identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03B387F5FF9EC16142C6FADDFF6BF831.text	03B387F5FF9EC16142C6FADDFF6BF831.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neoclypeodytes balkei Scheers & Hájek 2020	<div><p>Neoclypeodytes balkei sp. nov.</p><p>(Fig. 1)</p><p>Type material. Holotype: m#, “ Panama: Panama, Campana, / cascade below Anam office, / 270m, 19.vi.2007, / 08.40.818N 079.54.075W, / Balke (PAN 07) [p] // HOLOTYPE m# / NEOCLYPEODYTES / balkei sp. nov. / K. Scheers &amp; J. Hájek det. 2020 [p, red label]” (ZSMG) . Paratypes: 46 specimens; same label data as holotype (BMNH, CKSB, NMPC, ZSMG). Each paratype is provided with the appropriate red printed label .</p><p>Description of male holotype. Habitus (Fig. 1A). Ovoid-shaped, slightly attenuate anteriorly and strongly attenuate posteriorly; lateral outline nearly continuous between pronotum and elytra in dorsal aspect. Surface shiny.</p><p>Coloration. Head piceous. Antennae with antennomeres I–IV uniformly pale testaceous, antennomeres V–XI slightly infuscated apically. Palpi pale testaceous. Disc of pronotum piceous, medially slightly lightened; lateral parts of pronotum pale testaceous, broadly reaching inwards. Elytron dark piceous with two testaceous maculae; anterior (subbasal) macula subquadrangular, extending from lateral margin to one fifth of elytral width to suture, anterior margin straight with small emargination sublaterally, posterior margin largely concave with posteriorly protruding tooth near suture; posterior macula not reaching suture, anteriorly not reaching lateral margin, with concave emargination towards suture, posteriorly reaching lateral margin and apex of elytron (Fig. 1A); epipleuron completely testaceous. Legs including tarsi testaceous. Prosternum, mesepisterna and mesepimera rufotestaceous; metaventrite, metepisterna and metacoxae rufotestaceous, somewhat darkened at lateral portions. Abdominal ventrites piceous.</p><p>Head with fine but distinct microreticulation anterior of occipital line; punctation consisting of fine punctures near eyes and coarser punctures on clypeus; anterior clypeal margin medially distinctly flattened; occipital line narrowly separated from posterior margin of eyes.</p><p>Pronotum widest at base; lateral bead distinct; lateral margin evenly rounded; microreticulation absent; punctation consisting of punctures separated by 2–3× puncture diameter, punctation somewhat sparser laterally and distinctly denser near anterior and posterior margins; basal striae sinuated, distinctly curved inwards near base and slightly outwards anteriorly, length about 0.55× length of pronotum measured along same line as stria.</p><p>Elytra with basal striae straight, about same length than those on pronotum; sutural striae absent; microreticulation absent; punctation similar to that of pronotal disc, but with punctures slightly larger; epipleura sparsely punctate, with moderately oblique humeral carina; lateral epipleural rim anteriorly visible in dorsal view.</p><p>Legs. Pro- and mesotibiae club shaped. Pro- and mesotarsomeres 1–3 distinctly broadened, ventrally with adhesive setae. Metatarsomeres with well-developed natatorial setae on both sides.</p><p>Ventral surface (Fig. 1B). Prosternum medially narrowly convex, coarsely sculptured; prosternal process rather broad, subovate with apex broadly obtuse, laterally margined, slightly constricted medially. Metaventrite distinctly punctate over entire surface, punctures separated by 2–4× puncture diameter. Intralinear space of metacoxa broad (ML/MW = 2.40), sparsely and irregularly punctate with fine punctures, punctures separated by 1–4× puncture diameter; metacoxal plates distinctly punctate with coarse punctures, punctures separated by 1–3× puncture diameter, plates narrowly impunctate along anterior and posterior margins. Abdominal ventrite I coarsely punctate along margins, ventrite II sparsely and finely punctate; succeeding ventrites smooth with a row of setae bearing punctures along posterior margin and some sparse and fine punctures scattered over surface; ventrites IV and V with a tuft of long setae medially; ventrite VI unmodified.