taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03B187DCFFC7FFC4FF398604FD41FEAD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17889771/files/figure.png	https://doi.org/10.5281/zenodo.17889771	FIGURE 11. Distribution of pollen wasps in the Caucasus. A, Celonites cagrii Mauss & Yildirim, 2022; B, C. carpenteri Fateryga & Mauss, sp. nov.; C, C. ivanovi Mauss & Fateryga, 2022; D, C. laetus Panfilov, 1968; E, C. osseus Morawitz, 1888; F, C. tauricus Kostylev, 1935; G, C. tudesculptus Kostylev, 1935; H, Ceramius caucasicus André, 1884; I, Jugurtia eurycara Kostylev, 1935; J, J. irana Kostylev, 1935; K, Quartinia aliyevi Fateryga, Mauss, Proshchalykin & Maharramov, sp. nov.; L, Q. araxana Giordani Soika, 1960.	FIGURE 11. Distribution of pollen wasps in the Caucasus. A, Celonites cagrii Mauss & Yildirim, 2022; B, C. carpenteri Fateryga & Mauss, sp. nov.; C, C. ivanovi Mauss & Fateryga, 2022; D, C. laetus Panfilov, 1968; E, C. osseus Morawitz, 1888; F, C. tauricus Kostylev, 1935; G, C. tudesculptus Kostylev, 1935; H, Ceramius caucasicus André, 1884; I, Jugurtia eurycara Kostylev, 1935; J, J. irana Kostylev, 1935; K, Quartinia aliyevi Fateryga, Mauss, Proshchalykin & Maharramov, sp. nov.; L, Q. araxana Giordani Soika, 1960.	2025-11-03	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.		Zenodo	biologists	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.			
03B187DCFFC4FFC0FF398080FE4AFE41.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17889741/files/figure.png	https://doi.org/10.5281/zenodo.17889741	FIGURE 1. Celonites carpenteri Fateryga & Mauss, sp. nov. A–C, E, ♀, holotype; D, F–H, ♀, paratype.A, head and mesosoma in dorsal view; B, habitus in lateral view; C, labels; D, posterior part of mesosoma and metasoma in dorsal view; E, head in frontal view; F, apex of metasoma in dorsal view; G, emargination between lateral lamella and postero-lateral process of propodeum; H, apex of metasoma in ventral view. Scale bars 1 mm.	FIGURE 1. Celonites carpenteri Fateryga & Mauss, sp. nov. A–C, E, ♀, holotype; D, F–H, ♀, paratype.A, head and mesosoma in dorsal view; B, habitus in lateral view; C, labels; D, posterior part of mesosoma and metasoma in dorsal view; E, head in frontal view; F, apex of metasoma in dorsal view; G, emargination between lateral lamella and postero-lateral process of propodeum; H, apex of metasoma in ventral view. Scale bars 1 mm.	2025-11-03	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.		Zenodo	biologists	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.			
03B187DCFFC4FFC0FF398080FE4AFE41.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17889771/files/figure.png	https://doi.org/10.5281/zenodo.17889771	FIGURE 11. Distribution of pollen wasps in the Caucasus. A, Celonites cagrii Mauss & Yildirim, 2022; B, C. carpenteri Fateryga & Mauss, sp. nov.; C, C. ivanovi Mauss & Fateryga, 2022; D, C. laetus Panfilov, 1968; E, C. osseus Morawitz, 1888; F, C. tauricus Kostylev, 1935; G, C. tudesculptus Kostylev, 1935; H, Ceramius caucasicus André, 1884; I, Jugurtia eurycara Kostylev, 1935; J, J. irana Kostylev, 1935; K, Quartinia aliyevi Fateryga, Mauss, Proshchalykin & Maharramov, sp. nov.; L, Q. araxana Giordani Soika, 1960.	FIGURE 11. Distribution of pollen wasps in the Caucasus. A, Celonites cagrii Mauss & Yildirim, 2022; B, C. carpenteri Fateryga & Mauss, sp. nov.; C, C. ivanovi Mauss & Fateryga, 2022; D, C. laetus Panfilov, 1968; E, C. osseus Morawitz, 1888; F, C. tauricus Kostylev, 1935; G, C. tudesculptus Kostylev, 1935; H, Ceramius caucasicus André, 1884; I, Jugurtia eurycara Kostylev, 1935; J, J. irana Kostylev, 1935; K, Quartinia aliyevi Fateryga, Mauss, Proshchalykin & Maharramov, sp. nov.; L, Q. araxana Giordani Soika, 1960.	2025-11-03	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.		Zenodo	biologists	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.			
03B187DCFFC4FFC0FF398080FE4AFE41.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17889743/files/figure.png	https://doi.org/10.5281/zenodo.17889743	FIGURE 2. Habitats of Celonites carpenteri Fateryga & Mauss, sp. nov. (A) and Quartinia aliyevi Fateryga, Mauss, Proshchalykin & Maharramov, sp. nov. (B) in Azerbaijan.	FIGURE 2. Habitats of Celonites carpenteri Fateryga & Mauss, sp. nov. (A) and Quartinia aliyevi Fateryga, Mauss, Proshchalykin & Maharramov, sp. nov. (B) in Azerbaijan.	2025-11-03	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.		Zenodo	biologists	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.			
03B187DCFFC0FFCEFF3980ECFF32FD39.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17889751/files/figure.png	https://doi.org/10.5281/zenodo.17889751	FIGURE 4. Celonites ivanovi Mauss & Fateryga, 2022. A, female standing on a small stone; B, female at flower of Heliotropium styligerum Trautv.	FIGURE 4. Celonites ivanovi Mauss & Fateryga, 2022. A, female standing on a small stone; B, female at flower of Heliotropium styligerum Trautv.	2025-11-03	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.		Zenodo	biologists	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.			
03B187DCFFC0FFCEFF3980ECFF32FD39.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17889769/files/figure.png	https://doi.org/10.5281/zenodo.17889769	FIGURE 10. Characters of pollen wasps. A, I, Celonites tauricus Kostylev, 1935, ♀, Azerbaijan; B, C. ivanovi Mauss & Fateryga, 2022, ♀, Dagestan; C, J, C. tudesculptus Kostylev, 1935, ♀, Azerbaijan; D, H, C. laetus Panfilov, 1968, ♀, holotype; E, C. osseus Morawitz, 1888, ♀, Armenia; F, C. tauricus, ♂, Dagestan; G, C. osseus, ♂, Iran; K, C. ivanovi, ♂, paratype; L, Jugurtia eurycara Kostylev, 1935, ♀, Azerbaijan; M, J. irana Kostylev, 1935, ♀, Azerbaijan. A–C, emargination between lateral lamella and postero-lateral process of propodeum; D, E, tegula; F, G, apex of metasoma in dorsal view; H, metasomal terga II–VI; I–K, metasoma in dorsal view; L, M, metasomal terga II–IV. Scale bars 1 mm.	FIGURE 10. Characters of pollen wasps. A, I, Celonites tauricus Kostylev, 1935, ♀, Azerbaijan; B, C. ivanovi Mauss & Fateryga, 2022, ♀, Dagestan; C, J, C. tudesculptus Kostylev, 1935, ♀, Azerbaijan; D, H, C. laetus Panfilov, 1968, ♀, holotype; E, C. osseus Morawitz, 1888, ♀, Armenia; F, C. tauricus, ♂, Dagestan; G, C. osseus, ♂, Iran; K, C. ivanovi, ♂, paratype; L, Jugurtia eurycara Kostylev, 1935, ♀, Azerbaijan; M, J. irana Kostylev, 1935, ♀, Azerbaijan. A–C, emargination between lateral lamella and postero-lateral process of propodeum; D, E, tegula; F, G, apex of metasoma in dorsal view; H, metasomal terga II–VI; I–K, metasoma in dorsal view; L, M, metasomal terga II–IV. Scale bars 1 mm.	2025-11-03	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.		Zenodo	biologists	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.			
