taxonID	type	description	language	source
03B58785DE031E385EA4771CFB64854C.taxon	materials_examined	Type species — Laelaps claviger Berlese, 1883: 2, by original designation.	en	Joharchi, Omid, Ermilov, Sergey G., Khaustov, Alexander A. (2020): Two new species of Cosmolaelaps Berlese (Acari: Laelapidae) from Sri Lanka. Zootaxa 4743 (2): 151-166, DOI: 10.11646/zootaxa.4743.2.1
03B58785DE031E385EA4771CFB64854C.taxon	diagnosis	Diagnosis. The diagnosis of Cosmolaelaps used here is based on that of Moreira et al. (2014).	en	Joharchi, Omid, Ermilov, Sergey G., Khaustov, Alexander A. (2020): Two new species of Cosmolaelaps Berlese (Acari: Laelapidae) from Sri Lanka. Zootaxa 4743 (2): 151-166, DOI: 10.11646/zootaxa.4743.2.1
03B58785DE031E315EA470B3FDAE8442.taxon	description	(Figures 1 – 24)	en	Joharchi, Omid, Ermilov, Sergey G., Khaustov, Alexander A. (2020): Two new species of Cosmolaelaps Berlese (Acari: Laelapidae) from Sri Lanka. Zootaxa 4743 (2): 151-166, DOI: 10.11646/zootaxa.4743.2.1
03B58785DE031E315EA470B3FDAE8442.taxon	materials_examined	Type material: Holotype, female, Sri Lanka, Sabaragamuwa Province, Polgampola, 06 ° 27 ’ N, 080 ° 12 ’ E, alt. 42 m, 24 January 2019, O. Joharchi, S. G. Ermilov & A. A. Khaustov coll., from forest soil-litter near the Thambadola Ella waterfall (in TSUMZ). Paratypes, three females and seven males same data as holotype (two females and four males in TSUMZ, one female and three males in ZISP).	en	Joharchi, Omid, Ermilov, Sergey G., Khaustov, Alexander A. (2020): Two new species of Cosmolaelaps Berlese (Acari: Laelapidae) from Sri Lanka. Zootaxa 4743 (2): 151-166, DOI: 10.11646/zootaxa.4743.2.1
03B58785DE031E315EA470B3FDAE8442.taxon	diagnosis	Diagnosis (adult female and male). Dorsal shield with well reticulate ornamentation over whole surface, covering most of the idiosoma, narrowing from level of setae r 3, progressively tapering until rounded posteriorly, bearing 39 pairs of subequal oblong-mucronate setae (22 pairs of podonotal setae and 17 pairs of opisthonotal setae, including two pairs of Zx setae) except j 1, z 1 and Z 5 foliate-dentate and shorter. Adult female sternal shield smooth almost throughout; anterior margin hardly conspicuous and posterior margin concave, bearing three pairs of setae and two pairs of large slit shape pore-like structures, st 1 long enough to reach base of st 2, ratio of shield length / width (at broadest level) 1. Genital shield ratio of length / width (at broadest level) 2.4, surface without any distinct reticulation, almost ≃ completely smooth, except two diagonal lines in posterior part ≃ which confluence with another horizontal line and forming semi-triangular in posterior part of surface. Anal shield ratio of length / width (at broadest level) 1.4, cribrum well developed. Peritreme long, reaching mid-level of coxa I (near s 1). Fixed digit of chelicera with 10 ≃ – 11 teeth of various sizes. Some dorsal setae on tibia and genu of legs I – IV and pd 2, pd 3 on tarsi III – IV feather-like. Male with holoventral shield; shield with a strongly sclerotised median circular shape projection; spermatodactyl short and apically curved dorsally; av 1 on femur of leg II lanceolate, pv on tibia IV and av 2 on tarsus II, IV, pv 2 on tarsus III, IV spur-like (apically blunt) and trochanter bearing a large bump shape spur (Fig. 24).	en	Joharchi, Omid, Ermilov, Sergey G., Khaustov, Alexander A. (2020): Two new species of Cosmolaelaps Berlese (Acari: Laelapidae) from Sri Lanka. Zootaxa 4743 (2): 151-166, DOI: 10.11646/zootaxa.4743.2.1
