taxonID	type	description	language	source
03BE8789FFF39771FF47FA95E0A9A0DE.taxon	materials_examined	Material examined (HMIM). Ardabil province: 21 exs.: Moghan Protected Area, ca. 4 km W Aslanduz, Aras river, nr. Border Guard Police Station, N 39 ° 29 ’ 42.1 ”, E 48 ° 03 ’ 04.5 ”, 101 m, 28. VI. 2017, leg. H. Nasserzadeh & M. Mojdehi (Light trap); 1 ex.: Moghan Protected Area, Meshkin-Shahr to Parsabad rd., ca. 7 km N Gharehghaj, N 39 ° 08 ’ 38.8 ”, E 47 ° 38 ’ 43.3 ” E, 453 m, 25. V. 2017, leg. H. Nasserzadeh; 8 exs.: Moghan Protected Area, Aslanduz to Parsabad rd., ca. 5 km E Eslamabad, N 39 ° 27 ’ 05.05 ”, E 47 ° 23 ’ 08.6 ”, 140 m, 27. VI. 2017, leg. H. Nasserzadeh; 8 exs.: Moghan Protected Area, ca. 1 km W Aslanduz, nr. Aras river, N 39 ° 26 ’ 15.1 ”, E 47 ° 24 ’ 05.9 ”, 182 m, 25. VI. 2017, leg. H. Nasserzadeh & M. Mojdehi (Light trap); 1 ex.: “ ARDABIL, Moghan, 10 km. Pârsâbâd, 100 m., 23 - 24. V. 1997, [leg.] Badii / Safz. / Naz. V ”. Azarbaijan-e Sharghi province: 2 exs.: Arasbaran Protected Area, Jananlu to Kaleybar rd., ca. 28 km S Jananlu, N 38 ° 57 ’ 09.8 ”, E 47 ° 01 ’ 17.3 ”, 756 m, 26. VI. 2017, leg. H. Nasserzadeh; 6 exs.: Arasbaran Protected Area, N Kaleybar, Jananlu to Asheghlu rd., 500 m E Toali, N 39 ° 05 ’ 04.7 ”, E 46 ° 49 ’ 27.8 ”, 1194 m, 24. VI. 2015, leg. H. Nasserzadeh; 1 ex.: Arasbaran Protected Area, Jananlu to Kaleybar rd., ca. 28 km S Jananlu, N 38 ° 57 ’ 09.8 ”, E 47 ° 01 ’ 17.3 ”, 756 m, 26. VI. 2017, leg. H. Nasserzadeh; 1 ex.: “ AZARBAIDJAN, Ahar, Kalibar [Kaleybar], 14.6.1973, [leg.] Ragabi [Rajabi] ’. Climate types. Semi-arid, cold to cool winters and warm summers.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFF29770FF47FF5CE7A7A17C.taxon	materials_examined	Material examined (HMIM). Azarbaijan-e Gharbi province: 1 ex.: “ Mianduab [Miandoab], 11. VII. [19] 66, [leg.] Argand [Arghand] ”; 4 exs.: “ Mianduab [Miandoab], 9.8. [19] 66, [leg.] Argand [Arghand] ”. Climate types. Semi-arid, cold winters and warm summers. MAP 2. Distribution map of Aeoloides, Aeolosomus, Lacon and Lanelater species in northwestern Iran.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFF29777FF47F97EE0A9A57E.taxon	materials_examined	Material examined (HMIM). Ardabil province: 6 exs.: Moghan Protected Area, ca. 1 km W Aslanduz, nr. Aras river, N 39 ° 26 ’ 15.1 ”, E 47 ° 24 ’ 05.9 ”, 182 m, 25. VI. 2017, leg. H. Nasserzadeh & M. Mojdehi (Light trap); 4 exs.: Moghan Protected Area, ca. 4 km W Aslanduz, Aras river, nr. Border Guard Police Station, N 39 ° 29 ’ 42.1 ”, E 48 ° 03 ’ 04.5 ”, 101 m, 28. VI. 2017, leg. H. Nasserzadeh & M. Mojdehi (Light trap); 2 exs.: Moghan Protected Area, Sarband to Ardabil rd., ca. 20 Km S Sarband, nr. Alamlu vill., N 39 ° 37 ’ 30.6 ”, E 47 ° 46 ’ 57.8 ”, 58 m, 27. VI. 2017, leg. H. Nasserzadeh & M. Mojdehi (Light trap). Azarbaijan-e Gharbi province: 2 exs.: “ Mianduab [Miandoab], 11. VII. [19] 66, [leg.] Argand [Arghand] ”; 2 exs.: “ Mianduab [Miandoab], 29.6. [19] 66, [leg.] Argand [Arghand] ”. Azarbaijan-e Sharghi province: 1 ex.: Arasbaran Protected Area, Jananlu to Kaleybar rd., ca. 28 km S Jananlu, N 38 ° 57 ’ 09.8 ”, E 47 ° 01 ’ 17.3 ”, 756 m, 26. VI. 2017, leg. H. Nasserzadeh; 1 ex.: Heris to Ahar rd., 5 km N Yengejeh, N 38 ° 19 ’ 47.7 ”, E 46 ° 52 ’ 02.0 ”, 1709 m, 22. V. 2015, leg. H. Nasserzadeh; 1 ex.: Yari Ghari Protected Area, Heris, Khoshknab, N 38 ° 14 ’ 18.7 ”, E 47 ° 12 ’ 03.8 ”, 1942 m, 22. VI. 2015, leg. H. Nasserzadeh; 1 ex.: Abdol-Razagh Protected Area, Saqez, nr. Ashaab village, N 36 ° 14 ’ 57.1 ”, E 46 ° 46 ’ 19.5 ” E, 1881 m, 29. V. 2016, leg. H. Nasserzadeh; 1 ex.: Divandarreh to Hosseinabad rd., Bagherabad, N 35 ° 36 ’ 35.5 ”, E 47 ° 08 ’ 57.3 ”, 1803 m, 30. V. 2016, leg. H. Nasserzadeh. Zanjan province: 1 ex.: Sorkhabad Protected Area, Sohrin village, Sohrin Ranger Station, N 36 ° 52 ’ 05.9 ”, E 48 ° 19 ’ 20.9 ”, 1700 m, 21. V. 2016, leg. H. Nasserzadeh; 1 ex.: Zanjan to Miyaneh rd., 1 km S Kushkan, N 36 ° 43 ’ 37.8 ”, E 48 ° 19 ’ 10.1 ”, 1527 m, 21. VI. 2015, leg. H. Nasserzadeh. Climate types. Semi-arid, cool to cold winters and warm summers.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFF59776FF47FD78E778A626.taxon	materials_examined	Material examined (HMIM). Azarbaijan-e Gharbi province: 1 ex.: Sardasht, Mirabad to Alootan rd., ca. 3 km W Mirabad, N 36 ° 22 ’ 53.6 ”, E 45 ° 20 ’ 15.6 ”, 1650 m, 27. V. 2016, leg. H. Nasserzadeh. Ardabil province: 29 exs.: Moghan Protected Area, Sarband to Ardabil rd., ca. 20 Km S Sarband, nr. Alamlu vill., N 39 ° 37 ’ 30.6 ”, E 47 ° 46 ’ 57.8 ”, 58 m, 27. VI. 2017, leg. H. Nasserzadeh & M. Mojdehi (Light trap); 4 exs.: Moghan Protected Area, ca. 4 km W Aslanduz, Aras river, nr. Border Guard Police Station, N 39 ° 29 ’ 42.1 ”, E 48 ° 03 ’ 04.5 ”, 101 m, 28. VI. 2017, leg. H. Nasserzadeh & M. Mojdehi (Light trap); 12 exs.: Moghan Protected Area, Meshkin-Shahr to Parsabad rd., ca. 7 km N Gharehghaj, N 39 ° 08 ’ 38.8 ”, E 47 ° 38 ’ 43.3 ” E, 453 m, 25. V. 2017, leg. H. Nasserzadeh; 1 ex.: Moghan Protected Area, Aslanduz to Parsabad rd., ca. 5 km E Eslamabad, N 39 ° 27 ’ 05.05 ”, E 47 ° 23 ’ 08.6 ”, 140 m, 27. VI. 2017, leg. H. Nasserzadeh; 20 exs.: Moghan Protected Area, Ardabil to Parsabad rd., 1 km S Qaraghaj, N 38 ° 59 ’ 06.5 ”, E 47 ° 41 ’ 22.4 ”, 1322 m, 25. VI. 2017, leg. H. Nasserzadeh; 2 exs.: Moghan Protected Area, Parsabad to Bilehsavar rd., ca. 20 km SE Parsabad, N 39 ° 25 ’ 18.5 ”, E 47 ° 35 ’ 43.3 ”, 225 m, 28. VI. 2017, leg. H. Nasserzadeh; 10 exs.: Moghan Protected Area, ca. 1 km W Aslanduz, nr. Aras