03B187DCFFC0FFCEFF3980ECFF32FD39.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17889771/files/figure.png	https://doi.org/10.5281/zenodo.17889771	FIGURE 11. Distribution of pollen wasps in the Caucasus. A, Celonites cagrii Mauss & Yildirim, 2022; B, C. carpenteri Fateryga & Mauss, sp. nov.; C, C. ivanovi Mauss & Fateryga, 2022; D, C. laetus Panfilov, 1968; E, C. osseus Morawitz, 1888; F, C. tauricus Kostylev, 1935; G, C. tudesculptus Kostylev, 1935; H, Ceramius caucasicus André, 1884; I, Jugurtia eurycara Kostylev, 1935; J, J. irana Kostylev, 1935; K, Quartinia aliyevi Fateryga, Mauss, Proshchalykin & Maharramov, sp. nov.; L, Q. araxana Giordani Soika, 1960.	FIGURE 11. Distribution of pollen wasps in the Caucasus. A, Celonites cagrii Mauss & Yildirim, 2022; B, C. carpenteri Fateryga & Mauss, sp. nov.; C, C. ivanovi Mauss & Fateryga, 2022; D, C. laetus Panfilov, 1968; E, C. osseus Morawitz, 1888; F, C. tauricus Kostylev, 1935; G, C. tudesculptus Kostylev, 1935; H, Ceramius caucasicus André, 1884; I, Jugurtia eurycara Kostylev, 1935; J, J. irana Kostylev, 1935; K, Quartinia aliyevi Fateryga, Mauss, Proshchalykin & Maharramov, sp. nov.; L, Q. araxana Giordani Soika, 1960.	2025-11-03	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.		Zenodo	biologists	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.			
03B187DCFFC0FFCEFF3980ECFF32FD39.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17889749/files/figure.png	https://doi.org/10.5281/zenodo.17889749	FIGURE 3. Nests of Celonites ivanovi Mauss & Fateryga, 2022 in Dagestan. A, nesting site (arrow indicates the position of nest no. 1); B, position of nest no. 1 (indicated with an arrow); C, nest no. 1, close up; D, the same nest longitudinally dissected; E, nest no. 2; F, nest. no. 3; G, nest no. 4. Scale bars 1 cm.	FIGURE 3. Nests of Celonites ivanovi Mauss & Fateryga, 2022 in Dagestan. A, nesting site (arrow indicates the position of nest no. 1); B, position of nest no. 1 (indicated with an arrow); C, nest no. 1, close up; D, the same nest longitudinally dissected; E, nest no. 2; F, nest. no. 3; G, nest no. 4. Scale bars 1 cm.	2025-11-03	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.		Zenodo	biologists	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.			
03B187DCFFCEFFCEFF39820DFEC3FA62.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17889767/files/figure.png	https://doi.org/10.5281/zenodo.17889767	FIGURE 9. Characters of pollen wasps. A, D, Ceramius caucasicus André, 1884, ♂,Azerbaijan; B, Jugurtia eurycara Kostylev, 1935, ♀, Azerbaijan; C, J. irana Kostylev, 1935, ♀, Azerbaijan; E, J. eurycara, ♂,Azerbaijan; F, G, Celonites osseus Morawitz, 1888, ♀, Armenia; H, C. laetus Panfilov, 1968, ♀, holotype; I, C. tauricus Kostylev, 1935, ♀, Azerbaijan; J, C. tudesculptus Kostylev, 1935, ♀, Azerbaijan. A, clypeus; B, C, F, head in frontal view; D, E, antenna; G–J, head in lateral view. Scale bars 1 mm.	FIGURE 9. Characters of pollen wasps. A, D, Ceramius caucasicus André, 1884, ♂,Azerbaijan; B, Jugurtia eurycara Kostylev, 1935, ♀, Azerbaijan; C, J. irana Kostylev, 1935, ♀, Azerbaijan; E, J. eurycara, ♂,Azerbaijan; F, G, Celonites osseus Morawitz, 1888, ♀, Armenia; H, C. laetus Panfilov, 1968, ♀, holotype; I, C. tauricus Kostylev, 1935, ♀, Azerbaijan; J, C. tudesculptus Kostylev, 1935, ♀, Azerbaijan. A, clypeus; B, C, F, head in frontal view; D, E, antenna; G–J, head in lateral view. Scale bars 1 mm.	2025-11-03	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.		Zenodo	biologists	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.			
03B187DCFFCEFFCEFF39820DFEC3FA62.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17889769/files/figure.png	https://doi.org/10.5281/zenodo.17889769	FIGURE 10. Characters of pollen wasps. A, I, Celonites tauricus Kostylev, 1935, ♀, Azerbaijan; B, C. ivanovi Mauss & Fateryga, 2022, ♀, Dagestan; C, J, C. tudesculptus Kostylev, 1935, ♀, Azerbaijan; D, H, C. laetus Panfilov, 1968, ♀, holotype; E, C. osseus Morawitz, 1888, ♀, Armenia; F, C. tauricus, ♂, Dagestan; G, C. osseus, ♂, Iran; K, C. ivanovi, ♂, paratype; L, Jugurtia eurycara Kostylev, 1935, ♀, Azerbaijan; M, J. irana Kostylev, 1935, ♀, Azerbaijan. A–C, emargination between lateral lamella and postero-lateral process of propodeum; D, E, tegula; F, G, apex of metasoma in dorsal view; H, metasomal terga II–VI; I–K, metasoma in dorsal view; L, M, metasomal terga II–IV. Scale bars 1 mm.	FIGURE 10. Characters of pollen wasps. A, I, Celonites tauricus Kostylev, 1935, ♀, Azerbaijan; B, C. ivanovi Mauss & Fateryga, 2022, ♀, Dagestan; C, J, C. tudesculptus Kostylev, 1935, ♀, Azerbaijan; D, H, C. laetus Panfilov, 1968, ♀, holotype; E, C. osseus Morawitz, 1888, ♀, Armenia; F, C. tauricus, ♂, Dagestan; G, C. osseus, ♂, Iran; K, C. ivanovi, ♂, paratype; L, Jugurtia eurycara Kostylev, 1935, ♀, Azerbaijan; M, J. irana Kostylev, 1935, ♀, Azerbaijan. A–C, emargination between lateral lamella and postero-lateral process of propodeum; D, E, tegula; F, G, apex of metasoma in dorsal view; H, metasomal terga II–VI; I–K, metasoma in dorsal view; L, M, metasomal terga II–IV. Scale bars 1 mm.	2025-11-03	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.		Zenodo	biologists	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.			
