taxonID	type	description	language	source
03B38784B13F0F27DB94FCB6335BFCE6.taxon	type_taxon	Type species: Jacaena distincta Thorell, 1897	en	Mu, Yannan, Zhang, Feng (2020): The genus Jacaena Thorell, 1897 from southern China (Araneae: Liocranidae). Zootaxa 4819 (2): 335-348, DOI: 10.11646/zootaxa.4819.2.6
03B38784B13F0F25DB94FC6732A9FD68.taxon	description	Figs 1 D – F, 2 D – F, 3	en	Mu, Yannan, Zhang, Feng (2020): The genus Jacaena Thorell, 1897 from southern China (Araneae: Liocranidae). Zootaxa 4819 (2): 335-348, DOI: 10.11646/zootaxa.4819.2.6
03B38784B13F0F25DB94FC6732A9FD68.taxon	materials_examined	Type material. Holotype ♂ (YNPB 190809 - 01), CHINA, Yunnan Province, Hani-Yi Autonomous Prefecture of Honghe, Pingbian County, Daweishan Forest Park (103 ° 42 ′ 18.75 ″ E, 22 ° 54 ′ 38.91 ″ N), 1967 m a. s. l., 9 Aug 2019, leg. Yannan Mu. Paratype: 1 ♀, same data as the holotype; 1 ♀ (YNML 190815), Yunnan Province, Dai Autonomous Pre-fecture of Xishuangbanna, Mengla County, Menglun Town, Xishuangbanna Tropical Botanical Garden of Chinese Academy of Sciences (21 ° 55 ′ 09.01 ″ N, 101 ° 16 ′ 14.34 ″ E), 566 m a. s. l., 15 Aug 2019, leg. Yannan Mu; 1 ♀, Chongq-ing City, Simian Mountain National Nature Reserve, Dawopu Reserve Station, Chaqi Mountain (28 ° 34 ′ 47.4 ′′ N, 106 ° 21 ′ 05.58 ′′ E), 1336 m a. s. l., 10 Nov 2011, leg. Luyu Wang; 2 ♀, Yunnan Province, Hani-Yi Autonomous Prefec-ture of Honghe, Pingbian County, Daweishan Forest Park (22 ° 57 ′ 12.18 ′′ N, 103 ° 41 ′ 35.28 ′′ E), 1684 m a. s. l., 10 May 2011, leg. Zongxu Li.	en	Mu, Yannan, Zhang, Feng (2020): The genus Jacaena Thorell, 1897 from southern China (Araneae: Liocranidae). Zootaxa 4819 (2): 335-348, DOI: 10.11646/zootaxa.4819.2.6
03B38784B13F0F25DB94FC6732A9FD68.taxon	etymology	Etymology. The specific name is derived from the Latin word “ asper ”, meaning “ rough ”, referring to the rough surface of the tegular apophysis; adjective.	en	Mu, Yannan, Zhang, Feng (2020): The genus Jacaena Thorell, 1897 from southern China (Araneae: Liocranidae). Zootaxa 4819 (2): 335-348, DOI: 10.11646/zootaxa.4819.2.6
03B38784B13F0F25DB94FC6732A9FD68.taxon	diagnosis	Diagnosis. This new species can be distinguished from all other Jacaena species by the large and thick tegular apophysis, which has a granulated surface (Fig. 2 F).	en	Mu, Yannan, Zhang, Feng (2020): The genus Jacaena Thorell, 1897 from southern China (Araneae: Liocranidae). Zootaxa 4819 (2): 335-348, DOI: 10.11646/zootaxa.4819.2.6
03B38784B13F0F25DB94FC6732A9FD68.taxon	description	Description. Male (Holotype). Total body length 4.90: carapace 2.60 long, 1.94 wide; abdomen 2.30 long, 1.37 wide. Carapace black, radial granulated integument on carapace; cephalic area relatively broad; fovea small and round. Eye sizes and interdistances: AME 0.12, ALE 0.14, PME 0.08, PLE 0.13; AME – AME 0.07, AME – ALE 0.03, PME – PME 0.11, PME – PLE 0.14; MOA 0.34 long, anterior width 0.30, posterior width 0.28. Clypeal height 0.20, greater than diameter of AME. Chelicerae moderately long, deep red-brown, with three promarginal teeth, two retromarginal teeth. Endites and labium gray; endites longer than wide, widest anteriorly, with maxillar hair tuft prolaterally. Labium almost as long as wide (0.38: 0.37), tip depressed, with several setae. Sternum brown, shieldshaped, with intercoxal sclerites between coxae I and II, II and III, III and IV; precoxal triangles present; anterior margin truncated, posterior