identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03BE87B64A77FF9340C5CDF4FC8FE50B.text	03BE87B64A77FF9340C5CDF4FC8FE50B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Geholaspis Berlese 1918	<div><p>Genus Geholaspis Berlese, 1918</p><p>Macrocheles (Geholaspis) Berlese, 1918: 145 .</p><p>Type species. Gamasus longispinosus Kramer, 1876, by original designation.</p><p>Diagnosis of the genus (female and male). Dorsal shield oblong or widely oval, bearing 28 pairs of brush-like or smooth (setae j5, j6, z5, z6, J2 and J5) setae. Dorsal shield seta j1 short and plumose, z1 smooth or barbed. Dorsal shield seta j5 is displaced posteriorly to a position behind the base of seta z5. Metasternal plates bearing st4 and iv3 free (subgenus Geholaspis s. str.) or fused to endopodal plates (subgenus Cyrtocheles Valle, 1953). Ventri-anal shield usually wider than long, with five pairs of pre-anal setae and a pair of inguinal pores. Deutosternal groove with five wide rows of fine denticles. Corniculi elongate and narrow. Chelicerae robust, movable digit with 2–3 denticles, fixed digit with fewer than six teeth (rarely more as G. (G.) biperforatus). Epistome with triangular serrate base and an elongate, tapering bifid median process. Males with holoventral shield. The male chelicera is similar to that in the genus Macrocheles, with a short dorsally-directed spermatodactyl. Femur II has a small spur. The genus Geholaspis differs from Longicheles in that Longicheles has elongate and multidentate cheliceral digits, so that both fixed and movable digits have at least eight teeth, and the epistome of Longicheles has a transverse dentate base and a narrow distally bifid median process.</p><p>Notes on the genus. The genus Geholaspis is widely distributed in the Palaearctic region, including Europe (Austria, Switzerland, Netherlands, Belgium, England, Italy, Ireland, Hungary and Slovakia), Turkey and Iran (present study) (Krauss, 1970; Bregetova, 1977; Mašán, 2003; Özbek &amp; Bal, 2014). The presence of G. longispinosus (Kramer, 1876) in New Zealand (Emberson, 1973) and Longicheles mandibularis (Berlese, 1904) in Australia (Halliday, 2001) may be the result of human activity.</p></div>	https://treatment.plazi.org/id/03BE87B64A77FF9340C5CDF4FC8FE50B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Babaeian, Esmaeil;Halliday, Bruce;Saboori, Alireza	Babaeian, Esmaeil, Halliday, Bruce, Saboori, Alireza (2015): A new species of Geholaspis Berlese (Acari: Mesostigmata: Macrochelidae) from Northern Iran. Zootaxa 3925 (3): 422-430, DOI: 10.11646/zootaxa.3925.3.6
03BE87B64A74FF9640C5C9A9FB76E44A.text	03BE87B64A74FF9640C5C9A9FB76E44A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Geholaspis (Geholaspis) pennulatus	<div><p>Geholaspis (Geholaspis) pennulatus sp. nov.</p><p>(Figs 1‒16)</p><p>Diagnosis. Dorsal shield oval, with reticulate and micropunctate ornamentation. Dorsal setae z1 and j2 barbed, seta j5 displaced posteriorly and inserted at a level behind setae j6 and z6. Setae j5, j5, z5, z6, J2 and J5 smooth and needle-like. Sternal shield punctate-reticulate (some punctation linearly arranged), ventri-anal shield wide, bearing five pairs of smooth and needle-like pre-anal setae.</p><p>Material examined. Holotype, female, Kheyroodkenar forest. Nowshahr, Mazandaran province, E. Babaeian coll. Leaf-litter, 36˚35.265’ N, 051˚34.271’ E, alt. 532 m, 5 July 2014. Paratypes: seven females, two males, same data as holotype (in JAZM and ANIC).</p><p>Description. Female</p><p>Dorsal idiosoma (Fig. 1). Dorsal shield oblong, length 957–978, width 649–667 (n=4), punctate-reticulate over whole surface and a complement of 28 pairs of setae (j1–j6, z1, z2, z4–z6, s2, s4–s6, r2–r4, J2, J5, Z1, Z2, Z4, Z5, S1, S2, S4, S5) and 22 pairs of pores (including 16 pairs of lyrifissures and six pairs of gland pores), seta j1 with separate base, plumose for its entire length, seta z1 long and barbed, setae j5, j6, z5, z6, J2 and J5 smooth and needle-like, remaining dorsal setae plumose in the distal 2/3. Seta j5 displaced behind setae j6 and z6. Measurements of setae: j1 37–45, j2 92–97, z1 104–110, z5 93, J5 87–99, Z5 83–87.