taxonID	type	description	language	source
03B387FDD252FFCF99CFFBDB33501513.taxon	materials_examined	Material examined. – Refer to subspecies accounts (see later). Comparative material. Geothelphusa miyazakii (Miyake & Minei) (all specimens collected in Taiwan): 1 male (holotype), CL 24.6 mm, ZLKU 10983, Alilao Village, Shi-Men district, Taipei County, coll. Jui-Kuang Chiu, 28 Jan. 1964; 1 female (paratype), CL 26.4 mm, ZLKU 10984, data same as holotype; 3 males CLs 20.0 – 23.2 mm, 3 females, CLs 20.8 – 28.3 mm, ZLKU 10137, Shenkeng, Taipei County, coll. Jui-Kuang Chiu, 3 Oct. 1963; 1 male, CL 21.3 mm, 1 female, CL 26.7 mm, ZLKU 10985, Shenkeng, Taipei County, coll. Jui-Kuang Chiu, 3 Oct. 1963; 2 males, CLs 22.1, 23.3 mm, 1 female, CL 16.2 mm, ZLKU 13752, Patoutzu, Keelung City, coll. Hsiang-Ping Yu, 22 Jun. 1972; 2 males, CLs 15.6, 18.8 mm, 1 female, CL 27.0 mm, RUMF-ZC- 76, Tinmu, Taipei City, coll. Jhy-Yun Shy, 2 Sep. 2001; 1 male, CL 23.6 mm, 1 female, CL 22.3 mm, RUMF- ZC- 77, Tinmu, Taipei City, coll. Jhy-Yun Shy, Kari Lee & Tohru Naruse, 18 Feb. 2002; 1 male, CL 23.0 mm, NTOU F 10094, National Taiwan Ocean University, Keelung City, coll. Ping-Ho Ho, 1 Jul. 1986; 1 male, CL 23.5 mm, 1 female, CL 22.5 mm, NTOU F 10095, Shuangshi, Shihfan, Shangsi, Taipei City, 1 Jun. 1982; 1 female, CL 26.3 mm, NTOU F 10096, Wanli / Kingshan, Chishiangshan, Taipei City, 17 Jun. 1984; 1 male, CL 11.9 mm, 1 female, CL 22.4 mm, NTOU F 10116, National Taiwan Ocean University, Keelung City, coll. Dar-Yi Wang, 1 Jan. 1989; 1 male, CL 21.4 mm, NTOU F 10091, Rayping, Taipei County, coll. Jhy-Yun Shy, 8 May. 1993; 3 males, CLs 12.8 – 19.4 mm 1 female, CL 20.9 mm, NTOU F 10317, Tinmu, Taipei City, coll. Jhy-Yun Shy, 4 Sep. 2001; 2 males, CLs 14.3, 18.5 mm, NMNS 3999 - 001, Luchiaokeng, Yanmingshan, Taipei City, coll. Yu-Hsi Wang, 25 Jul. 1997; 3 males, CLs 10.3 – 26.6 mm, 2 females, CLs 13.0, 15.1 mm, NMNS 3999 - 002, Luchiaokeng, Yanmingshan, Taipei City, coll. L. - S. Chen, 9 Jun. 1997; 1 male, CL 10.5 mm, TMCD 2225, Alilan stream, Taipei County, coll. Tsun-Hsien Wu, Dec. 1986; 1 female, CL 20.0 mm, TMCD 2041, Pingteng Li, Yangmingshan, Taipei City, coll. Chiu-Chu Lu, 20 Jul. 1983; 1 male, CL 14.9 mm, TMCD 2043, Nankung, Taipei City, coll. Yuang Wei, 5 Aug. 1984; 1 male, CL 24.8 mm, 1 femlae, CL 21.9 mm, ZRC 1995.606, Yangmingshan, Taipei City, coll. C. - J. Leu, 26 Jul. 1983; 1 male, CL 25.3 mm, 1 female, CL 21.7 mm, ZRC 1998.213, Keelung City, 1 Jul. 1986. Geothelphusa nanshi Shy, Ng & Yu: male (holotype), CL 18.4 mm, NTOU F 10198, Nanshi, Tainan, Taiwan, coll. Jhy-Yun Shy & W. L. Tsay, 7 Aug. 1992. Geothelphusa shokitai Shy & Ng: male (holotype), CL 20.5 mm, NSMT-Cr 15176, Uotsuri Island, Senkaku Group, coll. Masatsugu Yokota, 28 May. 1991; 1 female (paratype), CL 29.7 mm, NSMT-Cr 15177, data same as holotype.	en	Naruse, Tohru, Shokita, Shigemitsu (2004): A New Species Of The Freshwater Crab, Previously Assigned To Geothelphusa Miyazakii (Miyake & Chiu, 1965) (Crustacea: Decapoda: Potamidae), From Yaeyama Group, Southern Ryukyus, Japan. Raffles Bulletin of Zoology 52 (2): 555-562, DOI: 10.5281/zenodo.13244414