margin not extending between coxae IV, only slightly narrower in front than in middle. Legs gray, tarsi with two dentate claws and conspicuous claw tufts, trochanters deeply notched. Leg spination: femur I with 2 strong pl spines (Fig. 2 E), other femora spineless; tibia I with 7 pv and 6 rv spines, tibia II with 7 pv and 5 rv spines, tibia III with 2 pl and 3 rv spines, tibia IV with 3 pv and 2 rv spines, and 1 rl spine; metatarsus I with 4 pairs spines ventrally, metatarsus II with 4 pairs spines ventrally, metatarsus III with 1 pl, 2 rl spine, metatarsus IV with 2 pl, 1 rl spines. Leg I 5.95 (1.78 + 0.63 + 1.44 + 1.31 + 0.79); Leg II 6.12 (1.74 + 0.71 + 1.51 + 1.34 + 0.82); Leg III 5.57 (1.55 + 0.69 + 1.15 + 1.30 + 0.88); Leg IV 8.06 (2.15 + 0.79 + 1.81 + 2.14 + 1.17). Leg formula: 4213. Abdomen ovoid and black, with small dorsal scutum (occupying less than half abdomen length); one pair of muscular impressions centrally, wrinkled on both sides. Palp as illustrated (Fig. 3 C – E). Tibia short, dark brown, RTA with two processes, sclerotized dorsal and membranous ventral surface; dRTA little, pRTA expanded. Tegulum white-yellow, reniform, with faint sperm duct and large tegular apophysis with strongly rough and granulated surface. Embolus long, with thin base, strongly coiled, tip along with tegular apophysis. Conductor indistinct, behind embolus. Female. One female paratype total body length 6.12: carapace 2.94 long, 2.31 wide; abdomen 3.18 long, 1.94 wide. Carapace deep brown, radial granulate integument on carapace; cephalic area relatively broad; fovea inconspicuous. Eye sizes and interdistances: AME 0.13, ALE 0.14, PME 0.11, PLE 0.13; AME – AME 0.11, AME – ALE 0.06, PME – PME 0.11, PME – PLE 0.16; MOA 0.35 long, anterior width 0.33, posterior width 0.33. Clypeal height 0.27, more than two times diameter of AME. Leg I 8.35 (2.38 + 0.87 + 2.24 + 1.81 + 1.05); Leg II 7.38 (2.14 + 0.94 + 1.8 5 + 1.52 + 0.93); Leg III 5.78 (1.64 + 0.85 + 1.32 + 1.26 + 0.71); Leg IV 8.67 (2.28 + 0.81 + 2.09 + 2.41 + 1.08). Leg formula: 4123. Other characters as in male. Abdomen ovoid and black, without dorsal scuta, with two pairs of muscular impressions centrally. Venter of abdomen light brown. Epigynum as in Fig. 3 A – B. Epigynal plate poorly sclerotized, two copulatory openings large and circular, located at center of epigynal plate. Copulatory ducts long and tapered off, coiled as conical; fertilization ducts short. Spermathecae small, separated into two parts.	en	Mu, Yannan, Zhang, Feng (2020): The genus Jacaena Thorell, 1897 from southern China (Araneae: Liocranidae). Zootaxa 4819 (2): 335-348, DOI: 10.11646/zootaxa.4819.2.6
03B38784B13F0F25DB94FC6732A9FD68.taxon	distribution	Distribution. China (Fig. 9, Yunnan Province, Chongqing City).	en	Mu, Yannan, Zhang, Feng (2020): The genus Jacaena Thorell, 1897 from southern China (Araneae: Liocranidae). Zootaxa 4819 (2): 335-348, DOI: 10.11646/zootaxa.4819.2.6
03B38784B13A0F23DB94F9EB3296F80C.taxon	description	Figs 1 A – C, 2 A, 4	en	Mu, Yannan, Zhang, Feng (2020): The genus Jacaena Thorell, 1897 from southern China (Araneae: Liocranidae). Zootaxa 4819 (2): 335-348, DOI: 10.11646/zootaxa.4819.2.6
