taxonID	type	description	language	source
03BE87F2FFB1C63478D9FE1AFE9A5C02.taxon	description	Wing length 0.35 – 1.8 mm. Antenna with 6 – 13 flagellomeres. AR 0.16 – 1.42. Anal point usually absent or rarely present.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFB6C63378D9FD20FE045A49.taxon	materials_examined	Material examined: P. R. CHINA: Tianjin City, Nankai University, 39 ° 8 ' N, 117 ° 12 ' E, alt. 2 – 5 m, sweep net, 5 males, 20. iv. 1986, Xinhua Wang; Tianjin City, Wuqing County, Dahuangpu wetland area, 39 ° 3 ' N, 117 ° 3 ' E, alt. 5 m, sweep net, 24 males, 26. iv. 2005, Xinhua Wang. Diagnostic characters. The species can be separated from other members of the genus by antenna with 10 flagellomeres, sternapodeme inverted V-shaped to U-shaped, and undeveloped inferior volsella.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFB6C63378D9FD20FE045A49.taxon	description	Addition to description. Chinese specimens had the following measurements: Total length 1.28 – 1.80, 1.36 mm. Wing length 1.00 – 1.25, 1.10 mm. Antenna with 10 flagellomeres; AR 0.77 – 1.1, 1.0. Apex of hind tibia with one weakly hooked seta near spur. Sternapodeme inverted V-shaped, attachment point with phallapodeme placed in caudal third of lateral sternapodeme and directed caudally, phallapodeme 85 – 90, 88 µm long, strongly curved with projection for joint with sternapodeme placed prelateral, and not extending beyond posterior margin of tergite IX. Gonocoxite 80 – 128, 100 µm long, with 4 – 6 setae apically. Gonostylus apically curved, with high basal lobe on inner margin.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFB6C63378D9FD20FE045A49.taxon	distribution	Distribution. Austria, Finland, France, Germany, Great Britain, Norway, Portugal, Spain, Switzerland, Russia including Novaya Zemlya and Russian Far East, Northern China, Canada, USA (Oliver et al. 1990, Saether & Spies 2004).	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFB6C63178D9FA4CFE6F5EB4.taxon	materials_examined	Type material: Holotype male (BDN No. 13103), P. R. CHINA: Sichuan Province, Kangding County, Wasigou, 30 ° 03 ' N, 101 ° 58 ' E, alt. 2560 m, light trap, 15. vi. 1996, Xinhua Wang.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFB6C63178D9FA4CFE6F5EB4.taxon	etymology	Etymology. From Latin, confidens, trusting, bold, referring to the stout ultimate flagellomere. Diagnostic characters. The species can be separated from other members of the genus by having AR of 0.16, inferior volsella reduced, and phallapodeme not extending beyond posterior margin of tergite IX.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFB6C63178D9FA4CFE6F5EB4.taxon	description	Description Male (n = 1). Total length 1.16 mm. Wing length 0.70 mm. Total length / wing length 1.65. Wing length / length of profemur 3.4. Coloration: Head dark brown. Antenna and palp yellow. Thorax brown. Legs yellowish-brown. Tergites I – V yellow, tergites VI – IX brown. Wings light yellow to hyaline with pale yellow clava. Head: Antenna (Fig. 3 B) with 10 flagellomeres; ultimate flagellomere stout, 55 µm long, with short sensilla chaetica subapically, apex distinctly swollen, 80 µm long; AR 0.16. Tentorium (Fig. 3 E) 115 µm long, 15 µm wide. Stipes 55 µm long. Cibarial pump slightly incurved. Palpomeres lengths (in µm): 13; 13; 20; 15; 50. Palpomere 2, 3 and 4 ellipsoid, 5 long and broad. Palpomere 5 / 3: 2.5. Thorax: Dorsocentrals more than 3, prealars 3, supraalar 1. Scutellum with 2 setae. Wing (Fig 3 A): VR 3.2. C length 170 µm, C / wing length 0.24. Cu length 420 µm. Cu / wing length 0.6. Wing width / length 0.46. C with 7 – 8 setae. Legs: Spur of fore tibia 15 µm long, of mid tibia 8 µm long, and of hind tibia 23 µm long. Apex of fore tibia 15 µm wide, of mid tibia 15 µm, of hind tibia 35 µm. Apex of hind tibia expanded, with comb of 12 setae, one seta near spur strongly S-shaped. Lengths and proportions of legs as in Table 1. TABLE 1. Lengths (in µm) and proportion of leg segments of Corynoneura confidens sp. n., male. fe ti ta 1 ta 2 ta 3 ta 4 ta 5 LR BV SV BR p 1 205 250 130 70 35 20 30 0.52 3.8 3.5 2.0 p 2 295 275 155 73 38 18 28 0.56 4.7 3.7 1.8 p 3 238 260 143 78 33 15 25 0.55 4.3 3.5 2.3 Hypopygium (Figs 3 C – D): Anal point absent. Tergite IX with posterior margin almost straight. Laterosternite IX with 1 long seta. Superior volsella narrow. Inferior volsella inconspicuous. Sternapodeme inverted V-shaped; coxapodeme 23 µm long; attachment point with phallapodeme placed in caudal third of lateral sternapodeme and directed caudally, phallapodeme 65 µm long, strongly curved with projection for joint with sternapodeme placed pre-laterally, not extending beyond posterior margin of tergite IX, with projection basally. Gonocoxite 63 µm long, with 1 seta apically. Gonostylus simple, 25 µm long; megaseta 3 µm long. HR 2.7, HV 4.6.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFB4C63178D9FE78FB1658DE.taxon	materials_examined	Type material: Holotype male (BDN No. 25092), P. R. CHINA: Xinjiang Province, Aletai City, Eerqisi river, 88 ° 07 ' E, 47 ° 55 ' N, alt. 400 m, sweep net, 3. xii. 2002, Hongqu Tang.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFB4C63178D9FE78FB1658DE.taxon	etymology	Etymology. From Latin, cylinder and cauda, tail, appendage, referring to the cylindrical anal point. Diagnostic characters. The species is similar to C. tokarapequea in having antenna with 11 flagellomeres and a similarly shaped sternapodeme, but can be separated by having a cylindrical anal point and broad inferior volsella.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFB4C63178D9FE78FB1658DE.taxon	description	Description Male (n = 1). Total length 1.32 mm. Wing length 0.84 mm. Total length / wing length 1.6. Wing length / length of profemur 3.2. Coloration: Head brown. Antenna and palp yellow. Thorax dark brown. Legs yellow. Abdomen blackish brown. Wings light yellow to hyaline, with pale yellow clava. Head: Antenna (Fig. 4 B) with 11 flagellomeres; ultimate flagellomere 140 µm long, with rosette of apical sensilla chaetica, apically expanded, thickening 60 µm long; AR 0.42. Tentorium (Fig. 4 E) 135 µm long, 15 µm wide. Stipes 63 µm long. Palpomeres length (in µm): 13; 15; 20; 25; 58. Palpomere 2, 3 and 4 ellipsoid, 5 long and slender. Palpomere 5 / 3: 2.9. Thorax: Dorsocentrals 5, prealars 4. Scutellum with 2 setae. Wing (Fig. 4 A): VR 3.2. C length 190 µm, C / wing length 0.23. Cu length 510 µm. Cu / wing length 0.61. Wing width / length 0.42. C with 7 setae. Legs: Spurs of fore tibia 20 µm and 13 µm long, of mid tibia 13 µm and 8 µm long, and of hind tibia 28 µm long. Width at apex of fore tibia 23 µm, of mid tibia 28 µm, of hind tibia 35 µm. Apex of hind tibia expanded, with comb of 17 setae, one seta near spur strongly S-shaped. Lengths and proportions of legs as in Table 2. TABLE 2. Lengths (in µm) and proportion of leg segments of Corynoneura cylindricauda sp. n., male. fe ti ta 1 ta 2 ta 3 ta 4 ta 5 LR BV SV BR p 1 265 300 165 100 58 28 33 0.55 3.4 3.4 1.7 p 2 375 310 210 93 45 20 33 0.68 4.7 3.3 2.7 p 3 310 315 185 100 43 20 33 0.59 4.2 3.4 2.0 Hypopygium (Figs 4 C – D): Tergite IX with posterior margin almost straight, with many short setae. Anal point present, cylindrical, 10 µm long, 10 µm wide. Superior volsella well developed, inferior volsella large, rounded posteriorly, partly united with superior volsella. Sternapodeme inverted U-shaped; coxapodeme 33 µm long, attachment point with phallapodeme placed in caudal position of sternapodeme and caudally directed; phallapodeme curved, not extending beyond posterior margin of tergite IX, 38 µm long. Gonocoxite 88 µm long. Gonostylus apically bent, 20 µm long; megaseta 5 µm long. HR 4.4, HV 6.6.