03B387FDD252FFCF99CFFBDB33501513.taxon	diagnosis	Diagnosis. – Carapace slightly convex longitudinally and transversely; anterolateral margin cristated, convex, faintly granulated, external orbital angle blunt, epibranchial tooth represented only by small granules. Eyes with relatively small cornea, subdistal part shorter to almost same height with base of peduncle in frontal view. Ambulatory legs relatively long, total length of second ambulatory legs more than two times as long as CL. Male telson bell-shaped, tip of telson reaching imaginary line joining posterior 2 / 3 of the coxae of chelipeds. G 1 sinuous to straight, proximal half of subterminal segment and terminal segment directing inward, and distal half of subterminal segment directing outward; G 1 slender, subterminal segment with a knob on proximal outer margin, with rounded lobe on proximal inner margin; synovial membrane variable in shape, distal margin forked, inner margin just swollen laterally or hooked proximally.	en	Naruse, Tohru, Shokita, Shigemitsu (2004): A New Species Of The Freshwater Crab, Previously Assigned To Geothelphusa Miyazakii (Miyake & Chiu, 1965) (Crustacea: Decapoda: Potamidae), From Yaeyama Group, Southern Ryukyus, Japan. Raffles Bulletin of Zoology 52 (2): 555-562, DOI: 10.5281/zenodo.13244414
03B387FDD252FFCF99CFFBDB33501513.taxon	description	Description. – Carapace slightly convex longitudinally and transversely; orbital and frontal margins cristated, infraorbital margin faintly granulated, frontal margin slightly concave medially, anterolateral margin cristated, convex, faintly granulated, present on anterior 2 / 5 of carapace, external orbital angle blunt, epibranchial tooth represented only by small granules; postorbital and postfrontal cristae low, gradually raised backward, anterolateral region of carapace with several oblique ridges and small pits, posterolateral region with short carinae; H-shaped gastric groove distinct, two large pits on anterior outer part of H-shaped gastric groove, on either side of cervical groove, one large pit located halfway between Hshaped gastric groove and epibranchial tooth, inside of cervical groove, branchial groove represented by wide and shallow transverse hollow. Eyes with relatively small cornea, subdistal part shorter to almost same height with base of peduncle in frontal view. Mouth parts similar to those of Geothelphusa miyakoensis Shokita, Naruse & Fujii, 2002, except for longer distal segment of mandibular palp and longer coxa of maxilla. Male chelae asymmetric in size and shape, outer surface of manus of major chela smooth, lower margin rough. Ambulatory legs relatively long, total length of second ambulatory legs more than two times as long as CL; propodus with elliptic cross-section and four longitudinal margins, only anterior dorsal margin lacking row of spines; propodus of first and second ambulatory legs lined by long setae along outer, inner margins, and along midline of ventral surface; dactylus with rectangular cross-section and four longitudinal margins, distal spine of outer dorsal margin of fourth ambulatory leg thin, sharp, locating about