taxonID	type	description	language	source
03BE87B64A77FF9340C5CDF4FC8FE50B.taxon	materials_examined	Type species. Gamasus longispinosus Kramer, 1876, by original designation.	en	Babaeian, Esmaeil, Halliday, Bruce, Saboori, Alireza (2015): A new species of Geholaspis Berlese (Acari: Mesostigmata: Macrochelidae) from Northern Iran. Zootaxa 3925 (3): 422-430, DOI: 10.11646/zootaxa.3925.3.6
03BE87B64A77FF9340C5CDF4FC8FE50B.taxon	diagnosis	Diagnosis of the genus (female and male). Dorsal shield oblong or widely oval, bearing 28 pairs of brush-like or smooth (setae j 5, j 6, z 5, z 6, J 2 and J 5) setae. Dorsal shield seta j 1 short and plumose, z 1 smooth or barbed. Dorsal shield seta j 5 is displaced posteriorly to a position behind the base of seta z 5. Metasternal plates bearing st 4 and iv 3 free (subgenus Geholaspis s. str.) or fused to endopodal plates (subgenus Cyrtocheles Valle, 1953). Ventri-anal shield usually wider than long, with five pairs of pre-anal setae and a pair of inguinal pores. Deutosternal groove with five wide rows of fine denticles. Corniculi elongate and narrow. Chelicerae robust, movable digit with 2 – 3 denticles, fixed digit with fewer than six teeth (rarely more as G. (G.) biperforatus). Epistome with triangular serrate base and an elongate, tapering bifid median process. Males with holoventral shield. The male chelicera is similar to that in the genus Macrocheles, with a short dorsally-directed spermatodactyl. Femur II has a small spur. The genus Geholaspis differs from Longicheles in that Longicheles has elongate and multidentate cheliceral digits, so that both fixed and movable digits have at least eight teeth, and the epistome of Longicheles has a transverse dentate base and a narrow distally bifid median process. Notes on the genus. The genus Geholaspis is widely distributed in the Palaearctic region, including Europe (Austria, Switzerland, Netherlands, Belgium, England, Italy, Ireland, Hungary and Slovakia), Turkey and Iran (present study) (Krauss, 1970; Bregetova, 1977; Mašán, 2003; Özbek & Bal, 2014). The presence of G. longispinosus (Kramer, 1876) in New Zealand (Emberson, 1973) and Longicheles mandibularis (Berlese, 1904) in Australia (Halliday, 2001) may be the result of human activity.	en	Babaeian, Esmaeil, Halliday, Bruce, Saboori, Alireza (2015): A new species of Geholaspis Berlese (Acari: Mesostigmata: Macrochelidae) from Northern Iran. Zootaxa 3925 (3): 422-430, DOI: 10.11646/zootaxa.3925.3.6
03BE87B64A74FF9640C5C9A9FB76E44A.taxon	diagnosis	Diagnosis. Dorsal shield oval, with reticulate and micropunctate ornamentation. Dorsal setae z 1 and j 2 barbed, seta j 5 displaced posteriorly and inserted at a level behind setae j 6 and z 6. Setae j 5, j 5, z 5, z 6, J 2 and J 5 smooth and needle-like. Sternal shield punctate-reticulate (some punctation linearly arranged), ventri-anal shield wide, bearing five pairs of smooth and needle-like pre-anal setae.	en	Babaeian, Esmaeil, Halliday, Bruce, Saboori, Alireza (2015): A new species of Geholaspis Berlese (Acari: Mesostigmata: Macrochelidae) from Northern Iran. Zootaxa 3925 (3): 422-430, DOI: 10.11646/zootaxa.3925.3.6