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFBBC63F78D9F956FDDA5D20.taxon	materials_examined	Material examined: P. R. CHINA: Xinjiang Province: Aketao County: Kalakuli Lake 39 ° 8 ' N, 75 ° 56 ' E, alt. 4.000 m, sweep net, 1 male, 19. viii. 2002, Hongqu Tang; Xinjiang Province, Fukang County, Tianchi Lake, 44 ° 8 ' N, 87 ° 56 ' E, alt. 1.980 m, sweep net, 1 male, 1. ix. 2002, Hongqu Tang. Diagnostic characters. Total length 1.47 – 1.76, 1.60 mm. Wing length 1.15 – 1.25, 1.2 mm. Antenna with 10 flagellomeres; AR 0.94 – 1.0, 0.97. Apex of hind tibia with one nearly straight seta near spur. Posterior margin of tergite IX weakly bilobed. Inferior volsella absent, sternapodeme inverted V-shaped, attachment point with phallapodeme placed in caudal third of lateral sternapodeme and directed caudally; phallapodeme strongly curved with projection for joint with sternapodeme placed pre-lateral. Gonostylus curved apically, subapical slender.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFBBC63F78D9F956FDDA5D20.taxon	discussion	Remarks. Hirvenoja and Hirvenoja (1988) redescribed the male, pupa and larva in detail. Distribution. Bulgaria, Denmark, Finland, France, Germany, Great Britain, Ireland, Italy, Lithuania, Macedonia, Norway, Romania, Russia including Far East, Switzerland, Spain, Sweden, The Netherlands; Japan; Xinjiang Province, Palaearctic China (Saether & Spies 2004).	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFBAC63F78D9FDEAFE1C5AD8.taxon	materials_examined	Material examined. Male holotype (no. 90: 81), Japan, Honshu, Yamanashi Prefecture, Lake Yamanaka, sweep net, 13. v. 1983, M. Sasa. Diagnostic characters. The species can be separated from other members of the genus by having an antenna with 12 flagellomeres, AR 0.62 – 0.70; superior volsella not clear, nearly fused with the gonocoxite, and inferior volsella broad. Posterior margin of tergite IX almost straight, sternapodeme inverted U-shaped.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFBAC63F78D9FDEAFE1C5AD8.taxon	description	Additional description and corrections. Scutellum and postnotum brown, abdominal tergites yellowish brown while the original description Sasa (1985) mentions scutellum and postnotum entirely black, abdominal tergites I – V yellowish brown, VI to hypopygium dark brown. " R 1, R 2 + 3 and R 4 + 5 fused and shortened " should be corrected to " R 1, R 2 + 3 and costa fused. " Some additional characters should be added: Total length 1.86 – 2.13 mm. Wing length 1.14 – 1.24 mm. Antenna 12 flagellomeres, AR 0.62 – 0.70. Wing width / wing length 0.35. Clypeus with 10 setae, Tentorium 138 µm long, 23 µm wide. VR 2.7. C length 350 µm, C / wing length 0.31. Cu length 700 µm. Cu / wing length 0.62. Wing width / wing length 0.35. C with 9 setae. Posterior margin of tergite IX almost straight, with several short setae. Sternapodeme inverted U-shaped, coxapodeme 25 µm long, attachment point with phallapodeme placed in caudal position of sternapodeme and caudally directed; phallapodeme short, straight and broad, not extending beyond posterior margin of tergite IX, 50 µm long. Gonocoxite 98 µm long, with 6 long setae apically. Gonostylus 50 µm long; megaseta 10 µm long. HR 1.95, HV 3.25. The wing and hypopygium of holotype are redrawn (Figs 5 A – C).	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFBAC63F78D9FDEAFE1C5AD8.taxon	distribution	Distribution. Japan.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFBAC63D78D9F9DDFEC85EEE.taxon	materials_examined	Material examined. P. R. CHINA: Sichuan Province, Yajiang County, Sandaoqiao, 30 ° 18 ' N, 101 ° E, alt. 2460 m, light trap, 1 male (BDN No. 11608), 9. vi. 1996, Xinhua Wang; Yunnan Province, Lijiang County, Heilongtan, 26 ° 52 ' N, 100 ° 15 ' E, alt. 2.400 m, light trap, 1 male (BDN No. 10553), 28. v. 1996, Xinhua Wang; Yunnan Province, Hutiaoxia County, 30 ° N, 98 ° E, alt. 1.700 – 5.369 m, light trap, 2 males, 26. v. 1996, Xinhua Wang; Ningxia Autonomous Region, Liupanshan County, 35 ° 14 ' – 39 ° 14 ' N, 104 ° 17 ' – 109 ° 39 ' E, alt. 1.100 – 1.200 m, light trap, 1 male (BDN No. 1177), 7. viii. 1987, Xinhua Wang. Diagnostic characters. This species is similar to T. celtica Tokunaga sharing an AR of about 0.20 – 0.30, and an inverted V-shaped sternapodeme, but can be separated by the digitiform inferior volsella and the antenna with 9 flagellomeres.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFBAC63D78D9F9DDFEC85EEE.taxon	description	Addition to description. The Chinese specimens had the following measurements: Total length 1.12 – 1.46, 1.34 mm. Wing length 0.7 – 0.85, 0.76 mm. Antenna with 9 flagellomeres; AR 0.27 – 0.34, 0.3. Wing width / wing length 0.41 – 0.44, 0.41, C with 6 – 8, 7 setae. Inferior volsella digitiform, sternapodeme inverted V-shaped, coxapodeme 25 – 35, 30 µm long, attachment point with phallapodeme placed in caudal third of lateral sternapodeme and directed caudally; phallapodeme strongly curved with projection for joint with sternapodeme placed pre-lateral. Gonocoxite 58 – 78, 68 µm long, with 2 setae apically. Gonostylus curved apically, 23 – 28, 27 µm long; megaseta 3 – 5, 3 µm long. HR 2.7 – 3.6, 3.1, HV 4.5 – 5.6, 5.3 (4).	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFBAC63D78D9F9DDFEC85EEE.taxon	discussion	Remarks. Sublette and Sasa (1994) described the species based on specimens from Guatemala. The Chinese specimens show some variations when compared with the original description. The Chinese specimens are paler, and the ultimate flagellomere is slightly curved. However, some variations also exist between specimens collected in southern and northern China.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFBAC63D78D9F9DDFEC85EEE.taxon	distribution	Distribution. Guatemala; Sichuan and Yunnan Provinces, Ningxia Autonomous Region in Oriental and Palaearctic China.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFB8C63D78D9FD97FCC25B0E.taxon	materials_examined	Material examined: P. R. CHINA: Liaoning Province, Panjin County, Guchengzi, 40 ° 41 ' – 41 ° 27 ' N, 121 ° 34 ' E, alt. 50 m, 1 male (BDN no. 03374), 13. iv. 1992, Juncai Wang. Diagnostic characters. Wing length 1.0 mm. Antenna with 10 flagellomeres; apex acute, with short subapical sensilla chaetica, AR 0.92. Hind tibia without S-shaped seta. Posterior margin of tergite IX bilobed, laterosternite IX with 1 seta. Inferior volsella absent, sternapodeme inverted U-shaped, attachment point with phallapodeme placed caudal and ventrally directed; phallapodeme strongly curved.