taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03B30955FFC2FFE8FF0C470DFDDEF8D6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13302877/files/figure.png	https://doi.org/10.5281/zenodo.13302877	Figure 12. Holotype of L. dysmicus (MHNG 1005.72) from Foulassi, South Region, Cameroon. A, Ventral and dorsal view of body. B, Details of cloacal region. C, Details of left pes. D, Detailed view of head in both lateral views and ventral and dorsal views (from top to bottom). Photos by J.L.R.	Figure 12. Holotype of L. dysmicus (MHNG 1005.72) from Foulassi, South Region, Cameroon. A, Ventral and dorsal view of body. B, Details of cloacal region. C, Details of left pes. D, Detailed view of head in both lateral views and ventral and dorsal views (from top to bottom). Photos by J.L.R.	2023-09-23	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli		Zenodo	biologists	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli			
03B30955FFC2FFE8FF0C470DFDDEF8D6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13302850/files/figure.png	https://doi.org/10.5281/zenodo.13302850	Figure 5. Gular and skull osteology comparisons between holotype material or representative material of all members of the L. gutturalis subgroup and the L. angularis group mentioned in this work.From top to bottom, ventral view gular section; dorsal, lateral, and ventral view of the skull; lateral, dorsal, medial, and ventral view of jaw; and number of tooth loci. PM = premaxilla, MX(R) = right maxilla, MX(L) = left maxilla, DN(R) = right dentary of the jaw.	Figure 5. Gular and skull osteology comparisons between holotype material or representative material of all members of the L. gutturalis subgroup and the L. angularis group mentioned in this work.From top to bottom, ventral view gular section; dorsal, lateral, and ventral view of the skull; lateral, dorsal, medial, and ventral view of jaw; and number of tooth loci. PM = premaxilla, MX(R) = right maxilla, MX(L) = left maxilla, DN(R) = right dentary of the jaw.	2023-09-23	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli		Zenodo	biologists	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli			
03B30955FFC2FFE8FF0C470DFDDEF8D6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13302854/files/figure.png	https://doi.org/10.5281/zenodo.13302854	Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi).	Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi).	2023-09-23	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli		Zenodo	biologists	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli			
03B30955FFC0FFE4FF7147FBFDD3F87F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13302879/files/figure.png	https://doi.org/10.5281/zenodo.13302879	Figure 13. Dorsolateral and ventral variation of two male specimens [RMCA R.8575/A (left) and RMCA R.8575/B (right)] of L. depressus, collected in sympatry at Flandria, Équateur Province, DRC.	Figure 13. Dorsolateral and ventral variation of two male specimens [RMCA R.8575/A (left) and RMCA R.8575/B (right)] of L. depressus, collected in sympatry at Flandria, Équateur Province, DRC.	2023-09-23	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli		Zenodo	biologists	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli			
03B30955FFC0FFE4FF7147FBFDD3F87F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13302883/files/figure.png	https://doi.org/10.5281/zenodo.13302883	Figure 14. Holotype of L. depressus (AMNH 10345) from Medje, Ituri Forest, Haut-Uele Province, DRC. A, Dorsal and ventral view of body. B, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom).C, Details of cloacal region.D, Details of left manus.Photos provided by Lauren Vonnahme:American Museum of Natural History, New York City (AMNH).	Figure 14. Holotype of L. depressus (AMNH 10345) from Medje, Ituri Forest, Haut-Uele Province, DRC. A, Dorsal and ventral view of body. B, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom).C, Details of cloacal region.D, Details of left manus.Photos provided by Lauren Vonnahme:American Museum of Natural History, New York City (AMNH).	2023-09-23	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli		Zenodo	biologists	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli			
03B30955FFC0FFE4FF7147FBFDD3F87F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13302887/files/figure.png	https://doi.org/10.5281/zenodo.13302887	Figure 15. Specimens of L. depressus in life or freshly euthanized from (A) Npenda, Équateur Province, DRC (UTEP 22580), (B) Katopa, Maniema Province, DRC (UTEP 22597), (C) Lake Tumba, Équateur Province, DRC (UTEP 22578), and (E–G) Barra do Cuanza, Cuanza Sul Province, Angola (MNCN 50772). Photo of habitat at (D) Lake Tumba, Équateur Province, DRC and (H) Barra do Cuanza, Cuanza Sul Province, Angola. Photographs by E.G.(A–C), C.K. (D), P.V.P.(E–G), and J.L.R. (H).	Figure 15. Specimens of L. depressus in life or freshly euthanized from (A) Npenda, Équateur Province, DRC (UTEP 22580), (B) Katopa, Maniema Province, DRC (UTEP 22597), (C) Lake Tumba, Équateur Province, DRC (UTEP 22578), and (E–G) Barra do Cuanza, Cuanza Sul Province, Angola (MNCN 50772). Photo of habitat at (D) Lake Tumba, Équateur Province, DRC and (H) Barra do Cuanza, Cuanza Sul Province, Angola. Photographs by E.G.(A–C), C.K. (D), P.V.P.(E–G), and J.L.R. (H).	2023-09-23	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli		Zenodo	biologists	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli			
