identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03B187D4DF1EFF96FF4F624DFC06DE94.text	03B187D4DF1EFF96FF4F624DFC06DE94.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aphaniidae	<div><p>Keys to the genera of  Aphaniidae</p><p>References to bars in the keys and subsequent text correspond to the dark brown or black vertical flank bars. The silvery-white alternating bars are referred to as interspaces. Several genera are identified by male colour pattern. The key is therefore designed to permit identification of nuptial adult males larger than 30 mm SL. This does not necessarily mean that smaller and younger males cannot be identified via the same characters, but it should be noted that some character states change with age and size of individuals. See Figure 1 for a two-way demonstration of character states against genera, with simplified cladograms provided by previous studies.</p><p>1a Head canals present, anterior supraorbital canal with 2–3 pores, posterior supraorbital canal with 2 pores, preopercular-mandibular canal with 6–7 pores............................................................................. 2</p><p>1b Head canals absent, cephalic sensory pores reduced to a series of neuromasts in small depressions, not connected to canals.. 3</p><p>2a In male caudal fin hyaline or white, often with yellow margin, with three, wide, bold black bars, first bar situated on caudal-fin base, shortly behind end of hypural complex, often faded (no bars in  A. furcatus). In nuptial female no dermal sheath around anterior anal-fin rays............................................................................  Aphaniops</p><p>2b In male caudal fin pale or deep yellow or orange, clearly distinct from silvery interspaces of flank bars; caudal fin with 0–2, often faint bars, no bar behind end of hypural complex. In nuptial female dermal sheath present around first few anal-fin rays.............................................................................................  Aphanius</p><p>3a Teeth conical, in three rows; caudal peduncle very narrow, its depth 2.5–3.2 times in its length; body naked.......................................................................................................  Kosswigichthys</p><p>3b Teeth tricuspid, in one row; caudal peduncle deep, its depth 1.2–2.0 times in its length; body covered by scales, but reduced or absent in some species.................................................................................. 4</p><p>4a Pelvic fin always absent; male with flank bars behind head to vertical through dorsal-fin origin, no bars on caudal peduncle; nuptial colour pattern comprising a bold, black band with wide yellow margin on dorsal, caudal and anal fins..........  Tellia</p><p>4b Pelvic fin present; male with flank bars along entire flank or flank without bars; nuptial colour pattern comprising black, darkgrey or bluish dorsal and anal fins with silvery or bright blue lines or spots, or with a bold black or white marginal band.... 5</p><p>5a Colour pattern in male comprising grey, bluish or almost black body with often irregular set and shaped, iridescent blue-white to silvery spots, or spots forming narrow vertical rows on flank, especially in juveniles; male with very narrow, blue-white or silvery rows of spots or small blotches forming bands on black or blue caudal fin. Female with many silvery spots or small blotches on flank; without diamond-shaped or roundish bold black blotch at mid-height of caudal-fin base...... Paraphanius</p><p>5b Colour pattern in male comprising silvery body with very regular set and shaped, rarely irregular, brown or black bars on flank, overlaid with iridescent silvery spots in some species; male with bold, black or brown bars or with narrow brown bands on hyaline, white or yellow caudal fin, or caudal fin hyaline, with white margin in some species. Female with numerous black or brown spots, blotches or bars on flank; with a diamond-shaped or roundish bold black blotch at mid-height of caudal-fin base.................................................................................................... 6</p><p>6a Caudal fin in male without bars or rows of spots, or with 1–5 indistinct vertical rows of small brown spots on proximal portion of caudal fin in some species; dorsal-fin margin white, caudal- and anal-fin margins often white; if dorsal and anal-fin margins black ( E. isfahanensis) caudal fin without rows of spots or bars.........................................  Esmaeilius</p><p>6b Caudal fin in male with 1–4 bold black bars or numerous vertical rows of small black or brown spots (4–14 in  A. villwocki, bars rarely absent in  A. saourensis); dorsal- and usually anal-fin margin black.......................................... 7</p><p>7a Dorsal fin in nuptial male completely black or with a wide greyish or black margin, a white proximal band or with a proximal row of white spots or blotches distinctly or slightly above dorsal-fin base; flank in male with 5–13 regularly shaped and set bars (13–25 in  A. villwocki); bars not overlaid with white spots (but irregular and overlaid with white spots in  A. irregularis)............................................................................................... Anatolichtys</p><p>7b Dorsal fin in nuptial male with a narrow black margin and a white or hyaline submargin; flank bars in male absent ( A. saourensis) or with 14–22 irregularly shaped and set bars; bars usually split vertically and overlaid with many minute whitish spots..........................................................................................  Apricaphanius</p><p>Remarks. The generic concept for  Aphaniidae suggested here is based on the criteria monophyly, morphological diagnosability, and reasonable compactness (Gill et al. 2005; Borkenhagen 2017). The proposed genera fulfil these criteria, because all of them are monophyletic groups that can be diagnosed by a combination of morphological characters (with few complications see below), and the genera each encompass one to 13 species, which are much more closely related to each other than to the species of another genus.</p><p>The grouping of  Aphaniidae into well-defined genera is based on phylogenies published by Hrbek et al. (2002) and Hrbek &amp; Mayer (2003), who analysed the genes encoding the 12S and 16S ribosomal RNAs, several transfer RNAs, and complete NADH dehydrogenase subunits I and II. These comprehensive analyses are partly congruent with the COI-based studies by Geiger et al. (2014) and Esmaeili et al. (2020). Esmaeili et al. (2020) separated  Aphaniops and Paraphanius and maintained all other species in  Aphanius, but this approach fails to account for morphological characters and ignores the ambiguous phylogenetic position of  Tellia (see Hrbek et al. 2002). Our concept of eight genera integrates morphological characters and the compactness of the monophyletic species groups in  Aphaniidae, even if this concept is challenged by the poor diagnosability of the proposed genus  Apricaphanius against  Anatolichthys .</p><p>Four of the eight aphaniid genera proposed in the present study are diagnosable by anatomical characters.  Aphaniops possesses head canals (vs. absent in the other genera except  Aphanius) and lacks a dermal sheath overlapping the base of the anterior anal-fin rays in the nuptial female (vs. present in other genera).  Tellia does not possess head canals or a pelvic fin (vs. pelvic fin present in other genera).  Kosswigichthys does not possess head canals and possesses three rows of conical teeth (vs. one row of tricuspid teeth in other genera).  Aphanius possesses a pelvic fin, a dermal sheath at the anal-fin base, and head canals.</p><p>The remaining genera  Anatolichthys, Apricaphanius, Esmaeilius, and Paraphanius share the combination of possessing a single row of tricuspid teeth, a pelvic fin, a dermal sheath at the anal-fin base, and not possessing head canals. To keep these groups of species in a single genus ( Anatolichthys) would violate the criterion of monophyly.</p><p>Anatolichthys, Apricaphanius, Esmaeilius, Paraphanius can be distinguished from one another by colour pattern. This phenomenon is particularly pronounced in Paraphanius, which exhibits an unusually derived colour pattern (see discussion below).  Anatolichthys, Apricaphanius, and Esmaeilius can in the main be distinguished by colour pattern, but three problematic species are discussed in detail below.</p></div>	https://treatment.plazi.org/id/03B187D4DF1EFF96FF4F624DFC06DE94	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Freyhof, Jörg;Yoğurtçuoğlu, Baran	Freyhof, Jörg, Yoğurtçuoğlu, Baran (2020): A proposal for a new generic structure of the killifish family Aphaniidae, with the description of Aphaniops teimorii (Teleostei: Cyprinodontiformes). Zootaxa 4810 (3): 421-451, DOI: 10.11646/zootaxa.4810.3.2
03B187D4DF18FF92FF4F60D5FC25DF1A.text	03B187D4DF18FF92FF4F60D5FC25DF1A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anatolichthys Kosswig & Sozer 1945	<div><p>Anatolichthys Kosswig &amp; Sözer, 1945</p><p>Figs. 2–3</p><p>Type species.  Anatolichthys splendens Kosswig &amp; Sözer 1945 .</p><p>Anatolichthys transgrediens is the type species of  Turkichthys Ermin, 1946, which is a synonym of  Anatolichthys (Wildekamp 1993).</p><p>Diagnosis.  Anatolichthys is distinguished from  Kosswigichthys by possessing a single (vs. three) row of tricuspid (vs. conical) teeth and presence (vs. absence) of black or dark-brown bars in the caudal fin of the male. It is distinguished from the remaining genera in the family  Aphaniidae by a combination of characters, none unique to the genus. Head canals absent (vs. present in  Aphanius and  Aphaniops); a dermal sheath at the anal-fin base in nuptial female present (vs. absent in  Aphaniops); pelvic fin present (vs. absent in  Tellia); black or dark-brown bars in the caudal fin of male present (vs. absent in Paraphanius); flank pattern in male comprising a series of black or brown bars (vs. small whitish or blue vertically-arranged spots or very narrow bars in Paraphanius); a bold, black spot at centre of the caudal-fin base in female present (vs. absent in Paraphanius); dorsal- and anal-fin margins black in male (vs. without black margins in  Aphaniops, except  A. sirhani, yellow in  Tellia, only dorsal-fin margin black in  Aphanius).</p><p>Included species.  Anatolichthys anatoliae,  A. danfordii,  A. fontinalis,  A. iconii,  A. irregularis,  A. maeandricus,  A. marassantensis,  A. meridionalis,  A. saldae,  A. splendens,  A. sureyanus,  A. transgrediens,  A. villwocki .</p><p>Distribution.  Anatolichthys is widely distributed in inland waters of central and southwestern Anatolia as well as the southern Black Sea basin, where it is found in the Sakarya, Kızılırmak and Yeşilırmak River drainages (Fig. 4, Yoğurtçuoğlu &amp; Ekmekçi 2017). Southern records from the Levant are discussed below.