river, N 39 ° 26 ’ 15.1 ”, E 47 ° 24 ’ 05.9 ”, 182 m, 25. VI. 2017, leg. H. Nasserzadeh & M. Mojdehi (Light trap). Kordestan province: 8 exs.: 50 km SW Sanandaj, Kusalan va Shaho Protected Area, Piyazeh MT., nr. Gelin village, N 35 ° 07 ’ 12.0 ”, E 46 ° 53 ’ 17.7 ”, ca. 2000 m, 1. VII. 2015, leg. H. Nasserzadeh; 3 exs.: Kusalan va Shaho Protected Area, Sarvabad, 2 km E Dezli, N 35 ° 22 ’ 34.5 ”, E 46 ° 09 ’ 36.4 ”, 1379 m, 30. VI. 2015, leg. H. Nasserzadeh (Light trap); 10 exs.: Kusalan va Shaho Protected Area, Oraman, Dezli to Daraki rd., 2 km N Daraki, N 35 ° 20 ’ 06.5 ”, E 46 ° 10 ’ 55.7 ”, 1720 m, 30. VI. 2015, leg. H. Nasserzadeh; 2 exs.: Marivan, Zarivar Wildlife Sanctuary, Tefli valley to Sardush rd., ca. 1 km E Sardush village, N 35 ° 32 ’ 33.3 ”, E 46 ° 05 ’ 01.5 ”, 1400 m, 22. VIII. 2017, leg. H. Nasserzadeh (light trap & searching before sunset); 2 exs.: Abdol-Razagh Protected Area, Saqez, nr. Ashaab village, N 36 ° 14 ’ 57.1 ”, E 46 ° 46 ’ 19.5 ” E, 1881 m, 29. V. 2016, leg. H. Nasserzadeh; 9 exs.: Saghez, Abdol-Razzagh Protected Area, ca. 1 km S Qaplantu village, nr. Patar mt., N 36 ° 13 ’ 10.7 ”, E 46 ° 42 ’ 37.6 ”, 1866 m, 29. V. 2016, leg. H. Nasserzadeh; 11 exs.: Badr va Parishan Protected Area, Qorveh, Meyhim-e Sofla to Meyhim-e Olia rd., ca. 1 km NW Meyhim-e Sofla, E 35 ° 03 ’ 26.4 ”, E 47 ° 52 ’ 13.9 ”, 2155 m, 31. V. 2016, leg. H. Nasserzadeh; 2 exs.: Saghez, Abdol-Razzagh Protected Area, ca. 3 km S Qaplantu village, N 36 ° 12 ’ 46.2 ”, E 46 ° 42 ’ 55.1 ”, 1864 m, 29. V. 2016, leg. H. Nasserzadeh; 3 exs.: Saghez to Marivan rd., ca. 13 km NW Bastam, N 35 ° 56 ’ 54.5 ”, E 46 ° 19 ’ 27.6 ”, 1516 m, 29. VI. 2015, leg. H. Nasserzadeh; 6 exs.: Saqez to Divandarreh rd., 29 km E Saqez, nr. Sonateh, nr. Khorkhoreh river, N 36 ° 10 ’ 38.9 ”, E 46 ° 34 ’ 01.6 ”, 1479 m, 26. V. 2016, leg. H. Nasserzadeh; 2 exs.: Baneh to Saqez rd., ca. 10 km NE Baneh, N 36 ° 02 ’ 52.6 ”, E 45 ° 57 ’ 29.3 ”, 1871 m, 28. V. 2016, leg. H. Nasserzadeh. Azarbaijan-e Sharghi province: 3 exs.: Kaleybar, Arasbaran Protected Area, Barzandigh to Peygham rd., 5 km W Peygham, N 38 ° 45 ’ 30.3 ”, E 46 ° 57 ’ 28.4 ”, 1621 m, 25. VI. 2015, leg. H. Nasserzadeh; 1 ex.: Yari Ghari Protected Area, Heris, Khoshknab, N 38 ° 14 ’ 18.7 ”, E 47 ° 12 ’ 03.8 ”, 1942 m, 22. VI. 2015, leg. H. Nasserzadeh; 2 exs.: Arasbaran Protected Area, N Kaleybar, Jananlu to Asheghlu rd., 500 m E Toali, N 39 ° 05 ’ 04.7 ”, E 46 ° 49 ’ 27.8 ”, 1194 m, 24. VI. 2015, leg. H. Nasserzadeh; 11 exs.: Arasbaran Protected Area, Jananlu to Kaleybar rd., ca. 28 km S Jananlu, N 38 ° 57 ’ 09.8 ”, E 47 ° 01 ’ 17.3 ”, 756 m, 26. VI. 2017, leg. H. Nasserzadeh. Zanjan province: 2 exs.: Zanjan to Miyaneh rd., 1 km S Kushkan, N 36 ° 43 ’ 37.8 ”, E 48 ° 19 ’ 10.1 ”, 1527 m, 21. VI. 2015, leg. H. Nasserzadeh; 2 exs.: Anguran Protected Area, Mahneshan, Behestan to Takaghaj rd., 2 km NW Behestan, nr. Anguran river, N 36 ° 40 ’ 30.3 ”, E 47 ° 43 ’ 15.5 ”, 1318 m, 25. V. 2016, leg. H. Nasserzadeh; 1 ex.: Anguran Protected Area, Dandi, Duzkand, N 36 ° 31 ’ 15.4 ”, E 47 ° 52 ’ 55.3 ”, 1527 m, 23. V. 2016, leg. H. Nasserzadeh. Climate types. Semi-arid and sub-humid, cool to cold winters and warm summers.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFF49776FF47FE10E063A596.taxon	description	(Fig. 1, Map 2)	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFF49776FF47FE10E063A596.taxon	materials_examined	Material examined (HMIM). Azarbaijan-e Gharbi province: 1 ex.: Mirabad Protected Area, Sardasht to Mirabad rd., ca. 5 km NW Nalas, Degeh village, N 36 ° 16 ’ E 45 ° 29 ’, ca. 1200 m, 27. V. 2016, leg. H. Nasserzadeh.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFF49776FF47FE10E063A596.taxon	discussion	Remarks: The diagnostic characters of the specimens show affinity to L. nadaii Platia & Németh, 2011 that is synonymyzed with L. unicolor by Kundrata et al. (2019 a). Climate types. Sub-humid, cold winters and warm summers.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFF79775FF47FD44E75CA08E.taxon	materials_examined	Material examined (HMIM). Ardabil province: 1 ex.: Moghan Protected Area, Aslanduz to Parsabad rd., ca. 5 km E Eslamabad, N 39 ° 27 ’ 05.05 ”, E 47 ° 23 ’ 08.6 ”, 140 m, 27. VI. 2017, leg. H. Nasserzadeh; 23 exs.: Moghan Protected Area, ca. 4 km W Aslanduz, Aras river, nr. Border Guard Police Station, N 39 ° 29 ’ 42.1 ”, E 48 ° 03 ’ 04.5 ”, 101 m, 28. VI. 2017, leg. H. Nasserzadeh & M. Mojdehi (Light trap). Azarbaigan-e Sharghi: 11 exs.: Arasbaran Protected Area, Jananlu to Kaleybar rd., ca. 28 km S Jananlu, N 38 ° 57 ’ 09.8 ”, E 47 ° 01 ’ 17.3 ”, 756 m, 26. VI. 2017, leg. H. Nasserzadeh. Climate types. Semi-arid, cool winters and warm summers. MAP 3. Distribution map of Adrastus, Drasterius and Synaptus species in northwestern Iran.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFF79775FF47FF14E07DA6C9.taxon	materials_examined	Material examined (HMIM). Kordestan province: 1 ex.: Kusalan va Shaho Protected Area, Sarvabad, ca. 1 km W Tefli village, N 35 ° 19 ’ E 46 ° 19 ’, 1300 m, 23. VIII. 2017, leg. H. Nasserzadeh (Light trap). Climate types. Semi-arid, cool winters and warm summers.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFF69774FF47FF14E07DA6B5.taxon	materials_examined	Material examined (HMIM). Ardabil province: 6 exs.: “ Moghan, 17.8.1967, [leg.] Daman [Damanabi ”; 2 exs.: “ Moghan, 15. IX. 1967, [leg.] D. Ab [Damanabi] ”; 2 exs.: “ Moghan, 24. VIII. 1967, [leg.] D. Ab [Damanabi] ”; 1 ex.: “ Moghan, 10. VII. 1967, [leg.] Damanabi ”; 1 ex.: “ Moghan, 15. VII. 1967, [leg.] Damanabi ”. Climate types. Semi-arid, cool winters and warm summers.