taxonID	type	description	language	source
03B7AD03FFA4FF98EDBCFBD082B9FAA4.taxon	materials_examined	Type material. The original type material from “ Bíly Potok, Trebušany ” (on Mt. “ Pop Ivan ” (today Pip Ivan, at that time Czechoslovakia, today Ukraine) is considered to be lost with certitude. It could not be located in any collection of the Czech Republic (J. Rusek in litt.) and no type specimen was found in the heritage of Stach (W. M. Weiner in litt.) who was in active contact with Mirko Kseneman (J. Rusek in litt.). In view of these, an eligible specimen collected at the type locality is designated herein as neotype for the species. Neotype adult female in alcohol by present designation, labelled: Ukraine, Marmarosh ridge, Pip Ivan Mt., near Dilove village (former Trebušany), Zakarpattia district, Rakhiv region, subalpine meadow, moss, 1750 m a. s. l., 18.0 8.1993, leg. Kaprus’ I. J., det. Kaprus’ I. J. (No. 5.1.4.5, collection of invertebrate animals in the State Natural History Museum of Ukrainian National Academy of Sciences, L'viv, Ukraine). Other material. Four females and three males: same data as neotype; One subadult: Romania, Igniş Mts, Baia Mare (Nagybánya), Izvoare Station, near Tăul lui Dumitru, 31. viii. 2004, N 47 º 49.431 ’ EO 23 º 41.870 ’, 1143 m a. s. l., turf bog, from moss, leg. D. Murányi & K. M. Orci; Four subadults: Romania, Igniş Mts, Baia Mare (Nagybánya), Izvoare Station, Cheile Tătarului 1. ix. 2004, N 47 º 48.438 ’ EO 23 º 45.580 ’, 738 m a. s. l., leg. D. Murányi & K. M. Orci; Six subadults: Romania, Piatra Mts, Săpânţa (Szaplonca), Valea Brazi, 2. ix. 2004, N 47 º 49.775 ’ EO 23 º 44.606 ’, 841 m a. s. l., leg. D. Murányi & K. M. Orci. (Figs 8 – 9, 17, 19, 24 b, 25); Two subadults: Romania, Rodnei Mts, Borşa (Borsa), Borşa Station, source area of the Aranyos-Beszterce under Gargaló 3. ix. 2004, 1688 – 1711 m a. s. l., leg. D. Murányi & K. M Orci.; one male and one female: Romania, Rodnei Mts, Borşa, N slope of Pietrosul Rodnei (2303 m), thereabout Iezer Meteorological Station, 1780 m a. s. l., under Pinus mugo, soil with Polytrichum sp., 26. vii. 2004, leg. J. M. Radwański (in Dányi et al. (2006) erroneously published as “ Munţii Rodnei, Borşa, soil under Picea abies and Pinus sylvestris with Vaccinium sp., 700 m a. s. l. “); one subadult: Romania, Rodnei Mts, Borşa, N slope of Pietrosul Rodnei (2303 m), c. a. 1200 m a. s. l. Plagiothecio-Piceetum association, litter and soil, 27. vii. 2004, leg. J. M. Radwański; five males, five females and one subadult: Romania, Rodnei Mts, V ậrful Ineu Peak (Ünőkő), 2279 m a. s. l., 1. viii. 2008, leg. Gy. Traser & S. Schiffer (males on Figs 1 – 7, 10 – 14, 16, 18, 21 – 23, 24 a, 26 – 30; female on Fig. 15 and 20).	en	Dányi, László, Traser, György, Kaprus’, Ighor J. (2010): Redescription of Friesea handschini Kseneman, 1938 (Collembola, Neanuridae) with notes on intraspecific variability of the species. Zootaxa 2620: 45-55, DOI: 10.5281/zenodo.198089