</p><p>Remarks.  Anatolichthys splendens, only known from Lake Gölcük, has not been found since the 1980s and is obviously extinct, following the stocking of the lake with alien fish species.</p><p>Our unpublished COI data agree with Hrbek et al. (2002) in the placement of  Anatolichthys close to  Kosswigichthys, and the sister-genus relationship of  Aphanius to these genera. With 13 member species,  Anatolichthys is the richest genus in the family  Aphaniidae, and there is significant diversity in body shape plus colour and scale patterns. In particular, the absence or reduction of scales in pelagic lacustrine species ( A. transgrediens,  A. saldae,  A. splendens, and  A. sureyanus) has been studied extensively (Ermin 1946, Grimm 1980, Villwock 1963), and they had been separated from  Aphanius in the genera  Anatolichthys and  Turkichthys by earlier authors (Kosswig &amp; Sözer 1945, Ermin 1946). Hrbek et al. (2002) found these pelagic species  Anatolichthys and  Turkichthys to be nested within a group of species previously identified as  Aphanius anatoliae and  A. danfordii .</p><p>The most atypical  Anatolichthys species is  A. villwocki, which lacks the bold, black caudal-fin bars present in all congeners. This character state is shared with  Esmaeilius sophiae, but  Anatolichthys villwocki is clearly distinguished from all  Esmaeilius species except  E. isfahanensis by possessing a black (vs. white) dorsal-fin margin. Geiger et al. (2014) placed  A. villwocki between  A. danfordii,  A. marassantensis and the other  Anatolichthys species, whereas Hrbek et al. (2002) considered it as sister to  A. meridionalis . The results of these studies suggest that the relatively high number of flank bars and the presence of rows of spots on the caudal-fin represent an apomorphic condition in  A. villwocki, since it is not shared with  A. danfordii,  A. marassantensis or  A. meridionalis . Due to its phylogenetic position within  Anatolichthys,  A. villwocki is not recognised as a distinct genus despite its higher number of flank bars and caudal-fin colour pattern.</p><p>Pellegrin (1911, 1923) published records of  Cyprinodon sophiae from the region of Damascus in Syria, and Bodenheimer (1935) and Tortonese (1938) found similar fishes elsewhere in Syria and Jordan. Krupp (1985) and JF searched for these fishes at some of these locations in the Barada River in Syria, and Suwaima in Jordan, but were unable to find them. Figures by Krupp (1985), depicting individuals collected in the 19 th century from Damascus, suggest that they belong to  Anatolichthys due to the presence of two black caudal-fin bars and a black dorsal fin with hyaline base in the male. It is clearly not a species of  Esmaeilius, which is found in the adjacent lower Tigris. Krupp (1985) discussed whether this species might represent an introduced population of  A. marassantensis but subsequently rejected this hypothesis.</p><p>Material examined.</p><p>Anatolichthys anatoliae: FSJF 2483, 15, 19–36 mm SL;   IUSHM 2017-1280, 48, 10–30 mm SL; Turkey: Aksaray prov.: spring in <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=33.5458&amp;materialsCitation.latitude=38.2414" title="Search Plazi for locations around (long 33.5458/lat 38.2414)">Köşk Park</a> within Sultanhanı, 38.2414 33.5458.  — FSJF 2485, 38, 25–45 mm SL;  IUSHM 2017-1282, 30, 21–40 mm SL; Turkey: Konya prov.: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=33.3514&amp;materialsCitation.latitude=37.9861" title="Search Plazi for locations around (long 33.3514/lat 37.9861)">Meyil Lake</a>, about 3 km southwest of Esentepe, 37.9861 33.3514.  — FSJF 2527, 9, 21–30 mm SL;  IUSHM 2017-1283, 25, 16–31 mm SL; Turkey: Konya prov.: stream north of Sarıyayla, draining to former <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=32.7592&amp;materialsCitation.latitude=39.1189" title="Search Plazi for locations around (long 32.7592/lat 39.1189)">Lake Samsam</a>, 39.1189 32.7592.  — FSJF 2610, 4, 21–38 mm SL;  IUSHM 2017-1279, 21, 12–46 mm SL; Turkey: Konya prov.: stream at Gölyazı at road from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=33.2009&amp;materialsCitation.latitude=38.5526" title="Search Plazi for locations around (long 33.2009/lat 38.5526)">Eskil</a> to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=33.2009&amp;materialsCitation.latitude=38.5526" title="Search Plazi for locations around (long 33.2009/lat 38.5526)">Cihanbeyli</a>, 38.5526 33.2009.   — FSJF 2514, 10, 16–26 mm SL; Turkey: Aksaray prov.: spring area at eastern shore of former <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=33.4427&amp;materialsCitation.latitude=38.2655" title="Search Plazi for locations around (long 33.4427/lat 38.2655)">Lake Büget</a> in Esmekaya north of the first mosque, 38.2655 33.4427.  — ZMH 3489, 10, 27–37 mm SL; Turkey: Niğde prov.: Niğde.  — ZMH 3495, 28, 28–39 mm SL; Turkey: Konya prov.: stream 20 km north of  Çumra .  — ZMH 3490, 10, 34–45 mm SL, Turkey: Aksaray prov.: Aksaray.</p><p>Anatolichthys danfordii: FSJF 2601, 16, 23–42 mm SL;   IUSHM 2017-1276, 30, 16–45 mm SL; Turkey: Kayseri prov.: spring <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.3656&amp;materialsCitation.latitude=38.3902" title="Search Plazi for locations around (long 35.3656/lat 38.3902)">Soysallı</a> west of Soysallı, west of Develi, 38.3902 35.3656.  — ZMH 3473, 1, 29 mm SL; ZMH 3474, 1, 32 mm SL;  ZMH 3475, 8, 19–27 mm SL; Turkey: Kayseri prov.: Çayırözü, about 15 km northwest of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.2894&amp;materialsCitation.latitude=38.4172" title="Search Plazi for locations around (long 35.2894/lat 38.4172)">Develi</a>, 38.4172 35.2894.   — ZMH 3476, 21, 19–36 mm SL; Turkey: Kayseri prov.: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.3631&amp;materialsCitation.latitude=38.2014" title="Search Plazi for locations around (long 35.3631/lat 38.2014)">Soysallı</a>, 38.2014 35.3631.   FSJF 2508, 2, 26–45 mm SL; Turkey: Kayseri prov.: Canal north of Senir at road to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.2525&amp;materialsCitation.latitude=38.2013" title="Search Plazi for locations around (long 35.2525/lat 38.2013)">Sultansazlığı National Park</a>, 38.2013 35.2525.  — ZMH 3477, 33, 19–43 mm SL; ZMH 3478; 1, 40.1 mm SL;  ZMH 3479, 1, 46 mm SL; Turkey: Kayseri prov.: Develi,  Ilipınar .</p><p>Anatolichthys fontinalis: FSJF 3456, 20, 22–38 mm SL; Turkey: Burdur prov.: spring in <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=29.9472&amp;materialsCitation.latitude=37.5806" title="Search Plazi for locations around (long 29.9472/lat 37.5806)">Lake Yarışlı</a> basin, 37.5806 29.9472.   — FSJF 3690, 23, 14–36 mm SL; Turkey: Burdur prov.: spring <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=30.4007&amp;materialsCitation.latitude=37.5797" title="Search Plazi for locations around (long 30.4007/lat 37.5797)">Karaevli</a>, 2 km southeast of Burdur, 37.5797 30.4007.</p><p>Anatolichthys iconii: FSJF 2325, 16, 25–32 mm SL;   IUSHM 2017-1273, 10, 23–28 mm SL; Turkey: Isparta prov.: spring Karaot at shore of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=30.9074&amp;materialsCitation.latitude=38.1349" title="Search Plazi for locations around (long 30.9074/lat 38.1349)">Lake Eğirdir</a>, about 4 km north of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=30.9074&amp;materialsCitation.latitude=38.1349" title="Search Plazi for locations around (long 30.9074/lat 38.1349)">Yenice</a> village, 38.1349 30.9074.  — FSJF 2476, 6, 28–39 mm SL;  IUSHM 2017-1274, 13, 26–34 mm SL; Turkey: Isparta prov.: lower stream Çayköy at <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=30.8916&amp;materialsCitation.latitude=37.8415" title="Search Plazi for locations around (long 30.8916/lat 37.8415)">Koysazı</a> bridge, southeast of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=30.8916&amp;materialsCitation.latitude=37.8415" title="Search Plazi for locations around (long 30.8916/lat 37.8415)">Eğirdir</a>, 37.8415 30.8916.  — FSJF 2271, 16, 18–35 mm SL;  IUSHM 2017-1281, 7, 19–32 mm SL; Turkey: Konya prov.: spring <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=31.6743&amp;materialsCitation.latitude=37.8251" title="Search Plazi for locations around (long 31.6743/lat 37.8251)">Eflatun Pınarı</a> at Sadıkhacı, 37.8251 31.6743.</p><p>Anatolichthys irregularis . FFR 08653, 1, 35 mm SL;  FFR 08654, 14, 22– 42 mm SL; FSJF 3460, 3, 29–31 mm SL;  FSJF 3461, 10, 25–30 mm SL; Turkey: Denizli prov.: spring <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=29.3852&amp;materialsCitation.latitude=37.856" title="Search Plazi for locations around (long 29.3852/lat 37.856)">Kaklık</a>, 37.8560 29.3852.</p><p>Anatolichthys maeandricus: FSJF 1876, 37, 16–40 mm SL;  FSJF 3027, 9, 27–40 mm SL; FSJF 3639, 39, 18–35 mm SL;  IUSHM 2017-1278, 9, 28–41 mm SL; Turkey: Denizli prov.: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=29.8511&amp;materialsCitation.latitude=38.3214" title="Search Plazi for locations around (long 29.8511/lat 38.3214)">Işıklı</a> spring at Işıklı, 38.3214 29.8511.  — FSJF 2470, 5, 26–38 mm SL;  IUSHM 2017-1277, 3, 25–27 mm SL; Turkey: Afyonkarahisar prov.: spring Düden, 5 km east of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=30.1756&amp;materialsCitation.latitude=38.0519" title="Search Plazi for locations around (long 30.1756/lat 38.0519)">Dinar</a>, 38.0519 30.1756.</p><p>Anatolichthys marassantensis: FSJF 3455, 29, 28–43 mm SL; Turkey: Ankara prov.: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=33.6367&amp;materialsCitation.latitude=39.1525" title="Search Plazi for locations around (long 33.6367/lat 39.1525)">Hirfanlı Reservoir</a>, 39.1525 33.6367.   — FSJF 3733, 20, 18–35 mm SL; Turkey: Ankara prov.: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=33.6327&amp;materialsCitation.latitude=39.1559" title="Search Plazi for locations around (long 33.6327/lat 39.1559)">Hirfanlı Reservoir</a> at Geçitli, 39.1559 33.6327.   — FSJF 3634, 66, 23–43 mm SL; Turkey: Kayseri prov.: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.2083&amp;materialsCitation.latitude=38.7749" title="Search Plazi for locations around (long 35.2083/lat 38.7749)">Kapuzatan</a> close to Kayseri, 38.7749 35.2083.  — ZMH 3480, 86, 19–55 mm SL; ZMH 3484, 1, 33 mm SL;  ZMH 3485, 1, 40 mm SL; Turkey: Kayseri prov.: Kayseri, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.4531&amp;materialsCitation.latitude=38.7736" title="Search Plazi for locations around (long 35.4531/lat 38.7736)">Karpuzatan</a>, 38.7736 35.4531.   — ZMH 3482, 20, 17–35 mm SL; Turkey: Samsun prov.: Lake Balık about 12 km east of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.0833&amp;materialsCitation.latitude=41.5792" title="Search Plazi for locations around (long 36.0833/lat 41.5792)">Bafra</a>, 41.5792 36.0833.  — ZMH 3481, 33, 18–28 mm SL; ZMH 26068, 18, 23–35 mm SL; Turkey: Kırşehir prov.: springs in Kırşehir.