03B58785DE031E315EA470B3FDAE8442.taxon	description	Description. Female (n = 4). Dorsal idiosoma (Figs 1, 7 & 9). Dorsal shield 295 – 305 long, 180 – 190 wide, covering most of dorsal idiosoma, well reticulated over whole surface, narrowing from level of setae r 3, progressively tapering until rounded posteriorly. Shield with 39 pairs of oblong-mucronate setae, 22 pairs of podonotal setae (j 1 – 6; z 1 – 6; s 1 – 6; r 2 – 5), and 17 pairs of opisthonotal setae (J 1 – 5; Z 1 – 5; S 1 – 5 and two pairs of Zx setae) except j 1, z 1 and Z 5 foliate-dentate; two unpaired supernumerary setae (Jx) present (Figs 1, 7 & 9). Almost all setae with 2 – 4 minute barbs (Fig. 9) and uniform in length (21 – 26) and thickness, except Z 5 (13 – 15), z 1 (8 – 10) and j 1 (15 – 17) length of setae slightly increasing from central to lateral (Figs 1, 7 & 9). Narrow strip of soft cuticle surrounding dorsal shield with two pairs of R setae (R 3, R 4). Shield with about 16 pairs of discernible pore-like structures, including 15 poroids (id 1, id 2, id 4, id 5, id 6, idm 1, idm 2, idm 3, idm 4, idm 5, idx, idl 3) and four gland openings (gd 1, gd 2, gd 5, gd 6), others indistinct (Figures 1 and 7). Ventral idiosoma (Figs 2 & 8). Tritosternum with paired pilose laciniae (53 – 58), fused basally (2 – 4), columnar base 20 – 24 × 8 – 11 wide; presternal area poorly sclerotised. Sternal shield (length 86 – 90) narrowest between coxae II (58 – 60), widest (88 – 90), anterior margin completely indistinct and posterior margin concave; shield with three pairs of smooth pointed setae (st 1 32 – 34, st 2 22 – 24, st 3 22 – 24, st 1 reaching base of next setae) and two pairs of large slit-shaped pore-like structures (iv 1 adjacent to setae st 1; iv 2 between st 2 and st 3); sternal shield smooth almost throughout and without distinct reticulation. Metasternal setae st 4 (19 – 21) and metasternal poroids located on soft integument; metasternal platelets absent. Endopodal plates II / III completely fused to sternal shield, endopodal plates III / IV elongate, narrow and curved. Genital shield broad, flask-shaped, not adjacent to anal shield, expanded laterally behind setae st 5, length 137 – 139, maximum width 54 – 58, posterior margin rounded, shield smooth almost throughout, except two diagonal lines in posterior part that fuse with another horizontal line and forming a faintly semi-triangular shape; bearing a pair of simple setae st 5 (19 – 21); paragenital poroids iv 5 located on soft cuticle lateral to shield near seta st 5. Shield flanked by two pairs of irregular small platelets. Anal shield subtriangular, rounded anteriorly, length 55 – 57, width 40 – 44, anterior half lineate-reticulate, para-anal setae and post-anal seta subequal in length (14 – 16), cribrum consisting of a terminal tuft and pair of anterior arms reaching about to midlevel of distance between post-anal and para-anal setae; anal poroids gv 3 on anterolateral margin of anal shield. Soft opisthogastric cuticle surrounding genital and anal shields with 12 pairs of subequal setae (12 – 16) (Jv 1 – Jv 5, Zv 1 – Zv 5, UR 1, UR 2). Setae Jv 5, Zv 5 and UR 2 spatulate, metapodal plates absent. Exopodal and parapodal platelets not fused, strip-like, extending narrowly behind