03B187DCFFCEFFCEFF39820DFEC3FA62.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17889771/files/figure.png	https://doi.org/10.5281/zenodo.17889771	FIGURE 11. Distribution of pollen wasps in the Caucasus. A, Celonites cagrii Mauss & Yildirim, 2022; B, C. carpenteri Fateryga & Mauss, sp. nov.; C, C. ivanovi Mauss & Fateryga, 2022; D, C. laetus Panfilov, 1968; E, C. osseus Morawitz, 1888; F, C. tauricus Kostylev, 1935; G, C. tudesculptus Kostylev, 1935; H, Ceramius caucasicus André, 1884; I, Jugurtia eurycara Kostylev, 1935; J, J. irana Kostylev, 1935; K, Quartinia aliyevi Fateryga, Mauss, Proshchalykin & Maharramov, sp. nov.; L, Q. araxana Giordani Soika, 1960.	FIGURE 11. Distribution of pollen wasps in the Caucasus. A, Celonites cagrii Mauss & Yildirim, 2022; B, C. carpenteri Fateryga & Mauss, sp. nov.; C, C. ivanovi Mauss & Fateryga, 2022; D, C. laetus Panfilov, 1968; E, C. osseus Morawitz, 1888; F, C. tauricus Kostylev, 1935; G, C. tudesculptus Kostylev, 1935; H, Ceramius caucasicus André, 1884; I, Jugurtia eurycara Kostylev, 1935; J, J. irana Kostylev, 1935; K, Quartinia aliyevi Fateryga, Mauss, Proshchalykin & Maharramov, sp. nov.; L, Q. araxana Giordani Soika, 1960.	2025-11-03	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.		Zenodo	biologists	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.			
03B187DCFFCEFFCEFF3984CDFB5BF8BB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17889767/files/figure.png	https://doi.org/10.5281/zenodo.17889767	FIGURE 9. Characters of pollen wasps. A, D, Ceramius caucasicus André, 1884, ♂,Azerbaijan; B, Jugurtia eurycara Kostylev, 1935, ♀, Azerbaijan; C, J. irana Kostylev, 1935, ♀, Azerbaijan; E, J. eurycara, ♂,Azerbaijan; F, G, Celonites osseus Morawitz, 1888, ♀, Armenia; H, C. laetus Panfilov, 1968, ♀, holotype; I, C. tauricus Kostylev, 1935, ♀, Azerbaijan; J, C. tudesculptus Kostylev, 1935, ♀, Azerbaijan. A, clypeus; B, C, F, head in frontal view; D, E, antenna; G–J, head in lateral view. Scale bars 1 mm.	FIGURE 9. Characters of pollen wasps. A, D, Ceramius caucasicus André, 1884, ♂,Azerbaijan; B, Jugurtia eurycara Kostylev, 1935, ♀, Azerbaijan; C, J. irana Kostylev, 1935, ♀, Azerbaijan; E, J. eurycara, ♂,Azerbaijan; F, G, Celonites osseus Morawitz, 1888, ♀, Armenia; H, C. laetus Panfilov, 1968, ♀, holotype; I, C. tauricus Kostylev, 1935, ♀, Azerbaijan; J, C. tudesculptus Kostylev, 1935, ♀, Azerbaijan. A, clypeus; B, C, F, head in frontal view; D, E, antenna; G–J, head in lateral view. Scale bars 1 mm.	2025-11-03	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.		Zenodo	biologists	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.			
03B187DCFFCEFFCEFF3984CDFB5BF8BB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17889769/files/figure.png	https://doi.org/10.5281/zenodo.17889769	FIGURE 10. Characters of pollen wasps. A, I, Celonites tauricus Kostylev, 1935, ♀, Azerbaijan; B, C. ivanovi Mauss & Fateryga, 2022, ♀, Dagestan; C, J, C. tudesculptus Kostylev, 1935, ♀, Azerbaijan; D, H, C. laetus Panfilov, 1968, ♀, holotype; E, C. osseus Morawitz, 1888, ♀, Armenia; F, C. tauricus, ♂, Dagestan; G, C. osseus, ♂, Iran; K, C. ivanovi, ♂, paratype; L, Jugurtia eurycara Kostylev, 1935, ♀, Azerbaijan; M, J. irana Kostylev, 1935, ♀, Azerbaijan. A–C, emargination between lateral lamella and postero-lateral process of propodeum; D, E, tegula; F, G, apex of metasoma in dorsal view; H, metasomal terga II–VI; I–K, metasoma in dorsal view; L, M, metasomal terga II–IV. Scale bars 1 mm.	FIGURE 10. Characters of pollen wasps. A, I, Celonites tauricus Kostylev, 1935, ♀, Azerbaijan; B, C. ivanovi Mauss & Fateryga, 2022, ♀, Dagestan; C, J, C. tudesculptus Kostylev, 1935, ♀, Azerbaijan; D, H, C. laetus Panfilov, 1968, ♀, holotype; E, C. osseus Morawitz, 1888, ♀, Armenia; F, C. tauricus, ♂, Dagestan; G, C. osseus, ♂, Iran; K, C. ivanovi, ♂, paratype; L, Jugurtia eurycara Kostylev, 1935, ♀, Azerbaijan; M, J. irana Kostylev, 1935, ♀, Azerbaijan. A–C, emargination between lateral lamella and postero-lateral process of propodeum; D, E, tegula; F, G, apex of metasoma in dorsal view; H, metasomal terga II–VI; I–K, metasoma in dorsal view; L, M, metasomal terga II–IV. Scale bars 1 mm.	2025-11-03	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.		Zenodo	biologists	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.			
03B187DCFFCEFFCEFF3984CDFB5BF8BB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17889771/files/figure.png	https://doi.org/10.5281/zenodo.17889771	FIGURE 11. Distribution of pollen wasps in the Caucasus. A, Celonites cagrii Mauss & Yildirim, 2022; B, C. carpenteri Fateryga & Mauss, sp. nov.; C, C. ivanovi Mauss & Fateryga, 2022; D, C. laetus Panfilov, 1968; E, C. osseus Morawitz, 1888; F, C. tauricus Kostylev, 1935; G, C. tudesculptus Kostylev, 1935; H, Ceramius caucasicus André, 1884; I, Jugurtia eurycara Kostylev, 1935; J, J. irana Kostylev, 1935; K, Quartinia aliyevi Fateryga, Mauss, Proshchalykin & Maharramov, sp. nov.; L, Q. araxana Giordani Soika, 1960.	FIGURE 11. Distribution of pollen wasps in the Caucasus. A, Celonites cagrii Mauss & Yildirim, 2022; B, C. carpenteri Fateryga & Mauss, sp. nov.; C, C. ivanovi Mauss & Fateryga, 2022; D, C. laetus Panfilov, 1968; E, C. osseus Morawitz, 1888; F, C. tauricus Kostylev, 1935; G, C. tudesculptus Kostylev, 1935; H, Ceramius caucasicus André, 1884; I, Jugurtia eurycara Kostylev, 1935; J, J. irana Kostylev, 1935; K, Quartinia aliyevi Fateryga, Mauss, Proshchalykin & Maharramov, sp. nov.; L, Q. araxana Giordani Soika, 1960.	2025-11-03	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.		Zenodo	biologists	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.			
