taxonID	type	description	language	source
03B58796B650FFC6FF7EFAC8FBE673BD.taxon	type_taxon	Type species: Phrynus cheiracanthus Gervais, 1842, by original designation. Entomologie. L’Institut, Journal Universal des Sciences et des Sociétés Savantes en France et a l’Étranger, 1 ère Section, 10, 76.	en	Chirivi-Joya, Daniel, Moreno-González, Jairo A., Fagua, Giovanny (2020): Two new species of the whip-spider genus Heterophrynus (Arachnida: Amblypygi) with complementary information of four species. Zootaxa 4803 (1): 1-41, DOI: 10.11646/zootaxa.4803.1.1
03B58796B650FFCCFF7EF987FB507402.taxon	description	Figures 1 – 3, 22 (in part). Table 1	en	Chirivi-Joya, Daniel, Moreno-González, Jairo A., Fagua, Giovanny (2020): Two new species of the whip-spider genus Heterophrynus (Arachnida: Amblypygi) with complementary information of four species. Zootaxa 4803 (1): 1-41, DOI: 10.11646/zootaxa.4803.1.1
03B58796B650FFCCFF7EF987FB507402.taxon	materials_examined	Type locality: Upper Suriname River, Suriname. Examined material (22 females, 25 males, 5 juveniles): BRAZIL: Amapá: Oiapoque: Taparabó (4 ° 00´50.3 ´´ N; 51 ° 41´22.8 ´´ W): Rain forest, over a wide trunk, 31 March 2015, D. Chirivi, J. Murienne, manual collection at night, 1 male, 1 female, 1 juvenile (MZUSP 74497); Rain forest, over a wide trunk, 31 March 2015, D. Chirivi, J. Murienne, manual collection at night, male (MZUSP 74496). Tumucumaque (3 ° 16´58.9 ´´ N; 52 ° 12´06.5 ´´ W): in front of Carbet Mitemps, Rain forest, over a wide trunk, 23 March 2015, D. Chirivi, J. Murienne, manual collection at night, 2 males, 1 female (MZUSP 74502); in front of Carbet Mitemps, Rain forest, over a wide trunk, 23 March 2015, D. Chirivi, J. Murienne, manual collection at night, 2 male, 1 female (MZUSP 74505); in front of Carbet Mitemps, Rain forest, over a wide trunk, 23 March 2015, D. Chirivi, J. Murienne, manual collection at night, 2 juveniles (MZUSP 74506); in front of Carbet Mitemps, Rain forest, over a wide trunk, 18 March 2015, D. Chirivi, J. Murienne, manual collection at night, male (MZUSP 74508); in front of Carbet Mitemps, Rain forest, over a wide trunk, 22 March 2015, D. Chirivi, J. Murienne, manual collection at night, 2 males, 1 female, 1 juvenile (MZUSP 74509); in front of Saut Maripa, Rain forest, over a wide trunk, 17 March 2015, D. Chirivi, J. Murienne, manual collection at night, 2 males (MZUSP 74510); in front of Saut Maripa, Rain forest, over a wide trunk, 17 March 2015, D. Chirivi, J. Murienne, Manual collection at night, 3 females (MZUSP 74516); in front of Saut Maripa, Rain forest, over a wide trunk, 18 March 2015, D. Chirivi, J. Murienne, manual collection at night, 2 males (MZUSP 74517). Pará: Almeirim: Estação Jari (0 ° 35´91.1 ´´ S; 52 ° 41´72.4 ´´ W): 12 – 19 June 2004, 1 female, 2 males (MPEG 00172); 22 June 2005, T. Gardner, A. Ribeiro, female (MPEG 00171). FRENCH GUIANA: Carbet Mitemps: Oiapoque River (3 ° 17´50,59 ´´ N, 52 ° 10´22,01 ´´ W): Rain forest, over a wide trunk, 20 March 2015, D. Chirivi, J. Murienne, manual collection at night, 1 male, 1 female (MZUSP 74500); Rain forest, over a wide trunk, 25 March 2015, D. Chirivi, J. Murienne, manual collection at night, 1 male, 2 females (MZUSP 74501); Rain forest, over a wide trunk, 24 March 2015, D. Chirivi, J. Murienne, manual collection at night, 1 male, 1 female (MZUSP 74507); Rain forest, over a wide trunk, 24 March 2015, D. Chirivi, J. Murienne, manual collection at night, 1 male, 3 females (MZUSP 74511); Rain forest, over a wide trunk, 26 March 2015, D. Chirivi, J. Murienne, manual collection at night, 1 female (MZUSP 74512); Rain forest, over a wide trunk, 26 March 2015, D. Chirivi, J. Murienne, manual collection at night, 2 males (MZUSP 74514). Saut Maripa: Oiapoque River (3 ° 18´54.8 ´´ N; 52 ° 11´37.5 ´´ W): Rain forest, over a wide trunk, 14 March 2015, D. Chirivi, J. Murienne, manual collection at night, female (MZUSP 74499); Rain forest, over a wide trunk, 14 March 2015, D. Chirivi, J. Murienne, manual collection at night, 2 females (MZUSP 74498); Rain forest, over a wide trunk, 15 March 2015, D. Chirivi, J. Murienne, manual collection at night, 2 males (MZUSP 74503); Rain forest, over a wide trunk, 15 March 2015, D. Chirivi, J. Murienne, manual collection at night, 1 female, 1 male (MZUSP 74504). Trois Palétuviers: Oiapoque River: Rain forest, over a wide trunk, 30 March 2015, D. Chirivi, J. Murienne, manual collection at night, juvenile (MZUSP 74513); Rain forest, over a wide trunk, 29 March 2015, D. Chirivi, J. Murienne, manual collection at night, 1 male, 1 female (MZUSP 74515).	en	Chirivi-Joya, Daniel, Moreno-González, Jairo A., Fagua, Giovanny (2020): Two new species of the whip-spider genus Heterophrynus (Arachnida: Amblypygi) with complementary information of four species. Zootaxa 4803 (1): 1-41, DOI: 10.11646/zootaxa.4803.1.1
03B58796B650FFCCFF7EF987FB507402.taxon	distribution	Distribution: Countries: Brazil, French Guyana, Guyana, Surinam. Other records from: Weygoldt (2002); Weygoldt (1977): Brasil, Amapá, Serra do Navío; Hammen (1986): Surinam, Saramacca river.	en	Chirivi-Joya, Daniel, Moreno-González, Jairo A., Fagua, Giovanny (2020): Two new species of the whip-spider genus Heterophrynus (Arachnida: Amblypygi) with complementary information of four species. Zootaxa 4803 (1): 1-41, DOI: 10.11646/zootaxa.4803.1.1
03B58796B650FFCCFF7EF987FB507402.taxon	diagnosis	Diagnosis. Heterophrynus alces can be differenced from other species by presents the following characters together: three teeth in the ectal portion of the basal segment of the chelicerae. Six long spines in the ventral femur, spine FIII is smaller than FII and FV; it presents a small spine TIII and T 3. Female genitalia with a rounded basal area and a distal portion curved. Seminal receptacles with a sclerotized area, which reach the apical part (Fig. 3 D). The male genital of H. alces differs from other evaluated species by presenting a LoL 1 shorter than the LaM and the Pi. The LoL 2 is smooth and, in some regions, covered by minute projections, while, in the other species the LoL 2 is wrinkled and without projections (Fig. 3 A, B, C). Similar species: No other species present all these characters together, the most similar are H. armiger and H. awa sp. nov., the only character shared with these species is that all present the spine FIII smaller than FII and FV. Complementary description. Female (MZUSP 74499): Total length 23 mm. Carapace, opisthosoma, and legs have a brown coloration, with yellow areas, pedipalps brown and red. Measures of the female are provided in Table 1. Carapace. Anterior margin straight without lobes, carapace surface with a widely spaced granulation, posterior margin of carapace with a concave medial region. Lateral and anteromedial eyes clearly visible, anteromedial ocular tubercle with black coloration (Fig. 1 A). Sternum. Tri-segmented, segments are well sclerotized; the area around segments is also sclerotized. Tritosternum projected anteriorly, elongated, conical, with 16 setae in the basal region, two in the medial region, five in the distal region and two more in the apex. The second segment (tetrasternum) rounded and with six setae; four in the basal region and two in the apical region. The third segment (pentasternum) rounded, smaller than the second segment, has four setae; one on each side of the apical region, and one on each side of the basal region. Metasternum longitudinally divided with two setae in each segment (Fig. 1 B). Abdomen. Oblong, color yellow and brown, with soft punctuations. Presents patches of light coloration at both sides of each tergite. Carapace wider than the Abdomen (Fig. 1 C). Chelicera. Presents a mesal row of three teeth on the basal segment of the chelicerae, with the first in the