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFF69774FF47FDE8E02AA0B1.taxon	materials_examined	Material examined (HMIM). Kordestan province: Abdol-Razagh Protected Area, Saqez, nr. Ashaab village, N 36 ° 14 ’ 57.1 ”, E 46 ° 46 ’ 19.5 ” E, 1881 m, 29. V. 2016, leg. H. Nasserzadeh. Climate types. Semi-arid, cold winters and warm summers. MAP 4. Distribution map of Agriotes species in northwestern Iran.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFF9977BFF47FF5CE700A505.taxon	materials_examined	Material examined (HMIM). Ardabil province: 68 exs.: Moghan Protected Area, ca. 1 km W Aslanduz, nr. Aras river, N 39 ° 26 ’ 15.1 ”, E 47 ° 24 ’ 05.9 ”, 182 m, 25. VI. 2017, leg. H. Nasserzadeh & M. Mojdehi (Light trap); 33 exs.: Moghan Protected Area, ca. 4 km W Aslanduz, Aras river, nr. Border Guard Police Station, N 39 ° 29 ’ 42.1 ”, E 48 ° 03 ’ 04.5 ”, 101 m, 28. VI. 2017, leg. H. Nasserzadeh & M. Mojdehi (Light trap); 1 ex.: Agh-Dagh Protected Area, Khalkhal to Hashjin rd., ca. 4 km SE Susahaab village, nr. Mirajin village, N 37 ° 23 ’ 19.8 ”, E 48 ° 31 ’ 01.7 ”, 30. VI. 2017, leg. H. Nasserzadeh & M. Mojdeh. Azarbaijan-e Sharghi province: 4 exs.: Arasbaran Protected Area, Jananlu to Kaleybar rd., ca. 28 km S Jananlu, N 38 ° 57 ’ 09.8 ”, E 47 ° 01 ’ 17.3 ”, 756 m, 26. VI. 2017, leg. H. Nasserzadeh. Kordestan province: 1 exs.: Saqez to Divandarreh rd., 29 km E Saqez, nr. Sonateh, nr. Khorkhoreh river, N 36 ° 10 ’ 38.9 ”, E 46 ° 34 ’ 01.6 ”, 1479 m, 26. V. 2016, leg. H. Nasserzadeh. Climate types. Semi-arid, cold to cool winters and mild to warm summers.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFF9977AFF47FD78E07DA79E.taxon	description	(Fig. 2, Map 4)	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFF9977AFF47FD78E07DA79E.taxon	materials_examined	Material examined (HMIM). Azarbaijan-e Sharghi province: 1 ex.: “ AZARBAIDJAN, Ahar, Kalibar [Kaleybar], 14.6.1973, [leg.] Ragabi [Rajabi]. Climate types. Semi-arid, cold winters and warm summers.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFF8977AFF47FE59E246A082.taxon	materials_examined	Material examined (HMIM). Kordestan province: 1 ex.: Baneh to Saqez rd., ca. 10 km NE Baneh, N 36 ° 02 ’ 52.6 ”, E 45 ° 57 ’ 29.3 ” E, 1871 m, 28. V. 2016, leg. H. Nasserzadeh. Climate types. Semi-arid, cold winters and warm summers. MAP 5. Distribution map of Ampedus, Anostirus, Reitterelater, Ectamenogonus, Mulsanteus and Pittonotus species in northwestern Iran.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFFB9779FF47FF14E07DA6C9.taxon	materials_examined	Material examined (HMIM). Kordestan province: 1 ex.: Kusalan va Shaho Protected Area, Sarvabad, Tarkhanabad village, N 35 ° 18 ’ 30.0 ” E 46 ° 22 ’ 47.8 ”, 1233 m, 30. VI. 2015, leg. H. Nasserzadeh (bait trap, rotten banana). Climate types. Semi-arid, cool winters and warm summers.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFFB9779FF47FD8CE07DA439.taxon	materials_examined	Material examined (HMIM). Azarbaijan-e Sharghi province: 4 exs.: Arasbaran ProtectedArea, Kaleybar, Barzandigh to Peygham rd., 5 km W Peygham, N 38 ° 45 ’ 30.3 ”, E 46 ° 57 ’ 28.4 ”, 1621 m, 25. VI. 2015, leg. H. Nasserzadeh; 1 ex.: Arasbaran Protected Area, nr. Vinagh, N 38 ° 59 ’ 56.6 ”, E 46 ° 51 ’ 45.4 ”, 23. VI. 2015, leg. Nasserzadeh, Hajesmailian, Montreuil (Light trap) Climate types. Semi-arid, cold winters and warm summers.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFFB9778FF47FC64E0A9A016.taxon	description	(Figs. 3 – 4, Map 6)	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFFB9778FF47FC64E0A9A016.taxon	materials_examined	Type locality. Kusalan va Shaho Protected Area, near Sarvabad city, Sarvabad to Dezli road, 2 km east of Dezli, southern Kordestan province. Type material. Holotype Ƌ (HMIM): “ Kordeatsn province, Kusalan va Shaho Protected Area, Sarvabad, 2 km E Dezli, N 35 ° 22 ’ 34.5 ”, E 46 ° 09 ’ 36.4 ”, 1379 m, 30. VI. 2015, leg. H. Nasserzadeh (L. T.) [Light trap] ”. Paratypes: 1 ♀ (HMIM): “ Ardabil [province], 14. VI. 2000, [leg.] Aghdam, on potato ”; 1 Ƌ (HMIM): “ Ardabil province, Moghan Protected Area, ca. 1 km W Aslanduz, nr. Aras river, N 39 ° 26 ’ 15.1 ”, E 47 ° 24 ’ 05.9 ”, 182 m, 25. VI. 2017, leg. H. Nasserzadeh & M. Mojdehi (L. T) [Light trap] ”; 40 ƋƋ, 4 ♀♀ (HMIM, CGP): “ Ardabil province, Moghan Protected Area, Sarband to Ardabil rd., ca. 20 Km S Sarband, nr. Alamlu vill., N 39 ° 37 ’ 30.6 ”, E 47 ° 46 ’ 57.8 ”, 58 m, 27. VI. 2017, leg. H. Nasserzadeh & M. Mojdehi (Light trap) ”; 2 ƋƋ (HMIM): “ Ardabil province, Moghan Protected Area, ca. 4 km W Aslanduz, Aras river, nr. Border Guard Police Station, N 39 ° 29 ’ 42.1 ”, E 48 ° 03 ’ 04.5 ”, 101 m, 28. VI. 2017, leg. H. Nasserzadeh & M. Mojdehi (L. T) [Light trap] ”; 3 ƋƋ (HMIM): “ [Ardabil province], Khalkhal, 25.7. [19] 49, [leg.] Mak. ”. (Azarbaijan-e Gharbi): 1 Ƌ (HMIM): “ W. Azarbayjan, Rezaieh [Urmia], [leg.] Abai (L. T) [Light trap] ”. Azarbaijan-e Sharghi: 10 ƋƋ (HMIM, CPG): “ Azarbaijan-e Sharghi province, Arasbaran Protected Area, nr. Vinagh, N 38 ° 59 ’ 56.6 ”, E 46 ° 51 ’ 45.4 ”, 23. VI. 2015, leg. Nasserzadeh, Hajesmailian, Montreuil (L. T) [Light trap] ”; 1 ♀: Arasbaran Protected Area, Jananlu to Kaleybar rd., ca. 28 km S Jananlu, N 38 ° 57 ’ 09.8 ”, E 47 ° 01 ’ 17.3 ”, 756 m, 26. VI. 2017, leg. H. Nasserzadeh Kordestan province): 2 ƋƋ, 1 ♀ (HMIM, CGP): same data as holotype; 1 Ƌ: “ Kordestan province, Marivân, Kanisânân, 1320 m, 9. VIII. 2004, [leg.] Gh. / Nem. ”. Additional material examined. The localities marked with an asterisk (*) include individual (s) with some morphological features intermediate between M. borumandi sp. n., M. kurdestanicus sp. n. and / or M. fusciceps (Gyllenhal, 1817) (see Remarks). Azarbaijan-e Gharbi province: * 80 exs.: Lake Urmia National Park, Rashakan Research Station, N 37 ° 20 ’ 38.8 ”, E 45 ° 17 ’ 37.4 ”, 1315 m, 08. VII- 6. VIII. 2016, leg. R. Vahabzadeh (Malaise trap); * 6 exs.: Lake Urmia National Park, Rashakan Research Station, N 37 ° 20 ’ 38.8 ”, E 45 ° 17 ’ 37.4 ”, 1315 m, 18 - 25. VI. 2016, leg. E. Gilasian & M. Parchami- Araghi (Malaise trap (. Azarbaijan-e Sharghi province: * 1 ex.: “ AZARBAIJAN, TABRIZ, BOSTANABAD / TIKMEHDASH, 2.8. [19] 83, [leg.] KAZEMI-NOSHAD ”.