identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03BF8784E378687AFF4FFB792238F9B5.text	03BF8784E378687AFF4FFB792238F9B5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudonortonia Giordani Soika 1936	<div><p>Genus Pseudonortonia Giordani Soika, 1936</p><p>Pseudonortonia Giordani Soika 1936: 268, genus.</p><p>Type species: Odynerus difformis de Saussure, 1853, by original designation.</p><p>Subancistroceroides Blüthgen 1938: 441, 460, subgenus of “ Subancistrocerus de Saussure ” sensu Blüthgen. Type species: Odynerus aegyptiacus de Saussure, 1863, by original designation.</p><p>Diagnosis. The genus Pseudonortonia can be distinguished from the other Oriental genera of Eumeninae by the following combination of characters: anterior face of pronotum with a deep oval pit in the middle, pretegular carina present, metanotum not tuberculate, submarginal carina of propodeum produced into a pointed lobe above valvula, T1 sub-petiolate and basally with one transverse carina, T2 not forming a basal acarinarium, second submarginal cell not petiolate.</p></div>	https://treatment.plazi.org/id/03BF8784E378687AFF4FFB792238F9B5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Selis, Marco;Nguyen, Lien Thi Phuong	Selis, Marco, Nguyen, Lien Thi Phuong (2024): The Oriental species of the genus Pseudonortonia Giordani Soika (Hymenoptera Vespidae: Eumeninae). Zootaxa 5477 (2): 171-194, DOI: 10.11646/zootaxa.5477.2.4, URL: http://dx.doi.org/10.11646/zootaxa.5477.2.4
03BF8784E3786878FF4FF91B2010FBF9.text	03BF8784E3786878FF4FF91B2010FBF9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudonortonia abbreviaticornis Giordani Soika 1941	<div><p>Pseudonortonia abbreviaticornis Giordani Soika, 1941</p><p>(Fig. 1)</p><p>Pseudonortonia abbreviaticornis Giordani Soika 1941: 246, fig. 21 (5), ♂ —“ China meridionale: East Kwantung, Tsin-Leng ” (lectotype MSNVE).</p><p>Pseudonortonia crassipunctata Gusenleitner 2001: 655, 657, ♂ —“ Laos: Louangnamtha Prov., Namtha—Muang Sing, 21°09’N 10119’E, 900-1200m ” (OLML). Syn. nov.</p><p>Diagnosis. The female of this species can be distinguished from the other Oriental species of Pseudonortonia by the following combination of characters: clypeus sparsely and finely punctate; occipital carina shallow and straight on gena; pronotal carina almost entirely absent dorsally; propodeal shelf as long as metanotum and deeply furrowed, about at same level as scutellum; T1 1.0–1.1× as long as apically wide and densely punctate, interspaces forming reticulation; T2 with punctures becoming abruptly coarser and denser preapically, interspaces mostly smooth and shiny, short smooth apical part not preceded by elongate pits; S2 evenly convex in basal half.</p><p>Material examined. LECTOTYPE OF PSEUDONORTONIA ABBREVIATICORNIS: ♂, labeled: “TsinLeong San / E. Kwantung. S. / China.VI-4-36 // L. Gressitt / Collector // Pseudonortonia / # 112. // HOLOTYPUS / Ps. / abbreviaticornis [red label] // ACTUALLY / LECTOTYPUS / Dal Pos, 2017 [red label]” (MSNVE). HOLOTYPE OF PSEUDONORTONIA CRASSIPUNCTATA: ♂, labeled: “HOLOTYPUS [red label] / LAOS, Louangnamtha pr. / 21°09’N 101°19’E. / Namtha --&gt; Muang Sing, 5–31. v. 1997, 900- / Vit Kubáň leg. - 1200 m // ♂ // Pseudonortonia / crassipunctata nov. spec. ♂ / J.Gusenleitner,det.2001 / Holotypus // Biologiezentrum/Linz / ex Coll. J. Halada / Eingang 1998/99” (OLML). CHINA: Guangdong, Guangzhou, South China University Campus, 4-24.VIII.2015, leg. P. Rosa, 2♂ 2♀ (MSVI); Guangdong, Shangshanbei, 23.4628N 113.7653E, 19.VIII.2016, leg. V. Soon, 1♂ (MSVI); same data of lectotype, 3.VI.1936, 1♂ (MSNVE, paralectotype). VIETNAM: Cao Bang, Tra Linh, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.895&amp;materialsCitation.latitude=20.743334" title="Search Plazi for locations around (long 104.895/lat 20.743334)">Thang Hen lake</a>, 22°45’53”N 106°17’46”E, 578 m, 4.VIII.2022, leg. Lien Thi Phuong Nguyen et al., 1♀ (IEBR); Hoa Binh, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.895&amp;materialsCitation.latitude=20.743334" title="Search Plazi for locations around (long 104.895/lat 20.743334)">Mai Chau</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.895&amp;materialsCitation.latitude=20.743334" title="Search Plazi for locations around (long 104.895/lat 20.743334)">Hang Kia—Pa Co Natural Reserve</a>, 20°44’36”N 104°53’42”E, 1340 m, 24.VI.2020, leg. Lien Thi Phuong Nguyen &amp; Ngat Thi Tran, 1♀ (IEBR); Son La, Muong La, Nam Pam, 660 m, 25.VII.2009, leg. Lien Thi Phuong Nguyen, Phong Huy Pham &amp; Junichi Kojima, 1♀ (IEBR).</p><p>Distribution. China: Guangdong; Laos; Vietnam * (Giordani Soika 1941; Gusenleitner 2001) (Fig. 10).</p><p>Notes. Gusenleitner (2001) described Pseudonortonia crassipunctata comparing it to P. bhamensis, but also mentioning that the very small F10–11 of the male were like those observed in P. abbreviaticornis . Comparison of the holotype of P. crassipunctata with specimens of P. abbreviaticornis from Guangdong, including the lectotype, revealed in fact that the two taxa correspond in all characters, making the former a junior synonym. Photos of the holotype of P. crassipunctata are available at ZOBODAT (2023).</p></div>	https://treatment.plazi.org/id/03BF8784E3786878FF4FF91B2010FBF9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Selis, Marco;Nguyen, Lien Thi Phuong	Selis, Marco, Nguyen, Lien Thi Phuong (2024): The Oriental species of the genus Pseudonortonia Giordani Soika (Hymenoptera Vespidae: Eumeninae). Zootaxa 5477 (2): 171-194, DOI: 10.11646/zootaxa.5477.2.4, URL: http://dx.doi.org/10.11646/zootaxa.5477.2.4
