruinose posteriorly; poststernum dark brown to blackish, partly bluish-white pruinose; sternum brownish with bluish-white pruinosity. Legs dark brown, coxae, trochanters, and tarsi paler, each femur basally with a whitish ring; coxae, trochanters, and femora slightly covered with bluish-white pruinosity; spines black. Wings Hyaline; venation dark brown, costa and radius black at base; pterostigma dark brown, with narrow pale margins, centrally a trace of bluish-white pruinosity, less than one cell long, rhomboidal; 15 / 15 postnodal crossveins in the left and 15 / 14 postnodal crossveins in the right forewing, 13 / 12 postnodal crossveins in hind wings. Abdomen Segment 1 dorsally dark brown to blackish, slightly covered with bluish-white pruinosity, brownish laterally; segment 2 dorsally blackish with greenish metallic shimmer, laterally pale to dark brown, centrally glaucous; secondary genital apparatus with terminal segment of ligula broadened towards its apical end, medially emarginated, basal third of terminal segment with longitudinal row of setae on each side (Fig. 2); segments 3 - 6 brown to dark brown, becoming paler laterally and turning yellowish ventrally, brown colouration darker at the segment borders, segments on the dorsum slightly metallic; segments 7 - 9 dark brown to blackish dorsally and laterally, ventrally yellow-brown; segment 10 dorsally brown, posterior margin blackish, elevated and dorsomedially excised, laterally and ventrally yellow-brown. Superior anal appendages brown, short, in dorsal view roughly resembling a paisley pattern (Fig. 4 C, F); in dorsocaudal view paisley-like shape more obvious, appendages broad, with convex, bulging upper margins and slightly concave, flattened lower margins, inner margins ending ventrally in bevelled edges, laterally each upper and lower margin is converging to a prominent tip directed posteriorly; in lateral view the appendages appear roughly triangular (Fig. 4 I, L). Inferior anal appendages yellow-brown, in dorsal view forceps-like, incurved, ending in a black tip which is directed inwards; about three times as long as the superior appendages (Fig. 4 C, F); in lateral aspect broad at base, narrowing to an acute tip (Fig. 4 I, L). Each inferior anal appendage bears basally on its inner side a strong, rounded process fringed with hairs (Fig. 4 C, F). Measurements Length of abdomen: c. 35 mm (not completely elongated), length of hind wings: 22.3 mm. VARIATION Colouration of mature males only with minor variations. Colour of face in two males pale greenish-blue, in four males light blue, and in three males dull aquamarine; the pruinose postocular spots are completely missing in four males. Yellow-brownish to olive-green colouration onprothoraxdorso-laterallysometimesmoreexpanded. Green metallic shimmer on dorsum of thorax in six specimens lacking; the lateral thoracic colouration varies between pale aquamarine and brownish-olivegreen; brown to black colouration on mesepimeron is differently developed and ranges from small, elongate, pale spots to broad, dark stripes, three males without any spots; extension of bluish-white pruinosity very variable, two males without any pruinescence. Segment 1 and 2 laterally often brighter pale aquamarine and less brown than in holotype. Segment 9 and 10 in three males dorsally and laterally with more extensive light brown to beige colouration. Superior and especially inferior anal appendages uniformly coloured. Three teneral males are straw-coloured with the exception of following parts: head dorsally green-black metallic, prothorax and synthorax dorsally brown to dark brown, abdominal segments 1 - 8 brown dorsally and at segment borders, segments 9 and 10 whitish dorsally. Males also differ slightly in the shape of pronotal structures. In two males the upper rim of pronotal hind lobe is curved with rounded apex, in four specimens the rim is more conspicuously curved forming a very acute tip medially. The median apex of the upper rim is connected with the lower rim by a more or less prominent keel in several males. There are also considerable differences in the extension of the inner process of inferior anal appendages. In six males the strong basal process on the inner side of each inferior anal appendage is only very small to small and more or less directed inwards. It