03B30955FFC0FFE4FF7147FBFDD3F87F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13302895/files/figure.png	https://doi.org/10.5281/zenodo.13302895	Figure 16. Dorsal and ventral views of head showing gular ornamentation and pholidosis variation in L. depressus. Red points denote males and blue points denote females.	Figure 16. Dorsal and ventral views of head showing gular ornamentation and pholidosis variation in L. depressus. Red points denote males and blue points denote females.	2023-09-23	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli		Zenodo	biologists	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli			
03B30955FFC0FFE4FF7147FBFDD3F87F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13302837/files/figure.png	https://doi.org/10.5281/zenodo.13302837	Figure 1. Maximum likelihood analysis based on 532 bp of the mitochondrial 16S rRNA gene, for all available samples of the Afro-American Lygodactylus Clade (Clade C of Gippner et al. 2021). Support values (ML BS = Maximum Likelihood bootstrap values; BI PP = Bayesian Inference posterior probabilities) are shown graphically at the nodes according to the colours shown in the inset key.	Figure 1. Maximum likelihood analysis based on 532 bp of the mitochondrial 16S rRNA gene, for all available samples of the Afro-American Lygodactylus Clade (Clade C of Gippner et al. 2021). Support values (ML BS = Maximum Likelihood bootstrap values; BI PP = Bayesian Inference posterior probabilities) are shown graphically at the nodes according to the colours shown in the inset key.	2023-09-23	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli		Zenodo	biologists	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli			
03B30955FFC0FFE4FF7147FBFDD3F87F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13302854/files/figure.png	https://doi.org/10.5281/zenodo.13302854	Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi).	Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi).	2023-09-23	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli		Zenodo	biologists	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli			
03B30955FFC0FFE4FF7147FBFDD3F87F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13302850/files/figure.png	https://doi.org/10.5281/zenodo.13302850	Figure 5. Gular and skull osteology comparisons between holotype material or representative material of all members of the L. gutturalis subgroup and the L. angularis group mentioned in this work.From top to bottom, ventral view gular section; dorsal, lateral, and ventral view of the skull; lateral, dorsal, medial, and ventral view of jaw; and number of tooth loci. PM = premaxilla, MX(R) = right maxilla, MX(L) = left maxilla, DN(R) = right dentary of the jaw.	Figure 5. Gular and skull osteology comparisons between holotype material or representative material of all members of the L. gutturalis subgroup and the L. angularis group mentioned in this work.From top to bottom, ventral view gular section; dorsal, lateral, and ventral view of the skull; lateral, dorsal, medial, and ventral view of jaw; and number of tooth loci. PM = premaxilla, MX(R) = right maxilla, MX(L) = left maxilla, DN(R) = right dentary of the jaw.	2023-09-23	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli		Zenodo	biologists	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli			
03B30955FFC0FFE4FF7147FBFDD3F87F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13302840/files/figure.png	https://doi.org/10.5281/zenodo.13302840	Figure 2. A, Multilocus species tree and Bayesian chronogram of divergences among Afro-American Lygodactylus clade (Clade C of Gippner et al. 2021), based on an alignment of 13 concatenated markers (including eight nuclear-encoded genes, four protein-coding mitochondrial genes, and the 16S mitochondrial rRNA gene, see Gippner et al.2021). Branches with posterior probability> 0.9 are denoted by asterisks (*) at relevant nodes. Blue bars depict 95% High Probability Density (HPD) intervals of estimated divergence dates.B, Median-joining nuclear allele network showing the relationships between the L. picturatus-fischeri groups, inferred from the RAG1 nuclear gene (262 bp including all available RAG1 sequences of the L. picturatus-fischeri groups).C, Median-joining nuclear allele network of the RAG1 nuclear gene (980 bp) showing the relationships within the L. gutturalis subgroup, excluding L. gutturalis s.s. In all networks, circle frequency of alleles and small lines represent mutational steps.	Figure 2. A, Multilocus species tree and Bayesian chronogram of divergences among Afro-American Lygodactylus clade (Clade C of Gippner et al. 2021), based on an alignment of 13 concatenated markers (including eight nuclear-encoded genes, four protein-coding mitochondrial genes, and the 16S mitochondrial rRNA gene, see Gippner et al.2021). Branches with posterior probability> 0.9 are denoted by asterisks (*) at relevant nodes. Blue bars depict 95% High Probability Density (HPD) intervals of estimated divergence dates.B, Median-joining nuclear allele network showing the relationships between the L. picturatus-fischeri groups, inferred from the RAG1 nuclear gene (262 bp including all available RAG1 sequences of the L. picturatus-fischeri groups).C, Median-joining nuclear allele network of the RAG1 nuclear gene (980 bp) showing the relationships within the L. gutturalis subgroup, excluding L. gutturalis s.s. In all networks, circle frequency of alleles and small lines represent mutational steps.	2023-09-23	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli		Zenodo	biologists	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli			