coxae IV. Peritreme extending anteriorly to mid-level of coxa I; peritrematal shield narrow, free from exopodal shields, protrusion band of cuticle lateral to the peritreme at the level of coxae II – III, a lyrifissure ip at level of coxa II, post-stigmatic section bearing two pore-like structures, a lyrifissures ip and a gland pore gp (Figs 2 & 8). Gnathosoma (Figs 4 – 6, 10 – 12). Hypostomal groove with six transverse rows of denticles, each row with about 11 – 25 small teeth (Figs 4 & 10), groove wider anteriorly. Hypostome with four pairs of setae, h 1 (28 – 30), h 2 (10 – 12), h 3 (28 – 30), palpcoxal setae (pc) (24 – 26) (Figs 4 & 10). Epistome trapezoidal, central part irregularly denticulate (Figs 5, 11). Corniculi robust and horn-like, extending slightly beyond palp trochanter. Internal malae with median and lateral projections, fringed, inner lobes touching with outer lobes; labrum with pilose surface. Chaetotaxy of palps: trochanter 2, femur 5, genu 6, tibia 14, tarsus 15, all setae smooth and needle-like except al on palp femur and genu thickened and apically spatulate, palp tarsal apotele two-tined (Fig. 12). Fixed digit of chelicera with an offset and most distal large tooth (gabelzhan) followed by 9 – 10 variously sized teeth, 4 nd, 6 th, 9 th larger, a setaceous pilus dentilis, dorsal cheliceral setae prostrate, arthrodial membrane with a rounded flap and normal filaments and cheliceral lyrifissures indistinct, movable digit with two teeth (Fig. 6). Legs (Figs 13 – 18). Legs II and III short (213 – 217, 195 – 204), I and IV longer (273 – 296, 294 – 301). Chaetotaxy normal for free-living Laelapidae: Leg I (Fig. 15): coxa 0 - 0 / 1, 0 / 1 - 0, trochanter 1 - 0 / 2, 1 / 1 - 1, femur 2 - 2 / 1, 3 / 3 - 2 (pv 1, pv 2, ad 1, pd 1, pd 2, pd 3 thickened, all dorsal setae with minute barbs), genu 2 - 3 / 2, 3 / 1 - 2 (all dorsal setae feather-like, lateral and ventral setae thickened except av 2), tibia 2 - 3 / 2, 3 / 1 - 2 (all dorsal setae feather-like, lateral setae thickened). Leg II (Fig. 16): coxa 0 - 0 / 1, 0 / 1 - 0, trochanter 1 - 0 / 1, 0 / 2 - 1, femur 2 - 3 / 1, 2 / 2 - 1 (ad 1 truncate, pd 1, pd 2 falcate and with minute barbs), genu 2 - 3 / 1, 2 / 1 - 2 (ad 1, ad 2, ad 3 feather-like, pd 1, pd 2 falcate and with minute barbs), tibia 2 - 2 / 1, 2 / 1 - 2 (ad 1, ad 2 feather-like, pd 1, pd 2 thickened and with minute barbs). Leg III (Fig. 17): coxa 0 - 0 / 1, 0 / 1 - 0, trochanter 1 - 1 / 1, 0 / 1 - 1, femur 1 - 2 / 1, 1 / 0 - 1 (ad 1, pd, pl thickened, ad 1 apically blunt), genu 2 - 2 / 1, 2 / 1 - 1 (pd 1, pd 2 feather-like, ad 1, ad 2 lanceolate, with minute barbs, pl thickened, Fig. 13), tibia: 2 - 1 / 1, 2 / 1 - 1 (pd 1, pd 2 feather-like, ad 1 lanceolate, with minute barbs, pl thickened, Fig. 13). Leg IV (Fig. 17): coxa 0 - 0 / 1, 0 / 0 - 0, trochanter 1 - 1 / 1, 0 / 1 - 1 (ad spatulate), femur 1 - 2 / 1, 1 / 0 - 1 (ad 1, ad 2 spatulate, pd thickened with minute barbs, Fig. 14), genu 2 - 2 / 1, 3 / 0 - 1 (pd 1, pd 2, pd 3 feather-like, ad 1, ad 2 lanceolate, with minute barbs, av thickened, Fig. 14), tibia 2 - 1 / 1, 3 / 1 - 2 (pd 1, pd 2, pd 3 feather-like, ad 1, ad 2 lanceolate, with minute barbs, av, al, pv thickened, Fig. 14). Tarsi II-IV with 18 setae (3 - 3 / 2, 3 / 2 - 3 + mv, md). All pretarsi with well-developed paired claws and rounded pulvilli and a long thin stalk, ventral and lateral setae on tarsus II and III