</p><p>Male genitalia. Median lobe in ventral view rather slender, parallel over whole length; apex distinctly truncate (Fig. 1C); ventral surface in apical part lacking scale-like setae (see under N. fortunensis sp. nov., and cf. Fig. 3D). Median lobe in lateral view broadest in apical third; ventral outline almost evenly curved; dorsal outline almost straight medially, distinctly curved inwards near apex (Fig. 1D). Lateral lobe (paramere) with apical segment elongated, nearly straight; in apical half on inner side with several long setae; apex distinctly curved but not protruding, subapical incision not perceptible (Fig. 1E).</p><p>Variability. With the exception of small variability in extent and shape of elytral color pattern, there is no significant variation present within the specimens of the type series.</p><p>Sexual dimorphism. The females have the dorsal surface completely reticulated with irregular polygonal meshes and thus they appear submatt; pro- and mesotarsomeres are less broadened than in males.</p><p>Measurements. TL: 2.05–2.15 mm (mean value: 2.10 ± 0.03 mm); holotype: 2.15 mm. TL-h: 1.85–1.95 mm (mean value: 1.85 ± 0.04 mm); holotype: 1.95 mm. TW: 1.10–1.20 mm (mean value: 1.15 ± 0.01 mm); holotype: 1.20 mm. Ratio TL/TW: 1.73–1.82 (mean value: 1.80 ± 0.02).</p><p>Differential diagnosis. Neoclypeodytes balkei sp. nov. is, together with N. curtulus and N. fortunensis sp. nov., one of only three representatives of the genus known south of Guatemala. The three species are easily distinguished from each other by the color pattern (cf. Figs 1A, 2 A–B and 3A), and the male genitalia with the median lobe parallel in ventral view, not medially expanded in lateral view (Figs 1 C–D) and lacking distinct scale-like setae at the apex (which are very distinct in both N. curtulus and N. fortunensis sp. nov.), and the apical part of the lateral lobe with appendage not protruding (Figs 1E, 2F and 3F).</p><p>The new species differs from the other species in the genus by the combination of the distinctive color pattern, the absence of sutural striae, the shape of the basal pronotal and elytral striae, and the male genitalia. In the identification key of Miller (2001), the new species falls into couplet 24, together with N. fryii (Clark, 1862) and N. quadrinotatus (Sharp, 1882); from both these species, N. balkei sp. nov. differs by the posteriorly strongly attenuated habitus, more strongly punctured metacoxal plates, and by the male genitalia (cf. Figs 1 C–E, and Figs 20C–E and 21C–E in Miller (2001)).</p><p>Etymology. The new species is named after Michael Balke, who collected the type series of this species. The specific epithet is a noun in the genitive case.</p><p>Collection circumstances. The specimens were collected in small water holes in the bedrock around a cascade of a forest stream (M. Balke, pers. comm. 2020) (Fig. 4). At this locality, the species was co-occurring with N. curtulus .</p><p>Distribution. To date the species is only known from the type locality at Campana, Panama province, Panama (Fig. 6).</p></div>	https://treatment.plazi.org/id/03B387F5FF9EC16142C6FADDFF6BF831	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Scheers, Kevin;Hájek, Jiří	Scheers, Kevin, Hájek, Jiří (2020): Neoclypeodytes Young from Panama, with description of two new species (Coleoptera: Dytiscidae: Hydroporinae: Bidessini). Zootaxa 4890 (2): 245-256, DOI: 10.11646/zootaxa.4890.2.5
