taxonID	type	description	language	source
03B7AE4BFFCFBC46FF1AFE4F62E5F9E1.taxon	description	In the redescription of his misidentified S. westwoodi Harold, Péringuey (1908) mentions that he saw one specimen amongst some S. galenus Westwood from the Z [S] outpansberg. This could only be a specimen of Scarabaeus geminogalenus sp. nov. This provides strong support for the misidentification as S. westwoodi Harold shows a highland distribution along the central and northeast Drakensberg escarpment. Subsequently, Janssens, 1940 again, synonymized Peringuey ’ s misidentified S. westwoodi (= Scarabaeus geminogalenus sp. nov.) with S. galenus Westwood [cited as a synonym under S. galenus: ‘ Sebasteos Westwoodi PERINGUEY (non HAROLD), Trans. s. Afr. Phil. Soc., XIII, 1908, p. 555 (nov. syn.) ’], an erroneous decision that was followed by all subsequent authors. Although we have been unable to locate type material for the misidentified S. westwoodi Harold, the description in Péringuey (1908) corresponds very well with that for a species that occurs on coastal sands of northeast South Africa, southeast Mozambique and along the Limpopo river valley. This new species is not a synonym of S. galenus Westwood but a very close relative that would require a new name (Scarabaeus geminogalenus sp. nov.) as S. westwoodi is unavailable.	en	Deschodt, Christian M., Davis, Adrian L. V., Scholtz, Clarke H. (2016): Three new species, a synonymy and taxonomic notes on southern African members of the dung beetle genera Scarabaeus s. str. Linnaeus, 1758 and Escarabaeus Zídek & Pokorný, 2011 (Scarabaeidae: Scarabaeinae). Journal of Natural History 51 (5 - 6): 299-316, DOI: 10.1080/00222933.2016.1261949, URL: http://dx.doi.org/10.1080/00222933.2016.1261949
03B7AE4BFFC8BC49FEF9FF1863B6FBD0.taxon	description	(Figures 1, 2, 4 and 5)	en	Deschodt, Christian M., Davis, Adrian L. V., Scholtz, Clarke H. (2016): Three new species, a synonymy and taxonomic notes on southern African members of the dung beetle genera Scarabaeus s. str. Linnaeus, 1758 and Escarabaeus Zídek & Pokorný, 2011 (Scarabaeidae: Scarabaeinae). Journal of Natural History 51 (5 - 6): 299-316, DOI: 10.1080/00222933.2016.1261949, URL: http://dx.doi.org/10.1080/00222933.2016.1261949
03B7AE4BFFC8BC49FEF9FF1863B6FBD0.taxon	description	Paratypes one spec, S. Afr; Kruger Nat Pk, Punda Maria, 22.41 S – 31.01 E, 3.2.1994; E – Y: 2977, elephant dung, leg. Endrödy-Younga (TMSA), two spec, S. Afr; KrugerNat. Pk, Malonga sands, 22.38 S – 31.17 E, 8.2.1994: E – Y: 2986, on red sand, Endrödy- Younga (TMSA), one spec, Masemola ♂ – 6: 03., 4593 (TMSA), one spec, S. Afr; KrugerNat. Pk, Pumbe sands, 24.12 S – 31.55 E, 22.11.1994: E – Y: 3063, groundtraps. 60 days, Endrödy. Bellamy, groundtrap with faece bait (TMSA), five spec, ST. LUCIA ESTUARY, Natal. 26. VIII. 78, G. F. Bornemissza, two with COLS 22352, one with COLS 22346 one with COLS 22345 and one with COLS 22348 (SANC), five spec, ST. LUCIA ESTUARY, Natal. 19. V. 78, G. F. Bornemissza, three with COLS 22343 and two with COLS 22344 (SANC), four spec, SOUTH AFRICA: KZN, Tembe Elephant Park, 27 ° 01 ‘ S 32 ° 24 ʹ E, 24 – 27. i., 1996, BJ van Rensburg, pitfall trap: elephant dung, COLS 28131 (SANC), one spec, SOUTH AFRICA, NTL, St Lucia Nat. Park, 28. [°] 22 [’] S 32. [°] 25 [‘] E, G Bernon, COLS 23082 (SANC), two spec, SOUTH AFRICA: LIMP, Greater Kuduland, Safaris, E of Tshipise, 22 ° 38 ʹ 28 “ S 30 ° 18 ʹ 50 ” E, 08 – 16. iv. 2007 SARCA, COLS 23080 (SANC), one spec, SOUTH AFRICA, Tvl, D ’ Nyala Nat. Res., Ellisras District, 23. [°] 45 [’] S 27. [°] 49 [‘] E, 10 – 14. xi. 1986, V. M. Uys, COLS 23081 (SANC), one