03B38784B13A0F23DB94F9EB3296F80C.taxon	materials_examined	Type material. Holotype ♂ (YNBN- 17 - 0515), CHINA, Yunnan Province, Dai Autonomous Prefecture of Xish-uangbanna, Mengla County, Menglun Town, Xishuangbanna Tropical Botanical Garden of Chinese Academy of Sciences, Lvshi Forest (21 ° 54 ′ 40.59 ′′ N, 101 ° 17 ′ 02.39 ′′ E), 683 m a. s. l., 15 May 2017, leg. Beibei Zhou. Paratype: 1 ♀, Yunnan Province, Dai Autonomous Prefecture of Xishuangbanna, Jinghong City, 23 Oct 1986, leg. Mingsheng Zhu; 1 ♀, Hainan Province, Chengmai County, Heling Farm, Dawangling, 17 May 2015, leg. Chi Jin; 2 ♀ 1 ♂ 2 j, Hainan Province, Qiongzhong County, on the road to Limu Mountain, 21 May 2015, leg. Li’na Fu; 2 ♀ 2 ♂, Hainan Province, Wuzhishan City, Shuiman Town, Wuzhishan tropical rainforest scenic spot, 25 Jan 2015, leg. Xiangbo Guo; 2 ♀ 2 ♂, Hainan Province, Changjiang County, Bawangling Mountain, 4 Feb 2015, leg. Baoshi Zhang; 1 ♀ 2 ♂, Hainan Province, Changjiang County, Bawangling Mountain, 4 Feb 2015, leg. Chi Jin; 1 ♀ 1 ♂ 2 j, Hainan Province, Ledong Li Autonomous County, Jianfengling Mountain, 31 Jan 2015, leg. Li’na Fu; 1 ♀, Hainan Province, Qiong-zhong County, Limushan Town, Qijiafang holiday village, 20 Jan 2015, leg. Chi Jin; 1 ♀, Hainan Province, Ledong Li Autonomous County, Jianfengling Mountain, 26 May 2011, leg. Chao Zhang; 1 ♂ (HaiN- 16 - 02), Hainan Prov-ince, Qiongzhong County, on the road to Limu Mountain (19 ° 10 ′ 37.62 ′′ N, 109 ° 45 ′ 00.84 ′′ E), 694 m a. s. l., 4 April 2016, leg. Chi Jin; 1 ♀ (HaiN- 16 - 04) Hainan Province, Qiongzhong County, Shuiman Town, Wuzhishan tourist area (18 ° 54 ′ 09.89 ″ N, 109 ° 41 ′ 18.27 ″ E), 930 m a. s. l., 6 April 2016, leg. Chi Jin; 1 ♀ (HaiN- 16 - 05), Hainan Province, Wuzhishan City, Shuiman Town, Wuzhishan Tropical Rainforest Scenic Spot (18 ° 52 ′ 11.64 ′′ N, 109 ° 40 ′ 22.56 ′′ E), 800 m a. s. l., 7 April 2016, leg. Chi Jin; 2 ♀ 1 ♂ (HaiN- 16 - 07), Hainan Province, Ledong Li Autonomous County, Jianfengling Mountain (18 ° 44 ′ 29.16 ′′ N, 108 ° 50 ′ 33.72 ′′ E), 1068 m a. s. l., 9 April 2016, leg. Chi Jin; 1 ♀ (HaiN- 1608), Hainan Province, Ledong Li Autonomous County, Jianfengling Mountain Forestry Bureau (18 ° 44 ′ 13.50 ′′ N, 108 ° 51 ′ 32.04 ′′ E), 885 m a. s. l., 10 April 2016, leg. Chi Jin; 1 ♂ (Y 940 IL 1), 1 ♀ (H 9403 L 3 b), no detailed collection information.	en	Mu, Yannan, Zhang, Feng (2020): The genus Jacaena Thorell, 1897 from southern China (Araneae: Liocranidae). Zootaxa 4819 (2): 335-348, DOI: 10.11646/zootaxa.4819.2.6
03B38784B13A0F23DB94F9EB3296F80C.taxon	etymology	Etymology. The species name is a noun in apposition, derived from the type locality.	en	Mu, Yannan, Zhang, Feng (2020): The genus Jacaena Thorell, 1897 from southern China (Araneae: Liocranidae). Zootaxa 4819 (2): 335-348, DOI: 10.11646/zootaxa.4819.2.6
03B38784B13A0F23DB94F9EB3296F80C.taxon	diagnosis	Diagnosis. This new species resembles Jacaena mihun (Deeleman-Reinhold, 2001), but can be easily distinguished by: (1) the long and circular embolus (vs narrower and shorter); (2) the thin base of the embolus (vs wider than all other species); (3) the simple coiled and folding copulatory duct (vs complex).	en	Mu, Yannan, Zhang, Feng (2020): The genus Jacaena Thorell, 1897 from southern China (Araneae: Liocranidae). Zootaxa 4819 (2): 335-348, DOI: 10.11646/zootaxa.4819.2.6