03B187DCFFCFFFCFFF3981A1FE4AF913.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17889767/files/figure.png	https://doi.org/10.5281/zenodo.17889767	FIGURE 9. Characters of pollen wasps. A, D, Ceramius caucasicus André, 1884, ♂,Azerbaijan; B, Jugurtia eurycara Kostylev, 1935, ♀, Azerbaijan; C, J. irana Kostylev, 1935, ♀, Azerbaijan; E, J. eurycara, ♂,Azerbaijan; F, G, Celonites osseus Morawitz, 1888, ♀, Armenia; H, C. laetus Panfilov, 1968, ♀, holotype; I, C. tauricus Kostylev, 1935, ♀, Azerbaijan; J, C. tudesculptus Kostylev, 1935, ♀, Azerbaijan. A, clypeus; B, C, F, head in frontal view; D, E, antenna; G–J, head in lateral view. Scale bars 1 mm.	FIGURE 9. Characters of pollen wasps. A, D, Ceramius caucasicus André, 1884, ♂,Azerbaijan; B, Jugurtia eurycara Kostylev, 1935, ♀, Azerbaijan; C, J. irana Kostylev, 1935, ♀, Azerbaijan; E, J. eurycara, ♂,Azerbaijan; F, G, Celonites osseus Morawitz, 1888, ♀, Armenia; H, C. laetus Panfilov, 1968, ♀, holotype; I, C. tauricus Kostylev, 1935, ♀, Azerbaijan; J, C. tudesculptus Kostylev, 1935, ♀, Azerbaijan. A, clypeus; B, C, F, head in frontal view; D, E, antenna; G–J, head in lateral view. Scale bars 1 mm.	2025-11-03	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.		Zenodo	biologists	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.			
03B187DCFFCFFFCFFF3981A1FE4AF913.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17889769/files/figure.png	https://doi.org/10.5281/zenodo.17889769	FIGURE 10. Characters of pollen wasps. A, I, Celonites tauricus Kostylev, 1935, ♀, Azerbaijan; B, C. ivanovi Mauss & Fateryga, 2022, ♀, Dagestan; C, J, C. tudesculptus Kostylev, 1935, ♀, Azerbaijan; D, H, C. laetus Panfilov, 1968, ♀, holotype; E, C. osseus Morawitz, 1888, ♀, Armenia; F, C. tauricus, ♂, Dagestan; G, C. osseus, ♂, Iran; K, C. ivanovi, ♂, paratype; L, Jugurtia eurycara Kostylev, 1935, ♀, Azerbaijan; M, J. irana Kostylev, 1935, ♀, Azerbaijan. A–C, emargination between lateral lamella and postero-lateral process of propodeum; D, E, tegula; F, G, apex of metasoma in dorsal view; H, metasomal terga II–VI; I–K, metasoma in dorsal view; L, M, metasomal terga II–IV. Scale bars 1 mm.	FIGURE 10. Characters of pollen wasps. A, I, Celonites tauricus Kostylev, 1935, ♀, Azerbaijan; B, C. ivanovi Mauss & Fateryga, 2022, ♀, Dagestan; C, J, C. tudesculptus Kostylev, 1935, ♀, Azerbaijan; D, H, C. laetus Panfilov, 1968, ♀, holotype; E, C. osseus Morawitz, 1888, ♀, Armenia; F, C. tauricus, ♂, Dagestan; G, C. osseus, ♂, Iran; K, C. ivanovi, ♂, paratype; L, Jugurtia eurycara Kostylev, 1935, ♀, Azerbaijan; M, J. irana Kostylev, 1935, ♀, Azerbaijan. A–C, emargination between lateral lamella and postero-lateral process of propodeum; D, E, tegula; F, G, apex of metasoma in dorsal view; H, metasomal terga II–VI; I–K, metasoma in dorsal view; L, M, metasomal terga II–IV. Scale bars 1 mm.	2025-11-03	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.		Zenodo	biologists	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.			
03B187DCFFCFFFCFFF3981A1FE4AF913.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17889771/files/figure.png	https://doi.org/10.5281/zenodo.17889771	FIGURE 11. Distribution of pollen wasps in the Caucasus. A, Celonites cagrii Mauss & Yildirim, 2022; B, C. carpenteri Fateryga & Mauss, sp. nov.; C, C. ivanovi Mauss & Fateryga, 2022; D, C. laetus Panfilov, 1968; E, C. osseus Morawitz, 1888; F, C. tauricus Kostylev, 1935; G, C. tudesculptus Kostylev, 1935; H, Ceramius caucasicus André, 1884; I, Jugurtia eurycara Kostylev, 1935; J, J. irana Kostylev, 1935; K, Quartinia aliyevi Fateryga, Mauss, Proshchalykin & Maharramov, sp. nov.; L, Q. araxana Giordani Soika, 1960.	FIGURE 11. Distribution of pollen wasps in the Caucasus. A, Celonites cagrii Mauss & Yildirim, 2022; B, C. carpenteri Fateryga & Mauss, sp. nov.; C, C. ivanovi Mauss & Fateryga, 2022; D, C. laetus Panfilov, 1968; E, C. osseus Morawitz, 1888; F, C. tauricus Kostylev, 1935; G, C. tudesculptus Kostylev, 1935; H, Ceramius caucasicus André, 1884; I, Jugurtia eurycara Kostylev, 1935; J, J. irana Kostylev, 1935; K, Quartinia aliyevi Fateryga, Mauss, Proshchalykin & Maharramov, sp. nov.; L, Q. araxana Giordani Soika, 1960.	2025-11-03	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.		Zenodo	biologists	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.			
03B187DCFFCFFFCAFF39861AFD51FD4D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17889755/files/figure.png	https://doi.org/10.5281/zenodo.17889755	FIGURE 5. Celonites tudesculptus Kostylev, 1935, ♂, Azerbaijan. A, habitus in dorsal view; B, habitus in lateral view; C, metasomal terga II–VII; D, genitalia in dorsal view and metasomal sternum VII+VIII in ventral view; E, head in frontal view. Scale bars 1 mm.	FIGURE 5. Celonites tudesculptus Kostylev, 1935, ♂, Azerbaijan. A, habitus in dorsal view; B, habitus in lateral view; C, metasomal terga II–VII; D, genitalia in dorsal view and metasomal sternum VII+VIII in ventral view; E, head in frontal view. Scale bars 1 mm.	2025-11-03	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.		Zenodo	biologists	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.			