spec, SOUTH AFRICA, Tvl, D ’ Nyala Nat. Res., Ellisras District, 23. [°] 45 [’] S 27. [°] 49 [‘] E, 10 – 14. xi. 1986, B. Grobbelaar, COLS 23081 (SANC), one spec, NDUMU G. R., Natal, 25. X. 73, Tribe & Temby, COLS 00177 (SANC), PAFURI, KNP, (24 kms S – Maohai Pan), 25. III. 73, I. D. Temby, COLS 22349 (SANC), FOMOTHINI PAN, N. Natal 8. X. 80, E. Klingelhoefer, COLS 22350 (SANC), one spec, SOUTH AFRICA, NATAL, Cape Vidal, 28. [°] 10 [’] S, 32. [°] 32 [‘] E. 13. i. 1981, I. M. Millar, COLS 23012 (SANC), one spec, HLUHLUWE, Natal, (15 km S) 31. X. 78, Bornemissza & Aschenborn, COLS 22347 (SANC), two spec (one spec gold coated for SEM imaging), Tembe Elephant park, Kwazulu-Natal, 27 ° 01 ’ S 32 ° 24 ʹ E, 24 – 29. I. 1996, B. J. Van Rensburg (UPSA), one spec, Tembe Elephant park, KZN, South Africa, 27 ° 01 ‘ S 32 ° 24 ʹ E, 26 – 27. VII. 1995, B. J. Van Rensburg (UPSA), one spec, Tembe Elephant park, KZN, South Africa, 27 ° 01 ’ S 32 ° 24 ʹ E, 17. iii. 1995, B. J. Van Rensburg (UPSA), three spec, Soutpansberg, Limpopo, S 22.9157 ° E 29.68756 °, 3 – 4. ii. 2009, L. J. Niemand (UPSA).	en	Deschodt, Christian M., Davis, Adrian L. V., Scholtz, Clarke H. (2016): Three new species, a synonymy and taxonomic notes on southern African members of the dung beetle genera Scarabaeus s. str. Linnaeus, 1758 and Escarabaeus Zídek & Pokorný, 2011 (Scarabaeidae: Scarabaeinae). Journal of Natural History 51 (5 - 6): 299-316, DOI: 10.1080/00222933.2016.1261949, URL: http://dx.doi.org/10.1080/00222933.2016.1261949
03B7AE4BFFC8BC49FEF9FF1863B6FBD0.taxon	diagnosis	Diagnosis. This species is similar to S. galenus, but can easily be separated by the much denser punctation on the pronotum.	en	Deschodt, Christian M., Davis, Adrian L. V., Scholtz, Clarke H. (2016): Three new species, a synonymy and taxonomic notes on southern African members of the dung beetle genera Scarabaeus s. str. Linnaeus, 1758 and Escarabaeus Zídek & Pokorný, 2011 (Scarabaeidae: Scarabaeinae). Journal of Natural History 51 (5 - 6): 299-316, DOI: 10.1080/00222933.2016.1261949, URL: http://dx.doi.org/10.1080/00222933.2016.1261949
03B7AE4BFFC8BC49FEF9FF1863B6FBD0.taxon	description	Description: Large beetle, completely black. Body size of holotype: Head capsule length 9.3 mm, combined length of pronotum and elytra over midline 23.5 mm, widest width over elytra 16.1 mm, height 11.8 mm. Head: Surface completely rugose, without granulation. No projection on dorsal surface of clypeus. Clypeal teeth slightly upturned, with anterior fourth of teeth smooth. Notch between two median clypeal teeth rounded; notch between first and second obtuse angled. Median two teeth protruding forward, with a downward pointing spine on each; a third downward pointing spine just under notch of median teeth present. Vertex slightly convex. Antennae dark brown to black. Pronotum: Punctate, with large, sometimes irregular, punctures almost contiguous. Surface between punctures very finely shagreened. Thin medial line present, withoutpunctures and not always traversing pronotum. Sometimes small lateral clear area at anterior half adjacent to medial line present. Lateral margins punctate, with punctures at anterolateral angle elongated and arranged in subparallel downward pointing lines. Anterior margin smooth ending with sharp triangular protrusion. Protibia: Outside margin with four teeth in distal two thirds, proximal tooth smallest. Margin between teeth serrate and somewhat crenulate in first third. Inside margin serrate, with dark long setae that are evenly spaced. Denticle on inside margin opposite third protibial tooth pointing upward at almost 90 degrees, and fourth