03BF8784E37A6878FF4FFBD72212F883.text	03BF8784E37A6878FF4FFBD72212F883.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudonortonia accliva (Gusenleitner 2007) Selis & Nguyen 2024	<div><p>Pseudonortonia accliva (Gusenleitner, 2007) comb. nov.</p><p>(Fig. 2)</p><p>Parancistrocerus acclivus Gusenleitner 2007: 97, 99, ♀ —“ Laos: Lao Pako env., 55 km NE of Vientiane ” (OLML).</p><p>Diagnosis. The female of this species can be distinguished from the other Oriental species of Pseudonortonia by the following combination of characters: clypeus sparsely and finely punctate; occipital carina shallow and straight on gena; pronotal carina entirely absent dorsally and forming spiniform teeth on humeri; propodeal shelf shorter than metanotum and shallowly depressed; T1 1.3× as long as apically wide, sparsely punctate with interspaces large and flattened, and with apical margin reflexed; T2 preapically depressed and densely punctate, with reflexed and crenate apical margin; S2 evenly convex.</p><p>Material examined. HOLOTYPE OF PARANCISTROCERUS ACCLIVUS: ♀, labeled: “HOLOTYPUS [red label] / LAOS, 4.v.2004 / LAO PAKO env. / 55 km NE of Vientiane / leg. F. Kantner // ♀ // Parancistrocerus / acclivus NOV. SPEC. ♀ / J.Gusenleitner,det.2006 / Holotypus // Biologiezentrum Linz, / Austria (LI) / ex Coll. J. &amp; M. Halada / Eingang 2006” (OLML). THAILAND: Chiang Mai, 100 km N of Chiang Mai, Chiang Dao Hill Resort, 540-640 m, 4-11.IX.2016, 1♀ (MSVI).</p><p>Distribution. Laos; Thailand * (Gusenleitner 2007) (Fig. 10).</p><p>Notes. Gusenleitner (2007) described this species in the genus Parancistrocerus Bequaert, but it clearly cannot be assigned to that genus due to the following characters: strongly elongate mesosoma, propodeum with a long horizontal shelf behind metanotum, subpetiolate T1 and absence of a basal acarinarium on T2. These characters support instead the placement of the taxon in the genus Pseudonortonia, where it is morphologically related to P. gujaratica (Nurse) .</p></div>	https://treatment.plazi.org/id/03BF8784E37A6878FF4FFBD72212F883	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Selis, Marco;Nguyen, Lien Thi Phuong	Selis, Marco, Nguyen, Lien Thi Phuong (2024): The Oriental species of the genus Pseudonortonia Giordani Soika (Hymenoptera Vespidae: Eumeninae). Zootaxa 5477 (2): 171-194, DOI: 10.11646/zootaxa.5477.2.4, URL: http://dx.doi.org/10.11646/zootaxa.5477.2.4
03BF8784E37D687EFF4FFA46270FF9B6.text	03BF8784E37D687EFF4FFA46270FF9B6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudonortonia bhamensis Giordani Soika 1941	<div><p>Pseudonortonia bhamensis Giordani Soika, 1941</p><p>(Fig. 3)</p><p>Pseudonortonia bhamensis Giordani Soika 1941: 245, fig. 21 (2, 4) [birmanensis [!] in explanation of fig.], ♀ —“ Birmania: Bhamò ” (lectotype MSNVE).</p><p>Diagnosis. The female of this species can be distinguished from the other Oriental species of Pseudonortonia by the following combination of characters: clypeus sparsely and finely punctate; occipital carina shallow and straight on gena; pronotal carina almost entirely absent dorsally; propodeal shelf as long as metanotum and deeply furrowed; T1 1.10–1.15× as long as apically wide and densely punctate, interspaces forming reticulation; T2 with punctures more or less regular from base to apex, interspaces mostly smooth and shiny, short smooth apical part not preceded by elongate pits; S2 strongly convex in basal fourth and then nearly straight.</p><p>Material examined. LECTOTYPE OF PSEUDONORTONIA BHAMENSIS: “Bhamò / Birmania / Fea VII 1886 // non oss. = / in Par. Mus / (1889) // 4 / 01 // HOLOTYPUS / Pseudonortonia / Bhamensis / Giord Ska [red label]” (MSNVE). INDIA: Assam, leg. S. Hartert, 1♀ (MSVI); Darjeeling, leg. Fruhstorfer, 1♀ (MSNVE).</p><p>Distribution. India: Sikkim, West Bengal; Myanmar (Giordani Soika 1941; Gusenleitner 2006a) (Fig. 10). Notes. The specimen labeled as holotype by Giordani Soika is in fact the lectotype.</p></div>	https://treatment.plazi.org/id/03BF8784E37D687EFF4FFA46270FF9B6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Selis, Marco;Nguyen, Lien Thi Phuong	Selis, Marco, Nguyen, Lien Thi Phuong (2024): The Oriental species of the genus Pseudonortonia Giordani Soika (Hymenoptera Vespidae: Eumeninae). Zootaxa 5477 (2): 171-194, DOI: 10.11646/zootaxa.5477.2.4, URL: http://dx.doi.org/10.11646/zootaxa.5477.2.4
03BF8784E37C6873FF4FF91A2247FDF1.text	03BF8784E37C6873FF4FF91A2247FDF1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudonortonia camarinensis Selis & Nguyen 2024	<div><p>Pseudonortonia camarinensis sp. nov.</p><p>(Fig. 4)</p><p>Diagnosis. The female of this species can be distinguished from the other Oriental species of Pseudonortonia by the following combination of characters: clypeus sparsely and finely punctate; occipital carina shallow and straight on gena; pronotal carina almost entirely absent dorsally; propodeal shelf longer than metanotum and shallowly furrowed, about at same level as scutellum; T1 1.15× as long as apically wide and densely punctate, interspaces forming reticulation; T2 with punctures becoming abruptly finer and sparser on disc, long smooth apical part not preceded by elongate pits and about 4× as long as apical lamella; S2 evenly convex.</p><p>Material examined. HOLOTYPE: ♀ labeled “P.I., CAMARINES / SUR, Mt. Iriga / 500m, 31.III.1962 // H.M. Torrevillas / Collector / BISHOP // Pseudonortonia / aff. bhamens / Giordani Soika det. 1981 // Pseudonortonia camarinensis / HOLOTYPUS ♀ / Det. Marco Selis [red label]” (MSNVE).</p><p>Description. Female holotype. Body length 8.6 mm; fore wing length 7.0 mm.