proximal region, with two cuspids follow by one tooth shorter in the medial portion, and the third bigger than others placed in the distal region (Fig. 1 D). The ectal row of the basal segment of the chelicerae have three conspicuous teeth, two with acuminated apex placed in the proximal and medial portions (tooth 1 and 2), and one with rounded apex placed on the medial portion (tooth 3) (Fig. 1 F). Mobile finger of the chelicerae with five teeth. Pedipalp. Trochanter: Prolateral face with four spines; spines Tr 1 and Tr 3 placed in the medial region, Tr 2 placed near to ventral margin. Tr 3 and Tr 4 with subequal lengths. Spines lengths: Tr 2> Tr 1> Tr 3 ≥ Tr 4. Dorsal oblique series of six setal tubercles. The dorsomedial area without spines but with one tubercle (Fig. 1 E). Femur: Ventral face with six major spines, there are two tubercles between FII – FIII, and FIII – FV, a small spine between FV – FVI, with two tubercles at each side, a small spine between FVI – FVII and three tubercles more distally than FVII. Spine lengths: FI> FII> FV> FVI> FIII> FVII (Fig. 2 A). Dorsal face with five major spines, there are two tubercles between F 3 – F 4, five between F 4 – F 5, and five more distally than F 5, F 1 and F 2 are joined at the base, F 3 clearly separated from F 2. Spine lengths: F 3> F 4> F 1 = F 2> F 5 (Fig. 2 B). Patella: Ventral face with five major spines, there are six tubercles between PV – PIV, one between PIV – PIII, two between PIII – PII and between PII – PI, PI wider than the other spines, PVI absent. Spine lengths: PIV> PV> PII> PI> PIII (Fig. 2 C). Dorsal face with six major spines, two tubercles between P 6 – P 5, P 7 is absent. Spine lengths: P 3> P 4 = P 5> P 1> P 2> P 6 (Fig. 2 D). Tibia: Ventral face with two major spines, and one very small spine TIII, with one tubercle between TII – TIII. Spine lengths: TII> TI> TIII (Fig. 2 E). Dorsal face with two major spines, and one very small spine T 3, with one tubercle between T 2 – T 3. Spine lengths: T 2> T 1> T 3 (Fig. 2 F). Tarsus-metatarsus: Internal face with the secondary line of dorsomedial bristles complete, the suture between the tarsus and the apotele is not visible. Legs. Femora brown. Femora lengths: I> II> III> IV (Table 1). Leg I: Tibia with 31 segments and tarsus with 68 segments in the right and left legs. Leg IV: Basitibia with three segments. Basitibia-distitibia lengths: BT 1> DT> BT 3> BT 2. Basitarsus shorter than telotarsus. Tarsus tetramerous. Female genitalia. Genital operculum pentagonal, reaching the second segment of opisthosoma, seminal receptacles with a rounded shape, completely separated at the medial portion, dorsal surface sclerotized at the proximal and medial region reaching a portion of the distal region, with orange coloration; distal portion white. Sclerites with a rounded base, medial and distal portion curved and thinner than the basal portion, with rounded apex; sclerites placed parallel, shorter than the width of the seminal receptacle, sclerites cover just a half of the total length of the seminal receptacle (Fig. 3 D).	en	Chirivi-Joya, Daniel, Moreno-González, Jairo A., Fagua, Giovanny (2020): Two new species of the whip-spider genus Heterophrynus (Arachnida: Amblypygi) with complementary information of four species. Zootaxa 4803 (1): 1-41, DOI: 10.11646/zootaxa.4803.1.1
03B58796B650FFCCFF7EF987FB507402.taxon	description	Variation. Variation in measures and segments of the basitibia IV, tibia I and tarsus I, are provided in Table 1. Sternum. The number of setae over all segments of the sternum was variable. Pedipalp. Trochanter: line of setal tubercles vary between four and eight. Patella. Ventral face: can be present small spines at each side of PV. Dorsal face: can be present small spines at each side of P 6. Male. Observed Males have a similar size with the females (Table 1), Spination pattern same as females. Genital operculum has an oval posterior margin, expanded to the third segment of the opisthosoma. Male genitalia. (MZUSP 74504): LaM as long as Pi. LoL 1 longer than LoL 2, LoL 1 is wider, densely covered with minute projections, placed close together with the base not visible; LoL 2 smooth and with few projections, the base is visible. In dorsal view, the LoD are rounded covering LoL 1 and LoL 2 (Fig. 3 A, B, C).	en	Chirivi-Joya, Daniel, Moreno-González, Jairo A., Fagua, Giovanny (2020): Two new species of the whip-spider genus Heterophrynus (Arachnida: Amblypygi) with complementary information of four species. Zootaxa 4803 (1): 1-41, DOI: 10.11646/zootaxa.4803.1.1
03B58796B65AFFC8FF7EFD1CFC53755A.taxon	description	Figures 4 – 7, 22 (in part). Table 2	en	Chirivi-Joya, Daniel, Moreno-González, Jairo A., Fagua, Giovanny (2020): Two new species of the whip-spider genus Heterophrynus (Arachnida: Amblypygi) with complementary information of four species. Zootaxa 4803 (1): 1-41, DOI: 10.11646/zootaxa.4803.1.1
03B58796B65AFFC8FF7EFD1CFC53755A.taxon	materials_examined	Type locality: Pambilar, Ecuador. Revised material (2 females, 7 males): Colombia: Cauca: Guapi municipality, Natural National Park (N. N. P.) Gorgona, Isla Gorgona (2 ° 57 ′ 53 ″ N, 78 ° 10 ′ 30 ″ W): Village 5 m, at 13 masl, inside a house, 5 March 1990, M. L. Baena, male (IAvH without number); in a house, April 23 of 1991, J. Portilla, manual collection, male (IAvH-I 2923); between bricks and construction material, 20 October 1991, E. Solis, manual collection, 2 males (IAvH-I 2925, IAvH-I 2924). Sendero Cerro Los Micos (2 ° 58´20,5 ´´ N, 78 ° 10´38,8 ´´ W): 21 October 2010, Zoology team, female (MUSENUV 24303). Playa Palmeras (2 ° 56´28,6 ´´ N, 78 ° 12´21,4 ´´ W): Forest behind of a house, 28 masl, 24 February 2011, 1 male, 1 male juvenile (MUSENUV 24304). El Poblado (2 ° 57´15,8 ´´ N, 78 ° 10´6,6 ´´ W): 2 December 1987, Grupo de Insectos, male (MUSENUV 24307); September 1989, M. Baena, female (MUSENUV 24306).	en	Chirivi-Joya, Daniel, Moreno-González, Jairo A., Fagua, Giovanny (2020): Two new species of the whip-spider genus Heterophrynus (Arachnida: Amblypygi) with complementary information of four species. Zootaxa 4803 (1): 1-41, DOI: 10.11646/zootaxa.4803.1.1
03B58796B65AFFC8FF7EFD1CFC53755A.taxon	distribution	Distribution. Countries: Colombia, Ecuador. Other records from: Víquez et al. (2014): Colombia, Gorgona; Ecuador, Pambilar, Durango River.	en	Chirivi-Joya, Daniel, Moreno-González, Jairo A., Fagua, Giovanny (2020): Two new species of the whip-spider genus Heterophrynus (Arachnida: Amblypygi) with complementary information of four species. Zootaxa 4803 (1): 1-41, DOI: 10.11646/zootaxa.4803.1.1
03B58796B65AFFC8FF7EFD1CFC53755A.taxon	diagnosis	Diagnosis. Heterophrynus armiger can be differentiated from other species by present the following characters together: four teeth in the ectal portion of the basal segment of the chelicerae. Five long spines in the ventral femur, spine FIII is smaller than FII and FV, FIV is absent or like as a tubercle. Female genitalia with gonopod sclerites elongated and with laminar shape, and an acuminated apex. Seminal receptacles oval with a sclerotized area at the base (Fig. 6 D). Similar species: H. armiger is similar to H. alces and H. awa sp. nov., however, H. alces presents a small spine TIII and T 3, which are absent in H. armiger. The female genital is completely different from the one of H. alces, which presents the sclerites of the gonopods with a rounded shape, and H. awa sp. nov., which have sclerites of the gonopods wider at the middle region. The male genital of H. armiger presents a spine in LOL 1, which is not present in any other species (Fig. 7). Complementary description. Male (IAvH-I 2924): Total length 40.2 mm. Carapace, opisthosoma, and legs have a black coloration, with brown areas, pedipalps black. Measures