</p><p>Anatolichthys meridionalis: FSJF 2460, 7, 21–36 mm SL;   IUSHM 2017-1275, 22, 20–39 mm SL; Turkey: Antalya prov.: small field canal south of Kırkpınar, north of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=29.9181&amp;materialsCitation.latitude=37.1392" title="Search Plazi for locations around (long 29.9181/lat 37.1392)">Kızılcadağ</a>, 37.1392 29.9181.  — FSJF 3669, 3, 27–29 mm SL; Turkey: Burdur prov.: reservoir south of Yeşilova, 37.4908 29.7433.  — FSJF 3465, 14, 26–29 mm SL; Turkey: Antalya prov.: spring Kocapınar, 2 km northwest of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=29.8023&amp;materialsCitation.latitude=36.6907" title="Search Plazi for locations around (long 29.8023/lat 36.6907)">Mursal</a>, 36.6907 29.8023.   — ZMH 3502, 17, 23–39 mm SL; Turkey:  Lake Gölhisar .</p><p>Anatolichthys saldae: ZMH 3507, 40, 32–38 mm SL; Turkey: Burdur prov.: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=29.6567&amp;materialsCitation.latitude=37.5296" title="Search Plazi for locations around (long 29.6567/lat 37.5296)">Lake Salda</a>, 37.5296 29.6567.</p><p>Anatolichthys splendens: ZMH 3505, lectotype, 31 mm SL;   ZMH 3506, 1, 38 mm SL; Turkey: Isparta prov.:  Lake Gölcük .</p><p>Anatolichthys sureyanus: ZMH 7850, 40, 16–34 mm SL; Turkey: Burdur prov.: shore of  Lake Burdur at road from Burdur to Yeşilova.</p><p>Anatolichthys transgrediens: ZMH 3509, 52, 23–33 mm SL; Turkey: Afyonkarahisar prov.: spring Akpınar at the south shore of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=29.9263&amp;materialsCitation.latitude=37.8172" title="Search Plazi for locations around (long 29.9263/lat 37.8172)">Lake Acıgöl</a>, 37.8172 29.9263.</p><p>Anatolichthys villwocki: FSJF 2626, 4, 31–55 mm SL;  IUSHM 2017-1285, 6, 24–48 mm SL; FSJF 3501, 17, 29–41 mm SL; Turkey: Afyonkarahisar prov.: spring about 11 km east of Emirdağ, 39.0481 31.3272.  — FSJF 3091, 12, 20–35 mm SL; Turkey: Konya prov.: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=31.8903&amp;materialsCitation.latitude=38.3208" title="Search Plazi for locations around (long 31.8903/lat 38.3208)">Lake Ilgın</a>, 38.3208 31.8903.   — FSJF 3741, 12, 20–35 mm SL; Turkey: Eskişehir prov.: spring Başkurt 20 km southeast of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=31.1463&amp;materialsCitation.latitude=39.266" title="Search Plazi for locations around (long 31.1463/lat 39.266)">Çifteler</a>, 39.2660 31.1463.</p></div>	https://treatment.plazi.org/id/03B187D4DF18FF92FF4F60D5FC25DF1A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Freyhof, Jörg;Yoğurtçuoğlu, Baran	Freyhof, Jörg, Yoğurtçuoğlu, Baran (2020): A proposal for a new generic structure of the killifish family Aphaniidae, with the description of Aphaniops teimorii (Teleostei: Cyprinodontiformes). Zootaxa 4810 (3): 421-451, DOI: 10.11646/zootaxa.4810.3.2
03B187D4DF1BFF9CFF4F661EFE37D8BE.text	03B187D4DF1BFF9CFF4F661EFE37D8BE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aphaniops Hoedeman 1951	<div><p>Aphaniops Hoedeman, 1951</p><p>Figs. 5–6</p><p>Type species.  Lebias dispar Rüppell 1829</p><p>Diagnosis.  Aphaniops is distinguished from other genera in the family  Aphaniidae by absence of a dermal sheath at the anal-fin base in the nuptial female (vs. presence). It is further distinguished from  Anatolichthys by presence of head canals (vs. absence), and absence of black dorsal- and anal-fin margins in the male in all species except  A. sirhani (vs. presence).</p><p>Included species.  Aphaniops dispar,  A. furcatus,  A. ginaonis,  A. kruppi,  A. richardsoni,  A. sirhani,  A. stiassnyae,  A. stoliczkanus,  A. teimorii .</p><p>Distribution.  Aphaniops occurs from the southeastern Mediterranean Sea basin in Egypt and Israel southwards to Somalia and eastwards to southwestern India including the Arabian Peninsula and the Gulf basin (Fig. 7).</p><p>Remarks. We were unable to examine specimens of  A. furcatus, and the description by Teimori et al. (2014) does not provide sufficient details to determine whether this species possesses the diagnostic characters of the genus. We follow Esmaeili et al. (2020) in including it in  Aphaniops .</p><p>As discussed by Hoedeman (1951),  Aphaniops lacks the dermal sheath overlapping the base of the anterior anal-fin rays present in the nuptial females of other Aphaniids. Discrepancies in fin-ray counts between  Aphaniops and  Aphanius published by Hoedeman (1951) could not be confirmed by Villwock et al. (1983) or ourselves.</p><p>Teimori et al. (2018a) described an  Aphaniops species from Hormozgan Province (Iran) as  Aphanius hormuzensis . However, Article 16.4. of the International Code for Zoological Nomenclature (ICZN, 1999) states that the fixation of name-bearing types for a new species has to be explicit: “Every new specific and subspecific name published after 1999, except a new replacement name…, must be accompanied in the original publication 16.4.1. by the explicit fixation of a holotype,…..and 16.4.2. where the holotype or syntypes are extant specimens, by a statement of intent that they will be (or are) deposited in a collection and a statement indicating the name and location of that collection.”</p><p>That means that for species described after 1999 it has been obligatory to indicate the name and location of the collection and holotype (if extant) in new species descriptions. In the description of  Aphanius hormuzensis, the authors designated “ ZM-FISBUK 157 “ as the holotype of this new species, without mentioning which collection ZM-FISBUK might be and where this collection is located. The holotype of A.  hormuzensis is therefore invalid, because it does not fulfil the requirements of Article 16.4.2, and the name  Aphanius hormuzensis Teimori, Esmaeili, Hamidan &amp; Reichenbacher 2018 is not available following the rules of zoological nomenclature. The lead author of this study (JF) communicated this issue to Bettina Reichenbacher, one of the authors of the species description, in 2018 but it has not been resolved, hence the taxon is briefly re-described here as  Aphaniops teimorii . Due to the limited availability of materials, the redescription is largely based on data presented by Teimori et al. (2018a) in addition to our own materials.</p></div>	https://treatment.plazi.org/id/03B187D4DF1BFF9CFF4F661EFE37D8BE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Freyhof, Jörg;Yoğurtçuoğlu, Baran	Freyhof, Jörg, Yoğurtçuoğlu, Baran (2020): A proposal for a new generic structure of the killifish family Aphaniidae, with the description of Aphaniops teimorii (Teleostei: Cyprinodontiformes). Zootaxa 4810 (3): 421-451, DOI: 10.11646/zootaxa.4810.3.2
03B187D4DF15FF9EFF4F624DFDD9DE62.text	03B187D4DF15FF9EFF4F624DFDD9DE62.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aphaniops teimorii Freyhof & Yoğurtçuoğlu 2020	<div><p>Aphaniops teimorii,  new species</p><p>Fig. 6</p><p>Aphanius hormuzensis 
Teimori, Esmaeili, Hamidan &amp; Reichenbacher, 2018a, nomen nudum, (Iran, Hormuzgan Province (S-Iran), 330 m, Mehran River, Gotab village, 15 km south of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=27.1441&amp;materialsCitation.latitude=54.2628" title="Search Plazi for locations around (long 27.1441/lat 54.2628)">Bastak</a>, 54.2628 27.1441)</p><p>Holotype. ZFMK-ICH 122627, 33 mm SL; Iran: Hormozgan prov.: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=55.639&amp;materialsCitation.latitude=27.199" title="Search Plazi for locations around (long 55.639/lat 27.199)">Govdar River</a> near Kahoorestan, 27.1990 55.6390.</p><p>Paratypes. FSJF 4021, 5, 31–35 mm SL; Iran: Hormozgan prov.: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=54.2667&amp;materialsCitation.latitude=27.136" title="Search Plazi for locations around (long 54.2667/lat 27.136)">Hormalin River</a> at Bastak, 27.1360 54.2667  .  FSJF 4100, 3, 26–39 mm SL; same data as holotype (captive-bred) .</p><p>Diagnosis.  Aphaniops teimorii is distinguished by a combination of non-unique characters. It is somewhat similar to  A. ginaonis from Iran and  A. kruppi from Oman in that males possess 12–17 brown bars on the flank, the anterior-most of which is located beneath the pectoral fin and the posterior-most on the caudal-fin base (vs. flank bars in males absent or restricted to the caudal peduncle in  A. dispar,  A. richardsoni and  A. stoliczkanus). It is distinguished from  A. kruppi by presence of a long narrow bar (vs. a diamond-shaped or vertically-elongate black or dark-brown blotch) at the caudal-fin base in females, and 12–17 (vs. 9–14) brown bars on the flank in the male. According to the molecular data presented by Teimori et al. (2018a),  A. teimorii is closely related to  A. ginaonis, which is endemic to a single spring connected to a stream in which  A. teimorii occurs (Reichenbacher et al. 2009b). The two species exhibit the same colour pattern in both males and females, and possess 4–5 scale rows on the caudal-fin base.  Aphaniops teimorii is distinguished from  A. ginaonis by possessing 7½–8½ (vs. 5) branched dorsal-fin rays. Diagnostic otolith and osteological characters are provided by Teimori et al. (2018a).</p><p>Description. See Figure 6 for general appearance. Dorsal head profile straight. Dorsal profile straight or slightly convex from nape to dorsal-fin origin. Ventral profile convex. Body deeper than wide, deepest at about dorsalfin origin and widest at pectoral-fin base or centre of belly in females. Lower jaw gently upturned, oriented about 45° to body axis. Caudal peduncle compressed laterally, its length 1.6 times in its depth in holotype, 1.5–1.6 times longer than deep. Pectoral fin rounded, reaching almost to or slightly beyond pelvic-fin base. Anal-fin origin below vertical through second or third branched dorsal-fin ray. Pelvic fin not reaching or reaching anus. One large scale present between pelvic-fin bases. Anus situated slightly anterior to anal-fin origin. Dorsal and anal fins roundish in females, tip of longest dorsal-fin ray reaching to a vertical through centre of posterior anal fin ray. Dorsal and anal fins elongated in males, posterior tip of dorsal fin reaching vertical of tip of anal fin or to a point slightly before. Caudal fin rounded to truncate. Largest individual examined 39 mm SL.</p><p>Dorsal and anal fins with 7½–8½ branched rays. Caudal fin with 8+7–8+8 branched rays. Pectoral fin with 15–16 and pelvic fin with 7–8 rays. Trunk and head entirely scaled. Scales large and cycloid in females, with small ctenae in males. Scale above pectoral-fin origin enlarged. One scale row on upper portion of opercle. Flank with 24–26 scales along lateral series. 4–5 additional rows of small scales on anterior caudal-fin base. Teimori et al. (2018a) counted 27–29 flank scales but did not describe the method used. Nine scale rows between dorsal- and pelvic-fin origins. 14 circumpeduncular scales. Teeth tricuspid, median tip longer than laterals.