middle of dorsal surface. Male telson bell-shaped, tip of telson reaching imaginary line joining posterior 2 / 3 of the coxae of chelipeds. G 1 sinuous to straight, proximal half of subterminal segment and terminal segment directing inward, and distal half of subterminal segment directing outward; G 1 slender, subterminal segment with a knob on proximal outer margin, with rounded lobe on proximal inner margin; synovial membrane variable in shape, distal margin forked, inner margin just swollen laterally or hooked proximally. Habitat. – Geothelphusa marginata is formed in shallow, narrow, slow-flowing streams and in small muddy swamps in forests. Crabs were collected under large stones and from burrows.	en	Naruse, Tohru, Shokita, Shigemitsu (2004): A New Species Of The Freshwater Crab, Previously Assigned To Geothelphusa Miyazakii (Miyake & Chiu, 1965) (Crustacea: Decapoda: Potamidae), From Yaeyama Group, Southern Ryukyus, Japan. Raffles Bulletin of Zoology 52 (2): 555-562, DOI: 10.5281/zenodo.13244414
03B387FDD252FFCF99CFFBDB33501513.taxon	distribution	Distribution. – Ishigaki and Iriomote Islands, Yaeyama Group, Southern Ryukyus, Japan.	en	Naruse, Tohru, Shokita, Shigemitsu (2004): A New Species Of The Freshwater Crab, Previously Assigned To Geothelphusa Miyazakii (Miyake & Chiu, 1965) (Crustacea: Decapoda: Potamidae), From Yaeyama Group, Southern Ryukyus, Japan. Raffles Bulletin of Zoology 52 (2): 555-562, DOI: 10.5281/zenodo.13244414
03B387FDD252FFCF99CFFBDB33501513.taxon	etymology	Etymology. – Geothelphusa marginata is named for the Latin “ marginatum ”, meaning “ margined ”, because of the distinctly convex anterolateral margins of carapace.	en	Naruse, Tohru, Shokita, Shigemitsu (2004): A New Species Of The Freshwater Crab, Previously Assigned To Geothelphusa Miyazakii (Miyake & Chiu, 1965) (Crustacea: Decapoda: Potamidae), From Yaeyama Group, Southern Ryukyus, Japan. Raffles Bulletin of Zoology 52 (2): 555-562, DOI: 10.5281/zenodo.13244414
03B387FDD252FFCF99CFFBDB33501513.taxon	discussion	Remarks. – Miyake & Chiu (1965) originally described Geothelphusa miyazakii on the basis of specimens obtained from Alilao Village, Shimen District, Taipei County, Taiwan. Minei (1973) was the first to report on specimens from Ishigaki and Iriomote Islands, Southern Ryukyus, which he identified as G. miyazakii. Later, Shy et al. (1994) doubted whether the specimens from Taiwan and Southern Ryukyuan were conspecific. The present study has idetified a number of differences between G. marginata and G. miyazakii sensu stricto, viz., the proximal inner margin of the subterminal segment of G 1 (roundly swollen in G. marginata vs. straight in G. miyazakii sensu stricto), the size of the knob on the proximal outer margin of G 1 (small vs. large), and the size of the epibranchial tooth (rudiment vs. absent). In addition, there are significant differences between G. marginata and G. miyazaki i in the ratio of the total length of the second ambulatory leg to CL, and in the ratio of the length to width of the propodus of the second ambulatory legs (Tables 1, 2). Shy et al. (1994) remarked that the terminal segment