03B7AD03FFA4FF98EDBCFBD082B9FAA4.taxon	diagnosis	Diagnosis. Friesea handschini Kseneman, 1938 belongs to the Polyacanthella Schäffer subgenus of Friesea Dalla Torre. Six well developed anal spines and one spiniform seta (according to the terminology of Palacios-Vargas (1987 )), anal spines straight, without papillae. 8 + 8 eyes, furcula vestigial: mucro absent, dens shortened with 2 – 3 setae (type 3 and 4 according to Cassagnau (1958) and Massoud (1967 )). Tita I – III with 2,3,3 knobbed tenent hairs dorsally and 2,2,2 ventrally.	en	Dányi, László, Traser, György, Kaprus’, Ighor J. (2010): Redescription of Friesea handschini Kseneman, 1938 (Collembola, Neanuridae) with notes on intraspecific variability of the species. Zootaxa 2620: 45-55, DOI: 10.5281/zenodo.198089
03B7AD03FFA4FF98EDBCFBD082B9FAA4.taxon	description	Redescription. Habitus as in Fig. 1 with dark blue colour. Body length 0.6 – 0.8 mm (without antennae). Head. Head 0.12 – 0.16 mm, length of antennae 0.09 – 0.12 mm. Head / antenna ratio: 1.1 – 1.2 (Fig. 1). Ant. I – IV ratio 24: 24: 20: 22 (ventrally). Ant. III and IV dorsally fused, ventrally weakly separated (Fig. 2). Ant. IV with deeply retracted unilobed apical bulb (Fig. 3), and 6 curved sensilla (Fig. 2). Ant. III with one curved sensillum ventrally and one dorsally (Fig 2). AOIII typical of the genus. Ant. I with 7, Ant. II with 13 setae. 8 + 8 pigmented ocelli, ‘ E’ and ‘ F’ smaller than the other ones (Fig. 6). Ocular area with 3 setae (oc 1 – 3 in Fig. 6). Prelabral / labral chaetotaxy as in Fig. 4 and Fig. 5. 4 prelabral and 10 labral setae (4 / 334), medial setae in first and second labral row longer. Two median prelabral setae situated close together in midline, two other much more laterally (Fig. 7). Labium as in Fig. 7. Mandible with 7 – 8 teeth (Fig. 8). Maxilla as in Fig. 9. Head dorsal chaetotaxy as in Fig. 6 and Tab. 1, setae a 0, d 0 and p 1 present. a) Cephalic chaetotaxy. Type of setae sd d a 0 oc c p Number of setae 5 1 + 4 1 3 3 4 Setae sd 1 – 5 d 0 + d 1 – 4 a 0 oc 1 – 3 c 1 – 3 p 1 – 4 b) Postcephalic chaetotaxy. Row of setae a m p Remarks Th. I – 4 – m 1 – 4 Th. II 5 2 5 a 1 – 5; m 4,5; p 1 – 5; m 5 and p 3 are ‘ s’, m 4 may be missing, m 5 can be considered also as m 6 Th. III 4 2 5 a 1,3,4,5; m 4,5; p 1 – 5; m 5 and p 3 are ‘ s’, a 2 always missing, sometimes a 3 too Abd. I – III 3 2 5 a 1,3,5; m 3,4; p 1 – 5; p 4 is ‘ s’; frequently asymmetrical arrangement of setae in these segments, m 4 often moved forward to the ‘ a’ row Abd. IV 3 2 5 a 1,2,5; m 3,4; p 1 – 5; p 4 is ‘ s’, a 3 sometimes present Abd. V 2 – 3 a 1,4; p 1,3,4; p 3 is ‘ s’ Abd. VI 2 – 1 + 2 a 1,2; p 0,1,3; a 1,2 and p 1 spines; p 0 spiniform seta Thorax, abdomen, legs. Dorsal chaetotaxy of Th. I – III as in Figs 6, 10, 12 and in Tab. 1. Dorsal chaetotaxy of Abd. I – VI as in Figs 11 – 16 and in Tab. 1. Dorsal chaetotaxy often with some variation (e. g. Figs 14 – 15). Body setae smooth and acuminate, but large macrosetae on Abd. V – VI often with knobbed tips (Fig. 17 and 19). Abd. VI with 6 anal spines without distinct papillae in positions of a 1, a 2, p 1 (Figs 17 – 20). Anal spines a 1 shorter. Seta p 0 on Abd. VI short, spiniform, thicker than other setae (Figs 17 – 20). Anal spines in subadult specimens more or less knobbed at the apex (Fig. 17 and 19), seem to tend to break down (Fig. 18 and 20). Legs’ chaetotaxy from I to III: subcoxae I: 0,2,2; subcoxae II: 0,2,2; coxae: 3,8,8; trochanters: 5,5,5 (4); femora: 12,11,10. Tita I – III with 18, 18, 17 setae respectively (11 in the distal verticil and 6 or 7 further ones more proximally as in Fig. 23 and Fig. 24). Tita I – III respectively with 2,2,2 knobbed tenent hairs ventrally and 2,3,3 dorsally (Fig. 