</p><p>Colouration. See Figure 6 for general appearance. Live and preserved males: all yellow, orange, silvery and blue colours faded in preserved specimens. Lateral head and flank silvery to whitish with brown or dark-grey pattern of bars and blotches, dorsal head and back brown or dark-grey. Lower cheek, breast and belly whitish or pale yellow. Lateral head and flank with a bluish hue in life. Flank between pectoral-fin base and vertical through pelvicfin origin with a brown or yellowish-brown network pattern forming roundish silvery or pale-bluish blotches, often just a few silvery blotches on grey or brown background. Flank with 12–17 bars, confluent with brown or dark-grey back. Bars brown in life, black in preserved individuals. Interspaces silvery, narrower than bars. Pectoral fin hyaline or greyish-blue in life, blackish in preserved individuals. Pelvic fin hyaline or white. Anal fin hyaline or yellow in life, whitish in preserved individuals, greyish-blue or yellow anteriorly with 2–4 narrow black bars. Dorsal fin with irregularly set and shaped bands, often restricted to blotches on rays. Caudal fin hyaline with 2 wide, black bars. A pale grey or black blotch on unbranched caudal-fin rays at upper and lower extremities. Living and preserved females: top of head and back pale-brown. Cheek, ventral surface of head, belly and flank silvery-grey or pale-brown. Flank with a series of 12–17 narrow, vertically-elongate bars along lateral midline, bars anterior to vertical through dorsal-fin origin often faded. A narrow dark-brown or black bar at centre of caudal-fin base. All fins hyaline in life, grey in preserved individuals.</p><p>Distribution.  Aphaniops teimorii has been collected from coastal rivers and streams between the Merhan and Minhab River drainages in southern Iran (Esmaeili et al. 2020) (Figure 7).</p><p>Etymology. Named for Azad Teimori (Kerman, Iran) for his many valuable contributions to the biology of Iranian killifishes. A noun in genitive, indeclinable.</p><p>Material examined.</p><p>Aphaniops cf. dispar: FSJF 3487, 4, 32–43 mm SL; Israel: salt marshes at <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=34.9403&amp;materialsCitation.latitude=32.6919" title="Search Plazi for locations around (long 34.9403/lat 32.6919)">Atlit</a>, 32.6919 34.9403.   — FSJF 3636, 4, 40–45 mm SL; Eritrea: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=40.3296&amp;materialsCitation.latitude=14.6202" title="Search Plazi for locations around (long 40.3296/lat 14.6202)">Shukoray River</a>, 14.6202 40.3296.   — FSJF DNA-2599, 2, 35–36 mm SL; Egypt: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=33.6778&amp;materialsCitation.latitude=27.3886" title="Search Plazi for locations around (long 33.6778/lat 27.3886)">el-Guna</a>, Qesm Hurghada, 27.3886 33.6778.   — TAUP 6318, 10, 27–37 mm SL; Egypt: En Mussa,  Sinai, about 60 km inland.   — ZMH 4379, 20, 28–38 mm SL; Egypt: creeks in  Siwa Oasis .  — ZMH 7549, 26, 21–44 mm SL; Egypt: Lakes in Wadi El-Raiyan about 65 km southwest of Faiyum.  — ZMH 7548, 26, 22–56 mm SL; Egypt:  Bitter Lake .  — ZMH 7550, 2, 46–62 mm SL; ZMH 7551, 2, 45–46 mm SL; ZMH 7552, 2, 37–49 mm SL; ZMH 7553, 2, 44–47 mm SL; ZMH 7554, 2, 47–49 mm SL;  ZMH 7555, 2, 36–55 mm SL; Egypt:  Lake Timsah south of Ismailia.  — ZMH 13176, 2, 51–54 mm SL; Egypt: Cairo. — ZMH 13177, 4, 27–41 mm SL; Egypt: Suez.  — ZMH 4376, 14, 24–44 mm SL; Saudi Arabia:  Sarso Island in Farasan Archipelago.  — ZMH 13181, 4, 24–31 mm SL; ZMH 13206, 14, 22–37 mm SL; Yemen: Socotra. — ZMH 13183, 4, 46–49 mm SL;  ZMH 13196, 1, 46 mm SL; Ethiopia: north-west of  Harrar .</p><p>Aphaniops ginaonis: FSJF 3274, 40, 16–36 mm SL;   FSJF 3275, 37, 16–34 mm SL; Iran: spring <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=56.3056&amp;materialsCitation.latitude=27.4411" title="Search Plazi for locations around (long 56.3056/lat 27.4411)">Geno</a>, 27.4411 56.3056.</p><p>Aphaniops teimorii: FSJF 4021, 5, 31–35 mm SL;   FSJF 3275, 37, 16–34 mm SL; Iran: Hormuzgan prov.: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=54.2667&amp;materialsCitation.latitude=27.136" title="Search Plazi for locations around (long 54.2667/lat 27.136)">Hormalin</a> river at Bastak, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=54.2667&amp;materialsCitation.latitude=27.136" title="Search Plazi for locations around (long 54.2667/lat 27.136)">Mirahmad</a>, 27.1360 54.2667.</p><p>Aphaniops kruppi: ZFMK-ICH 103668, holotype, 46 mm SL;   FSJF 3671, paratypes, 69, 21– 51 mm SL; Oman: spring in <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=58.6752&amp;materialsCitation.latitude=22.6129" title="Search Plazi for locations around (long 58.6752/lat 22.6129)">Al Mudayrib</a>, 22.6129 58.6752.   FSJF 3537, 2, 26–32 mm SL; Oman: Wadi Bani Khalid at <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=59.0869&amp;materialsCitation.latitude=22.5953" title="Search Plazi for locations around (long 59.0869/lat 22.5953)">Sayh al Hayl</a>, 22.5953 59.0869.  — FSJF 4088, 25, 27–39 mm SL; Oman: Al Mudayrib, 22.6128 58.6675.  — FSJF 4091, 10, 26–43 mm SL; Oman: Falaj in <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=59.3179&amp;materialsCitation.latitude=22.0239" title="Search Plazi for locations around (long 59.3179/lat 22.0239)">Bani Bu Ali</a>, 22.0239 59.3179.   — FSJF 4101, 50, 18–38 mm SL; Oman: Wadi Bani Khalid north of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=59.093&amp;materialsCitation.latitude=22.619" title="Search Plazi for locations around (long 59.093/lat 22.619)">Muqal</a>, 22.619 59.093.</p><p>Aphaniops cf. kruppi: FSJF DNA-2590, 4, 25–54 mm SL; Oman: stream <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=57.3583&amp;materialsCitation.latitude=23.0756" title="Search Plazi for locations around (long 57.3583/lat 23.0756)">Al Hoota</a> below <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=57.3583&amp;materialsCitation.latitude=23.0756" title="Search Plazi for locations around (long 57.3583/lat 23.0756)">Al Hoota Cave</a>, 23.0756 57.3583.</p><p>Aphaniops richardsoni: ZMH 2407, 30, 30–45 mm SL;  ZMH 4377, 30, 30–42 mm SL;  ZMH 4378, 20, 18–43 mm SL; Israel:  Ein Feshka .</p><p>Aphaniops sirhani: FSJF 3672, 12, 23–44 mm SL; Jordan: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.8201&amp;materialsCitation.latitude=31.8342" title="Search Plazi for locations around (long 36.8201/lat 31.8342)">Azraq Oasis</a>, 31.8342 36.8201  .</p><p>Aphaniops stoliczkanus: FSJF 3532, 34, 27–44 mm SL; Oman: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=58.1069&amp;materialsCitation.latitude=23.4581" title="Search Plazi for locations around (long 58.1069/lat 23.4581)">Wadi Fanja</a> in Fanja, 23.4581 58.1069.—   FSJF 3533, 19, 23–35 mm SL; Oman: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=58.5208&amp;materialsCitation.latitude=23.54" title="Search Plazi for locations around (long 58.5208/lat 23.54)">Wadi</a> northeast of Al Amarat, 23.5400 58.5208.—   FSJF 3670, 16, 21–31 mm SL; India: Gujarat: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=69.7174&amp;materialsCitation.latitude=22.4428" title="Search Plazi for locations around (long 69.7174/lat 22.4428)">Narara-Salt Pans</a> at Vadinar, 22.4428 69.7174.—   FSJF 3673, 2, 36–39 mm SL; Iran: Southern Sistan and Baluchistan prov.: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=60.7464&amp;materialsCitation.latitude=26.2596" title="Search Plazi for locations around (long 60.7464/lat 26.2596)">Kajou River</a> at <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=60.7464&amp;materialsCitation.latitude=26.2596" title="Search Plazi for locations around (long 60.7464/lat 26.2596)">Ghasr-Ghand</a>, 26.2596 60.7464.—   FSJF 4002, 6, 33–42 mm SL; Iraq: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=47.8312&amp;materialsCitation.latitude=30.5395" title="Search Plazi for locations around (long 47.8312/lat 30.5395)">Shatt al Arab</a> at Basra, 30.5395 47.8312.—   FSJF DNA-2600, 2, 35–38 mm SL; UAE: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=56.3614&amp;materialsCitation.latitude=25.0142" title="Search Plazi for locations around (long 56.3614/lat 25.0142)">Khor Kalba</a>, 25.0142 56.3614.—   FSJF 4096, 14, 29–37 mm SL; Oman: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=56.1862&amp;materialsCitation.latitude=24.7129" title="Search Plazi for locations around (long 56.1862/lat 24.7129)">Hatta</a> pools, 24.7129 56.1862.—   FSJF 4074, 30, 27–44 mm SL; Oman: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=56.1057&amp;materialsCitation.latitude=24.5491" title="Search Plazi for locations around (long 56.1057/lat 24.5491)">Al Juwayf</a>, 24.5491 56.1057.—   FSJF 4082, 2, 32–34 mm SL; Oman: outflow of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=57.3681&amp;materialsCitation.latitude=23.0718" title="Search Plazi for locations around (long 57.3681/lat 23.0718)">Ghubrat Tanuf Cave</a>, 23.0718 57.3681.—   FSJF 4087, 2, 28–30 mm SL; Oman: Wadi Kabbah about 3 km east of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=58.5943&amp;materialsCitation.latitude=22.9835" title="Search Plazi for locations around (long 58.5943/lat 22.9835)">Saman</a>, 22.9835 58.5943.—   FSJF DNA-2603, 2, 31–45 mm SL; Bahrain: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=50.6167&amp;materialsCitation.latitude=26.1" title="Search Plazi for locations around (long 50.6167/lat 26.1)">Jirdab</a>, 26.1000 50.6167.—   ZMH 4374, 23, 19–39 mm SL; Pakistan:  Salt Pans of Karachi and  Hawks Bay. —  ZMH 4373, 30, 21–41 mm SL; Iraq: south of Karbala (likely from Euphrates).—  ZMH 4380, 4, 20–30 mm SL; Iraq:  Al Fallűjah .</p></div>	https://treatment.plazi.org/id/03B187D4DF15FF9EFF4F624DFDD9DE62	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Freyhof, Jörg;Yoğurtçuoğlu, Baran	Freyhof, Jörg, Yoğurtçuoğlu, Baran (2020): A proposal for a new generic structure of the killifish family Aphaniidae, with the description of Aphaniops teimorii (Teleostei: Cyprinodontiformes). Zootaxa 4810 (3): 421-451, DOI: 10.11646/zootaxa.4810.3.2