03B38784B13A0F23DB94F9EB3296F80C.taxon	description	Description. Male (Holotype). Total body length 4.88: carapace 2.45 long, 1.95 wide; abdomen 2.43 long, 1.55 wide. Carapace black brown, darker in eye area; radial granulated integument on carapace; cephalic area relatively broad; fovea deep and round. Eye sizes and interdistances: AME 0.15, ALE 0.15, PME 0.10, PLE 0.13; AME – AME 0.18, AME – ALE 0.13, PME – PME 0.20, PME – PLE 0.20, ALE – PLE 0.20; MOA 0.25 long, anterior width 0.18, posterior width 0.23. Clypeal height 0.15, equal to the diameter of AME. Labium longer than wide, tip cambered and with several setae. Leg spination: femur I with 2 strong pl spines, other femora spineless; femur I with 2 strong pl spines (Fig. 1), other femora spineless; tibia I with 7 pv and 8 rv spines, tibia II with 6 pv and 6 rv spines, tibia III with 2 pl and 1 rv spines, tibia IV with 3 pv and 2 rv spines, and 1 rl spine; metatarsus I with 5 pairs spines ventrally, metatarsus II with 4 pairs spines ventrally, metatarsus III with 1 pl, 1 rl and 1 pair ventral spines, metatarsus IV with 1 pl, 1 rl and 2 pairs ventral spines. Leg I 6.81 (1.28 + 0.85 + 1.95 + 1.80 + 0.93); Leg II 6.03 (1.68 + 0.69 + 1.29 + 1.45 + 0.9 2); Leg III 3.19 (0.59 + 0.56 + 0.69 + 0.92 + 0.43); Leg IV 7.32 (1.52 + 0.96 + 1.91 + 1.95 + 0.98). Leg formula: 4123. Other characters as same as J. aspera sp. n., except color. Abdomen ovoid and gray, with small distinct dorsal scutum (occupying less than half the length of the abdomen), wrinkled at both sides. Palp as illustrated (Fig. 4 D – F). Tibia short, red-brown, RTA with two processes, sclerotized dorsal and membranous ventral surface; dRTA wide, pRTA small. Color of tegulum same as tibia, reniform, with distinct sperm duct and spiral and cuspidal tegular apophysis. Embolus long, with thin base, strongly coiled, tip along with tegular apophysis. Conductor membranous, behind embolus. Female. One female paratype total body length 5.83: carapace 2.90 long, 2.30 wide; abdomen 2.93 long, 1.93 wide. Carapace black brown, with metallic lustre, darker in eye area; radial granulate integument on carapace; cephalic area relatively broad; fovea deep and round. Eye sizes and interdistances: AME 0.16, ALE 0.15, PME 0.12, PLE 0.14; AME – AME 0.25, AME – ALE 0.18, PME – PME 0.26, PME – PLE 0.30, ALE – PLE 0.20; MOA 0.25 long, anterior width 0.18, posterior width 0.23. Clypeal height 0.16, equal to the diameter of AME. Leg spination: tibia I with 7 pv and 8 rv spines, tibia II with 8 pv and 9 rv spines, tibia III with 1 pl and 2 rv spines, tibia IV with 1 pv and 4 rv spines; metatarsus I with 4 pairs spines ventrally, metatarsus II with 4 pairs spines ventrally, metatarsus III with 1 pl, 1 pv and 1 rv spines, metatarsus IV with 3 pv and 3 rv spines. Leg I 7.49 (1.90 + 0.95 + 2.30 + 1.35 + 0.99); Leg II 7.03 (1.80 + 0.90 + 1.80 + 1.49 + 1.04); Leg III 6.38 (1.71 + 0.81 + 1.35 + 1.48 + 1.03); Leg IV 9.00 (2.30 + 0.90 + 2.16 + 2.43 + 1.21). Leg formula: 4123. Other characters same as male. Abdomen ovoid, without dorsal scutum, brownish gray, with two pairs of muscular impressions centrally. Venter of abdomen light brown. Epigynum as in Fig. 4 A – C. Epigynal plate sclerotized, two copulatory openings concave and oval, located at center of epigynal plate. Copulatory ducts long and coiled; fertilization ducts short. Spermathecae large, separated into two parts.	en	Mu, Yannan, Zhang, Feng (2020): The genus Jacaena Thorell, 1897 from southern China (Araneae: Liocranidae). Zootaxa 4819 (2): 335-348, DOI: 10.11646/zootaxa.4819.2.6
03B38784B13A0F23DB94F9EB3296F80C.taxon	distribution	Distribution. China (Fig. 9, Yunnan Province, Hainan Province).	en	Mu, Yannan, Zhang, Feng (2020): The genus Jacaena Thorell, 1897 from southern China (Araneae: Liocranidae). Zootaxa 4819 (2): 335-348, DOI: 10.11646/zootaxa.4819.2.6