</p><p>Head in frontal view about 1.2× as wide as high. Inner eye margins converging below, interocular distance about 1.5× wider at vertex than at clypeus. Clypeus in frontal view about 1.1× as wide as high; basal margin very weakly convex, narrowly separated from antennal toruli; apical margin shallowly emarginate, 0.3× as wide as clypeus, lateral teeth apically rounded and slightly pointing out; clypeus in lateral view weakly convex, shallowly depressed near apex. Gena 0.65× as wide as eye at level of bottom of ocular sinus; distance from posterior ocellus to occipital carina about 1.65× as distance from posterior ocellus to inner eye margin; cephalic foveae very small and close to each other, placed in a barely developed smooth convex area which continues posteriorly in an irregular longitudinal carina; occipital carina complete, higher and shallowly reflexed on gena and middle of vertex, much weaker on sides of vertex, weakly and evenly curved on gena. Mandible with five teeth. Scape 3.2× as long as apically wide; F1 1.35× as long as wide, F2 subquadrate, F3–9 wider than long. Mesosoma 1.55× as long as wide. Pronotal carina strong and sharp on lateral faces, strongly produced and angled on humeri, where it forms a lobe that is triangular in frontal view and spiniform in dorsal view; pronotal carina absent on the dorsal face, except for a very weak indication in the middle. Mesoscutum 1.1× as long as wide; notauli visible in the posterior third; mesoscutum in lateral view weakly convex, somewhat flattened posteriorly. Scutellum smoothly passing into mesoscutum, entirely flattened; anterior margin crenate and forming five pits, the median one larger; median longitudinal furrow present but very shallow. Metanotum almost horizontal, very weakly sloping posteriorly and shallowly furrowed medially. Tegula broadly rounded, with anterior third of outer margin nearly straight, posterior lobe more or less right-angled and reaching apex of parategula; parategula digitiform and curved. Epicnemial carina complete, well-developed but blunt, almost reaching pronotal margin. Posterior face of propodeum with sharp median carina running from base to apex; dorsal faces of propodeum meeting along mid-line and forming a propodeal shelf, which is longer than the metanotum and shallowly furrowed; dorsal carinae present but irregular due to coarse sculpture, ventral carinae present and sharp, lateral carinae absent; submarginal carina produced into a subtriangular but apically rounded lobe above valvula; propodeal valvula with deeply emarginate posterior margin, not completely fused to submarginal carina. T1 subpetiolate and narrower than T2, in dorsal view about 1.15× as long as apically wide, sides evenly diverging from carina to apex; T 1 in lateral view with a barely developed basal petiole, then abruptly swollen with evenly convex dorsal margin, barely depressed apically; transverse carina strong on the dorsal face, distinctly angled on the sides and then shortly continued on lateral face, where it is blunt. S1 subtriangular with concave apical margin, completely separated from T1 and bearing a pair of strong but blunt carinae running from basal petiole to posterior corners. T 2 in dorsal view with weakly and evenly convex sides, barely constricted apically; in lateral view more or less evenly convex, with shallowly depressed apical margin; apical margin with a series of coarse punctures basally delimiting a long smooth area, which is 4× as long as apical hyaline lamella. S 2 in lateral view barely longer than T2 and evenly convex; apical margin similar to T2, but hyaline lamella longer. Prestigma subquadrate, about 0.20× as long as pterostigma; second recurrent vein nearly interstitial.</p><p>Clypeus with very sparse and fine punctures, interspaces equal to several puncture diameters and strongly shiny, with very sparse micropunctures. Frons with dense deep punctures, interspaces narrower than puncture diameter and forming a shiny and shallowly convex reticulation. Vertex and gena similar to frons but with sparser punctures, some interspaces reaching one puncture diameter in length; lower narrow part of gena with irregular and sparse punctures; occipital carina crenate on middle of vertex and less evidently on gena, with a short longitudinal carina in the middle. Scape with sparse micropunctures and few scattered fine macropunctures. Pronotum similar to frons but punctures larger; anterior face sculpted like dorsal faces but with larger smooth interspaces; lateral faces with traces of longitudinal striation. Mesoscutum with larger and coarser punctures, interspaces forming irregular longitudinal ridges in the area between the notauli. Scutellum similar to mesoscutum, longitudinal ridges more irregular. Metanotum with few large and coarse punctures, sides mostly smooth and shiny. Tegula strongly shiny, completely smooth on disc, with dense micropunctures on margins. Mesepisternum with punctures similar to mesoscutum, partly arranged in oblique series; epicnemium crenate along border of foramen, with deep punctures on mesosternum. Dorsal plate of metaepisternum with strong longitudinal ridges, median ridge particularly developed; ventral plate largely shiny and micropunctate, with crenate anterior margin and small deep punctures on anteroventral corner. Dorsal faces of propodeum coarsely punctate, punctures large and deep, interspaces strongly raised and forming reticulation; lateral faces with shallower and smaller flat-bottomed punctures, mostly arranged in longitudinal series; posterior face shagreened and silky-shiny, with large shallow punctures along dorsolateral margins and oblique striae covering most of the remaining surface. Post-carinal area of T1 with large and coarse punctures, slightly sparser basally, interspaces very narrow and forming a strong reticulation; pre-carinal area mostly smooth with a curved series of deep punctures in the upper half; apical swollen margin completely smooth. T2 on basal third with deep punctures becoming denser laterally, punctures becoming much finer and sparser posteriorly on disc; T2 preapically bearing a narrow area where punctures become rapidly coarser; area between coarse punctures and apical lamella completely smooth. T3–6 shagreened and with minute punctures becoming progressively sparser and less visible. S1 with coarse and deep punctures in the area between the longitudinal carinae, transversely ridged on sides. S2 with sparse fine punctures, similar to those on disc of T2; apical smooth area preceded by an irregular series of sparse larger punctures. S3–6 similar to respective tergites. Pale golden setae on whole body, longer on frons and sides of mesosoma, shorter and very sparse on tergites; short and dense pubescence on lower part of face including clypeus, gena, sides of mesosoma and posterior face of propodeum; metasoma with dense and darker pubescence, scattered setae denser on sternites than on tergites.