03B187D4DF10FF98FF4F60D5FAB2DB2E.text	03B187D4DF10FF98FF4F60D5FAB2DB2E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aphanius Nardo 1827	<div><p>Aphanius Nardo, 1827</p><p>Fig. 8</p><p>Type species.  Aphanius nanus Nardo 1827 is the type species of  Aphanius, but is a synonym of  A. fasciatus (Kottelat et al. 2007) .  Micromugil timidus Gulia 1861, also a synonym of  Aphanius fasciatus, is the type species of  Micromugil Gulia, 1861 .</p><p>Diagnosis.  Aphanius is distinguished from other genera in the family  Aphaniidae by the unique male colour pattern, comprising a pale to deep yellow or orange caudal fin, well distinct from silvery interspaces between grey or brown flank bars. Although the caudal fin can also be yellow in some  Anatolichthys species, particularly  A. iconii, the interspaces of the flank bars are the same colour as the caudal fin.  Aphanius is further distinguished by presence of a dermal sheath at the anal-fin base in the nuptial female (vs. absence in  Aphaniops); presence of a black dorsalfin margin in the male (vs. absence in all  Aphaniops except  A. sirhani); and presence of head canals (vs. absence in Anatolichtys).</p><p>Included species.  Aphanius almiriensis,  A. fasciatus</p><p>Distribution. The genus  Aphanius is widespread in coastal lagoons and estuaries of the Mediterranean Sea basin (Fig. 9), where it is only absent from southern France (extirpated), Spain (introduced) and Morocco west of the Moulouya River estuary (Valdesalici et al. 2019).</p><p>Remarks.  Aphanius almiriensis and  A. fasciatus are superficially very similar and difficult to identify in the field. Kottelat et al. (2007) and Valdesalici et al. (2019) discussed the characters distinguishing these two species in detail. Cavraro et al. (2017) documented the existence of intraspecific genetic diversity between different populations of  A. fasciatus . These differences are clearly intraspecific and specimens we examined could not be distinguished morphologically.</p><p>Material examined.</p><p>Aphanius almiriensis: FSJF 3738, 23, 18–40 mm SL; Turkey: Çanakkale prov.: estuary of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=26.1695&amp;materialsCitation.latitude=39.5693" title="Search Plazi for locations around (long 26.1695/lat 39.5693)">Tuzla River</a>, 39.5693 26.1695.</p><p>Aphanius fasciatus: FSJF 153, 3, 18–28 mm SL; Italy: Sardinia, Brackish lagoon north of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=9.6833&amp;materialsCitation.latitude=39.95" title="Search Plazi for locations around (long 9.6833/lat 39.95)">Arbarax</a>, 39.9500 09.6833.   — FSJF 209, 6, 11–29 mm SL; Italy: Sardinia, Brackish lagoon at <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=9.0333&amp;materialsCitation.latitude=39.2667" title="Search Plazi for locations around (long 9.0333/lat 39.2667)">Cagliari</a>, 39.2667 09.0333.   — FSJF 2141, 32, 13–28 mm SL; Italy: Laguna di Comacchio, southeast of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.1847&amp;materialsCitation.latitude=44.6612" title="Search Plazi for locations around (long 12.1847/lat 44.6612)">Comacchio</a>, 44.6612 12.1847.   — FSJF 2149, 40, 18–48 mm SL; Italy: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.225&amp;materialsCitation.latitude=45.215" title="Search Plazi for locations around (long 12.225/lat 45.215)">Laguna Veneto</a>, at road to Chioggia, 45.2150 12.2250.   — FSJF 3012, 50, 15–47 mm SL; Tunisia: Oued Zahzah two km south of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=10.16112&amp;materialsCitation.latitude=35.823" title="Search Plazi for locations around (long 10.16112/lat 35.823)">Bechechema</a>, 35.8230 10.16112.   — FSJF 3278, 30, 17–33 mm SL; Algeria: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=6.0992&amp;materialsCitation.latitude=33.205" title="Search Plazi for locations around (long 6.0992/lat 33.205)">Lake at Meggarine</a>, 33.2050 06.0992.  — FSJF 4038, 2, 27–35 mm SL; Croatia: Lagoon in the city of Pag, 44.4397 15.0558.</p></div>	https://treatment.plazi.org/id/03B187D4DF10FF98FF4F60D5FAB2DB2E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Freyhof, Jörg;Yoğurtçuoğlu, Baran	Freyhof, Jörg, Yoğurtçuoğlu, Baran (2020): A proposal for a new generic structure of the killifish family Aphaniidae, with the description of Aphaniops teimorii (Teleostei: Cyprinodontiformes). Zootaxa 4810 (3): 421-451, DOI: 10.11646/zootaxa.4810.3.2
03B187D4DF11FF9BFF4F662CFE72DB73.text	03B187D4DF11FF9BFF4F662CFE72DB73.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kosswigichthys Sozer 1942	<div><p>Kosswigichthys Sözer, 1942</p><p>Fig. 10</p><p>Type species.  Kosswigichthys asquamatus Sözer 1942</p><p>Diagnosis.  Kosswigichthys is distinguished from all other genera in the family  Aphaniidae by possessing three (vs. one) rows of conical (vs. tricuspid) teeth. It is further distinguished from  Anatolichthys,  Aphanius, and  Aphaniops by absence of black or dark-brown bars in the caudal fin of the male (vs. presence, except in  Aphaniops furcatus), possession of a hyaline caudal fin in the male (vs. yellow or orange in  Aphanius), absence of head canals (vs. presence in  Aphanius and  Aphaniops), presence of a dermal sheath at the anal-fin base in the nuptial female (vs. absence in  Aphaniops), and possession of a naked body (vs. covered by scales in  Aphanius).</p><p>Included species.  Kosswigichthys asquamatus</p><p>Distribution.  Kosswigichthys is endemic to Lake Hazar in Eastern Turkey (Fig. 4).</p><p>Remarks. The phylogenetic analysis by Hrbek et al. (2002) and our own unpublished COI data place  K. asquamatus as the sister clade to  Anatolichthys . The morphological differences between  Kosswigichthys and  Anatolichthys are very clear, and might be related to the specialised pelagic ecology of  K. asquamatus . The slender body shape and lack of scales are comparable to the pelagic  Anatolichthys species  A. saldae and  A. splendens, but in contrast  K. asquamatus has conical teeth, no bars on the caudal fin in males, and a slightly upturned (vs. distinctly superior) lower jaw. As in the case of the sympatric loach  Oxynoemacheilus hazerensis,  K. asquamatus seems to be a relict species in the ancient rift lake Hazar (Freyhof et al. 2019).</p><p>Material examined.</p><p>Kosswigichthys asquamatus: FSJF 2511, 30, 25–36 mm SL; Turkey: Elazığ prov.: north-eastern shore of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=39.3016&amp;materialsCitation.latitude=38.4733" title="Search Plazi for locations around (long 39.3016/lat 38.4733)">Lake Hazar</a>, 38.4733 39.3016.</p></div>	https://treatment.plazi.org/id/03B187D4DF11FF9BFF4F662CFE72DB73	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Freyhof, Jörg;Yoğurtçuoğlu, Baran	Freyhof, Jörg, Yoğurtçuoğlu, Baran (2020): A proposal for a new generic structure of the killifish family Aphaniidae, with the description of Aphaniops teimorii (Teleostei: Cyprinodontiformes). Zootaxa 4810 (3): 421-451, DOI: 10.11646/zootaxa.4810.3.2
03B187D4DF12FF87FF4F6635FBDBD89A.text	03B187D4DF12FF87FF4F6635FBDBD89A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Paraphanius Esmaeili, Teimori, Zarei & Sayyadzadeh 2020	<div><p>Paraphanius Esmaeili, Teimori, Zarei &amp; Sayyadzadeh, 2020</p><p>Fig. 11</p><p>Type species.  Lebias mento Heckel, 1843</p><p>Diagnosis.  Paraphanius is distinguished from all other genera in the family  Aphaniidae by the unique nuptial male colour pattern, comprising a dark grey, bluish or almost black background colour with irregular or regular iridescent bluish, white or silvery spots, often arranged in vertical rows or narrow bars (vs. background colour silvery with brown or black bars in  Anatolichthys,  Aphanius,  Kosswigichthys and most  Aphaniops species, males with silvery vermiculation or roundish to ovoid silvery spots or blotches on grey background in other  Aphaniops species). Male Paraphanius are further distinguished by absence of bars in the caudal fin and instead possess very narrow, blue-white or silvery rows of spots or small blotches forming bands on a grey, black or blue background (vs. bars presence in caudal fin of  Anatolichthys,  Aphanius, and all  Aphaniops except  A. furcatus). Female Paraphanius are distinguished from other Aphaniid genera by presence of numerous silvery spots or small blotches on the flank (vs. absence, usually with narrow bars or brown or black blotches or spots) and absence of a bar or diamond-shaped to roundish bold black blotch at the centre of the caudal-fin base (vs. presence in  Anatolichthys,  Aphaniops, and  Aphanius).</p><p>Paraphanius is further distinguished by presence of a dermal sheath at the anal-fin base in the nuptial female (vs. absence in  Aphaniops), absence of head canals (vs. presence in  Aphaniops and  Aphanius), and absence of black dorsal- and anal-fin margins in the male (vs. presence in  Anatolichthys and  Kosswigichthys, presence of black dorsal-fin margin only in  Aphanius), possession of a single row of tricuspid teeth (vs. three rows of conical teeth in  Kosswigichthys), and the body being completely covered by scales (vs. naked in  Kosswigichthys).</p><p>Included species.  Paraphanius alexandri,  P. boulengeri,  P. mento,  P. mentoides,  P. orontis,  P. similis,  P. striptus .</p><p>Distribution. Paraphanius species are widespread from Antalya to the Seyhan, Ceyhan and Orontes drainages on the Mediterranean coast of Turkey (Kara et al. 2010, Erk’akan &amp; Özdemir 2011, Wildekamp et al. 1999), and southwards to the northern Dead Sea basin in the Levant. Paraphanius is also widespread in the Tigris and Euphrates river drainages from northern Syria to the Iraqi Marshes (Krupp 1985). A translocated population of  P. mentoides inhabits the crater lake Nemrut in eastern Turkey, and some landlocked populations exist in central Turkey (Geiger et al. 2014) (Fig. 12).</p><p>Remarks. The colour pattern of Paraphanius species is clearly distinct from that exhibited by other Aphaniids, which typically comprises a series of bars on a silvery background. This pattern could be interpreted as being a reverse (silvery bars on brown background in Paraphanius vs. brown bars on a silvery background) of the situation found in other aphaniid genera. These bars can be partly (in some  Aphaniops and  Apricaphanius species) or completely (in some  Aphaniops spp. and  Apricaphanius saourensis) dissociated into spots or a vermiculate flank pattern. In juvenile individuals the flank is silvery with a brown mottled pattern in all Aphaniids analysed for this character ( A. anatoliae,  A. irregularis,  A. maeandricus,  A. marassantensis,  A. meridionalis,  A. villwocki,  A. fasciatus,  A. dispar,  A. kruppi,  A. stoliczkanus,  P. mento,  P. mentoides,  P. orontis,  P. similis,  P. striptus .  T. apoda,  A. baeticus,  A. saourensis,  E. sophiae,  E. isfahanensis,  E. persicus, and  E. vladykovi). As they grow, male Paraphanius develop indistinct dark-brown bars with pale brown interspaces, which are often only visible in live individuals with aggressive mood. The brown bars later become wider, often partly confluent to each other, and silvery spots appear in the interspaces, which often form narrow bars.  Paraphanius striptus is a valid species, and the only member of the genus to maintain a distinct pattern of wide dark-bluish bars with narrow silvery interspaces throughout ontogeny. In other Paraphanius species, the development of silvery spots is not restricted to males and or interspaces between the flank bars, and some species, particularly  P. alexandri and  P. mentoides, develop minute silvery spots over the entire flank which entirely mask the barred pattern. The silvery spots may be organised in vertical rows or narrow bars in juvenile individuals, but these typically dissociate in adults except in  P. mentoides,  P. striptus, and some populations of  P. mento .