</p><p>Ventral idiosoma (Fig. 2). Tritosternum with columnar base 27–30 and two pilose laciniae 97–104. Sternal shield (Fig. 11) ornamented with micropunctate sculpture and with distinct sculptural lines, 174–186 long, 196–205 wide at level of setae st2–st2, slightly wider than long (ratio of length/width ≈ 0.93), bearing pilose st1 (78–87), st2 (55–68), st3 (54–74) and two pairs of lyrifissures; st4 (48–50) smooth, and third pair of sternal lyrifissures iv3 on oval and free metasternal plates. Epigynal shield (210‒223 long, 260–307 wide at level of epigynal setae) as defined in genus, surface of the shield with lines bordered by fine punctation (Fig. 12); st5 52–58 long and smooth. Ventri-anal shield (Fig. 13) ornate with lines and fine punctation, large and greatly widened (411–428 long, and 501‒517 wide at greatest point, ratio of length/width ≈ 0.84); with five pairs of smooth preanal setae (Jv1 67–85; Jv2 102–114; Jv3 94–99; Zv1 106–119; Zv2 70–77), a pair of para-anal setae (33–37) and a post-anal seta (12–14). Cribrum well developed, anterolateral extensions of cribrum reaching bases of para-anal setae. Metapodal platelets small. Peritrematal shields (Fig. 6) with three pairs of gland pores gp and two pairs of lyrifissures ip, peritremes long and extending anteriorly and laterally and reaching to level of insertions of z1 and with three pairs of lyrifissures.</p><p>Gnathosoma (Figs 3–5, 14). Epistome with triangular and sharply denticulate base, and a bifid median process (Fig. 4); subcapitulum typical for genus (Fig. 3). Hypostomatal (h1, h2, h3) and capitular setae simple and smooth, 70–74, 21–25, 50 and 20–22 long, respectively. Deutosternal groove with five wide transverse rows of fine denticles. Internal malae slender, pointed and densely pubescent, labrum prominent, blade-like and pilose on surface. Corniculi slender and horn-like, 88–94 long, salivary styli conspicuous and flanked by corniculi. Chelicerae robust, with tridentate movable digit (100–104 long), a blunt proximal tooth, a larger backwardlydirected subdistal tooth, and a large tooth adjacent to minute distal hook; middle segment 245–270 long, fixed digit (101–106 long) with four teeth, external face with two adjoining teeth and a minute distal one, a terminal hook and smooth dorsal seta (Figs 5, 14). Arthrodial brushes pilose. Chaetotaxy of palp trochanter, femur and genu 2–5–6. Palp apotele with three subequal teeth.</p><p>Legs. Tarsus I without ambulacrum and claw; tarsi II–IV with well developed ambulacra and claws. Length of legs (except ambulacrum): leg I, 582‒594, leg II, 630–686, leg III, 515‒599, leg IV, 866‒903; tarsus II as in Fig. 7. Chaetotaxy: Leg I: coxa 0 0/1 0/1 0, trochanter 1 0/1 0/2 1, femur 2 3/1 2/3 2, genu 2 3/1 2/1 1, tibia 2 3/2 2/1 2. Leg II: coxa 0 0/1 0/1 0, trochanter 1 0/1 0/2 1, femur 2 3/1 2/2 2, genu 2 3/1 2/1 2, tibia 2 2/1 2/1 2. Leg III: coxa 0 0/1 0/1 0, trochanter 1 1/1 0/1 1, femur 1 2/1 1/0 1, genu 1 2/1 2/0 1, tibia 1 1/1 2/1 1. Leg IV: coxa 0 0/0 0/1 0, trochanter 1 1/2 0/1 0, femur 1 2/1 1/0 1, genu 1 2/1 2/0 0, tibia 1 1/1 2/1 1. Tarsi II–IV with 18 setae: 3 3/2 3/2 3 + mv, md. All leg setae pilose, except for some smooth setae on tibia I (ad, pd, al and pl) and tibia II (al and av). All tarsal setae smooth.</p><p>Male. Dorsal idiosoma (Fig. 8). Dorsal shield oblong, length 834‒850, width 611‒638 (n=2), similar in ornamentation and setation to female.</p><p>Ventral idiosoma (Figs 9, 16). Holoventral shield punctate-reticulate throughout, widened posteriorly. Ventral adenotaxy and poroidotaxy as in female.</p><p>Gnathosoma . As in female, except chelicera (Figs 10, 15) with movable digit length 79–84, bidentate, with a blunt proximal tooth and a larger backwardly directed subdistal one. Middle segment length 214–220, fixed digit length 81–85, with five teeth and ending a terminal hook, external face with two teeth, dorsal seta smooth. Spermatodactyl short. Arthrodial brushes pilose.</p><p>Legs. Length of legs (except ambulacrum): leg I, 511‒527, leg II, 553–577, leg III, 515‒522, leg IV, 763‒794.