of the G 1 of G. marginata is slender and more elongated than G. miyazakii sensu stricto. However, the ratio of the total length to width of G 1 of G. miyazakii sensu stricto is in almost the same range as G. marginata, and the length to the width ratio of the terminal segment of G 1 is wider than in G. marginata (Table 1). The respective populations of G. marginata occurring on Ishigaki and Iriomote Islands differ slightly from one another in morphology and live colouration. We regard these differences as infraspecific in nature because each population does not show any differences in the shape of G 1, which is reproductively and taxonomically most important character. As such, we regard two subspecies for the two islands. The subspecific accounts follow (see later). Geothelphusa marginata also resembles G. nanshi Shy, Ng & Yu, 1994. However, G. marginata can be distinguished from G. nanshi by the shape of G 1 (sinuous in G. marginata vs. gently curved outward in G. nanshi), the shape of the proximal inner margin of the subterminal segment of G 1 (roundly swollen vs. straight), the shape of the external orbital angle (blunt vs. sharp), the form of the anterolateral margin (barely granulated vs. certainly granulated), the condition of the cervical groove (distinct vs. invisible), and the accessory of the propodus of the second ambulatory leg (three rows of spines vs. only two rows of spines on the inner margins). Geothelphusa marginata is also close to G. shokitai Shy & Ng. However, G marginata is easily differentiated from G. shokitai by the shape of G 1 (sinuous in G. marginata vs. almost straight in G. shokitai), the shape of the dorsal surface of the carapace (convex vs. almost flat), the form of the anterolateral margin (very low granules vs. distinct granules), the shape of the external orbital angle (blunt vs. acute), and the shape and the relative position of the male telson (tip narrow, reaching imaginary line joining proximal 2 / 3 of the coxae of the chelipeds vs. tip obtuse, reaching imaginary line joining proximal 1 / 2 of the coxae of the chelipeds).	en	Naruse, Tohru, Shokita, Shigemitsu (2004): A New Species Of The Freshwater Crab, Previously Assigned To Geothelphusa Miyazakii (Miyake & Chiu, 1965) (Crustacea: Decapoda: Potamidae), From Yaeyama Group, Southern Ryukyus, Japan. Raffles Bulletin of Zoology 52 (2): 555-562, DOI: 10.5281/zenodo.13244414
03B387FDD253FFC99A9EF9C934491153.taxon	materials_examined	Material examined. – All specimens of G. m. marginata were collected in Ishigaki Island, Yaeyama Group, Southern Ryukyus, Japan. Holotype – male, CL 27.7 mm, RUMF-ZC- 59, a stream of Mt. Omoto-Dake (type locality), coll. Tohru Naruse & Yoko Okamoto, 11 Jul. 2001. Paratypes – 2 females, CLs 27.7, 27.9 mm, RUMF-ZC- 87, data same as holotype; 2 males, CLs 22.4, 26.8 mm, 1 female, CL 28.0 mm, NSMT-Cr 15172, upstream of Sukubaru Dam Reserve, coll. Tohru Naruse, 9 Aug. 2001; 1 male, CL 26.7 mm, 1 female, CL 26.0 mm, NSMT-Cr 15173, a stream of Mt. Omoto-Dake, coll. Tohru Naruse, 11 Aug. 2001; 1 male, CL 29.9 mm, CBM-ZC 6723, a stream of Mt. Omoto-Dake, coll. Hsi-Te Shih & Tohru Naruse, 