taxonID	type	description	language	source
03BE87970056FFB398B1FB66FCB4C413.taxon	description	Fig. 1 Mortoniella acauda is apparently related to a group of species, including M. albolineata; M. dolonis, n. sp.; M. latispina, n. sp.; and M. teutona. Character similarities for this group are presented in the diagnosis for M. albolineata. All of these species have a dorsal phallic spine with a depressed, apically rounded, somewhat spatulate apex, and an endophallic membrane with distinct membranous lobes. Mortoniella acauda differs diagnostically from all the other species in this group by lacking a ventromesal projection on the fused inferior appendages; additionally, the apicomesal excision of tergum X is distinctly wider than in any of these other species. Like M. latispina and M. teutona, M. acauda lacks scale-like setae on its hind wings. Also like those species, it has a short, curved endophallic spine and elongate paramere appendages that are somewhat widened preapically. Adult. Length of forewing: male 3.2 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 4: 4. Overall color yellowish-brown. Legs yellowish, tibial spurs darker in color, contrasting with legs. Wing bar at anastamosis relatively indistinct, interrupted, marked with whitish setae. Male genitalia. Ventral process of segment VI laterally compressed, ventrally projecting, subtriangular, wide basally, acute apically. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin weakly convexly rounded dorsally, narrowing ventrally; segment deeply excised dorsomesally and ventromesally, forming lateral lobes, lobes separated dorsomesally by much less than 1 / 2 width of segment. Tergum X with very wide, uniformly rounded mesal excision; lateral lobes moderately elongate, apices rounded as viewed dorsally, subacute as viewed laterally. Inferior appendages fused, without ventromesal projection; laterally, on each side, with setose, tapering, dorsally-directed lobes. Mesal pockets of fused inferior appendages with apical processes short, dorsally curved. Paramere appendages elongate, slender, slightly widened in apical 1 / 4 th, apices acute. Dorsal phallic spine, as viewed laterally, with basal part more or less uniform in width, upturned near base, more strongly in apical 1 / 3 rd, apex narrowed and acute; in dorsal view, somewhat widened in middle, apical 1 / 3 rd narrow, apex rounded, spatulate (depressed). Phallicata with sclerotized, anteriorly directed process subtending dorsal phallic spine, and elongate, narrow, projecting lateral processes. Endophallic membrane with membranous dorsal lobe, paralleling dorsal phallic spine, membranous lateral lobes on either side, and sclerotized ventromesal spine; ventromesal spine short, curved, apex acute; phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970056FFB398B1FB66FCB4C413.taxon	materials_examined	Holotype male: BRAZIL: Santa Catarina: Urubici, Cachoeira Avencal, 28 ° 02 ' 50 " S, 049 ° 37 ' 00 " W, 1260 m, 6. iii. 1998, Holzenthal, Froehlich & Paprocki (UMSP 000068088) (pinned) (MZUSP). Etymology. This species is named M. acauda, without a tail, from the Latin word cauda or tail and referring to the absence of a narrow, ventromesal projection from the fused inferior appendages, which characterizes the group of species to which it appears to be most closely related.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970054FFBD98B1FD06FC0AC4E3.taxon	description	Fig. 2 This species is easily distinguished from other members of the M. albolineata subgroup by the distinctive dorsolateral processes of the phallicata, which are very distinctly sclerotized, elongate and arm-like. Other characters, useful in combination for separating it from members of this group include the following: Fused inferior appendages without a distinct ventromesal process; endophallic spine relatively elongate and nearly straight apically; paramere appendages elongate, slightly widened preapically; and tergum X with a V-shaped mesal excision. Specimens from Minas Gerais have the dorsolateral processes of the phallicata more rounded apically (Fig. 2 D) and less angularly bent, as viewed ventrally (Fig. 2 E). The basal part of the tergum X in these specimens also seems to be somewhat more bulbously rounded, as viewed laterally. These differences, while apparently constant for the geographic areas compared, are also relatively minor. There also seems to be some variation in these characters even within populations; we consider this variation to be intraspecific. Adult. Length of forewing: male 3.1 – 4.1 mm, female 3.5 – 4.4 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 4: 4. Overall color dark brown. Legs brown, apices of tarsi whitish, tibial spurs somewhat darker than legs, but not strongly contrasting in color. Antennae with apical part of basal antennal segments whitish. Wing bar at anastamosis relatively indistinct, interrupted, marked with whitish setae, more strongly developed at arculus on anal margin. Male genitalia. Ventral process of segment VI laterally compressed, ventrally projecting, subtriangular, wide basally, rounded to subacute apically. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin forming rounded to slightly angular projection dorsally, narrowing ventrally; segment deeply excised dorsomesally and ventromesally, forming lateral lobes, lobes separated dorsomesally by much less than 1 / 2 width of segment. Tergum X with broad V-shaped mesal excision; lateral lobes moderately elongate, apices subacute. Inferior appendages without ventromesal projection; laterally, on each side, with setose, tapering, dorsally-directed lobes. Mesal pockets of fused inferior appendages with apical processes short, dorsally curved. Paramere appendages elongate, narrow, slightly widened in apical 1 / 3 rd, apices acute. Dorsal phallic spine, as viewed laterally, with basal part more or less uniform in width, strongly upturned in apical 1 / 2, apex narrowed and acute; in dorsal view, slightly widened in middle, apex narrowly acuminate. Phallicata with distinctly sclerotized dorsolateral processes and lightly sclerotized, rounded, apicolateral lobes; dorsolateral processes very elongate, arm-like, ventrally curved, apices rounded or subacute. Endophallic membrane with prominent, sclerotized ventromesal spine; ventral spine elongate, curved at base, apex nearly straight; phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970054FFBD98B1FD06FC0AC4E3.taxon	materials_examined	Holotype male: BRAZIL: Rio de Janeiro: Rio Macaé, Macaé de Cima, 22 ° 23 ' 41 " S, 042 ° 30 ' 08 " W, 1000 m, 8. iii. 2002, Holzenthal, Blahnik, Paprocki, & Prather (UMSP 000085777) (pinned) (MZUSP). Paratypes: BRAZIL: Minas Gerais: Parque Estadual do Itacolomi, trib. to Rio Belchior, 20 ° 25 ' 18 " S, 043 ° 25 ' 42 " W, 700 m, 6. xi. 2001, Holzenthal, Amarante, Blahnik, & Paprocki — 1 male, 3 females (alcohol) (MZUSP); Corrego das Aguas Pretas & tribs., ca. 15 km S Aiuruoca, 22 ° 03 ' 42 " S, 044 ° 38 ' 14 " W, 1386 m, 21. xi. 2001, Holzenthal, Blahnik, Neto, & Paprocki — 1 male, 42 females (pinned) (UMSP); Parque Estadual de São Gonçalo do Rio Preto, Córrego das Eguas, 18 ° 08 ' 43 " S, 043 ° 22 ' 09 " W, 891 m, 14. x. 2000, Paprocki, Amarante & Isaac — 1 male, 3 females (pinned) (UMSP); Estação Ecológica do Tripuí, Córrego Tripuí, 20 ° 23 ' 22 " S, 043 ° 32 ' 32 " W, 1070 m, 21. ii. 1999, Paprocki, Braga & Amarante — 4 males, 40 females (pinned) (UMSP); Cachoeira do Abacaxi, Vale do Tropeiro, 20 ° 12 ' 16 " S, 043 ° 38 ' 10 " W, 1120 m, 7. xi. 2001, Holzenthal, Amarante, Blahnik & Paprocki — 1 male, 1 female (alcohol) (UMSP); Córrego da Serra de Ouro Fino, Vale do Tropeiro, 20 ° 12 ' 22 " S, 043 ° 38 ' 35 " W, 1000 m, 8. x. 2000, Paprocki, Salgado & Isaac — 1 male (alcohol) (UMSP); Rio de Janeiro: same locality and date as holotype — 63 males, 127 females (pinned), 180 males, 304 females (alcohol) (MZUSP, UMSP, NMNH); Rio das Flores, Macaé de Cima, 10 km SE Mury, 1000 m, 9. iii. 2002, Holzenthal, Blahnik, Paprocki & Prather — 75 males, 81 females (pinned), 26 males, 52 females (alcohol) (UMSP); Rio Bengalas, Hotel Bucsky, Nova Friburgo, 22 ° 18 ' 49 " S, 042 ° 31 ' 09 " W, 895 m, Paprocki — 47 males, 67 females (alcohol) (UMSP); Nova Friburgo, 22 ° 16 ' 00 " S, 042 ° 31 ' 59 " W, 950 m, 20. iv. 1977, C & O Flint — 1 male, 9 females (pinned), 5 males, 26 females (alcohol) (NMNH); Nova Friburgo, mun. water supply, 22 ° 16 ' 00 " S, 042 ° 31 ' 59 " W, 950 m, 24. iv. 1977, C & O. Flint — 1 male, 13 females (pinned) (NMNH); Encontro dos Rios (Macaé / Bonito), 6 km S Lumiar, 22 ° 23 ' 29 " S, 042 ° 18 ' 42 " W, 600 m, 10. iii. 2002, Holzenthal, Blahnik, Paprocki, & Prather — 7 males, 9 females (pinned) 35 males (alcohol) (UMSP). Etymology. This species is named M. agosta from the Greek word agostos, a bent arm, and referring to the elongate, bent, dorsolateral projections of the phallicata in this species.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE8797005AFFBF98B1FC93FD2EC517.taxon	description	Figs. 3, 38	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE8797005AFFBF98B1FC93FD2EC517.taxon	materials_examined	Mortoniella albolineata belongs to a group of closely related species, including M. dolonis, n. sp.; M. latispina, n. sp.; and M. teutona. All of these species have the character combination of inferior appendages with a distinct, asymmetrical ventromesal process, dorsal phallic spine with a depressed, apically rounded, somewhat spatulate apex, and endophallic membrane with membranous lobes. Despite the name given to the type for this group, none of these species has a particularly distinct wing bar, although a small white spot is present at the arculus on the forewing. Perhaps the name given by Ulmer refers to the more or less hyaline crossveins present at the anastamosis, evident in denuded or alcohol preserved material, a general feature within the genus. In contrast to M. latispina and M. teutona, males of both M. albolineata and M. dolonis have hind wings densely covered with modified, somewhat flattened and scale-like setae; however, this character could escape casual attention. In M. albolineata, the setae of the hind wing are more distinctly scale-like than in M. dolonis. In other characters, M. albolineata is also similar to M. dolonis; in both species the ventral endophallic spine is elongate and prominent, and both species also have paramere appendages that are uniformly narrow throughout their length, as well as distinctly separated, paired lateral lobes on the endophallic membrane (in addition to a dorsal lobe paralleling the dorsal phallic spine). The lateral lobes on the endophallic membrane in the other 2 species are either unitary or more or less contiguous. In both M. teutona and M. latispina the endophallic spine is relatively short and curved and the paramere appendages are somewhat widened preapically. Mortoniella albolineata differs from M. dolonis in a number of details, the most diagnostic of which is that the paramere appendages are much more elongate. Other differences include an endophallic spine that is strongly curved, rather than nearly straight, and dorsolateral processes on the phallicata that are narrower, almost peg-like. The notched or bifid apex of the apicomesal process of the inferior appendages of M. albolineata is a character featured in the illustration of the lectotype by Flint (1966) and was also observed in the specimens examined. It may be diagnostic for this species. Adult. Length of forewing: male 3.4 – 4.4 mm, female 3.7 – 4.6 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 4: 4. Overall color, of male, grayish-brown, of female, brown. Legs brown, apices of tarsi whitish, tibial spurs somewhat darker than legs, but not strongly contrasting in color. Wing bar at anastamosis scarcely evident, marked with whitish setae at arculus on anal margin. Setae of forewing dense, decumbent, forewing of female with erect setae along major veins, that of male with slightly widened scale-like setae. Hind wing of male covered with modified scale-like setae. Male genitalia. Ventral process of segment VI laterally compressed, ventrally projecting, short, wide basally, rounded apically. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin forming rounded to slightly angular projection in dorsal 1 / 2, narrowing ventrally; segment deeply excised dorsomesally and ventromesally, forming lateral lobes, lobes separated dorsomesally by much less than 1 / 2 width of segment. Tergum X with broad V-shaped mesal excision; lateral lobes weakly developed, apices subacute. Inferior appendages with prominent and somewhat asymmetrically developed ventromesal projection, apex of projection notched, as viewed laterally; laterally, on each side, with setose, tapering, dorsally-directed lobes. Mesal pockets of fused inferior appendages with apical processes short, dorsally curved. Paramere appendages elongate, narrow, uniform in width, apices acute. Dorsal phallic spine, as viewed laterally, more or less uniform in width, gradually upturned in apical 1 / 2, apex narrowed and acute; in dorsal view, somewhat widened in middle, apex rounded, spatulate (depressed). Phallicata with short peg-like dorsolateral processes. Endophallic membrane with membranous dorsal lobe, paralleling dorsal phallic spine, 2 membranous lateral lobes on either side, and sclerotized ventromesal spine; ventral spine stout, elongate, curved, apex acute; phallotremal spines absent. Material examined. BRAZIL: Santa Catarina: Riberão Gaspar, Belchior Alto, Gaspar, 26 ° 48 ' 22 " S, 049 ° 02 ' 28 " W, 120 m, 27. xi. 2003, Holzenthal, Paprocki & Calor — 1 male, 7 females (pinned) (UMSP); Mun. Ilhota, Morro do Bau, 3. xii. 1975 — 4 males, 10 females (alcohol) (NMNH); São Paulo: Parque Estadual Intervales, Rio do Carmo, 24 ° 18 ' 59 " S, 048 ° 25 ' 15 " W, 560 m, 29. ix. 2002, Blahnik, Prather, Melo & Calor — 1 male (pinned) (UMSP). Distribution. Argentina, Brazil, Uruguay.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970058FFB998B1FC05FF44C438.taxon	description	Fig. 4 Mortoniella asymmetris is most similar to, and undoubtedly most closely related to M. truncata, n. sp. Both species are characterized by having paramere appendages asymmetrically developed and differing in length, the left one shorter than the right. Other character similarities include upright processes bordering the paramere appendages that emerge from the ventral margin of the phallicata, and mesal pocket-like structures of the inferior appendages with very elongate, sinuous, spine-like apical processes. The latter structures appear to be fused or semi-fused mesally. Mortoniella asymmetris is diagnostically distinguished from M. truncata by having a tergum X, as viewed dorsally, with acute, rather than truncate, apicolateral projections, and also by having the ventral spine of the endophallic membrane modified into a structure with minute apical spines. Both of these species seem to be closely related to the group of species discussed under M. unota, agreeing in having a dorsal phallic spine sharply upturned apically, with a ventral deflection at the point of inflection, and with the spine, in dorsal view, distinctly widened at the point of inflection and very narrow basally and apically. In M. asymmetris the lateral margins of the dorsal phallic spine are distinctively upturned at the inflection point for the spine. Adult. Length of forewing: male 2.5 – 2.7 mm, female 2.6 – 3.4 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 3: 4. Overall color, in alcohol, light brown. Tibial spurs somewhat darker than legs, contrasting in color. Wing bar at anastamosis not evident (in alcohol). Male genitalia. Ventral process of segment VI laterally compressed, ventrally projecting, subtriangular, wide basally, acute apically. