taxonID	type	description	language	source
522B6BC06B8B718C251D42A611E394EA.taxon	distribution	Distribution. Palaearctic, north Oriental.	en	Wang, Cheng-Bin, Ruzicka, Jan, Perreau, Michel, Nishikawa, Masaaki, Park, Sun-Jae (2016): Revision of the genus Ptomaphagus Hellwig (Coleoptera, Leiodidae, Cholevinae) from the Russian Far East and the Korean Peninsula. ZooKeys 637: 33-45, DOI: http://dx.doi.org/10.3897/zookeys.637.9384, URL: http://dx.doi.org/10.3897/zookeys.637.9384
8DEF8A32A746E5BD872850B68BD1BFD7.taxon	distribution	Distribution. Holarctic, north Oriental, north Neotropical.	en	Wang, Cheng-Bin, Ruzicka, Jan, Perreau, Michel, Nishikawa, Masaaki, Park, Sun-Jae (2016): Revision of the genus Ptomaphagus Hellwig (Coleoptera, Leiodidae, Cholevinae) from the Russian Far East and the Korean Peninsula. ZooKeys 637: 33-45, DOI: http://dx.doi.org/10.3897/zookeys.637.9384, URL: http://dx.doi.org/10.3897/zookeys.637.9384
C8FCCFC2D8533E2F4BC6BFE9A606810F.taxon	description	Description. Male. EBL: 3.8 - 4.3 mm (4.2 mm in holotype). Length of different body parts: HL: AL: PL: ELL = 0.7: 1.1: 1.1: 2.3 mm; width: HW: EW: PW: ELW = 1.1: 0.1: 1.6: 1.8 mm. Proportion of antennomeres from base to tip in μm (length x width): 149 x 75, 109 x 67, 95 x 70, 64 x 79, 58 x 92, 41 x 100, 91 x 128, 39 x 128, 87 x 151, 90 x 156, 161 x 143. Habitus (Fig. 1 B) elongated oval, regularly convex and sublustrous. Well pigmented: mostly blackish brown; mouthparts, antennae (apical half of ultimate antennomere yellowish) and tarsi reddish brown. Dorsum continuously clothed with fine, recumbent, yellowish pubescence. Insertions of pubescence on dorsal surfaces of pronotum, elytra and femora aligned along transverse striolations; interspace between two striolations glabrous. Head transverse, HW / HL = 1.6. Clypeofrontal suture absent. Clypeus with anterior margin slightly rounded. Compound eyes well developed, EW / HW = 0.11. Antennae (Fig. 2 A) slender, AL / HW = 1.0; antennomere III shorter than II; VI with length / width = 0.4; XI peach-shape. Pronotum (Fig. 2 B) transverse, widest just before hind angles, PW / PL = 1.5. Sides gently arched, simply narrowing from posterior to anterior; hind angles slightly projected backwards and acute. Posterior margin widely protruded in middle part, emarginate near hind angles. Elytra oval, widest at about basal 1 / 4, ELL / EW = 1.3. Sides weakly arched, gradually narrowing from widest part to apices; apices (Fig. 2 G) narrowly rounded. Sutural striae present. Metathoracic wings fully developed. Prolegs robust, with basal three protarsomeres (Fig. 2 C) strongly expanded: TW / BTW = 1.2. Protibiae (Fig. 2 E) strongly expanded towards apex. Profemora broad. Mesotibiae arcuate, mesotarsi simply linear. Metatibiae slender and straight. Abdominal ventrite VIII (Fig. 2 I) round at posterior edge and with an inconspicuous median notch. Spiculum gastrale (Fig. 2 J) of genital segment with about 1 / 5 of length protruding beyond anterior edge of epipleurite IX. Aedeagus (Fig. 3 A) long and slender, with median lobe gradually narrowing towards a lanceolate apex and terminated to a widely rounded knob in dorsal view; opening of genital orifice situated on dorsal surface, deeply cut inwards on preapical left margin of median lobe. Ventral surface of the apex of the median lobe (Fig. 3 C) inserted with 5 ventrally oriented setae on both sides; parameres narrow, reaching about apical 1 / 5 of median lobe, each apex (Fig. 3 D) with 2 lateral setae and 1 apical seta distinctly shorter. In lateral view (Fig. 3 B), median lobe regularly bent ventrad and gradually tapering to a flat apex. Endophallus with stylus quite slender, a subelliptical nodule in middle region, a cheliform complex just below base of stylus, and a circular complex in the basal