03B30955FFCCFFE2FC6E4462FF66F92B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13302897/files/figure.png	https://doi.org/10.5281/zenodo.13302897	Figure 17. Holotype of Lygodactylus kibera sp. nov. (UTEP 22566) from Mpishi, near Kibira National Park, Bubanza Province, Burundi. A, Dorsal and ventral view of body. B, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Details of cloacal region (C), (D) tail tip, and (E) left manus.	Figure 17. Holotype of Lygodactylus kibera sp. nov. (UTEP 22566) from Mpishi, near Kibira National Park, Bubanza Province, Burundi. A, Dorsal and ventral view of body. B, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Details of cloacal region (C), (D) tail tip, and (E) left manus.	2023-09-23	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli		Zenodo	biologists	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli			
03B30955FFCCFFE2FC6E4462FF66F92B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13302899/files/figure.png	https://doi.org/10.5281/zenodo.13302899	Figure 18. Specimens of Lygodactylus kibera sp. nov. in life or freshly euthanized from (A) Mpishi (holotype: UTEP 22566), near Kibira National Park, Bubanza Province, Burundi, (B) N'Komo River (UTEP 22586), South Kivu Province, DRC, and (C) Bujumbura city, Bujumbura Mairie Province (UTEP 22576), Burundi.Photo of habitat at (D) Mpishi, Bubanza Province, Burundi. Photographs by E.G. (A–C) and C.K. (D).	Figure 18. Specimens of Lygodactylus kibera sp. nov. in life or freshly euthanized from (A) Mpishi (holotype: UTEP 22566), near Kibira National Park, Bubanza Province, Burundi, (B) N'Komo River (UTEP 22586), South Kivu Province, DRC, and (C) Bujumbura city, Bujumbura Mairie Province (UTEP 22576), Burundi.Photo of habitat at (D) Mpishi, Bubanza Province, Burundi. Photographs by E.G. (A–C) and C.K. (D).	2023-09-23	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli		Zenodo	biologists	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli			
03B30955FFCCFFE2FC6E4462FF66F92B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13302854/files/figure.png	https://doi.org/10.5281/zenodo.13302854	Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi).	Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi).	2023-09-23	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli		Zenodo	biologists	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli			
03B30955FFCCFFE2FC6E4462FF66F92B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13302840/files/figure.png	https://doi.org/10.5281/zenodo.13302840	Figure 2. A, Multilocus species tree and Bayesian chronogram of divergences among Afro-American Lygodactylus clade (Clade C of Gippner et al. 2021), based on an alignment of 13 concatenated markers (including eight nuclear-encoded genes, four protein-coding mitochondrial genes, and the 16S mitochondrial rRNA gene, see Gippner et al.2021). Branches with posterior probability> 0.9 are denoted by asterisks (*) at relevant nodes. Blue bars depict 95% High Probability Density (HPD) intervals of estimated divergence dates.B, Median-joining nuclear allele network showing the relationships between the L. picturatus-fischeri groups, inferred from the RAG1 nuclear gene (262 bp including all available RAG1 sequences of the L. picturatus-fischeri groups).C, Median-joining nuclear allele network of the RAG1 nuclear gene (980 bp) showing the relationships within the L. gutturalis subgroup, excluding L. gutturalis s.s. In all networks, circle frequency of alleles and small lines represent mutational steps.	Figure 2. A, Multilocus species tree and Bayesian chronogram of divergences among Afro-American Lygodactylus clade (Clade C of Gippner et al. 2021), based on an alignment of 13 concatenated markers (including eight nuclear-encoded genes, four protein-coding mitochondrial genes, and the 16S mitochondrial rRNA gene, see Gippner et al.2021). Branches with posterior probability> 0.9 are denoted by asterisks (*) at relevant nodes. Blue bars depict 95% High Probability Density (HPD) intervals of estimated divergence dates.B, Median-joining nuclear allele network showing the relationships between the L. picturatus-fischeri groups, inferred from the RAG1 nuclear gene (262 bp including all available RAG1 sequences of the L. picturatus-fischeri groups).C, Median-joining nuclear allele network of the RAG1 nuclear gene (980 bp) showing the relationships within the L. gutturalis subgroup, excluding L. gutturalis s.s. In all networks, circle frequency of alleles and small lines represent mutational steps.	2023-09-23	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli		Zenodo	biologists	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli			