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFB8C63D78D9FD97FCC25B0E.taxon	discussion	Remarks. Schlee (1968 b) described the species based on specimen from Stuttgart, Germany. C. scutellata listed by Wang (2000) from Liaoning province was wrongly identified and should be corrected to C. gratias.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFB8C63D78D9FD97FCC25B0E.taxon	distribution	Distribution. Czech Republic, Finland, France, Germany, Norway, Russia, Slovakia, Spain, Sweden; Liaoning Province, Palaearctic China (Saether & Spies 2004).	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFB8C63D78D9FBF7FE1C58CC.taxon	materials_examined	Material examined. Male holotype (no. 389: 96), JAPAN: Honshu, Fukushima Prefecture, Lake Inawashiro, light trap, 20. viii. 1999, K. Kitami. Diagnostic characters. The species can be separated from other members of the genus by having an antenna with 10 flagellomeres; AR 0.66; superior volsella almost fused with gonocoxite, inferior volsella digitiform; sternapodeme inverted V- to U-shaped; and gonostylus with triangular crista dorsalis.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFB8C63D78D9FBF7FE1C58CC.taxon	description	Additional description and corrections. R 1, R 2 + 3 and costa fused to a short and thick vein with VR 3.2, while the original description Sasa et al. (1999) mentions R 1 and R 4 + 5 fused to a short and thick vein and VR 2.08. Apex of hind tibia with comb of 17 setae while the original description mentions apex of hind tibia with a comb of 12 setae. Gonocoxite with digitiform inferior volsella while the original description mentions gonocoxite without inner lobe, but with a small rounded and hyaline process. Some additional characters should be added: Clypeus with 12 setae. Tentorium 118 µm long, 10 µm wide. VR 3.2. C length 180 µm, C / wing length 0.27. Cu length 420 µm. Cu / wing length 0.64. Wing width / wing length 0.38. C with 12 setae. Posterior margin of tergite IX almost straight. Sternapodeme inverted V- to Ushaped, coxapodeme 38 µm long, attachment point with phallapodeme placed in caudal third of lateral sternapodeme and directed caudally; phallapodeme weakly curved, extending beyond posterior margin of tergite IX, 65 µm long. Gonocoxite 85 µm long, with 5 long setae apically. Gonostylus 33 µm long, megaseta 3 µm long. HR 2.6, HV 3.7. The wing and hypopygium of holotype are redrawn (Figs 6 A – C).	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFB8C63D78D9FBF7FE1C58CC.taxon	distribution	Distribution. Japan.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFBFC63878D9F9AAFACD5C79.taxon	materials_examined	Type material: Holotype male (BDN no. 03375), P. R. CHINA: Liaoning Province, Kuandian County, 40 ° 45 ' N, 124 ° 46 ' E, alt. 400 – 1336 m, light trap, 22. ix. 1992, Xinhua Wang.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFBFC63878D9F9AAFACD5C79.taxon	etymology	Etymology. From Latin, inefligiata, formless, shapeless, referring to the large, broad inferior volsella with dented edge. Diagnostic characters. Similar to C. yoshimurai in coloration and in having antenna with 10 flagellomeres, but can be separated by the weak anal lobe of wing, phallapodeme with small inner projection and the broad inferior volsella with dented edge.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFBFC63878D9F9AAFACD5C79.taxon	description	Description Male (n = 1). Total length 1.45 mm. Wing length 0.88 mm. Total length / wing length 1.65. Wing length / length of profemur 3.45. Coloration: Head dark brown. Antenna and palpomere yellow. Thorax brown. Abdominal segments yellow to brown. Legs yellowish brown. Wings light yellow to hyaline. Head: Antenna (Fig. 7 B) with 10 flagellomeres; ultimate flagellomere 130 µm long; apex with rosette of short sensilla chaetica; apically expanded, thickening 70 µm long, 30 µm wide; AR 0.43. Clypeus with 6 setae. Tentorium (Fig. 7 E) 125 µm long, 25 µm wide. Stipes 50 µm long. Palpomere lengths (in µm): 15; 13; 18; 25; 50. Palpomere 2 and 3 ellipsoid, 4 rectangular, 5 long and slender. Palpomere 5 / 3: 2.86. Thorax: Damaged. Wing (Fig. 7 A): Anal lobe weak. R 4 + 5 clearly separate from M 1 in full length. VR 3.4. C length 220 µm. Cu / wing length 0.6. Wing width / length 0.44. C with 7 setae. Legs: Spur of fore tibia 27 µm long, spurs of mid tibia 10 and 13 µm long, and of hind tibia 30 µm long. Width at apex of fore tibia 20 µm, of mid tibia 20 µm, and of hind tibia 38 µm. Apex of hind tibia expanded, with comb of 12 setae and one hook-like seta near spur. Lengths and proportions of legs as in Table 3. Hypopygium (Figs 7 C – D): Posterior margin of tergite IX straight. Anal point absent. Inferior volsella large and broad with dented edge. Sternapodeme inverted U-shaped, attachment point with phallapodeme placed in caudal position of sternapodeme and ventrally directed; phallapodeme weakly curved, not extending beyond posterior margin of tergite IX, with small inner projection. Gonocoxite 70 µm long. Gonostylus slightly curved apically, 35 µm long; megaseta 3 µm long. HR 2.0, HV 4.1. TABLE 3. Lengths (in µm) and proportion of leg segments of Corynoneura inefligiata sp. n., male. fe ti ta 1 ta 2 ta 3 ta 4 ta 5 LR BV SV BR p 1 255 325 158 93 60 30 35 0.49 3.0 3.7 2.0 p 2 370 340 195 93 50 23 35 0.57 4.5 3.6 1.6 p 3 300 360 193 110 45 25 35 0.54 3.8 3.4 1.5	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFBDC63878D9FC4AFE1C599B.taxon	materials_examined	Material examined: JAPAN: Honshu, Kyoto Prefecture, Kibune River, sweep net, male holotype (No. 166: 61), 11. x. 1988, M. Sasa. Diagnostic characters. The species can be separated from other members of the genus by having an antenna with 10 flagellomeres; AR 0.31; superior volsella (not clear in the available material) probably fused with the gonocoxite; inferior volsella broad and with a small prominent part; sternapodeme inverted Ushaped; and gonostylus medially expanded.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFBDC63878D9FC4AFE1C599B.taxon	description	Additional description and corrections. The holotype possesses 10 flagellomeres and an AR of 0.31, while the original description by Sasa (1989) mentions antenna with 9 flagellomeres and an AR of 0.43 – 0.56. Some additional characters should be added: Tentorium 125 µm long, 23 µm wide. Posterior margin of tergite IX almost straight. Sternapodeme inverted U-shaped, coxapodeme 38 µm long, small attachment point with phallapodeme placed in caudal position of sternapodeme and caudally directed; phallapodeme short, weakly curved, not extending beyond margin of tergite IX, 35 µm long. Gonocoxite 68 µm long, with 2 long setae apically. Gonostylus 35 µm long; megaseta 5 µm long. HR 1.9, HV 3.5. The wing and the hypopygium of the holotype are redrawn (Figs 8 A – C).	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFBDC63878D9FC4AFE1C599B.taxon	distribution	Distribution. Japan.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFBCC62778D9F887FF1E5E5D.taxon	materials_examined	Material examined: JAPAN: Honshu, Kyoto Prefecture, Kibune River, sweep net, male holotype (No. 166: 50 B), 11. x. 1988, M. Sasa. Diagnostic characters. The species can be separated from other East Asian Corynoneura by having an antenna with 10 flagellomeres, AR 0.27; posterior margin of tergite IX slightly bilobed, sternapodeme inverted V-shaped, and inferior volsella digitiform.