</p><p>Black; following parts yellow: spots on basal and apical corners of clypeus fused to form an irregularly Ushaped marking, mandible except inner margin, elongate interantennal spot, lower face of scape, small spot behind upper lobe of eye, narrowly separated triangular spots on middle of pronotum, suffused spots on tegula, parategula, narrowly separated spots on metanotum, narrow apical band on T1 shortly curved on sides, whole apical area of T2 and S2, apical third of fore femur, outer face of all tibiae (except apex of hind tibia), short line on fore basitarsus; following parts yellowish-ferruginous: lower face of flagellum, tegula, part of fore tibia, most of fore tarsi. Wings fusco-hyaline, with suffused spot at apex of marginal cell.</p><p>Male. Unknown.</p><p>Distribution. Philippines: Luzon (Fig. 10).</p><p>Etymology. The species name refers to the province of Camarines Sur, where this species was collected. It is an adjective.</p></div>	https://treatment.plazi.org/id/03BF8784E37C6873FF4FF91A2247FDF1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Selis, Marco;Nguyen, Lien Thi Phuong	Selis, Marco, Nguyen, Lien Thi Phuong (2024): The Oriental species of the genus Pseudonortonia Giordani Soika (Hymenoptera Vespidae: Eumeninae). Zootaxa 5477 (2): 171-194, DOI: 10.11646/zootaxa.5477.2.4, URL: http://dx.doi.org/10.11646/zootaxa.5477.2.4
03BF8784E3716873FF4FFDD72606F91C.text	03BF8784E3716873FF4FFDD72606F91C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudonortonia flavolimbata (Giordani Soika 1986) Giordani Soika 1986	<div><p>Pseudonortonia flavolimbata Giordani Soika, 1986 stat. nov.</p><p>(Fig. 5)</p><p>Pseudonortonia bhamensis flavolimbata Giordani Soika 1986: 77, ♀ —“ India: “ Benkgalis ” [Sumatra: Bengkalis]” (MNHN).</p><p>Diagnosis. The female of this species can be distinguished from the other Oriental species of Pseudonortonia by the following combination of characters: clypeus sparsely and finely punctate; occipital carina shallow and straight on gena; pronotal carina almost entirely absent dorsally; propodeal shelf shorter than metanotum and shallowly depressed, placed below level of scutellum; T1 1.10–1.15× as long as apically wide and densely punctate, interspaces forming reticulation; T2 with punctures becoming progressively and weakly coarser in front of apical smooth part, interspaces mostly micropunctate and matte, short smooth apical part not preceded by elongate pits; S2 evenly convex in basal half or basal third.</p><p>Material examined. INDONESIA: Bengkalis, 1885, leg. Maindron, 1♀ (MSNVE, paratype) ; E. Sumatra, Riouw Res., Inderagiri, Pangkalan, Kasai, IV.1939, leg. P. Buwalda, 1♀ (MSVI) . MALAYSIA: British N. Borneo, Ranau, 11.X.1958, leg. L.W. Quate, 1♀ (MSVI) ; Borneo, Sandakan, leg. Baker, 1♀ (MSNVE) .</p><p>Distribution. Indonesia: Sumatra; Malaysia: Sabah * (Giordani Soika 1986) (Fig. 10).</p><p>Notes. Giordani Soika (1986) described this taxon as a subspecies of Pseudonortonia bhamensis, differentiated by color characters only. Comparison of the two taxa, including the lectotype of P. bhamensis and a paratype of P. bhamensis flavolimbata, revealed a series of differences that allow to consider P. flavolimbata a distinct species: pronotal humeri sharp spiniform, propodeal shelf less developed and sloping, shorter smooth apical part of T2, S2 more evenly convex and not flattened posteriorly, finer punctures on head and mesosoma, legs more extensively ferruginous, markings pale yellow and much reduced.</p><p>Giordani Soika (1986) reports an unspecified “Benkgalis” in India as the type locality, but the examination of a paratype shows that the correct locality is Bengkalis, a small island just off the coast of Sumatra. A second specimen from Pangkalan Kasai, a village in the central part of Sumatra, further confirms the Sumatran origin of Pseudonortonia flavolimbata .</p><p>Two specimens from Sabah (Fig. 5F) present a darker color pattern, with yellow markings of the meso- and metasoma reduced (apical band of T2 reduced to two barely visible lateral lines) and the tegulae brown-red instead of ferruginous, and the punctures of the pronotum and mesoscutum subtly denser with less shiny interspaces. These differences, however, are very subtle and apparently gradual between the four available specimens, thus leading us to tentatively consider the specimens from Sabah as a mere dark variety of P. flavolimbata . Examination of more material and the discovery of the males of both forms will help to clarify the situation. One of the two specimens from Sabah was labeled by Giordani Soika as paratype of “ Pseudonortonia borneana ”, an unpublished name.</p></div>	https://treatment.plazi.org/id/03BF8784E3716873FF4FFDD72606F91C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Selis, Marco;Nguyen, Lien Thi Phuong	Selis, Marco, Nguyen, Lien Thi Phuong (2024): The Oriental species of the genus Pseudonortonia Giordani Soika (Hymenoptera Vespidae: Eumeninae). Zootaxa 5477 (2): 171-194, DOI: 10.11646/zootaxa.5477.2.4, URL: http://dx.doi.org/10.11646/zootaxa.5477.2.4