</p><p>This pattern of ontogenetic development clearly demonstrates that the flank pattern of silvery bars or spots on a dark background exhibited by Paraphanius species is not a reverse condition of that seen in other Aphaniids. In reality, the dark-grey, brown or black bars are so wide that the silvery interspaces appear to form bars, or they become confluent, with the silvery spots formed by small gaps in the dark pigmentation. Comparable colour patterns are exhibited by  Anatolichthys irregularis (Fig. 3), in which the adult male appear black with a pattern of yellowish blotches (Yoğurtçuoğlu &amp; Freyhof 2018), and some male  Apricaphanius which appear blackish with whitish spots.</p><p>The distribution map of Paraphanius species (Fig. 11) suggests that  P. mento is geographically separated from  P. striptus (Fig. 12). However, Goren (1974) suggested that both species occur in Israel, and in some cases occur in sympatry. This situation has not been revisited, because we did not possess sufficient material to confirm the occurrence of  P. mento in the Jordan River drainage within the framework of this study.</p><p>Esmaeili et al. (2020) clearly based their diagnosis of Paraphanius on Parenti (1981), because they examined only two juvenile individuals (24, 27 mm SL) from Beirut (Lebanon). Parenti (1981) examined eight individuals, but it remains unclear of which species because her specimens originated from the aquarium trade. In AMNH I-28610 there are three cleared and stained individuals, and we expect that the published osteological characters were based only on these specimens.</p><p>Parenti (1981) stated that  P. mento possesses an upturned lower jaw (vs. not upturned in other Aphaniid species). However, there exists a wide variation in mouth morphology throughout the family, particularly among Paraphanius and  Anatolichthys species, rendering this character unsuitable for diagnosis of Aphaniid genera.</p><p>Esmaeili et al. (2020) distinguished Paraphanius from  Aphaniops by possession of 9–14 (vs. 8–9) dorsal-fin rays. However, Villwock et al. (1983) counted 8–12 dorsal-fin rays in  Aphaniops, while Akşiray (1948) counted 7– 11 branched dorsal-fin rays in some Turkish Paraphanius species. The large overlap in dorsal-fin ray counts indicate that this character cannot be used to distinguish these genera. Parenti (1981) stated that  P. mento possesses a unique pattern of neuromasts on the head, but we did not find it distinctive from other Aphaniids lacking head canals.</p><p>Parenti (1981) also stated: “In  A. mento, as well as in  Orestias,  Kosswigichthys, and  Anatolichthys, the interhyal is cartilaginous and the urohyal is embedded ….”. Esmaeili et al. (2020), who treated  Kosswigichthys and  Anatolichthys as synonyms of  Aphanius, misinterpreted this information when they wrote: “Also, according to Parenti [1981],  Aphanius mento possesses several diagnostic features not found in other members of the genus  Aphanius, including a cartilaginous interhyal (ossified in other  Aphanius species), an embedded urohyal (not embedded in other  Aphanius),…”. However, Parenti (1981) clearly confirmed that she examined  Paraphanius mento,  Kosswigichthys asquamatus,  Anatolichthys transgrediens and  A. splendens, and that all share these two osteological characters. Teimori et al. (2018b) examined the urohyal bone in several aphaniid species and found Paraphanius to be nested within  Aphaniops in a cladogram, but mention that  Paraphanius mento can be distinguished from the studied  Aphaniops species by morphometry of the urohyal bone. More research is needed to verify if the urohyal bone in Paraphanius is a unique shape.</p><p>Esmaeili et al. (2020) additionally distinguished Paraphanius by certain otolith characters and a thin or less developed epural (vs. thick and well developed in  Aphanius and  Aphaniops) but give no details, which species were examined and how variable this character is. Esmaeili et al. (2020) only examined two juvenile Paraphanius. We recommend that these characters should be re-examined using a larger series of material.</p><p>While we fully support the conclusion of Esmaeili et al. (2020) to treat Paraphanius as a distinct genus, the only definitive diagnostic characters confirmed during the present study were elements of the unique colour pattern.</p><p>Material examined.</p><p>Paraphanius alexandri: FSJF 2318, 21, 22–33 mm SL; Turkey: Hatay prov.: stream <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.886&amp;materialsCitation.latitude=36.3992" title="Search Plazi for locations around (long 35.886/lat 36.3992)">Arsuz east of Arsuz</a>, 36.3992 35.8860.   — FSJF 2605, 12, 34–46 mm SL; Turkey: Kahramanmaraş prov.: spring <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=37.1248&amp;materialsCitation.latitude=37.2812" title="Search Plazi for locations around (long 37.1248/lat 37.2812)">Çöçelli north of Çöçelli</a>, south of Kahramanmaraş, 37.2812 37.1248.   — FSJF 3464, 13, 22–39 mm SL; Turkey: Kahramanmaraş prov.: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=37.2194&amp;materialsCitation.latitude=38.1831" title="Search Plazi for locations around (long 37.2194/lat 38.1831)">Elbistan</a>, 38.1831 37.2194.</p><p>Paraphanius boulengeri: FSJF 2506, 12, 24–37 mm SL;   FSJF 3737, 3, 32–39 mm SL; Turkey: Adıyaman prov.: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=37.6263&amp;materialsCitation.latitude=37.7903" title="Search Plazi for locations around (long 37.6263/lat 37.7903)">River connecting lakes Gölbaşı</a> and <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=37.6263&amp;materialsCitation.latitude=37.7903" title="Search Plazi for locations around (long 37.6263/lat 37.7903)">Azaplı</a> south of Gölbaşı, 37.7903 37.6263.</p><p>Paraphanius mento: NMW 59832, 5 35–40 mm SL; Iraq:  Mossul .   — FSJF 2650, 57, 16–52 mm SL; Syria: spring of stream <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.0555&amp;materialsCitation.latitude=33.6753" title="Search Plazi for locations around (long 36.0555/lat 33.6753)">Barada</a> north-west of Damascus, 33.6753 36.0555.   — FSJF 2689, 17, 29–44 mm SL; Syria: spring <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.9248&amp;materialsCitation.latitude=33.2332" title="Search Plazi for locations around (long 35.9248/lat 33.2332)">Al Fawwar</a>, 33.2332 35.9248.   — FSJF 2701, 21, 27–46 mm SL; Syria: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.9707&amp;materialsCitation.latitude=33.2935" title="Search Plazi for locations around (long 35.9707/lat 33.2935)">Nahr al Tammasiyyar</a> near Magsoofa, 33.2935 35.9707.   — FSJF 2657, 8, 22–32 mm SL; Syria: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.5628&amp;materialsCitation.latitude=35.2716" title="Search Plazi for locations around (long 36.5628/lat 35.2716)">River Orontes</a> at Shayzar, 35.2716 36.5628.   — FSJF 2659, 2, 31–43 mm SL; Syria: spring south of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.3106&amp;materialsCitation.latitude=35.3303" title="Search Plazi for locations around (long 36.3106/lat 35.3303)">Qala’</a> at al Jarras, 35.3303 36.3106.   — FSJF 2683, 16, 21–36 mm SL; Syria: River Orontes at Mashr’a al <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.3958&amp;materialsCitation.latitude=35.9508" title="Search Plazi for locations around (long 36.3958/lat 35.9508)">Bouz</a>, 35.9508 36.3958.   — FSJF 2691, 14, 22–34 mm SL; Syria: Reservoir of Nahr al <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.0156&amp;materialsCitation.latitude=35.3419" title="Search Plazi for locations around (long 36.0156/lat 35.3419)">Hawaiz</a>, 35.3419 36.0156.   — FSJF 2758, 54, 16–40 mm SL; Syria: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.5336&amp;materialsCitation.latitude=34.5938" title="Search Plazi for locations around (long 36.5336/lat 34.5938)">River Orontes</a> at Al Ghassaniyya, 34.5938 36.5336.  — FSJF 4001, 2, 31–32 mm SL; Iraq: Shatt al Arab at Basra, 30.5395 47.832.</p><p>Paraphanius mentoides: FSJF 2270, 39, 26–61 mm SL; Turkey: Antalya prov.: spring <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=30.5805&amp;materialsCitation.latitude=37.1098" title="Search Plazi for locations around (long 30.5805/lat 37.1098)">Kırkgöz</a>, 37.1098 30.5805.   — FSJF 3108, 7, 35–42 mm SL; Turkey: Antalya prov.: stream at <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=30.5918&amp;materialsCitation.latitude=37.0239" title="Search Plazi for locations around (long 30.5918/lat 37.0239)">Döşemealtı</a>, 37.0239 30.5918.   — FSJF 3678, 26, 28–66 mm SL; Turkey: Bitlis prov.: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=42.2367&amp;materialsCitation.latitude=38.6444" title="Search Plazi for locations around (long 42.2367/lat 38.6444)">Nemrut Lake</a>, 38.6444 42.2367.</p><p>Paraphanius orontis: FSJF 2431, 9, 18–29 mm SL; Turkey: Hatay prov.: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.0785&amp;materialsCitation.latitude=36.0974" title="Search Plazi for locations around (long 36.0785/lat 36.0974)">River Orontes</a> at Sinanlı, 36.0974 36.0785.   — FSJF 3490, 40, 25–37 mm SL; Turkey: Antalya prov.: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=31.4516&amp;materialsCitation.latitude=36.7524" title="Search Plazi for locations around (long 31.4516/lat 36.7524)">Titreyengöl</a>, 36.7524 31.4516.</p><p>Paraphanius similis: FSJF 3126, 14, 29–37 mm SL; Turkey: Konya prov.: Shallow pool east of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=34.0865&amp;materialsCitation.latitude=37.539" title="Search Plazi for locations around (long 34.0865/lat 37.539)">Ereğli</a>, 37.5390 34.0865.   — FSJF 2434, 9, 15–24 mm SL; Turkey: Adana prov.: Seyhan below water regulation doors at <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.3354&amp;materialsCitation.latitude=36.9754" title="Search Plazi for locations around (long 35.3354/lat 36.9754)">Yüreyir</a>, south of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.3354&amp;materialsCitation.latitude=36.9754" title="Search Plazi for locations around (long 35.3354/lat 36.9754)">Adana</a>, 36.9754 35.3354.   — FSJF 4012, 5, 26–30 mm SL; Turkey: Niğde prov.: spring in <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=34.2499&amp;materialsCitation.latitude=37.8299" title="Search Plazi for locations around (long 34.2499/lat 37.8299)">Zengen</a>, 37.8299 34.2499.</p><p>Paraphanius striptus: FSJF 2680, 1, 22 mm SL; Syria: Canal draining from spring at <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.0092&amp;materialsCitation.latitude=32.7389" title="Search Plazi for locations around (long 36.0092/lat 32.7389)">Al Asha’ari</a>, 32.7389 36.0092.   — FSJF 3495, 62, 19–31 mm SL; Israel: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=34.9239&amp;materialsCitation.latitude=32.551" title="Search Plazi for locations around (long 34.9239/lat 32.551)">Einot Timsach</a> east of Ma’agan Micha’el, 32.5510 34.9239.   — FSJF 3529, 42, 23–36 mm SL; Israel: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.5247&amp;materialsCitation.latitude=32.5031" title="Search Plazi for locations around (long 35.5247/lat 32.5031)">Nachal HaKibbutzim</a>, 32.5031 35.5247.</p></div>	https://treatment.plazi.org/id/03B187D4DF12FF87FF4F6635FBDBD89A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Freyhof, Jörg;Yoğurtçuoğlu, Baran	Freyhof, Jörg, Yoğurtçuoğlu, Baran (2020): A proposal for a new generic structure of the killifish family Aphaniidae, with the description of Aphaniops teimorii (Teleostei: Cyprinodontiformes). Zootaxa 4810 (3): 421-451, DOI: 10.11646/zootaxa.4810.3.2