03B30955FFCCFFE2FC6E4462FF66F92B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13302846/files/figure.png	https://doi.org/10.5281/zenodo.13302846	Figure 4. A, PCA plots of the first principal component (PC1) vs. the second principal component (PC2). Species are represented by different colours included in the legend. B, Boxplots (top whisker–maximum value; lower whisker–minimum value; dark horizontal line–median) of the body size (SVL) and linear measurements analysed in this study.Values in all the variables with the exception of SVL, are represented using the Least-squares means interaction plot (function Lsmip, package lsmeans). For abbreviations and further details see Material and methods section.	Figure 4. A, PCA plots of the first principal component (PC1) vs. the second principal component (PC2). Species are represented by different colours included in the legend. B, Boxplots (top whisker–maximum value; lower whisker–minimum value; dark horizontal line–median) of the body size (SVL) and linear measurements analysed in this study.Values in all the variables with the exception of SVL, are represented using the Least-squares means interaction plot (function Lsmip, package lsmeans). For abbreviations and further details see Material and methods section.	2023-09-23	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli		Zenodo	biologists	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli			
03B30955FFCAFFDEFF1F46B0FD56FE68.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13302905/files/figure.png	https://doi.org/10.5281/zenodo.13302905	Figure 19. Holotype of L. karamojasp. nov. (UTEP 22590) from Agoro Town, Imatong Foothills, Northern Region, Uganda.A, Dorsal and ventral view of body, (B) details of cloacal section, (C) regenerated tail tip, and (D) left pes. E, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R.	Figure 19. Holotype of L. karamojasp. nov. (UTEP 22590) from Agoro Town, Imatong Foothills, Northern Region, Uganda.A, Dorsal and ventral view of body, (B) details of cloacal section, (C) regenerated tail tip, and (D) left pes. E, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R.	2023-09-23	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli		Zenodo	biologists	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli			
03B30955FFCAFFDEFF1F46B0FD56FE68.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13302907/files/figure.png	https://doi.org/10.5281/zenodo.13302907	Figure 20. Specimens of L. karamoja sp. nov. in life from (A–B) Agoro Town (holotype: UTEP 22590), Imatong Foothills and (D–E) Nakapiripirit (UTEP 22594), Mount Kadam, Northern Region, Uganda.Photo of habitat at (C) Imatong Foothills, and (F) Mount Kadam, Northern Region, Uganda. Photographs by D.F.H.	Figure 20. Specimens of L. karamoja sp. nov. in life from (A–B) Agoro Town (holotype: UTEP 22590), Imatong Foothills and (D–E) Nakapiripirit (UTEP 22594), Mount Kadam, Northern Region, Uganda.Photo of habitat at (C) Imatong Foothills, and (F) Mount Kadam, Northern Region, Uganda. Photographs by D.F.H.	2023-09-23	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli		Zenodo	biologists	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli			
03B30955FFCAFFDEFF1F46B0FD56FE68.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13302858/files/figure.png	https://doi.org/10.5281/zenodo.13302858	Figure 7. Holotype of L. heeneni [RMCA (= MRAC) R.8477] from Kapiri, Lualaba Province, DRC. A, Ventral and dorsal view of body. B, Details of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R.	Figure 7. Holotype of L. heeneni [RMCA (= MRAC) R.8477] from Kapiri, Lualaba Province, DRC. A, Ventral and dorsal view of body. B, Details of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R.	2023-09-23	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli		Zenodo	biologists	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli			
03B30955FFCAFFDEFF1F46B0FD56FE68.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13302840/files/figure.png	https://doi.org/10.5281/zenodo.13302840	Figure 2. A, Multilocus species tree and Bayesian chronogram of divergences among Afro-American Lygodactylus clade (Clade C of Gippner et al. 2021), based on an alignment of 13 concatenated markers (including eight nuclear-encoded genes, four protein-coding mitochondrial genes, and the 16S mitochondrial rRNA gene, see Gippner et al.2021). Branches with posterior probability> 0.9 are denoted by asterisks (*) at relevant nodes. Blue bars depict 95% High Probability Density (HPD) intervals of estimated divergence dates.B, Median-joining nuclear allele network showing the relationships between the L. picturatus-fischeri groups, inferred from the RAG1 nuclear gene (262 bp including all available RAG1 sequences of the L. picturatus-fischeri groups).C, Median-joining nuclear allele network of the RAG1 nuclear gene (980 bp) showing the relationships within the L. gutturalis subgroup, excluding L. gutturalis s.s. In all networks, circle frequency of alleles and small lines represent mutational steps.	