03BE87B64A74FF9640C5C9A9FB76E44A.taxon	materials_examined	Material examined. Holotype, female, Kheyroodkenar forest. Nowshahr, Mazandaran province, E. Babaeian coll. Leaf-litter, 36 ˚ 35.265 ’ N, 051 ˚ 34.271 ’ E, alt. 532 m, 5 July 2014. Paratypes: seven females, two males, same data as holotype (in JAZM and ANIC).	en	Babaeian, Esmaeil, Halliday, Bruce, Saboori, Alireza (2015): A new species of Geholaspis Berlese (Acari: Mesostigmata: Macrochelidae) from Northern Iran. Zootaxa 3925 (3): 422-430, DOI: 10.11646/zootaxa.3925.3.6
03BE87B64A74FF9640C5C9A9FB76E44A.taxon	description	Description. Female Dorsal idiosoma (Fig. 1). Dorsal shield oblong, length 957 – 978, width 649 – 667 (n = 4), punctate-reticulate over whole surface and a complement of 28 pairs of setae (j 1 – j 6, z 1, z 2, z 4 – z 6, s 2, s 4 – s 6, r 2 – r 4, J 2, J 5, Z 1, Z 2, Z 4, Z 5, S 1, S 2, S 4, S 5) and 22 pairs of pores (including 16 pairs of lyrifissures and six pairs of gland pores), seta j 1 with separate base, plumose for its entire length, seta z 1 long and barbed, setae j 5, j 6, z 5, z 6, J 2 and J 5 smooth and needle-like, remaining dorsal setae plumose in the distal 2 / 3. Seta j 5 displaced behind setae j 6 and z 6. Measurements of setae: j 1 37 – 45, j 2 92 – 97, z 1 104 – 110, z 5 93, J 5 87 – 99, Z 5 83 – 87. Ventral idiosoma (Fig. 2). Tritosternum with columnar base 27 – 30 and two pilose laciniae 97 – 104. Sternal shield (Fig. 11) ornamented with micropunctate sculpture and with distinct sculptural lines, 174 – 186 long, 196 – 205 wide at level of setae st 2 – st 2, slightly wider than long (ratio of length / width ≈ 0.93), bearing pilose st 1 (78 – 87), st 2 (55 – 68), st 3 (54 – 74) and two pairs of lyrifissures; st 4 (48 – 50) smooth, and third pair of sternal lyrifissures iv 3 on oval and free metasternal plates. Epigynal shield (210 ‒ 223 long, 260 – 307 wide at level of epigynal setae) as defined in genus, surface of the shield with lines bordered by fine punctation (Fig. 12); st 5 52 – 58 long and smooth. Ventri-anal shield (Fig. 13) ornate with lines and fine punctation, large and greatly widened (411 – 428 long, and 501 ‒ 517 wide at greatest point, ratio of length / width ≈ 0.84); with five pairs of smooth preanal setae (Jv 1 67 – 85; Jv 2 102 – 114; Jv 3 94 – 99; Zv 1 106 – 119; Zv 2 70 – 77), a pair of para-anal setae (33 – 37) and a post-anal seta (12 – 14). Cribrum well developed, anterolateral extensions of cribrum reaching bases of para-anal setae. Metapodal platelets small. Peritrematal shields (Fig. 6) with three pairs of gland pores gp and two pairs of lyrifissures ip, peritremes long and extending anteriorly and laterally and reaching to level of insertions of z 1 and with three pairs of lyrifissures. Gnathosoma (Figs 3 – 5, 14). Epistome with triangular and sharply denticulate base, and a bifid median process (Fig. 4); subcapitulum typical for genus (Fig. 3). Hypostomatal (h 1, h 2, h 3) and capitular setae simple and smooth, 70 – 74, 21 – 25, 50 and 20 – 22 long, respectively. Deutosternal groove with five wide transverse rows of fine denticles. Internal malae slender, pointed and densely pubescent, labrum prominent, blade-like and pilose on surface. Corniculi slender and horn-like, 88 – 94 long, salivary styli conspicuous and flanked by corniculi. Chelicerae robust, with tridentate movable digit (100 – 104 long), a blunt proximal tooth, a larger backwardlydirected