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin weakly, convexly rounded dorsally, narrowing ventrally; segment deeply excised dorsomesally and ventromesally, forming lateral lobes, lobes separated dorsomesally by much less than 1 / 2 width of segment. Tergum X with U-shaped mesal excision, extending less than 1 / 2 length of segment, and projecting lateral lobes; lateral lobes with apices acute to subacute, as viewed both laterally and dorsally. Inferior appendages without ventromesal projection; laterally, on each side, with setose, tapering, dorsally-directed lobes, lobes very narrowed dorsally. Mesal pockets of fused inferior appendages with apical processes elongate, sinuous, more or less fused mesally. Paramere appendages uniformly narrow, asymmetrically developed in length and orientation, the left short and upturned, the right long and downturned. Dorsal phallic spine, as viewed laterally, with lateral margins broadened and somewhat upturned, apical 1 / 3 rd sharply upturned, with distinct sinuous deflection on ventral margin before point of upturn; in dorsal view, very distinctly widened in middle, apical part abruptly narrowed, apex bluntly rounded, subacute. Phallicata with sclerotized, posteriorly-directed dorsomesal process articulating with dorsal phallic spine, and paired, upturned, lightly sclerotized processes arising from basoventral margin; dorsolateral processes only suggestively developed, apparently absent. Endophallic membrane with membranous apical lobe bearing minute spines and modified ventral spine; ventral spine of endophallic membrane somewhat rod-like, lightly sclerotized basally, apically slightly curved and with minute spines; phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970058FFB998B1FC05FF44C438.taxon	materials_examined	Holotype male: PARAGUAY: Amambay: Cerro Cora, Río Aquidaban, 29. xi. 1973, Flint, O S (UMSP 000124876) (alcohol) (NMNH). Paratypes: PARAGUAY: Amambay: same locality and date as holotype — 6 males, 13 females (alcohol) (NMNH, UMSP); Concepción: Concepción, 23 ° 25 ' 00 " S, 057 ° 17 ' 00 " W, 26. viii. 1989, Kochalka, J — 1 male (alcohol) (NMNH). Etymology. This species is named M. asymmetris for the asymmetrically developed paramere appendages of the male.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE8797005EFFB998B1FDE5FEE4C199.taxon	description	Fig. 5 Mortoniella crescentis is diagnostically distinguished from any other species of Mortoniella by the structure of the male dorsal phallic spine, which has its apex laterally compressed, but widened and crescentic in structure, as viewed laterally. The extent of development varies in the material available and is especially strongly developed in the specimens from Teresopolis (Fig. 5 D). Despite, its unique appearance, M. crescentis is somewhat similar in structure to both M. hystricosa, n. sp., and M. parauna, n. sp., both of which also have similar dorsolateral processes of the phallicata, relatively short paramere appendages, a tergum X with lateral lobes relatively narrowly separated, and a strongly upturned phallic spine. Like M. hystricosa, M. crescentis has a distinct, asymmetrical mesal process on the fused inferior appendages, lacking in M. parauna. Adult. Length of forewing: male 3.8 - 4.8 mm, female 4.3 - 5.0 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 4: 4. Overall color very dark brown. Legs same color, apices of tarsi whitish, tibial spurs somewhat darker than legs, but not strongly contrasting in color. Wing bar at anastamosis distinctly marked with white, contrasting setae, bar narrow, but more or less continuous. Male genitalia. Ventral process of segment VI laterally compressed, ventrally projecting, subtriangular, wide basally, acute apically (Fig. 5 E). Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin convexly rounded, narrowing ventrally; segment deeply excised dorsomesally and ventromesally, forming lateral lobes, lobes separated dorsomesally by much less than 1 / 2 width of segment. Tergum X with narrow, V-shaped mesal excision and projecting lateral lobes; lateral lobes with apices subacute, as viewed dorsally, bluntly rounded, as viewed laterally. Inferior appendages with prominent and somewhat asymmetrically developed mesal projection; laterally, on each side, with setose, tapering, dorsally-directed lobes. Mesal pockets of fused inferior appendages with apical processes short, dorsally curved. Paramere appendages of moderate length, as viewed laterally, straight, narrow, uniform in width, apices acute; in dorsal view, paramere appendages slightly curved. Dorsal phallic spine, as viewed laterally, of distinctive shape, relatively stout, apical 1 / 3 rd sharply upturned, apex laterally compressed and crescentic in shape. Phallicata with short, rounded, depressed dorsolateral processes, emerging from upturned, sclerotized, dorsal margin of phallicata. Endophallic membrane relatively simple in structure, with sclerotized ventromesal spine; ventral spine short, curved, apex acute; phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE8797005EFFB998B1FDE5FEE4C199.taxon	materials_examined	Holotype male: BRAZIL: Rio de Janeiro: Rio Campo Belo, trail to Veu da Noiva, 22 ° 25 ' 42 " S, 044 ° 37 ' 10 " W, 1310 m, 24. xi. 2001, Holzenthal, Blahnik, Neto & Paprocki (UMSP 000081785) (pinned) (MZUSP). Paratypes: BRAZIL: Rio de Janeiro: same locality and date as holotype — 2 females (pinned) (UMSP); Parque Nacional do Itatiaia, Rio Campo Belo, 22 ° 27 ' 02 " S, 044 ° 36 ' 49 " W, 1300 m, 23. xi. 2001, Holzenthal, Blahnik, Neto, & Paprocki — 5 males, 1 female (pinned), 1 male (alcohol) (UMSP, MZUSP); same locality, 7. iii. 2002, Holzenthal, Blahnik, Paprocki & Prather — 5 males, 1 female (pinned) (UMSP, NMNH); Teresopolis, 18 km S, Km 17, 1180 m, 18 - 19. iv. 1977, C & O Flint — 7 males, 1 female (pinned) (NMNH). Etymology. This species is named M. crescentis for the crescentic apical development of the dorsal phallic spine of the male.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE8797005CFFBB98B1FF1EFE4DC0F0.taxon	description	Fig. 6 As discussed under M. albolineata, Mortoniella dolonis belongs to a group of closely related species, including M. albolineata; M. latispina, n. sp.; and M. teutona. All of these species have the character combination of inferior appendages with a distinct, asymmetrical mesal process, dorsal phallic spine with a depressed, apically rounded, somewhat spatulate apex, and endophallic membrane with membranous lobes. Mortoniella dolonis is most similar to M. albolineata; in both species the ventral endophallic spine is elongate and prominent, and both species also have paramere appendages that are uniformly narrow throughout their length and also distinctly separated, paired lateral lobes on the endophallic membrane (in addition to a lobe paralleling the dorsal phallic spine). Also, in contrast to M. latispina and M. teutona, males of both M. albolineata and M. dolonis have hind wings densely covered with modified, somewhat flattened and scale-like setae. Mortoniella dolonis differs from M. albolineata in several details, the most diagnostic of which are that the paramere appendages are short and curved apically, rather than elongate and straight, and the endophallic spine is nearly straight and dagger-like, rather than strongly curved. An additional difference is that the dorsolateral processes on the phallicata are more prominent. Adult. Length of forewing: male 3.2 – 3.6 mm, female 3.8 – 4.1 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 4: 4. Overall color, of male, grayish-brown, of female, brown. Legs brown, apices of tarsi whitish, tibial spurs somewhat darker than legs, but not strongly contrasting in color. Wing bar at anastamosis scarcely evident, marked with whitish setae at arculus on anal margin. Setae of forewing dense, decumbent; forewing of female with erect setae along major veins, that of male with slightly widened scale-like setae. Hind wing of male covered with modified scale-like setae. Male genitalia. Ventral process of segment VI laterally compressed, ventrally projecting, short, wide basally, rounded apically. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin convexly rounded dorsally, narrowing ventrally; segment deeply excised dorsomesally and ventromesally, forming lateral lobes, lobes separated dorsomesally by much less than 1 / 2 width of segment. Tergum X with shallow V-shaped mesal excision; lateral lobes very weakly developed, apices broadly rounded. Inferior appendages with prominent and somewhat asymmetrically developed ventromesal projection; laterally, on each side, with setose, tapering, dorsally-directed lobes. Mesal pockets of fused inferior appendages with apical processes short, dorsally curved. Paramere appendages short, narrow, uniform in width, upturned apically, apices acute. Dorsal phallic spine, as viewed laterally, more or less uniform in width, gradually upturned in apical 1 / 2, apex narrowed and acute; in dorsal view, somewhat widened in middle, apex rounded, spatulate (depressed). Phallicata with prominent, sclerotized, dorsolateral processes. Endophallic membrane with membranous dorsal lobe, paralleling dorsal phallic spine, 1 pair of membranous dorsolateral lobes, and sclerotized ventromesal spine; ventromesal spine elongate, nearly straight, apex acute.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE8797005CFFBB98B1FF1EFE4DC0F0.taxon	materials_examined	Holotype male: BRAZIL: São Paulo: Pedregulho, Riberão São Pedro, 20 ° 09 ' 07 " S, 047 ° 30 ' 38 " W, 617 m, 16. xi. 2003, Holzenthal, Paprocki & Calor (UMSP 000085343) (pinned) (MZUSP). Paratypes: BRAZIL: Minas Gerais: Córrego da Serra de Ouro Fino, Vale do Tropeiro, 20 ° 12 ' 22 " S, 043 ° 38 ' 35 " W, 1000 m, 8. x. 2000, Paprocki, Salgado & Isaac — 1 male, 1 female (alcohol) (UMSP); São Paulo: same locality and date as holotype — 1 male, 2 females (pinned) (UMSP); 10 males (alcohol) (MZUSP, UMSP, NMNH); Pedregulho, Sitio Bruninho, 20 ° 09 ' 14 " S, 047 ° 30 ' 42 " W, 630 m, 17. xi. 2003, Holzenthal, Paprocki & Calor — 2 males (pinned) (UMSP); Pedregulho, 20 ° 09 ' 07 " S, 047 ° 30 ' 38 " W, 617 m, 6. x. 2000, Paprocki & Froehlich — 1 male, 1 female (pinned) (UMSP). Etymology. This species is named M. dolonis, from the Greek word dolon, a dagger or stiletto, and referring to the nearly straight, dagger-like spine of the endophallic membrane, which helps to distinguish this species from its most closely related congeners.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970041FFA698B1FF1EFABFC2B1.taxon	description	Fig. 7 This species is distinguished from other members of the M. albolineata subgroup by the following diagnostic character set: Fused inferior appendages without distinct ventromesal process; paramere appendages short, narrow throughout, upcurved apically; and tergum X with a narrow, V-shaped mesal excision. Additional characters include a strongly inflected dorsal phallic spine and an endophallic spine that is short and curved. Adult. Length of forewing: male 3.5 – 3.7 mm, female 3.8 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 4: 4. Overall color very dark brown. Legs brown, apices of tarsi whitish, tibial spurs somewhat darker than leg, but not strongly contrasting in color. Antenna with apical part of basal segments whitish. Wing bar at anastamosis distinctly marked with white, contrasting setae, band narrow, but more or less continuous. Male genitalia. Ventral process of segment VI laterally compressed, ventrally projecting, subtriangular, wide basally, acute apically. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin weakly, convexly rounded dorsally, narrowing ventrally; segment deeply excised dorsomesally and ventromesally, forming lateral lobes, lobes separated dorsomesally by much less than 1 / 2 width of segment. Tergum X with narrow V-shaped mesal excision and projecting lateral lobes; lateral lobes tapering, acute, slightly mesally curved. Inferior appendages without ventromesal projection; laterally, on each side, with setose, tapering, dorsally-directed lobes. Mesal pockets of fused inferior appendages with apical processes short, dorsally curved. Paramere appendages of moderate length, narrow, uniform in width, upcurved apically, apices acute. Dorsal phallic spine, as viewed laterally, more or less uniform in width, strongly upturned in apical 1 / 2, apex narrowed and acute in both lateral and dorsal views. Phallicata with short, rounded, sclerotized dorsolateral processes. Endophallic membrane simple, with bulging membranous lateral lobes and ventromesal spine; ventral spine very short, curved, apex acute; phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970041FFA698B1FF1EFABFC2B1.taxon	materials_examined	Holotype male: BRAZIL: São Paulo: Parque Estadual de Campos do Jordão, Rio Galharada, 22 ° 41 ' 40 " S, 045 ° 27 ' 47 " W, 1530 m, 4 - 5. iii. 1996, Holzenthal & Guahyba (UMSP 000001575) (pinned) (MZUSP). Paratypes: BRAZIL: São Paulo: same locality and date as holotype — 1 male, 2 females (pinned) (UMSP). Etymology. This species is named M. guahybae in memory of Rosalys Guahyba, who helped to collect the type specimens and under whose invitation the second author initiated his studies of Trichoptera in Brazil.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970041FFA098B1FB7BFCD3C488.taxon	description	Fig. 8 Mortoniella hystricosa is diagnostically distinguished from any other species of Mortoniella by the structure of its dorsal phallic spine, which has its apex both distinctly spinose and strongly upturned, and also by the structure of tergum X, which is deeply and narrowly incised mesally and has the apices of the lateral lobes truncate in lateral view. Despite, its unique appearance, M. hystricosa is somewhat similar in structure to both M. crescentis, n. sp., and M. parauna, n. sp., both of which have similar dorsolateral processes of the phallicata, relatively short paramere appendages, a tergum X with lateral lobes narrowly separated, and a strongly upturned dorsal phallic spine. Like M. crescentis, but unlike M. parauna, M. hystricosa has a distinct, asymmetrical mesal process on the inferior appendages. Adult. Length of forewing: male 4.0 – 4.4 mm, female 4.6 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 4: 4. Overall color very dark brown. Legs same color, apices of tarsi whitish, tibial spurs somewhat darker than leg, but not strongly contrasting in color. Wing bar at anastamosis distinctly marked with whitish, setae, band narrow, but more or less continuous. Male genitalia. Ventral process of segment VI laterally compressed, ventrally projecting, subtriangular, wide basally, acute apically. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin forming rounded to slightly angular projection in dorsal 1 / 2, narrowing ventrally; segment deeply excised dorsomesally and ventromesally, forming lateral lobes, lobes separated dorsomesally by much less than 1 / 2 width of segment. Tergum X with narrow, deep, V-shaped mesal excision and projecting lateral lobes; lateral lobes with apices acute and slightly mesally curved in dorsal view, wide and broadly truncate in lateral view. Inferior appendages with prominent and somewhat asymmetrically developed ventromesal projection; laterally, on each side, with setose, tapering, dorsally-directed lobes. Mesal pockets of fused inferior appendages with apical processes short, dorsally curved. Paramere appendages moderately elongate, narrow, uniform in width, upcurved apically, apices acute. Dorsal phallic spine, as viewed laterally, relatively stout, broadened in middle, strongly upturned in apical 1 / 2, apex narrowed and acute, with numerous minute spines; in dorsal view gradually broadened at midlength, narrowed apically. Phallicata with short, rounded, sclerotized dorsolateral processes. Endophallic membrane simple, with bulging membranous lateral lobes and ventromesal spine; ventral spine very short, curved, apex acute; phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970041FFA098B1FB7BFCD3C488.taxon	materials_examined	Holotype male: BRAZIL: Santa Catarina: Parque Ecológica Spitzkopf, confl. Rio Ouro & Rio Caeté, 27 ° 00 ' 21 " S, 049 ° 06 ' 42 " W, 140 m, 25. xi. 2003, Holzenthal, Paprocki & Calor (UMSP 000085400) (pinned) (MZUSP). Paratypes: BRAZIL: Santa Catarina: Urubici, Cachoeira Avencal, 28 ° 02 ' 50 " S, 049 ° 37 ' 00 " W, 1260 m, 6. iii. 1998, Holzenthal, Froehlich & Paprocki — 5 males, 1 female (pinned) (UMSP). Etymology. This species is named M. hystricosa from the Latin word hystrix, a porcupine, and referring to the spiny apex of the dorsal phallic spine of the male in this species.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970047FFA098B1FD75FBEBC109.taxon	description	Fig. 9 This is a distinctive species within the M. albolineata subgroup, easily diagnosed by the very elongate apicolateral projections of tergum X, ventrally recurved apex of the dorsal phallic spine, and presence of a ventromesal projection on the inferior appendages. Adult. Length of forewing: male 3.8 – 4.8 mm, female 4.3 – 5.0 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 4: 4. Overall color dark brown. Legs same color, apices of tarsi whitish, tibial spurs somewhat darker than leg, but not strongly contrasting in color. Wing bar at anastamosis distinctly marked with white setae, band narrow, but more or less continuous. Male genitalia. Ventral process of segment VI laterally compressed, ventrally projecting, subtriangular, moderately elongate, wide basally, acute apically (Fig. 9 D). Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin convexly rounded, narrowing ventrally; segment deeply excised dorsomesally and ventromesally, forming lateral lobes, lobes separated dorsomesally by much less than 1 / 2 width of segment. Tergum X with deep, V-shaped mesal excision and projecting apicolateral lobes, basolaterally with short rounded setose lobes; lateral lobes very elongate, tapering, acute apically. Inferior appendages with prominent and somewhat asymmetrically developed ventromesal projection; laterally, on each side, with setose, tapering, dorsally-directed lobes. Mesal pockets of fused inferior appendages with apical processes short, dorsally curved. Paramere appendages short, uniform in width, apices acute and somewhat upturned. Dorsal phallic spine, as viewed laterally, more or less uniform in width, gradually upturned in apical 1 / 2, apex acute and distinctly ventrally recurved. Phallicata rather distinctly sclerotized, with broadly rounded dorsolateral lobes and weakly projecting, rounded ventral lobes; apicodorsally with 2 pairs additional sclerotized lobes, the anterior ones elongate and fingerlike, the posterior ones short and rounded. Endophallic membrane relatively short and simple, with ventromesal spine; ventral spine short, stout, curved, apex acute; phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970047FFA098B1FD75FBEBC109.taxon	materials_examined	Holotype male: BRAZIL: São Paulo: Parque Estadual Intervales, Rio do Carmo, 24 ° 18 ' 59 " S, 048 ° 25 ' 15 " W, 560 m, 29. ix. 2002, Blahnik, Prather, Melo & Calor (UMSP 000088142) (pinned) (MZUSP). Paratypes: BRAZIL: São Paulo: same locality and date as holotype — 10 males, 9 females (pinned) (MZUSP, UMSP, NMNH). Etymology. This species is named M. intervales, with the epithet used as a noun in apposition, for Parque Estadual Intervales, the very beautiful park where the type specimens were collected.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970044FFA398B1FF1EFB20C0B8.taxon	description	Fig. 10 As discussed under M. albolineata, Mortoniella latispina belongs to a group of closely related species, including M. dolonis, n. sp.; M. teutona; and M. albolineata. All of these species have the character combination of inferior appendages with a distinct, asymmetrical mesal process, dorsal phallic spine with a depressed, apically rounded, spatulate apex, and endophallic membrane with membranous lobes. It is most similar to M. teutona; in both species the ventral endophallic spine is relatively short and curved and the paramere appendages are somewhat widened preapically. The endophallic spine is much more prominent in the other 2 species of this group and both of these species have paramere appendages that are uniform in width throughout their length. Mortoniella latispina differs from M. teutona in several details, the most diagnostic of which is a dorsal phallic spine that is distinctly widened at the point of inflection, as viewed dorsally. This also distinguishes M. latispina from M. albolineata and M. dolonis. The major flexion of this spine is also more apical than in M. teutona. Other differences include a slightly wider mesal excavation of tergum X, appearing more U-shaped than V-shaped, and differences in the structure of the membranous lobes of the endophallic membrane. In M. latispina the dorsal lobe, which parallels the dorsal phallic spine, is short, rather than elongate, and the lateral lobe is not subdivided and is also distinctly sclerotized. However, this is only likely to be evident in specimens in which this structure is expanded. Adult. Length of forewing: male 3.3 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 4: 4. Overall color dark brown. Legs brown, apices of tarsi whitish, tibial spurs darker in color, contrasting with legs. Antennae with apical part of basal segments whitish. Wing bar at anastamosis more or less distinctly marked with whitish setae, bar narrow, discontinuous. Male genitalia. Ventral process of segment VI laterally compressed, ventrally projecting, subtriangular, wide basally, rounded to subacute apically. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin weakly, convexly rounded dorsally, narrowing ventrally; segment deeply excised dorsomesally and ventromesally, forming lateral lobes, lobes separated dorsomesally by much less than 1 / 2 width of segment. Tergum X with U-shaped mesal excision; lateral lobes moderately elongate, apices subacute as viewed dorsally and laterally. Inferior appendages with prominent and somewhat asymmetrically developed ventromesal projection; laterally, on each side, with setose, tapering, dorsally-directed lobes. Mesal pockets of fused inferior appendages with apical processes short, slightly dorsally curved. Paramere appendages elongate, narrow, slightly widened in apical 1 / 3 rd, upturned apically, apices acute. Dorsal phallic spine, as viewed laterally, more or less uniform in width, gradually upturned, weakly near base, more strongly in apical 1 / 3 rd, apex narrowed and acute; in dorsal view, very distinctly widened at about 2 / 3 rds length, apical 1 / 3 rd narrow, apex rounded, spatulate (depressed). Phallicata with sclerotized, anteriorly directed process, subtending dorsal phallic spine, and moderately elongate, narrow, projecting lateral processes. Endophallic membrane with short membranous dorsal lobe, subtending dorsal phallic spine, weakly sclerotized, paired dorsolateral lobes, and sclerotized ventromesal spine; ventral spine relatively short, curved, apex acute; phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970044FFA398B1FF1EFB20C0B8.taxon	materials_examined	Holotype male: BRAZIL: Rio de Janeiro: Parque Nacional do Itatiaia, Rio Campo Belo, trail to Veu da Noiva, 22 ° 25 ' 42 " S, 044 ° 37 ' 10 " W, 1310 m, 24. xi. 2001, Holzenthal, Blahnik, Neto & Paprocki (UMSP 000047063) (pinned) (MZUSP). Paratypes: BRAZIL: Rio de Janeiro: same locality and date as holotype — 1 male, 2 females (pinned) (UMSP); Nova Friburgo, 22 ° 16 ' 00 " S, 042 ° 31 ' 59 " W, 950 m, 20. iv. 1977, C & O Flint — 1 male (pinned) (NMNH). Etymology. This species is named M. latispina, Latin for wide spine, in reference to the widened dorsal phallic spine of the male, which helps to distinguish this species from its closely related congeners.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970044FFAD98B1F963FD99C5C3.taxon	description	Fig. 11 This member the M. albolineata subgroup of species is distinguished by the following diagnostic character set: Fused inferior appendages without distinct ventromesal process; endophallic spine very large, prominent, and strongly curved; paramere appendages short, uniform in width; and dorsal phallic spine angularly upturned apically. It is perhaps most similar overall to Mortoniella guahybae, n. sp., but differs particularly in its much more prominent endophallic spine and in that the apicolateral processes of tergum X are less narrowed and the mesal incision of that structure is wider and V-shaped. Adult. Length of forewing: male 3.6 mm, female 3.6 – 4.0 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 4: 4. Overall color medium brown. Legs brown, apices of tarsi whitish, tibial spurs somewhat darker than legs, but not strongly contrasting in color. Antennae with apical part of basal segments whitish. Wing bar at anastamosis more or less distinctly marked with whitish setae, bar narrow, discontinuous. Male genitalia. Ventral process of segment VI laterally compressed, ventrally projecting, subtriangular, wide basally, rounded to subacute apically. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin convexly rounded, narrowing ventrally; segment deeply excised dorsomesally and ventromesally, forming lateral lobes, lobes separated dorsomesally by much less than 1 / 2 width of segment. Tergum X with broad V-shaped mesal excision and weakly projecting lateral lobes; lateral lobes acute apically, as viewed dorsally, short and rounded as viewed laterally. Inferior appendages without ventromesal projection; laterally, on each side, with setose, tapering, dorsally-directed lobes. Mesal pockets of fused inferior appendages with apical processes moderately elongate, curved, distinctly thickened through most of length. Paramere appendages short, narrow, uniform in width, apices acute, appendages somewhat downcurved. Dorsal phallic spine, as viewed laterally, more or less uniform in width, strongly upturned in apical 1 / 3 rd, apex cute. Phallicata with weakly projecting, broadly rounded, sclerotized dorsolateral processes; ventral margin moderately projecting, lightly sclerotized, with rounded apical projection. Endophallic membrane with very elongate, sclerotized spine; spine curved at base, straight apically, apex acute; phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970044FFAD98B1F963FD99C5C3.taxon	materials_examined	Holotype male: BRAZIL: Santa Catarina: Urubici, Rio Canoas, road to Campo dos Padres, 28 ° 00 ' 15 " S, 049 ° 22 ' 24 " W, 1100 m, 7. iii. 1998, Holzenthal, Froehlich & Paprocki (UMSP 000029822) (pinned) (MZUSP). Paratypes: BRAZIL: Santa Catarina: same data as holotype — 25 females (pinned) (MZUSP, UMSP). Etymology. This species is named M. longispina, Latin for long spine, in reference to the very long spine of the endophallic membrane of the male.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE8797004AFFAF98B1FBB6FA38C438.taxon	description	Fig. 12 Mortoniella paraguaiensis is closely related to a group of species including M. paraunota, n. sp., M. unota, and M. uruguaiensis, n. sp. All of these species are characterized by a dorsal phallic spine that, in lateral view, has a sharply upturned, blade-like apex, accompanied by a slight ventral deflection at the point of inflexion. In dorsal view the spine is very narrow apically and distinctly widened at the point of inflection. Other useful characters defining the group include the absence of a ventromesal process on the inferior appendages and a tergum X with sharply pointed apicolateral processes and a relatively narrow mesal incision. All of the species except M. paraunota also have short paramere appendages and inferior appendages with the dorsal apices at least somewhat recurved. Mortoniella paraguaiensis is most similar to M. unota, especially in having the apex of the dorsal phallic spine very sharply upturned and in having a 0: 3: 4 spur formula. It differs in the more robust dorsolateral processes of the phallicata and in having the lateral lobes of the inferior appendages with only a minute recurved apex, rather than a strongly bent one. Adult. Length of forewing: male 2.7 – 2.8 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 3: 4. Overall color medium brown. Legs same color, apices of tarsi whitish, tibial spurs somewhat darker than legs, but not strongly contrasting in color. Wing bar at anastamosis more or less distinctly marked with whitish setae, bar discontinuous Male genitalia. Ventral process of segment VI laterally compressed, ventrally projecting, wide basally, narrowed and subacute apically, length about 2 times width at base. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin convexly rounded, narrowing ventrally; segment deeply excised dorsomesally and ventromesally, forming lateral lobes, lobes separated dorsomesally by much less than 1 / 2 width of segment. Tergum X with somewhat angular, U-shaped mesal excision and distinctly projecting lateral lobes; lateral lobes acute apically, as viewed both dorsally and laterally. Inferior appendages without ventromesal projection; laterally, on each side, with setose, tapering, dorsally-directed lobes, apices of lobes each with short, spine-like, posteriorly angled process. Mesal pockets of fused inferior appendages with apical processes short, dorsally curved. Paramere appendages short, narrow, uniform in width, apices acute, appendages somewhat downcurved. Dorsal phallic spine, as viewed laterally, with apical 1 / 3 rd sharply upturned, apex slightly recurved, spine with distinct sinuous deflection on ventral margin at point of upturn; in dorsal view, very distinctly widened from base to point of inflection, apical part abruptly narrowed, making overall appearance of apex blade-like. Phallicata with strongly sclerotized dorsal process, subtending dorsal phallic spine, and broad, sclerotized, posteriorly directed dorsolateral processes. Endophallic membrane relatively simple, with short, stout, curved, ventromesal spine; phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE8797004AFFAF98B1FBB6FA38C438.taxon	materials_examined	Holotype male: PARAGUAY: Alto Parana: SE Naranja, ca. 20 km S. Pto. Stroessner, 18 - 24. viii. 1988, L E Peña G (UMSP 000118564) (pinned) (NMNH). Paratypes: PARAGUAY: Alto Parana: same locality and date as holotype — 1 male (pinned) (NMNH). Etymology. This species is named M. paraguaiensis for the country of origin of the holotype of this species.