is not clear at present if this is an artefact of preservation. The female of V. santoensis n. sp. could not be identified with certainty. Therefore we refrained from designing female paratypes. However, we tentatively assign four females to the new species, as they generally match the male colouration with following exceptions: aquamarine colouration of face paler or more glaucous, legs yellowish-brown, and apical two-thirds of segment 9, and segment 10 creamy or glaucous dorsally (light blue in life, see Fig. 3). One female has a slightly darker face and greenish eyes, and the pruinosity is more developed than in the other females. Pronotal hind lobe of females erected, rounded, not curved and not forming an apex medially; on the posterior rim of each mesostigmal lamina there is only a very inconspicuous protuberance. Cerci shorter than segment 10; ovipositor finely serrate at ventral border and slightly extending beyond segment 10. The paratype series varies in dimensional and venational characters as follows: abdomen length: c. 30 - 37.5 mm, hind wing length: 19.6 - 22.9 mm, 12 - 17 postnodal crossveins in forewings, 10 - 14 postnodal crossveins in hind wings.	en	Ober, Stefan V., Staniczek, Arnold H. (2009): A new genus and species of coenagrionid damselflies (Insecta, Odonata, Zygoptera, Coenagrionidae) from Vanuatu. Zoosystema 31 (3): 485-497, DOI: 10.5252/z2009n3a6, URL: http://www.bioone.org/doi/abs/10.5252/z2009n3a6
03B687F62B28FF89FF05C93A5D035AC0.taxon	discussion	REMARKS At first glance, males of V. santoensis n. sp. resemble males of V. malekulana n. comb., but a closer view reveals differences in the shapes of pronotal hind lobe, mesostigmal laminae, and superior anal appendages. In V. santoensis n. sp. the upper hind rim of the prothorax is more raised and only slightly curved, the lower ridge is not or only marginally protruding posteriorly. The protuberance of each mesostigmal hind ridge is less prominent and the superior anal appendages are paisley-shaped, contrary to the crescent-like form in V. malekulana n. comb. In lateral view, the flattened inner angle of the superior anal appendages is not visible in V. santoensis n. sp. Moreover, V. santoensis n. sp. is larger in size and the pterostigmata are dark brown versus light brown in V. malekulana n. comb. The shape of the cerci is also a significant difference to V. bidens n. comb., in which the superior anal appendages are ovate and bidentate.	en	Ober, Stefan V., Staniczek, Arnold H. (2009): A new genus and species of coenagrionid damselflies (Insecta, Odonata, Zygoptera, Coenagrionidae) from Vanuatu. Zoosystema 31 (3): 485-497, DOI: 10.5252/z2009n3a6, URL: http://www.bioone.org/doi/abs/10.5252/z2009n3a6
03B687F62B2FFF8BFD74CA5C58595C6B.taxon	materials_examined	HOLOTYPE. — Vanuatu. Tafea Province, Aneityum, 3 miles NE of Anelcauhat, Red Crest, 1200 ft, VI. 1955, leg. L. E. Cheesman, ♂ (BMNH). LABELS. — Holotype: Type; NEW HEBRIDES: / Aneityum. / Red Crest. 1,200 ft. / 3 m. N. E. of Anelgauhat. / vi vii 1955.; L. E. Cheesman. / B. M. 1955 - 217.; NesobasIs / ♂ bIdens KIm / D. E. Kimmins det. 1957 / TYPE.	en	Ober, Stefan V., Staniczek, Arnold H. (2009): A new genus and species of coenagrionid damselflies (Insecta, Odonata, Zygoptera, Coenagrionidae) from Vanuatu. Zoosystema 31 (3): 485-497, DOI: 10.5252/z2009n3a6, URL: http://www.bioone.org/doi/abs/10.5252/z2009n3a6
03B687F62B2FFF8BFD74CA5C58595C6B.taxon	description	ADDITIONAL MATERIAL. — Not available.	en	Ober, Stefan V., Staniczek, Arnold H. (2009): A new genus and species of coenagrionid damselflies (Insecta, Odonata, Zygoptera, Coenagrionidae) from Vanuatu. Zoosystema 31 (3): 485-497, DOI: 10.5252/z2009n3a6, URL: http://www.bioone.org/doi/abs/10.5252/z2009n3a6
03B687F62B2FFF8BFD74CA5C58595C6B.taxon	distribution	DISTRIBUTION. — The species is only known from the type locality.	en	Ober, Stefan V., Staniczek, Arnold H. (2009): A new genus and species of coenagrionid damselflies (Insecta, Odonata, Zygoptera, Coenagrionidae) from Vanuatu. Zoosystema 31 (3): 485-497, DOI: 10.5252/z2009n3a6, URL: http://www.bioone.org/doi/abs/10.5252/z2009n3a6