of the male are provided in Table 2. Carapace. Anterior margin straight without lobes, carapace surface with a widely spaced granulation, posterior margin of carapace with two lobes. Lateral and anteromedial eyes clearly visible, anteromedial ocular tubercle with black coloration (Fig. 4 A). Sternum. Tri-segmented, segments are well sclerotized; the area around segments is also sclerotized. Tritosternum projected anteriorly, elongated, conical, with 10 setae in the basal region, four in the medial region two in distal region and two more in the apex. The second segment (tetrasternum) rounded and with four setae; two in the basal region and two in the apical region. The third segment (pentasternum) rounded, smaller than the second segment, has four setae; one on each side of the apical region, one on each side of the basal region. Metasternum longitudinally divided with two setae in each segment (Fig. 4 B). Abdomen. Oblong, color black and brown, with soft punctuations. Carapace wider than the Abdomen (Fig. 4 C). Chelicera. Presents a mesal row of three teeth on the basal segment of the chelicerae with the first in the proximal region, with two cuspids, follow by one tooth shorter in the medial portion, and the third bigger than others placed in the distal region (Fig. 4 D). The ectal row of the basal segment of the chelicerae have four conspicuous teeth, two with acuminated apex placed in the proximal and medial portions (tooth 1 and 2), and two with acuminated apex placed on the medial portion (tooth 3 and 4) (Fig. 4 F). Mobile finger of the chelicerae with five teeth. Pedipalp. Trochanter: Prolateral face with four spines; spines Tr 1 and Tr 3 placed in the medial region, Tr 2 placed near to ventral margin. Tr 3 and Tr 4 with subequal lengths. Spines lengths: Tr 2> Tr 1> Tr 4 ≥ Tr 3. Dorsal oblique series of six setal tubercles. The dorsomedial area without spines but with one tubercle (Fig. 4 E). Femur: Ventral face with five major spines, there are three tubercles between FII – FIII, and FIII – FV, a small spine and 12 tubercles, between FV – FVI, ten tubercles more distally than FVI, FIV absent. Spine lengths: FI> FII> FV> FVI> FIII (Fig. 5 A). Dorsal face with five major spines, there are two tubercles between F 2 – F 3, a very small spine with two tubercles at each side between F 3 – F 4, a very small spine with three tubercles more proximal and two more distally between F 4 – F 5, and more than ten tubercles more distally than F 5. F 1 and F 2 are joined at the base, F 3 clearly separated from F 2. Spine lengths: F 3> F 4> F 1 = F 2> F 5 (Fig. 5 B). Patella: Ventral face with five major spines, there are three tubercles between PV – PIV, three between PIV – PIII, four between PIII – PII and between PII – PI, PI wider than the other spines, PVI is absent. Spine lengths: PIV> PII> PV> PI> PIII (Fig. 5 C). Dorsal face with five major spines, seven tubercles between P 6 – P 5; P 7, and P 2 are absent. Spine lengths: P 3 = P 4 ≥ P 5> P 6> P 1 (Fig. 5 D). Tibia: Ventral face with two major spines. Spine lengths: TII> TI (Fig. 5 E). Dorsal face with two major spines, with one tubercle between T 2 – T 3. Spine lengths: T 2> T 1 (Fig. 5 F). Tarsus-metatarsus: Internal face with the secondary line of dorsomedial bristles complete, the suture between the tarsus and the apotele is not visible. Legs. Femora brown. Femora lengths: I> III> II> IV (Table 2). Leg I: Tibia with 31 segments and tarsus with 70 segments in the right and left leg. Leg IV: Basitibia with three segments. Basitibia-distitibia lengths: BT 1> DT> BT 3> BT 2. Basitarsus shorter than telotarsus. Tarsus tetramerous. Female genitalia. (MUSENUV 24303): Genital operculum pentagonal, reaching the second segment of the opisthosoma, seminal receptacles with an oval shape, completely separated at the medial portion, dorsal surface sclerotized at the proximal region, with brown coloration, medial and distal portion white. Sclerites with laminar shape, distal portion a little curved and thinner than the basal portion, with acuminated apex, sclerites placed parallel, longer than the width of the seminal receptacle, sclerites cover almost the total length of the seminal receptacle (Fig. 6 D).	en	Chirivi-Joya, Daniel, Moreno-González, Jairo A., Fagua, Giovanny (2020): Two new species of the whip-spider genus Heterophrynus (Arachnida: Amblypygi) with complementary information of four species. Zootaxa 4803 (1): 1-41, DOI: 10.11646/zootaxa.4803.1.1
03B58796B65AFFC8FF7EFD1CFC53755A.taxon	description	Variation. Variation in measures and segments of the basitibia IV, tibia I and tarsus I, are provided in Table 2. Sternum. The number of setae over all segments of the sternum was variable. Pedipalp. Trochanter: line of setal tubercles vary between five and six. Femur: Ventral face: number of tubercles between spines are variable, the small spine between FV and FVI, and tubercles can be more conspicuous. In the specimen MUSENUV 24306, there is an FIV with the same size of FIII. Dorsal face: tubercles between spines are variable, can be present a very small spine between F 4 and F 5. Patella. Ventral face: number and size of the tubercles are variable, PVI can be present as a small spine. Dorsal face: number of tubercles are variable, P 2 can be present. Tibia. Ventral and Dorsal face: number of tubercles between spines are variable. Legs. leg I: Number of segments of tibia vary between 31 and 32 and tarsus between 66 and 71 (Table 2). Male. Observed Males were bigger than females (Table 2); Spination pattern same as females. Genital operculum has an oval posterior margin, expanded to the third segment of the opisthosoma. Male genitalia. (IAvH-I 2924): LaM longer than Pi. LoL 1 longer than LoL 2, LoL 1 is wider, densely covered with minute conical projections, placed close together with the base not visible; LoL 2 is wrinkled and does not present projections. In both LoL 1, there is present a spine sticking out from the projections. In dorsal view, the LoD are elongated and do not cover LoL 1 and LoL 2 (Fig. 6 A, B, C; 7 A-B).	en	Chirivi-Joya, Daniel, Moreno-González, Jairo A., Fagua, Giovanny (2020): Two new species of the whip-spider genus Heterophrynus (Arachnida: Amblypygi) with complementary information of four species. Zootaxa 4803 (1): 1-41, DOI: 10.11646/zootaxa.4803.1.1
03B58796B65FFFD6FF7EFC84FB19755A.taxon	description	Figures 8 – 10, 22 (in part). Table 3.	en	Chirivi-Joya, Daniel, Moreno-González, Jairo A., Fagua, Giovanny (2020): Two new species of the whip-spider genus Heterophrynus (Arachnida: Amblypygi) with complementary information of four species. Zootaxa 4803 (1): 1-41, DOI: 10.11646/zootaxa.4803.1.1
03B58796B65FFFD6FF7EFC84FB19755A.taxon	description	Admetus pumilio (C. L. Koch): Kraepelin 1899: 245 – 246 (in part).	en	Chirivi-Joya, Daniel, Moreno-González, Jairo A., Fagua, Giovanny (2020): Two new species of the whip-spider genus Heterophrynus (Arachnida: Amblypygi) with complementary information of four species. Zootaxa 4803 (1): 1-41, DOI: 10.11646/zootaxa.4803.1.1
03B58796B65FFFD6FF7EFC84FB19755A.taxon	materials_examined	Type locality: Georgetown (as Démérara), Demerara-Mahaica, Guyana. Examined material (3 females, 5 males, 1 juvenile): BRASIL: Mato Grosso: 40 km from Sidrolândia, near to Campo Grande, 1 June 1960, Miki, female (MZUSP 9459). Chapada dos Guimarães, Vale do Jamacá (15 º 28´30.3 ´´ S; 55 º 41´21.7 ´´ W): 749 masl, 6 November 2015, J. Cabra, male (DUSP 131). Mato Grosso do Sul: Bonito (21 ° 8 ′ 0 ″ S, 56 ° 29 ′ 0 ″ W): Gruta João Arruja, 18 October 1990, E. Trajano, male (MZUSP 15859); Gruta do Carneiro, 6 - 8 April 1998, R. Pinto-da-Rocha, S. Sessegolo, 2 females (MZUSP 68458). Goias: Santana, November 1945, F. Curado, male (MZUSP 9450). Padre Bernardo (15 ° 9 ′ 36 ″ S, 48 ° 17 ′ 2 ″ W): Gruta qualquer coisa, S. Gregeo, male juvenile (MZUSP 68459). VENEZUELA: Miranda (10 ° 15 ′ 1.72 ″ N, 66 ° 25 ′ 37.74 ″ W): Quebrada de Chamburales, 20 December 2002, A. Perez, A. Giupponi, male (MNRJ 09140).	en	Chirivi-Joya, Daniel, Moreno-González, Jairo A., Fagua, Giovanny (2020): Two new species of the whip-spider genus Heterophrynus (Arachnida: Amblypygi) with complementary information of four species. Zootaxa 4803 (1): 1-41, DOI: 10.11646/zootaxa.4803.1.1