</p><p>Etymology. The name " pennulatus " refers to the feather-like appearance of dorsal shield setae j1 and z1.</p><p>Notes. Geholaspis (G.) pennulatus sp. nov. differs from most other species in the subgenus in the position of dorsal shield seta j5, which is strongly displaced posteriorly to a position behind seta j6. Only G. (G.) pauperior shares this character state. In the other species, seta j5 is at a level between setae z5 and j6. In G. (G.) pennulatus sp. nov. the dorso-central setae on the dorsal shield (z5, j5, j6, z6, J2 and J5) and the pre-anal setae on the ventrianal shield are needle-like, while in G. (G.) pauperior these setae are flattened and blade-like.</p></div>	https://treatment.plazi.org/id/03BE87B64A74FF9640C5C9A9FB76E44A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Babaeian, Esmaeil;Halliday, Bruce;Saboori, Alireza	Babaeian, Esmaeil, Halliday, Bruce, Saboori, Alireza (2015): A new species of Geholaspis Berlese (Acari: Mesostigmata: Macrochelidae) from Northern Iran. Zootaxa 3925 (3): 422-430, DOI: 10.11646/zootaxa.3925.3.6
03BE87B64A70FF9440C5CD35FA46E4DD.text	03BE87B64A70FF9440C5CD35FA46E4DD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Geholaspis (Geholaspis)	<div><p>Key to the known species of Geholaspis (Geholaspis) (adult females)</p><p>1. Seta j5 inserted at a level posterior to both z5 and j6 .......................................................... 2</p><p>- Seta j5 inserted at a level between z5 and j6 ................................................................. 3</p><p>2. Dorso-central setae (including j5, j6, z5, z6, J2 and J5) and pre-anal setae smooth and blade-like; seta J5 as long as Z5; epis- tome with long triangular base and short and dissected median process................. G. (G.) pauperior (Berlese, 1918)</p><p>- Dorso-central setae (including j5, j6, z5, z6, J2 and J5) and pre-anal setae smooth and needle-like; seta J5 slightly longer than Z5; epistome with short triangular base and long and bifid median process................... G. (G.) pennulatus sp. nov.</p><p>3. Dorsal setae j6 and z6 short, much shorter than the distance j6-z6 ...................... G. (G.) biperforatus Krauss, 1970</p><p>- Dorsal setae j6 and z6 long, clearly longer than the distance j6-z6 ................................................ 4</p><p>4. Seta j1 with widely separated base; ventri-anal setae Zv1, Jv2 and Jv3 elongate................ G. (G.) berlesei Valle, 1953</p><p>- Seta j1 with adjacent base; ventri-anal setae Zv1, Jv2 and Jv3 not elongate........................................ 5</p><p>5. Ventri-anal shield only slightly wider than long, sternal shield longer than wide............ G. (G.) alpinus (Berlese, 1887)</p><p>- Ventri-anal shield distinctly wider than long, ratio length:width at least 1.2:1; sternal shield wider than long or as long as wide.................................................................................................... 6</p><p>6. Dorsal setae j2 and s2 smooth, st1-st3 short, not reaching base of next posterior seta; movable digit bidentate; with a large backwardly directed tooth, a minute distal tooth, and a terminal hook; fixed digit with a large median tooth, a small blunt prox- imal tooth and a minute subdistal tooth, ventri-anal shield smoothly reticulate......... G. (G.) longispinosus (Kramer, 1876)</p><p>- Dorsal setae j2 and s2 plumose, st1-st3 long, reaching base of next posterior seta; movable digit tridentate; with a large back- wardly directed tooth, a minute distal tooth, and a minute blunt proximal tooth, and a terminal hook; fixed digit with a forked distal tooth and about eight small proximal teeth, ventri-anal shield punctate-reticulate......... G. (G.) aeneus Krauss, 1970</p></div>	https://treatment.plazi.org/id/03BE87B64A70FF9440C5CD35FA46E4DD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Babaeian, Esmaeil;Halliday, Bruce;Saboori, Alireza	Babaeian, Esmaeil, Halliday, Bruce, Saboori, Alireza (2015): A new species of Geholaspis Berlese (Acari: Mesostigmata: Macrochelidae) from Northern Iran. Zootaxa 3925 (3): 422-430, DOI: 10.11646/zootaxa.3925.3.6