03B187DCFFCFFFCAFF39861AFD51FD4D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17889767/files/figure.png	https://doi.org/10.5281/zenodo.17889767	FIGURE 9. Characters of pollen wasps. A, D, Ceramius caucasicus André, 1884, ♂,Azerbaijan; B, Jugurtia eurycara Kostylev, 1935, ♀, Azerbaijan; C, J. irana Kostylev, 1935, ♀, Azerbaijan; E, J. eurycara, ♂,Azerbaijan; F, G, Celonites osseus Morawitz, 1888, ♀, Armenia; H, C. laetus Panfilov, 1968, ♀, holotype; I, C. tauricus Kostylev, 1935, ♀, Azerbaijan; J, C. tudesculptus Kostylev, 1935, ♀, Azerbaijan. A, clypeus; B, C, F, head in frontal view; D, E, antenna; G–J, head in lateral view. Scale bars 1 mm.	FIGURE 9. Characters of pollen wasps. A, D, Ceramius caucasicus André, 1884, ♂,Azerbaijan; B, Jugurtia eurycara Kostylev, 1935, ♀, Azerbaijan; C, J. irana Kostylev, 1935, ♀, Azerbaijan; E, J. eurycara, ♂,Azerbaijan; F, G, Celonites osseus Morawitz, 1888, ♀, Armenia; H, C. laetus Panfilov, 1968, ♀, holotype; I, C. tauricus Kostylev, 1935, ♀, Azerbaijan; J, C. tudesculptus Kostylev, 1935, ♀, Azerbaijan. A, clypeus; B, C, F, head in frontal view; D, E, antenna; G–J, head in lateral view. Scale bars 1 mm.	2025-11-03	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.		Zenodo	biologists	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.			
03B187DCFFCFFFCAFF39861AFD51FD4D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17889769/files/figure.png	https://doi.org/10.5281/zenodo.17889769	FIGURE 10. Characters of pollen wasps. A, I, Celonites tauricus Kostylev, 1935, ♀, Azerbaijan; B, C. ivanovi Mauss & Fateryga, 2022, ♀, Dagestan; C, J, C. tudesculptus Kostylev, 1935, ♀, Azerbaijan; D, H, C. laetus Panfilov, 1968, ♀, holotype; E, C. osseus Morawitz, 1888, ♀, Armenia; F, C. tauricus, ♂, Dagestan; G, C. osseus, ♂, Iran; K, C. ivanovi, ♂, paratype; L, Jugurtia eurycara Kostylev, 1935, ♀, Azerbaijan; M, J. irana Kostylev, 1935, ♀, Azerbaijan. A–C, emargination between lateral lamella and postero-lateral process of propodeum; D, E, tegula; F, G, apex of metasoma in dorsal view; H, metasomal terga II–VI; I–K, metasoma in dorsal view; L, M, metasomal terga II–IV. Scale bars 1 mm.	FIGURE 10. Characters of pollen wasps. A, I, Celonites tauricus Kostylev, 1935, ♀, Azerbaijan; B, C. ivanovi Mauss & Fateryga, 2022, ♀, Dagestan; C, J, C. tudesculptus Kostylev, 1935, ♀, Azerbaijan; D, H, C. laetus Panfilov, 1968, ♀, holotype; E, C. osseus Morawitz, 1888, ♀, Armenia; F, C. tauricus, ♂, Dagestan; G, C. osseus, ♂, Iran; K, C. ivanovi, ♂, paratype; L, Jugurtia eurycara Kostylev, 1935, ♀, Azerbaijan; M, J. irana Kostylev, 1935, ♀, Azerbaijan. A–C, emargination between lateral lamella and postero-lateral process of propodeum; D, E, tegula; F, G, apex of metasoma in dorsal view; H, metasomal terga II–VI; I–K, metasoma in dorsal view; L, M, metasomal terga II–IV. Scale bars 1 mm.	2025-11-03	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.		Zenodo	biologists	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.			
03B187DCFFCFFFCAFF39861AFD51FD4D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17889771/files/figure.png	https://doi.org/10.5281/zenodo.17889771	FIGURE 11. Distribution of pollen wasps in the Caucasus. A, Celonites cagrii Mauss & Yildirim, 2022; B, C. carpenteri Fateryga & Mauss, sp. nov.; C, C. ivanovi Mauss & Fateryga, 2022; D, C. laetus Panfilov, 1968; E, C. osseus Morawitz, 1888; F, C. tauricus Kostylev, 1935; G, C. tudesculptus Kostylev, 1935; H, Ceramius caucasicus André, 1884; I, Jugurtia eurycara Kostylev, 1935; J, J. irana Kostylev, 1935; K, Quartinia aliyevi Fateryga, Mauss, Proshchalykin & Maharramov, sp. nov.; L, Q. araxana Giordani Soika, 1960.	FIGURE 11. Distribution of pollen wasps in the Caucasus. A, Celonites cagrii Mauss & Yildirim, 2022; B, C. carpenteri Fateryga & Mauss, sp. nov.; C, C. ivanovi Mauss & Fateryga, 2022; D, C. laetus Panfilov, 1968; E, C. osseus Morawitz, 1888; F, C. tauricus Kostylev, 1935; G, C. tudesculptus Kostylev, 1935; H, Ceramius caucasicus André, 1884; I, Jugurtia eurycara Kostylev, 1935; J, J. irana Kostylev, 1935; K, Quartinia aliyevi Fateryga, Mauss, Proshchalykin & Maharramov, sp. nov.; L, Q. araxana Giordani Soika, 1960.	2025-11-03	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.		Zenodo	biologists	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.			
03B187DCFFCAFFCAFF398275FEB4F81B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17889767/files/figure.png	https://doi.org/10.5281/zenodo.17889767	FIGURE 9. Characters of pollen wasps. A, D, Ceramius caucasicus André, 1884, ♂,Azerbaijan; B, Jugurtia eurycara Kostylev, 1935, ♀, Azerbaijan; C, J. irana Kostylev, 1935, ♀, Azerbaijan; E, J. eurycara, ♂,Azerbaijan; F, G, Celonites osseus Morawitz, 1888, ♀, Armenia; H, C. laetus Panfilov, 1968, ♀, holotype; I, C. tauricus Kostylev, 1935, ♀, Azerbaijan; J, C. tudesculptus Kostylev, 1935, ♀, Azerbaijan. A, clypeus; B, C, F, head in frontal view; D, E, antenna; G–J, head in lateral view. Scale bars 1 mm.	FIGURE 9. Characters of pollen wasps. A, D, Ceramius caucasicus André, 1884, ♂,Azerbaijan; B, Jugurtia eurycara Kostylev, 1935, ♀, Azerbaijan; C, J. irana Kostylev, 1935, ♀, Azerbaijan; E, J. eurycara, ♂,Azerbaijan; F, G, Celonites osseus Morawitz, 1888, ♀, Armenia; H, C. laetus Panfilov, 1968, ♀, holotype; I, C. tauricus Kostylev, 1935, ♀, Azerbaijan; J, C. tudesculptus Kostylev, 1935, ♀, Azerbaijan. A, clypeus; B, C, F, head in frontal view; D, E, antenna; G–J, head in lateral view. Scale bars 1 mm.	2025-11-03	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.		Zenodo	biologists	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.			