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFFB9778FF47FC64E0A9A016.taxon	diagnosis	Differential diagnosis. The length of the third antennomere in this species is less than half length of fourth (0.3 – 0.4 ×) this ratio is slightly more than 0.5 in the similar species Melanotus kurdestanicus sp. n., M. fusciceps and M. zagrosensis Platia 2016. It can also be separated from M. fusciceps and M. zagrosensis by the longer antennae that exceed the apicis of the posterior angles of pronotum by 1.5 - 2 antennomeres. It is one antennomere or less than one antennomere in M. fusciceps and M. zagrosensis respectively. The aedeagus and median lobe in M. borumandi sp. n. is not as slender and elongated as M. kurdestanicus sp. n. The length of the sclerified part of the bursa copulatrix in M. borumandi is almost as long as (or often clearly shorter than) the length of the ovipositor while it is clearly longer in M. fusciceps.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFFB9778FF47FC64E0A9A016.taxon	description	Description. Male. Body and legs unicolorous, rufous to rufo-testaceus, covered with dense and testaceous pubescence; blackened along marginal edges of contours and striae. Body length 12.2 – 16.5 mm, width 3.5 – 5.0 mm. Head as wide as anterior margin of pronotum near outer margin of eyes; covered with dense, umblicate punctures with interspaces mainly narrower than their own diameter; frons declining anteriorly, flat between eyes without a distinct depression, anterior margin thickened and slightly sinuate or straight at middle and protruding sub-horizontally above clypeus; antennae exceeding the apices of posterior angles of pronotum by one and a half to two antennomeres, antennomeres serrated from fourth to tenth; second and third moderately subconical, the latter slightly longer than half the length of second (second antennomere 0.7 × as long as third one, third antennomere slightly shorter than half length of fourth antennomere third antennomere 0.3 – 0.4 × as long as fourth) (Figure 6 b). Pronotum slightly (1.1 ×) broader than long, moderately and regularly convex, covered with coarse punctures, rather uniformly distributed, umbilicate, with interspaces mainly narrower than their own diameter, sides arcuate in posterior third, posterior angles slightly divergent with subparallel lateral sided, angles with raised carinae, carina longer than the angles, exceeding posterior third and sometimes reaching half the length of the lateral sides of pronotum. Scutellum shield-shaped, almost 1.5 × longer than wide, flat, punctures coarse and pubescent. Elytra 2.8 – 3.1 × longer than pronotum and as wide as or slightly (1.1 ×) wider at widest point, moderately convex; sides subparallel in anterior third, moderately narrowing to apex, striae regularly punctured, interstriae moderately and finely punctured. Last abdominal sternite entire and rounded along apical margin. Tarsomeres simple, first to fourth gradually decreasing in length, apical tarsomere slightly longer than the first and two times longer than the penultimate tarsomere. Aedeagus as in Figure 6 c, moderately elongated, length 2.0 – 2.2 mm, median lobe rather thickened, longer than parameres (by less than 1.25 times) (Figure 6 c), narrowing from the midlength and pointed at the apex, posterior angles of parameres apically rounded. Female. Similar to males except as follows: body length 14.0 – 18.4 mm, width 4.4 – 5.4 mm; antenna reaching apices of posterior angles of pronotum, second antennomere half or slightly longer than half the length of the third, third antennomere half as long as fourth; apical tarsomere almost as long as first tarsomere. Female genitalia as in Figure 4. Bursa copulatrix covered with sclerotized spines inside the memberanous wall; sclerified part of the bursa shorter than the length of the ovipositor.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFFB9778FF47FC64E0A9A016.taxon	etymology	Etymology. This species is named in honour of Hooshang Borumand (1936 – 2009), the first curator and the founder of the Coleoptera collection at the Hayk Mirzayans Insect Museum.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFFB9778FF47FC64E0A9A016.taxon	discussion	Remarks. The localities marked with an asterisk (*) include individuals with some variable morphological features intermediate between M. borumandi sp. n. and M. kurdestanicus sp. n. or M. fusciceps. Some specimens have a more thickened median lobe and are rather parallel sided or have a longer and slimmer median lobe; the ratio of the length of second and third antennomeres is intermediate between M. borumandi sp. n. and M. kurdestanicus sp. n. These specimens are also have fewer coarse punctures on the pronotum and show affinities to M. fusciceps and / or M. kurdestanicus sp. n. in the mentioned diagnostic features. Habitat. Specimens were collected in different habitat types ranging from mountainous oak forest in Arasbaran and Zagross to alluvial plain in the Moghan protected area and to saline land near Urmia lake. Specimens were collected by light trap or Malaise traps that were installed for collecting Hymenoptera and Hemiptera (Fig. 15). Climate types. Semi-arid, cold to cool winters and warm summers.