Figure 2. A, Multilocus species tree and Bayesian chronogram of divergences among Afro-American Lygodactylus clade (Clade C of Gippner et al. 2021), based on an alignment of 13 concatenated markers (including eight nuclear-encoded genes, four protein-coding mitochondrial genes, and the 16S mitochondrial rRNA gene, see Gippner et al.2021). Branches with posterior probability> 0.9 are denoted by asterisks (*) at relevant nodes. Blue bars depict 95% High Probability Density (HPD) intervals of estimated divergence dates.B, Median-joining nuclear allele network showing the relationships between the L. picturatus-fischeri groups, inferred from the RAG1 nuclear gene (262 bp including all available RAG1 sequences of the L. picturatus-fischeri groups).C, Median-joining nuclear allele network of the RAG1 nuclear gene (980 bp) showing the relationships within the L. gutturalis subgroup, excluding L. gutturalis s.s. In all networks, circle frequency of alleles and small lines represent mutational steps.	2023-09-23	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli		Zenodo	biologists	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli			
03B30955FFCAFFDEFF1F46B0FD56FE68.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13302854/files/figure.png	https://doi.org/10.5281/zenodo.13302854	Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi).	Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi).	2023-09-23	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli		Zenodo	biologists	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli			
03B30955FFF6FFDDFF0E427BFC3FFEEA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13302911/files/figure.png	https://doi.org/10.5281/zenodo.13302911	Figure 21. Holotype of L. mirabundus sp. nov. (UTEP 22585) from Katako Kombe, Sankuru Province, DRC. A, Dorsal and ventral view of body. B, Details of cloacal region and (C) right pes. D, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R.	Figure 21. Holotype of L. mirabundus sp. nov. (UTEP 22585) from Katako Kombe, Sankuru Province, DRC. A, Dorsal and ventral view of body. B, Details of cloacal region and (C) right pes. D, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R.	2023-09-23	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli		Zenodo	biologists	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli			
03B30955FFF6FFDDFF0E427BFC3FFEEA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13302840/files/figure.png	https://doi.org/10.5281/zenodo.13302840	Figure 2. A, Multilocus species tree and Bayesian chronogram of divergences among Afro-American Lygodactylus clade (Clade C of Gippner et al. 2021), based on an alignment of 13 concatenated markers (including eight nuclear-encoded genes, four protein-coding mitochondrial genes, and the 16S mitochondrial rRNA gene, see Gippner et al.2021). Branches with posterior probability> 0.9 are denoted by asterisks (*) at relevant nodes. Blue bars depict 95% High Probability Density (HPD) intervals of estimated divergence dates.B, Median-joining nuclear allele network showing the relationships between the L. picturatus-fischeri groups, inferred from the RAG1 nuclear gene (262 bp including all available RAG1 sequences of the L. picturatus-fischeri groups).C, Median-joining nuclear allele network of the RAG1 nuclear gene (980 bp) showing the relationships within the L. gutturalis subgroup, excluding L. gutturalis s.s. In all networks, circle frequency of alleles and small lines represent mutational steps.	Figure 2. A, Multilocus species tree and Bayesian chronogram of divergences among Afro-American Lygodactylus clade (Clade C of Gippner et al. 2021), based on an alignment of 13 concatenated markers (including eight nuclear-encoded genes, four protein-coding mitochondrial genes, and the 16S mitochondrial rRNA gene, see Gippner et al.2021). Branches with posterior probability> 0.9 are denoted by asterisks (*) at relevant nodes. Blue bars depict 95% High Probability Density (HPD) intervals of estimated divergence dates.B, Median-joining nuclear allele network showing the relationships between the L. picturatus-fischeri groups, inferred from the RAG1 nuclear gene (262 bp including all available RAG1 sequences of the L. picturatus-fischeri groups).C, Median-joining nuclear allele network of the RAG1 nuclear gene (980 bp) showing the relationships within the L. gutturalis subgroup, excluding L. gutturalis s.s. In all networks, circle frequency of alleles and small lines represent mutational steps.	2023-09-23	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli		Zenodo	biologists	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli			
03B30955FFF6FFDDFF0E427BFC3FFEEA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13302850/files/figure.png	https://doi.org/10.5281/zenodo.13302850	Figure 5. Gular and skull osteology comparisons between holotype material or representative material of all members of the L. gutturalis subgroup and the L. angularis group mentioned in this work.From top to bottom, ventral view gular section; dorsal, lateral, and ventral view of the skull; lateral, dorsal, medial, and ventral view of jaw; and number of tooth loci. PM = premaxilla, MX(R) = right maxilla, MX(L) = left maxilla, DN(R) = right dentary of the jaw.	Figure 5. Gular and skull osteology comparisons between holotype material or representative material of all members of the L. gutturalis subgroup and the L. angularis group mentioned in this work.From top to bottom, ventral view gular section; dorsal, lateral, and ventral view of the skull; lateral, dorsal, medial, and ventral view of jaw; and number of tooth loci. PM = premaxilla, MX(R) = right maxilla, MX(L) = left maxilla, DN(R) = right dentary of the jaw.	2023-09-23	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli		Zenodo	biologists	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli			