subdistal tooth, and a large tooth adjacent to minute distal hook; middle segment 245 – 270 long, fixed digit (101 – 106 long) with four teeth, external face with two adjoining teeth and a minute distal one, a terminal hook and smooth dorsal seta (Figs 5, 14). Arthrodial brushes pilose. Chaetotaxy of palp trochanter, femur and genu 2 – 5 – 6. Palp apotele with three subequal teeth. Legs. Tarsus I without ambulacrum and claw; tarsi II – IV with well developed ambulacra and claws. Length of legs (except ambulacrum): leg I, 582 ‒ 594, leg II, 630 – 686, leg III, 515 ‒ 599, leg IV, 866 ‒ 903; tarsus II as in Fig. 7. Chaetotaxy: Leg I: coxa 0 0 / 1 0 / 1 0, trochanter 1 0 / 1 0 / 2 1, femur 2 3 / 1 2 / 3 2, genu 2 3 / 1 2 / 1 1, tibia 2 3 / 2 2 / 1 2. Leg II: coxa 0 0 / 1 0 / 1 0, trochanter 1 0 / 1 0 / 2 1, femur 2 3 / 1 2 / 2 2, genu 2 3 / 1 2 / 1 2, tibia 2 2 / 1 2 / 1 2. Leg III: coxa 0 0 / 1 0 / 1 0, trochanter 1 1 / 1 0 / 1 1, femur 1 2 / 1 1 / 0 1, genu 1 2 / 1 2 / 0 1, tibia 1 1 / 1 2 / 1 1. Leg IV: coxa 0 0 / 0 0 / 1 0, trochanter 1 1 / 2 0 / 1 0, femur 1 2 / 1 1 / 0 1, genu 1 2 / 1 2 / 0 0, tibia 1 1 / 1 2 / 1 1. Tarsi II – IV with 18 setae: 3 3 / 2 3 / 2 3 + mv, md. All leg setae pilose, except for some smooth setae on tibia I (ad, pd, al and pl) and tibia II (al and av). All tarsal setae smooth. Male. Dorsal idiosoma (Fig. 8). Dorsal shield oblong, length 834 ‒ 850, width 611 ‒ 638 (n = 2), similar in ornamentation and setation to female. Ventral idiosoma (Figs 9, 16). Holoventral shield punctate-reticulate throughout, widened posteriorly. Ventral adenotaxy and poroidotaxy as in female. Gnathosoma. As in female, except chelicera (Figs 10, 15) with movable digit length 79 – 84, bidentate, with a blunt proximal tooth and a larger backwardly directed subdistal one. Middle segment length 214 – 220, fixed digit length 81 – 85, with five teeth and ending a terminal hook, external face with two teeth, dorsal seta smooth. Spermatodactyl short. Arthrodial brushes pilose. Legs. Length of legs (except ambulacrum): leg I, 511 ‒ 527, leg II, 553 – 577, leg III, 515 ‒ 522, leg IV, 763 ‒ 794.	en	Babaeian, Esmaeil, Halliday, Bruce, Saboori, Alireza (2015): A new species of Geholaspis Berlese (Acari: Mesostigmata: Macrochelidae) from Northern Iran. Zootaxa 3925 (3): 422-430, DOI: 10.11646/zootaxa.3925.3.6
03BE87B64A74FF9640C5C9A9FB76E44A.taxon	etymology	Etymology. The name " pennulatus " refers to the feather-like appearance of dorsal shield setae j 1 and z 1. Notes. Geholaspis (G.) pennulatus sp. nov. differs from most other species in the subgenus in the position of dorsal shield seta j 5, which is strongly displaced posteriorly to a position behind seta j 6. Only G. (G.) pauperior shares this character state. In the other species, seta j 5 is at a level between setae z 5 and j 6. In G. (G.) pennulatus sp. nov. the dorso-central setae on the dorsal shield (z 5, j 5, j 6, z 6, J 2 and J 5) and the pre-anal setae on the ventrianal shield are needle-like, while in G. (G.) pauperior these setae are flattened and blade-like.	en	Babaeian, Esmaeil, Halliday, Bruce, Saboori, Alireza (2015): A new species of Geholaspis Berlese (Acari: Mesostigmata: Macrochelidae) from Northern Iran. Zootaxa 3925 (3): 422-430, DOI: 10.11646/zootaxa.3925.3.6