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970048FFAF98B1FDE5FD3DC095.taxon	description	Fig. 13 This is a somewhat nondescript species, best diagnosed by the following set of characters considered together: Fused inferior appendages without an apicomesal process; elongate paramere appendages, somewhat widened preapically; and an upturned dorsal phallic spine with an acute apex, as viewed both laterally and dorsally. The lightly sclerotized ventral projection of the phallicata is also probably diagnostic, but may be more difficult to discern, depending on how the specimen has been cleared. Adult. Length of forewing: male 2.5 – 2.9 mm, female 2.5 – 3.3 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 4: 4. Overall color light brown. Legs yellowish, tibial spurs darker in color, contrasting with legs. Wing bar at anastamosis more or less distinctly marked with whitish setae, bar discontinuous. Male genitalia. Ventral process of segment VI laterally compressed, ventrally projecting, subtriangular, wide basally, acute apically. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin weakly, convexly rounded dorsally, narrowing ventrally; segment deeply excised dorsomesally and ventromesally, forming lateral lobes, lobes separated dorsomesally by much less than 1 / 2 width of segment. Tergum X with relatively shallow, U-shaped mesal excision and projecting lateral lobes; lateral lobes moderately elongate, apices acute to subacute as viewed both dorsally and laterally. Inferior appendages without ventromesal projection; laterally, on each side, with setose, dorsally-directed, non-tapering lobes, rounded apically. Mesal pockets of fused inferior appendages with apical processes short, dorsally curved. Paramere appendages elongate, narrow, slightly widened in apical 1 / 3 rd, apices acute. Dorsal phallic spine, as viewed laterally, more or less uniform in width, upturned in apical 1 / 2, apex narrowed and acute in both lateral and dorsal views. Phallicata with sclerotized dorsal process subtending dorsal phallic spine, and, sclerotized, laterally directed dorsolateral processes; ventrally with short, lightly sclerotized mesal projection. Endophallic membrane apparently simple in structure, with sclerotized ventromesal spine; ventral spine moderately elongate, stout, curved, apex acute; phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970048FFAF98B1FDE5FD3DC095.taxon	materials_examined	Holotype male: BRAZIL: Minas Gerais: Rio Paraúna, 3 km S Santana do Riacho, 19 ° 10 ' 59 " S, 043 ° 43 ' 29 " W, 650 m, 16. ii. 1998, Holzenthal & Paprocki (UMSP 000031878) (pinned) (MZUSP). Paratypes: BRAZIL: Minas Gerais: same locality as holotype, 11. xi. 2001, Holzenthal, Amarante, Blahnik, & Paprocki — 5 males, 21 females (alcohol) (UMSP, NMNH). Etymology. This species is named M. parauna, used as a noun in apposition, for Rio Parauna, near which the type specimen for the species was collected.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE8797004DFFAA98B1FF1EFDC7C061.taxon	description	Fig. 14 As discussed under Mortoniella paraguaiensis, n. sp., M. paraunota, n. sp., is closely related to a group of species, including M. paraguaiensis; M. unota; and M. uruguaiensis, n. sp. All of these species are characterized by a dorsal phallic spine that, in lateral view, has a sharply upturned, blade-like apex, accompanied by a slight ventral deflection at the point of inflection. In dorsal view the spine is very narrow apically and distinctly widened at the point of inflection. Mortoniella paraunota differs from these other species in having longer paramere appendages, inferior appendages with dorsal projections whose apices are not at all recurved, and the presence of dorsolateral lobe-like extensions on the endophallic membrane, which serve as guides for the apices of the paramere appendages. The apical inflection of the dorsal phallic spine is most similar to M. uruguaiensis (less strongly recurved than either M. paraguaiensis or M. unota). However, it differs significantly from M. paraunota in the structure of the dorsolateral processes of the phallicata. Adult. Length of forewing: male 2.5 – 2.9 mm; female 2.5 – 3.3 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 4: 4. Overall color light brown. Legs yellowish, tibial spurs darker than legs, contrasting in color. Wing bar at anastamosis more or less distinctly marked with whitish setae, bar discontinuous. Male genitalia. Ventral process of segment VI laterally compressed, ventrally projecting, subtriangular, wide basally, acute apically. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin convexly rounded, narrowing ventrally; segment deeply excised dorsomesally and ventromesally, forming lateral lobes, lobes separated dorsomesally by much less than 1 / 2 width of segment. Tergum X with narrow, U-shaped mesal excision, extending less than 1 / 2 length of segment, and projecting lateral lobes; lateral lobes acute apically, as viewed both dorsally and laterally. Inferior appendages without ventromesal projection; laterally, on each side, with setose, tapering, dorsally-directed lobes, apices of lobes unmodified. Mesal pockets of fused inferior appendages with apical processes short, dorsally curved. Paramere appendages elongate, narrow, uniform in width, apices acute. Dorsal phallic spine, as viewed laterally, with apical 1 / 3 rd gradually upturned, apex posteriorly directed, spine with distinct sinuous deflection on ventral margin at point of upturn; in dorsal view, very distinctly widened at inflection, apical part abruptly narrowed, making overall appearance of apex blade-like. Phallicata with sclerotized, subtruncate dorsal process, subtending dorsal phallic spine, somewhat projecting laterally; laterally with weakly projecting, broadly rounded, depressed projections on either side, subtending paramere appendages. Endophallic membrane (or dorsal extension of phallicata), with lightly sclerotized, depressed projections, projecting over apices of paramere appendages, and short, stout, curved, ventromesal spine; phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE8797004DFFAA98B1FF1EFDC7C061.taxon	materials_examined	Holotype male: BRAZIL: Santa Catarina: Seara (Nova Teutônia), 27 ° 11 ' S, 052 ° 23 ' W, 300 - 500 m, 10. x. 1964, F Plaumann (UMSP 000118566) (pinned) (MZUSP). Paratypes: ARGENTINA: Entre Rios: Ao. Piray Mini W, Dos Hermanas, 33 ° 10 ' 00 " S, 058 ° 53 ' 59 " W, 23. xi. 1973, O S Flint — 4 males, 17 females (alcohol) (NMNH, UMSP); BRAZIL: Santa Catarina: same locality as holotype, ix. 1964, F Plaumann — 1 male (pinned) (NMNH); same locality, 12. x. 1964, F Plaumann — 1 male (pinned) (NMNH). Etymology. This species is named M. paraunota, from the Greek word para, meaning near or by, and referring to the similarity of this species to M. unota.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE8797004DFF9598B1F92BFD2EC7A7.taxon	description	Fig. 15	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE8797004DFF9598B1F92BFD2EC7A7.taxon	description	Adult. Length of forewing: male 2.6 – 3.8 mm, female 3.3 – 4.8 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 4: 4. Overall color medium brown. Legs brown, apices of tarsi whitish, tibial spurs somewhat darker than legs, but not strongly contrasting in color. Antennae with apical part of basal segments whitish. Wing bar at anastamosis indistinct, marked with whitish setae on anal margin. Male genitalia. Ventral process of segment VI laterally compressed, ventrally projecting, short, wide basally, rounded apically. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin convexly rounded dorsally, narrowing ventrally; segment deeply excised dorsomesally and ventromesally, forming lateral lobes, lobes separated dorsomesally by much less than 1 / 2 width of segment. Tergum X with broad, V-shaped mesal excision and weakly projecting lateral lobes; lateral lobes subtriangular, broad basally, apices subacute as viewed both dorsally and laterally. Inferior appendages with prominent and somewhat asymmetrically developed ventromesal projection; laterally, on each side, with setose, tapering, dorsally-directed lobes. Mesal pockets of fused inferior appendages with apical processes short, dorsally curved. Paramere appendages elongate, narrow, slightly widened in apical 1 / 3 rd, apices acute. Dorsal phallic spine, as viewed laterally, more or less uniform in width, gradually upturned in apical 1 / 2, apex narrowed and acute; in dorsal view, slightly widened in middle, apex rounded, spatulate (depressed). Phallicata with sclerotized, anteriorly directed process, subtending dorsal phallic spine, and moderately elongate, narrow, projecting lateral processes. Endophallic membrane with membranous dorsal lobe, subtending dorsal phallic spine, membranous, divided or subcontiguous lateral lobes on either side, and sclerotized ventromesal spine; ventral spine relatively short, curved, apex acute; phallotremal spines absent Material examined. BRAZIL: Parana: Rio Mãe Catira, 10 km N Porto de Cima, 25 ° 21 ' 49 " S, 048 ° 52 ' 28 " W, 200 m, 8 - 9. xii. 1997, Holzenthal & Huisman — 1 male, 5 females (pinned) (UMSP), 2 males, 6 females (pinned) (MZUSP); Rio Jacarei, ca 5 km S BR 277, 25 ° 33 ' 56 " S, 048 ° 42 ' 10 " W, 80 m, 9. xi. 1997, Holzenthal & Huisman — 4 males, 5 females (pinned) (UMSP); Rio De Janeiro: Rio Sousa, in Cachoeiras de Macacú, 22 ° 26 ' 34 " S, 042 ° 37 ' 57 " W, 150 m, 16. iii. 1996, Holzenthal, Rochetti & Oliveira — 13 males, 52 females (pinned); 120 males, 627 females (alcohol) (UMSP; MZUSP); Parque Nacional da Serra dos Orgãos, Trilha das Ruínas, Guapimirim, 22 ° 29 ' 41 " S, 042 ° 59 ' 44 " W, 940 m, 28. ii. 2002, Blahnik & Paprocki — 5 males, 5 females (alcohol) (UMSP); Encontro dos Rios (Macaé / Bonito), 6 km S Lumiar, 22 ° 23 ' 29 " S, 042 ° 18 ' 42 " W, 600 m, 10. iii. 2002, Holzenthal, Blahnik, Paprocki & Prather — 13 males, 11 females (pinned), 118 males (alcohol) (UMSP; MZUSP); Parati, Riacho PerequÍ-açu, Sitio Cachoeira Grande, 23 ° 13 ' 14 " S, 044 ° 47 ' 24 " W, 120 m, 25. ix. 2002, Blahnik, Prather, Melo, Froehlich & Silva — 37 males, 59 females (pinned), 17 males, 57 females (alcohol) (UMSP); Parati, trib. to Riacho PerequÍ-açu, 23 ° 12 ' 50 " S, 044 ° 47 ' 29 " W, 190 m, 26. ix. 2002, Blahnik, Prather, Melo, Froehlich & Silva — 27 males (alcohol), (UMSP); Parati, Riacho PerequÍ-açu, 23 ° 13 ' 27 " S, 044 ° 46 ' 09 " W, 30 m, 24. ix. 2002, Blahnik, Prather, Melo, Froehlich & Silva — 8 males, 72 females (pinned), 11 males, 42 females (alcohol) (UMSP; MZUSP); Santa Catarina: Parque Ecológica Spitzkopf, confl. Rio Ouro & Rio Caeté, 27 ° 00 ' 21 " S, 049 ° 06 ' 42 " W, 140 m, 3. iii. 1998, Holzenthal, Froehlich & Paprocki — 16 males, 14 females (pinned), 13 males, 34 females (alcohol), (UMSP); same locality, 25. xi. 2003, Holzenthal, Paprocki & Calor — 3 males, 14 females (pinned), 27 males, 54 females (alcohol), (UMSP); Rio Caeté above 1 st falls, 27 ° 00 ' 21 " S, 049 ° 06 ' 42 " W, 170 m, 4. iii. 1998, Holzenthal, Froehlich & Paprocki — 5 males, 5 females (pinned), 4 males, 5 females (alcohol) (UMSP); Riberão Gaspar, Belchio Alto, Gaspar, 26 ° 48 ' 22 " S, 049 ° 02 ' 28 " W, 120 m, 27. xi. 2003, Holzenthal, Paprocki & Calor — 3 males, 1 female pinned) (UMSP); Nova Teutonia, 27 ° 11 ' S, 052 ° 23 ' W, 300 - 500 m, ix. 1963, F Plaumann — 48 males (alcohol) (NMNH); São Paulo: Pedregulho, Riberão São Pedro, 20 ° 09 ' 07 " S, 047 ° 30 ' 38 " W, 617 m, 16. xi. 2003, Holzenthal, Paprocki & Calor — 29 males, 1 female (alcohol) (UMSP); 11 km SE Bananal, small stream on São Paulo Route 247, 22 ° 45 ' 41 " S, 044 ° 23 ' 11 " W, 675 m, 23. ix. 2002, Blahnik, Prather, Melo, Froehlich & Silva — 5 males, 15 females (pinned) (UMSP); Altinópolis, Cachoeira Dos Macacos, 20 ° 55 ' 23 " S, 047 ° 22 ' 45 " W, 759 m, 18. xi. 2003, Holzenthal, Paprocki & Calor — 1 male (alcohol) (UMSP); Altinópolis, Fazenda São João da Mata, Rio Baguassu, 21 ° 00 ' 35 " S, 047 ° 28 ' 54 " W, 745 m, 19 - 21. xi. 2003, Holzenthal, Paprocki & Calor — 8 males, 14 females (pinned), 164 males, 114 females (alcohol) (UMSP; MZUSP). Distribution. Argentina, Brazil, Uruguay.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970072FF9598B1FE55FB50C12D.taxon	description	Fig. 16 Mortoniella truncata is most closely related to M. asymmetris, n. sp. Both species are characterized by having paramere appendages asymmetrically developed and differing in length, the left one shorter than the right. Other character similarities include upright processes bordering the paramere appendages than emerge near the ventral margin of the phallicata, and mesal pocket-like structures of the inferior appendages with very elongate, sinuous, spine-like apical processes which appear to be fused or semi-fused mesally. Mortoniella truncata is most easily and diagnostically distinguished from M. asymmetris by having a tergum X with truncate, rather than acute, apicolateral projections. As discussed in the diagnosis for M. asymmetris, both of these species seem to be closely related to the group of species discussed under M. unota, agreeing in having a dorsal phallic spine that is sharply upturned apically, with a slight, but distinct, ventral deflection at the point of inflection, and with the spine, in dorsal view, distinctly widened at the inflection point and narrow apically. Adult. Length of forewing: male 2.2 – 2.4 mm, female 2.3 – 2.5 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 3: 4. Overall color, in alcohol, medium brown. Legs brown, apices of tarsi whitish, tibial spurs somewhat darker than legs, but not strongly contrasting in color. Wing bar at anastamosis indistinct (in alcohol), marked with whitish setae on anal margin. Male genitalia. Ventral process of segment VI laterally compressed, somewhat posteriorly directed, subtriangular, wide basally, acute apically. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin weakly, convexly rounded, narrowing ventrally; segment deeply excised dorsomesally and ventromesally, forming lateral lobes, lobes separated dorsomesally by much less than 1 / 2 width of segment. Tergum X with narrow, U-shaped mesal excision, extending about 1 / 2 length of segment, and projecting lateral lobes; lateral lobes with apices broadly truncate as viewed dorsally, subacute as viewed laterally. Inferior appendages without ventromesal projection; laterally, on each side, with setose, tapering, dorsally-directed lobes. Mesal pockets of fused inferior appendages with apical processes moderately elongate, sinuous, more or less fused mesally. Paramere appendages uniformly narrow, asymmetrically developed in length and orientation, the left short and downturned, the right long and nearly straight. Dorsal phallic spine, as viewed laterally, sharply upturned in apical 1 / 3 rd, with distinct sinuous deflection on ventral margin at point of upturn; in dorsal view, very distinctly widened in middle, apical part abruptly narrowed, apex acute. Phallicata with weakly sclerotized dorsomesal process and paired, upturned, lightly sclerotized processes arising from basoventral margin; laterally with asymmetrical, lightly sclerotized areas, paralleling paramere appendages. Endophallic membrane somewhat globular in shape, simple in structure, without apparent ventromesal spine; phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970072FF9598B1FE55FB50C12D.taxon	materials_examined	Holotype male: BRAZIL: Minas Gerais: spring trib to Rio Macauba, near Pandeiros, 15 ° 28 ' 38 " S, 044 ° 44 ' 38 " W, 525 m, 17. xi. 2001, Paprocki & Blahnik (UMSP 000208501) (alcohol) (MZUSP). Paratypes: BRAZIL: Minas Gerais: same locality and date as holotype — 3 males, 17 females (alcohol) (UMSP). Etymology. This species is named M. truncata for the truncate apices of tergum X of the male.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970077FF9098B1FF1EFD9CC39B.taxon	description	Fig. 17	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970077FF9098B1FF1EFD9CC39B.taxon	description	Adult. Length of forewing: male 2.8 – 3.0 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 3: 4. Overall color (in alcohol) yellowish-brown. Wing bar at anastamosis not evident (in alcohol). Male genitalia. Ventral process of segment VI laterally compressed, ventrally projecting, subtriangular, wide basally, acute apically. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin weakly, convexly rounded dorsally, narrowing ventrally; segment deeply excised dorsomesally and ventromesally, forming lateral lobes, lobes separated dorsomesally by much less than 1 / 2 width of segment. Tergum X with narrow, U-shaped mesal excision, extending less than 1 / 2 length of segment, and distinctly projecting lateral lobes; lateral lobes acute apically, as viewed both dorsally and laterally. Inferior appendages without ventromesal projection, but mesal margin somewhat produced and rounded; laterally, on each side, with setose, tapering, dorsally-directed lobes, apices of lobes strongly posteriorly bent, apex acute. Mesal pockets of fused inferior appendages with apical processes moderately elongate, posteriorly curved. Paramere appendages short, narrow, nearly uniform in width, apices acute, appendages somewhat downcurved. Dorsal phallic spine, as viewed laterally, with apical 1 / 3 rd strongly upturned, spine with distinct sinuous deflection on ventral margin at point of upturn; in dorsal view, very distinctly widened at point of inflection, apical part strongly narrowed, apex acute, making overall appearance of apex somewhat blade-like. Phallicata with mesally fused, ovately rounded dorsolateral processes. Endophallic membrane apparently simple in structure, with stout, curved, ventromesal spine; phallotremal spines absent Material examined. ARGENTINA: Entre Rios: Ao. Piray Mini W, Dos Hermanas, 33 ° 10 ' 00 " S, 058 ° 53 ' 59 " W, 23. xi. 1973, O S Flint — 1 male (alcohol) (NMNH); BRAZIL: Santa Catarina: Seara (Nova Teutônia), 27 ° 11 ' S, 052 ° 23 ' W, 300 - 500 m, x-xi. 1971, F Plaumann - 1 male, 3 females (alcohol) (NMNH). Distribution. Argentina, Brazil.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970077FF9298B1F98EFA85C5E8.taxon	description	Fig. 18 As discussed in the diagnosis for Mortoniella paraguaiensis, n. sp., M. uruguaiensis, n. sp., is closely related to a group of species, including M. paraguaiensis; M. paraunota, n. sp.; and M. unota. All of these species are characterized by a dorsal phallic spine that, in lateral view, has a sharply upturned, blade-like apex, accompanied by a slight ventral deflection at the point of inflection. In dorsal view the spine is very narrow apically and distinctly widened at the point of inflection. All of the species except M. paraunota also have short paramere appendages and inferior appendages with the apices at least somewhat recurved. Mortoniella uruguaiensis is diagnostically characterized by the shape of the dorsolateral processes of the phallicata, which are very elongate and somewhat elevated basally. It resembles M. paraunota in that the apical inflection of the dorsal phallic spine is less pronounced than in the other 2 species of this group. It differs in the shorter paramere appendages, and in having the apex of the inferior appendages forming a short, but very strongly bent projection. Adult. Length of forewing: male 2.4 - 2.9 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 4: 4. Overall color (in alcohol) yellowish-brown. Wing bar at anastamosis not evident (in alcohol). Male genitalia. Ventral process of segment VI laterally compressed, ventrally projecting, subtriangular, wide basally, acute apically. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin convexly rounded dorsally, narrowing ventrally; segment deeply excised dorsomesally and ventromesally, forming lateral lobes, lobes separated dorsomesally by much less than 1 / 2 width of segment. Tergum X with narrow, U-shaped mesal excision and distinctly projecting lateral lobes; lateral lobes acute apically, as viewed both dorsally and laterally. Inferior appendages without ventromesal projection, but mesal margin weakly, angularly produced; laterally, on each side, with setose, tapering, dorsally-directed lobes, apices of lobes with short, angularly recurved projection, apex acute. Mesal pockets of fused inferior appendages with apical processes short, dorsally curved. Paramere appendages very short, narrow, uniform in width, apices acute, appendages somewhat downcurved. Dorsal phallic spine, as viewed laterally, upturned in apical 1 / 2, spine with distinct sinuous deflection on ventral margin at point of upturn; in dorsal view, slightly widened from base to point of inflection, apical part abruptly narrowed, making overall appearance of apex blade-like. Phallicata with mesally fused dorsolateral processes; dorsolateral processes distinctly sclerotized, elongate, narrow, posteriorly curved. Endophallic membrane simple, with short, stout, curved, ventromesal spine; phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970077FF9298B1F98EFA85C5E8.taxon	materials_examined	Holotype male: URUGUAY: Artigas: San Gregorio, 30 ° 33 ' S, 057 ° 52 ' W, Carbonell, Mesa, & San Martin (UMSP 000124875) (alcohol) (NMNH). Paratypes: BRAZIL: Santa Catarina: Nova Teutonia, 27 ° 11 ' S, 052 ° 23 ' W, 300 - 500 m, xi. 1963, F Plaumann — 2 males (alcohol) (NMNH, MZUSP); URUGUAY: Artigas: same locality and date as holotype — 4 males (alcohol) (NMNH, UMSP). Etymology. This species is named M. uruguaiensis for the country of origin of the holotype specimen.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970075FF9C98B1F9ECFD8EC550.taxon	description	Fig. 19	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970075FF9C98B1F9ECFD8EC550.taxon	description	Adult. Length of forewing: male 2.5 – 2.9 mm, female 2.6 – 3.5 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 3: 4. Overall color, in alcohol, yellowish-brown. Wing bar at anastamosis not evident (in alcohol). Male genitalia. Ventral process of segment VI laterally compressed, somewhat posteriorly directed, subtriangular, wide basally, acute apically. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin forming rounded to slightly angular projection in dorsal 1 / 2, narrowing ventrally; segment deeply excised dorsomesally and ventromesally, forming lateral lobes, lobes separated dorsomesally by much less than 1 / 2 width of segment. Tergum X with U-shaped mesal excision, extending less than 1 / 2 length of segment, and projecting lateral lobes; lateral lobes, as viewed dorsally, with apical projections subtriangular, narrowed and acute, extreme apices incurved, as viewed laterally, projecting and subacute; lobes somewhat depressed. Inferior appendages with very short, acute ventromesal projection; laterally, on each side, with basally rounded, posteriorly directed lobes; lobes moderately elongate, tapering, outwardly curved, apices acute. Mesal pockets of fused inferior appendages with apical processes relatively short, posteriorly curved. Paramere appendages bulbous basally, strongly dorsally curved near base, stout at inflection, tapering to acute apex. Dorsal phallic spine, as viewed laterally, somewhat inflated in middle, strongly dorsally curved at about apical 1 / 4 th, with projecting, rounded lobe at inflection, apex acute; in dorsal view, very distinctly widened in middle, apical part narrowed, acute (apex laterally compressed). Phallicata with elongate narrow dorsolateral processes, basal part of each process anteriorly directed and curved around base of paramere appendage, apical part posteriorly directed, apex acute; phallicata laterally with lightly sclerotized, broadly rounded, strongly depressed projections. Endophallic membrane with apex forming projecting, rounded, laterally compressed lobe; ventromesal spine absent; phallotremal spines short, acute, anterolaterally directed Material examined. ARGENTINA: Salta: Río Pescado, W Oran, 14. x. 1973, OS Flint — 2 males, 118 females (Paratypes, alcohol) (NMNH). Distribution. Argentina, Bolivia.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE8797007BFF9E98B1FA2FFD25C253.taxon	description	Fig. 20 This is a distinctive species, similar only to M. pusilla, n. sp. Both species have a distinctively shaped phallic spine, very broad in the middle as viewed laterally, and sharply upturned apically. Mortoniella pumila is most readily distinguished from M. pusilla by its longer paramere appendages. These are distinctly longer than the dorsal phallic spine in M. pumila, bowed upward, and distinctly shorter than the dorsal phallic spine in M. pusilla. Other differences include a shorter apicolateral, seta-bearing process on tergum X, a less pronounced mesal notch of tergum X, a dorsal process of the phallotheca that is more upright, and a longer and distinctly acute spine on the endophallic membrane. The dorsal phallic spine in M. pumila is also more distinctly upturned and its apical part narrower and more acuminate than M. pusilla (dorsal view). The latter character may be somewhat variable, but was especially evident in the site where the 2 species were found sympatrically. Adult. Length of forewing: male 2.0 – 2.8 mm, female 2.3 – 2.9 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 3: 4. Overall color light brown. Legs light brown, apices of tarsi whitish, tibial spurs somewhat darker than legs, but not strongly contrasting in color. Antennae with apical part of basal segments whitish. Wing bar at anastamosis distinct, marked with whitish setae. Male genitalia. Ventral process of segment VI laterally compressed, ventrally projecting, subtriangular, length greater than width at base, acute to subacute apically. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin convexly rounded, narrowing ventrally; segment deeply excised dorsomesally and ventromesally, forming lateral lobes, lobes separated dorsomesally by much less than 1 / 2 width of segment. Tergum X with mesal margin very shallowly, concavely incised, lateral lobes only weakly developed, apices broadly rounded in dorsal view, each with single, prominent ventrolateral seta. Inferior appendages without apicomesal projection; laterally, on each side, with short, setose, dorsally-directed lobes, apices of lobes subacute. Mesal pockets of fused inferior appendages with apical processes prominent, posteriorly curved, distinctly projecting beyond inferior appendages. Paramere appendages elongate (as long or longer than dorsal phallic spine), narrow, mesally curved, slightly widened preapically, apices acute; in dorsal view strongly mesally curved preapically. Dorsal phallic spine, as viewed laterally, distinctly upturned in apical 1 / 2, apex narrowed and acute, spine very distinctly, bulbously enlarged in middle. Phallicata short, tubular, with prominent, sclerotized, raised, anteriorlydirected, dorsomesal process subtending dorsal phallic spine; ventral margin weakly sclerotized, somewhat protruding. Endophallic membrane apparently short and simple in structure, with short, stout, curved ventromesal spine; phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE8797007BFF9E98B1FA2FFD25C253.taxon	materials_examined	Holotype male: BRAZIL: Rio de Janeiro: Encontro dos Rios (Macaé / Bonito), 6 km S Lumiar, 22 ° 23 ' 29 " S, 042 ° 18 ' 42 " W, 600 m, 10. iii. 2002, Holzenthal, Blahnik, Paprocki & Prather (UMSP 000088069) (pinned) (MZUSP). Paratypes: BRAZIL: Minas Gerais: Parque Estadual de São Gonçalo do Rio Preto, Córrego das Eguas, 18 ° 08 ' 43 " S, 043 ° 22 ' 09 " W, 891 m, 14. x. 2000, Paprocki, Amarante & Isaac — 1 male (alcohol) (UMSP); Rio Santo Antônio, downstream from Morro do Pilar, 19 ° 08 ' 08 " S, 043 ° 21 ' 15 " W, 530 m, 17. x. 2000, Paprocki & Ferreira — 18 males, 74 females (alcohol) (UMSP); Corrego Pitanga in Braúnas, 19 ° 03 ' 25 " S, 042 ° 43 ' 05 " W, 353 m, 18. x. 2000, Paprocki & Ferreira — 1 male, 3 females (UMSP); Corrego Pitanga, upstream of confl. with Rio Santo Antônio, 19 ° 05 ' 40 " S, 042 ° 39 ' 54 " W, 238 m, 19. x. 2000, Paprocki & Ferreira — 15 males (alcohol) (UMSP); Rio de Janeiro: same locality and date as holotype — 1 male, 5 females (pinned) 23 males, 54 females (alcohol) (MZUSP, UMSP, NMNH). Etymology. This species is named M. pumila from the Latin word pumilis, meaning diminutive or little, and referring to the diminutive size of this species.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970079FF9898B1FB46FE1BC4C0.taxon	description	Fig. 21 This is a distinctive and diminutive species, similar only to M. pumila. Character similarities are discussed in the diagnosis for M. pumila. Mortoniella pusilla is most readily distinguished from M. pusilla by its shorter paramere appendages. These are distinctly shorter than the dorsal phallic spine in M. pusilla. Other differences include a much longer apicolateral, seta-bearing process on tergum X, a more pronounced mesal notch of tergum X, dorsal processes of the phallotheca that are more flattened and anvil-like, and an apically rounded or short angular, as opposed to more prominent and apically acute spine on the endophallic membrane. The dorsal phallic spine in M. pusilla is also less distinctly upturned and its apical part wider, as viewed dorsally. Adult. Length of forewing: male 2.4 – 2.8 mm, female 2.5 – 3.0 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 3: 4. Overall color light brown. Legs light brown, apices of tarsi whitish, tibial spurs somewhat darker than legs, but not strongly contrasting in color. Antennae with apical part of basal segments whitish. Wing bar at anastamosis distinct, marked with whitish setae. Male genitalia. Ventral process of segment VI laterally compressed, ventrally projecting, subtriangular, length greater than width at base, acute to subacute apically. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin convexly rounded, narrowing ventrally; segment deeply excised dorsomesally and ventromesally, forming lateral lobes, lobes separated dorsomesally by much less than 1 / 2 width of segment. Tergum X with mesal margin shallowly, concavely incised, lateral lobes only weakly developed, apices truncately rounded in dorsal view, each with single, prominent ventrolateral seta on posteriorly curved, finger-like protrusion. Inferior appendages without apicomesal projection; laterally, on each side, with short, setose, dorsallydirected lobes, apices of lobes subacute. Mesal pockets of fused inferior appendages with apical processes prominent, posteriorly curved, projecting beyond inferior appendages. Paramere appendages relatively short (shorter than dorsal phallic spine), narrow, posteriorly-directed. Dorsal phallic spine, as viewed laterally, distinctly upturned in apical 1 / 2, apex narrowed and acute, spine very distinctly, bulbously enlarged in middle. Phallicata short, tubular, with prominent, sclerotized, raised and flattened, anteriorly-directed dorsomesal process subtending dorsal phallic spine; ventral margin weakly sclerotized, distinctly projecting. Endophallic membrane apparently short and simple in structure, with very short, curved, bluntly rounded or subacute ventromesal sclerite; phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970079FF9898B1FB46FE1BC4C0.taxon	materials_examined	Holotype male: BRAZIL: Minas Gerais: Corrego Pitanga, upstream of confl. with Rio Santo Antônio, 19 ° 05 ' 40 " S, 042 ° 39 ' 54 " W, 238 m, 19. x. 2000, Paprocki & Ferreira — (UMSP 000208515) (pinned, wings mounted, body in glycerin) (MZUSP). Paratypes: BRAZIL: Minas Gerais: same locality and date as holotype — 22 males (alcohol) (UMSP); Parque Estadual de São Gonçalo do Rio Preto, Rio Preto, 18 ° 07 ' 50 " S, 043 ° 20 ' 15 " W, 791 m, 12. x. 2000, Paprocki, Amarante & Salgado — 36 males, 576 females (alcohol) (UMSP, NMNH, MZUSP); Rio Mainarte, bridge on Cibrão road, 20 ° 27 ' 15 " S, 043 ° 24 ' 06 " W, 700 m, 18. viii. 1998. Paprocki & Amarante — 1 male, 1 female (pinned) (UMSP). Etymology. This species is named M. pusilla from a Latin word for very small or little, referring to the diminutive size of this species.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE8797007FFF9A98B1FB47FD8EC438.taxon	description	Fig. 22	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE8797007FFF9A98B1FB47FD8EC438.taxon	description	Adult. Length of forewing: male 4.2 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 4: 4. Overall color medium brown. Tibial spurs somewhat darker than legs, contrasting in color. Wing bar at anastamosis more or less distinctly marked with whitish setae, bar discontinuous. Male genitalia. Ventral process of segment VI laterally compressed, somewhat posteriorly directed, relatively wide basally, narrowed and subacute apically, length about 2 times width at base (Fig. 22 E). Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin convexly rounded in dorsal 1 / 2, narrowing ventrally; segment deeply excised dorsomesally and ventromesally, forming lateral lobes, lobes separated dorsomesally by much less than 1 / 2 width of segment. Tergum X convexly rounded mesally, with short, projecting dorsolateral lobes and rounded ventrolateral lobes; dorsolateral lobes, as viewed dorsally, somewhat mesally curved, narrow basally, subacute apically. Inferior appendages short, without distinct ventromesal projection, lateral lobes rounded dorsally, not forming tapering dorsal projections. Mesal pockets of fused inferior appendages with apical processes relatively short, posteriorly curved. Paramere appendages short, narrow, uniform in width, acute apically, each mesally turned so that appendages from opposite sides cross over one another mesally. Dorsal phallic spine, as viewed laterally, somewhat widened in middle, rather gradually upturned from base, more strongly in apical 1 / 4 th, tapering apically; in dorsal view, uniform in width in basal part, somewhat inflated before apical inflection, apex weakly bifid, apices acute. Phallicata with elongate narrow, sinuous dorsolateral processes, each curved over corresponding paramere appendage; laterally with broad, subtruncate, dorsallydirected lobes, each paralleling assemblage formed by paramere appendage and dorsolateral lobe of phallicata. Endophallic membrane apparently short and simple, without ventromesal spine; phallotremal spines absent. Material examined. ARGENTINA: Tucuman: Rt. 307, La Angostura, 11. x. 1973, OS Flint — 1 male (paratype, pinned) (NMNH). Distribution. Argentina, Bolivia.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE8797007DFF8498B1FB0FFC16C438.taxon	description	Fig. 23 From other members of the M. ormina species group, this species is notably distinguished by the structure of its dorsal phallic spine, which has pronounced T-shaped apicolateral expansions and an apex that is upturned and posteriorly recurved. The long narrow paramere appendages are also distinctive as compared to other species described from this region. Among described species, M. alicula is undoubtedly most closely related to M. ormina (Mosely, 1939), which also has the apex of the dorsal phallic spine similarly narrowed and posteriorly bent. Diagnostic differences include the much longer paramere appendages of M. alicula, as well as the more greatly widened dorsal phallic spine and the different armature of the phallicata. Adult. Length of forewing: male 2.1 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 3: 4 (preapical spur of mesotibia very small). Overall color medium brown. Legs yellowish, tibial spurs darker in color, contrasting with legs. Wing bar at anastamosis marked with white, contrasting setae, apices of forewings with small white setal spots at apices of veins. Male genitalia. Ventral process of segment VI posteriorly projecting, elongate, narrow, length more than 2 times width at base; apex dorsoventrally compressed, acute as viewed laterally (Fig. 23 E), rounded as viewed ventrally. Segment IX rounded anterolaterally, length greatest in ventral 1 / 2, posterolateral margin nearly linear; segment deeply excised dorsomesally and ventromesally, forming lateral lobes, lobes separated dorsomesally by more than 1 / 2 width of segment. Tergum X elongate, simple in structure, with very deep, narrow U-shaped mesal excision; apical lobes broadly rounded. Inferior appendages short, inconspicuous, without dorsal or ventromesal projections, more or less fused ventromesally. Mesal pockets of inferior appendages with prominent, posteriorlydirected, spine-like projections, apparently fused to ventral margin of inferior appendages. Paramere appendages very narrow, elongate, slightly widened preapically, apices acute; appendages with broad U-shaped curvature, curved downward at base and upward at apex. Dorsal phallic spine of very distinctive structure, laterally with rounded, depressed projections; apically, as viewed dorsally, greatly widened, with T-shaped lateral projections and narrow, acute, strongly posteriorly curved dorsomesal projection. Phallicata with 2 pairs of relatively short spinelike processes, dorsally with mesally-directed, attenuate processes, crossing each other mesally, laterally with shorter, laterally-directed processes. Base of endophallic membrane (or apex of phallicata) with rounded, membranous (or lightly sclerotized) lateral projections, apex of endophallic membrane with paired, rounded, membranous projections, each with needle-like, sclerotized phallotremal spine, mesally with additional short spine (possibly modified apex of phallotremal sclerite). Phallotremal sclerite evident as small rounded sclerite.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE8797007DFF8498B1FB0FFC16C438.taxon	materials_examined	Holotype male: BRAZIL: Rio de Janeiro: Rio Macaé, Macaé de Cima, 22 ° 23 ' 41 " S, 042 ° 30 ' 08 " W, 1000 m, 8. iii. 2002, Holzenthal, Blahnik, Paprocki & Prather (UMSP 000087906) (pinned) (MZUSP). Etymology. This species is named M. alicula, from a Latin word for wing and referring to the wing-like lateral expansions of the apical part of the dorsal phallic spine of this species.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970063FF8498B1FDE5FE1AC005.taxon	description	Fig. 24	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970063FF8498B1FDE5FE1AC005.taxon	description	Adult. Length of forewing: male 2.0 mm. Venation difficult to discern; forewing (probably) with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 3: 4. Specimen in alcohol and totally cleared, color not evident. Male genitalia: Ventral process of segment VI posteriorly projecting, elongate, narrow, length more than 2 times width at base; apex dorsoventrally compressed, acute as viewed laterally (Fig. 24 E), rounded as viewed ventrally. Segment IX rounded anterolaterally, length greatest in ventral 1 / 2, posterolateral margin nearly linear; segment deeply excised dorsomesally and ventromesally, forming lateral lobes, lobes separated dorsomesally by about 1 / 2 width of segment. Tergum X elongate, simple in structure, with V-shaped mesal excision extending about 1 / 2 length of segment; apical lobes acute. Inferior appendages inconspicuous, subtriangular as viewed laterally, each with acute apicoventral projection and short dorsal lobe; posteromesal margin convex. Mesal pockets of fused inferior appendages with apical processes elongate, narrow, sinuous, posteriorly directed, extending beyond inferior appendages. Paramere appendages (apparently) composed of 3 pairs of elongate, acute processes (lateral 2 possibly from base of phallicata). Dorsal phallic spine of distinctive structure, laterally with wing-like, depressed, lateral projections, basal margin of each wing rounded, apical margins acute; apical 1 / 3 rd of dorsal phallic spine sharply upturned, apex acute. Phallicata lightly sclerotized, without obvious spines. Endophallic membrane bulbous, with lightly sclerotized, paired apicodorsal projections (probably phallotremal spines). Material examined. BRAZIL: Santa Catarina: i. 1963, F Plaumann — 1 male (holotype, in alcohol, USNM type 72739) (NMNH). Distribution. Brazil	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970061FF8098B1FF1EFDC4C733.taxon	description	Fig. 25	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970061FF8098B1FF1EFDC4C733.taxon	materials_examined	The illustrated specimen is from Chile. Although differing from the illustration of the type in several points, the essential features correspond and the differences seem to represent variable features. Diagnostic features are found in the strongly upturned dorsal phallic spine with a deep apicomesal incision and the apically incised lateral margins of tergum X. Although the apex of the dorsal phallic spine characteristically has the margin and mesal surface with small spines, these may be absent on the margin or completely absent. This is variable within the same population in the specimens recorded from Argentina. They are present in the specimens from Chile. A characteristic feature of this species is the outward curved spines at the base of the phallicata, which emerge from an enlarged, sclerotized collar. In the specimens examined, these spines are not as thick as those illustrated for the holotype. However, the development of the basal collar and the length of the spines also seem to be variable; the spines were even absent or vestigial in 1 of the specimens examined from Chile. What we have interpreted as paramere appendages are very lightly sclerotized, short, spine-like processes, which appear to be vestigial. These vary in length in the material examined; they are easily overlooked and it is possible that they may be absent in some specimens; they were not illustrated in the holotype specimen. The last notable difference of the specimens examined from the holotype are in the apical lobes of the endophallic membrane; there are 2 such lobes in the material examined, with sclerotized apical spines (probably phallotremal spines), and 4 prominent lobes with apical spines in the holotype. We have observed lobes similar to those of the holotype in some specimens from Bolivia (which, however, completely lack small apical spines on the dorsal phallic spine). In as much as this is a relatively minor feature, and only easily observed in specimens in which the endophallic membrane is expanded, we are considering this to be variation. All of the specimens we examined (for which the features could be ascertained) had a posteriorly directed dorsomesal spine from the posterior margin of the phallicata (or base of the endophallic membrane), which apparently articulates with the cleft apex of the dorsal phallic spine, and also lightly sclerotized (or nearly membranous) balloon-like projections from the lateral margin of the endophallic membrane. These were not featured or described for the holotype; they are also relatively minor features that may not have been present on the holotype specimen, or may not have been evident. Although we believe the variation described here can be accommodated in 1 variable species, we acknowledge that the species status of these populations may need to be reassessed when more material is available. Adult. Length of forewing: male 2.1 - 2.5 mm, female 2.5 - 3.0. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 3: 4. Overall color medium brown, legs slightly paler, tibial spurs darker, but not strongly contrasting with legs. Wing bar at anastamosis marked with white, contrasting setae. Males with modified scale-like setae on forewing, scales not quite extending to apex. Male genitalia: Ventral process of segment VI posteriorly projecting, elongate, narrow, length more than 2 times width at base; apex slightly dorsoventrally compressed, acute as viewed laterally, subacute as viewed ventrally. Segment IX rounded anterolaterally, length generally greatest in ventral 1 / 2, posterolateral margin nearly linear; segment deeply excised dorsomesally and ventromesally, forming lateral lobes, lobes separated dorsomesally by 1 / 2 or more width of segment. Tergum X moderately elongate, relatively simple in structure, with V-shaped mesal excision; apical lobes with rounded lateral excision, producing acute dorsal and ventral projections, as viewed laterally. Inferior appendages very small, inconspicuous, with acute posterior projection. Mesal pockets of fused inferior appendages with apical processes short, dorsally curved, extending beyond inferior appendages. Paramere appendages hyaline, very narrow, apices acute, variable in length and possibly sometimes absent. Dorsal phallic spine of distinctive structure, laterally with narrow, depressed projections, apically strongly upturned in apical 1 / 3 rd; apex somewhat inflated and deeply incised mesally, producing paired lobes, each with minute spines on apical and mesal margin (minute spines absent in some specimens or populations). Phallicata with base strongly sclerotized and somewhat dorsally produced, forming basal collar; collar laterally with outwardly curved spine-like process on each side, apices of spines often somewhat swollen, with very small scale-like projections. Endophallic membrane (or apex of phallicata) with dorsomesal, posteriorly-oriented spine-like process; laterally, at midlength with lightly sclerotized or nearly membranous balloon-like projections; apicodorsally with either 2 or 4 lobes, each with small, sclerotized spine-like apex (probably phallotremal spines). Material examined. ARGENTINA: Neuquen: Río Agrio, N. Zapala, 9 - 11. xii. 1983, L E Peña G. — 4 males, 8 females (pinned), 6 males, 29 females (alcohol) (NMNH, UMSP); CHILE: Curico: El Coigo, 1. iii. 1968, Flint & Peña — 2 males (alcohol) (NMNH). Distribution. Argentina, Bolivia, Chile.