03B187DCFFCAFFCAFF398275FEB4F81B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17889771/files/figure.png	https://doi.org/10.5281/zenodo.17889771	FIGURE 11. Distribution of pollen wasps in the Caucasus. A, Celonites cagrii Mauss & Yildirim, 2022; B, C. carpenteri Fateryga & Mauss, sp. nov.; C, C. ivanovi Mauss & Fateryga, 2022; D, C. laetus Panfilov, 1968; E, C. osseus Morawitz, 1888; F, C. tauricus Kostylev, 1935; G, C. tudesculptus Kostylev, 1935; H, Ceramius caucasicus André, 1884; I, Jugurtia eurycara Kostylev, 1935; J, J. irana Kostylev, 1935; K, Quartinia aliyevi Fateryga, Mauss, Proshchalykin & Maharramov, sp. nov.; L, Q. araxana Giordani Soika, 1960.	FIGURE 11. Distribution of pollen wasps in the Caucasus. A, Celonites cagrii Mauss & Yildirim, 2022; B, C. carpenteri Fateryga & Mauss, sp. nov.; C, C. ivanovi Mauss & Fateryga, 2022; D, C. laetus Panfilov, 1968; E, C. osseus Morawitz, 1888; F, C. tauricus Kostylev, 1935; G, C. tudesculptus Kostylev, 1935; H, Ceramius caucasicus André, 1884; I, Jugurtia eurycara Kostylev, 1935; J, J. irana Kostylev, 1935; K, Quartinia aliyevi Fateryga, Mauss, Proshchalykin & Maharramov, sp. nov.; L, Q. araxana Giordani Soika, 1960.	2025-11-03	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.		Zenodo	biologists	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.			
03B187DCFFCBFFCBFF3981E9FEE2FB72.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17889767/files/figure.png	https://doi.org/10.5281/zenodo.17889767	FIGURE 9. Characters of pollen wasps. A, D, Ceramius caucasicus André, 1884, ♂,Azerbaijan; B, Jugurtia eurycara Kostylev, 1935, ♀, Azerbaijan; C, J. irana Kostylev, 1935, ♀, Azerbaijan; E, J. eurycara, ♂,Azerbaijan; F, G, Celonites osseus Morawitz, 1888, ♀, Armenia; H, C. laetus Panfilov, 1968, ♀, holotype; I, C. tauricus Kostylev, 1935, ♀, Azerbaijan; J, C. tudesculptus Kostylev, 1935, ♀, Azerbaijan. A, clypeus; B, C, F, head in frontal view; D, E, antenna; G–J, head in lateral view. Scale bars 1 mm.	FIGURE 9. Characters of pollen wasps. A, D, Ceramius caucasicus André, 1884, ♂,Azerbaijan; B, Jugurtia eurycara Kostylev, 1935, ♀, Azerbaijan; C, J. irana Kostylev, 1935, ♀, Azerbaijan; E, J. eurycara, ♂,Azerbaijan; F, G, Celonites osseus Morawitz, 1888, ♀, Armenia; H, C. laetus Panfilov, 1968, ♀, holotype; I, C. tauricus Kostylev, 1935, ♀, Azerbaijan; J, C. tudesculptus Kostylev, 1935, ♀, Azerbaijan. A, clypeus; B, C, F, head in frontal view; D, E, antenna; G–J, head in lateral view. Scale bars 1 mm.	2025-11-03	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.		Zenodo	biologists	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.			
03B187DCFFCBFFCBFF3981E9FEE2FB72.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17889769/files/figure.png	https://doi.org/10.5281/zenodo.17889769	FIGURE 10. Characters of pollen wasps. A, I, Celonites tauricus Kostylev, 1935, ♀, Azerbaijan; B, C. ivanovi Mauss & Fateryga, 2022, ♀, Dagestan; C, J, C. tudesculptus Kostylev, 1935, ♀, Azerbaijan; D, H, C. laetus Panfilov, 1968, ♀, holotype; E, C. osseus Morawitz, 1888, ♀, Armenia; F, C. tauricus, ♂, Dagestan; G, C. osseus, ♂, Iran; K, C. ivanovi, ♂, paratype; L, Jugurtia eurycara Kostylev, 1935, ♀, Azerbaijan; M, J. irana Kostylev, 1935, ♀, Azerbaijan. A–C, emargination between lateral lamella and postero-lateral process of propodeum; D, E, tegula; F, G, apex of metasoma in dorsal view; H, metasomal terga II–VI; I–K, metasoma in dorsal view; L, M, metasomal terga II–IV. Scale bars 1 mm.	FIGURE 10. Characters of pollen wasps. A, I, Celonites tauricus Kostylev, 1935, ♀, Azerbaijan; B, C. ivanovi Mauss & Fateryga, 2022, ♀, Dagestan; C, J, C. tudesculptus Kostylev, 1935, ♀, Azerbaijan; D, H, C. laetus Panfilov, 1968, ♀, holotype; E, C. osseus Morawitz, 1888, ♀, Armenia; F, C. tauricus, ♂, Dagestan; G, C. osseus, ♂, Iran; K, C. ivanovi, ♂, paratype; L, Jugurtia eurycara Kostylev, 1935, ♀, Azerbaijan; M, J. irana Kostylev, 1935, ♀, Azerbaijan. A–C, emargination between lateral lamella and postero-lateral process of propodeum; D, E, tegula; F, G, apex of metasoma in dorsal view; H, metasomal terga II–VI; I–K, metasoma in dorsal view; L, M, metasomal terga II–IV. Scale bars 1 mm.	2025-11-03	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.		Zenodo	biologists	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.			
03B187DCFFCBFFCBFF3981E9FEE2FB72.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17889771/files/figure.png	https://doi.org/10.5281/zenodo.17889771	FIGURE 11. Distribution of pollen wasps in the Caucasus. A, Celonites cagrii Mauss & Yildirim, 2022; B, C. carpenteri Fateryga & Mauss, sp. nov.; C, C. ivanovi Mauss & Fateryga, 2022; D, C. laetus Panfilov, 1968; E, C. osseus Morawitz, 1888; F, C. tauricus Kostylev, 1935; G, C. tudesculptus Kostylev, 1935; H, Ceramius caucasicus André, 1884; I, Jugurtia eurycara Kostylev, 1935; J, J. irana Kostylev, 1935; K, Quartinia aliyevi Fateryga, Mauss, Proshchalykin & Maharramov, sp. nov.; L, Q. araxana Giordani Soika, 1960.	FIGURE 11. Distribution of pollen wasps in the Caucasus. A, Celonites cagrii Mauss & Yildirim, 2022; B, C. carpenteri Fateryga & Mauss, sp. nov.; C, C. ivanovi Mauss & Fateryga, 2022; D, C. laetus Panfilov, 1968; E, C. osseus Morawitz, 1888; F, C. tauricus Kostylev, 1935; G, C. tudesculptus Kostylev, 1935; H, Ceramius caucasicus André, 1884; I, Jugurtia eurycara Kostylev, 1935; J, J. irana Kostylev, 1935; K, Quartinia aliyevi Fateryga, Mauss, Proshchalykin & Maharramov, sp. nov.; L, Q. araxana Giordani Soika, 1960.	2025-11-03	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.		Zenodo	biologists	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.			