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFFF977DFF47FF5CE063A6ED.taxon	materials_examined	Material examined (HMIM). Kordestan province: 3 exs.: Marivan, Zarivar Wildlife Sanctuary, Tefli valley to Sardush rd., ca. 1 km E Sardush village, N 35 ° 32 ’ 33.3 ”, E 46 ° 05 ’ 01.5 ”, 1400 m, 22. VIII. 2017, leg. H. Nasserzadeh (Light trap). Climate types. Sub-humid, cold winters and warm summers.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFFF977DFF47FE10E07DA5A9.taxon	materials_examined	Material examined (HMIM). Kordestan province: 2 exs.: Kusalan va Shaho Protected Area, Sarvabad, Tarkhanabad village, N 35 ° 18 ’ 30.0 ” E 46 ° 22 ’ 47.8 ”, 1233 m, 30. VI. 2015, leg. H. Nasserzadeh (bait trape, rotten banana); 1 ex. Marivan, Zarivar Wildlife Sanctuary, N 35 ° 32 ’ E 46 ° 08 ’, 1300 m, 21. VIII. 2017, leg. H. Nasserzadeh. Climate types. Semi-arid, cool winters and warm summers.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFFF977DFF47FCD4E049A489.taxon	materials_examined	Material examined (HMIM). Ardabil province: 3 exs.: “ ARDABIL, Meshkinshahr, Ilându, 1800 m., 3. VII. 1997, Bar. / Mof. ”. Climate types. Semi-arid, cold winters and mild summers.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFFF977CFF47FBF4E74DA45E.taxon	description	(Fig. 5, Map 6)	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFFF977CFF47FBF4E74DA45E.taxon	materials_examined	Type material. Holotype Ƌ (HMIM): 1 ex.: IRAN, Kordestan province prov., Marivan, Zarivar Wildlife Sanctuary, Tefli valley to Sardush rd., ca. 1 km E Sardush village, N 35 ° 32 ’ 33.3 ”, E 46 ° 05 ’ 01.5 ”, 1400 m, 22. VIII. 2017, leg. H. Nasserzadeh (Light trap). Paratype Ƌ (CGP): 1 ex.: IRAN, Kordestan province prov., Oramanat region, July 2015, leg. Nasserzadeh (certain locality unknown).	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFFF977CFF47FBF4E74DA45E.taxon	diagnosis	Differential diagnosis. The species is very similar to Melanotus borumandi sp. n., M. fusciceps and M. zagrosensis. It can be separated from M. fusciceps and M. zagrosensis by the longer antennae that exceed posterior angles by 1.5 antennomeres. The antennae exceed the posterior angles by one antennomere or less than one antennomere in M. fusciceps and M. zagrosensis respectively. The third antennomere in M. kurdestanicus is approximately half the length of the fourth or slightly longer, but in M. borumandi it is clearly shorter than half the length of the fourth (0.3 – 0.4 ×). The aedeagus and median lobe in M. kurdestanicus is rather elongated and slimmer than other three species, and the posterior angles of the apical portion of the parameres is curved contrary to other three species which is rather sharp.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFFF977CFF47FBF4E74DA45E.taxon	description	Description. Male. Body and legs unicolor, rufous, covered with testaceous pubescence; blackened along marginal edges of contours and striae. Body length 13.4 – 13.5 mm, width 4.0 – 4.1 mm. Head as wide as anterior margin of pronotum near outer margin of eyes; covered with dense, umbilicate punctures with interspaces mainly narrower than their own diameter; frons declining anteriorly, flat between eyes without a distinct depression, anterior margin thickened and slightly sinuate or straight at middle and protruding sub-horizontally above clypeus; antennae exceeding apicis of posterior angles of pronotum by one and a half antennomeres, antennomeres serrated from fourth to tenth; second and third moderately subconical, the latter 1.5 × longer than the second (second antennomere 0.6 × as long as third one), third antennomere about half the length of fourth (or slightly longer) (Figure 8 c). Pronotum slightly (1.1 – 1.2 ×) broader than long, moderately and regularly convex, covered with coarse punctures, rather uniformly distributed, umbilicate, with interspaces mainly narrower than their own diameter or sometimes equal, sides arcuate on posterior third, posterior angles slightly divergent with lateral side subparallel, angles with raised carinae (Figure 8 b), carina longer than the angles, reaching to posterior third of lateral sides of pronotum. Scutellum shield-shaped, almost 1.5 × longer than wide, flat, punctures coarse and pubescent. Elytra 3.0 – 3.2 × longer than pronotum and as wide as or slightly (1.1 times) wider in their widest size, moderately convex; sides subparallel in anterior third, moderately narrowing to apex, striae regularly punctured, interstriae moderately and finely punctured. Last abdominal sternite entire and with rounded apical margin. Tarsomeres simple, first to fourth gradually decreasing in length, apical tarsomere slightly longer than the first one and two times longer than penultimate tarsomere. Aedeagus as in figure 8 d, distinctly elongated, length 2.4 mm, median lobe slim, 1.25 times the length of parameres (which is exposed between parameres), narrowing from the midlength and sharply pointed at apex. Female. Unknown.