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970067FF8098B1FEE6FE12C368.taxon	description	Fig. 26	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970067FF8098B1FEE6FE12C368.taxon	description	This species most closely resembles M. alicula, n. sp., particularly in the structure of its dorsal phallic spine, which has it apex distinctively narrowed, upturned and posteriorly curved. However, among other differences, in M. ormina, the phallic spine is not nearly so widely explanate preapically and the paramere appendages are much shorter. Adult. Length of forewing: male 1.8 - 2.0 mm, female 2.1 - 2.5. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 3: 4. Overall color (in alcohol) pale yellowish-brown, appendages pale, without pigmentation. Wing bar at anastamosis evident in some specimens. Male genitalia: Ventral process of segment VI posteriorly projecting, elongate, narrow, length more than 2 times width at base, apex acute (Fig. 26 D). Segment IX rounded anterolaterally, length greatest in ventral 1 / 2, posterolateral margin weakly, convexly rounded in dorsal 1 / 2, narrowing ventrally; segment deeply excised dorsomesally and ventromesally, forming lateral lobes, lobes separated dorsomesally by 1 / 2 or more width of segment. Tergum X moderately elongate, simple in structure, with very deep, narrow, V-shaped mesal excision; apical lobes rounded, as viewed both laterally and dorsally. Inferior appendages very small, inconspicuous, with acute posterior projection. Mesal pockets of fused inferior appendages with apical processes short, posterodorsally curved, extending beyond inferior appendages. Paramere appendages moderate in length, extending about as far as dorsal phallic spine, uniformly narrow in width, acute apically. Dorsal phallic spine of distinctive structure, somewhat widened at middle, strongly dorsally curved in apical 1 / 4 th, apex abruptly narrowed and posteriorly recurved. Phallicata bulbous, lightly sclerotized, dorsally with mesal, posteriorly-directed, spine-like process. Endophallic membrane short, simple in structure, with lightly sclerotized, rounded to subquadrate lateral processes; ventral spine and phallotremal spines absent. Material examined. BRAZIL: Santa Catarina: Nova Teutonia, 27 ° 03 ' S, 052 ° 24 ' W, i. 1964, F Plaumann — 12 males, 35 females (alcohol) (NMNH, UMSP). Distribution. Brazil.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970065FF8298B1FEE6FDF8C3B5.taxon	description	Figs. 27, 32 Like the other 2 new species of the M. velasquezi group from Brazil described here, this species lacks scales on the hind wings. It is perhaps most likely to be confused with M. tripuiensis, n. sp., since both species lack the elongate and diagnostic endophallic spines found in M. froehlichi, n. sp. However, M. bocaina does have a pair of short, apically rounded, endophallic spines, not found in M. tripuiensis, which can help to diagnose it. Adult. Length of forewing: male 2.9 – 3.3, female 3.0 – 3.2 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 3: 4. Overall color dark brown. Legs brown, tibial spurs somewhat darker in color, but not strongly contrasting with legs. Wing bar at anastamosis marked with white, contrasting setae. Male genitalia. Ventral process of segment VI posteriorly projecting, elongate, narrow, length more than 2 times width at base (Fig. 27 E). Segment IX moderately rounded anterolaterally, length greatest in ventral 1 / 2 (or near middle), posterolateral margin nearly linear; segment deeply excised dorsomesally and ventromesally, forming lateral lobes, lobes separated dorsomesally by about 1 / 2 width of segment. Tergum X moderate in length, simple in structure, with deep, broad, U-shaped mesal excision; apical lobes, as viewed laterally, distinctly projecting, rounded to subacute apically. Inferior appendages almost completely fused to phallic ensemble, basally with short, setose, anteriorly-directed lobes, apically apparently fused to phallicata and endophallic membrane, forming membranous or very lightly sclerotized lobes. Mesal pockets of fused inferior appendages very bulbously enlarged, with elongate, spine-like, posteriorly-directed apicoventral projections. Paramere appendages very narrow, elongate, about as long as dorsal phallic spine. Phallotheca very short; ventral rod-like appendages elongate, with widely flared apices. Dorsal phallic spine, as viewed dorsally, broad in middle, gradually narrowed and rounded apically; as viewed laterally, relatively narrow, sharply upturned at about apical 1 / 3 rd. Phallicata membranous, continuous with endophallic membrane. Endophallic membrane simple in structure, with membranous apical and dorsal lobes, laterally with short, sclerotized, ventrally curved lobes.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970065FF8298B1FEE6FDF8C3B5.taxon	materials_examined	Holotype male: BRAZIL: São Paulo: Parque Nacional da Serra da Bocaina, Cachoeira dos Posses, 22 ° 46 ' 26 " S, 044 ° 36 ' 15 " W, 1250 m, 3. iii. 2002, Holzenthal, Blahnik, Paprocki, & Prather (UMSP 000069696) (pinned) (MZUSP). Paratypes: BRAZIL: São Paulo: same locality and date as holotype — 1 male, 3 females (pinned) (UMSP). Etymology. This species is named M. bocaina, as a noun in apposition, for the name of the beautiful national park where this species was collected.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970065FF8D98B1FA67FE32C5E7.taxon	description	Figs. 28, 31, 40 This is the most distinctive of the 3 new species of the M. velasquezi group from Brazil and unlikely to be confused because of its elongate, spine-like endophallic spines. Adult. Length of forewing: male 2.5 – 2.9, female 2.6 – 3.3 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 3: 4. Overall color dark brown. Legs brown, tibial spurs somewhat darker in color, but not strongly contrasting with legs, apices of tarsi whitish. Wing bar at anastamosis marked with white, contrasting setae. Male genitalia. Ventral process of segment VI posteriorly projecting, elongate, narrow, length more than 2 times width at base (Fig. 28 E). Segment IX moderately rounded anterolaterally, length greatest in ventral 1 / 2, posterolateral margin nearly linear; segment deeply excised dorsomesally and ventromesally, forming lateral lobes, lobes separated dorsomesally by about 1 / 2 width of segment. Tergum X moderate in length, simple in structure, with deep, broad, U-shaped mesal excision; apical lobes, as viewed laterally, distinctly projecting, rounded to subacute apically. Inferior appendages almost completely fused to phallic ensemble, basally with short, setose, anteriorly-directed lobes, apically with elongate narrow lobes. Mesal pockets of fused inferior appendages very bulbously enlarged, with elongate, spine-like, posteriorly-directed apicoventral projections. Paramere appendages very narrow, elongate, slightly dorsally curved, about as long as dorsal phallic spine. Phallotheca very short; ventral rod-like appendages elongate, with widely flared apices. Dorsal phallic spine, as viewed dorsally, broad in middle, gradually narrowed and rounded apically; as viewed laterally, relatively narrow, sharply upturned at about apical 1 / 3 rd. Phallicata with lightly sclerotized, slightly scabrous, rounded lateral lobes. Endophallic membrane relatively simple in structure, dorsally with small, rounded sclerite, apparently articulating with ventral margin of dorsal phallic spine; laterally, on either side, with very prominent, sclerotized, anteriorly-directed spine-like processes.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970065FF8D98B1FA67FE32C5E7.taxon	materials_examined	Holotype male: BRAZIL: Rio de Janeiro: Parati, Riacho Perequê-açu, Sitio Cachoeira Grande, 23 ° 13 ' 14 " S, 044 ° 47 ' 24 " W, 120 m, 25. ix. 2002, Blahnik, Prather, Melo, Froehlich & Silva (UMSP 000086573) (pinned) (MZUSP). Paratypes: BRAZIL: Rio de Janeiro: same locality and date as holotype — 13 males, 12 females (pinned) (MZUSP, UMSP, NMNH), 12 males, 2 females (alcohol) (UMSP); Parati, trib. to Riacho Perequê-açu, 23 ° 12 ' 50 " S, 044 ° 47 ' 29 " W, 190 m, 26. ix. 2002, Blahnik, Prather, Melo, Froehlich & Silva — 3 males, 7 females (pinned), 9 males, 6 females (alcohol) (UMSP); Parati, Riacho Perequê-açu, 23 ° 13 ' 27 " S, 044 ° 46 ' 09 " W, 30 m, 24. ix. 2002, Blahnik, Prather, Melo, Froehlich & Silva — 9 males, 3 females (pinned) (UMSP), 3 males, 1 female (alcohol) (MZUSP); Parque Estadual Intervales, Rio do Carmo, 24 ° 18 ' 59 " S, 048 ° 25 ' 15 " W, 560 m, 29. ix. 2002, Blahnik, Prather, Melo & Calor — 3 males, 3 females (pinned) (UMSP). Etymology. We take great pleasure in naming this species for Dr. Claudio Froehlich, both in recognition of his long career of working with aquatic insects in Brazil and also for his personal assistance during our collecting efforts in the country.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE8797006AFF8898B1FB95FD5FC783.taxon	description	Figs. 29, 30 This species is perhaps most likely to be confused with M. bocaina, since both species lack the elongate endophallic spines found in M. froehlichi. Unlike M. bocaina, this species lacks any sclerotized endophallic spines or lobes at all, but has a lightly sclerotized, rounded lateral lobe on the phallicata, similar to that found in M. froehlichi. The apical part of the inferior appendages is apparently adnate to or continuous with the apical spine-like projections of the mesal pockets of the fused inferior appendages, giving the ventral profile of the phallic ensemble an open appearance. Adult. Length of forewing: male 2.8 – 3.1 mm, female 3.0 – 3.5 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 3: 4. Overall color dark brown. Legs brown, tibial spurs somewhat darker in color, but not strongly contrasting with legs. Wing bar at anastamosis marked with white, contrasting setae. Male genitalia. Ventral process of segment VI posteriorly projecting, elongate, narrow, length more than 2 times width at base (Fig. 29 E). Segment IX moderately rounded anterolaterally, length greatest in ventral 1 / 2 (or near middle), posterolateral margin nearly linear; segment deeply excised dorsomesally and ventromesally, forming lateral lobes, lobes separated dorsomesally by about 1 / 2 width of segment. Tergum X moderate in length, simple in structure, with deep, broad, U-shaped mesal excision; apical lobes, as viewed laterally, distinctly projecting, rounded to subacute apically. Inferior appendages almost completely fused to phallic ensemble, basally with short, setose, anteriorly-directed lobes, apically apparently fused to spine-like apical projections of mesal pockets of fused inferior appendages. Mesal pockets of fused inferior appendages very bulbously enlarged, with elongate, spine-like, posteriorly-directed apicoventral projections. Paramere appendages very narrow, elongate, about as long as dorsal phallic spine. Phallotheca very short; ventral rod-like appendages elongate, with widely flared apices. Dorsal phallic spine, as viewed dorsally, broad in middle, gradually narrowed and rounded apically; as viewed laterally, relatively narrow, upturned, more distinctly at about apical 1 / 3 rd. Phallicata with lightly sclerotized, rounded lateral lobes. Endophallic membrane simple in structure, without sclerotized lateral processes or lobes. Phallotremal sclerite evident as small, rounded sclerite.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE8797006AFF8898B1FB95FD5FC783.taxon	materials_examined	Holotype male: BRAZIL: Minas Gerais: Estação Ecológica do Tripuí, Córrego Tripuí, 20 ° 23 ' 22 " S, 043 ° 32 ' 32 " W, 1070 m, 21. xi. 1998, Paprocki, Braga & Amarante (UMSP 000047049) (pinned) (MZUSP). Paratypes: BRAZIL: Minas Gerais: same locality and date as holotype — 1 female (pinned) (MZUSP); same locality, 16. viii. 1998, Paprocki & Amarante — 2 males, 1 female (pinned) (UMSP); same locality, 20. ix. 1998, Paprocki & Braga — 2 males (pinned) (UMSP, NMNH); Córrego Tripuí, 20 ° 23 ' 19 " S, 043 ° 32 ' 37 " W, 1100 m, 27. vii. 1998, Paprocki Braga & Amarante — 1 female (pinned) (UMSP). Etymology. This species is named M. tripuiensis after Estação Ecológica do Tripuí, the beautiful reserve where the holotype and paratype specimens were collected.