24). Setae on legs as in Figs 21 – 24. One longer seta on the ventral side of each femur (Figs 22 a – c). Claws without teeth (Figs 23 – 24). Empodial appendage absent. Ventral chaetotaxy as in Figs 25 – 26. Thoracic sterna without setae (Fig. 21). Ventral tube with 4 + 4 (3 – 5) distal setae (Fig. 30). Furcula reduced, mucro absent, dens with 3 (2) + 3 (2) setae (Figs 27 – 28). Chaetotaxy of manubrium as in Fig. 28. Retinaculum with two teeth, without seta (Figs 27 – 28). Male genital plate with 4 (5) + 4 (5) eugenital and about 30 circumgenital setae (Fig. 26). Chaetotaxy of anal lobes as in Fig. 29.	en	Dányi, László, Traser, György, Kaprus’, Ighor J. (2010): Redescription of Friesea handschini Kseneman, 1938 (Collembola, Neanuridae) with notes on intraspecific variability of the species. Zootaxa 2620: 45-55, DOI: 10.5281/zenodo.198089
03B7AD03FFA4FF98EDBCFBD082B9FAA4.taxon	discussion	Discussion. Using the key of Massoud (1967) we can identify Friesea mauriesi Cassagnau, 1958 having larger size (about 1.3 mm, instead of 0.6 – 0.8 mm in F. handschini) and 8 knobbed anal spines (6 spines in F. handschini). Among the North-American Friesea species of 8 + 8 eyes we find Friesea fara Christiansen & Bellinger, 1974 with highly similar character composition, but differing in seta a 2 on Abd. V (present in F. fara, missing in F. handschini) and in the spine-like feature of seta p 0 on Abd. VI (simple seta in F. f a r a, spiniform seta in F. handschini).	en	Dányi, László, Traser, György, Kaprus’, Ighor J. (2010): Redescription of Friesea handschini Kseneman, 1938 (Collembola, Neanuridae) with notes on intraspecific variability of the species. Zootaxa 2620: 45-55, DOI: 10.5281/zenodo.198089
03B7AD03FFA4FF98EDBCFBD082B9FAA4.taxon	distribution	Distribution and ecology. Till date F. handschini was reported from Austria (Törne 1958, Querner 2008), France (Poinsot-Balaguer 1976), Germany (Schulz 1991), Poland (Skarżyński et al. 2002), Romania (Fiera 2007), Slovakia (Rusek 1963) and Ukraine (Kaprus’ et al. 2006). The records suggest a Central European montane chorotype, but the misinterpreted characters of the species question the reliability of all West- European data. Thus, the identities of the Austrian, French and German specimens need to be confirmed. The necessity for this is also indicated by the species’ Spanish data (Selga 1958, Jordana et al. 1990) that were later proved to be Friesea nigrimontana Cassagnau (Jordana et al. 1997). According to this, the species was considered as Carpathian endemic by Kaprus’ (1999), Kaprus’ et al. (2006) and Sterzyńska and Kaprus’ (2000).	en	Dányi, László, Traser, György, Kaprus’, Ighor J. (2010): Redescription of Friesea handschini Kseneman, 1938 (Collembola, Neanuridae) with notes on intraspecific variability of the species. Zootaxa 2620: 45-55, DOI: 10.5281/zenodo.198089
03B7AD03FFADFF98EDBCFC5E83DFF90C.taxon	materials_examined	Material examined. Eleven males, four females and six juveniles: Romania, Rodnei Mts, V ậrful Ineu Peak (Ünőkő), 2279 m a. s. l., 1. viii. 2008, leg. Gy. Traser & S. Schiffer; together with F. handschini.	en	Dányi, László, Traser, György, Kaprus’, Ighor J. (2010): Redescription of Friesea handschini Kseneman, 1938 (Collembola, Neanuridae) with notes on intraspecific variability of the species. Zootaxa 2620: 45-55, DOI: 10.5281/zenodo.198089