protibial tooth pointing forward at about 45 degrees. Elytra: Slightly ribbed. Interstriae shagreened, with small widespread punctures. Striae clearly discernible with the naked eye, with small, regular punctures visible only from 20 x magnification. Sterna: Clear median longitudinal groove over meta- and mesosternum, forming deep foveae on meta- and mesosternum. Meta- and mesosternum with setigerous punctures. Mesosternum sloping at about 45 degrees anteriorly. Pygidium: Wrinkled with small irregular punctures far apart. Aedeagus: See Figure 2 (C) Variability: There is very little morphological variation between the specimens except for the variable size (smallest and largest specimens in type series: head capsule length 8.4 – 12.0 mm, combined length of pronotum and elytra over midline 20.3 – 28.3 mm, widest width over elytra 14.1 – 20.0 mm, height 9.8 – 13.6 mm.).	en	Deschodt, Christian M., Davis, Adrian L. V., Scholtz, Clarke H. (2016): Three new species, a synonymy and taxonomic notes on southern African members of the dung beetle genera Scarabaeus s. str. Linnaeus, 1758 and Escarabaeus Zídek & Pokorný, 2011 (Scarabaeidae: Scarabaeinae). Journal of Natural History 51 (5 - 6): 299-316, DOI: 10.1080/00222933.2016.1261949, URL: http://dx.doi.org/10.1080/00222933.2016.1261949
03B7AE4BFFC8BC49FEF9FF1863B6FBD0.taxon	etymology	Etymology: The species ’ long association and incorporation with S. galenus is echoed by renaming it as the twin of S. galenus	en	Deschodt, Christian M., Davis, Adrian L. V., Scholtz, Clarke H. (2016): Three new species, a synonymy and taxonomic notes on southern African members of the dung beetle genera Scarabaeus s. str. Linnaeus, 1758 and Escarabaeus Zídek & Pokorný, 2011 (Scarabaeidae: Scarabaeinae). Journal of Natural History 51 (5 - 6): 299-316, DOI: 10.1080/00222933.2016.1261949, URL: http://dx.doi.org/10.1080/00222933.2016.1261949
03B7AE4BFFC8BC49FEF9FF1863B6FBD0.taxon	distribution	Distribution: This species occurs on the coastal sand plains of eastern South Africa and southern Mozambique, extending inland along the Limpopo River valley.	en	Deschodt, Christian M., Davis, Adrian L. V., Scholtz, Clarke H. (2016): Three new species, a synonymy and taxonomic notes on southern African members of the dung beetle genera Scarabaeus s. str. Linnaeus, 1758 and Escarabaeus Zídek & Pokorný, 2011 (Scarabaeidae: Scarabaeinae). Journal of Natural History 51 (5 - 6): 299-316, DOI: 10.1080/00222933.2016.1261949, URL: http://dx.doi.org/10.1080/00222933.2016.1261949
03B7AE4BFFC8BC49FEF9FF1863B6FBD0.taxon	materials_examined	Type specimens of Scarabaeus galenus Westwood, 1847, Scarabaeus westwoodi (Von Harold, 1869) and Scarabaeus vicinus Janssens, 1940 Figures 1, 2, 5 and 6 Zídek and Pokorný (2004); listed the holotype specimen of Scarabaeus galenus Westwood, 1847; as being deposited in the Natural History Museum, London, but it could not be located after an extensive search by Mr Malcolm Kerley, recently retired curator in the Coleoptera Section. Mr Darren Mann, Head of Life Collections at the Oxford University Museum of Natural History, also unsuccessfully searched the Hope Collection. The holotype could not be traced and is believed to be lost. The type of Scarabaeus westwoodi Von Harold (1869); is listed (Zídek and Pokorný 2004) as being in the Muséum National d ’ Histoire Naturelle, Paris. Janssens (1940, p. 46) reported to have examined this type, and noted that it is in the Oberthür Collection in the MNHN. It is, however, also presumed lost since it could not be found after an extensive search by the current curator, Dr Antoine Mantilleri. Article 75.2 of