9 Apr. 2002; 2 females, CLs 28.3 32.1 mm, CBM-ZC 6724, a stream of Mt. Omoto-Dake, coll. Ryoko Segawa, 19 Mar. 1997. Others – 2 males, CLs 21.7, 22.2 mm, RUMF-ZC- 60, a stream of Mt. Omoto-Dake, coll. Ryoko Segawa, 19 Mar. 1997; 3 males, CLs 14.4 – 20.9 mm, 1 female, CL 21.1 mm, ZRC 2003.0596, a stream of Mt. Omoto-Dake, coll. Tohru Naruse, 22 Mar. 2001; 1 male, CL 26.2 mm, 2 females, CLs 25.6, 27.2 mm, ZLKU 12648, a stream of Mt. Omoto-Dake, coll. Y. Hashiguchi, 31 Jul. 1968; 3 males, CLs 19.2 – 27.1 mm, 3 females, CLs 20.1 – 23.4 mm, ZLKU m 1092, a stream of Mt. Omoto-Dake, coll. Y. Hashiguchi, 31 Jul. 1968; 1 male, CL 27.5 mm, HMNH-C 9, Ohama, coll. Hidenori Yoshigou, 11 Aug. 1999.	en	Naruse, Tohru, Shokita, Shigemitsu (2004): A New Species Of The Freshwater Crab, Previously Assigned To Geothelphusa Miyazakii (Miyake & Chiu, 1965) (Crustacea: Decapoda: Potamidae), From Yaeyama Group, Southern Ryukyus, Japan. Raffles Bulletin of Zoology 52 (2): 555-562, DOI: 10.5281/zenodo.13244414
03B387FDD253FFC99A9EF9C934491153.taxon	diagnosis	Diagnosis. – Carapace with convex anterolateral margin, frontal margin slightly concave medially, cervical groove distinct as Geothelphusa species, barely connected from distinct H-shaped gastric groove to epibranchial tooth. G 1 sinuous, subterminal segment with small knob on proximal outer margin, proximal inner margin roundly swollen. Carapace, chelipeds, and ambulatory legs purple, fingers of chelipeds, articulations of ambulatory, and about distal half of dactylus yellow / orange.	en	Naruse, Tohru, Shokita, Shigemitsu (2004): A New Species Of The Freshwater Crab, Previously Assigned To Geothelphusa Miyazakii (Miyake & Chiu, 1965) (Crustacea: Decapoda: Potamidae), From Yaeyama Group, Southern Ryukyus, Japan. Raffles Bulletin of Zoology 52 (2): 555-562, DOI: 10.5281/zenodo.13244414
03B387FDD253FFC99A9EF9C934491153.taxon	distribution	Distribution. – Ishigaki Island, Yaeyama Group, Southern Ryukyus, Japan.	en	Naruse, Tohru, Shokita, Shigemitsu (2004): A New Species Of The Freshwater Crab, Previously Assigned To Geothelphusa Miyazakii (Miyake & Chiu, 1965) (Crustacea: Decapoda: Potamidae), From Yaeyama Group, Southern Ryukyus, Japan. Raffles Bulletin of Zoology 52 (2): 555-562, DOI: 10.5281/zenodo.13244414
03B387FDD253FFC99A9EF9C934491153.taxon	discussion	Remarks. – Geothelphusa marginata marginata, can be easily differentiated from G. m. fulva by their live colouration. However, morphological differences are limited to the size of the proximal outer knob of the subterminal segment of G 1 (small in G. m. marginata vs. large in G. m. fulva), the condition of the cervical groove (barely connected from Hshaped gastric groove to epibranchial tooth vs. visible between H-shaped gastric region to hepatic region), and the shape of the frontal margin (slightly concaved medially vs. almost straight). Because G. m. marginata and G. m. fulva do not have large differences even in G 1, subspecific status are considered for these two taxa. Geothelphusa marginata marginata has a similar colouration to G. miyazakii, but the former tend to have softer colour than