region. Female. Similar to male in general appearance (Fig. 1 C), including elytral apices (Fig. 2 H), but distinguished by the following characteristics: protarsi (Fig. 2 D) simply linear; protibiae (Fig. 2 F) narrower; abdominal ventrite VIII (Fig. 3 E) narrowly rounded at posterior edge; genital segment and ovipositor as shown in Fig. 3 F; spermatheca (Fig. 3 F) curved in distal part and coiled in proximal part.	en	Wang, Cheng-Bin, Ruzicka, Jan, Perreau, Michel, Nishikawa, Masaaki, Park, Sun-Jae (2016): Revision of the genus Ptomaphagus Hellwig (Coleoptera, Leiodidae, Cholevinae) from the Russian Far East and the Korean Peninsula. ZooKeys 637: 33-45, DOI: http://dx.doi.org/10.3897/zookeys.637.9384, URL: http://dx.doi.org/10.3897/zookeys.637.9384
C8FCCFC2D8533E2F4BC6BFE9A606810F.taxon	diagnosis	Diagnosis. Ptomaphagus (s. str.) sibiricus is sympatric with Ptomaphagus (s. str.) hayashii sp. n. in some locations of the Russian Far East, and they are very similar to each other (including spermatheca (Figs 3 F; 5 F), which is only more roundly curved in the stem part in Ptomaphagus (s. str.) hayashii sp. n.). For external characters, the body size of Ptomaphagus (s. str.) sibiricus (Figs 1 A, D-F) is a little smaller than Ptomaphagus (s. str.) hayashii sp. n. (Figs 1 B, C). However, their aedeagi provide critical characters to distinguish the two species: in Ptomaphagus (s. str.) hayashii sp. n., the aedeagus is much larger and more slender (Fig. 3 A), the right apicoventral piece of the median lobe is slenderly lanceolate (Fig. 3 C), the apical half of median lobe much flatter in lateral view (Fig. 3 B); while in Ptomaphagus (s. str.) sibiricus, the aedeagus is stouter (Fig. 5 A), the right apicoventral piece of median lobe is much wider and subpentagonal (Fig. 5 C), the apical half of the median lobe thicker in lateral view (Fig. 5 B). Furthermore, based on specimens examined, Ptomaphagus (s. str.) sibiricus seems to be much more widely distributed, extending southward to South Korea; while Ptomaphagus (s. str.) hayashii sp. n. is presently known only in the Russian Far East.	en	Wang, Cheng-Bin, Ruzicka, Jan, Perreau, Michel, Nishikawa, Masaaki, Park, Sun-Jae (2016): Revision of the genus Ptomaphagus Hellwig (Coleoptera, Leiodidae, Cholevinae) from the Russian Far East and the Korean Peninsula. ZooKeys 637: 33-45, DOI: http://dx.doi.org/10.3897/zookeys.637.9384, URL: http://dx.doi.org/10.3897/zookeys.637.9384
C8FCCFC2D8533E2F4BC6BFE9A606810F.taxon	etymology	Etymology. The specific epithet is dedicated to Mr. Yasuhiko Hayashi (Kawanishi, Japan), a famous independent taxonomist on Staphylinoidea, for his continual generous help to our study.	en	Wang, Cheng-Bin, Ruzicka, Jan, Perreau, Michel, Nishikawa, Masaaki, Park, Sun-Jae (2016): Revision of the genus Ptomaphagus Hellwig (Coleoptera, Leiodidae, Cholevinae) from the Russian Far East and the Korean Peninsula. ZooKeys 637: 33-45, DOI: http://dx.doi.org/10.3897/zookeys.637.9384, URL: http://dx.doi.org/10.3897/zookeys.637.9384
C8FCCFC2D8533E2F4BC6BFE9A606810F.taxon	distribution	Distribution. Russia (Far East) (Fig. 6).	en	Wang, Cheng-Bin, Ruzicka, Jan, Perreau, Michel, Nishikawa, Masaaki, Park, Sun-Jae (2016): Revision of the genus Ptomaphagus Hellwig (Coleoptera, Leiodidae, Cholevinae) from the Russian Far East and the Korean Peninsula. ZooKeys 637: 33-45, DOI: http://dx.doi.org/10.3897/zookeys.637.9384, URL: http://dx.doi.org/10.3897/zookeys.637.9384