the ICZN (1999) states that ‘ A neotype is not to be designated as an end in itself, or as a matter of curatorial routine, and any such neotype designation is invalid ’. Both these species are well known and without any taxonomic or nomenclatural complications other than those resolved in the present paper. Hence we do not designate neotypes for the two abovementioned species. During preparation for this paper, specimens believed to be S. galenus were placed next to each other and subtle differences were observed between individuals from the drier Kalahari and the moister savannah areas of southern Africa. Characterization of the Kalahari individuals corresponded well with the description of Scarabaeus vicinus Janssens, 1940. However, two extensive searches in the MNHN by Dr Antoine Mantilleri also failed to locate the type specimen. Although we could not compare our specimens with type material, we are confident that we have rediscovered a species for which there have been no records since its description in 1940. There is a history of previous misidentification since specimens of S. vicinus in the SANC and TMSA were found among those determined as S. galenus by zur Strassen in 1993. Some published data also appear to reflect misidentifications, e. g. from Burghersdorp (sic) [S 30.996 ° E 26.328 °], Kimberley [S 28.728 ° E 24.749 °], Prieska [S 29.670 ° E 22.745 °] (Péringuey 1901), Twee Rivieren [S 20.611 ° E 26.470 °] (Ferreira 1959), from the Waterberg plateau [S 20.652 ° E 22.146 °] (Delatour 2008) and in extensive survey data from the Northern Cape (Davis et al. 2008). Scarabaeus vicinus can be easily distinguished from the morphologically similar S. galenus by the more velutinous appearance of the elytra and pronotum compared to the more shiny S. galenus. The fore tibiae of S. vicinus are also more inwardly curved than those of S. galenus. See Figure 4 for SEM images of the elytra and pronota of both species. Locality records show that S. galenus is widespread in the drier savannah of south-eastern Africa with a range extending north-eastwards to southern Tanzania whereas the newly rediscovered, close relative, S. vicinus is Kalahari-centred in the drier, sandy areas of south western Africa (Figure 2). Again with reference to Article 75.2 (ICZN 1999), we refrain from defining a neotype for S. vicinus because it is relatively easy to identify and the original type specimen might still be found. Type specimen of Scarabaeus rixosus Péringuey 1901 As the type specimen of Scarabaeus rixosus Péringuey, 1901: 50, is not listed in the holdings of the Iziko South African Museum where the majority of Péringuey ’ s type specimens are housed, it is likely to have been deposited in his personal collection (R. Stals pers. comm.). Péringuey ’ s private collection was acquired by the UPSA and subsequently donated to the SANC (Cochrane 2006). We propose that a specimen recently found amongst undetermined Scarabaeinae material in the SANC is the type specimen of S. rixosus (Figure 3 A – C) as it conforms to the original description. It also differs from all other known Scarabaeus species, including Scarabaeu s zambesianus Péringuey, 1901, and its putative closest relative Scarabaeu s proboscideus (Guérin- Méneville 1844) (Figure 3 (D )), which is the only other Scarabaeus species known to bear a spur on the mentum. Although the locality label on the re-discovered specimen (‘ Upper Limpopo ’) is in Péringuey ’ s handwriting (R. Stals pers. comm.), it differs from the published type locality for S. rixosus (‘ Southern Rhodesia (Middle Limpopo) ’) However, it is known that Péringuey seldom