03BF8784E3706870FF4FF9F627A6FCD1.text	03BF8784E3706870FF4FF9F627A6FCD1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudonortonia gujaratica (Nurse 1902)	<div><p>Pseudonortonia gujaratica (Nurse, 1902)</p><p>(Fig. 6)</p><p>Montezumia gujaratica Nurse 1902: 90, pl. fig. 16, ♀ — “Deesa” (NHMUK, type no. 18.211).</p><p>Labus campanulatus Wickwar 1908: 118, 121, fig. 10–13, ♀ —“ Ceylon ” (NHMUK, type no. 18.109).</p><p>Diagnosis. The female of this species can be distinguished from the other Oriental species of Pseudonortonia by the following combination of characters: clypeus sparsely and finely punctate; occipital carina shallow and straight on gena; pronotal carina shortly developed in the middle and forming obtuse teeth on humeri; propodeal shelf as long as metanotum and deeply furrowed; T1 1.4–1.55× as long as apically wide, sparsely punctate with interspaces large and flattened, and with apical margin not reflexed; T2 with barely reflexed and regular apical margin; S2 evenly convex.</p><p>Material examined. HOLOTYPE OF MONTEZUMIA GUJARATICA: ♀ labeled “Type [circular label with red border] // Deesa / 10.98 // ♀ // Type // Montezumia / gujaratica / (Nurse) // B.M. TYPE / HYM. 18.211 // [QR code] / NHMUK015610159” (NHMUK). HOLOTYPE OF LABUS CAMPANULATUS: ♀ labeled “Type [circular label with red border] // CEYLON, // Ceylon. / O.S.Wickwar. / 1912–189. // Labus / campanulatus / Type. Wickwar // Is a / Nortonia / CD. // B.M. TYPE / HYM. / 18.109 // [QR code] / NHMUK015610160” (NHMUK). INDIA: Deesa, VIII.1897, 1♀ (MSNVE); Mysore, Bandipur, 1100 m, 3.IV.1970, leg. R.T. Simon Thomas, 1♀ (MSNVE).</p><p>Distribution. India: Gujarat, Karnataka *; Sri Lanka (Nurse 1902; Wickwar 1908) (Fig. 10).</p><p>Notes. The specimen from Deesa in MSNVE (Figs 6 A-B) was labelled by Giordani Soika as paratype (Fig. 6C). Nurse (1902) does not report the exact number of specimens examined to describe this species, but the labels of the specimen examined indicate that it came from the Nurse collection and the collecting data are in line with what is reported in the description; on the other hand, the type in NHMUK bears original type labels, while the specimen in MSNVE does not. It is therefore unclear whether the specimen in MSNVE belongs to the original type series, but, in any case, it would be a paralectotype and not a paratype.</p><p>The types of Montezumia gujaratica and Labus campanulatus show some slight differences, shared by the two specimens found in MSNVE: the type of Labus campanulatus and the specimen from Bandipur have slightly larger punctures on clypeus (Fig. 5G), generally more elongate appearance, T1 1.55× as long as apically wide (1.4× in the type of M. gujaratica and the other specimen from Deesa), apical margin of T2 more reflexed (Fig. 5H). These differences are however very subtle, and we prefer to tentatively consider them as intraspecific variability probably linked to the different size of the specimens (8.5 mm for the specimen from Deesa, 10.0 mm for the one from Bandipur), accepting the synonymy of the two taxa as already proposed by Giordani Soika (1941). The examination of more material and the discovery of the males will provide useful data to clarify the situation.</p></div>	https://treatment.plazi.org/id/03BF8784E3706870FF4FF9F627A6FCD1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Selis, Marco;Nguyen, Lien Thi Phuong	Selis, Marco, Nguyen, Lien Thi Phuong (2024): The Oriental species of the genus Pseudonortonia Giordani Soika (Hymenoptera Vespidae: Eumeninae). Zootaxa 5477 (2): 171-194, DOI: 10.11646/zootaxa.5477.2.4, URL: http://dx.doi.org/10.11646/zootaxa.5477.2.4
03BF8784E3726877FF4FFC3F2793FD89.text	03BF8784E3726877FF4FFC3F2793FD89.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudonortonia henrica (Cameron 1908)	<div><p>Pseudonortonia henrica (Cameron, 1908)</p><p>(Fig. 7)</p><p>Odynerus henricus Cameron 1908: 306, ♂ —“ Bombay ” (NHMUK, type no. 18.296).</p><p>Diagnosis. Distinguished from the other Oriental species of Pseudonortonia by the short T1 and the basally concave S2.</p><p>Material examined. HOLOTYPE OF ODYNERUS HENRICUS: ♂ labeled “Type [circular label with red border] // Ancistrocerus / henricus Cam / Type Bombay // P. Cameron Coll. / 1914-110. // B.M. TYPE / HYM. / 18.296 // [QR code] NHMUK015610166” (NHMUK).</p><p>Distribution. India: Maharastra (Cameron 1908) (Fig. 10).</p><p>Notes. Pseudonortonia henrica is distinct among the Oriental species of Pseudonortonia in possessing propodeum and T1 more similar to some African and Arabian species of the genus. The placement in Pseudonortonia, originally proposed by Giordani Soika (1941), is here provisionally maintained. The club-shaped antenna and the morphology of the second sternite place this species close to other genera with carinate T1 such as Subancistrocerus de Saussure and Stenancistrocerus de Saussure, and are also observed in an undescribed Eumenine wasp from Pakistan with uncertain generic affinities (M. Selis, unpublished data). The actual generic attribution of Pseudonortonia henrica will have to be discussed in the light of a phylogenetic study of Pseudonortonia and related genera.</p></div>	https://treatment.plazi.org/id/03BF8784E3726877FF4FFC3F2793FD89	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Selis, Marco;Nguyen, Lien Thi Phuong	Selis, Marco, Nguyen, Lien Thi Phuong (2024): The Oriental species of the genus Pseudonortonia Giordani Soika (Hymenoptera Vespidae: Eumeninae). Zootaxa 5477 (2): 171-194, DOI: 10.11646/zootaxa.5477.2.4, URL: http://dx.doi.org/10.11646/zootaxa.5477.2.4
03BF8784E3756875FF4FFD072653FA0D.text	03BF8784E3756875FF4FFD072653FA0D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudonortonia incola Selis & Nguyen 2024	<div><p>Pseudonortonia incola sp. nov.</p><p>(Fig. 8)</p><p>Diagnosis. The female of this species can be distinguished from the other Oriental species of Pseudonortonia by the following combination of characters: clypeus sparsely and finely punctate; occipital carina shallow and straight on gena; pronotal carina almost entirely absent dorsally; propodeal shelf as long as metanotum and shallowly furrowed, slightly below level of scutellum; T1 1.05× as long as apically wide and densely punctate, interspaces forming reticulation; T2 with a series of large and elongate pits preceding the short smooth apical part, apical lamella with long and pointed basal costulation; S2 evenly convex basally.