03B38784B1380F21DB94FA71323DFD80.taxon	description	Figs 1 G, 2 B, 5 A – C	en	Mu, Yannan, Zhang, Feng (2020): The genus Jacaena Thorell, 1897 from southern China (Araneae: Liocranidae). Zootaxa 4819 (2): 335-348, DOI: 10.11646/zootaxa.4819.2.6
03B38784B1380F21DB94FA71323DFD80.taxon	materials_examined	Type material. Holotype ♂, CHINA, Guangdong Province, Chaozhou City, Chao’an County, Jiaoshuikeng Vil-lage, 8 Aug 2011, leg. Shuwen Xing.	en	Mu, Yannan, Zhang, Feng (2020): The genus Jacaena Thorell, 1897 from southern China (Araneae: Liocranidae). Zootaxa 4819 (2): 335-348, DOI: 10.11646/zootaxa.4819.2.6
03B38784B1380F21DB94FA71323DFD80.taxon	etymology	Etymology. The specific name is a Latin adjective meaning “ tawny ”, referring to the color of the palp.	en	Mu, Yannan, Zhang, Feng (2020): The genus Jacaena Thorell, 1897 from southern China (Araneae: Liocranidae). Zootaxa 4819 (2): 335-348, DOI: 10.11646/zootaxa.4819.2.6
03B38784B1380F21DB94FA71323DFD80.taxon	diagnosis	Diagnosis. This new species is similar to J. jinxini, but can be distinguished by: 1) the short embolus (vs longer embolus); 2) the expanded and thumb-shaped pRTA (vs truncated); 3) the different color of whole palp fulvous.	en	Mu, Yannan, Zhang, Feng (2020): The genus Jacaena Thorell, 1897 from southern China (Araneae: Liocranidae). Zootaxa 4819 (2): 335-348, DOI: 10.11646/zootaxa.4819.2.6
03B38784B1380F21DB94FA71323DFD80.taxon	description	Description. Male (Holotype). Total body length 5.89: carapace 2.97 long, 2.32 wide; abdomen 2.92 long, 1.72 wide. Carapace nearly circular, deep red-brown, with radial granulated integument; eye area relatively wide; fovea deep and round. Eye sizes and interdistances: AME 0.15, ALE 0.15, PME 0.10, PLE 0.14; AME – AME 0.10, AME – ALE 0.06, PME – PME 0.14, PME – PLE 0.15, ALE – PLE 0.10; MOA 0.36 long, anterior width equal to poste-rior width (0.36). Clypeal height 0.23. Leg spination: femur I with 2 strong pl spines, other femora spineless; femur I with 2 strong pl spines, other femora spineless; tibia I with 8 pv and 7 rv spines, tibia II with 6 pv and 8 rv spines, tibia III with 2 pl and 1 rv spines, tibia IV absent; metatarsus I with 5 pl and 4 rl spines, metatarsus II with 3 pl and 4 rl spines, metatarsus III with 1 pl spines, metatarsus IV absent. Leg I 7.85 (2.16 + 0.90 + 2.13 + 1.81 + 0.85), Leg II 7.07 (1.98 + 0.87 + 1.76 + 1.50 + 0.96), Leg III 6.20 (1.73 + 0.81 + 1.29 + 1.42 + 0.95), Leg IV absent. Leg formula: 4123. Other characters as for J. aspera sp. n., except for coloration. Abdomen oval and gray, with small distinct dorsal scutum (occupying less than half the length of the abdomen), two pairs of muscular impressions centrally. Palp as illustrated (Fig. 5 A – C). Tibia short, yellowish-brown, RTA with two processes, dRTA rounded distally and strongly sclerotized, pRTA expanded and sclerotized. Tegulum same color as tibia, reniform, with distinct sperm duct and short thin tegular apophysis. Embolus short (not exceeding tip of TA), strongly coiled, tip alongside tegular apophysis. Conductor membranous, behind embolus. Female: unknown.	en	Mu, Yannan, Zhang, Feng (2020): The genus Jacaena Thorell, 1897 from southern China (Araneae: Liocranidae). Zootaxa 4819 (2): 335-348, DOI: 10.11646/zootaxa.4819.2.6
03B38784B1380F21DB94FA71323DFD80.taxon	distribution	Distribution. China (Fig. 9, Guangdong Province).	en	Mu, Yannan, Zhang, Feng (2020): The genus Jacaena Thorell, 1897 from southern China (Araneae: Liocranidae). Zootaxa 4819 (2): 335-348, DOI: 10.11646/zootaxa.4819.2.6