03B187DCFFCBFFCBFF39843AFC82F9F4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17889767/files/figure.png	https://doi.org/10.5281/zenodo.17889767	FIGURE 9. Characters of pollen wasps. A, D, Ceramius caucasicus André, 1884, ♂,Azerbaijan; B, Jugurtia eurycara Kostylev, 1935, ♀, Azerbaijan; C, J. irana Kostylev, 1935, ♀, Azerbaijan; E, J. eurycara, ♂,Azerbaijan; F, G, Celonites osseus Morawitz, 1888, ♀, Armenia; H, C. laetus Panfilov, 1968, ♀, holotype; I, C. tauricus Kostylev, 1935, ♀, Azerbaijan; J, C. tudesculptus Kostylev, 1935, ♀, Azerbaijan. A, clypeus; B, C, F, head in frontal view; D, E, antenna; G–J, head in lateral view. Scale bars 1 mm.	FIGURE 9. Characters of pollen wasps. A, D, Ceramius caucasicus André, 1884, ♂,Azerbaijan; B, Jugurtia eurycara Kostylev, 1935, ♀, Azerbaijan; C, J. irana Kostylev, 1935, ♀, Azerbaijan; E, J. eurycara, ♂,Azerbaijan; F, G, Celonites osseus Morawitz, 1888, ♀, Armenia; H, C. laetus Panfilov, 1968, ♀, holotype; I, C. tauricus Kostylev, 1935, ♀, Azerbaijan; J, C. tudesculptus Kostylev, 1935, ♀, Azerbaijan. A, clypeus; B, C, F, head in frontal view; D, E, antenna; G–J, head in lateral view. Scale bars 1 mm.	2025-11-03	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.		Zenodo	biologists	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.			
03B187DCFFCBFFCBFF39843AFC82F9F4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17889769/files/figure.png	https://doi.org/10.5281/zenodo.17889769	FIGURE 10. Characters of pollen wasps. A, I, Celonites tauricus Kostylev, 1935, ♀, Azerbaijan; B, C. ivanovi Mauss & Fateryga, 2022, ♀, Dagestan; C, J, C. tudesculptus Kostylev, 1935, ♀, Azerbaijan; D, H, C. laetus Panfilov, 1968, ♀, holotype; E, C. osseus Morawitz, 1888, ♀, Armenia; F, C. tauricus, ♂, Dagestan; G, C. osseus, ♂, Iran; K, C. ivanovi, ♂, paratype; L, Jugurtia eurycara Kostylev, 1935, ♀, Azerbaijan; M, J. irana Kostylev, 1935, ♀, Azerbaijan. A–C, emargination between lateral lamella and postero-lateral process of propodeum; D, E, tegula; F, G, apex of metasoma in dorsal view; H, metasomal terga II–VI; I–K, metasoma in dorsal view; L, M, metasomal terga II–IV. Scale bars 1 mm.	FIGURE 10. Characters of pollen wasps. A, I, Celonites tauricus Kostylev, 1935, ♀, Azerbaijan; B, C. ivanovi Mauss & Fateryga, 2022, ♀, Dagestan; C, J, C. tudesculptus Kostylev, 1935, ♀, Azerbaijan; D, H, C. laetus Panfilov, 1968, ♀, holotype; E, C. osseus Morawitz, 1888, ♀, Armenia; F, C. tauricus, ♂, Dagestan; G, C. osseus, ♂, Iran; K, C. ivanovi, ♂, paratype; L, Jugurtia eurycara Kostylev, 1935, ♀, Azerbaijan; M, J. irana Kostylev, 1935, ♀, Azerbaijan. A–C, emargination between lateral lamella and postero-lateral process of propodeum; D, E, tegula; F, G, apex of metasoma in dorsal view; H, metasomal terga II–VI; I–K, metasoma in dorsal view; L, M, metasomal terga II–IV. Scale bars 1 mm.	2025-11-03	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.		Zenodo	biologists	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.			
03B187DCFFCBFFCBFF39843AFC82F9F4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17889771/files/figure.png	https://doi.org/10.5281/zenodo.17889771	FIGURE 11. Distribution of pollen wasps in the Caucasus. A, Celonites cagrii Mauss & Yildirim, 2022; B, C. carpenteri Fateryga & Mauss, sp. nov.; C, C. ivanovi Mauss & Fateryga, 2022; D, C. laetus Panfilov, 1968; E, C. osseus Morawitz, 1888; F, C. tauricus Kostylev, 1935; G, C. tudesculptus Kostylev, 1935; H, Ceramius caucasicus André, 1884; I, Jugurtia eurycara Kostylev, 1935; J, J. irana Kostylev, 1935; K, Quartinia aliyevi Fateryga, Mauss, Proshchalykin & Maharramov, sp. nov.; L, Q. araxana Giordani Soika, 1960.	FIGURE 11. Distribution of pollen wasps in the Caucasus. A, Celonites cagrii Mauss & Yildirim, 2022; B, C. carpenteri Fateryga & Mauss, sp. nov.; C, C. ivanovi Mauss & Fateryga, 2022; D, C. laetus Panfilov, 1968; E, C. osseus Morawitz, 1888; F, C. tauricus Kostylev, 1935; G, C. tudesculptus Kostylev, 1935; H, Ceramius caucasicus André, 1884; I, Jugurtia eurycara Kostylev, 1935; J, J. irana Kostylev, 1935; K, Quartinia aliyevi Fateryga, Mauss, Proshchalykin & Maharramov, sp. nov.; L, Q. araxana Giordani Soika, 1960.	2025-11-03	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.		Zenodo	biologists	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.			
03B187DCFFCBFFD7FF3987E0FD2BFA9D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17889757/files/figure.png	https://doi.org/10.5281/zenodo.17889757	FIGURE 6. Quartinia aliyevi Fateryga, Mauss, Proshchalykin & Maharramov, sp. nov., ♀, holotype. A, habitus in dorsal view; B, habitus in lateral view; C, apex of metasoma in dorsal view; D, head in frontal view; E, labels. Scale bars 0.5 mm.	FIGURE 6. Quartinia aliyevi Fateryga, Mauss, Proshchalykin & Maharramov, sp. nov., ♀, holotype. A, habitus in dorsal view; B, habitus in lateral view; C, apex of metasoma in dorsal view; D, head in frontal view; E, labels. Scale bars 0.5 mm.	2025-11-03	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.		Zenodo	biologists	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.			