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFBCC62778D9F887FF1E5E5D.taxon	description	Additional description and corrections. AR 0.27 while the original description by Sasa (1989) mentions AR 0.41. Some additional characters should be added: Tentorium 113 µm long, 15 µm wide. Posterior margin of tergite IX slightly bilobed. Sternapodeme inverted V-shaped; coxapodeme 38 µm long; attachment point with phallapodeme placed in caudal third of lateral sternapodeme and directed caudally; phallapodeme strongly curved with projection for joint with sternapodeme placed pre-lateral, extending beyond margin of tergite IX, 88 µm long. Gonocoxite 65 µm long with 3 long setae apically. Gonostylus bent, 30 µm long; megaseta 8 µm long. HR 2.2, HV 3.4. The wing and the hypopygium of the holotype are redrawn (Figs 9 A – C).	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFA1C62478D9FF1AFE1D5CA8.taxon	materials_examined	Material examined: JAPAN: Honshu, Aichi, Bisai, sweep net, male holotype (No. 222: 100), 15. xi. 1988, M. Sasa. Diagnostic characters: The species can be separated from other East Asian Corynoneura by having antenna with 12 flagellomeres, AR 0.33; phallapodeme straight and erect; sternapodeme inverted U-shaped, and no inferior volsella.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFA1C62478D9FF1AFE1D5CA8.taxon	description	Additional description and corrections. The holotype possesses 12 flagellomeres with AR 0.33, while the original description by Sasa & Kondo (1993) mentions antenna with 10 flagellomeres and AR 0.46. The total length is 1.12 mm and the wing length 0.83 mm, while the original description mentions a total length of 1.20 mm and a wing length of 0.74 mm. The leg ratio of the mid leg is 0.52, of the hind leg 0.54, while the original description mentions 0.64 and 0.46 respectively. Some additional characters should be added: Tentorium 125 µm long, 15 µm wide. Sternapodeme inverted U-shaped, coxapodeme 20 µm long, attachment point with phallapodeme placed in caudal position of sternapodeme and caudally directed; phallapodeme short and straight, 33 µm long. Gonocoxite 73 µm long. Gonostylus 38 µm long. HR 2.1, HV 3.0. The wing and hypopygium of the holotype are redrawn (Figs 10 A – C).	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFA1C62478D9FF1AFE1D5CA8.taxon	distribution	Distribution. Japan.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFA1C62278D9FC52FAC25D0F.taxon	materials_examined	Type material: Holotype male (BDN no. 1180), P. R. CHINA: Hainan Province, Bawangling County, 19 ° 5 ' N, 109 ° 8 ' E, alt. 350 – 1438 m, sweep net, 10. v. 1988, Xinhua Wang.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFA1C62278D9FC52FAC25D0F.taxon	etymology	Etymology. From Latin, korema, broom, referring to hind ta 1 having a row of stout brush-like setae. Diagnostic characters. Similar to C. prima in having antenna with 7 flagellomeres and AR about 0.5; but can be separated by the thorax with longitudinal vittae, VR 5.0, hind ta 1 with row of stout setae, and sternapodeme inverted V-shaped.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFA1C62278D9FC52FAC25D0F.taxon	description	Description Male (n = 1). Total length 0.74 mm. Wing length 0.46 mm. Total length / wing length 1.6. Wing length / length of profemur 3.0. Coloration: Head dark brown. Antenna and palpomere yellow. Thorax and legs yellowish brown. Tergites yellow to brown. Wing hyaline, with pale yellow clava. Head: Antenna (Fig. 11 B) with 7 flagellomeres; ultimate flagellomere 83 µm long, apex with short sensilla chaetica subapically, apically slightly expanded, thickening 43 µm long, AR 0.5. Tentorium (Fig. 11 E) 98 µm long, 8 µm wide. Stipes 35 µm long. Palpomeres length (in µm): 8; 10; 13; 15; 35. Palpomere 2, 3 and 4 ellipsoid, 5 long and slender, palpomere 5 / 3: 2.8. Thorax (Fig. 11 F): With two longitudinal vittae. Wing (Fig. 11 A): VR 5.0. C length 100 µm, C / wing length 0.22. Cu length 325 µm. Cu / wing length 0.7. Wing width / wing length 0.46. C with 5 setae. Legs: Spur of fore tibia 20 µm long, of mid tibia 13 µm long, and of hind tibia 23 µm long. Width at apex of fore tibia 13 µm, of mid tibia 13 µm, of hind tibia 23 µm. Apex of hind tibia expanded, with comb of 15 setae and one seta near spur strongly S-shaped, hind ta 1 (Fig. 9 G) with row of stout setae. Lengths and proportions of legs as in Table 4. Hypopygium (Figs 11 C – D): Posterior margin straight, and with many microtriches, laterosternite IX without long seta. Anal point absent. Superior volsella not in good show. like small triangular. Inferior volsella developed. Sternapodeme inverted V-shaped, coxapodeme 25 µm long, small attachment point with phallapodeme placed in caudal third of lateral sternapodeme and directed caudally; phallapodeme strongly curved and broad with lateral apex bifid and embracing knob on lateral sternapodeme, 50 µm long, base bifurcate. Gonocoxite 63 µm long with 2 setae apically. Gonostylus small and strongly curved, 13 µm long; megaseta 3 µm long. HR 5.0, HV 5.9. TABLE 4. Lengths (in µm) and proportion of leg segments of Corynonrura korema sp. n., male. fe ti ta 1 ta 2 ta 3 ta 4 ta 5 LR BV SV BR p 1 153 170 88 53 28 13 20 0.52 3.6 3.7 – p 2 205 185 95 45 25 15 23 0.51 3.6 4.1 1.3 p 3 163 180 88 40 15 13 20 0.49 4.9 3.9 2.0	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFA7C62078D9FD12FA885ED5.taxon	materials_examined	Type material: Holotype male (BDN No. 22248), P. R. CHINA: Guizhou Province, Daozhen County, Dashahe, 29 ° 00 ' – 29 ° 13 ' N, 107 ° 22 ' – 107 ° 48 ' E, alt. 1.300 – 1.900 m, sweep net, 25. v. 2004, Hongqu Tang. Paratypes: 8 males, Guizhou Province, Daozhen County, Dashahe, 29 ° 00 ' – 29 ° 13 ' N, 107 ° 21 ' – 107 ° 47 ' E, alt. 1.300 – 1.900 m, sweep net, 25. v. 2004, Hongqu Tang.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFA7C62078D9FD12FA885ED5.taxon	etymology	Etymology. From Latin, latus, broad, wide; and atra, black, referring to the antenna being expanded and black distally Diagnostic characters. Similar to C. secunda in having sternapodeme inverted strongly V-shaped, and phallapodeme well developed and strongly curved, but the two species can easily be separated as C. latusatra has 7 flagellomeres and an AR of 0.31.