published labels verbatim (R. Stals pers. comm.) Thus, we propose that the rediscovered specimen is the holotype of Scarabaeus rixosus Péringuey, 1901. It is curated in the SANC, with accession numbers COLS 23260 and TYPH 02071. No other specimens of this species are known. Descriptions of new species Scarabaeus karae Davis and Deschodt sp. nov. Figures 1, 2 and 6 Specimens examined Holotype: ♂ Abe Bailey NR, Gauteng, S 26 ° 17 ʹ - 21 ʹ E 027 ° 16 ʹ - 18 ʹ, 1496 m- 1544 m, 09 – 10. xii. 2002. A Davis. (TMSA). Paratypes: three spec same as holotype, (one SANC, one TMSA, one UPSA). Figure 5. Map showing the known distribution of Scarabaeus galenus Westwood () (the type localities of Ateuchus (Actinophorus) paradoxus Boheman, and Sebasteos adelphus Kolbe, are not shown here), Scarabaeus vicinus Janssens (▲) and Scarabaeus geminogalenus Davis and Deschodt (▪). Note that the record for Scarabaeus galenus in Namibia and the two easterly records for Scarabaeus vicinus are seen as errors.	en	Deschodt, Christian M., Davis, Adrian L. V., Scholtz, Clarke H. (2016): Three new species, a synonymy and taxonomic notes on southern African members of the dung beetle genera Scarabaeus s. str. Linnaeus, 1758 and Escarabaeus Zídek & Pokorný, 2011 (Scarabaeidae: Scarabaeinae). Journal of Natural History 51 (5 - 6): 299-316, DOI: 10.1080/00222933.2016.1261949, URL: http://dx.doi.org/10.1080/00222933.2016.1261949
03B7AE4BFFC8BC49FEF9FF1863B6FBD0.taxon	etymology	Etymology: This species is named in honour of Kara, the only daughter of CD.	en	Deschodt, Christian M., Davis, Adrian L. V., Scholtz, Clarke H. (2016): Three new species, a synonymy and taxonomic notes on southern African members of the dung beetle genera Scarabaeus s. str. Linnaeus, 1758 and Escarabaeus Zídek & Pokorný, 2011 (Scarabaeidae: Scarabaeinae). Journal of Natural History 51 (5 - 6): 299-316, DOI: 10.1080/00222933.2016.1261949, URL: http://dx.doi.org/10.1080/00222933.2016.1261949
03B7AE4BFFC8BC49FEF9FF1863B6FBD0.taxon	description	Description: Medium sized beetle, completely black except for tan brown coloured setae on ventral side. Body size of holotype: Head capsule length 4.0 mm, combined length of pronotum and elytra over midline 11.9 mm, widest width over elytra 8.1 mm, height 5.6 mm. Head: Frons punctate, clypeus rugose. No projection on dorsal or ventral surface of clypeus. Clypeal teeth slightly upturned, with anterior apices of teeth smooth. Notch between median clypeal teeth and between first and second clypeal teeth right angled. Vertex slightly convex. Antennae black. Pronotum: Punctate with irregularly spaced punctures that are sometimes far apart and sometimes contiguous. Surfaces between punctures smooth. Row of irregular punctures on posterior margin. Lateral margins crenulate. Protibia: Inside margin smooth, with dark setae of irregular length and uneven spacing. No tooth at distal inside end. Outside margin with four teeth in distal four fifths, proximal tooth smallest. Serrate between teeth and crenulate in first fifth. Elytra: Ribbed. Shagreened on interstriae with irregularly spaced big punctures that are far apart. Punctures sometimes touch outside margin of interstriae. Striae acutely edged and distinct to the naked eye with small punctures close together giving a stippled appearance. Sterna: Shallow and indistinct median longitudinal groove over meta- and mesosternum, area next to groove smooth. Otherwise meso and metasternum with long thin tan brown setae and small and close together punctures. Mesosternum strongly inclined at about 90 degrees anteriorly. Pygidium: Smooth with very few large irregular punctures. Aedeagus: See Figure 2 (D). Variability: Except for the variable size (smallest and largest specimens in type series: head capsule length 3.5 – 5.1 mm, combined length of pronotum and elytra over midline 10.5 – 13.1 mm, widest width over elytra 7.0 – 9.0 mm, height 5.0 – 6.1 mm.), there is very little variation between the specimens.	