03B58796B65FFFD6FF7EFC84FB19755A.taxon	distribution	Distribution: Countries: Brazil, Guyana, Venezuela. Other records from: Mello-Leitão 1931: mentioned that is recorded from central America but did not mentioned a specific locality, this record do not make sense with known distribution of Heterophrynus for which we consider it as a mistake.	en	Chirivi-Joya, Daniel, Moreno-González, Jairo A., Fagua, Giovanny (2020): Two new species of the whip-spider genus Heterophrynus (Arachnida: Amblypygi) with complementary information of four species. Zootaxa 4803 (1): 1-41, DOI: 10.11646/zootaxa.4803.1.1
03B58796B65FFFD6FF7EFC84FB19755A.taxon	diagnosis	Diagnosis. Heterophrynus cheiracanthus can be differentiated from other species by present the following characters together: three teeth in the ectal portion of the basal segment of the chelicerae. Seven spines in the dorsal Patella, and six in ventral Patella, P 7 and PVI are conspicuous. Five long spines in the ventral femur, spine FIII is smaller than FII and longer than FV, FIV is absent. Female genitalia with gonopod sclerites short, thick and curved at the distal portion. Seminal receptacles with a square shape (Fig. 10 D). Similar species: This species is similar to H. batesii but it presents a FIII longer than FII, also is similar to H. elaphus but this species does not present the spines P 7 and PVI. In both cases, the female genital of H. cheiracanthus is very different (Figs. 13 D, 20 D). H. guacharo is the most similar species, but differs in the genital, H. guacharo presents a more elongated sclerite of the female gonopod, which rises out from a conical seminal receptacle (see Armas 2015, figs 1 - 2). Other similar species could be H. vesanicus which also presents five spines in the ventral femur, however, the original description does not describe the sizes of the spines, the female genital is not described either, the original diagnosis says that the spines on the tibia of the pedipalp have an equal size, which differs from H. cheiracanthus (Mello-Leitão 1931). Male genital of H. cheiracanthus differs from other species by presenting a very elongated and wide LaM, longer than the Pi, and LoL 1 and LoL 2 together (Fig. 10 A, B, C). Complementary description. Female (MZUSP 9459): Total length 21.3 mm. Carapace, opisthosoma, pedipalps, and legs have a brown coloration, with red tonalities. Measures of the female are provided in Table 3. Carapace. Anterior margin straight without lobes, carapace surface with a widely spaced granulation, posterior margin of with a concave medial region. Lateral and anteromedial eyes clearly visible, anteromedial ocular tubercle with black coloration (Fig. 8 A). Sternum. Tri-segmented, segments are well sclerotized, the area around segments is also sclerotized. Tritosternum projected anteriorly, elongated, conical, with 11 setae in the basal region, two in the medial region and two more in the apex. The second segment (tetrasternum) rounded and with two setae on medial region. The third segment (pentasternum) rounded, smaller than the second segment, has four setae, one on each side of the apical region, one on each side of the basal region. Metasternum longitudinally divided with two setae in each segment (Fig. 8 B). Abdomen. Oblong, color brown, with soft punctuations. Carapace wider than the Abdomen (Fig. 8 C). Chelicera. Presents a mesal row of three teeth on the basal segment of the chelicerae with the first in the proximal region, with two cuspids, follow by one tooth shorter in the medial portion, and the third bigger than others placed in the distal region (Fig. 8 D). The ectal row of the basal segment of the chelicerae have three teeth, two with acuminated apex placed in the proximal and medial portions (tooth 1 and 2), and one smaller with rounded apex placed on the medial portion (tooth 3) (Fig. 8 F). Mobile finger of the chelicerae with four teeth. Pedipalp. Trochanter: Prolateral face with four spines; spines Tr 1 and Tr 3 placed in the medial region, Tr 2 placed near to ventral margin. Spines lengths: Tr 2> Tr 3> Tr 4 ≥ Tr 1. Dorsal oblique series of 11 setal tubercles. The dorsomedial area without spines but with one tubercle (Fig. 8 E). Femur: Ventral face with five major spines, FIV absent. Spine lengths: FI> FII> FIII> FV> FVI (Fig. 9 A). Dorsal face with six major spines. Spines F 1 and F 2 share the same base and are clearly separated from F 3. Spine lengths: F 3> F 4> F 1 = F 2> F 5> 6 (Fig. 9 B). Patella: Ventral face with six major spines, PI wider than the other spines. Spine lengths: PIII> PIV> PI> PII> PV> PVI (Fig. 9 C). Dorsal face with seven major spines. Spine lengths: P 5> P 4 = P 3> P 6> P 1> P 7> P 2 (Fig. 9 D). Tibia: Ventral face with two major spines. Spine lengths: TII> TI (Fig. 9 E). Dorsal face with two major spines. Spine lengths: T 2> T 1 (Fig. 9 F). Tarsus-metatarsus: Internal face with the secondary line of dorsomedial bristles complete, the suture between the tarsus and the apotele is not visible. Legs. Femora brown and red. Femora lengths: I> III> II> IV (Table 3). Leg I: Tibia with 39 segments and tarsus with 75 segments in the right leg; left leg is missed. Leg IV: Basitibia with three segments. Basitibia-distitibia lengths: BT 1> DT> BT 3> BT 2. Basitarsus shorter than telotarsus. Tarsus tetramerous. Female genitalia. Genital operculum pentagonal, reaching the second segment of the opisthosoma, seminal receptacles with a square shape, joined at the medial portion, dorsal surface sclerotized at the proximal portion, with brown coloration, medial and distal portion white with some brown areas. Sclerites of the gonopod short and thick, with claw-like shape, distal portion a little curved, with rounded apex, sclerites placed parallel, shorter than the width of the seminal receptacle, sclerites cover almost the total length of the seminal receptacle (Fig. 10 D).	en	Chirivi-Joya, Daniel, Moreno-González, Jairo A., Fagua, Giovanny (2020): Two new species of the whip-spider genus Heterophrynus (Arachnida: Amblypygi) with complementary information of four species. Zootaxa 4803 (1): 1-41, DOI: 10.11646/zootaxa.4803.1.1
03B58796B65FFFD6FF7EFC84FB19755A.taxon	description	Variation. Variation in measures and segments of the basitibia IV, tibia I and tarsus I, are provided in Table 3. Sternum. The number of setae over all segments of the sternum was variable. Pedipalp. Trochanter: line of setal tubercles vary between four and eight. Patella. Dorsal face: relative size between P 4 and P 3 is variable it can be almost equal. Legs. In leg I, the number of segments of the tibia vary between 33 and 39 and tarsus vary between 69 and 82. The relative sizes of the femora II, III and IV is variable. (Table 3). Male. Observed males have a similar size to the females (Table 3). Spination pattern is the same as females. Genital operculum has an oval posterior margin, expanded to the third segment of the opisthosoma. Male genitalia. (MZUSP 15859): LaM longer than Pi. LoL 1 longer than LoL 2, LoL 1 is wider, densely covered with minute projections, placed close together with the base not visible; LoL 2 is wrinkled does not present projections. In dorsal view, the LoD are elongated, rounded and do not cover LoL 1 (Fig. 10 A, B, C).	en	Chirivi-Joya, Daniel, Moreno-González, Jairo A., Fagua, Giovanny (2020): Two new species of the whip-spider genus Heterophrynus (Arachnida: Amblypygi) with complementary information of four species. Zootaxa 4803 (1): 1-41, DOI: 10.11646/zootaxa.4803.1.1