03B387F5FF9BC16642C6FF65FC57F8DA.text	03B387F5FF9BC16642C6FF65FC57F8DA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neoclypeodytes curtulus (Sharp 1887)	<div><p>Neoclypeodytes curtulus (Sharp, 1887)</p><p>(Fig. 2)</p><p>Bidessus curtulus Sharp, 1887: 753 (orig. descr., Panama).</p><p>Neoclypeodytes curtulus: Young 1969: 3 (new combination).</p><p>Type material. Lectotype: m# (designated by Miller 2001), “ Bidessus cur- / tulus. Types D.S. / Taboga island . / Champion [Sharp’s hw] // B.C.A. Col. I.2. / Bidessus / curtulus, / Sharp. [p] // Taboga Isl., / Panama. / Champion. [p] // Sharp Coll / 1905-313. [p] // Type [p, round label with red frame] // Lectotype specimen / dissected and on red / point. Other specimen / (White point) is a / paralectotype. [p, pink label] KBM [hw] // LECTOTYPE / Bidessus curtulus / Sharp 1887 / designated by / K.B. Miller 1997 [p, pink label with double black frame]” . Paralectotypes: 1 specimen with same label data as lectotype; 3 specimens, “ Bidessus cur- / tulus D.S. / Taboga island . / Champion [ Sharp’s hw] // Co- / type [p, round label with yellow frame] // Taboga Isl., / Panama. / Champion. [p] // Sharp Coll / 1905-313. [p] // B.C.A. Col. I.2. / Bidessus / curtulus, / Sharp. [p] // PARALECTOTYPE / Bidessus curtulus Sharp / 1887, designated by / K.B. Miller 1997 [p, blue label with double black frame]” ; 1 specimen, “ Bidessus cur- / tulus. / Taboga island / Panama. Champion [Sharp’s hw] // Co- / type [p, round label with yellow frame] // Sharp Coll / 1905-313. [p] // PARALECTOTYPE / Bidessus curtulus Sharp / 1887, designated by / K.B. Miller 1997 [p, blue label with double black frame]” ; 21 specimens, “Co- / type [p, round label with yellow frame] // Taboga Isl., / Panama. / Champion. [p] // B.C.A. Col. I.2. / Bidessus / curtulus, / Sharp. [p]” ; 3 specimens, “ San Miguel, / Pearl Isl. / Champion. [p] // B.C.A. Col. I.2. / Bidessus / curtulus, / Sharp. [p]” (all BMNH) .</p><p>Additional material examined. 7 specimens: “ Panama: Panama, Campana, / cascade below Anam office, / 270m, 19.vi.2007, / 08.40.818N 079.54.075W, / Balke (PAN 07) [p]” (CKSB, NMPC, ZSMG) .</p><p>Diagnosis. Small, ovoid-shaped species, only slightly attenuated posteriorly; lateral outline nearly continuous between pronotum and elytra in dorsal aspect (Figs 2 A–B). Surface shiny. Elytron rufopiceous with two testaceous maculae; subbasal macula transversely quadrangular, longitudinally extended at lateral margin, narrower basomedially, not reaching suture; posterior macula longitudinal-oblique, expanded medially in apical half, reaching elytral apex. Head finely reticulated and punctate. Pronotum widest at base; lateral bead distinct; finely punctate over entire surface, basal striae sinuated, distinctly curved inwards near base, length about 0.60× length of pronotum measured along same line as stria. Elytra with basal striae straight, slightly shorter than those on pronotum; sutural striae absent; epipleura with moderately oblique humeral carina. Metaventrite sparsely and rather finely punctate over entire surface; metacoxal plates with punctate medial and lateral portions, broadly impunctate along anterior and posterior margins (Fig. 2C). Abdominal ventrites I and II coarsely punctate; ventrite VI unmodified. Median lobe in lateral view expanded medially, apical third linear with lateral margins convergent to acute apex (Fig. 2E); in ventral view with attenuation in apical fifth, then evenly convergent to obtusely pointed apex; ventral surface in apical part with numerous scale-like setae (Fig. 2D). Lateral lobe with apical segment almost straight and parallel-sided, apex slightly protruding (Fig. 2F).</p><p>Neoclypeodytes curtulus is easily distinguished from the other two Panamanian species by the smaller body size (TL: 1.65–1.80 mm; TL-h: 1.45–1.60; TW: 0.90–1.00 mm), characteristic elytral color pattern (Figs 2 A–B), and the male genitalia with the median lobe expanded medially, and the apex of the lateral lobe protruding but less distinctly than in N. fortunensis sp. nov. (Figs 2 D–F).</p><p>Note. Miller (2001) describes the subbasal elytral maculae as triangular in N. curtulus, which is however not completely accurate: although the macula is usually somewhat attenuated basomedially, it is never acute triangular (cf. Figs 2 A–B).</p><p>Collection circumstances. The recently collected specimens were found together with N. balkei sp. nov. (see above).</p><p>Distribution. The species was described and known so far only from two islands in Pacific Gulf of Panama; here we present the first record from continental Central Panama (Fig. 6).</p></div>	https://treatment.plazi.org/id/03B387F5FF9BC16642C6FF65FC57F8DA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Scheers, Kevin;Hájek, Jiří	Scheers, Kevin, Hájek, Jiří (2020): Neoclypeodytes Young from Panama, with description of two new species (Coleoptera: Dytiscidae: Hydroporinae: Bidessini). Zootaxa 4890 (2): 245-256, DOI: 10.11646/zootaxa.4890.2.5