03B687F62B2FFF8BFD74CA5C58595C6B.taxon	description	REDESCRIPTION OF THE HOLOTYPE Preserved dry, pinned. In good condition, but right foreleg and left hind leg missing. Right forewing and abdomen between segments 6, 7 and 8 glued. Head Labium pale fulvous; labrum dark fulvous, with a small, central dark brown spot at the basal margin; genae, anteclypeus, and postclypeus dark fulvous, postclypeus with a denoted dark brown T-mark; frons brown; antennae brown, inner side of each scapus blackish; vertex and occiput black except for a small brown spot in front of the median ocellus; rear of head brownish; eyes dark brown. Thorax Prothorax dorsally bright black, laterally fulvous; hind lobe of pronotum rounded, prominent, with distinct, fulvous median keel connecting upper and inconspicuous lower pronotal hind ridge. Synthorax: mesanepisternum with shiny black median band reaching the mesopleural suture in its hindmost fourth, but elsewhere separated from it by a narrow, pale fulvous band, dorsal carina brown; mesostigmal lamina slightly triangular, anterior and posterior ridge convergent and ending in a protuberance, posterior rim with small median tubercle; mesokatepisternum fulvous with dark brown spot in its upper half; mesepimeron fulvous; metanepisternum fulvous, posteriorly with two small, elongate, brown spots at the intersegmental and metapleural suture; metakatepisternum fulvous; metepimeron fulvous becoming yellowish posteriorly; poststernum yellowish, posterior margin black; sternum yellowish. Legs yellowish, basal inner surface of protibia brownish; spines dark brown. Wings Hyaline; venation dark brown; pterostigma brownish with narrow pale margin, less than one cell long, rhomboidal; 15 / 14 postnodal crossveins in forewings and 13 / 12 postnodal crossveins in hind wings. Abdomen Segments 1 and 2 yellowish, dorsally brown to dark brown; segments 3 - 6 yellowish, dorsally light brown to brown, dorsal brown colouration extending downwards at the posterior part of each segment but missing anteriorly, there the yellowish colouration reaches the dorsal carina in a spot of almost triangular shape; on segment 7 the yellowish colouration turns creamy and the brown colour gets darker and extends posteriorly to form a dark triangle as seen in lateral view, brown of segments 1 - 6 with a slight greenish metallic shimmer getting bronze on segment 7; segment 8 with the exception of a narrow creamy line on the lower lateral border dark brown to blackish with a metallic sheen; the colouration of segment 9 is divided in a ratio of 5: 3, being bluish-white in the upper, and brown in the lower part; segment 10 almost entirely bluish-white, only ventrally a bit creamy. Superior anal appendages short, in dorsal view broad and ovate, upper surface slightly concave, dark brown, lateral part paler, the apical margins with two small, acute, shiny black teeth, one at the inner angle, the other just in the centre (Fig. 4 A, D); in lateral view the inner tooth appears as an elevated knob; lateral part of superior anal appendages bulged (Fig. 4 G, J); in posterior view the appendages are straightened diagonally, forming a rounded roof. Inferior anal appendages in dorsal view forceps-like, but only slightly incurved, ending in black tips which are directed inwards, about two and a half times as long as the superior appendages (Fig. 4 A, D), yellowish-white at base, forceps brown to dark brown; in lateral aspect broad at base, abruptly narrowed to an acute apex (Fig. 4 G, J). In posterior view a small protuberance is visible basally on the inner side of each appendage. Measurements Length of abdomen: c. 35 mm (not completely elongated), length of left hind wing: 22.1 mm, length of right hind wing: 21.8 mm. VARIATION The species is only known from the holotype.	en	Ober, Stefan V., Staniczek, Arnold H. (2009): A new genus and species of coenagrionid damselflies (Insecta, Odonata, Zygoptera, Coenagrionidae) from Vanuatu. Zoosystema 31 (3): 485-497, DOI: 10.5252/z2009n3a6, URL: http://www.bioone.org/doi/abs/10.5252/z2009n3a6