en	Deschodt, Christian M., Davis, Adrian L. V., Scholtz, Clarke H. (2016): Three new species, a synonymy and taxonomic notes on southern African members of the dung beetle genera Scarabaeus s. str. Linnaeus, 1758 and Escarabaeus Zídek & Pokorný, 2011 (Scarabaeidae: Scarabaeinae). Journal of Natural History 51 (5 - 6): 299-316, DOI: 10.1080/00222933.2016.1261949, URL: http://dx.doi.org/10.1080/00222933.2016.1261949
03B7AE4BFFC8BC49FEF9FF1863B6FBD0.taxon	diagnosis	Diagnosis: This new species keys out to Scarabaeus funebris in Zur Strassen (1967), but can easily be separated from this allied species, which sometimes overlaps in distribution and habitat, by having larger punctures on the pronotum and clear and round punctures on the elytra. The parameres of the aedeagus are basally slightly more concave for S. karae than for S. funebris. The serrations on the inside front tibiae are reduced to small round scallops to almost smooth as opposed to S. funebris. It is also generally smaller than S. funebris. We have seen images of the type specimen of Scarabaeus funebris pretoriensis Janssens, 1940 (IRSNB) and compared it with images of the type specimen of Scarabaeus funebris Boheman, 1857; in the NHRS and confirm the synonymy of S. funebris pretoriensis with S. funebris as proposed by Zur Strassen, 1967 even though the pronotal punctation is less pronounced in the subspecies.	en	Deschodt, Christian M., Davis, Adrian L. V., Scholtz, Clarke H. (2016): Three new species, a synonymy and taxonomic notes on southern African members of the dung beetle genera Scarabaeus s. str. Linnaeus, 1758 and Escarabaeus Zídek & Pokorný, 2011 (Scarabaeidae: Scarabaeinae). Journal of Natural History 51 (5 - 6): 299-316, DOI: 10.1080/00222933.2016.1261949, URL: http://dx.doi.org/10.1080/00222933.2016.1261949
03B7AE4BFFC8BC49FEF9FF1863B6FBD0.taxon	distribution	Distribution: This species is probably restricted to the high grassland of central South Africa.	en	Deschodt, Christian M., Davis, Adrian L. V., Scholtz, Clarke H. (2016): Three new species, a synonymy and taxonomic notes on southern African members of the dung beetle genera Scarabaeus s. str. Linnaeus, 1758 and Escarabaeus Zídek & Pokorný, 2011 (Scarabaeidae: Scarabaeinae). Journal of Natural History 51 (5 - 6): 299-316, DOI: 10.1080/00222933.2016.1261949, URL: http://dx.doi.org/10.1080/00222933.2016.1261949
03B7AE4BFFC0BC55FEC1FB19601EF925.taxon	description	(Figure 1, 2 and 7) Specimens examined Holotype: ♂ Botswana, Okavango, Thamalakane riv., P. Reavell, XII. 1973. (TMSA) Paratypes: two ♀ Tau Lodge Botswana., S 20.42679 ° E 24.52141 °, 20. xii. 2003 Light, C. Deschodt & L. Niemand. (UPSA), one ♂ spec and ten spec unsexed same as holotype, (TMSA), one ♂ spec and one spec unsexed, Namibia; Rundu Distr., 17 km E Rundu 1059 m, 17. [°] 51 [’] S – 19. [°] 54 [’] E, 14.3.2006; E-Y: 3722, light trap, leg Ruth Müller (TMSA), one spec unsexed, Botswana, Kasane, groundtraps, 1 – 19.9.85; Beyleveld (TMSA), one spec unsexed, Kwandu River, WestCaprivi, SE 1723 Cd, 14 - XI- 1979, C. H. Scholtz COLS 2073 (SANC), one spec unsexed, 20 km N MAUN, BOTSWANA, SE 1923, Dc, 16 - I- 1978, Holm, Jacobs, Kirsten, Scholtz, COLS 23072 (SANC), one spec unsexed BOTSWANA, Makgadikgadi Pans, Res, Khumaga, 20. [°] 26 [’] S 24. [°] 35 [’] E, 04. xii. 1980, P. Reavell, COLS 23074 (SANC), one spec unsexed, BOTSWANA, Sepupa, SE 18 22 Ca 5 – 6. I. 1985, University of Pretoria, Dept. Entomology Tour ’ 85, COLS 22492 (SANC), one spec unsexed, MAUN. Botswana, (16 kms N) 18. I. 78, Grey sand, Open, Mopane Woodland, C (Bait), H. H. Aschenborn, COLS 08770 (SANC).	