03B187D4DF0EFF86FF4F6191FF32DE49.text	03B187D4DF0EFF86FF4F6191FF32DE49.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tellia Gervais 1853	<div><p>Tellia Gervais, 1853</p><p>Fig. 13</p><p>Type species.  Tellia apoda Gervais 1853</p><p>Diagnosis.  Tellia is distinguished from all other genera in the family  Aphaniidae by absence (vs. presence) of the pelvic fin. It is further distinguished by presence of a wide, black sub-marginal bar bordered by a yellow marginal band in the anal, dorsal and caudal fins of the male (vs. 0–14 bars or bands in the caudal fin, no distinct yellow margin in anal, dorsal and caudal fins, caudal fin entirely yellow in  Aphanius species), plus a combination of non-unique characters as follows: head canals absent (vs. present in  Aphaniops and  Aphanius); dermal sheath at the anal-fin base in the nuptial female present (vs. absent in  Aphaniops); flank pattern in the male comprising a series of black or brown bars (vs. small whitish or blue spots, arranged in vertical series or very narrow bars in Paraphanius); a bold, black spot at centre of the caudal-fin base in the female present (vs. absent in Paraphanius).</p><p>Included species.  Tellia apoda .</p><p>Distribution.  Tellia is restricted to northern Algeria (Pellegrin 1921) (Fig. 14) but only a captive population originating from a single collection by Van Der Zee &amp; Vonk (1991) remains. Despite intensive field work, no additional populations have been found and this species appears to be extinct in the wild.</p><p>Remarks.  Tellia is clearly diagnosed by absence of the pelvic fin as well as by a unique nuptial male colour pattern. Hoedeman (1951) synonymised  Tellia with  Aphanius, because absence of pelvic fins seems not to be enough for an own genus. We support this view, since the pelvic fin is occasionally absent in individuals, but never entire populations, belonging to other genera, particularly  Apricaphanius and  Anatolichthys . In addition, molecular data presented by Hrbek et al. (2002) and Geiger et al. (2014) as well as the unique colour pattern of male  Tellia strongly support its recognition as a distinct genus.</p><p>Hrbek et al. (2002) and Esmaeili et al. (2020) discussed the phylogenetic position of  Tellia and suggested it to be the sister of all Aphaniid genera except  Aphaniops and Paraphanius. However, Hrbek et al. (2002) and Geiger et al. (2014) alternatively recovered  Tellia as the sister taxon to all other Aphaniid species. There were no statistical differences between these contrasting placements (Hrbek et al. 2002), therefore further research is required to resolve the phylogenetic position of  Tellia within  Aphaniidae .</p><p>Material examined.</p><p>Tellia apoda: FSJF 3462, 21, 26–44 mm SL; Algeria: stream south of Ain M’Lila, east of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=6.5667&amp;materialsCitation.latitude=36.0" title="Search Plazi for locations around (long 6.5667/lat 36.0)">Fourchi</a>, 36.0000 6.5667.</p></div>	https://treatment.plazi.org/id/03B187D4DF0EFF86FF4F6191FF32DE49	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Freyhof, Jörg;Yoğurtçuoğlu, Baran	Freyhof, Jörg, Yoğurtçuoğlu, Baran (2020): A proposal for a new generic structure of the killifish family Aphaniidae, with the description of Aphaniops teimorii (Teleostei: Cyprinodontiformes). Zootaxa 4810 (3): 421-451, DOI: 10.11646/zootaxa.4810.3.2
03B187D4DF08FF80FF4F60D5FE6DDF66.text	03B187D4DF08FF80FF4F60D5FE6DDF66.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apricaphanius Freyhof & Yoğurtçuoğlu 2020	<div><p>Apricaphanius,  new genus</p><p>Fig. 15</p><p>Type species.  Lebias iberus Valenciennes in Cuvier &amp; Valenciennes, 1846 .</p><p>Diagnosis.  Apricaphanius is superficially similar to  Anatolichthys, from which it is distinguished by having a narrow, black dorsal-fin margin (vs. wide black margin in  Anatolichthys) followed by a narrow or wide white or hyaline sub-marginal band in nuptial males. Nuptial male  Anatolichthys never have a white sub-marginal band in the dorsal fin. In the male  Anatolichthys the dorsal fin is completely black or it has a wide greyish or black margin, followed by a white proximal band ( A. iconii) or by a proximal row of white spots or blotches distinctly or slightly above the dorsal-fin base, or by a hyaline field with single or rows of brown spots. Furthermore, male  Apricaphanius are diagnosed by the absence of flank bars (in  A. saourensis) or possession of 14–22 irregular, usually vertically split, flank bars, overlaid with numerous minute whitish spots in the adult male (vs. 5–13 regular flank bars, except  A. villwockii, and not overlaid by spots in all species except  A. irregularis).  Apricaphanius is distinguished from the remaining genera in the family  Aphaniidae by the following combination of non-unique characters: head canals absent (vs. present in  Aphanius and  Aphaniops); dermal sheath at the anal-fin base in the nuptial female present (vs. absent in  Aphaniops); possession of a single row of tricuspid teeth (vs. three rows of conical teeth in  Kosswigichthys); body covered by scales (vs. naked in  Kosswigichthys); pelvic fin present (vs. absent in  Tellia); black or dark-brown bars in the caudal fin of the male present, rarely absent in  A. saourensis (vs. absent in Paraphanius); flank pattern in the male comprising a series of black or brown bars or intricate silvery vermiculation (vs. small whitish or blue sports arranged in vertical series or very narrow bars in Paraphanius); a bold, black spot at the centre of the caudal-fin base in the female present (vs. absent in Paraphanius); dorsal- and anal-fin margins in the male black (vs. without black margins in  Aphaniops, yellow in  Tellia, only dorsal-fin margin black in  Aphanius); background colour of caudal fin identical to interspaces between flank bars (vs. caudal fin pale or deep yellow or orange, distinct from silvery interspaces between flank bars in  Aphanius).</p><p>Included species.  Apricaphanius iberus,  A. baeticus,  A. saourensis .</p><p>Distribution.  Apricaphanius species are distributed along the coastline of the Iberian Peninsula from the southern Atlantic slope of Spain (lower Guadalquivir region) to Catalonia, plus the Oued Saoura basin in northwestern Algeria (Blanco et al. 2006; Doadrio et al. 2002, Gonzales et al. 2014) (Fig. 14). Other records from Algeria represented in Figure 14 are based on historic records of fishes identified as  A. iberus by Pellegrin (1921) and may have been populations of  A. saourensis, but none have been confirmed since.</p><p>Etymology. Based on the Latin substantive Apricus, shining, for the many small white spots on the flanks of the male, which give them a shiny appearance. Gender masculine.</p><p>Remarks. The molecular phylogeny presented by Hrbek et al. (2002) placed  Apricaphanius as sister genus to all Aphaniids, except  Aphaniops, Paraphanius and  Tellia . The molecular tree published by Esmaeili et al. (2020) placed  Apricaphanius close to  Esmaeilius, while Geiger et al. (2014), placed  Apricaphanius as sister to  Aphanius and  Anatolichthys in their analysis of Mediterranean species (Fig. 1). The results of these studies clearly demonstrate that  Apricaphanius cannot be placed in  Anatolichthys, and that it represents a distinct, well-distinguished phylogenetic group. To avoid paraphyly,  Apricaphanius must be treated as a distinct genus despite its superficial similarity to  Anatolichthys .</p><p>Apricaphanius and  Anatolichthys species lack head canals and possess pelvic fins, a dermal sheath at the analfin base in the nuptial female, and a single row of tricuspid teeth.</p><p>Male  Apricaphanius and  Anatolichthys possess a black dorsal-fin margin, which is narrow in  Apricaphanius (vs. wide, often the entire dorsal fin is black in  Anatolichthys). Male  Apricaphanius are well distinguished from all  Anatolichthys by presence of a white or hyaline sub-marginal band in the dorsal fin often with some isolated brown spots or blotches (vs. absence in  Anatolichthys). Male  Apricaphanius either lack flank bars ( A. saourensis) or possess 12–22 narrow, irregular set and shaped flank bars which are usually split vertically and overlaid with numerous minute whitish spots in adults (vs. 5–13 regularly set and shaped bars, not overlaid by spots in all but one  Anatolichthys species). Male  Anatolichthys villwocki possess 13–25 narrow brown flank bars, but also 4–14 vertical rows of small black or brown spots in the caudal fin (vs. 1–4 bold black bars in  Apricaphanius species).</p><p>The flank bars are not very easy to see in male  Apricaphanius, since the flank bars and dorsal-fin colour pattern often dissociate in adult individuals larger than 25 mm SL. The flank pattern in the adult male  Anatolichthys irregularis also comprises numerous small white spots (Yoğurtçuoğlu &amp; Freyhof 2018) and this is also the case in some species of Paraphanius such as  P. alexandri and  P. mentoides . We suspect the presence of small white spots on the flank has evolved at least four times in  Aphaniidae, and its occurrence in  Apricaphanius,  Anatolichthys irregularis, Paraphanius and  Tellia is the result of convergent evolution.</p><p>Material examined.</p><p>Apricaphanius baeticus: FSJF 4015, 1, 28 mm SL; Spain: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-5.994935&amp;materialsCitation.latitude=36.95773" title="Search Plazi for locations around (long -5.994935/lat 36.95773)">Arroyo del Moscardo</a> between Lebrija and Las Cabezas de San Juan, 36.957729 -5.994935.   — FSJF 4102, 25, 20–33 mm SL; Spain: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-4.8825&amp;materialsCitation.latitude=37.4176" title="Search Plazi for locations around (long -4.8825/lat 37.4176)">Arroyo Salado</a>, 37.4176 - 4.8825.   — ZMH 9281, 10, 23–28 mm SL; Spain:  Arroyo de Mascardo, Lebrija.</p><p>Apricaphanius iberus: ZFMK-ICH 53133, 1, 23 mm SL; Spain: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-0.8297&amp;materialsCitation.latitude=37.6795" title="Search Plazi for locations around (long -0.8297/lat 37.6795)">Mar Menor</a> at Estrella del Mar, 37.6795 - 0.8297.   — ZFMK-ICH 59739-59740, 2, 23 – 28 mm SL; Spain: canals at <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=3.1122&amp;materialsCitation.latitude=42.2526" title="Search Plazi for locations around (long 3.1122/lat 42.2526)">Empuriabrava</a>, 42.2526 3.1122.   — ZFMK-ICH 59745, 1, 28 mm SL; Spain: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=3.1926&amp;materialsCitation.latitude=42.0471" title="Search Plazi for locations around (long 3.1926/lat 42.0471)">Estartit</a>, 42.0471 3.1926.   — ZFMK-ICH 59746, 59747, 2, 20 – 33 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=0.9312&amp;materialsCitation.latitude=40.9922" title="Search Plazi for locations around (long 0.9312/lat 40.9922)">Riu de Llastres</a> at Miami Platja, 40.9922 0.9312.   — ZMH 4372, 10, 21–24 mm SL; Spain: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-0.6074&amp;materialsCitation.latitude=38.1857" title="Search Plazi for locations around (long -0.6074/lat 38.1857)">Saltworks at Santa Pola</a>, (about 38.1857 -0.6074).</p><p>Apricaphanius saourensis: FSJF 3635, 10, 22–36 mm SL; Algeria: Mazzer prov.: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-2.2685&amp;materialsCitation.latitude=30.3168" title="Search Plazi for locations around (long -2.2685/lat 30.3168)">La tombe de Moché</a>, Oued Saoura, 30.3168 -2.2685.</p></div>	https://treatment.plazi.org/id/03B187D4DF08FF80FF4F60D5FE6DDF66	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Freyhof, Jörg;Yoğurtçuoğlu, Baran	Freyhof, Jörg, Yoğurtçuoğlu, Baran (2020): A proposal for a new generic structure of the killifish family Aphaniidae, with the description of Aphaniops teimorii (Teleostei: Cyprinodontiformes). Zootaxa 4810 (3): 421-451, DOI: 10.11646/zootaxa.4810.3.2