thickened, pd 2, pd 3 on tarsi III-IV feather-like. Insemination structures (Fig. 3). Laelapid-type sperm access system, tubulus long, wider at the solenostome level of coxa III and entering sacculus via a pair of circular openings. Sacculus reniform, the proximal ends of the tubulus slightly swollen at junction with ramus. Male (n = 7). (Figs 19 – 24). Dorsal idiosoma. Dorsal shield (268 – 295) long, (154 – 172) wide; ornamentation and chaetotaxy as in female. Ventral idiosoma (Figs 19, 21 & 23). Sternal, genital, endopodal, ventral and anal shields fused into holoventral shield, weakly reticulate, more distinct behind ZV 1, bearing st 1 – 5, five pairs of opisthogastric setae (Jv 1 – Jv 3, Zv 1, Zv 2) in addition to circumanal setae; six pairs of poroids and a pore-like (gv 3) laterad of para-anal setae, gland pore gv 2 behind coxa IV not discerned; shield with a strongly sclerotised median circular shape projection (Fig. 22), cribrum with 3 – 4 irregular rows of spicules and a pair of anterior arms reaching about to mid-level of distance between post-anal and para-anal setae. Soft opisthogastric cuticle with five pairs setae. Gnathosoma. Epistome and subcapitulum similar to female. Fixed digit of chelicera with seven teeth of various sizes, pilus dentilis setaceous. Movable digit of chelicera with one tooth, spermatodactyl short and apically curved dorsally, with blunt tip, free portion of spermatodactyl shorter than movable digit, fringed hyaline arthrodial process at base of movable digit (Fig. 20). Palps similar to those of female. Legs. Chaetotaxy as in female. Seta av 1 on femur of leg II lanceolate (Fig. 24), pv on tibia IV (Fig. 23) and av 2 on tarsus II, IV, pv 2 on tarsus III, IV spur-like (apically blunt) and tochanter II bearing a large dome-shaped spur (Fig. 24).	en	Joharchi, Omid, Ermilov, Sergey G., Khaustov, Alexander A. (2020): Two new species of Cosmolaelaps Berlese (Acari: Laelapidae) from Sri Lanka. Zootaxa 4743 (2): 151-166, DOI: 10.11646/zootaxa.4743.2.1
03B58785DE031E315EA470B3FDAE8442.taxon	etymology	Etymology. Named after the type locality, Sri Lanka, previously known as Ceylon.	en	Joharchi, Omid, Ermilov, Sergey G., Khaustov, Alexander A. (2020): Two new species of Cosmolaelaps Berlese (Acari: Laelapidae) from Sri Lanka. Zootaxa 4743 (2): 151-166, DOI: 10.11646/zootaxa.4743.2.1
03B58785DE031E315EA470B3FDAE8442.taxon	diagnosis	Differential diagnosis. The dorsal setae in Cosmolaelaps ceylonensis are similar in shape to those of C. rectangularis Sheals, 1962, C. serratus Trägårdh, 1952, C. claviger (Berlese, 1883), C. diversus (Karg, 1994) and C. bengalensis (Bhattacharyya, 1968), described from Argentina, Saint Lucia, Tahiti, Galápagos Islands and India, respectively. It differs from all these species by the different shape of j 1, z 1 and Z 5 setae, foliate-dentate in C. ceylonensis (Fig. 1) (vs. j 1, z 1 and Z 5 setae obovate-spatulate or simple and smooth in the other species, never foliate-dentate). The sternal and genital setae (st 1 – st 3 and st 5) are simple and the shields are smooth, without distinct reticulation, except for two diagonal lines in the posterior part of the genital shield which fuse with another horizontal line to form a semi-triangular figure in posterior part of the genital shield surface) in C. ceylonensis (Fig. 2). Both or at least one of the sternal or genital shields have distinct reticulation