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFA7C62078D9FD12FA885ED5.taxon	description	Description. Male (n = 9). Total length 0.78 – 0.91, 0.85 mm. Wing length 0.55 – 0.60, 0.51 mm. Total length / wing length 1.2 – 1.59, 1.42. Wing length / length of profemur 3.14 – 3.43, 3.30. Coloration: Head yellowish brown with brown frontal vertex. Antenna pale yellow with black apex, palpomere pale yellow. Thorax pale brown to hyaline with vittae, scutum, scutellum and postnotum brown, anterior anepisternum II, preepisternum, epimeron II light brown. Tergites I – IV yellow, tergites V – IX brown. Legs yellow. Wing yellowish to hyaline. Head: Antenna (Fig. 12 B) with 7 flagellomeres; ultimate flagellomere 53 – 60, 57 µm long, antenna with rosette of apical sensilla chaetica, apically expanded, thickening 50 – 56, 53 µm long, with maximum width 18 – 25, 20 µm; AR 0.28 – 0.33, 0.31 (7). Clypeus with 8 – 12, 9 (6) setae. Tentorium (Fig. 12 E) 93 – 105, 97 µm long. Stipes 40 – 50, 45 µm long. Palpomeres length (in µm): 10 – 15, 12; 10 – 15, 13; 13 – 18, 14; 13 – 20, 15; 28 – 38, 35. Palpomere 2, 3 and 4 ellipsoid, palpomere 5 rectangular, Palpomere 5 / 3: 1.7 – 3.0, 2.5. Thorax: Dorsocentrals 3 – 5, 4 (5), prealars 2 – 5, 3. Scutellum with 1 – 2, 1 setae. Wing (Fig. 10 A): VR 2.9 – 3.6, 3.2. C length 120 – 150, 140 µm, C / wing length 0.22 – 0.25, 0.24. Cu length 295 – 330, 321 µm. Cu / wing length 0.54 – 0.58, 0.56. Wing width / wing length 0.4 – 0.43, 0.42. C with 3 setae. Legs: Spurs of fore tibia 13 – 20, 16 µm and 9 – 15, 11 (5) µm long, spurs of mid tibia 6 – 10, 8 (7) µm and 5 – 8, 6 (3) µm long, and of hind tibia 25 – 30, 27 µm and 23 (2) µm long. Width at apex of fore tibia 13 – 20, 15 µm, of mid tibia 10 – 18, 13 µm, of hind tibia 28 – 33, 30 µm. Apex of hind tibia expanded, with comb of 14 – 17, 15 setae and one hook-like seta near spur. Lengths and proportions of legs as in Table 5. Hypopygium (Figs 10 C – D): Tergite IX large, covering almost the whole gonocoxite, with weakly bilobed posterior margin, and many short setae. Laterosternite IX with 1 long seta. Superior volsella narrow. Inferior volsella absent. Sternapodeme inverted strongly V-shaped, apically acute, 63 – 68, 65 (6) µm long; coxapodeme 13 – 20, 15 (6) µm long; attachment point with phallapodeme placed in caudal third of lateral sternapodeme and directed caudally; phallapodeme well developed, strongly curved with projection for joint with sternapodeme placed pre-lateral, not extending beyond tergite IX, 85 – 90, 86 µm long, 8 – 13, 10 (6) µm wide. Gonocoxite 45 – 55, 50 µm long, with 2 setae apically. Gonostylus strongly curved, 18 – 25, 20 µm long; megaseta 5 – 8, 6 µm long. HR 2 – 2.9, 2.3, HV 3.4 – 4.4, 3.9 (6). TABLE 5. Lengths (in µm) and proportion of leg segments of Corynoneura latusatra sp. n., male. fe ti ta 1 ta 2 ta 3 ta 4 p 1 165 – 185, 176 213 – 235, 226 95 – 105, 101 45 – 58, 51 20 – 35, 22 18 – 23, 20 p 2 223 – 245, 234 203 – 233, 218 108 – 120, 115 50 – 58, 54 25 – 30, 28 15 – 20, 18 p 3 175 – 200, 192 155 – 225, 208 88 – 103, 98 48 – 58, 54 20 – 25, 23 13 – 18, 14 ta 5 LR BV SV BR p 1 25 – 30, 28 0.43 – 0.47, 0.45 3.6 – 4.5, 3.9 3.8 – 4.2, 3.9 1.3 – 1.8, 1.5 p 2 25 – 28, 26 0.52 – 0.55, 0.53 4.4 – 4.7, 4.5 3.8 – 4.1, 4.0 1.2 – 1.5, 1.3 p 3 23 – 30, 26 0.43 – 0.58, 0.48 3.7 – 4.6, 4.1 3.7 – 4.3, 4.0 0.9 – 1.2, 1.0	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFA5C62078D9FDD8FD2D5AF6.taxon	materials_examined	Material examined: CHINA: Liaoning Province, Kuandian County, 40 ° 45 ' N, 124 ° 46 ' E, alt. 400 – 1.336 m, light trap, 1 male (BDN No. 07197), 22. iv. 1992, Xinhua Wang; Ningxia Autonomous Region, Liupanshan County, 35 ° 14 ' – 39 ° 14 ' N, 104 ° 17 ' – 109 ° 39 ' E, alt. 1.100 – 1.200 m, light trap, 1 male (BDN No. 05345), 7. viii. 1987, Xinhua Wang; Sichuan Province, Daocheng County, Sangdui, 29 ° 11 ' N, 100 ° 06 ' E, alt. 3943 m, light trap, 1 male (BDN No. 13410), 11. vi. 1996, Xinhua Wang. Diagnostic characters. Total length 1.34 – 1.65, 1.50 (2) mm. Wing length 0.85 – 1.05, 0.95 mm. Antenna with 10 flagellomeres, apex with short sensilla chaetica subapically, apically acute. AR 0.35 – 0.43, 0.39. C / wing length 0.25 – 0.29. Posterior margin of tergite IX weakly bilobed, inferior volsella digitiform, sternapodeme inverted V-shaped, attachment point with phallapodeme placed in caudal third of lateral sternapodeme and directed caudally; phallapodeme strongly curved with projection for joint with sternapodeme placed pre-lateral. Gonostylus curved apically.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFA5C62078D9FDD8FD2D5AF6.taxon	discussion	Remarks. Edwards (1924) established this species based on specimens from England. Sasa 1988; Sasa & Suzuki (1998, 1999) recorded the species from Japan. Schlee (1968 b) redescribed the male in detail based on material from Germany. Tokunaga (1936) redescribed the male and female based on Japanese specimen.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFA5C62078D9FDD8FD2D5AF6.taxon	distribution	Distribution. Austria, Belgium, Denmark including the Faeroes, Estonia, Finland, France, Germany, Great Britain, Ireland, Italy, Luxembourg, Macedonia, Norway, Poland, Portugal, Romania, Russia, Slovakia, Spain, Sweden, Switzerland, The Netherlands; the Near East; North Africa; Canada, USA; Japan; Liaoning Province and Ningxia Autonomous Region in Palaearctic China and Sichuan Province in Oriental China (Oliver et al. 1990, Saether & Spies 2004).	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFA5C62E78D9F9BFFAC25C2D.taxon	materials_examined	Type material: Holotype male (BDN No. 12635) P. R. CHINA: Sichuan Province, Ganzi State, Yajiang County, 31 ° 38 ' N, 99 ° 58 ' E, alt. 2000 m, light trap, 14. vi. 1996, Xinhua Wang.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFA5C62E78D9F9BFFAC25C2D.taxon	etymology	Etymology. Named after McDonald's hamburger stores, whose brand like “ M ” resembles the M-shaped sternapodeme. Diagnostic characters. The species is similar to C. lobata in having antenna with 10 flagellomeres and AR about 0.35, but can easily be separated by the subrectangular inferior volsella with round corner; and the inverted M-shaped sternapodeme.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFA5C62E78D9F9BFFAC25C2D.taxon	description	Description Male (n = 1). Total length 1.05 mm. Wing length 0.80 mm. Total length / wing length 1.30. Wing length / length of profemur 3.27. Coloration: Head dark brown. Antenna and palpomere yellow. Thorax brown with yellow anterior anepisternum II. Legs yellowish brown. Tergites I – V yellow, tergites VI – IX brown. Wings yellowish to hyaline, with pale yellow clava. Head: Antenna (Fig. 13 B) with 10 flagellomeres; ultimate flagellomere 113 µm long, apex with rosette of apical sensilla chaetica, slightly expanded apically. AR 0.36. Clypeus with 9 setae. Tentorium (Fig. 13 E) 118 µm long, 18 µm wide. Stipes 45 µm long. Palpomeres length (in µm): 13; 13; 18; 20; 53. Palpomere 2, 3 and 4 ellipsoid, 5 long. Palpomere 5 / 3: 3.0. Thorax: Dorsocentrals 5. Wing (Fig. 13 A): VR 3.2. C length 190 µm, C / wing length 0.24. Cu length 450 µm. Cu / wing length 0.56. Wing width / wing length 0.43. C with 5 setae. Legs: Spur of fore tibia 20 µm long, of mid tibia 8 µm long, and of hind tibia 15 µm long. Width at apex of fore tibia 20 µm, of mid tibia 15 µm, of hind tibia 38 µm. Apex of hind tibia expanded, with comb of 16 setae, one seta near spur strongly S-shaped. Lengths and proportions of legs as in Table 6. Hypopygium (Figs 13 C – D): Tergite IX large, almost covering gonocoxite. Posterior margin of tergite IX slightly bilobed, with 4 setae on each side; laterosternite IX with 1 long seta. Superior volsella broad, partly united with inferior volsella. Inferior volsella subrectangular, with round corners. Sternapodeme inverted Mshaped; coxapodeme 25 µm long, small attachment point with phallapodeme placed in caudal position of sternapodeme and ventrally directed; phallapodeme short, weakly curved, 24 µm long, with medial projection. Gonocoxite 70 µm long with 2 setae apically. Gonostylus slender, 28 µm long, with single setae near apex; megaseta 3 µm long. HR 2.5, HV 3.8. fe ti ta 1 ta 2 ta 3 ta 4 ta 5 LR BV SV BR p 1 233 290 155 80 50 25 28 0.53 3.7 3.4 1.7 p 2 330 280 193 83 40 20 28 0.69 4.7 3.2 2.2 p 3 265 280 155 80 38 23 30 0.55 4.1 3.5 2.0	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFABC62F78D9FCFEFACD5EFD.taxon	materials_examined	Type material: Holotype male (BDN No. 05368), P. R. CHINA: Sichuan Province, Emeishan County, Medicine School, 29 ° 21 ' N, 103 ° 17 ' E, alt. 1.500 m, sweep net, 13. v. 1986, Xinhua Wang.