G. miyazakii. Furthermore, G. miyazakii has bright a, G. m. marginata (holotype, RUMF-ZC- 59), dorsal view; b, G. m. marginata (holotype, RUMF-ZC- 59), ventral view; c, G. m. fulva (paratype, RUMF-ZC- 75); d, G. miyazakii (RUMF-ZC- 76). yellow rings on articulation of ambulatory legs and their dactylus are totally yellow. Yoshigou (1999) recorded Geothelphusa sp. 2 from Ishigaki and Iriomote Islands and G. sp. 3 from Iriomote Island. As far as I examined his materials, G. sp. 2 and G. sp. 3 correspond to G. m. marginata and G. m. fulva, respectively. Geothelphusa sp. 2 of Iriomote Island is most likely a misidentification due the choice of unsuitable characters, such as relative width of the cornea and the direction of the projection of anterolateral margin. Furthermore, Yoshigou (1999) reported ambulatory leg proportions that are beyond the values found in the present study (Table 3).	en	Naruse, Tohru, Shokita, Shigemitsu (2004): A New Species Of The Freshwater Crab, Previously Assigned To Geothelphusa Miyazakii (Miyake & Chiu, 1965) (Crustacea: Decapoda: Potamidae), From Yaeyama Group, Southern Ryukyus, Japan. Raffles Bulletin of Zoology 52 (2): 555-562, DOI: 10.5281/zenodo.13244414
03B387FDD255FFCB9992FD0936D913F3.taxon	materials_examined	Material examined. – All specimens of G. m. fulva were collected in Iriomote Island, Southern Ryukyus, Japan. Holotype – male, CL 30.7 mm, RUMF-ZC- 62, upstream of Nakama River (near Nishifunatsuki, Ohtomi forest road) (type locality), coll. Tohru Naruse, 12 Aug. 2001. Paratypes – 2 males, CLs 18.2, 20.1 mm, 2 females, CLs 17.0, 30.0 mm, RUMF-ZC- 88, data same as holotype; 1 male, CL 22.8 mm, 1 female, CL 25.2 mm, NTOU F 10220, data same as holotype; 1 male, CL 22.8 mm, 1 female, CL 24.0 mm, NSMT-Cr 15174, coll. Masayuki Osawa, Near Nadara Bridge, 4 Dec. 2000; 1 male, CL 28.6 mm, NSMT-Cr 15175, Aira River, coll. Shigemitsu Shokita, Jul. 2001; 1 female, CL 31.1 mm, CBM-ZC 4594, Near Kanpireh Waterfall, Urauchi River, coll. Tomoyuki Komai, 3 Jul. 1998; 1 male, CL 23.3 mm, CBM-ZC 6725, Ohmija River, coll. Tohru Naruse, Mohamed Idha Salim, Yoshihisa Fujita and Akane Ito, 19 Mar. 2001. Others – 4 males, CLs 17.7 – 25.0 mm, RUMF-ZC- 63, upstream of Nishifunatsuki, Nakama River, coll. Tohru Naruse & Mohamed Idha Salim, 12 Mar. 2001; 1 female, CL 31.3 mm, RUMF-ZC- 64, Midara River, coll. Tohru Naruse & Mohamed Idha Salim, 14 Mar. 2001; 3 males, CLs 15.7 – 22.2 mm, RUMF-ZC- 65, Ohmija River, coll. Tohru Naruse, 16 Mar. 2001; 2 males, CL 15.2, 21.5 mm, 1 female, CL 22.6 mm, RUMF-ZC- 66, Cape Kasa-saki, coll. Tohru Naruse, 17 Mar. 2001; 1 male, CL 20.9 mm, 2 females, CLs 17.0, 25.5 mm, RUMF-ZC- 67, upstream of Nishifunatsuki, Nakama River, coll. Tohru Naruse, 3 Dec. 1999; 1 female, CL 24.1 mm, RUMF-ZC- 68, Shirahama forest road, coll. Tohru Naruse, 5 Aug. 2001; 1 male, CL 15.2 mm, RUMF-ZC- 69, Midara River, coll. Tohru Naruse, 6 Aug. 2001; 1 male, CL 21.4 mm, RUMF-ZC- 70, Urauchi River (near the confruence with Itajiki River), coll. Tohru Naruse, 12 Oct. 1999; 1 female, CL 21.9 mm, RUMF-ZC- 71, Urauchi River (near the confluence with Itajiki River), coll. Tohru Naruse, 13 Oct. 