03B38784B1360F2EDB94FAB03451F869.taxon	description	Fig. 6	en	Mu, Yannan, Zhang, Feng (2020): The genus Jacaena Thorell, 1897 from southern China (Araneae: Liocranidae). Zootaxa 4819 (2): 335-348, DOI: 10.11646/zootaxa.4819.2.6
03B38784B1360F2EDB94FAB03451F869.taxon	materials_examined	Material examined. 2 ♂ 1 ♀ (GD 180516), CHINA, Guangdong Province, Shaoguan City, Ruyuan Yao Autono-mous County, Tianjingshan National Forest Park (24 ° 40 ’ 56.25 ’’ N, 112 ° 59 ’ 49.24 ’’ E), 483 m a. s. l., 15 May 2018, leg. Yannan Mu; 1 ♀ (GD 180517), Guangdong Province, Shaoguan City, Ruyuan Yao Autonomous County, Tian-jingshan National Forest Park (24 ° 40 ’ 56.25 ’’ N, 112 ° 59 ’ 49.24 ’’ E), 483 m a. s. l., 17 May 2018, leg. Xuyun Gong; 2 ♀ (GZJK 190727 - 02), Guizhou Province, Guizhou City, Jiangkou County, Fanjing Mountain Zoology Station (27 ° 50 ′ 51.72 ″ N, 108 ° 46 ′ 19.48 ″ E), 565 m a. s. l., 27 July 2019, leg. Yannan Mu; 2 ♂ 1 ♀ (GXBL- 18 - 11), Guangxi Province, Beiliu City, Minle Town, Darong Mountain National Forest Park (20 ° 51 ’ 59.02 ’’ N, 110 ° 16 ’ 10.42 ’’ E), 745 m a. s. l., 21 Aug 2018, leg. Yannan Mu; 1 ♂ (GXXA- 18 - 05), Guangxi Province, Guilin City, Xing’an County, Gaozhai Village, Liyuan Waterfall (25 ° 52 ’ 41.34 ’’ N, 110 ° 28 ’ 29.03 ’’ E), 636 m a. s. l., 13 Aug 2018, leg. Lingchen Zhao; 1 ♀ (GXGG 180817), Guangxi Province, Guigang City, Gangbei District, Pingtian Mountain National Forest Park (23 ° 10 ′ 38.54 ′′ N, 109 ° 28 ′ 12.38 ′′ E), 253 m a. s. l., 17 Aug 2018, leg. Yang Chen; 1 ♂, Hunan Province, Changsha City, Qiaoyi Town, Heimi Mountain Foundation (28 ° 26 ′ 48.10 ′′ N, 112 ° 59 ′ 50.02 ′′ E), 369 m a. s. l., 20 July 2018, leg. Zhaoyi Li; 1 ♀, Fujian Province, Nanping City, Jianyang County, Aotou Village, Dazhulan Region, 13 July 2011, leg. Feng Zhang; 1 ♀, Guangdong Province, Nan’ao County, 19 Sep 2014, leg. Shuwen Xing.	en	Mu, Yannan, Zhang, Feng (2020): The genus Jacaena Thorell, 1897 from southern China (Araneae: Liocranidae). Zootaxa 4819 (2): 335-348, DOI: 10.11646/zootaxa.4819.2.6
03B38784B1360F2EDB94FAB03451F869.taxon	description	Description. For description of both sexes see Liu et al. (2020). Habitus and genitalia of both sexes in Fig. 6.	en	Mu, Yannan, Zhang, Feng (2020): The genus Jacaena Thorell, 1897 from southern China (Araneae: Liocranidae). Zootaxa 4819 (2): 335-348, DOI: 10.11646/zootaxa.4819.2.6
03B38784B1360F2EDB94FAB03451F869.taxon	distribution	Distribution. China (Fig. 9, Guangxi Province, Guangdong Province, Hunan Province, Fujian Province).	en	Mu, Yannan, Zhang, Feng (2020): The genus Jacaena Thorell, 1897 from southern China (Araneae: Liocranidae). Zootaxa 4819 (2): 335-348, DOI: 10.11646/zootaxa.4819.2.6
03B38784B1360F2EDB94FF3033EFFB5D.taxon	description	Figs 1 H, 2 C, 5 D – E	en	Mu, Yannan, Zhang, Feng (2020): The genus Jacaena Thorell, 1897 from southern China (Araneae: Liocranidae). Zootaxa 4819 (2): 335-348, DOI: 10.11646/zootaxa.4819.2.6
03B38784B1360F2EDB94FF3033EFFB5D.taxon	materials_examined	Type material. Holotype ♂, CHINA, Yunnan Province, Dai Autonomous Prefecture of Xishuangbanna, Mengla County, Menglun Town, Xishuangnanna Tropical Botanical Garden of ChineseAcademy of Sciences (21 ° 54 ’ 36.41 ’’ N, 101 ° 16 ’ 11.17 ’’ E), 526 m a. s. l., 12 July 2018, leg. Chi Jin.	en	Mu, Yannan, Zhang, Feng (2020): The genus Jacaena Thorell, 1897 from southern China (Araneae: Liocranidae). Zootaxa 4819 (2): 335-348, DOI: 10.11646/zootaxa.4819.2.6