03B187D4DF09FF8DFF4F6685FD9ADED8.text	03B187D4DF09FF8DFF4F6685FD9ADED8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Esmaeilius Freyhof & Yoğurtçuoğlu 2020	<div><p>Esmaeilius,  new genus</p><p>Fig. 16–17</p><p>Type species.  Lebias sophiae Heckel, 1847 .</p><p>Diagnosis.  Esmaeilius is distinguished from  Anatolichthys and  Apricaphanius by possession of a hyaline caudal fin, with a white margin in some species, without bars or rows of spots, or with 1–5 indistinct vertical rows of small brown spots usually restricted to the proximal portion in  E. sophiae, (vs. caudal fin with 1–4 bold black or brown bars, or 4–14 vertical rows of small brown spots in  Anatolichthys villwocki), plus a white dorsal- and often caudaland anal-fin margins in the male, except  E. isfahanensis which possesses black dorsal- and anal-fin margins (vs. dorsal and anal-fin margins black, caudal-fin margin hyaline).</p><p>Esmaeilius is distinguished from other genera in the family  Aphaniidae by the following combination of nonunique characters: head canals absent (vs. present in  Aphanius and  Aphaniops); dermal sheath at the anal-fin base in the nuptial female present (vs. absent in  Aphaniops); a single row of tricuspid teeth (vs. three rows of conical teeth in  Kosswigichthys); body covered by scales (vs. naked in  Kosswigichthys); pelvic fin present (vs. absent in  Tellia); flank pattern in male comprising a series of black or brown bars (vs. small whitish or blue spots arranged in vertical series or very narrow bars in Paraphanius); a bold black spot at centre of the caudal-fin base in female present (vs. absent in Paraphanius); dorsal- and anal-fin margin white or black in male (vs. without white or black margins in all  Aphaniops except  A. sirhani, yellow in  Tellia, only dorsal-fin margin black in  Aphanius); background colour of caudal fin identical to interspaces between flank bars (vs. caudal fin pale or deep yellow or orange, distinct from silvery interspaces between flank bars in  Aphanius).</p><p>Included species.  Esmaeilius sophiae,  E. darabensis,  E. isfahanensis,  E. persicus,  E. shirini,  E. vladykovi .</p><p>Distribution.  Esmaeilius species are widespread in inland waters of Iran and the lower Shat-al-Arab River drainage in Iraq (Fig. 18). Esmaeili et al. (2020) provided a detailed map showing the distribution of the genus.</p><p>Etymology. Named for Hamid Reza Esmaeili (Shiraz) for his extensive contribution to the understanding of diversity within this genus. Gender masculine.</p><p>Remarks.  Esmaeilius is distinguished from  Anatolichthys and  Apricaphanius by colour pattern, specifically the absence of bars in the caudal fin (vs. presence of bars in all species except  A. villwocki) and presence of white (vs. black) dorsal- and usually caudal- and anal-fin margins in all species except  E. isfahanensis . This combination of characters never occurs in  Anatolichthys or  Apricaphanius species.</p><p>Anatolichthys villwocki is similar to  E. sophiae in possessing rows of small brown spots in the caudal fin but it differs by possessing black (vs. white) dorsal- and anal-fin margins.  Esmaeilius isfahanensis is similar to  Anatolichthys and  Apricaphanius species in possessing black dorsal- and anal-fin margins but it differs by lacking bold black bars in the caudal fin.</p><p>Brachylebias persicus Priem, 1908 was transferred to  Aphanius by Gaudant (2011), becoming a senior secondary homonym of  Cyprinodon persicus Jenkins, 1910 (=  Aphanius persicus), and  Aphanius farsicus was proposed as the replacement name by Teimori et al. (2011). When the phylogenetic groups of  Aphanius are separated into different genera,  Brachylebias must be considered as incertae sedis as we find no arguments to place it in one of the genera recognised here and it may represent a distinct (extinct) evolutionary lineage.  Brachylebias,  Aphaniops and  Aphanius share the presence of a preopercular canal (Gaudant (2011), and therefore  B. persicus might have been a Miocene species of  Aphaniops or  Aphanius (see Discussion below). While there is a need to re-examine the materials of  Brachylebias, we currently find it highly unlikely that  Brachylebias might have been a species of  Esmaeilius .</p><p>Brachylebias persicus and  Cyprinodon persicus became secondary homonyms when both were placed in  Aphanius . Because the two names are no longer congeneric, and because the replacement name was raised after 1960, the older name  Cyprinodon persicus Jenkins must be reinstated as the valid name (ICZN 1999: Art. 59.4). This makes  Aphanius farsicus a junior synonym of  Cyprinodon persicus .</p><p>The molecular phylogeny and results of four different molecular species delimitation methods published by Esmaeili et al. (2020) strongly suggest that an excessive number of  Esmaeilius populations have been recognised as species.  Esmaeilius darabensis,  E. persicus,  E. isfahanensis,  E. shirini, and  E. vladykovi were supported by all four methods, but  E. arakensis is only supported by two methods while  E. pluristriatus is paraphyletic with only a single population supported by one method. Teimori et al. (2012), Esmaeili et al. (2012), Gholami et al. (2014), and Esmaeili et al. (2014a, b) find it difficult to clearly distinguish  E. arakensis,  E. kavirensis,  E. mesopotamicus, and  E. pluristriatus from  E. sophiae by morphology and used a series of vague or overlapping character states, while there is little molecular distance between them. Gholami et al. (2013) recognised that geographic isolation of these species might have occurred in the Holocene, thus explaining the minor molecular differences. We recognise all four of these nominal species as populations of  E. sophiae because they have largely been diagnosed by otolith shape, a character likely to vary between individual populations in the family  Aphaniidae (Schulz-Mirbach et al. 2006, Reichenbacher et al. 2009 a, Annabi et al. 2013). The same might be true for scale surface microstructure as suggested by Gholami et al. (2013). We see no reason to treat  E. arakensis,  E. kavirensis,  E. mesopotamicus, and  E. pluristriatus as distinct species and therefore, we synonymise them with  E. sophiae .</p><p>It must be noted that we fully support distinguishing species with little or no molecular separation, provided they can be clearly distinguished by non-overlapping morphological characters, including colour pattern, and by patterns in their independent evolutionary histories. The same can be said for morphologically-congruous populations separated by deep molecular differences. Such ‘cryptic’ species are uncommon, but  E. darabensis appears to be a genuine example. Esmaeili et al. (2014 b, 2020) suggested that it is most closely related to  E. persicus, from which it is well distinguished by colour pattern, while it is morphologically identical to  E. sophiae .</p><p>Material examined.</p><p>Esmaeilius persicus: FSJF 2229, 5, 30–37 mm SL;   FSJF 4017, 18, 25–33 mm SL; Iran: Fars prov.: spring Pirbanoo about 10 km south of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=52.4656&amp;materialsCitation.latitude=29.5189" title="Search Plazi for locations around (long 52.4656/lat 29.5189)">Shiraz</a>, 29.5189 52.4656.</p><p>Esmaeilius isfahanensis: FSJF 3497, 13, 31–38 mm SL; Iran: Esfahan prov.: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=52.6538&amp;materialsCitation.latitude=32.4255" title="Search Plazi for locations around (long 52.6538/lat 32.4255)">Zayandeh River at Varzaneh</a> bridge, 32.4255 52.6538.</p><p>Esmaeilius sophiae: FSJF 2225, 44, 16–42 mm SL; Iran: Fars prov.: spring Ghadamgah near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=52.4246&amp;materialsCitation.latitude=30.2548" title="Search Plazi for locations around (long 52.4246/lat 30.2548)">Dorodzan Dam</a>, about 88 km north of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=52.4246&amp;materialsCitation.latitude=30.2548" title="Search Plazi for locations around (long 52.4246/lat 30.2548)">Shiraz</a>, 30.2548 52.4246.  — FSJF 3217, 44, 23–42 mm SL; FSJF 3245, 37, 19–34 mm SL; FSJF 3251, 23, 20–35 mm SL; Iran: Shiraz prov.: spring Malous, 10 km of Shiraz city, 29.8770 52.4806.  — FSJF 4064, 55, 17–42 mm SL; Iran: spring <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=54.0838&amp;materialsCitation.latitude=36.2793" title="Search Plazi for locations around (long 54.0838/lat 36.2793)">Cheshme Ali</a>, 36.2793 54.0838.   — FSJF 4103, 6, 30–33 mm SL; Iran:  Talabe-Gavkhuni .</p><p>Esmaeilius vladykovi: FSJF 4039, 4, 22–32 mm SL; Iran: Shahrestan-e Bakhtiari va <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=50.9047&amp;materialsCitation.latitude=31.9256" title="Search Plazi for locations around (long 50.9047/lat 31.9256)">Chahar Mahall</a>, 3 kilometers west of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=50.9047&amp;materialsCitation.latitude=31.9256" title="Search Plazi for locations around (long 50.9047/lat 31.9256)">Boldaji</a>, 31.9256 50.9047.</p></div>	https://treatment.plazi.org/id/03B187D4DF09FF8DFF4F6685FD9ADED8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Freyhof, Jörg;Yoğurtçuoğlu, Baran	Freyhof, Jörg, Yoğurtçuoğlu, Baran (2020): A proposal for a new generic structure of the killifish family Aphaniidae, with the description of Aphaniops teimorii (Teleostei: Cyprinodontiformes). Zootaxa 4810 (3): 421-451, DOI: 10.11646/zootaxa.4810.3.2