1999; 1 female, CL 19.9 mm, RUMF-ZC- 72, upstream of Nishifunatsuki, Nakama River, coll. Tohru Naruse, 30 Nov. 1999; 1 male, CL 23.8 mm, RUMF-ZC- 73, Urauchi River, coll. Chieko Matsumoto, 29 Apr. 2002; 1 male, CL 19.3 mm, RUMF-ZC- 74, Shirahama forest road, coll. Shinichi Watanabe & Nozomi Nakanishi, 25 Apr. 2002; 2 males, CLs 14.8, 21.8 mm 2 females, CLs 19.8, 27.1 mm, ZRC 2003.0597, a stream near Kanokawa Bay, coll. Tohru Naruse, 30 Dec. 2001; 5 males, CLs 19.2 – 30.0, 2 females, CLs 18.3, 19.9 mm, ZLKU m 1117, Urauchi River, coll. Shigemitsu Shokita, 2 May. 1971; 1 female, CL 25.81 mm, HMNH-C 81, Urauchi River (near the confluence with Itajiki R.), coll. Hidenori Yoshigou, 17 Aug. 2000.	en	Naruse, Tohru, Shokita, Shigemitsu (2004): A New Species Of The Freshwater Crab, Previously Assigned To Geothelphusa Miyazakii (Miyake & Chiu, 1965) (Crustacea: Decapoda: Potamidae), From Yaeyama Group, Southern Ryukyus, Japan. Raffles Bulletin of Zoology 52 (2): 555-562, DOI: 10.5281/zenodo.13244414
03B387FDD255FFCB9992FD0936D913F3.taxon	diagnosis	Diagnosis. – Carapace with convex anterolateral margin, frontal margin straight, cervical groove visible between Hshaped gastric region to hepatic region. G 1 sinuous to straight, subterminal segment with large knob on proximal outer margin, and proximal inner margin roundly swollen. Carapace and ambulatory legs dark brown to yellowish brown, outer surface of chelae orange, brown, or reddish, and inner surface of manus and fingers white to yellow.	en	Naruse, Tohru, Shokita, Shigemitsu (2004): A New Species Of The Freshwater Crab, Previously Assigned To Geothelphusa Miyazakii (Miyake & Chiu, 1965) (Crustacea: Decapoda: Potamidae), From Yaeyama Group, Southern Ryukyus, Japan. Raffles Bulletin of Zoology 52 (2): 555-562, DOI: 10.5281/zenodo.13244414
03B387FDD255FFCB9992FD0936D913F3.taxon	distribution	Distribution. – Iriomote Island, Yaeyama Group, Southern Ryukyus, Japan.	en	Naruse, Tohru, Shokita, Shigemitsu (2004): A New Species Of The Freshwater Crab, Previously Assigned To Geothelphusa Miyazakii (Miyake & Chiu, 1965) (Crustacea: Decapoda: Potamidae), From Yaeyama Group, Southern Ryukyus, Japan. Raffles Bulletin of Zoology 52 (2): 555-562, DOI: 10.5281/zenodo.13244414
03B387FDD255FFCB9992FD0936D913F3.taxon	etymology	Etymology. – Geothelphusa marginata fulva is named after the Latin “ fulvus ” meaning “ brown colour ”, because of their colouration.	en	Naruse, Tohru, Shokita, Shigemitsu (2004): A New Species Of The Freshwater Crab, Previously Assigned To Geothelphusa Miyazakii (Miyake & Chiu, 1965) (Crustacea: Decapoda: Potamidae), From Yaeyama Group, Southern Ryukyus, Japan. Raffles Bulletin of Zoology 52 (2): 555-562, DOI: 10.5281/zenodo.13244414
03B387FDD255FFCB9992FD0936D913F3.taxon	discussion	Remarks. – See “ Remarks ” of Geothelphusa marginata marginata.	en	Naruse, Tohru, Shokita, Shigemitsu (2004): A New Species Of The Freshwater Crab, Previously Assigned To Geothelphusa Miyazakii (Miyake & Chiu, 1965) (Crustacea: Decapoda: Potamidae), From Yaeyama Group, Southern Ryukyus, Japan. Raffles Bulletin of Zoology 52 (2): 555-562, DOI: 10.5281/zenodo.13244414