03B30955FFF5FFDBFC534104FEB5FB28.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13302915/files/figure.png	https://doi.org/10.5281/zenodo.13302915	Figure 22. Holotype of L. leopardinus sp. nov. (UTEP 22577) from Balolombo Village, Busira River, Équateur Province, DRC.A, Dorsal and ventral view of body. B, Details of right manus and (C) pes. D, Detailed view of head in ventral, dorsal, and lateral views (from top to bottom). Photos by J.L.R.	Figure 22. Holotype of L. leopardinus sp. nov. (UTEP 22577) from Balolombo Village, Busira River, Équateur Province, DRC.A, Dorsal and ventral view of body. B, Details of right manus and (C) pes. D, Detailed view of head in ventral, dorsal, and lateral views (from top to bottom). Photos by J.L.R.	2023-09-23	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli		Zenodo	biologists	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli			
03B30955FFF5FFDBFC534104FEB5FB28.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13302917/files/figure.png	https://doi.org/10.5281/zenodo.13302917	Figure 23. A–B, Holotype (UTEP 22577) and (C–D) paratype (UTEP 22596) of L. leopardinus sp. nov. in life and photo of habitat from (E) type locality at Balolombo Village, Busira River, Équateur Province, DRC. F–G, Holotype (UTEP 22587) and (H–I) paratype (UTEP 22584) of L. gamblei sp. nov. in life and photo of habitat from (J) type locality at Manono, Tanganyika Province, DRC. Photographs by E.G.	Figure 23. A–B, Holotype (UTEP 22577) and (C–D) paratype (UTEP 22596) of L. leopardinus sp. nov. in life and photo of habitat from (E) type locality at Balolombo Village, Busira River, Équateur Province, DRC. F–G, Holotype (UTEP 22587) and (H–I) paratype (UTEP 22584) of L. gamblei sp. nov. in life and photo of habitat from (J) type locality at Manono, Tanganyika Province, DRC. Photographs by E.G.	2023-09-23	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli		Zenodo	biologists	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli			
03B30955FFF5FFDBFC534104FEB5FB28.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13302840/files/figure.png	https://doi.org/10.5281/zenodo.13302840	Figure 2. A, Multilocus species tree and Bayesian chronogram of divergences among Afro-American Lygodactylus clade (Clade C of Gippner et al. 2021), based on an alignment of 13 concatenated markers (including eight nuclear-encoded genes, four protein-coding mitochondrial genes, and the 16S mitochondrial rRNA gene, see Gippner et al.2021). Branches with posterior probability> 0.9 are denoted by asterisks (*) at relevant nodes. Blue bars depict 95% High Probability Density (HPD) intervals of estimated divergence dates.B, Median-joining nuclear allele network showing the relationships between the L. picturatus-fischeri groups, inferred from the RAG1 nuclear gene (262 bp including all available RAG1 sequences of the L. picturatus-fischeri groups).C, Median-joining nuclear allele network of the RAG1 nuclear gene (980 bp) showing the relationships within the L. gutturalis subgroup, excluding L. gutturalis s.s. In all networks, circle frequency of alleles and small lines represent mutational steps.	Figure 2. A, Multilocus species tree and Bayesian chronogram of divergences among Afro-American Lygodactylus clade (Clade C of Gippner et al. 2021), based on an alignment of 13 concatenated markers (including eight nuclear-encoded genes, four protein-coding mitochondrial genes, and the 16S mitochondrial rRNA gene, see Gippner et al.2021). Branches with posterior probability> 0.9 are denoted by asterisks (*) at relevant nodes. Blue bars depict 95% High Probability Density (HPD) intervals of estimated divergence dates.B, Median-joining nuclear allele network showing the relationships between the L. picturatus-fischeri groups, inferred from the RAG1 nuclear gene (262 bp including all available RAG1 sequences of the L. picturatus-fischeri groups).C, Median-joining nuclear allele network of the RAG1 nuclear gene (980 bp) showing the relationships within the L. gutturalis subgroup, excluding L. gutturalis s.s. In all networks, circle frequency of alleles and small lines represent mutational steps.	2023-09-23	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli		Zenodo	biologists	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli			
03B30955FFF5FFDBFC534104FEB5FB28.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13302842/files/figure.png	https://doi.org/10.5281/zenodo.13302842	Figure 3. Comparative figure including dorsal (or dorsolateral) and ventral views of preserved holotypes (with exception of Lygodactylus gutturalis that is represented by a paratype) of all species within the L.gutturalis subgroup used in the current study.	Figure 3. Comparative figure including dorsal (or dorsolateral) and ventral views of preserved holotypes (with exception of Lygodactylus gutturalis that is represented by a paratype) of all species within the L.gutturalis subgroup used in the current study.	2023-09-23	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli		Zenodo	biologists	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli			