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFABC62F78D9FCFEFACD5EFD.taxon	etymology	Etymology. The name refers to the type locality, the Emeishan Medicine School of China. Diagnostic characters. Similar to C. scutellata in having antenna with 10 flagellomeres and AR 0.74, but can be separated by the inverted U-shaped sternapodeme and lack of inferior volsella.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFABC62F78D9FCFEFACD5EFD.taxon	description	Description Male (n = 1). Total length 1.08 mm. Wing length 0.87 mm. Total length / wing length 1.24. Wing length / length of profemur 3.35. Coloration: Head dark brown. Antenna and palpomere yellow. Thorax yellowish brown with brown vittae, scutum, scutellum, preepisternum, postnotum and yellow anterior anepisternum II, median anepisternum II, posterior anepisternum II. Abdominal segments brown. Legs yellowish brown. Wing light yellow to hyaline, with pale yellow clava. Head: Antenna with 10 flagellomeres; ultimate flagellomere 215 µm long, apex with short sensilla chaetica subapically, apically acute. AR 0.74. Clypeus with 4 setae. Tentorium 120 µm long, 13 µm wide. Stipes 55 µm long. Palpomeres length (in µm): 13; 13; 23; 33; 48. Palpomere 2 and 3 ellipsoid, 4 rectangular, 5 curved. Palpomere 5 / 3: 2.1. Wing (Fig. 12 A): VR 2.6. C length 240 µm, C / wing length 0.3. Cu length 520 µm. Cu / wing length 0.6. Wing width / wing length 0.37. C with 8 setae. Legs: Spurs of fore tibia 20 and 13 µm long, of mid tibia 10 µm long, and of hind tibia 33 µm long. Width at apex of fore tibia 18 µm, of mid tibia 18 µm, of hind tibia 30 µm. Apex of hind tibia expanded, with comb of 12 setae and one setae near spur near straight. Lengths and proportions of legs as in Table 7. Hypopygium (Figs 14 B – C): Posterior margin of tergite IX bilobed, 5 – 6 setae on each side; laterosternite IX with 2 setae. Anal point absent. Superior volsella almost fused anteriomedially. Inferior volsella absent. Sternapodeme inverted U-shaped, 33 µm wide; coxapodeme 25 µm long, attachment point with phallapodeme placed in caudal third of lateral sternapodeme and directed caudally; phallapodeme well developed, strongly curved with projection for joint with sternapodeme placed pre-lateral, not extending beyond margin of tergite IX, 55 µm long, 3 µm wide. Gonocoxite 63 µm long with 2 setae apically. Gonostylus hooked, 30 µm long, with basal lobe on inner margin; megaseta 5 µm long. HR 2.1, HV 3.6. TABLE 7. Lengths (in µm) and proportion of leg segments of Corynoneura medicinea sp. n., male. fe ti ta 1 ta 2 ta 3 ta 4 ta 5 LR BV SV BR p 1 260 325 185 90 48 20 32 0.57 4.1 3.0 2.0 p 2 365 335 195 88 44 22 34 0.58 4.8 3.8 2.0 p 3 285 340 175 85 30 15 35 0.51 4.8 3.6 1.7	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFA9C62D78D9FF1AFDBB595D.taxon	materials_examined	Type material: Holotype male (BDN No. 05361), CHINA: Tianjin City, Nankai University, 39 ° 8 ' N, 117 ° 1 ' E, alt. 2 – 5 m, sweep net, 20. iv. 1986, Xinhua Wang. Paratypes: 3 males as holotype (BDN slides no. 0 5356, 0 5351, 05357).	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFA9C62D78D9FF1AFDBB595D.taxon	etymology	Etymology. Named after the type locality. Diagnostic characters. Similar to C. scutellata in having antenna with 10 flagellomeres, inferior volsella digitiform, and sternapodeme inverted V-shaped, but can be separated by tergite IX with a posterior hyaline structure and anal point.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFA9C62D78D9FF1AFDBB595D.taxon	description	Description Male (n = 4). Total length 1.32 – 1.82, 1.63 mm. Wing length 0.95 – 1.05, 1.02 mm. Total length / wing length 1.3 – 1.9, 1.6. Wing length / length of profemur 2.6 – 3.3, 3.0. Coloration: Head blackish brown. Antenna and palpomere pale yellowish brown. Thorax and abdomen brown. Legs yellowish brown. Wings yellowish to hyaline. Head: Antenna with 10 flagellomeres; ultimate flagellomere 235 – 310, 268 µm long, apex with short sensilla chaetica subapically, apically acute. AR 0.72 – 0.94, 0.84. Clypeus with 6 – 10, 8 (3) setae. Tentorium (Fig. 15 E) 103 – 148, 129 (3) µm long, 13 – 20, 17 (3) µm wide. Stipes 65 – 75, 70 (3) µm long. Palpomeres length (in µm): 13 – 18, 15; 20 – 23, 21; 23 – 28, 26; 18 – 35, 30; 68 (2). Palpomere 2 and 3 ellipsoid, 4 rectangular, 5 long and slender. Palpomere 5 / 3: 2.45 – 3.0, 2.7. Thorax: Dorsocentrals 6 – 7. Wing (Fig. 15 A): VR 2.6 – 3.0, 2.8. C length 300 – 400 µm, C / wing length 0.29 – 0.4, 0.33 (3). Cu length 600 – 710, 697 µm. Cu / wing length 0.57 – 0.71, 0.64. Wing width / wing length 0.32 – 0.34, 0.33 (2). C with 7 setae. Legs: Spurs of fore tibia 25 – 35, 31 µm and 15 – 18, 16 µm long, of mid tibia 13 – 20, 15 (3) µm long, and of hind tibia 28 – 35, 31 µm long. Width at apex of fore tibia 25 – 30, 27 µm, of mid tibia 23 – 28, 25 µm, of hind tibia 45 – 50, 48 µm. Apex of hind tibia expanded, with comb of 12 – 18, 14 setae and one seta near spur strongly S-shaped. Lengths and proportions of legs as in Table 8. Hypopygium (Figs 15 C – D): Tergite IX with posterior margin slightly bilobed, with a hyaline structure, each lobe with about 4 setae, anal point present. Laterosternite IX with 2 setae. Inferior volsella digitiform, with glandular setae; sternapodeme inverted V-shaped; coxapodeme 45 – 55, 50 µm long, attachment point with phallapodeme placed in caudal third of lateral sternapodeme and directed caudally; phallapodeme strongly curved with projection for joint with sternapodeme placed pre-lateral, almost extending beyond tergite IX, 80 – 85, 83 µm long, with projection basally. Gonocoxite 88 – 100, 91 µm long with 2 setae apically. Gonostylus strongly curved, 18 – 25, 22 (4) µm long, with protuberance basally; megaseta 8 – 11 µm long. HR 2.9 – 4.0, 3.3, HV 4.4 – 4.6, 4.5 (2).	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFA8C62B78D9F940FB135AA2.taxon	materials_examined	Type material: Holotype male (BDN No. 24734), P. R. CHINA: Jiangxi Province, Wuyi Mountain, 27 ° 28 ' N, 118 ° 1 ' E, alt. 400 m, sweep net, 15. vi. 2004, Chuncai Yan. Paratypes: 2 males, as holotype (BDN No. 24583, 24735); 1 male (BDN No. 05363), Chongqi City, Nanchuan County, peak of Jinfo Mountain, 29 ° 9 ' N, 107 ° 8 ' E, alt. 1.200 m, sweep net, 9. v. 1986, Xinhua Wang; 1 male (BDN No. 25152), Hunan Province, Hengyang County, Heng Mountain, 112 ° 52 ' E, 27 ° 15 ' N, alt. 1.290 m, sweep net, 20. vii. 2004, Chuncai Yan.