</p><p>Material examined. HOLOTYPE: ♀ labeled “Kolambugan / Mindanao // HOLOTYPUS / bhamensis / incola [red label] // Pseudonortonia incola / HOLOTYPUS ♀ / Det. Marco Selis [red label]” (MSNVE).</p><p>Description. Female holotype. Body length 8.3 mm; fore wing length 6.9 mm.</p><p>Head in frontal view about 1.2× as wide as high. Inner eye margin converging below, interocular distance about 1.5× wider at vertex than at clypeus. Clypeus in frontal view 1.15× as wide as high; basal margin very weakly convex, narrowly separated from antennal toruli; apical margin shallowly emarginate, 0.25× as wide as clypeus, lateral teeth apically rounded and slightly pointing out; clypeus in lateral view weakly and evenly convex, shallowly depressed near apex. Gena 0.55× as wide as eye at level of bottom of ocular sinus; distance from posterior ocellus to occipital carina about 1.65× as distance from posterior ocellus to inner eye margin; cephalic foveae very small and close to each other, placed in a barely developed smooth area just behind ocellar triangle; occipital carina complete, higher on gena and middle of vertex, almost disappearing on sides of vertex, weakly and evenly curved on gena. Mandible with five teeth. Scape 3.2× as long as apically wide; F1 1.45× as long as wide, F2–3 subquadrate, F4–9 wider than long. Mesosoma 1.5× as long as wide. Pronotal carina strong and sharp on lateral faces of pronotum, strongly produced and angled on humeri, where it forms a lobe that is triangular in frontal view and spiniform in dorsal view; pronotal carina barely indicated on dorsal face, slightly sharper for a short median portion. Mesoscutum 1.1× as long as wide; notauli deep but present only in the posterior fourth; mesoscutum in lateral view weakly convex, somewhat flattened posteriorly. Scutellum smoothly passing into mesoscutum, barely convex from side to side; anterior margin crenate and forming five pits, the median and the most lateral ones larger; median longitudinal furrow present but very shallow. Metanotum weakly sloping, with convex sides and shallow median furrow. Tegula broadly rounded, with anterior third of outer margin nearly straight, posterior lobe more or less right-angled and reaching apex of parategula; parategula digitiform and angled. Epicnemial carina complete and strong but not sharp, almost reaching pronotal margin. Posterior face of propodeum with barely developed and irregular median carina; dorsal faces of propodeum meeting along mid-line and forming a propodeal shelf, which is as long as the metanotum and shallowly furrowed; dorsal carinae present but weak and irregular, ventral carinae sharp, lateral carinae barely indicated by a rounded fold of the surface; submarginal carina produced into a subtriangular but apically rounded lobe above valvula; propodeal valvula with emarginate posterior margin, not completely fused to submarginal carina. T1 subpetiolate and narrower than T2, in dorsal view about 1.05× as long as apically wide, sides evenly diverging from carina to apex; T 1 in lateral view with a barely developed basal petiole, then abruptly swollen with evenly convex dorsal margin, shallowly depressed apically; transverse carina strong on dorsal face and pointed in the middle, angled on sides and rapidly disappearing on lateral face. S1 subtriangular with concave apical margin, completely separated from T1 and bearing a pair of large but shallow carinae running from basal petiole to posterior corners. T 2 in dorsal view with sides convex and diverging in basal third, then almost straight and parallel, barely constricted apically; T 2 in lateral view convex in basal third, then flattened with shallow apical depression; apical margin with a series of large and dense elongate pits, delimiting a smooth area which is about as long as the pits and slightly longer than the apical hyaline lamella; apical lamella with elongate and sharp costulation, covering almost the whole length of the lamella. S 2 in lateral view as long as T2 and evenly convex, less bulging basally than T2; apical margin similar to T2 but pits much smaller and sparse and apical lamella shorter. Prestigma subquadrate and very short; second recurrent vein almost interstitial.</p><p>Clypeus with very sparse and barely impressed punctures, interspaces equal to several puncture diameters and strongly shiny, with very sparse micropunctures. Frons with dense deep punctures, interspaces narrower than puncture diameter and forming a shiny and shallowly convex reticulation. Vertex and gena similar to frons but with sparser punctures, most interspaces equal to puncture diameter in width; lower narrow part of gena with irregular and denser punctures; occipital carina crenate on middle of vertex and less evidently on gena, with a short longitudinal carina in the middle. Scape with sparse micropunctures and few scattered fine macropunctures. Pronotum similar to frons but punctures larger and interspaces slightly wider; anterior face with deep rounded punctures in the dorsal two thirds, dense with interspaces variable in width. Mesoscutum similar to pronotum but punctures denser, interspaces shiny and forming distinct longitudinal ridges in the area between the notauli. Scutellum similar to mesoscutum, interspaces generally as wide as puncture diameter. Metanotum with very few large punctures in the middle of posterior margin, sides mostly smooth and shiny. Tegula strongly shiny and micropunctate, more densely on the anterior half. Mesepisternum with punctures similar to mesoscutum, partly arranged in oblique series; epicnemium crenate along border of foramen, with deep punctures on mesosternum. Dorsal plate of metaepisternum with strong longitudinal ridges, median ridge particularly developed; ventral plate largely shiny and micropunctate, with crenate anterior margin and small deep punctures on anteroventral corner. Dorsal faces of propodeum coarsely punctate, punctures large and deep, interspaces strongly raised and forming