03B38784B1360F2EDB94FF3033EFFB5D.taxon	etymology	Etymology. The species name is a noun in apposition, derived from the type locality.	en	Mu, Yannan, Zhang, Feng (2020): The genus Jacaena Thorell, 1897 from southern China (Araneae: Liocranidae). Zootaxa 4819 (2): 335-348, DOI: 10.11646/zootaxa.4819.2.6
03B38784B1360F2EDB94FF3033EFFB5D.taxon	diagnosis	Diagnosis. This new species is similar to J. aspera sp. n., but can be distinguished by: 1) the thin and smooth tegular apophysis (vs thicker and rough); 2) the salient dRTA (vs small).	en	Mu, Yannan, Zhang, Feng (2020): The genus Jacaena Thorell, 1897 from southern China (Araneae: Liocranidae). Zootaxa 4819 (2): 335-348, DOI: 10.11646/zootaxa.4819.2.6
03B38784B1360F2EDB94FF3033EFFB5D.taxon	description	Description. Male (Holotype). Total body length 6.06: carapace 3.21 long, 2.49 wide; abdomen 2.85 long, 1.73 wide. Carapace oval, deep black brown, with radial granulated integument; fovea deep and round. Eye sizes and interdistances: AME 0.12, ALE 0.14, PME 0.12, PLE 0.12; AME – AME 0.11, AME – ALE 0.08, PME – PME 0.12, PME – PLE 0.20, ALE – PLE 0.20; MOA 0.33 long, anterior width slowly short than posterior width (0.33: 0.42). Clypeal height 0.27, more than two times of the diameter of AME. Leg spination: femur I with 2 strong pl spines, other femora spineless; tibia I with 6 pv and 7 rv spines, tibia II with 6 pairs of ventral spines, tibia III with 3 pl and 2 rv spines, and 1 pl and 1 rl spines, tibia IV with 3 pv and 1 rv spines; metatarsus I with 4 pairs of ventral spines, metatarsus II with 4 pairs of ventral spines, metatarsus III with 1 pl, 1 rl and 1 rv spines, metatarsus IV with 1 pv, 2 rv, and 1 pl, 1 rl spines. Leg I 8.53 (2.36 + 0.99 + 2.23 + 1.85 + 1.10), Leg II 7.51 (2.17 + 0.95 + 1.83 + 1.53 + 1.03), Leg III 6.63 (1.82 + 0.88 + 1.37 + 1.60 + 0.96), Leg IV 9.56 (2.48 + 0.98 + 2.13 + 2.57 + 1.40). Leg formula: 4123. Other characters see J. aspera sp. n., except for different coloration. Abdomen oval and dark gray, with small dorsal scutum and pair of muscular impressions centrally. Palp as illustrated (Fig. 5 D – E). Tibia short, dark brown, RTA with two processes, dRTA triangular, strongly sclerotized, tip milky; pRTA short and sclerotized. Tegulum yellow-brown, reniform, with faint sperm duct and long, spiral and cuspidal tegular apophysis; subtegulum thin. Embolus thin, strongly coiled, tip along with tegular apophysis. Conductor membranous, behind embolus. Female: unknown.	en	Mu, Yannan, Zhang, Feng (2020): The genus Jacaena Thorell, 1897 from southern China (Araneae: Liocranidae). Zootaxa 4819 (2): 335-348, DOI: 10.11646/zootaxa.4819.2.6
03B38784B1360F2EDB94FF3033EFFB5D.taxon	discussion	Comment: This new species and one of the paratypes of another new species (J. aspera sp. n.) were both collected from same location (Xishuangbanna Tropical Botanical Garden of the Chinese Academy of Sciences). However, the distribution range of J. aspera sp. n. is larger than J. menglaensis sp. n..	en	Mu, Yannan, Zhang, Feng (2020): The genus Jacaena Thorell, 1897 from southern China (Araneae: Liocranidae). Zootaxa 4819 (2): 335-348, DOI: 10.11646/zootaxa.4819.2.6
03B38784B1360F2EDB94FF3033EFFB5D.taxon	distribution	Distribution. China (Fig. 9, Yunnan Province).	en	Mu, Yannan, Zhang, Feng (2020): The genus Jacaena Thorell, 1897 from southern China (Araneae: Liocranidae). Zootaxa 4819 (2): 335-348, DOI: 10.11646/zootaxa.4819.2.6