03B58796B640FFD2FF7EFC24FF5D75E2.taxon	description	Figures 11 – 13, 22 (in part). Table 4.	en	Chirivi-Joya, Daniel, Moreno-González, Jairo A., Fagua, Giovanny (2020): Two new species of the whip-spider genus Heterophrynus (Arachnida: Amblypygi) with complementary information of four species. Zootaxa 4803 (1): 1-41, DOI: 10.11646/zootaxa.4803.1.1
03B58796B640FFD2FF7EFC24FF5D75E2.taxon	materials_examined	Type locality: Marcapata Valley, Cuzco, Perú. Examined material (1 female, 1 male, 1 juvenil): PERÚ: Pasco: Parque Nacional Yanachaga-Chemillén, Paujil (10 ° 19´22.5 ´´ S; 75 ° 15´44.7 ´´ W): 360 masl, 1 October 2015, J. Cabra, B. Portuguéz, three males, 1 female, 1 male juvenile (MZUSP 74571).	en	Chirivi-Joya, Daniel, Moreno-González, Jairo A., Fagua, Giovanny (2020): Two new species of the whip-spider genus Heterophrynus (Arachnida: Amblypygi) with complementary information of four species. Zootaxa 4803 (1): 1-41, DOI: 10.11646/zootaxa.4803.1.1
03B58796B640FFD2FF7EFC24FF5D75E2.taxon	distribution	Distribution: Countries: Bolivia, Perú. Other records from: Peru: Igelmund & Wendler (1991 a): Pucallpa; Roewer (1952): Tingo María; Roewer (1957): Cascas, near to Trujillo; La Merced; Saposoa; Puente Cayumba: Rio Huallaga; Tingo Maria: Cueva de Las Lechuzas; Ballón-Estacio & Armas (2019): Cusco, Madre de Dios. Bolivia: Réveillion et al. (2014): Beni.	en	Chirivi-Joya, Daniel, Moreno-González, Jairo A., Fagua, Giovanny (2020): Two new species of the whip-spider genus Heterophrynus (Arachnida: Amblypygi) with complementary information of four species. Zootaxa 4803 (1): 1-41, DOI: 10.11646/zootaxa.4803.1.1
03B58796B640FFD2FF7EFC24FF5D75E2.taxon	diagnosis	Diagnosis. Heterophrynus elaphus can be differentiated from other species by present the following characters together: three teeth in the ectal portion of the basal segment of the chelicerae. Six spines in the dorsal Patella, P 7 and PIII are absent. Four long spines in the ventral femur, spine FIII is smaller than FII and longer than FV, FIV is absent. Female genitalia with gonopod sclerites with claw-like shape, a little curved at the distal portion. Seminal receptacles with a square shape (Fig. 13 D). Similar species: the more similar species is H. cheiracanthus, however, this species present the spines P 7, and PIII. The female genital differs by presenting gonopod sclerites shorter and more curved (Fig. 10 D). Male genital of H. elaphus presents a Pi, LaM, and LoL 1 with a very similar length and an LoD very elongated and wide, which differs from the other species (Fig. 13, A, B, C). Complementary description. Female (MZUSP 74571): Total length 32.4 mm. Carapace, opisthosoma, and legs have a brown coloration, pedipalps brown with red tonalities. Measures of the female are provided in Table 4. Carapace. Anterior margin straight without lobes, carapace surface with a widely spaced granulation, posterior margin of with a concave medial region. Lateral and anteromedial eyes clearly visible, anteromedial ocular tubercle with black coloration (Fig. 11 A). Sternum. Tri-segmented, segments are well sclerotized, the area around segments is also sclerotized. Tritosternum projected anteriorly, elongated, conical, with 30 setae in the basal region, eighth in the medial region, 11 in distal region and two more in the apex. The second segment (tetrasternum) oval and with three setae on medial region. The third segment (penta- sternum) rounded, smaller than the second segment, has two setae on medial region. Metasternum longitudinally divided (Fig. 11 B). Abdomen. Oblong, color brown, with soft punctuations. Carapace wider than the Abdomen (Fig. 11 C). Chelicera. Presents a mesal row of three teeth on the basal segment of the chelicerae with the first in the proximal region, with two cuspids, follow by one tooth shorter in the medial portion, and the third bigger than others placed in the distal region (Fig. 11 D). The ectal row of the basal segment of the chelicerae have three teeth, two with acuminated apex placed in the proximal portions, fused at the base (tooth 1 and 2), and one more with acuminate apex placed on the medial portion (tooth 3) (Fig. 11 F). Mobile finger of the chelicerae with five teeth. Pedipalp. Trochanter: Prolateral face with four spines; spines Tr 1 and Tr 3 placed in the medial region, Tr 2 placed near to ventral margin. Tr 3 and Tr 4 with subequal lengths. Spines lengths: Tr 2> Tr 1> Tr 3 ≥ Tr 4. Dorsal oblique series of nine setal tubercles. The dorsomedial area without spines but with three tubercles (Fig. 11 E). Femur: Ventral face with four major spines, with four tubercles between FII – FIII, seven between FIII – FV, and 11 more distally than FV, FIV is absent. Spine lengths: FI> FII> FIII> FV (Fig. 12 A). Dorsal face with five major spines, with four tubercles between F 3 – F 4, five between F 4 – F 5, and 11 more distally than F 5. F 1 and F 2 share the same base and are clearly separated from F 3. Spine lengths: F 3> F 4> F 1 = F 2> F 5 (Fig. 12 B). Patella: Ventral face with four major spines, PVI and PIII are absent, six tubercles more proximally than PV, four between PV – PIV, six between PIV – PII, five between PII – PI, PI wider than the other spines. Spine lengths: PII> PIV> PV> PI (Fig. 12 C). Dorsal face with six major spines, P 7 is absent, with six tubercles more proximally than P 6 and five between P 6 and P 5. Spine lengths: P 5 = P 4> P 3> P 6> P 1> P 2 (Fig. 12 D). Tibia: Ventral face with two major spines, five very small tubercles more distally than TII. Spine lengths: TII> TI (Fig. 12 E). Dorsal face with two major spines, six very small tubercles more distally than T 2. Spine lengths: T 2> T 1 (Fig. 12 F). Tarsus-metatarsus: Internal face with the secondary line of dorsomedial bristles complete, the suture between the tarsus and the apotele is not visible. Legs. Femora brown. Femora lengths: I> II> III> IV (Table 4). Leg I: Tibia with 55 segments and tarsus with 97 segments in the right leg; in left leg, tibia with 26 segments and tarsus is broken. Leg IV: Basitibia with three segments. Basitibia-distitibia lengths: BT 1> DT> BT 3> BT 2. Basitarsus shorter than the telotarsus. Tarsus tetramerous. Female genitalia. Genital operculum pentagonal, reaching the second segment of the opisthosoma, seminal receptacles with a rectangular shape, separated at the medial portion, dorsal surface sclerotized at the proximal region and with an additional sclerotized layer around the base of the sclerite of the gonopod, basal region with orange coloration, medial and distal portion white. Sclerites of the gonopod with claw-like shape, basal portion with some transversal grooves, distal portion a little curved, with rounded apex, sclerites placed parallel, longer than the width of the seminal receptacle, sclerites cover almost the total length of the seminal receptacle (Fig. 13 D).	en	Chirivi-Joya, Daniel, Moreno-González, Jairo A., Fagua, Giovanny (2020): Two new species of the whip-spider genus Heterophrynus (Arachnida: Amblypygi) with complementary information of four species. Zootaxa 4803 (1): 1-41, DOI: 10.11646/zootaxa.4803.1.1