03B387F5FF99C16A42C6FF65FECBFC9B.text	03B387F5FF99C16A42C6FF65FECBFC9B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neoclypeodytes fortunensis Scheers & Hájek 2020	<div><p>Neoclypeodytes fortunensis sp. nov.</p><p>(Fig. 3)</p><p>Type material. Holotype: m#, “ PANAMA: CHIRIQUI PROVINCE / La Fortuna: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.22227&amp;materialsCitation.latitude=8.697267" title="Search Plazi for locations around (long -82.22227/lat 8.697267)">Rio Hornito Trail</a> / 08°41.836′N, 82°13.336′W, / 1165 m; montane forest, / stream pools, 3.ix.2017 / Fikáček, Hájek, Siedel &amp; Sekerka lgt. [p] // HO-LOTYPE m# / NEOCLYPEODYTES / fortunensis sp. nov. / K. Scheers &amp; J. Hájek det. 2020 [p, red label]” (NMPC) . Paratypes: 16 specimens; same label data as holotype (BMNH, CKSB, NMPC, ZSMG). Each paratype is provided with the appropriate red printed label .</p><p>Description of male holotype. Habitus (Fig. 3A). Ovoid-shaped, slightly attenuate anteriorly and more so posteriorly; lateral outline continuous between pronotum and elytra in dorsal aspect. Surface shiny.</p><p>Coloration. Head piceous. Antennae with antennomeres I–IV uniformly pale testaceous, antennomeres V–XI slightly infuscated apically. Palpi pale testaceous. Disc of pronotum dark rufous, distinctly and broadly darker piceous along anterior and posterior margins; sides pale testaceous. Elytron dark piceous with two testaceous maculae; anterior (subbasal) macula quadrangular, extending from lateral margin to one fifth of elytral width to suture, anterior margin convex, posterior margin concave; posterior macula not reaching suture, anteriorly not reaching lateral margin with emargination towards suture, posteriorly reaching lateral margin and apex of elytron; epipleuron completely testaceous. Legs including tarsi testaceous. Prosternum, mesepisterna and mesepimera rufotestaceous; metaventrite, metepisterna and metacoxae rufotestaceous, somewhat darkened at lateral portions. Abdominal ventrites rufotestaceous medially and broadly piceous anteriorly and laterally.</p><p>Head with fine but distinct microreticulation anterior of occipital line; punctation consisting of fine puncture near eyes and coarser punctures on clypeus; anterior clypeal margin medially distinctly flattened; occipital line narrowly separated from posterior margin of eyes.</p><p>Pronotum widest at base; lateral bead distinct; lateral margin nearly straight in basal third, very slightly round-ed in anterior two thirds; microreticulation absent; punctation consisting of punctures separated by 2–4× puncture diameter, punctation somewhat sparser laterally and distinctly denser near anterior margin; basal striae sinuate, distinctly curved inwards near base and slightly outwards anteriorly, length about 0.65× length of pronotum measured along same line as stria.</p><p>Elytra with basal striae very slightly and evenly curved, slightly shorter than those on pronotum; sutural striae absent; microreticulation absent; punctation similar to that of pronotal disc, with punctures slightly larger; epipleura finely punctate, with moderately oblique humeral carina; epipleural carina not visible in dorsal view.</p><p>Legs. Pro- and mesotibiae club shaped. Pro- and mesotarsomeres 1–3 slightly broadened, ventrally with adhesive setae. Metatarsomeres with well-developed natatorial setae on both sides.