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFA8C62B78D9F940FB135AA2.taxon	etymology	Etymology. From Latin, prominens, projecting, standing out, referring to the broad and protruding inferior volsella. Diagnostic characters. Similar to C. prima in having antenna with 8 flagellomeres, but can be separated by having an anal point.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFA8C62B78D9F940FB135AA2.taxon	description	Description Male (n = 5). Total length 0.82 – 0.95, 0.91 (4) mm. Wing length 0.50 – 0.64, 0.57 (4) mm. Total length / wing length 1.5 – 1.55, 1.53 (3). Wing length / length of profemur 2.7 – 3.0, 3.0 (3). Coloration: Head yellow brown with brown frontal vertex. Antenna and palpomere pale yellowish brown. Thorax yellowish brown, abdomen brown. Legs yellow. Wings light yellow to hyaline. Head: Antenna with 8 flagellomeres; ultimate flagellomere 83 – 133, 108 (4) µm long, antenna apically expanded, with rosette of apical sensilla chaetica, thickening 55 – 60, 57 µm long, with maximum width 15 – 20, 18 µm; AR 0.42 – 0.60, 0.52 (4). Clypeus with 4 – 6, 5 (3) setae. Tentorium (Fig. 16 E) 92 – 115, 105 µm long, 10 – 15, 13 µm wide. Stipes 35 – 40, 38 (3) µm long. Palpomeres length (in µm): 10; 10 – 13, 11; 13 – 18, 15; 18 – 20, 19; 33 – 43, 39 (4); Palpomere 2, 3 and 4 ellipsoid, palpomere 5 long and slender. Palpomere 5 / 3: 2.1 – 2.8, 2.6 (4). Thorax: Dorsocentrals 5. Wing (Fig. 16 A): VR 3.2 – 3.4, 3.3 (3). C length 100 – 140 µm, C / wing length 0.20 – 0.23, 0.22 (4). Cu length 310 – 375, 337 (3) µm. Cu / wing length 0.53 – 0.59, 0.57. Wing width / wing length 0.44 – 0.48, 0.46. C with 5 – 7, 6 setae. Leg: Spurs of fore tibia 10 – 18, 13 (4) µm and 8 – 10, 8 (2) µm long; of mid tibia 8 – 10, 8 (2) µm long; and of hind tibia 25 – 35, 29 µm long. Width at apex of fore tibia 13 – 18, 15 µm; of mid tibia 13 – 20, 15 µm; of hind tibia 23 – 33, 28 µm. Apex of hind tibia expanded, with comb of 14 – 16, 15 setae and one seta near spur strongly S-shaped. Lengths and proportions of legs as in Table 9. fe ti ta 1 ta 2 ta 3 ta 4 p 1 175 – 218, 197 210 – 240, 227 105 – 125, 114 55 – 75, 68 28 – 43, 35 13 – 25, 19 p 2 235 – 290, 260 210 – 260, 229 110 – 165, 135 58 – 73, 64 30 – 38, 33 15 – 23, 24 p 3 198 – 245, 215 200 – 275, 230 100 – 130, 110 60 – 70, 64 25 – 33, 28 15 – 20, 18 ta 5 LR BV SV BR p 1 20 – 30, 26 0.48 – 0.52, 0.5 3.3 – 4.6, 3.8 3.6 – 3.9, 3.7 1.5 – 2.3, 1.8 p 2 25 – 28, 26 0.51 – 0.63, 0 69 4.2 – 4.9, 4.5 3.3 – 3.7, 3.5 1.7 – 2.0, 1.85 p 3 20 – 30, 25 0.48 – 0.51, 0.49 3.8 – 4.2, 4.0 3.8 – 4.0, 3.9 1.6 – 1.8, 1.7 Hypopygium (Figs 16 C – D): Tergite IX posterior margin slightly bilobed, laterosternite IX with 1 long seta. Anal point present. Superior volsella conspicuously projecting; inferior volsella broad and protruding, with glandular setae. Sternapodeme inverted U-shaped; coxapodeme 18 µm long, attachment point with phallapodeme placed in caudal position of sternapodeme and caudally directed; phallapodeme weakly curved, not extending beyond tergite IX, 30 – 35, 33 µm long, with projection basally. Gonocoxite 50 – 65, 58 µm long with 1 seta apically. Gonostylus strongly curved, 18 – 25, 22 (4) µm long; megaseta 3 – 5, 4 µm long. HR 2.0 – 3.4, 2.7 (4), HV 3.8 – 4.2, 4.0.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFA8C62B78D9F940FB135AA2.taxon	discussion	Remarks. Anal point very small and paler, several of the paratypes hard to examined.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFAEC62878D9FA56FC0D5EC9.taxon	materials_examined	Material examined: P. R. CHINA: Fujian Province, Liancheng County, Guanzhi Mountain, 25 ° 43 ' N, 116 ° 45 ' E, alt. 660 m, sweep net, 1 male, 4. v. 1993, Xinhua Wang; Yunnan Province, Lijiang County, Heilongtan, 26 ° 52 ' N, 100 ° 15 ' E, alt. 2.400 m, sweep net, 1 male, 28. v. 1996, Xinhua Wang; Tianjin City, Ji County, 40 ° 3 ' N, 117 ° 24 ' E, alt. 857 m, sweep net, 1 male, 15. ix. 1987, Xinhua Wang; Tianjin City, Nankai University, 39 ° 8 ' N, 117 ° 12 ' E, alt. 2 – 5 m, sweep net, 6 males, 20. iv. 1986, Xinhua Wang. Diagnostic characters. Total length 1.07 – 1.82, 1.50 mm. Wing length 0.85 – 1.25, 1.03 mm. Antenna with 10 flagellomeres; apically acute; AR 0.74 – 1.1, 0.82. Posterior margin of tergite IX bilobed. Inferior volsella digitiform, with small inner lobe in middle part. Sternapodeme inverted V-shaped, attachment point with phallapodeme placed in caudal third of lateral sternapodeme and directed caudally, phallapodeme strongly curved with projection for joint with sternapodeme placed pre-lateral. Gonostylus curved apically.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFAEC62878D9FA56FC0D5EC9.taxon	discussion	Remarks. Winnertz (1846) established this species based on specimens from Poland. Hirvenoja and Hirvenoja (1988) redescribed the male, pupa and larva in detail. Schlee (1968 b) redescribed the male based on specimens from Germany. Wang (2000: 635) listed this species from Liaoning Province, China.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFAEC62878D9FA56FC0D5EC9.taxon	distribution	Distribution. Austria, Belgium, Bulgaria, Czech Republic, Croatia, Denmark, Estonia, Finland, Germany, Great Britain, Hungary, Ireland, Italy, Luxembourg, Macedonia, Moldova, Norway, Poland, Romania, Russia including Novaya Zemlya and The Far East, Slovakia, Spain, Sweden, Switzerland, The Netherlands, Turkey; Near East; Greenland, Canada, USA; Australia; Chile; Japan, Russian Far East and Tianjin, Fujian, and Yunnan Provinces, China. (Oliver et al. 1990, Saether & Spies 2004). The species thus is recorded from all zoogeographical regions except the Afrotropical.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFACC62978D9FF1AFE1C5C16.taxon	materials_examined	Material examined: JAPAN: Shikoku, Kochi Prefecture, Ekawasaki, Nishitosa-Mura, sweep net, male holotype (No. 358: 81), 26. iv. 1998, H. Suzuki. Diagnostic characters. The species can be separated from other East Asian Corynoneura by having antenna 8 flagellomeres, AR 0.47; phallapodeme short, almost straight; sternapodeme inverted U-shaped; and gonostylus apically curved.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFACC62978D9FF1AFE1C5C16.taxon	description	Additional description and corrections. The holotype possesses 8 flagellomeres, AR 0.47 and leg ratio of the mid leg of 0.44; while the original description Sasa et al. (1998) mentions antenna with only 7 flagellomeres, AR 0.69 and mid leg ratio of 0.51. The following characters should be added: Tentorium 100 µm long, 15 µm wide. Sternapodeme inverted U-shaped; coxapodeme 25 µm long; attachment point with phallapodeme placed in caudal position of sternapodeme and caudally directed; phallapodeme short, straight and broad, 20 µm long. Gonocoxite 63 µm long with 2 long setae apically. Gonostylus 20 µm long; megaseta 8 µm long. HR 3.1, HV 5.75. The wing and hypopygium of the holotype are redrawn (Figs 17 A – C).	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFACC62978D9FF1AFE1C5C16.taxon	distribution	Distribution. Japan.