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFFF977CFF47FBF4E74DA45E.taxon	etymology	Etymology. The name of the species refers to the geographical distribution in Kurdestan province in Iran. Habitat. Mountainous region with typical flora of central Zagross region like Oak, Almond and Salix trees and barberry shrubs. Climate types. Sub-humid to semi-arid, cool to cold winters and warm summers.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFE19763FF47FF5CE07DA6ED.taxon	materials_examined	Material examined (HMIM). Azarbaijan-e Sharghi province: 1 ex.: “ AZARBAIDJAN, Ahar, Kalibar [Kaleybar], 14.6.1973, [leg.] Ragabi [Rajabi]. Kordestan province: 2 exs: Saghez, Abdol-Razzagh Protected Area, ca. 1 km S Qaplantu village, nr. Patar mt., N 36 ° 13 ’ 10.7 ”, E 46 ° 42 ’ 37.6 ”, 1866 m, 29. V. 2016, leg. H. Nasserzadeh. Climate types. Semi-arid, cold winters and warm summers.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFE19763FF47FDE8E047A0B8.taxon	materials_examined	Material examined (HMIM). Kordestan province: 3 exs.: Kusalan va Shaho Protected Area, Sarvabad, 2 km E Dezli, N 35 ° 22 ’ 34.5 ”, E 46 ° 09 ’ 36.4 ”, 1379 m, 30. VI. 2015, leg. H. Nasserzadeh (Light trap). Climate types. Semi-arid, cool winters and warm summers. MAP 6. Distribution map of Melanotus species in northwestern Iran.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFE09762FF47FF14E764A691.taxon	description	(Fig. 6, Map 5)	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFE09762FF47FF14E764A691.taxon	materials_examined	Material examined (HMIM). Kordestan province: 2 exs.: Kusalan va Shaho Protected Area, Sarvabad, ca. 1 km W Tefli village, N 35 ° 19 ’ E 46 ° 19 ’, 1300 m, 23. VIII. 2017, leg. H. Nasserzadeh (Light trap); 3 exs.: Marivan, Zarivar Wildlife Sanctuary, Tefli valley to Sardush rd., ca. 1 km E Sardush village, N 35 ° 32 ’ 33.3 ”, E 46 ° 05 ’ 01.5 ”, 1400 m, 22. VIII. 2017, leg. H. Nasserzadeh (Light trap). Climate types. Sub-humid and Semi-arid, cold to cool winters and warm summers.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFE09760FF47FD44E063A4E6.taxon	description	(Fig. 7, Map 5)	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFE09760FF47FD44E063A4E6.taxon	materials_examined	Type locality. Mirabad Protected Area, Degeh village, Sardasht to Mirabad road, southern Azarbaijan-e Gharbi province. Type material. Holotype Ƌ (HMIM): Azarbaijan-e Gharbi prov., Sardasht to Mirabad rd., ca. 5 km NW Nalas, Degeh village, N 36 ° 16 ’ E 45 ° 29 ’, ca. 1200 m, 27. V. 2016, leg. H. Nasserzadeh (Light trap).	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFE09760FF47FD44E063A4E6.taxon	diagnosis	Differential diagnosis. The species can be separated from the species in neighboring regions, Reitterelater fulvus Reitter, 1891 and R. kovalenkoi Prosvirov, 2017 by the longer antenna that exceeds the apex of the posterior angles by less than one antennomere while the antenna in none of the mentioned species reaches the apex. It also differs from R. dubius Platia & Cate, 1990 by the acuminate hind angles of the pronotum, the general morphology of the prosternal process and the more elongated parameres.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFE09760FF47FD44E063A4E6.taxon	description	Description. Male. Body uniformly dark, rufo-piceus; scutellum rufous in central part; antennae, legs and ventral side brown to reddish brown; covered with testaceous, semi-recumbent pubescence, rather dense on ventral parts, antennomeres with long and dense pubescence on ventral half. Body length 11.2 mm, width 3.4 mm. Head in outer margin of eyes as wide as anterior margin of pronotum, covered with dense, umbilicate punctures with interspaces narrower than their own diameter; frons pentagonal, declining anteriorly, flat between eyes, anterior margin thickened, prominent at middle, rather triangular and thicker in apex; antennae (Figure 4 c) exceeding the apex of posterior angles by less than apical antennomere, antennomeres serrated from fourth to tenth, first antennomere thick, curved and cylindrical, as long as both second and third combined, two latter antennomeres subconical, third 1.3 × longer than second, fourth antennomere 1.3 × longer than both second and third combined and slightly (1.1 times) longer than all serrated antennomeres. Pronotum 1.3 × broader than length, moderately convex, widest behind middle, narrowing anteriorly from middle, very weakly sinuate in posterior third near base of posterior angles; the latter very weakly divergent with two divergent carinae (Figure 4 b) (external and internal) on each angle, external carina almost as long as or slightly longer than internal carina, both exceeding the base of angle and reaching to posterior third of lateral side; punctures on disc coarse, umbilicate with intervals mainly narrower than their diameters, denser and rather ovalshape on lateral sides; pronotosternal suture gaping in anterior third; prosternal lobe (collar) short and not covering mouthparts, broadly rounded in anterior margin, the latter thickened and prominent; prosternum rather parallel sided, covering with umbilicated double punctures, larger punctures dense, with interspace narrower than their own diameter, disperse in anterior and lateral portions, smaller punctures with interspace wider than their own diameter, dispersed mainly in posterior and central parts; hypomeron covered with dense and oval-shape punctures dispersed densely except in posterior fourth; prosternal process (Figures 4 d, e) (0.7 mm long), exceeding posteriorly as long as diameter of procoxa, roundly