in C. rectangularis, C. serratus, C. claviger, C. diversus; and setae st 1 – st 3 and st 5 are spatulate and with some barbs respectively in C. bengalensis. Some dorsal setae on the tibia and genu of legs I – IV and pd 2, pd 3 on tarsi III – IV are feather-like in C. ceylonensis (Figs 13 – 18); while the dorsal setae on tibia and genu of legs I – IV in all the other species mentioned are sometimes spatulate or thickened but never feather-like, and pd 2, pd 3 on tarsi III – IV simple. In the classification of Karg (1981), C. ceylonensis falls into the claviger species group of Hypoaspis (Cosmolaelaps), because almost all dorsal shield setae oblong-mucronate, but C. ceylonensis is distinguished easily from all species in the claviger species group by the following combination of unique characters: (1) dorsal shield with well reticulate ornamentation over whole surface and narrowing from level of setae r 3, progressively tapering until rounded posteriorly, bearing 39 pairs of oblong-mucronate setae and almost all setae with 2 – 4 minute barbs and uniform in length and thickness except j 1, z 1 and Z 5 foliate-dentate and shorter; (2) both sternal and genital shields obviously smooth and without distinct reticulation (except two diagonal lines in posterior part of genital shield which confluence with another horizontal line and forming semi-triangular in posterior part of genital shield surface) and sternal shield bearing two pairs of large slit shape pore-like structures and st 1 length enough to well reach base of st 2, (3) fixed digit of chelicera with 10 – 11 various size of teeth; (4) some dorsal setae on tibia and genu of legs I – IV (antero-dorsals and postero-dorsals setae on tibia and genu I – II and postero-dorsals setae on tibia and genu III – IV) and pd 2, pd 3 on tarsi III – IV feather-like; (5) Male with holoventral shield; shield with a strongly sclerotised median circular shape projection and spermatodactyl short and apically curved dorsally; (6) ad on tibia IV and ad 2, pd 2 on tarsus IV spur-like (apically blunt) in male.	en	Joharchi, Omid, Ermilov, Sergey G., Khaustov, Alexander A. (2020): Two new species of Cosmolaelaps Berlese (Acari: Laelapidae) from Sri Lanka. Zootaxa 4743 (2): 151-166, DOI: 10.11646/zootaxa.4743.2.1
03B58785DE0A1E345EA471B9FC0D82F2.taxon	description	(Figures 25 – 40)	en	Joharchi, Omid, Ermilov, Sergey G., Khaustov, Alexander A. (2020): Two new species of Cosmolaelaps Berlese (Acari: Laelapidae) from Sri Lanka. Zootaxa 4743 (2): 151-166, DOI: 10.11646/zootaxa.4743.2.1
03B58785DE0A1E345EA471B9FC0D82F2.taxon	materials_examined	Type material: Holotype, female, Sri Lanka, Sabaragamuwa Province, Polgampola, 06 ° 27 ’ N, 080 ° 12 ’ E, alt. 42 m, 24 January 2019, O. Joharchi, S. G. Ermilov & A. A. Khaustov coll., from forest soil-litter near the Thambadola Ella waterfall (in TSUMZ). Paratype, one female, same data as holotype (in TSUMZ).	en	Joharchi, Omid, Ermilov, Sergey G., Khaustov, Alexander A. (2020): Two new species of Cosmolaelaps Berlese (Acari: Laelapidae) from Sri Lanka. Zootaxa 4743 (2): 151-166, DOI: 10.11646/zootaxa.4743.2.1