en	Deschodt, Christian M., Davis, Adrian L. V., Scholtz, Clarke H. (2016): Three new species, a synonymy and taxonomic notes on southern African members of the dung beetle genera Scarabaeus s. str. Linnaeus, 1758 and Escarabaeus Zídek & Pokorný, 2011 (Scarabaeidae: Scarabaeinae). Journal of Natural History 51 (5 - 6): 299-316, DOI: 10.1080/00222933.2016.1261949, URL: http://dx.doi.org/10.1080/00222933.2016.1261949
03B7AE4BFFC0BC55FEC1FB19601EF925.taxon	etymology	Etymology: This species is named in honour of Remi the only son of CD.	en	Deschodt, Christian M., Davis, Adrian L. V., Scholtz, Clarke H. (2016): Three new species, a synonymy and taxonomic notes on southern African members of the dung beetle genera Scarabaeus s. str. Linnaeus, 1758 and Escarabaeus Zídek & Pokorný, 2011 (Scarabaeidae: Scarabaeinae). Journal of Natural History 51 (5 - 6): 299-316, DOI: 10.1080/00222933.2016.1261949, URL: http://dx.doi.org/10.1080/00222933.2016.1261949
03B7AE4BFFC0BC55FEC1FB19601EF925.taxon	description	Description: Large beetle, black. Body size of holotype: Head capsule length 5.4 mm, combined length of pronotum and elytra over midline 15.3 mm, widest width over elytra 12.1 mm, height 8.0 mm. Head: Surface anteriorly rugose, without granulation. Rugosity changes to punctures posterioriad. No projection on dorsal or ventral surface of clypeus. Clypeal teeth slightly upturned with anterior half of teeth smooth. Notch between innermost clypeal teeth rounded; notch between first and second clypeal teeth right angled. Vertex slightly convex. Antennae tan brown. Pronotum: Punctate with punctures far apart, extending to margins. Setae in punctures very short and only seen under at least 60 x magnification. Surfaces between punctures with microgranules. Single row of punctures on posterior margin. Protibia: Inside margin smooth with dark long setae evenly spaced. Outside margin with four teeth in distal three quarters, proximal tooth smallest. Fine serrations all along inside margin and between teeth. Elytra: Ribbed. Shagreened on interstriae. Striae clearly discernible with the naked eye. Margin between striae and interstriae well defined. Interstriae without setae. Sterna: Median longitudinal groove over meta- and mesosternum, deepening to form a fovea on each of the sterna. Mesosternum with long thin setae. Mesosternum sloping at about 45 degrees anteriorly. Metasternum smooth. Pygidium: Shagreened with small irregular punctures far apart. Aedeagus: See Figure 2 (E). Although very similar to that of Escarabaeus satyrus Boheman, 1860 (Figure 2 (F )), it is generally more slender and the spike is broader. Variability: Some size variation is observed (smallest and largest specimens in type series: head capsule length 5.0 – 6.5 mm, combined length of pronotum and elytra over midline 14.7 – 18.9 mm, widest width over elytra 11.2 – 14.2 mm, height 7.1 – 9.2 mm.). The ribbing in some specimens is not as clear.	en	Deschodt, Christian M., Davis, Adrian L. V., Scholtz, Clarke H. (2016): Three new species, a synonymy and taxonomic notes on southern African members of the dung beetle genera Scarabaeus s. str. Linnaeus, 1758 and Escarabaeus Zídek & Pokorný, 2011 (Scarabaeidae: Scarabaeinae). Journal of Natural History 51 (5 - 6): 299-316, DOI: 10.1080/00222933.2016.1261949, URL: http://dx.doi.org/10.1080/00222933.2016.1261949