03B687F62B2FFF8BFD74CA5C58595C6B.taxon	discussion	REMARKS Due to the colouring it is likely that the specimen is immature. A comparison with the original description (Kimmins 1958) reveals that the colouration has lost its intensity over the last 53 years. According to Kimmins (1958) it could be quite possible that the yellowish-fuscous areas become bluish or greenish during the maturation process. Males of V. bidens n. comb. can be easily distinguished from males of the other two species of the genus by the bidentate apical margins of the superior anal appendages. These appendages appear in the dried specimen in a very upright position, in life, they may be in a more oblique position. The length of the abdomen was stated by Kimmins (1958) as 25 mm, this seems to be a literal error.	en	Ober, Stefan V., Staniczek, Arnold H. (2009): A new genus and species of coenagrionid damselflies (Insecta, Odonata, Zygoptera, Coenagrionidae) from Vanuatu. Zoosystema 31 (3): 485-497, DOI: 10.5252/z2009n3a6, URL: http://www.bioone.org/doi/abs/10.5252/z2009n3a6
03B687F62B2DFF86FF2FCFE65A0D5E54.taxon	materials_examined	HOLOTYPE. — Vanuatu. Malampa Province, Malekula, Unua, III-IV. 1929, leg. L. E. Cheesman, ♂ (BMNH). PARATYPES. — Vanuatu. Malampa Province, Malekula, Unua, II. 1929, leg. L. E. Cheesman, 5 ♂♂ (BMNH).	en	Ober, Stefan V., Staniczek, Arnold H. (2009): A new genus and species of coenagrionid damselflies (Insecta, Odonata, Zygoptera, Coenagrionidae) from Vanuatu. Zoosystema 31 (3): 485-497, DOI: 10.5252/z2009n3a6, URL: http://www.bioone.org/doi/abs/10.5252/z2009n3a6
03B687F62B2DFF86FF2FCFE65A0D5E54.taxon	description	LABELS. — Holotype: Holo- / type; New Hebrides: / Malekula, / Ounua. / Mar. & Apl. 1929. / Miss L. E. Cheesman. / B. M. 1929 - 343.; 331.; NESOBASIS / malekulana / ♂ sp. n. / Holotype / det. D. E. Kimmins. Paratypes: Para- / type; New Hebrides: / Malekula, / Ounua. / Feb. 1929. / Miss L. E. Cheesman. / B. M. 1929 - 234.; 231.; NESOBASIS / malekulana / ♂ sp. n. / det. D. E. Kimmins. ADDITIONAL MATERIAL. — Not available.	en	Ober, Stefan V., Staniczek, Arnold H. (2009): A new genus and species of coenagrionid damselflies (Insecta, Odonata, Zygoptera, Coenagrionidae) from Vanuatu. Zoosystema 31 (3): 485-497, DOI: 10.5252/z2009n3a6, URL: http://www.bioone.org/doi/abs/10.5252/z2009n3a6
03B687F62B2DFF86FF2FCFE65A0D5E54.taxon	distribution	DISTRIBUTION. — The species is only known from the type locality.	en	Ober, Stefan V., Staniczek, Arnold H. (2009): A new genus and species of coenagrionid damselflies (Insecta, Odonata, Zygoptera, Coenagrionidae) from Vanuatu. Zoosystema 31 (3): 485-497, DOI: 10.5252/z2009n3a6, URL: http://www.bioone.org/doi/abs/10.5252/z2009n3a6
03B687F62B2DFF86FF2FCFE65A0D5E54.taxon	description	REDESCRIPTION OF THE HOLOTYPE Preserved dry, pinned. In fairly good condition. Head, prothorax, left foreleg, right hind wing, and abdomen between segments 5, 6 and 7 glued. Head Labium pale fulvous; labrum dark blue-green, centrally more glaucous, with a semicircular, central dark brown spot at the basal margin; genae glaucous; anteclypeus greyish-brown; postclypeus blackish, laterally at the basal margin glaucous; frons below a semicircular line from the basal joints of the antennae to the postclypeus glaucous, above blackish; antennae at the basal joints glaucous, each scapus dark brown, anteriorly with a small glaucous stripe, pedicelli and flagella brown; vertex and occiput black except for a small brown spot in front of the median ocellus, diminutive postocular spots, composed of whitish pruinosity; back of head yellowish with a diagonal dark brown stripe; eyes dark brown. Thorax Prothorax black with a touch of a greenish metallic shimmer, laterally with slight bluish-white pruinosity; hind lobe of pronotum slightly raised, each side of upper rim sinusoidally shaped, medially forming a slightly rounded apex. The apex is connected with the prominent, lower ridge by a median keel (Fig. 5 A). Synthorax: mesanepisternum with black median band, slightly greenish metallic, reaching the mesopleural suture in its hindmost fifth, but elsewhere separated from it by a brownish band, dorsal carina black; mesostigmal lamina slightly triangular, posterior margin proximally with a distinctive, rounded, slightly quadrangular protuberance, apex of lamina forming a slightly overhanging process (Fig. 5 A); mesokatepisternum brownish to dark brown in the upper, pale greenish-grey in the lower half; mesepimeron pale greenish-grey with a dark brown to blackish triangular streak in the anterior part; metanepisternum pale greenish-grey, posteriorly with a small, elongate, dark brown to blackish spot at the metapleural suture; metakatepisternum light ochre with slight bluish-white pruinosity; metepimeron pale greenish-grey becoming creamy anteriorly; poststernum