03B58796B640FFD2FF7EFC24FF5D75E2.taxon	description	Variation. Variation in measures and segments of the basitibia IV, tibia I and tarsus I, are provided in Table 4. Sternum. The number of setae over all segments of the sternum was variable. Pedipalp. Trochanter: line of setal tubercles vary between seven and nine. Femur: Ventral and Dorsal face: number of tubercles between the spines is variable; tubercles can be more conspicuous and, in some cases, can look as small spines. Patella. Ventral face: can be present a very small spine PVI, number of tubercles is variable. Dorsal face: number and size of the tubercles are variable. Tibia. Ventral and Dorsal face: tubercles between spines can be more conspicuous. Legs. In leg I, the number of segments of the tibia vary between 26 and 65 and tarsus vary between 97 and 103. The relative sizes of the femora II, III and IV is variable. (Table 4). Male. Observed Males have a similar size than the female (Table 4), Spination pattern same as females. Genital operculum has an oval posterior margin, expanded to the third segment of the opisthosoma. Male genitalia. (MZUSP 74571): LaM with similar size to Pi. LoL 1 longer than LoL 2, LoL 1 is wider, densely covered with minute projections, placed close together with the base not visible; LoL 2 is wrinkled and does not present projections. In dorsal view, the LoD are elongated, wide, rounded and cover LoL 1 and LoL 2 (Fig. 13 A, B, C).	en	Chirivi-Joya, Daniel, Moreno-González, Jairo A., Fagua, Giovanny (2020): Two new species of the whip-spider genus Heterophrynus (Arachnida: Amblypygi) with complementary information of four species. Zootaxa 4803 (1): 1-41, DOI: 10.11646/zootaxa.4803.1.1
03B58796B645FFDEFF7EFF30FAA27492.taxon	description	Figures 14 – 16, 22 (in part). Table 5.	en	Chirivi-Joya, Daniel, Moreno-González, Jairo A., Fagua, Giovanny (2020): Two new species of the whip-spider genus Heterophrynus (Arachnida: Amblypygi) with complementary information of four species. Zootaxa 4803 (1): 1-41, DOI: 10.11646/zootaxa.4803.1.1
03B58796B645FFDEFF7EFF30FAA27492.taxon	materials_examined	Type locality: Colombia, Nariño. Type material: Holotype (male): COLOMBIA: Nariño: Barbacoas: km 95 via Tumaco-Pasto, Reserva Natural Biotopo (1 ° 23´00 ´´ N; 78 ° 15´00 ´´ W): Rain Forest, 500 masl, 14 September 2009, D, Martinez, H. Herreño, manual collection, male (ICN-Am- 108). Paratypes (1 female, 1 male): COLOMBIA: Nariño: Barbacoas: km 95 via Tumaco-Pasto, Reserva Natural Biotopo (1 ° 23´00 ´´ N; 78 ° 15´00 ´´ W): Rain Forest, 500 masl, 14 September 2009, D, Martinez, H. Herreño, manual collection, male (ICN-Am- 108); Vereda Altaquer (1 ° 14´45.60 ´´ N, 78 ° 06´55.12 ´´ W): on the ground, April 15 of 2010, 1270 masl, manual collection, female (ICN-Am- 75).	en	Chirivi-Joya, Daniel, Moreno-González, Jairo A., Fagua, Giovanny (2020): Two new species of the whip-spider genus Heterophrynus (Arachnida: Amblypygi) with complementary information of four species. Zootaxa 4803 (1): 1-41, DOI: 10.11646/zootaxa.4803.1.1
03B58796B645FFDEFF7EFF30FAA27492.taxon	distribution	Distribution: Colombia, Nariño.	en	Chirivi-Joya, Daniel, Moreno-González, Jairo A., Fagua, Giovanny (2020): Two new species of the whip-spider genus Heterophrynus (Arachnida: Amblypygi) with complementary information of four species. Zootaxa 4803 (1): 1-41, DOI: 10.11646/zootaxa.4803.1.1
03B58796B645FFDEFF7EFF30FAA27492.taxon	etymology	Etymology: This species is dedicated to the Colombian native community Awá (Kwaiker or Awa-Kwaiker), which inhabit between the boundaries of Colombia and Ecuador, and some they settle in the type locality (Nariño). We consider that this species deserves to be considered as part of his natural richness. Name in apposition.	en	Chirivi-Joya, Daniel, Moreno-González, Jairo A., Fagua, Giovanny (2020): Two new species of the whip-spider genus Heterophrynus (Arachnida: Amblypygi) with complementary information of four species. Zootaxa 4803 (1): 1-41, DOI: 10.11646/zootaxa.4803.1.1
03B58796B645FFDEFF7EFF30FAA27492.taxon	diagnosis	Diagnosis. Heterophrynus awa sp. nov. can be differentiated from other species by present the following characters together: four teeth in the ectal portion of the basal segment of the chelicerae. Five long spines in the ventral femur, spine FIII is smaller than FII and FV, FIV is small, presents a small spine PVI. Female genitalia with gonopod sclerites elongated and with laminar shape, a medial portion wider than the basal and distal portion, distal portion thick and with a rounded apex. Seminal receptacles oval with a sclerotized area at the base (Fig. 16 D). Similar species: This species is similar to H. alces but it presents a small spine TIII and T 3, which is absent in H. awa sp. nov. and the female genitalia is completely different, presenting the sclerites of the gonopods with a rounded shape. The species H. armiger is very similar, however, it presents female genitalia with sclerites not widened at the medial portion, and with an acuminated apex (Fig. 6 D). The male genital of H. armiger and H. awa sp. nov, presents Pi rounded and so smaller than the LaM, the LoL 1 has a similar length with the LaM. Both species differ because H. armiger presents a spine on LOL 1 which is absent in H. awa sp. nov (Fig. 6 A, B, C; 7; 16 A, B, C).	en	Chirivi-Joya, Daniel, Moreno-González, Jairo A., Fagua, Giovanny (2020): Two new species of the whip-spider genus Heterophrynus (Arachnida: Amblypygi) with complementary information of four species. Zootaxa 4803 (1): 1-41, DOI: 10.11646/zootaxa.4803.1.1
03B58796B645FFDEFF7EFF30FAA27492.taxon	description	Description. Male (ICN-Am- 108): Total length 20.9 mm. Carapace, opisthosoma, and legs have a brown coloration, with red tonalities, pedipalps brown. Measures of the male are provided in Table 5. Carapace. Anterior margin straight without lobes, carapace surface with a widely spaced granulation, posterior margin with a concave area in the medial region. Lateral and anteromedial eyes clearly visible, anteromedial ocular tubercle with black coloration (Fig. 14 A). Sternum. Tri-segmented, segments are well sclerotized, the area around segments is also sclerotized. Tritosternum projected anteriorly, elongated, conical, with more than 10 setae in the basal region, five in the medial region three in distal region and two more in the apex. The second segment (tetrasternum) rounded and with four setae; two on the medial region and two on the apical region. The third segment (pentasternum) rounded, smaller than the second segment, has two setae on medial region. Metasternum longitudinally divided with two setae in each segment (Fig. 14 B). Abdomen. Oblong, color brown, with soft punctuations, and darker areas in the medial area of the sclerites. Carapace wider than the Abdomen (Fig. 14 C). Chelicera. Presents a mesal row of three teeth on the basal segment of the chelicerae with the first in the proximal region, with two cuspids, follow by one tooth shorter in the medial portion, and the third bigger than others placed in the distal region (Fig. 14 D). The ectal row of the basal segment of the chelicerae have four conspicuous teeth, two with acuminated apex placed in the proximal and medial portions (tooth 1 and 2), and two with acuminated apex placed on the medial portion (tooth 3 and 4) (Fig. 14 F). Mobile finger of the chelicerae with five teeth. Pedipalp. Trochanter: Prolateral face with four spines; spines Tr 1 and Tr 3 placed in the medial region, Tr 2 placed near to ventral margin. Tr 3 and Tr 4 with subequal lengths. Spines lengths: Tr 2> Tr 1> Tr 4 ≥ Tr 3. Dorsal oblique series of six setal tubercles. The dorsomedial area without spines but with one tubercle (Fig. 14 E). Femur: Ventral face with five major spines, there are three tubercles between FII – FIII, one small spine FIV with two tubercles at each side between FIII – FV, a small spine with three tubercles at each side between FV – FVI, ten tubercles more distally than FVI, FIV has almost the size of a tubercle. Spine lengths: FI> FII> FV> FVI> FIII> FIV (Fig. 15 A). Dorsal face with five major spines, there are three tubercles between F 2 – F 3, a very small spine and four tubercles