reticulation; lateral faces with shallower and smaller flat-bottomed punctures, mostly arranged in longitudinal series; posterior face shagreened and silky-shiny, with large shallow punctures along dorsolateral margins and irregular short striae on sides of median carina. Postcarinal area of T1 with deep and coarse punctures, larger and coarser along carina, interspaces very narrow and forming a strong reticulation; pre-carinal area mostly smooth with deep punctures arranged in several curved series; apical swollen margin mostly smooth. T2 with deep punctures, slightly denser and more oblique basally, interspaces generally as wide or wider than puncture diameter; T2 preapically bearing a narrow area where punctures become rapidly denser with sharp ridge-like interspaces, followed by the preapical series of elongate pits; area between pits and apical lamella completely smooth and shiny. T3–6 shagreened and with deep oblique punctures becoming progressively sparser and weaker. S1 irregularly rugose in the area between the longitudinal carinae, transversely ridged on sides. S2 with sparse fine punctures, similar to those on disc of T2; apical smooth area preceded by an irregular series of larger punctures. S3–6 similar to respective tergites. Pale golden setae on whole body, longer on frons and sides of mesosoma, shorter and very sparse on tergites; short and dense pubescence on lower part of face including clypeus, gena, sides of mesosoma and posterior face of propodeum; metasoma with dense and darker pubescence, scattered setae denser on sternites than on tergites.</p><p>Black; following parts yellow (turned to red post-mortem, probably for KCN): clypeus except longitudinal band originating from basal margin and reaching the middle, mandible except inner margin, elongate interantennal spot, lower face of scape, spot behind upper lobe of eye, wide band along pronotal carina not reaching humeri and fused to line along posterior pronotal margin, tegula, parategula, metanotum, narrow apical band on T1 shortly curved on sides, spots on lateral angles of basal carina of T1, wide bisinuate band on T2 and S2, apical third of fore femur, apex of mid femur, outer face of all tibiae (except apex of hind tibia); flagellum orange below. Wings hyaline with weakly darkened anterior margin and suffused spot at apex of marginal cell.</p><p>Male. Unknown.</p><p>Distribution. Philippines: Mindanao (Fig. 10).</p><p>Etymology. The holotype of this taxon was labeled as holotype of “ bhamensis incola ” by Giordani Soika. Although treated as a species and not a subspecies, the name incola is retained. It is a noun in apposition.</p></div>	https://treatment.plazi.org/id/03BF8784E3756875FF4FFD072653FA0D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Selis, Marco;Nguyen, Lien Thi Phuong	Selis, Marco, Nguyen, Lien Thi Phuong (2024): The Oriental species of the genus Pseudonortonia Giordani Soika (Hymenoptera Vespidae: Eumeninae). Zootaxa 5477 (2): 171-194, DOI: 10.11646/zootaxa.5477.2.4, URL: http://dx.doi.org/10.11646/zootaxa.5477.2.4
03BF8784E377686BFF4FF9832644F886.text	03BF8784E377686BFF4FF9832644F886.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudonortonia lamellata Selis & Nguyen 2024	<div><p>Pseudonortonia lamellata sp. nov.</p><p>(Fig. 9)</p><p>Diagnosis. The female of this species can be distinguished from the other Oriental species of Pseudonortonia by the following combination of characters: clypeus with dense and strong punctures, interspaces between punctures smaller than puncture diameter, punctures in central part larger than those in other parts; occipital carina strongly developed and forming a distinct angle in lower half; pronotal carina fully developed, strongly raised; propodeal shelf shorter than metanotum; T1 1.1× as long as apically wide and densely punctate, interspaces forming reticulation; T2 and S2 strongly bulging preapically and bearing a long reflexed hyaline lamella, S2 evenly convex and distinctly longer than T2.</p><p>Material examined. HOLOTYPE: ♀ labeled “VIETNAM; Lao Cai, Van Ban, / Nam Xe, Hoang Lien-Van Ban NR, / B 22°2’47.9”, E 103°58’13”; 900 m / 23.iv.2021; NTP Lien, TT Ngat // Pseudonortonia lamellata / HOLOTYPE ♀ / Det. Nguyen T. P. Lien [red label]” (IEBR) . PARATYPES. VIETNAM: same data as holotype, 4♀ (IEBR); Cao Bang, Tra Linh, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.29611&amp;materialsCitation.latitude=22.759167" title="Search Plazi for locations around (long 106.29611/lat 22.759167)">Thang Hen Lake</a>, 22°45’33”N 106°17’46”E, 578 m, 4.VIII.2022, leg. Lien Thi Phuong Nguyen et al., 2♀ (IEBR) ; Lang Son, Huu Lung, Huu Lien, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.41011&amp;materialsCitation.latitude=21.5635" title="Search Plazi for locations around (long 106.41011/lat 21.5635)">Lan Nghe</a>, Hu Lien NR, 21°33’48.6”N 106°24’36.4”E, 289 m, leg. Lien Thi Phuong Nguyen et al., 1♀ (IEBR) .</p><p>Description. Female. Body length 7.3–8.4 mm (holotype 8.0 mm); fore wing length 7.0–8.0 mm (holotype 7.6 mm).</p><p>Head in frontal view about 1.1× as wide as high. Inner eye margins strongly converging below, interocular distance about 1.5× wider at vertex than at clypeus. Clypeus in frontal view wider than long, about 1.1× as wide as high; basal margin almost straight and not touching antennal toruli; apical margin very shallowly emarginate, 0.35× as wide as clypeus, lateral teeth blunt and shortly carinate; clypeus in lateral view evenly convex, shallowly depressed near apex. Gena as wide as eye at bottom of ocular sinus; distance from posterior ocellus to occipital carina about 2.2× as distance from posterior ocellus to inner eye margin; vertex with very small cephalic foveae; occipital carina complete, strongly reflexed on upper three fourths of gena, angled and forming a blunt tooth in lower third. Mandible with five teeth. Scape 3.5× as long as apically wide; F1 1.3× as long as wide, F2 