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE8797006FFF8A98B1FE76FEA6C390.taxon	description	Sykora (1999) divided the known taxa into 5 " tentative " subgroups (the M. argentinica, bilineata, enchrysa, flinti, and wygodzinskii subgroups), and also suggested that concepts of relationships are likely to change once the taxa are better known. About 1 / 2 of these species were placed in his M. bilineata subgroup. Although the included taxa were listed for each group, the defining characters for the subgroups were not discussed. An examination of illustrations for the species and representative specimens from the subgroups, suggests that the majority of species in the M. bilineata species group do have a very coherent " gestalt " similarity, although individual species may vary somewhat from this idealized form. These characters apply especially to the M. bilineata, enchrysa, and wygodzinskii subgroups of Sykora (including the majority of species), all of which have a very consistent morphological pattern. The illustration of M. wygodzinskii (Fig. 34) can be used to point out the characters uniting this group. Character similarities include the following: A more or less elongate tergum X (as suggested by Flint), usually with the mesal margin characteristically notched (as in Fig. 34 B), although sometimes more deeply divided; the presence of sclerotized and setose ventral lobes on tergum X, bordering the dorsal phallic spine; an angulate ventral margin of the dorsal phallic spine, articulating with a sclerotized process on the phallicata [the process usually without dorsolateral processes acting as guides for the paramere appendages (M. roldani Flint an exception)]; a dorsal phallic spine that is sharply upturned apically and also has it apex rounded, as viewed laterally; a characteristic shape of segment IX, with the anteroventral margin produced and with the posterior margin distinctly angulate in its dorsal 1 / 2. Primary differences between the M. bilineata and enchrysa subgroups seem to be based largely on color. A number of the species in the M. bilineata subgroup of Sykora have the forewing distinctively marked with 2 wing bars, 1 at the anastamosis, as is common in Mortoniella, and another more proximal band (a unique character within Mortoniella). This contrasts with the distinctive color of the M. enchrysa group, with males uniformly golden in color (another unique character within Mortoniella), without any apparent wing bands, and with the membrane of the wing, beneath the golden setae, darkened or fuscous. Several of the species in the M. bilineata subgroup were described from pharate pupae or from alcohol, making it difficult to determine the original color of the wings, including M. angulata Flint, M. apiculata Flint, and M. hodgesi Flint. Species with 2 wing bands include M. bilineata Ulmer, M. roldani Flint, M. similis Sykora, M. chicana Sykora, and M. iridescens Flint, the first 3 with dark wings and white bands, M. chicana with light brown wings and white bands, and M. iridescens with dark wings and iridescent bands, only visible at some light angles. The only species placed in the M. enchrysa subgroup are M. enchrysa and M. paraenchrysa, although Flint (1996) recorded 2 additional undescribed species from Peru. Based on color considerations, and also genitalic characters, M. denticulata Sykora, which Sykora placed in his M. flinti subgroup, should probably be moved to the M. enchrysa subgroup. Males are nearly golden in color, without wing bars, and females have an indistinct or incomplete wing bar at the usual position, indicating the probable ancestral state for the group. Similarly, M. paralineata, which Sykora described in the M. bilineata subgroup, is described as being yellow with a fuscous wing membrane, suggesting its probable placement in the M. enchrysa subgroup. Mortoniella squamata Sykora, 1999, which Sykora placed in the M. bilineata subgroup, is described as being golden beige and unmarked, or not strictly conforming to either species group based on color considerations. Similarly, Mortoniella foersteri (Schmid, 1964) is discussed as being dark with a single white wing band. Although the M. enchrysa subgroup seems to represent a monophyletic assemblage, based on color considerations, monophyly of the entire M. bilineata group needs to be more clearly demonstrated. The M. flinti group of Sykora, as originally conceived, is quite heterogeneous and species differ significantly from the 3 species subgroups discussed above. With M. denticulata removed, only 4 species remain. We have not directly examined any of these species, although we have examined specimens of a species from Venezuela that is apparently very closely related to M. bifurcata Sykora, or possibly conspecific. It differs from the species discussed above in lacking an angulate ventral margin of the dorsal phallic spine and also a somewhat different overall structure of the dorsal phallic spine, as well as lacking an angulate posterior margin to segment IX. In most other respects, however, including the general structure of tergum X and also female genitalia with a deeply incised tergum VIII, it is characteristic of the M. bilineata species group. We believe that it is correctly placed here and that this will also prove to be true for M. flinti Sykora and M. quinas Harper & Turcotte. Based on genitalic considerations, we suspect that M. santiaga Sykora may not only not be a member of the M. flinti subgroup, but may be misplaced in the M. bilineata species group. However, this needs to be confirmed. Mortoniella wygodzinskii, the only species placed by Sykora in his M. wygodzinskii group, is discussed below in the redescription of that species. Morphologically it is very similar to the M. bilineata and enchrysa subgroups, but lacks the distinctive color attributes of either subgroup and has some peculiar morphological distinctions of its own. Based on genitalic considerations, M. argentinica, type species of the M. argentinica subgroup of Sykora, is being removed from the M. bilineata species group, as discussed in the species description for that species in the section on ' species unplaced to species group'. Both the overall form of the male genitalia, and also that of the female, more closely resemble species in the M. leroda species group. The only other species " speculatively " placed in the M. argentinica subgroup by Sykora is M. unilineata Sykora, which we believe may also be misplaced in the M. bilineata species group. However, this also needs verification. It appears that these species may have been placed in the M. bilineata group based mainly on venational considerations, which as discussed above, may be plesiomorphic.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE8797006DFFF498B1F980FCA5C5E8.taxon	description	Fig. 34	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE8797006DFFF498B1F980FCA5C5E8.taxon	description	Adult. Length of forewing: male 4.4 mm. Forewing with forks I, II, and III present, hind wing with forks II, III, and V. Spur formula 0: 4: 4. Overall color medium brown. Legs same color, tibial spurs darker, contrasting with legs. Wing without distinct bar at anastamosis; setae of wings slightly flattened. Male genitalia. Ventral process of segment VI posteriorly projecting, short, narrow basally, length subequal to width at base. Segment IX rounded anterolaterally, length greatest in ventral 1 / 2, posterolateral margin with moderately angular projection in dorsal 1 / 2, also abruptly, angularly narrowed ventrally; segment deeply excised dorsomesally and ventromesally, forming lateral lobes, lobes separated dorsomesally by more than 1 / 2 width of segment. Tergum X elongate, bulbously inflated, with narrow mesal excision, extending less than 1 / 4 th length; apical lobes, as viewed dorsally, bluntly rounded, as viewed laterally, more broadly rounded; ventromesally with strongly sclerotized, preapical projections, very narrowly separated mesally, and membranously connected, setose lobes, opposing dorsal phallic spine on either side. Inferior appendages with short, setose basal lobes and elongate, narrow, posteriorly-directed ventral projections, curved outward apically. Mesal pockets of fused inferior appendages with moderately elongate, spine-like, posteriorly-directed apicoventral projections. Paramere appendages very short, sclerotized, acute apically (possibly vestigial). Phallotheca with evident, rounded, laterally compressed, dorsomesal apodeme. Dorsal phallic spine, as viewed dorsally, broad in middle, narrowed apically; as viewed laterally, greatly narrowed in basal and apical parts, very bulbously inflated in middle, strongly upturned in apical 1 / 4 th, apex rounded; ventral margin distinctly angulate, articulating with rounded projection on phallicata. Phallicata with distinctly sclerotized basodorsal projection and lightly sclerotized, rounded, depressed, lateral lobes, partially microsetose on ventral surface. Endophallic membrane lightly sclerotized dorsally, simple in structure, without evident spines; phallotremal spines absent. Material examined. VENEZUELA: Truillo: Quebrada Potrerito, 7.5 km NE Boconó, 9 ° 16 ' 26 '' N, 70 ° 13 ' 06 " W, 1530 m, 29 - 30. iv. 1995, Holzenthal, Cressa, Gulic — 1 male (UMSP). Distribution. Argentina, Bolivia, Ecuador, Venezuela.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970011FFF698B1FF1EFE69C048.taxon	description	Figs. 35, 39	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970011FFF698B1FF1EFE69C048.taxon	description	Adult. Length of forewing: male 4.0 - 4.8 mm, female 4.6 mm. Forewing with forks I, II, and III present, hind wing with forks II, III, and V. Spur formula 0: 4: 4. Overall color dark brown. Legs same color, tibial spurs somewhat darker than legs, but not strongly contrasting in color. Wing bar at anastamosis indistinct, discontinuous, most strongly marked with whitish setae near arculus on anal margin. Male genitalia. Ventral process of segment VI posteriorly projecting, narrow, elongate, length about twice width at base (Fig. 35 E). Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin convexly rounded, narrowing ventrally; segment deeply excised dorsomesally and ventromesally, forming lateral lobes, lobes separated dorsomesally by much less than 1 / 2 width of segment. Tergum X with Ushaped mesal excision, extending less than 1 / 2 length of tergum, and distinctly projecting lateral lobes; lateral lobes acute apically, as viewed both dorsally and laterally. Inferior appendages short basally, with elongate, narrow, posteriorly arched, dorsolateral lobes; lobes with short papillae or papillate-like structures at apex and usually also at base. Mesal pockets of fused inferior appendages with apical processes very elongate, lance-like, posteriorly directed. Paramere appendages apparently with 2 processes on each side; 1 pair elongate, narrow, distinctly bowed outward in apical 1 / 2, slightly widened preapically, apex acute; 2 nd pair short, ventrally curved, narrowing apically, apex very narrow and acute. Dorsal phallic spine, as viewed laterally, more or less uniform in width, distinctly sinuously undulate in contour, apical 1 / 3 rd strongly upturned; in dorsal view, narrow throughout, apex narrowly acuminate. Phallicata moderately elongate, tubular, without dorsal processes, ventral margin lightly sclerotized. Endophallic membrane somewhat ballooned, simple in structure, without distinct lateral or ventral spines; phallotremal spines very short, curved outward. Material examined. ARGENTINA: Catamarca: N. Aconquija, 1 - 2. x. 1968, LE Peña G. — 1 male, 1 female (paratypes) (NMNH); Tucuman: Rt. 307, 7 km W. Acheral, 11. x. 1973, OS Flint, Jr., — 7 males (NMNH, UMSP). Distribution. Argentina.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970011FFF098B1F935FE7EC5C3.taxon	description	Fig. 36	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970011FFF098B1F935FE7EC5C3.taxon	description	Adult. (specimen totally cleared) Length of forewing: male 2.5 mm. Venation not discernible. Spur formula 0: 4: 4 (probably). Male genitalia. (Genital capsule separated from abdomen). Ventral process of segment VI very small, laterally compressed, ventrally projecting, subtriangular, widest basally, acute apically. Segment IX concavely rounded anterolaterally, sinuously produced and length greatest in ventral 1 / 2; segment deeply excised dorsomesally and ventromesally, forming lateral lobes, lobes separated dorsomesally by more than 1 / 2 width of segment, lobes discontinuous ventrally (probably broken). Tergum X very greatly inflated and rounded basally; apicomesal incision relatively short, narrow, U-shaped, forming projecting, acute, apicodorsal lobes; ventrolateral margin rounded, setose; tergum ventromesally with pair of converging, sclerotized, anteriorly-curved lobes. Inferior appendages very short, with only slight ventromesal projection, dorsally with relatively elongate, tapering, apically acute lobes, each strongly posteriorly bent at about midlength. Mesal pockets of fused inferior appendages with apical processes short, posteroventrally curved. Paramere appendages intermediate in length (shorter than dorsal phallic spine), very narrow, uniform in width, apices acute and slightly upturned. Dorsal phallic spine, as viewed laterally, widened in middle, obtusely angled upward from ventral margin, dorsal margin gradually curved, apical part nearly vertical in orientation, apex acute, slightly reflexed. Phallicata with very narrow, pencil-like dorsolateral processes, ventral margin weakly sclerotized, slightly projecting. Endophallic membrane very short, simple in structure, without ventromesal spine; phallotremal spines absent. Material examined. PARAGUAY: Salto del Guaira, 24 ° 02 ' S, 054 ° 16 ' W, 4. xii. 1971, L E Peña G - 1 male (holotype, alcohol, USNM type # 72741) (NMNH). Distribution. Paraguay	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970017FFF398B1FBB6FB6EC733.taxon	description	Fig. 37 This species is very distinctive and unlikely to be confused with any other species of Mortoniella. The nearly straight anterior margin of segment IX and synsclerous posterior margin of segment VIII are especially distinctive. Mortoniella meloi is also characterized by having slightly widened, but not exactly scale-like, setae along the costal margin of the forewings of the male, and on abdominal segments anterior to segment VIII. Adult. Length of forewing: male 3.5 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 4: 4. Overall color grayish-brown. Wing bar at anastamosis scarcely evident, marked only by whitish-brown setae at arculus on hind margin. Male genitalia. Ventral process of segment VI short, laterally compressed, ventrally projecting, bluntly rounded apically, slightly narrowed basally from anterior margin (Fig. 37 E). Segment VIII more or less synsclerous (more distinctly so on posterior margin); tergum with pronounced anterolateral apodemes. Segment IX very short, anterior margin nearly straight, posterior margin curved, narrowing dorsally; segment excised dorsomesally and ventromesally, forming lateral lobes, lobes separated dorsomesally by more than 1 / 2 width of segment, lobes convergent ventrally and very narrowly joined. Tergum X short; apicomesal incision relatively narrow, U-shaped, forming projecting, acute, apicodorsal lobes; ventrolateral margin subtruncate, setose; tergum ventromesally with convergent, mesally-fused, sclerotized, anteriorly-curved lobes. Inferior appendages very short, with short, paired, acute ventromesal projections, dorsally with short, subacute, outwardly-flared lobes, Mesal pockets of fused inferior appendages with apical processes short, dorsally curved. Paramere appendages relatively elongate (extending about as far as dorsal phallic spine), narrow, uniform in width, apices acute. Dorsal phallic spine, as viewed laterally, more or less uniform in width, strongly dorsally oriented, apical 1 / 4 th very sharply bent, apex acute and somewhat reflexed. Phallicata with sclerotized dorsolateral processes, each concavely curved to accommodate paramere appendage. Endophallic membrane very short, simple in structure; phallotremal sclerite composed of short mesal spine and 2 longer, diverging, lateral spines (possibly fused phallotremal spines).	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
03BE87970017FFF398B1FBB6FB6EC733.taxon	materials_examined	Holotype male: BRAZIL: São Paulo: Parque Estadual Intervales, Rio do Carmo, 24 ° 18 ' 59 " S, 048 ° 25 ' 15 " W, 560 m, 29. ix. 2002, Blahnik, Prather, Melo & Calor (UMSP 000088168) (pinned) (MZUSP). Etymology. We are very pleased to name this species M. meloi for Adriano Melo, who helped to collect the type specimens, in gratitude for the assistance he rendered during our collecting efforts in Brazil.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2011): Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851. Zootaxa 2851 (1): 1-75, DOI: 10.11646/zootaxa.2851.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1