03B38784B1340F2DDB94FF3033EFF9E6.taxon	description	Figs 7, 8	en	Mu, Yannan, Zhang, Feng (2020): The genus Jacaena Thorell, 1897 from southern China (Araneae: Liocranidae). Zootaxa 4819 (2): 335-348, DOI: 10.11646/zootaxa.4819.2.6
03B38784B1340F2DDB94FF3033EFF9E6.taxon	materials_examined	Other material examined. 1 ♂ 1 ♀ (YNJH- 16 - 08), Yunnan Province, Dai Autonomous Prefecture of Xish-uangbanna, Mengla County, Menglun Town, Xishuangbanna Tropical Botanical Garden of Chinese Academy of Sciences, Lvshi Forest (21.911417 ° N, 101.283798 ° E), 673 m a. s. l., 19 Oct 2016, leg. Xiangbo Guo; 1 ♂ 1 ♀ (YNJH- 16 - 06), Yunnan Province, Dai Autonomous Prefecture of Xishuangbanna, Xishuangbanna Virgin Forest Park (22.033023 ° N, 100.88974 ° E), 805 m a. s. l., 17 Oct 2016, leg. Xiangbi Guo; 4 ♂ 7 ♀ (YNJH- 16 - 09), Yunnan Province, Dai Autonomous Prefecture of Xishuangbanna, Mengla County, Menglun Town, Xishuangbanna Tropical Botanical Garden of Chinese Academy of Sciences, tropical forest (21.91947 ° N, 101.269045 ° E), 554 m a. s. l., 20 Oct 2016, leg. Xiangbo Guo; 1 ♀ (YNML- 16 - 10), Yunnan Province, Dai Autonomous Prefecture of Xishuangban-na, Mengla County, Nangong Mountain (21.621613 ° N, 101.460492 ° E), 1068 m a. s. l., 21 Oct 2016, leg. Xiangbo Guo; 4 ♀ 1 ♂ (YNBN- 16 - 05), Yunnan Province, Dai Autonomous Prefecture of Xishuangbanna, Mengla County, Menglun Town, Xishuangbanna Tropical Botanical Garden of the Chinese Academy of Sciences, tropical forest (21 ° 54.982 ′ N, 101 ° 16.216 ′ E), 558 m a. s. l., 29 Nov 2016, leg. Luyu Wang; 1 ♀ (YNBN- 16 - 09), Yunnan Province, Dai Autonomous Prefecture of Xishuangbanna, Mengla County, Menglun Town, Xishuangbanna Tropical Botanical Garden of Chinese Academy of Sciences, tropical forest (21 ° 54.982 ′ N, 101 ° 16.216 ′ E), 1 Dec 2016, leg. Guiqiang Huang; 1 ♀, Yunnan Province, Dai Autonomous Prefecture of Xishuangbanna, Mengla County, Menglun Town, Xishuangbanna Tropical Botanical Garden of Chinese Academy of Sciences, 18 July 2010, leg. Chao Zhang; 1 ♀ 2 ♂ (Y 9401 L 1), 1 ♀ 1 ♂ (Y 9403 L 1), 3 ♂ (Y 8311 L 5), no detailed collection information; 1 ♀ (YNBN- 16 - 09), Yunnan Province, Dai Autonomous Prefecture of Xishuangbanna, Mengla County, Menglun Town, Xishuangbanna Tropical Botanical Garden of Chinese Academy of Sciences, Lvshi Forest (21 ° 54.533 ′ N, 101 ° 16.958 ′ E), 632 m a. s. l., 18 Oct 2016, leg. Guiqiang Huang; 4 ♀ 1 ♂ (YNPE- 16 - 05), Yunnan Province, Pu’er City, Simao District, Ximahe Reservoir (22 ° 47.426 ′ N, 100 ° 59.378 ′ E), 1333 m a. s. l., 16 Oct 2016, leg. Yanchao Wang.	en	Mu, Yannan, Zhang, Feng (2020): The genus Jacaena Thorell, 1897 from southern China (Araneae: Liocranidae). Zootaxa 4819 (2): 335-348, DOI: 10.11646/zootaxa.4819.2.6
03B38784B1340F2DDB94FF3033EFF9E6.taxon	description	Description. See Zhang & Fu (2011) for description of both sexes. Habitus and genitalia of both sexes in Figs 7 and 8.	en	Mu, Yannan, Zhang, Feng (2020): The genus Jacaena Thorell, 1897 from southern China (Araneae: Liocranidae). Zootaxa 4819 (2): 335-348, DOI: 10.11646/zootaxa.4819.2.6
03B38784B1340F2DDB94FF3033EFF9E6.taxon	distribution	Distribution. China (Fig. 9, Yunnan Province).	en	Mu, Yannan, Zhang, Feng (2020): The genus Jacaena Thorell, 1897 from southern China (Araneae: Liocranidae). Zootaxa 4819 (2): 335-348, DOI: 10.11646/zootaxa.4819.2.6