03B58785DE0A1E345EA471B9FC0D82F2.taxon	diagnosis	Diagnosis (adult female). Dorsal shield covering most of the idiosoma, with reticulate ornamentation over whole surface, narrowing from level of setae r 3, progressively tapering until rounded posteriorly, bearing 39 pairs setae. Most setae long enough to reach well past base of next posterior seta, all setae abruptly taper apically (Figs 25, 30, 32), except j 1, z 1, Z 5 very short (7 – 10) and S 5 longest (44 – 46). Sternal shield smooth almost throughout; anterior margin indistinct and posterior margin almost straight, bearing three pairs of setae and two pairs of large slit-shape pore-like structures (Fig. 33), ratio of shield length / width (at broadest level) 1. Genital shield elongated, posterior margin of genital shield almost reaches anterior margin of anal shield ≃ (Figs 26 & 31), ratio of length / width (at broadest level) 3.2, surface without any distinct reticulation, almost completely smooth (Fig. 34). Anal shield ratio of length / width ≃ (at broadest level) 1.8 and cribrum small, length of post-anal seta almost twice ≃ para-anal setae. Peritreme long, extending to posterior of coxa I, post-stigmatal section of peritrematal shield oval, its tip extending to mid-level of coxa IV. Fixed digit of chelicera with six teeth in two rows, dorsal cheliceral setae thickened, arthrodial membrane reduced to a lobe and without filaments. Femur I and genu III with 12 and 8 setae, respectively.	en	Joharchi, Omid, Ermilov, Sergey G., Khaustov, Alexander A. (2020): Two new species of Cosmolaelaps Berlese (Acari: Laelapidae) from Sri Lanka. Zootaxa 4743 (2): 151-166, DOI: 10.11646/zootaxa.4743.2.1
03B58785DE0A1E345EA471B9FC0D82F2.taxon	description	Description. Female (n = 2). Dorsal idiosoma (Figs 25, 30 & 32). Dorsal shield 316 – 318 long, 160 – 162 wide, covering most of dorsal idiosoma, reticulated over whole surface, narrowing from level of setae r 3, progressively tapering until rounded posteriorly. Shield with 39 pairs of long setae, 22 pairs of podonotal setae on shield (j 1 – 6; z 1 – 6; s 1 – 6; r 2 – 5), and 17 pairs of opisthonotal setae on shield (J 1 – 5; Z 1 – 5; S 1 – 5; and two pairs of Zx), supernumerary setae Jx between J - series absent, most setae long enough to reach well past base of next posterior seta (28 – 38 long), all setae smooth and uniform in thickness and abruptly tapered apically (Fig. 32), except j 1, z 1, Z 5 very short (7 – 10) and S 5 longest (44 – 46). Soft cuticle surrounding dorsal shield with six pairs setae (r 6, R 2 – R 4 and UR 5 – 6) setae. Shield with about 14 pairs of discernible pore-like structures, including ten poroids (id 1, id 2, id 4, id 5, idm 1, idm 2, idm 3, idm 5, idm 6, idl 1) and four gland openings (gd 1, gd 2, gd 6, gd 9), others indistinct (Figures 25 and 30). Ventral idiosoma (Figs 26, 31, 33 & 34). Tritosternum with paired pilose laciniae (58 – 60), fused basally (2 – 4), columnar base 15 – 17 × 10 – 12 wide; presternal area poorly sclerotised, granulate-lineate (Figs 26 & 31). Sternal shield (length 99 – 102) narrowest between coxae II (73 – 75), widest (104 – 106), anterior margin completely indistinct and posterior margin almost straight; bears three pairs of smooth pointed setae (st 1 22 – 24, st 2 21 – 23, st 3 21 – 23), not reaching base of next setae, one pair of lyrifissures adjacent to setae st 1, and a pair of poroids between st 2 and st 3; sternal shield smooth almost throughout (Fig. 33). Metasternal setae st 4 (22 – 24) and metasternal poroids located on soft integument; metasternal platelets absent. Endopodal plates II / III completely fused to sternal shield, endopodal plates III / IV elongate, narrow