between F 3 – F 4, two very small spines and three tubercles between F 4 – F 5, and eight tubercles more distally than F 5, F 1 and F 2 are joined at the base, F 3 clearly separated from F 2. Spine lengths: F 3> F 4> F 1 = F 2> F 5 (Fig. 15 B). Patella: Ventral face with six major spines, presents a small PVI, there are three tubercles more proximally than PVI, one small spine and three tubercles between PVI – PV, four tubercles between PV – PIV, and PIV – PIII, three tubercles between PIII – PII and five between PII – PI, PI wider than the other spines. Spine lengths: PIV> PII> PV> PI> PIII> PVI (Fig. 15 C). Dorsal face with six major spines, seven tubercles between P 6 – P 5, P 7 is absent. Spine lengths: P 3 = P 4 ≥ P 5> P 6> P 1> P 2 (Fig. 15 D). Tibia: Ventral face with two major spines. Spine lengths: TII> TI (Fig. 15 E). Dorsal face with two major spines. Spine lengths: T 2> T 1 (Fig. 15 F). Tarsus-metatarsus: Internal face with the secondary line of dorsomedial bristles complete, the suture between the tarsus and the apotele is not visible. Legs. Femora brown. Femora lengths: I> III> II> IV (Table 5). Leg I: Tibia with 31 segments and tarsus with 70 segments in the right and left leg. Leg IV: Basitibia with three segments. Basitibia-distitibia lengths: BT 1> DT> BT 3> BT 2. Basitarsus as long as the telotarsus. Tarsus tetramerous. Female genitalia (ICN-Am- 75): Genital operculum pentagonal, reaching the second segment of the opisthosoma, seminal receptacles with an oval shape, completely separated at the medial portion, dorsal surface sclerotized at the proximal region, with brown coloration, medial and distal portion white. Sclerites with laminar shape, the medial region is wider than the basal and distal region, distal portion a little curved and thinner than the basal portion, with rounded apex, sclerites placed parallel, longer than the width of the seminal receptacle, sclerites longer than the total length of the seminal receptacle (Fig. 16 D). Variation. Variation in measures and segments of the basitibia IV, Tibia I and tarsus I, are provided in Table 5. Sternum. The number of setae over all segments of the sternum was variable. Pedipalp. Trochanter: line of setal tubercles vary between five and six. Femur: Ventral face: FIV can be present as a spine or tubercle, number of tubercles between spines is variable, and can be more conspicuous. Dorsal face: number of tubercles between spines is variable, tubercles can be more conspicuous. Patella. Ventral face: relative size between PI and PIII is variable, number and size of small spines and tubercles between major spines are variable. Dorsal face: number of tubercles are variable. Legs. Leg I: Number of segments of tibia vary between 31 and 33 and in the tarsus between 66 and 76, the relative size of femora is variable (Table 5). Male. Observed Males were bigger than females (Table 5). Spination pattern same as females. Genital operculum has an oval posterior margin, expanded to the third segment of the opisthosoma. Male genitalia. (ICN-Am- 108): LaM longer than Pi. LoL 1 longer than LoL 2, LoL 1 is wider, densely covered with minute projections, placed close together with the base not visible; LoL 2 is wrinkled and does not present projections. In dorsal view, the LoD are, long, rounded and do not cover LoL 1 and LoL 2 (Fig. 16 A, B, C).	en	Chirivi-Joya, Daniel, Moreno-González, Jairo A., Fagua, Giovanny (2020): Two new species of the whip-spider genus Heterophrynus (Arachnida: Amblypygi) with complementary information of four species. Zootaxa 4803 (1): 1-41, DOI: 10.11646/zootaxa.4803.1.1
03B58796B649FFE7FF7EFABEFAA17722.taxon	description	Figures 17 – 19, 22 (in part). Table 6.	en	Chirivi-Joya, Daniel, Moreno-González, Jairo A., Fagua, Giovanny (2020): Two new species of the whip-spider genus Heterophrynus (Arachnida: Amblypygi) with complementary information of four species. Zootaxa 4803 (1): 1-41, DOI: 10.11646/zootaxa.4803.1.1
03B58796B649FFE7FF7EFABEFAA17722.taxon	materials_examined	Type locality: Brazil, Rondônia, Porto Velho. Type material: Holotype (female): BRAZIL: Rondônia: Porto Velho: ESEC-Cuniã (8 ° 7´31 ´´ S, 63 ° 3´3 ´´ W): 14 February 2005, female (MZUSP 68451). Paratypes (13 females, 9 males, 2 juveniles): BRAZIL: Pará: Santarem (2 ° 26´34 ″ S, 54 ° 42´28 ´´ W): Anumã, 30 November 2008, Yamaguti, Pinto da Rocha, 1 male, 1 female (MZUSP 68457). Rondônia: Porto Velho: ESEC-Cuniã (8 ° 7´31 ´´ S, 63 ° 3´3 ´´ W): 14 February 2005, 1 female, 1 male (MZUSP 68451); 13 February 2005, 1 female, 1 juvenile (MZUSP 68453). Abunã (9 ° 41´41.91 ´´ S, 65 ° 21´23.90 ´´ W): 23 May 2013, A. Nogueira, male (MZUSP 54473); 2050 masl, 23 May 2013, A. Nogueira, male (MZUSP 54473); 1050 masl, 24 May 2013, S. Outeda-Jorge, manual collection at night, male (MZUSP 54589); 1050 masl, 25 May 2013, A. Nogueira, female (MZUSP 55979); 2050 masl, 24 May 2013, A. Nogueira, male juvenile (MZUSP 55848); 22 August 2011 M. C. Silveira, female (MZUSP 39834); 22 August 2011 M. C. Silveira, female (MZUSP 42301). Mutum (9 ° 28´52.536 ´´ S, 64 ° 52´6.492 ´´ W): 200 masl, 20 May 2013, S. Outeda-Jorge, manual collection at night, female (MZUSP 55797); 1300 masl, 21 May 2013, S. Outeda-Jorge, manual collection at night, male (MZUSP 54820); 200 masl, 20 May 2013, S. Outeda-Jorge, manual collection at night, male (MZUSP 55798); 400 masl, 20 May 2013, A. Nogueira, manual collection at night, female (MZUSP 55827). Alto paraíso (9 ° 42´47 ´´ S, 63 ° 19´15 ´´ W): 23 - 26 December 1983, female (MZUSP 68455); 23 - 26 December 1983, female (MZUSP 68456). Caiçara (9 ° 26´21.03 ´´ S; 64 ° 47´32.16 ´´ W): 24 October 2010, S. Outeda-Jorge, M. C. Silveira, R. Y. Lemos, G. P. Perroni, male (MZUSP 35370); 2 September 2011, M. C. Silveira, female (MZUSP 39093); 13 November 2011, M. C. Silveira, juvenile (MZUSP 44068); 23 April 2012, R. Indicatti, female (MZUSP 47611). COLOMBIA: Caquetá: Rio Yamilla (0 ° 1´33.03 ´´ S; 72 ° 6´38.87 ´´ W): flooded forest, at the base of a three, Manual collection at night, 26 November 2010, A. Sánchez, female (MPUJ _ ENT 0007616).	en	Chirivi-Joya, Daniel, Moreno-González, Jairo A., Fagua, Giovanny (2020): Two new species of the whip-spider genus Heterophrynus (Arachnida: Amblypygi) with complementary information of four species. Zootaxa 4803 (1): 1-41, DOI: 10.11646/zootaxa.4803.1.1
03B58796B649FFE7FF7EFABEFAA17722.taxon	distribution	Distribution: Colombia, Brazil.	en	Chirivi-Joya, Daniel, Moreno-González, Jairo A., Fagua, Giovanny (2020): Two new species of the whip-spider genus Heterophrynus (Arachnida: Amblypygi) with complementary information of four species. Zootaxa 4803 (1): 1-41, DOI: 10.11646/zootaxa.4803.1.1
03B58796B649FFE7FF7EFABEFAA17722.taxon	etymology	Etymology: This species is dedicated to the biologist Andrés Sánchez also known as “ Origami ”, which collected the first exemplar of this species in Colombia. Besides being an excellent entomologist, and friend, he is a virtuous in the art of origami, to which he owes his pseudonym.	en	Chirivi-Joya, Daniel, Moreno-González, Jairo A., Fagua, Giovanny (2020): Two new species of the whip-spider genus Heterophrynus (Arachnida: Amblypygi) with complementary information of four species. Zootaxa 4803 (1): 1-41, DOI: 10.11646/zootaxa.4803.1.1