dilated between procoxae, rather parallel sided, with a longitudinal carina ventrally, carina smoothly concave and sharply bent at its end, process becoming narrower from apical third and sharply rounded at apex. Scutellum almost 1.4 × longer than wide, shield-shaped, wider apically and narrowing posteriorly, rather convex in anterior margin and rounded in posterior end, flat, punctures coarse and pubescent. Elytra 3.1 × longer than pronotum and as wide as pronotum at its widest point; lateral margins sub-parallel at anterior half and widest at anterior quarter; base slightly narrower than width of pronotum at apices of hind angles, margin of base to shoulders thickened and slightly prominent anteriorly; ninth stria densely and deeply punctured, interstriae with sparse irregular punctures, distinctly smaller than punctures of the stria, also with short irregular and transversal linear impression. Tarsomeres simple, first to fourth decreasing in length, apical tarsomere almost two times longer than penultimate and as long as two-thirds of first tarsomere. Aedeagus as in figures 4 f and 4 g, length 1.2 mm, elongate; median lobe narrowing anteriorly and pointed at apex, exceeding apices of parameres by more than one-eight of its length. Female. unknown	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFE09760FF47FD44E063A4E6.taxon	etymology	Etymology. The epithet refers to the morphology of the prosternal process that is slim, rather parallel sided with smoothly concave carina in comparison with other similar species with oval sided processes and more concave carinae. Habitat. In a valley of mountainous region covered by Oak forest and near a village with old and large walnut trees.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFE09760FF47FD44E063A4E6.taxon	discussion	Remarks. The validity of the genus Reitterelater has been discussed by different authors (Chassain 1992; Delnatte et al. 2011; Prosvirov 2017 b). Further biological studies or systematic analysis may clarify this taxonomic ambiguity. The genus is new to Iran. Climate types. Sub-humid, cold winters and warm summers.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFE59767FF47FF14E07DA6C9.taxon	description	(Fig. 8, Map 3)	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFE59767FF47FF14E07DA6C9.taxon	materials_examined	Material examined (HMIM). Ardabil province: 3 exs.: “ ARDABIL, Moghan, 10 km. Pârsâbâd, 100 m., 23 - 24. V. 1997, [leg.] Badii / Safz. / Naz. V ”. Climate types. Semi-arid, cool winters and warm summers.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFE49766FF47FEECE049A57D.taxon	description	(Fig. 9, Map 5)	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFE49766FF47FEECE049A57D.taxon	materials_examined	Material examined (HMIM). Azarbaijan-e Sharghi province: 7 exs.: Arasbaran Protected Area, Kaleybar, Asheghlu, Marzrud to Kalan Ranger Station rd., 3 Km W Kalan Ranger Station, N 38 ° 46 ’ 07.5 ”, E 46 ° 49 ’ 35.7 ”, 2600 m, 25. VI. 2015, leg. H. Nasserzadeh. Climate types. Semi-arid, cold winters and mild summers.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFE79765FF47FD8CE07FA471.taxon	description	(Fig. 10, Map 7)	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFE79765FF47FD8CE07FA471.taxon	materials_examined	Material examined (HMIM). Ardabil province: “ AZARBAIDJAN [Ardabil prov.], Nour-Lake [Neor lake], 2363 - 2450 [m], 21.6.1973, [leg.] Ragabi [Rajabi] ”. Climate types. Sub-humid, cold winters and mild summers.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFE79765FF47FF14E07FA6C9.taxon	materials_examined	Material examined (HMIM). Ardabil province: “ ARDABIL, Heyrân, [N 38 ° 25 E 48 ° 33], 1600 m, 18. VI. 2002, [leg.] Deuve / Montreuil / Mof. / Hâj. ”. Climate types. Sub-humid, cold winters and mild summers.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFE69764FF47FEECE778A5A9.taxon	materials_examined	Material examined (HMIM). Ardabil province: 8 exs.: Moghan Protected Area, Meshkin-Shahr to Parsabad rd., ca. 7 km N Gharehghaj, N 39 ° 08 ’ 38.8 ”, E 47 ° 38 ’ 43.3 ” E, 453 m, 25. V. 2017, leg. H. Nasserzadeh; 1 ex.: Agh-Dagh Protected Area, Hashjin, Kivi-Zavieh, N 37 ° 21 ’ 14.8 ”, E 48 ° 24 ’ 55.4 ”, 1167 m, 1. VII. 2017, leg. H. Nasserzadeh & M. Mojdehi. Azarbaijan-e Sharghi province: 15 exs.: Arasbaran Protected Area, Jananlu to Kaleybar rd., ca. 28 km S Jananlu, N 38 ° 57 ’ 09.8 ”, E 47 ° 01 ’ 17.3 ”, 756 m, 26. VI. 2017, leg. H. Nasserzadeh. Zanjan province: 12 exs.: Anguran Protected Area, Mahneshan, Behestan to Takaghaj rd., 2 km NW Behestan, nr. Anguran river, N 36 ° 40 ’ 30.3 ”, E 47 ° 43 ’ 15.5 ”, 1318 m, 25. V. 2016, leg. H. Nasserzadeh; 3 exs.: Anguran Protected Area, 18 km NW Dandi, Qareh- Bugh Ranger Station, N 36 ° 38 ’ 16.7 ”, E 47 ° 40 ’ 57.5 ”, 1541 m, 24. V. 2016, leg. H. Nasserzadeh. Climate types. Sub-humid and semi-arid, cold to cool winters and warm summers.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFE69764FF47FCD4E79CA08F.taxon	materials_examined	Material examined (HMIM). Zanjan province: 1 ex.: Anguran Protected Area, Mahneshan, Behestan to Takaghaj rd., 2 km NW Behestan, nr. Anguran river, N 36 ° 40 ’ 30.3 ”, E 47 ° 43 ’ 15.5 ”, 1318 m, 25. V. 2016, leg. H. Nasserzadeh; 9 exs.: Anguran Protected Area, 18 km NW Dandi, Qareh-Bugh Ranger Station, N 36 ° 38 ’ 16.7 ”, E 47 ° 40 ’ 57.5 ”, 1541 m, 24. V. 2016, leg. H. Nasserzadeh. Climate types. Semi-arid, cold winters and warm summers. MAP 7. Distribution map of Pristilophus, Selatosomus and Zorochros species in northwestern Iran.