and curved. Genital shield tongue-shaped, sides almost parallel, elongated, length 134 – 136, maximum width 40 – 42, surface without any distinct reticulation, almost completely smooth (Fig. 34), posterior margin of genital shield almost reaches anterior margin of anal shield (Figs 26 & 31). Shield flanked by pair of minute, slightly elongate paragenital plaetelets; paragenital poroids iv 5 located on soft cuticle lateral to shield near seta st 5. Anal shield subtriangular, rounded anteriorly, length 52 – 54, width 36 – 38, anterior half faintly lineate-reticulate, post-anal seta (20 – 22) twice as long as para-anal setae (10 – 12), cribrum small, with 3 – 4 irregular rows of spicules, limited to region posterior to post-anal seta; anal poroids gv 3 on anterolateral margin of anal shield. Soft opisthogastric cuticle with pair of oval and slightly elongate metapodal plates (10 – 12 long × 2 – 4 wide), and 12 pairs of smooth setae (Jv 1 – Jv 5, Zv 1 – Zv 3) and four pairs of poroids including iv 5. Exopodal and parapodal platelets not fused, strip-like extending narrowly behind coxae IV. Peritreme extending anteriorly to posterior level of coxa I, peritrematal shield narrow, expanded anteriorly, free from exopodal shields, each shield bearing five pore-like structures, a lyrifissure ip and a gland pore gp at level of coxa III, two lyrifissures ip and a gland pore gp on post-stigmatic section (Fig. 26). Gnathosoma (Figs 27 – 29, 35 – 36). Epistome subtriangular and irregularly denticulate (Fig. 28). Hypostomal groove with seven transverse rows of denticles, each row with 8 – 15 small denticles, with smooth anterior transverse lines. Hypostome with four pairs of setae, internal posterior hypostomal setae h 3 (30 – 32), h 1 (24 – 26), h 2 (13 – 15), palpcoxal pc (22 – 24) (Fig. 27). Corniculi robust and horn-like, extending slightly beyond palptrochanter. Chaetotaxy of palps: trochanter 2, femur 5, genu 6, tibia 14, tarsus 15, all setae smooth and needle-like except except al 1 on palp femur and al 1, d 1 on palp genu thickened and apically spatulate, palp tarsal apotele two-tined (Fig. 35). Fixed digit of chelicera with an offset and most distal large tooth (gabelzhan), followed five variously sized teeth in two rows, a setaceous pilus dentilis, dorsal cheliceral setae thickened prostrate, arthrodial membrane reduced to a lobe and without filaments; cheliceral lyrifissures indistinct (Fig. 29). Legs (Figs 37 – 40). Legs II (232 – 237) and III, (206 – 195) short, I (306 – 309) and IV (278 – 286) longer. Chaetotaxy normal for free-living Laelapidae: Leg I (Fig. 37): coxa 0 - 0 / 1, 0 / 1 - 0, trochanter 1 - 0 / 1, 1 / 2 - 1, femur 2 - 2 / 1, 3 / 2 - 2 (pv 3 absent), genu 2 - 3 / 2, 3 / 1 - 2 (pv slightly thickened), tibia 2 - 3 / 2, 3 / 1 - 2 (ventral setae slightly thickened). Leg II (Fig. 38): coxa 0 - 0 / 1, 0 / 1 - 0, trochanter 1 - 0 / 1, 0 / 2 - 1, femur 2 - 3 / 1, 2 / 2 - 1, genu 2 - 3 / 1, 2 / 1 - 2, tibia 2 - 2 / 1, 2 / 1 - 2 (ventral setae slightly thickened). Leg III (Fig. 39): coxa 0 - 0 / 1, 0 / 1 - 0, trochanter 2 - 1 / 1, 0 / 1 - 0, femur 1 - 2 / 1, 1 / 0 - 1, genu 2 - 2 / 1, 2 / 0 - 1 (pv absent), tibia: 2 - 1 / 1, 2 / 1 - 1 (ventral setae thickened). Leg IV (Fig. 40): coxa 0 - 0 / 1, 0 / 0 - 0, trochanter 2 - 1 / 1, 0 / 1 - 1, femur 1 - 2 / 1, 1 / 0 - 1, genu 2 - 2 / 1, 3 / 0 - 1, tibia 2 - 1 / 1, 3 / 1 - 2. Tarsi II-IV with 18 setae (3 - 3 / 2, 3 / 2 - 3 + mv, md); some ventral and lateral setae on tarsi II-III thickened. All pretarsi with well-developed paired claws, rounded pulvilli and normal ambulacral stalk. Insemination structures: Not seen, apparently unsclerotised. Male and immatures. Unknown.	en	Joharchi, Omid, Ermilov, Sergey G., Khaustov, Alexander A. (2020): Two new species of Cosmolaelaps Berlese (Acari: Laelapidae) from Sri Lanka. Zootaxa 4743 (2): 151-166, DOI: 10.11646/zootaxa.4743.2.1