24E4CA88822C25B44C7CA6789421B9EB.taxon	materials_examined	Material examined. Type material. Holotype: ♀, [RUSSIA, Far East] Wladiwostok [ca. 43 ° 10 ' N 132 ° 00 ' E] // Reitter // Coll. Koltze // Pt. variicornis / Rosenh. // Ptomaphagus sibiricus Jeann. / type / R. Jeannel det. // DEI Muencheberg / Col - 07069 (SDEI). Additional material. RUSSIA, Far East: 1 ♀, FE. Russia, SW Khabarovsky / kray reg., Strel'nikova / range Mts., 46 ° 43 ' N 134 ° 08 ' E / Samur Riv., 130 - 550 m alt. / 25. VI. - 28. VII. 2014 / A. Plutenko leg. (CMNE); 1 ♀, Vladivostok [ca. 43 ° 10 ' N 132 ° 00 ' E] / Christov [leg.] IX. [18] 76 // Ptomaphagus sibiricus / Jeannel det. // Ptomaphagus / (Ptomaphagus) sibiricus / Jeannel, 1934 / Ex. M. Nishikawa, 2008 / # MNHN 103377 Ch 1 S ♀ (MNHN); 10 ♂♂ 10 ♀♀, RUSSIA: Primorsky / 30 km NE Vladivostok / Tajvaza / 29. VII- 5. VIII. 1992 / B. D. Gill [leg.] (CJRZ); 1 ♀, RUSSIA: Primoryi / Nakhodka [ca. 42 ° 49 ' N 132 ° 52 ' E] / 6 - 8. VIII. 1999 / B. D. Gill [leg.] (CJRZ); 1 ♂, USSR: Sadgorod [ca. 43 ° 15 ' N 132 ° 03 ' E] / (in forest; trap with / bait), Vladivostok // Primorskyi Kray / 16. VI. 1978 / E. Berlov leg. (CMNE); 1 ♂, Primorskiy Kray, Ussuriyskiy Region, Uss. [uriyskiy] Zapov. [ednik, = Reserve], Staraya Baza, 43.64 ° N, 132.34 ° E, poch. l. [= pitfall trap], V. Zinchenko & A. Korshunov [leg.], 9. - 19. viii. 2011 // Ptomaphagus / sibiricus / V. Zinchenko det. 2002 (CCBW). South Korea: 1 ♂, KOREA: Gangwon Prov., / Pyeongchang-gun, Jinbu-myeon, / Dongsan-ri, Mt. Odaesan, Sangwonsa [ca. 37 ° 47 ' N 128 ° 33 ' E], / 4. VI. - 22. VI. 2001, K. - J. Ahn, S. - J. / Park, M. - S. Kim, M. - J. Jeon [leg.], ex FIT // Ptomaphagus / baekamsanensis / n. sp. / det. S. - J. Park 2005 // 1 (CNUIC); 1 ♂, KOREA: Gangwon Prov., / Hongcheon-gun [ca. 37 ° 41 ' N 127 ° 50 ' E], Mt. Baekamsan, / 25. V. - 20. VI. 2002, K J Ahn, / C W Shin, J S Park, ex FIT // Ptomaphagus koreanus / new species / det. Sun-Jae Park (CNUIC); 1 ♀, KOREA, Chungbuk Prov., / Danyanggun, Youngchunmyeon, / Namcheonri, Namcheon valley, / Mt. Sobaeksan [ca. 36 ° 57 ' N 128 ° 26 ' E], (M. T), / 25. V. - 6. VII. 2006 (CNUIC); 1 ♂, same data as previous except: 2006. VII. 6 - VII. 28 (CNUIC); 1 ♀, Korea, Kyungsangbuk-do / Yuongdong-gun / Sangchon-myun, Mulhan-ri / Mt. Minjujisan [ca. 36 ° 02 ' N 127 ° 50 ' E] / VI. 16 - 18.2000, / Y. B. Cho / coll. / ex bait trap // ♀ No. 9 (CNUIC); 1 ♀, same data as previous except: ♀ 10 (CNUIC); 4 ♂♂ 1 ♀, Korea, Chonrabukdo / Selchonmyun / Mt. Deokyusan / Baekyeonsa Temple [ca. 35 ° 26 ' N 126 ° 52 ' E] // vi. 16.1999 / D. S. Kim coll. / ex bait trap // Ptomaphagus / sp. / Det. M. Nishikawa // [5, 6, 13, 14 & 15, respectively] (CNUIC); 1 ♂ 2 ♀♀, KOREA: Chungbuk Prov., / Boeun-gun, Mt. Sokrisan [ca. 36 ° 32 ' N 127 ° 54 ' E], / Bubjusa, 2004. VI. 1 - 28, / Y. - B. Cho [leg.], FIT (CMNE).	en	Wang, Cheng-Bin, Ruzicka, Jan, Perreau, Michel, Nishikawa, Masaaki, Park, Sun-Jae (2016): Revision of the genus Ptomaphagus Hellwig (Coleoptera, Leiodidae, Cholevinae) from the Russian Far East and the Korean Peninsula. ZooKeys 637: 33-45, DOI: http://dx.doi.org/10.3897/zookeys.637.9384, URL: http://dx.doi.org/10.3897/zookeys.637.9384
24E4CA88822C25B44C7CA6789421B9EB.taxon	diagnosis	Diagnosis. See under Ptomaphagus (s. str.) hayashii sp. n. above. Other remarks on this species see Wang et al. (2016 a).	en	Wang, Cheng-Bin, Ruzicka, Jan, Perreau, Michel, Nishikawa, Masaaki, Park, Sun-Jae (2016): Revision of the genus Ptomaphagus Hellwig (Coleoptera, Leiodidae, Cholevinae) from the Russian Far East and the Korean Peninsula. ZooKeys 637: 33-45, DOI: http://dx.doi.org/10.3897/zookeys.637.9384, URL: http://dx.doi.org/10.3897/zookeys.637.9384
24E4CA88822C25B44C7CA6789421B9EB.taxon	distribution	Distribution. Russia (Far East), South Korea (Fig. 6).	en	Wang, Cheng-Bin, Ruzicka, Jan, Perreau, Michel, Nishikawa, Masaaki, Park, Sun-Jae (2016): Revision of the genus Ptomaphagus Hellwig (Coleoptera, Leiodidae, Cholevinae) from the Russian Far East and the Korean Peninsula. ZooKeys 637: 33-45, DOI: http://dx.doi.org/10.3897/zookeys.637.9384, URL: http://dx.doi.org/10.3897/zookeys.637.9384