03B30955FFF5FFDBFC534104FEB5FB28.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13302850/files/figure.png	https://doi.org/10.5281/zenodo.13302850	Figure 5. Gular and skull osteology comparisons between holotype material or representative material of all members of the L. gutturalis subgroup and the L. angularis group mentioned in this work.From top to bottom, ventral view gular section; dorsal, lateral, and ventral view of the skull; lateral, dorsal, medial, and ventral view of jaw; and number of tooth loci. PM = premaxilla, MX(R) = right maxilla, MX(L) = left maxilla, DN(R) = right dentary of the jaw.	Figure 5. Gular and skull osteology comparisons between holotype material or representative material of all members of the L. gutturalis subgroup and the L. angularis group mentioned in this work.From top to bottom, ventral view gular section; dorsal, lateral, and ventral view of the skull; lateral, dorsal, medial, and ventral view of jaw; and number of tooth loci. PM = premaxilla, MX(R) = right maxilla, MX(L) = left maxilla, DN(R) = right dentary of the jaw.	2023-09-23	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli		Zenodo	biologists	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli			
03B30955FFF5FFDBFC534104FEB5FB28.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13302858/files/figure.png	https://doi.org/10.5281/zenodo.13302858	Figure 7. Holotype of L. heeneni [RMCA (= MRAC) R.8477] from Kapiri, Lualaba Province, DRC. A, Ventral and dorsal view of body. B, Details of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R.	Figure 7. Holotype of L. heeneni [RMCA (= MRAC) R.8477] from Kapiri, Lualaba Province, DRC. A, Ventral and dorsal view of body. B, Details of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R.	2023-09-23	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli		Zenodo	biologists	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli			
03B30955FFF5FFDBFC534104FEB5FB28.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13302860/files/figure.png	https://doi.org/10.5281/zenodo.13302860	Figure 8. Paratype of L. paurospilus [RMCA (=MRAC) R.27409] from Haute Lubitshako, South Kivu Province, DRC. A, Ventral and dorsal views of body. B, Details of head in ventral, dorsal, and lateral views (from top to bottom). C, Details of cloacal region. D, Details of right manus. Photos by J.L.R.	Figure 8. Paratype of L. paurospilus [RMCA (=MRAC) R.27409] from Haute Lubitshako, South Kivu Province, DRC. A, Ventral and dorsal views of body. B, Details of head in ventral, dorsal, and lateral views (from top to bottom). C, Details of cloacal region. D, Details of right manus. Photos by J.L.R.	2023-09-23	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli		Zenodo	biologists	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli			
03B30955FFF5FFDBFC534104FEB5FB28.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13302846/files/figure.png	https://doi.org/10.5281/zenodo.13302846	Figure 4. A, PCA plots of the first principal component (PC1) vs. the second principal component (PC2). Species are represented by different colours included in the legend. B, Boxplots (top whisker–maximum value; lower whisker–minimum value; dark horizontal line–median) of the body size (SVL) and linear measurements analysed in this study.Values in all the variables with the exception of SVL, are represented using the Least-squares means interaction plot (function Lsmip, package lsmeans). For abbreviations and further details see Material and methods section.	Figure 4. A, PCA plots of the first principal component (PC1) vs. the second principal component (PC2). Species are represented by different colours included in the legend. B, Boxplots (top whisker–maximum value; lower whisker–minimum value; dark horizontal line–median) of the body size (SVL) and linear measurements analysed in this study.Values in all the variables with the exception of SVL, are represented using the Least-squares means interaction plot (function Lsmip, package lsmeans). For abbreviations and further details see Material and methods section.	2023-09-23	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli		Zenodo	biologists	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli			
03B30955FFF5FFDBFC534104FEB5FB28.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13302854/files/figure.png	https://doi.org/10.5281/zenodo.13302854	Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi).	Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi).	2023-09-23	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli		Zenodo	biologists	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli			