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFE9976BFF47FF5CE07DA601.taxon	materials_examined	Material examined (HMIM). Kordestan province: 4 exs.: Saghez to Marivan rd., nr. Ghomchiyan, N 35 ° 42 ’ 53.4 ”, E 46 ° 22 ’ 45.1 ”, 2080 m, 29. VI. 2015, leg. H. Nasserzadeh. Climate types. Semi-arid, cold winters and warm summers.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFE9976AFF47FD8CE70CA602.taxon	description	(Figs 11 – 12, Map 8)	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFE9976AFF47FD8CE70CA602.taxon	materials_examined	Material examined (HMIM). Ardabil province: 5 exs: “ ARDABIL, Daryâcheh-e Neour [Neor lake], [N 37 ° 59 ’ E 48 ° 33 ’], 2250 m, 1. VII. 1997, [leg.] Bar. / Mof. ”. Azarbaijan-e Gharbi province: 2 exs.: Mirabad Protected Area, Sardasht to Mirabad rd., nr. Nalas, N 36 ° 20 ’ 14.6 ”, E 45 ° 25 ’ 05.8 ”, 1357 m, 27. V. 2016, leg. H. Nasserzadeh. Zanjan province: 1 ex.: Sorkhabad Protected Area, Mari to Golchin rd., ca. 8 km NE Golchin, Solodarreh, N 37 ° 04 ’ 17.5 ”, E 48 ° 27 ’ 36.6 ”, 1898 m, 22. V. 2016, leg. H. Nasserzadeh. Climate types. Sub-humid and semi-arid, cold winters and warm summers.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFE8976AFF47F927E0DAA0B8.taxon	materials_examined	Material examined (HMIM). Azarbaijan-e Gharbi province: 3 exs.: Sardasht, Mirabad to Alootan rd., ca. 3 km W Mirabad, N 36 ° 22 ’ 53.6 ”, E 45 ° 20 ’ 15.6 ”, 1650 m, 27. V. 2016, leg. H. Nasserzadeh. Ardabil province: 5 exs.: “ ARDABIL, Daryâcheh-e Neour [Neor lake], [N 37 ° 59 ’ E 48 ° 33 ’], 2250 m, 1. VII. 1997, [leg.] Bar. / Mof. ”. Climate types. Sub-humid, cold winters and mild to warm summers.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFEB9769FF47FF14E75BA113.taxon	materials_examined	Material examined (HMIM). Ardabil province: 3 exs.: Moghan Protected Area, Sarband to Ardabil rd., ca. 20 Km S Sarband, nr. Alamlu vill., N 39 ° 37 ’ 30.6 ”, E 47 ° 46 ’ 57.8 ”, 58 m, 27. VI. 2017, leg. H. Nasserzadeh & M. Mojdehi (light trap (; 5 exs.: Moghan Protected Area, ca. 1 km W Aslanduz, nr. Aras river, N 39 ° 26 ’ 15.1 ”, E 47 ° 24 ’ 05.9 ”, 182 m, 25. VI. 2017, leg. H. Nasserzadeh & M. Mojdehi (Light trap). Azarbaijan-e Gharbi province: 1 ex.: ca. 35 km N Mahabad, nr. Khurkhureh village, Kaniborazan swamp, N 36 ° 59 ’ 04.6 ”, E 45 ° 45 ’ 48.8 ”, 1328 m, 28. VI. 2015, leg. H. Nasserzadeh (Light trap); 1 ex.: Lake Urmia National Park, Rashakan Research Station, N 37 ° 20 ’ 38.8 ”, E 45 ° 17 ’ 37.4 ”, 1315 m, 18 - 25. VI. 2016, leg. E. Gilasian & M. Parchami-Araghi (Malaise trap (. Climate types. Semi-arid, cold to cool winters and warm summers. MAP 8. Distribution map of Cardiophorus and Craspedostethus species in northwestern Iran.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
03BE8789FFEB9768FF47F8C2E70CA596.taxon	materials_examined	Material examined (HMIM). Azarbaijan-e Gharbi province: 2 exs.: Mirabad Protected Area, Sardasht To Mirabad rd., nr. Nalas, N 36 ° 20 ’ 14.6 ”, E 45 ° 25 ’ 05.8 ”, 1357 m, 27. V. 2016, leg. H. Nasserzadeh. Azarbaijan-e Sharghi province: 20 exs.: Zanjan to Tabriz Highway, 5 km W Azizkandy, N 37 ° 27 ’ 44.9 ”, E 46 ° 55 ’ 14.4 ”, 1621 m, 21. VI. 2015, leg. H. Nasserzadeh. Kordestan province: 39 exs.: Abdol-Razagh Protected Area, Saqez, nr. Ashaab village, N 36 ° 14 ’ 57.1 ”, E 46 ° 46 ’ 19.5 ” E, 1881 m, 29. V. 2016, leg. H. Nasserzadeh; 6 exs.: Saghez, Abdol-Razzagh Protected Area, ca. 1 km S Qaplantu village, nr. Patar mt., N 36 ° 13 ’ 10.7 ”, E 46 ° 42 ’ 37.6 ”, 1866 m, 29. V. 2016, leg. H. Nasserzadeh; 2 exs.: Badr va Parishan Protected Area, Qorveh, Meyhim-e Sofla to Meyhim-e Olia rd., ca. 1 km NW Meyhim-e Sofla, E 35 ° 03 ’ 26.4 ”, E 47 ° 52 ’ 13.9 ”, 2155 m, 31. V. 2016, leg. H. Nasserzadeh; 2 exs.: Badr va Parishan Protected Area, Qorveh, ca. 10 km S Kani-Ganji, nr. Selseleh village, N 35 ° 06 ’ 38.2 ”, E 47 ° 30 ’ 33.4 ”, 2276 m, 31. V. 2016, leg. H. Nasserzadeh; 11 exs.: Saghez, Abdol-Razzagh Protected Area, ca. 3 km S Qaplantu village, N 36 ° 12 ’ 46.2 ”, E 46 ° 42 ’ 55.1 ”, 1864 m, 29. V. 2016, leg. H. Nasserzadeh; 1 ex.: Saqez to Divandarreh rd., 29 km E Saqez, nr. Sonateh, nr. Khorkhoreh river, N 36 ° 10 ’ 38.9 ”, E 46 ° 34 ’ 01.6 ”, 1479 m, 26. V. 2016, leg. H. Nasserzadeh. Zanjan province: 1 ex.: Anguran Protected Area, Mahneshan, Behestan to Takaghaj rd., 2 km NW Behestan, nr. Anguran river, N 36 ° 40 ’ 30.3 ”, E 47 ° 43 ’ 15.5 ”, 1318 m, 25. V. 2016, leg. H. Nasserzadeh; 1 ex.: Sorkhabad Protected Area, Sohrin village, nr. Sohrin Ranger Station, N 36 ° 51 ’ 36.6 ”, E 48 ° 24 ’ 11.9 ”, 1829 m, 22. V. 2016, leg. H. Nasserzadeh; 1 ex.: Anguran Protected Area, Dandi, Duzkand, N 36 ° 31 ’ 15.4 ”, E 47 ° 52 ’ 55.3 ”, 1527 m, 23. V. 2016, leg. H. Nasserzadeh; 2 exs.: Anguran Protected Area, 18 km NW Dandi, Qareh-Bugh Ranger Station, N 36 ° 38 ’ 16.7 ”, E 47 ° 40 ’ 57.5 ”, 1541 m, 24. V. 2016, leg. H. Nasserzadeh; 32 exs.: Anguran Protected Area, 18 km NW Dandi, Qareh-Bugh Ranger Station, N 36 ° 38 ’ 16.7 ”, E 47 ° 40 ’ 57.5 ”, 1541 m, 24. V. 2016, leg. H. Nasserzadeh; 10 exs.: Anguran Protected Area, Dandi, Moheb Ranger Station, N 35 ° 39 ’ 34.9, E 47 ° 40 ’ 14.8 ”, 1713 m, 25. V. 2016, leg. H. Nasserzadeh. Climate types. Sub-humid and semi-arid, cold winters and warm summers.	en	Nasserzadeh, Hiva (2021): A study of the click beetles (Coleoptera: Elateridae) of northwestern Iran with three new species Reitterelater prosternalis sp. n., Melanotus borumandi sp. n. and M. kurdestanicus sp. n. Zootaxa 4941 (3): 338-368, DOI: 10.11646/zootaxa.4941.3.2