03B58785DE0A1E345EA471B9FC0D82F2.taxon	etymology	Etymology. The species name trichiurus (Greek thrix, hair + oura, tail) refers to the tail-like appearance of the setae S 5.	en	Joharchi, Omid, Ermilov, Sergey G., Khaustov, Alexander A. (2020): Two new species of Cosmolaelaps Berlese (Acari: Laelapidae) from Sri Lanka. Zootaxa 4743 (2): 151-166, DOI: 10.11646/zootaxa.4743.2.1
03B58785DE0A1E345EA471B9FC0D82F2.taxon	diagnosis	Differential diagnosis. Cosmolaelaps trichiurus is similar in general appearance and the shape of the dorsal setae to C. gladii (Karg, 1993) and Cosmolaelaps dioscorea Joharchi, Fan & Arjomandi, 2017 described from Galápagos Islands and Fiji, respectively. Cosmolaelaps trichiurus can be easily distinguished from these two species by S 5 much longer than other dorsal setae (vs. S 5 in the same size with other dorsal setae in both species), genital shield elongated, posterior margin of genital shield almost reaches anterior margin of anal shield (genital shield not elongated in both species), length of post-anal seta almost twice that of para-anal setae (vs. length of post-anal seta shorter or at most subequal of para-anal setae but never longer in both species), fixed digit of chelicera with six teeth, dorsal cheliceral setae thickened, arthrodial membrane reduced to a lobe and without filaments (fixed digit of chelicera with 11 and eight teeth in C. gladii and C. dioscorea, respectively, cheliceral setae not thickened and arthrodial membrane with a rounded flap and normal filaments in both species). According to shape of dorsal setae, C. trichiurus in the classification of Karg (1981), falls into the nasoseta species group of Hypoaspis (Cosmolaelaps), but C. trichiurus is distinguished easily from all species in nasoseta species group by following combination of unique characters: (1) dorsal shield narrowing from level of setae r 3, progressively tapering until rounded posteriorly and most setae long enough to reach well past base of next posterior seta, all setae abruptly taper apically (except j 1, z 1, Z 5 very short) and setae S 5 much longer than other dorsal setae; (2) sternal shield smooth almost throughout; anterior margin indistinct and posterior margin almost straight; (3) genital shield elongated, posterior margin of genital shield almost reaches anterior margin of anal shield and surface almost completely smooth; (4) length of post-anal seta almost twice of para-anal setae; (5) fixed digit of chelicera with six teeth, dorsal cheliceral setae thickened and arthrodial membrane reduced to a lobe (filaments absent); (6) Femur I (2 - 2 / 1, 3 / 2 - 2) and genu III (2 - 2 / 1, 2 / 0 - 1) with 12 (pv 3 absent) and 8 (pv absent) setae, respectively.	en	Joharchi, Omid, Ermilov, Sergey G., Khaustov, Alexander A. (2020): Two new species of Cosmolaelaps Berlese (Acari: Laelapidae) from Sri Lanka. Zootaxa 4743 (2): 151-166, DOI: 10.11646/zootaxa.4743.2.1