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFACC62978D9FC34FE1C59E0.taxon	materials_examined	Material examined. JAPAN: Hokkaido, Sorachi River, Sorachi, sweep net, male holotype (No. 404: 69), 5. ix. 2000, H. Suzuki. Diagnostic characters. The species can be separated from other East Asian Corynoneura by having antenna with 10 flagellomeres, apically acute, AR 0.56 – 0.61, sternapodeme inverted V-shaped, phallapodeme strongly curved, no inferior volsella, gonostylus with small inner lobe in subapical part.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFACC62978D9FC34FE1C59E0.taxon	description	Additional description and corrections. Abdominal tergites with few setae, 0 on T I, 2 on each of T II – IX, while in the original description Sasa & Suzuki (2001) mentions abdominal tergites with small number of setae, 0 on I, 1 on II to IV, 2 on VI and VII, 1 on VIII, and 2 on IX. The ninth tergite without an anal point like Y-shaped ridge on the ventral side. The ventral Y-shaped ridge mentioned by Sasa apparently is the phallapodeme. Some additional characters should be added: Tentorium 125 µm long, 13 µm wide. Sternapodeme inverted V-shaped; coxapodeme 30 µm long; attachment point with phallapodeme placed in caudal position of sternapodeme and ventrally directed; phallapodeme 75 µm long, strongly curved with projection for joint with sternapodeme placed pre-lateral. Gonocoxite 80 µm long with 4 long setae apically. Gonostylus 38 µm long; megaseta very short. HR 2.1, HV 3.3. The wing and hypopygium of holotype are redrawn (Figs 18 A – C).	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FFACC62978D9FC34FE1C59E0.taxon	distribution	Distribution. Japan.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FF93C61778D9F8C5FE1C5DE1.taxon	materials_examined	Material examined: JAPAN: Kagoshima Prefecture, Nakanoshima Island, sweep net, male holotype (No. 289: 36), 20. v. 1994, H. Suzuki. Diagnostic characters. The species can be separated from other East Asian Corynoneura by having antenna with 11 flagellomeres, AR 0.62; sternapodeme inverted U-shaped; phallapodeme short, almost straight; gonocoxite with well triangular superior volsella; and nearly rectangular inferior volsella.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FF93C61778D9F8C5FE1C5DE1.taxon	description	Additional description and corrections. The anal point is absent and the gonocoxite has a well developed superior volsella and a rectangular inferior volsella, while the original description by Sasa & Suzuki (1995) mentions a triangular anal point and a gonocoxite with two inner lobes. The original description mentions a mid leg ratio of 0.57 while we found 0.61. Some additional characters should be added: Tergites I – V yellow, tergites VI – IX brown. Tentorium 125 µm long, 18 µm wide. Sternapodeme inverted U-shaped; coxapodeme 25 µm long; weak attachment point with phallapodeme placed in caudal position of sternapodeme and caudally directed; phallapodeme 35 µm long, weakly curved. Gonocoxite 65 µm long, with 2 long setae apically. Gonostylus 33 µm long; megaseta 8 µm long. HR 2.0, HV 4.4. Wing damaged. Hypopygium of holotype redrawn (Figs 19 A – B).	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FF93C61778D9F8C5FE1C5DE1.taxon	distribution	Distribution. Japan.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FF92C61578D9F964FE1C5E5D.taxon	materials_examined	Material examined: JAPAN: Kagoshima Prefecture, Nakanoshima Island, sweep net, male holotype (No. 289: 36), 20. v. 1994, H. Suzuki. Diagnostic characters. The species can be separated from other East Asian members of the genus by having antenna with 11 flagellomeres, apically acute, AR of about 0.5, inferior volsella reduced, sternapodeme inverted U-shaped, and gonostylus with inner lobe.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FF92C61578D9F964FE1C5E5D.taxon	description	Additional description and corrections. Antennal with short, subapical sensilla chaetica in lieu of no sensory setae as stated by Sasa & Suzuki (1995). The leg ratio of the mid leg is 0.63 while the original description mentions a leg ratio of 0.60. Some additional characters should be added: Tentorium 130 µm long, 15 µm wide. VR 3.4. C length 190 µm, C / wing length 0.27. Cu length 425 µm. Cu / wing length 0.60. Wing width / wing length 0.41. C with 5 setae. Sternapodeme inverted U-shaped; coxapodeme 45 µm long; attachment point with phallapodeme placed in caudal third of lateral sternapodeme and directed caudally; phallapodeme 75 µm long, strongly curved with projection for joint with sternapodeme placed pre-lateral. Gonocoxite 88 µm long, with 3 long setae apically. Gonostylus 38 µm long, megaseta 8 µm long. HR 2.3. The wing and hypopygium of holotype are redrawn (Figs 20 A – C).	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FF92C61578D9F964FE1C5E5D.taxon	distribution	Distribution. Japan.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FF90C61578D9FE40FB565A06.taxon	materials_examined	Material examined: P. R. CHINA: Sichuan Province, Emeishan County, Medicine School, 29 ° 21 ' N, 103 ° 17 ' E, alt. 1500 m, light trap, 6 males (BDN No. 1178, 1179, 0 5364, 0 5365, 0 5366, 05367), 17. v. 1986, Xinhua Wang; Sichuan Province, Emeishan County, Jinding, 29 ° 21 ' N, 103 ° 17 ' E, alt. 3.077 m, light trap, 1 male (BDN No. 05362), 18. v. 1985, Xinhua Wang; Tibet Autonomous Region, Chayu County, 29 ° N, 90 ° E, alt. 2500 m, light trap, 1 male (BDN No. 07845), 13. vi. 1988, Xinhua Wang. Diagnostic characters. The species can be separated from other East Asian members of the genus by having antenna with 9 or 10 flagellomeres; an AR of about 0.3 – 0.5; sternapodeme inverted U-shaped; and gonostylus slender and nearly straight.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FF90C61578D9FE40FB565A06.taxon	description	Addition to description. The Chinese specimens had the following measurements: Total length 0.87 – 1.15, 0.98 mm. Wing length 0.57 – 0.69, 0.62 mm. Posterior margin of tergite IX weakly bilobed. Inferior volsella large, nearly rectangular. Sternapodeme inverted U-shaped; attachment point with phallapodeme placed in caudal position of sternapodeme and ventrally directed; phallapodeme strongly curved with lateral apex bifid and embracing knob on lateral sternapodeme, not extending beyond tergite IX. Gonostylus slightly curved apically.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FF90C61578D9FE40FB565A06.taxon	discussion	Remarks. Tokunaga (1936) described the species based on specimens from Japan. When we examined the specimens from Sichuan province, we found that C. celtica listed in Wang (2000) was wrongly identified; the males from Sichuan all belong to C. yoshimurai.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
03BE87F2FF90C61578D9FE40FB565A06.taxon	distribution	Distribution. Japan, and Sichuan Province and Tibet Autonomous Region, Oriental China.	en	Fu, Yue, Saether, Ole A., Wang, Xinhua (2009): Corynoneura Winnertz from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae). Zootaxa 2287: 1-44, DOI: 10.5281/zenodo.191288