03B187DCFFCBFFD7FF3987E0FD2BFA9D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17889761/files/figure.png	https://doi.org/10.5281/zenodo.17889761	FIGURE 7. Quartinia aliyevi Fateryga, Mauss, Proshchalykin & Maharramov, sp. nov., ♂, paratype. A, habitus in dorsal view; B, habitus in lateral view; C, metasomal terga III–VII; D, head in frontal view; E, genitalia in dorsal view; F, right foretibia, view on the outside; G, right foretibia and basitarsus of foreleg in posterior view. Scale bars 0.5 mm.	FIGURE 7. Quartinia aliyevi Fateryga, Mauss, Proshchalykin & Maharramov, sp. nov., ♂, paratype. A, habitus in dorsal view; B, habitus in lateral view; C, metasomal terga III–VII; D, head in frontal view; E, genitalia in dorsal view; F, right foretibia, view on the outside; G, right foretibia and basitarsus of foreleg in posterior view. Scale bars 0.5 mm.	2025-11-03	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.		Zenodo	biologists	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.			
03B187DCFFCBFFD7FF3987E0FD2BFA9D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17889765/files/figure.png	https://doi.org/10.5281/zenodo.17889765	FIGURE 8. Quartinia araxana Giordani Soika, 1960. A, B, ♀, Azerbaijan; C–F, ♂, Azerbaijan. A, mesosoma in dorsal view; B, C, head in frontal view; D, right foretibia, view on the outside; E, right foretibia in posterior view; F, genitalia in dorsal view. Scale bars 0.5 mm.	FIGURE 8. Quartinia araxana Giordani Soika, 1960. A, B, ♀, Azerbaijan; C–F, ♂, Azerbaijan. A, mesosoma in dorsal view; B, C, head in frontal view; D, right foretibia, view on the outside; E, right foretibia in posterior view; F, genitalia in dorsal view. Scale bars 0.5 mm.	2025-11-03	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.		Zenodo	biologists	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.			
03B187DCFFCBFFD7FF3987E0FD2BFA9D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17889771/files/figure.png	https://doi.org/10.5281/zenodo.17889771	FIGURE 11. Distribution of pollen wasps in the Caucasus. A, Celonites cagrii Mauss & Yildirim, 2022; B, C. carpenteri Fateryga & Mauss, sp. nov.; C, C. ivanovi Mauss & Fateryga, 2022; D, C. laetus Panfilov, 1968; E, C. osseus Morawitz, 1888; F, C. tauricus Kostylev, 1935; G, C. tudesculptus Kostylev, 1935; H, Ceramius caucasicus André, 1884; I, Jugurtia eurycara Kostylev, 1935; J, J. irana Kostylev, 1935; K, Quartinia aliyevi Fateryga, Mauss, Proshchalykin & Maharramov, sp. nov.; L, Q. araxana Giordani Soika, 1960.	FIGURE 11. Distribution of pollen wasps in the Caucasus. A, Celonites cagrii Mauss & Yildirim, 2022; B, C. carpenteri Fateryga & Mauss, sp. nov.; C, C. ivanovi Mauss & Fateryga, 2022; D, C. laetus Panfilov, 1968; E, C. osseus Morawitz, 1888; F, C. tauricus Kostylev, 1935; G, C. tudesculptus Kostylev, 1935; H, Ceramius caucasicus André, 1884; I, Jugurtia eurycara Kostylev, 1935; J, J. irana Kostylev, 1935; K, Quartinia aliyevi Fateryga, Mauss, Proshchalykin & Maharramov, sp. nov.; L, Q. araxana Giordani Soika, 1960.	2025-11-03	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.		Zenodo	biologists	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.			
03B187DCFFCBFFD7FF3987E0FD2BFA9D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17889743/files/figure.png	https://doi.org/10.5281/zenodo.17889743	FIGURE 2. Habitats of Celonites carpenteri Fateryga & Mauss, sp. nov. (A) and Quartinia aliyevi Fateryga, Mauss, Proshchalykin & Maharramov, sp. nov. (B) in Azerbaijan.	FIGURE 2. Habitats of Celonites carpenteri Fateryga & Mauss, sp. nov. (A) and Quartinia aliyevi Fateryga, Mauss, Proshchalykin & Maharramov, sp. nov. (B) in Azerbaijan.	2025-11-03	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.		Zenodo	biologists	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.			
03B187DCFFD7FFD4FF398491FB16FDD1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17889765/files/figure.png	https://doi.org/10.5281/zenodo.17889765	FIGURE 8. Quartinia araxana Giordani Soika, 1960. A, B, ♀, Azerbaijan; C–F, ♂, Azerbaijan. A, mesosoma in dorsal view; B, C, head in frontal view; D, right foretibia, view on the outside; E, right foretibia in posterior view; F, genitalia in dorsal view. Scale bars 0.5 mm.	FIGURE 8. Quartinia araxana Giordani Soika, 1960. A, B, ♀, Azerbaijan; C–F, ♂, Azerbaijan. A, mesosoma in dorsal view; B, C, head in frontal view; D, right foretibia, view on the outside; E, right foretibia in posterior view; F, genitalia in dorsal view. Scale bars 0.5 mm.	2025-11-03	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.		Zenodo	biologists	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.			
03B187DCFFD7FFD4FF398491FB16FDD1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17889771/files/figure.png	https://doi.org/10.5281/zenodo.17889771	FIGURE 11. Distribution of pollen wasps in the Caucasus. A, Celonites cagrii Mauss & Yildirim, 2022; B, C. carpenteri Fateryga & Mauss, sp. nov.; C, C. ivanovi Mauss & Fateryga, 2022; D, C. laetus Panfilov, 1968; E, C. osseus Morawitz, 1888; F, C. tauricus Kostylev, 1935; G, C. tudesculptus Kostylev, 1935; H, Ceramius caucasicus André, 1884; I, Jugurtia eurycara Kostylev, 1935; J, J. irana Kostylev, 1935; K, Quartinia aliyevi Fateryga, Mauss, Proshchalykin & Maharramov, sp. nov.; L, Q. araxana Giordani Soika, 1960.	FIGURE 11. Distribution of pollen wasps in the Caucasus. A, Celonites cagrii Mauss & Yildirim, 2022; B, C. carpenteri Fateryga & Mauss, sp. nov.; C, C. ivanovi Mauss & Fateryga, 2022; D, C. laetus Panfilov, 1968; E, C. osseus Morawitz, 1888; F, C. tauricus Kostylev, 1935; G, C. tudesculptus Kostylev, 1935; H, Ceramius caucasicus André, 1884; I, Jugurtia eurycara Kostylev, 1935; J, J. irana Kostylev, 1935; K, Quartinia aliyevi Fateryga, Mauss, Proshchalykin & Maharramov, sp. nov.; L, Q. araxana Giordani Soika, 1960.	2025-11-03	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.		Zenodo	biologists	Fateryga, Alexander V.;Mauss, Volker;Proshchalykin, Maxim Yu.;Maharramov, Mahir M.			