bluish-white pruinose, posterior margin black; sternum creamy to light brown with bluish-white pruinosity. Legs creamy, except outer surface of coxae, trochanteres, and femora, and inner median parts of tibiae, which are light to dark brown, especially the coxae with bluish-white pruinosity; spines dark brown. Wings Hyaline; venation brown; pterostigma pale brown, anteriorly with small pale margin, less than one cell long, rhomboidal; 13 / 12 postnodal crossveins in the left and 15 / 14 postnodal crossveins in the right forewing, 11 / 10 postnodal crossveins in the A left and 12 / 11 postnodal crossveins in the right hind wing. Abdomen Segment 1 dorsally dark brown with bluish-white pruinosity, creamy and greenish-grey laterally; segment 2 dorsally blackish with slight greenish metallic shimmer, laterally greenish-grey and partly creamy; segments 3 - 6 brown to dark brown, getting paler laterally and pale ochre ventrally, brown colouration darker at the segment borders; segments 7 - 9 dark brown dorsally and at sides, ventrally reddish brown; segment 9 dorsally with a reddish-brown, centrally creamy bar, incurved anteriorly; segment 10 reddish-brown, dark brown dorsally, apical margin blackish, raised and excised at its centre. Superior anal appendages reddish-brown, short, in dorsal view crescent-shaped (Fig. 4 B, E); in dorsocaudal view broad with convex upper margins and slightly concave lower margins, both laterally and medially converging to strong, posteriorly projected, rounded tips; in lateral view each tip appears triangular, the inner tip being higher and more protruded than the outer one (Fig. 4 H, K). Inferior anal appendages reddish-brown, in dorsal view forceps-like, incurved, ending in a black tip which is directed inwards, about three times as long as the superior appendages (Fig. 4 B, E); in lateral aspect broad at base, narrowing to an acute tip (Fig. 4 H, K). Each inferior anal appendage bears basally on its inner side a strong, quadrate process fringed with hairs (Fig. 4 B, E). Measurements Length of abdomen: c. 33 mm (not completely elongated), length of hind wings: 20.3 mm. VARIATION Only two of the five designated paratypes were available for study. One paratype nearly resembles the holotype in colouration, but face dull aquamarine and thorax somewhat paler with a smaller and less intensive brown spot on mesepimeron. The pruinosity is a little more intensive posteriorly. Abdominal segments 1 and 2 laterally glaucous, segments 9 and 10 dorsally creamy with reduced brown markings. Pronotal hind lobe less raised than in holotype and in other paratype. The other, somewhat smaller paratype is more intensively coloured: face aquamarine, thorax bluish-green with dark brown to blackish stripes on mesepimeron and metepimeron; mesokatepisternum and metakatepisternum dark brown. Brown colouration on abdominal segments 1 and 2 generally more developed, bluish-green colouration reduced, but brighter than in the other two specimens at hand. Only a small pale brown spot dorsally on segment 9. Venation in both paratypes only slightly different from holotype: 12 - 15 postnodal crossveins in forewings, 11 - 13 postnodal crossveins in hind wings.	en	Ober, Stefan V., Staniczek, Arnold H. (2009): A new genus and species of coenagrionid damselflies (Insecta, Odonata, Zygoptera, Coenagrionidae) from Vanuatu. Zoosystema 31 (3): 485-497, DOI: 10.5252/z2009n3a6, URL: http://www.bioone.org/doi/abs/10.5252/z2009n3a6
03B687F62B2DFF86FF2FCFE65A0D5E54.taxon	discussion	REMARKS This species is only known from six males found at the type locality. All paratypes have identical label data (see above) (D. Goodger pers. comm.). Vanuatubasis malekulana n. comb. differs from V. bidens n. comb. in the wavy, less erected pronotal hind rigde and the strong, medio-posterior protuberance of the mesostigmal laminae. The shape of the superior anal appendages is crescent-like, the forcipate inferior anal appendages are more incurved than in V. bidens n. comb. For differences to the closely related V. santoensis n. sp. see above.	en	Ober, Stefan V., Staniczek, Arnold H. (2009): A new genus and species of coenagrionid damselflies (Insecta, Odonata, Zygoptera, Coenagrionidae) from Vanuatu. Zoosystema 31 (3): 485-497, DOI: 10.5252/z2009n3a6, URL: http://www.bioone.org/doi/abs/10.5252/z2009n3a6