</p><p>Ventral surface (Fig. 3B). Prosternum medially narrowly convex, coarsely sculptured; prosternal process rath-er broad, subovate with apex broadly obtuse, lateral rim indistinct. Metaventrite sparsely and rather finely punctate over entire surface, punctures separated by 2–4× puncture diameter. Intralinear space of metacoxae broad (ML/MW = 2.15), sparsely and irregularly punctate with fine punctures, punctures separated by 2–4× puncture diameter; metacoxal plates distinctly punctate with moderately coarse punctures, punctures separated by 1–3× puncture diameter, plates broadly impunctate along anterior and posterior margins.Abdominal ventrites I and II coarsely punctate; succeeding ventrites smooth with a row of setae bearing punctures along posterior margin and some sparse and fine punctures scattered over surface; ventrites IV and V with a tuft of long setae medially; ventrite VI unmodified.</p><p>Male genitalia. Median lobe in ventral view broad and subparallel in basal half, slightly attenuating in third quarter of its length to short parallel subapical part, then attenuating to obtusely pointed apex (Fig. 3C); ventral surface in apical part with numerous scale-like setae (Fig. 3D). Median lobe in lateral view with ventral outline distinctly expanded medially, curved, apical third straight with lateral outline attenuating to pointed apex (Fig. 3E). Lateral lobe with apical segment elongated, very slightly curved; on inner side with numerous long setae; apex distinctly curved and protruding (Fig. 3F).</p><p>Variability. There is no significant variation present in the limited number of specimens available.</p><p>Sexual dimorphism. The females have the anterior clypeal margin less strongly modified but otherwise agree with the description of the male; pro- and mesotarsomeres not distinctly broadened.</p><p>Measurements. TL: 1.80–2.00 mm (mean value: 1.85 ± 0.05 mm); holotype: 1.90 mm. TL-h: 1.50–1.75 mm (mean value: 1.65 ± 0.05 mm); holotype: 1.70 mm. TW: 1.00– 1.15 mm (mean value: 1.10 ± 0.03 mm); holotype: 1.10 mm. Ratio TL/TW: 1.68–1.74 (mean value: 1.72 ± 0.02).</p><p>Differential diagnosis. Neoclypeodytes fortunensis sp. nov. is distinguished from N. balkei sp. nov. and N. curtulus by the color pattern (cf. Figs 1A, 2 A–B and 3A). Additionally, it differs from N. balkei sp. nov. by the male genitalia with the median lobe in lateral view expanded medially (Fig. 3E), apical part with numerous scale like setae and apex obtusely pointed in ventral view (Figs 3C, D), and the apical segment of the lateral lobe with the apex protruding and subapical incision distinct in N. fortunensis sp. nov. (Fig. 3F); from N. curtulus, it differs by the larger body size, and by the median lobe of aedeagus – in ventral view with distinct attenuation in apical fifth, and apical part less narrowed to apex in N. fortunensis sp. nov. (Fig. 3E).</p><p>The new species differs from the other species in the genus by the combination of the distinctive color pattern, the absence of sutural striae, the shape of the basal pronotal and elytral striae, and the male genitalia. Due to inaccurate interpretation of elytral subbasal macula in N. curtulus as triangular (couplet 19) in the identification key of Miller (2001), the new species falls into couplet 24, together with N. fryii and N. quadrinotatus . However, based on the overall similar shape of male genitalia, N. fortunensis sp. nov. is most closely related with N. curtulus; see above for distinguishing of both species.</p><p>Etymology. The new species is named after its area of occurrence – La Fortuna. The specific epithet is an adjective in the nominative singular.</p><p>Collection circumstances. The specimens of the type series were collected in remnant pools of a forest streamlet (Figs 5 A–B), and among gravel in the riverbed of the larger river Rio Hornito (Fig. 5C); the two collecting sites were ca. 100 m apart.</p><p>Distribution. To date only known from the type locality at the Rio Hornito in La Fortuna, Chiriqui province, Panama (Fig. 6).</p></div>	https://treatment.plazi.org/id/03B387F5FF99C16A42C6FF65FECBFC9B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Scheers, Kevin;Hájek, Jiří	Scheers, Kevin, Hájek, Jiří (2020): Neoclypeodytes Young from Panama, with description of two new species (Coleoptera: Dytiscidae: Hydroporinae: Bidessini). Zootaxa 4890 (2): 245-256, DOI: 10.11646/zootaxa.4890.2.5