03B7AE4BFFC0BC55FEC1FB19601EF925.taxon	diagnosis	Diagnosis: This new species keys out to E. satyrus in Zur Strassen, 1967. The elytra are however strongly ribbed. The punctures on the pronotum are also much further apart.	en	Deschodt, Christian M., Davis, Adrian L. V., Scholtz, Clarke H. (2016): Three new species, a synonymy and taxonomic notes on southern African members of the dung beetle genera Scarabaeus s. str. Linnaeus, 1758 and Escarabaeus Zídek & Pokorný, 2011 (Scarabaeidae: Scarabaeinae). Journal of Natural History 51 (5 - 6): 299-316, DOI: 10.1080/00222933.2016.1261949, URL: http://dx.doi.org/10.1080/00222933.2016.1261949
03B7AE4BFFC0BC55FEC1FB19601EF925.taxon	distribution	Distribution: This species has a distribution following the Okavango and Zambezi River floodplains of north western Botswana, extending northwards into the Caprivi Strip of Namibia and possibly into southern Angola and Zambia. New synonymy	en	Deschodt, Christian M., Davis, Adrian L. V., Scholtz, Clarke H. (2016): Three new species, a synonymy and taxonomic notes on southern African members of the dung beetle genera Scarabaeus s. str. Linnaeus, 1758 and Escarabaeus Zídek & Pokorný, 2011 (Scarabaeidae: Scarabaeinae). Journal of Natural History 51 (5 - 6): 299-316, DOI: 10.1080/00222933.2016.1261949, URL: http://dx.doi.org/10.1080/00222933.2016.1261949
03B7AE4BFFC0BC55FEC1FB19601EF925.taxon	description	Species requiring further investigation Relationships and validity of Scarabaeus convexus (Hausmann, 1807), Scarabaeus spretus Zur Strassen, 1962 and Scarabaeus rusticus (Boheman, 1857) require further investigation. They are currently considered as close relatives showing combined distributions from the southwest Cape along the southeast seaboard to uplands and highlands of northeast South Africa, where, S. convexus is found on cool moist highlands and their characters. S. rusticus on drier warmer uplands. However, there may be some discrepancy between distribution pattern and type localities for S. convexus and S. spretus, which are distributed primarily on southwest winter rainfall uplands and the southern seaboard of South Africa.	en	Deschodt, Christian M., Davis, Adrian L. V., Scholtz, Clarke H. (2016): Three new species, a synonymy and taxonomic notes on southern African members of the dung beetle genera Scarabaeus s. str. Linnaeus, 1758 and Escarabaeus Zídek & Pokorný, 2011 (Scarabaeidae: Scarabaeinae). Journal of Natural History 51 (5 - 6): 299-316, DOI: 10.1080/00222933.2016.1261949, URL: http://dx.doi.org/10.1080/00222933.2016.1261949
03B7AE4BFFC0BC55FEC1FB19601EF925.taxon	description	Relationships and validity of Scarabaeus ebenus Klug, 1855; and Scarabaeus interstitialis Boheman, 1857; have also been investigated. Clines from larger to smaller-bodied and denser to sparser punctation occur from the northern edge of the Highveld grasslands to lowland savannah. However, it is currently considered that smaller-bodied, more sparsely punctate inland savannah populations may be separated from eastern lowland and east coastal populations on the basis of larger (S. interstitialis) versus very small punctures (S. ebenus). Further investigation is justified.	en	Deschodt, Christian M., Davis, Adrian L. V., Scholtz, Clarke H. (2016): Three new species, a synonymy and taxonomic notes on southern African members of the dung beetle genera Scarabaeus s. str. Linnaeus, 1758 and Escarabaeus Zídek & Pokorný, 2011 (Scarabaeidae: Scarabaeinae). Journal of Natural History 51 (5 - 6): 299-316, DOI: 10.1080/00222933.2016.1261949, URL: http://dx.doi.org/10.1080/00222933.2016.1261949