03B30955FFF3FFD9FF3444C6FC56F8A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13302917/files/figure.png	https://doi.org/10.5281/zenodo.13302917	Figure 23. A–B, Holotype (UTEP 22577) and (C–D) paratype (UTEP 22596) of L. leopardinus sp. nov. in life and photo of habitat from (E) type locality at Balolombo Village, Busira River, Équateur Province, DRC. F–G, Holotype (UTEP 22587) and (H–I) paratype (UTEP 22584) of L. gamblei sp. nov. in life and photo of habitat from (J) type locality at Manono, Tanganyika Province, DRC. Photographs by E.G.	Figure 23. A–B, Holotype (UTEP 22577) and (C–D) paratype (UTEP 22596) of L. leopardinus sp. nov. in life and photo of habitat from (E) type locality at Balolombo Village, Busira River, Équateur Province, DRC. F–G, Holotype (UTEP 22587) and (H–I) paratype (UTEP 22584) of L. gamblei sp. nov. in life and photo of habitat from (J) type locality at Manono, Tanganyika Province, DRC. Photographs by E.G.	2023-09-23	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli		Zenodo	biologists	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli			
03B30955FFF3FFD9FF3444C6FC56F8A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13302919/files/figure.png	https://doi.org/10.5281/zenodo.13302919	Figure 24. Holotype of L. gamblei sp. nov. (UTEP 22587) from Manono, Tanganyika Province, DRC.A, Dorsal and ventral view of body.B, Details of the cloacal region.C, Detailed view of head in ventral, dorsal, and lateral views (from top to bottom). Photos by J.L.R.	Figure 24. Holotype of L. gamblei sp. nov. (UTEP 22587) from Manono, Tanganyika Province, DRC.A, Dorsal and ventral view of body.B, Details of the cloacal region.C, Detailed view of head in ventral, dorsal, and lateral views (from top to bottom). Photos by J.L.R.	2023-09-23	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli		Zenodo	biologists	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli			
03B30955FFF3FFD9FF3444C6FC56F8A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13302840/files/figure.png	https://doi.org/10.5281/zenodo.13302840	Figure 2. A, Multilocus species tree and Bayesian chronogram of divergences among Afro-American Lygodactylus clade (Clade C of Gippner et al. 2021), based on an alignment of 13 concatenated markers (including eight nuclear-encoded genes, four protein-coding mitochondrial genes, and the 16S mitochondrial rRNA gene, see Gippner et al.2021). Branches with posterior probability> 0.9 are denoted by asterisks (*) at relevant nodes. Blue bars depict 95% High Probability Density (HPD) intervals of estimated divergence dates.B, Median-joining nuclear allele network showing the relationships between the L. picturatus-fischeri groups, inferred from the RAG1 nuclear gene (262 bp including all available RAG1 sequences of the L. picturatus-fischeri groups).C, Median-joining nuclear allele network of the RAG1 nuclear gene (980 bp) showing the relationships within the L. gutturalis subgroup, excluding L. gutturalis s.s. In all networks, circle frequency of alleles and small lines represent mutational steps.	Figure 2. A, Multilocus species tree and Bayesian chronogram of divergences among Afro-American Lygodactylus clade (Clade C of Gippner et al. 2021), based on an alignment of 13 concatenated markers (including eight nuclear-encoded genes, four protein-coding mitochondrial genes, and the 16S mitochondrial rRNA gene, see Gippner et al.2021). Branches with posterior probability> 0.9 are denoted by asterisks (*) at relevant nodes. Blue bars depict 95% High Probability Density (HPD) intervals of estimated divergence dates.B, Median-joining nuclear allele network showing the relationships between the L. picturatus-fischeri groups, inferred from the RAG1 nuclear gene (262 bp including all available RAG1 sequences of the L. picturatus-fischeri groups).C, Median-joining nuclear allele network of the RAG1 nuclear gene (980 bp) showing the relationships within the L. gutturalis subgroup, excluding L. gutturalis s.s. In all networks, circle frequency of alleles and small lines represent mutational steps.	2023-09-23	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli		Zenodo	biologists	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli			
03B30955FFF3FFD9FF3444C6FC56F8A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13302923/files/figure.png	https://doi.org/10.5281/zenodo.13302923	Figure 25. A, Dorsal and ventral views of head, showing gular ornamentation and pholidosis variation in L. gamblei sp. nov. B, Specimens of L. gamblei sp. nov. in lateral view, showing its characteristic scapular pattern. Red points denote males and blue points denote females. Photos by J.L.R.	Figure 25. A, Dorsal and ventral views of head, showing gular ornamentation and pholidosis variation in L. gamblei sp. nov. B, Specimens of L. gamblei sp. nov. in lateral view, showing its characteristic scapular pattern. Red points denote males and blue points denote females. Photos by J.L.R.	2023-09-23	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli		Zenodo	biologists	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli			
03B30955FFF3FFD9FF3444C6FC56F8A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/13302854/files/figure.png	https://doi.org/10.5281/zenodo.13302854	Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi).	Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi).	2023-09-23	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli		Zenodo	biologists	Lobón-Rovira, Javier;Bauer, Aaron M.;Pinto, Pedro Vaz;Trape, Jean-Francois;Conradie, Werner;Kusamba, Chifundera;Júlio, Timóteo;Cael, Garin;Stanley, Edward L.;Hughes, Daniel F.;Behangana, Mathias;Masudi, Franck M.;Pauwels, Olivier S. G.;Greenbaum, Eli			