03B58796B649FFE7FF7EFABEFAA17722.taxon	diagnosis	Diagnosis. Heterophrynus origamii sp. nov. can be differentiated from other species by present the following characters together: three teeth in the ectal portion of the basal segment of the chelicerae. Five long spines in the ventral femur, spine FII is smaller than FIII and FI, FIII longer than FV, FIV is absent. Female genitalia with a sclerite of the gonopod elongated with a medial portion wider, a constriction between the medial and distal portion, apex rounded and wider than the basal portion. Seminal receptacles are rounded, with a sclerotized area that covers only the basal part (Fig. 19 D). Similar species: This species is similar to H. batesii, H. caribensis and H. longicornis, this three species, together with H. origamii sp. nov., present almost the same pattern of spines in the pedipalp; however, the female genital has visible differences, in H. batesii and H. caribensis the sclerite of the gonopod is laminar and uniform, without a wider medial portion, or constrictions near to the apex. Almost in all adult specimens of H. batesii and H. caribensis, the sclerite is completely red without white regions, in contrast with H. origamii sp. nov. Seminal receptacles are more rectangular and elongated in H. batesii and H. caribensis, as well as the sclerites, are longer (Fig. 20 D). Heterophrynus longicornis has a sclerite of the female gonopod with a base more than two times wider than the medial and distal portion and presents a very strong curvature at the medial portion (Fig. 21 D), this sclerite is completely different with the one of H. origamii sp. nov. The male genital of H. batesii, H. caribensis, H. longicornis and H. origami sp. nov. are characterized by having a Pi thin and elongated, but smaller than the LaM; the LaM has a similar length with the of LoL 1 and 2 together (Fig. 19 A, B, C; 20 A, B, C; 21 A, B, C); it differs from the other evaluated species, however, male genital of this four species is almost equal.	en	Chirivi-Joya, Daniel, Moreno-González, Jairo A., Fagua, Giovanny (2020): Two new species of the whip-spider genus Heterophrynus (Arachnida: Amblypygi) with complementary information of four species. Zootaxa 4803 (1): 1-41, DOI: 10.11646/zootaxa.4803.1.1
03B58796B649FFE7FF7EFABEFAA17722.taxon	description	Description. Female (MZUSP 68451): Total length 18.5 mm. Carapace, opisthosoma, pedipalps, and legs have a brown coloration, with red tonalities. Measures of the female are provided in Table 6. Posterior margin of carapace with a concave medial region. Lateral and anteromedial eyes clearly visible, anteromedial ocular tubercle with black coloration (Fig. 17 A). Sternum. Tri-segmented, segments are well sclerotized, the area around segments is also sclerotized. Tritosternum projected anteriorly, elongated, conical, with 24 setae in the basal region, two in the medial region, two in distal region and two more in the apex. The second segment (tetrasternum) rounded and with four setae, two in the basal region and two in the medial region. The third segment (pentasternum) rounded, smaller than the second segment, has two setae on medial region. Metasternum longitudinally divided with two setae in each segment (Fig. 17 B). Abdomen. Oblong, color brown with red tonalities, with soft punctuations. Carapace wider than the Abdomen (Fig. 17 C). Chelicera. Presents a mesal row of three teeth on the basal segment of the chelicerae with the first in the proximal region, with two cuspids, followed by one tooth shorter in the medial portion, and the third bigger than others placed in the distal region (Fig. 17 D). The ectal row of the basal segment of the chelicerae have three conspicuous teeth, two with acuminated apex placed in the proximal and medial portions (tooth 1 and 2), and one with acuminated apex placed on the medial portion (tooth 3) (Fig. 17 F). Mobile finger of the chelicerae with four teeth. Pedipalp. Trochanter: Prolateral face with four spines; spines Tr 1 and Tr 3 placed in the medial region, Tr 2 placed near to ventral margin. Spines lengths: Tr 2> Tr 4> Tr 3 ≥ Tr 1. Dorsal oblique series of seven setal tubercles. The dorsomedial area without spines but with one tubercle (Fig. 17 E). Femur: Ventral face with five major spines, spine FIV is absent, there are three tubercles between FI – FII, four between FII – FIII, five between FIII – FV, eight tubercles between FV – FVI, and six tubercles more distally than FVI. Spine lengths: FI> FIII> FII> FV> FVI (Fig. 18 A). Dorsal face with six major spines, there are three tubercles between F 2 – F 3 and F 3 – F 4, five between F 4 – F 5 and F 5 – F 6, and six more distally than F 6, F 1 and F 2 are joined at the base, F 3 clearly separated from F 2. Spine lengths: F 3> F 4> F 1 = F 2> F 5> F 6 (Fig. 18 B). Patella: Ventral face with six major spines, there are two tubercles more proximally than PVI, three between PVI – PV, five between PV – PIV, four between PIV – PIII, three between PIII – PII, and three between PII – PI, PI wider than the other spines. Spine lengths: PIII> PIV> PII> PV> PI> PVI (Fig. 18 C). Dorsal face with seven major spines, four tubercles more proximally than P 7, six between P 7 – P 6, five between P 6 – P 5, four between P 5 – P 4, and P 4 – P 3. Spine lengths: P 3 = P 4> P 5> P 6> P 1> P 2> P 7 (Fig. 18 D). Tibia: Ventral face with two major spines, with one tubercle between TI – TII. Spine lengths: TII> TI (Fig. 18 E). Dorsal face with two major spines. Spine lengths: T 2> T 1 (Fig. 18 F). Tarsus-metatarsus: Internal face with the secondary line of dorsomedial bristles complete, the suture between the tarsus and the apotele is not visible. Legs. Femora brown. Femora lengths: I> II> III> IV (Table 6). Leg I: Tibia with 39 segments and tarsus with 89 segments in the right leg; in left leg ha 39 segments in the tibia and 88 segments in the tarsus. Leg IV: Basitibia with three segments. Basitibia-distitibia lengths: BT 1> DT> BT 3 = BT 2. Basitarsus as long as the telotarsus. Tarsus tetramerous. Female genitalia. Genital operculum pentagonal, reaching the second segment of the opisthosoma, seminal receptacles with a rounded shape, completely touching to each other at the medial portion, dorsal surface sclerotized only at the proximal portion, with orange coloration, medial and distal portion white. Sclerites elongated with a medial and distal portion wide, present a constriction between medial and distal portion, distal portion wider than the basal portion, with rounded apex, sclerites placed opposite to each other, shorter than the width of the seminal receptacle, the sclerite of the gonopod cover the half of the length of the seminal receptacle, present a brown-orange coloration with a white area at the base (Fig. 19 D). Variation. Variation in measures and segments of the basitibia IV, Tibia I and tarsus I, are provided in Table 6. Sternum. The number of setae over all segments of the sternum was variable. Pedipalp. Trochanter: line of setal tubercles vary between six and eight. Femur: Ventral face: more distally than FVI can be present one or two spines so much smaller than the others are. Dorsal face: spine F 6 can be more conspicuous. Patella. Dorsal face: P 7 can be as long as P 2. Female genital. in the sclerites of the gonopod, the brown coloration and white coloration is variable, in some cases brown coloration can cover a bigger or smaller portion of the sclerite. Chelicerae. Mobile segment: the number of teeth can be variable. Male. Observed Males are as big as females (Table 6). Spination pattern same as females. Genital operculum has an oval posterior margin, expanded to the third segment of the opisthosoma. Male genitalia. (MZUSP 54589): LaM longer than Pi. LoL 1 longer than LoL 2, LoL 1 is wider, densely covered with minute projections, placed close together with the base not visible; LoL 2 is winkled and does not present projections. In dorsal view, the LoD are rounded very long and covering LoL 1 and LoL 2 (Fig. 19 A, B, C).	en	Chirivi-Joya, Daniel, Moreno-González, Jairo A., Fagua, Giovanny (2020): Two new species of the whip-spider genus Heterophrynus (Arachnida: Amblypygi) with complementary information of four species. Zootaxa 4803 (1): 1-41, DOI: 10.11646/zootaxa.4803.1.1