subquadrate, F3–9 wider than long. Mesosoma 1.55× as long as wide. Pronotal carina complete and sharp, produced on humeri but not spiniform. Mesoscutum 1.2× as long as wide; notauli impressed and complete, forming a shallow pit on anterior margin of mesoscutum; mesoscutum in lateral view weakly convex, somewhat flattened posteriorly. Scutellum smoothly passing into mesoscutum, weakly convex posteriorly; anterior margin crenate and forming 7-8 pits, the median one larger; median longitudinal furrow complete but shallow. Metanotum oblique and evenly sloping to propodeum. Epicnemial carina strong and complete, almost reaching pronotal margin. Posterior face of propodeum with strong median carina running from base to apex; dorsal faces of propodeum meeting along mid-line and forming a propodeal shelf shorter than metanotum, depressed in the middle and forming median pit; all carinae present but not sharp, crenate due to the coarse sculpture; submarginal carina produced into a sharp and pointed lobe above valvula; propodeal valvula with emarginate posterior margin, not completely fused to submarginal carina. T1 subpetiolate and narrower than T2, in dorsal view 1.1× as long as apically wide, sides evenly diverging from carina to apex; T 1 in lateral view with a thin short basal petiole, then abruptly swollen with evenly convex dorsal margin, shallowly depressed apically; transverse carina strong on the dorsal face, becoming blunter on sides, where it curves and almost reaches the posterior corners of the tergite. S1 subtriangular with concave apical margin, completely separated from T1. T 2 in dorsal view with weakly and evenly convex sides, weakly constricted apically; in lateral view weakly convex basally and then almost flattened, with preapical bulge preceding apical lamella; apical margin forming a long and thick subhyaline lamella, shallowly reflexed. S 2 in lateral view longer and more strongly and evenly convex than T2, with preapical bulge less evident; apical margin lamellate like T2, but more strongly costulate basally. T3 with shallowly carinate apical margin.</p><p>Clypeus with dense and strong punctures, larger in the center and bearing short setae; interspaces shorter than puncture diameter and sparsely micropunctate. Frons with dense and coarse flat-bottomed punctures, interspaces very narrow and raised to form reticulation. Vertex and gena similar to frons but punctures deeper and slightly sparser; lower part of gena with small and shallow punctures; occipital carina crenate along vertex. Scape with sparse and small punctures. Pronotum similar to frons but punctures larger; anterior face with very dense and coarse punctures in the upper half, smooth ventrolaterally. Mesoscutum with coarse punctures, similar to those on pronotum; interspaces sparsely micropunctate, narrower and forming reticulation anteriorly, becoming much wider posteriorly. Scutellum similar to pronotum, interspaces forming longitudinal carinae along posterior margin. Metanotum with smaller but denser punctures, interspaces reduced to longitudinal carinae along anterior margin. Tegula with sparse minute punctures. Mesepisternum with flat-bottomed punctures, larger and denser posterodorsally; epicnemium smooth. Dorsal plate of metaepisternum rugose, with short weak striation; ventral plate mostly smooth, with sparse shallow punctures. Dorsal faces of propodeum coarsely punctate, interspaces strongly raised and forming a sharp reticulation; lateral faces similar to dorsal faces but punctures much shallower and smaller; posterior face mostly smooth and shiny, with coarse punctures along dorsolateral margins and some fine striae along median carina. T1 with large and coarse punctures, slightly sparser and weaker basally; interspaces very narrow and forming a strong reticulation. T2 densely and deeply punctate, interspaces equal to slightly larger than puncture diameter and densely micropunctate; preapical swollen margin mostly smooth. T3–6 with minute punctures. S1 with coarse and flat-bottomed punctures, interspaces raised to form sharp reticulation. S2 similar to T2 but punctures sparser. Silvery setae on whole body, longer on head and mesosoma, shorter on metasoma, particularly dense on T1; face and posterior face of propodeum with dense pubescence; T2 and S2 with very short pubescence in addition to short oblique setae.</p><p>Black; following parts yellow to orange-yellow: basal spot on clypeus, small spot above interantennal space, ventral face of scape, small spot behind dorsal lobe of eye, transverse lines along anterior margin of pronotum, apical margin of T1, preapical bulge of T2, widely interrupted preapical line on S2, apical spot on fore and mid femora, outer face of fore and mid tibia, basal spot on hind tibia; tegula with suffused brown markings; apical lamella of T2 and S2 hyaline with yellowish-brown tinge. Wings hyaline with slightly darkened anterior margin.</p><p>Male. Unknown.</p><p>Distribution. Vietnam (Fig. 10).</p><p>Etymology. The name refers to the strongly lamellate apical margin of T2 and S2. It is an adjective.</p></div>	https://treatment.plazi.org/id/03BF8784E377686BFF4FF9832644F886	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Selis, Marco;Nguyen, Lien Thi Phuong	Selis, Marco, Nguyen, Lien Thi Phuong (2024): The Oriental species of the genus Pseudonortonia Giordani Soika (Hymenoptera Vespidae: Eumeninae). Zootaxa 5477 (2): 171-194, DOI: 10.11646/zootaxa.5477.2.4, URL: http://dx.doi.org/10.11646/zootaxa.5477.2.4
03BF8784E3686869FF4FFC7826FFFC63.text	03BF8784E3686869FF4FFC7826FFFC63.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudonortonia Giordani Soika 1936	<div><p>Key to the Oriental species of Pseudonortonia</p><p>[The key is mostly based on the females, as the males are described for P. abbreviaticornis and P. henrica only. In the case of male specimens, the characters relating to the pronotal carina may not be applicable, since in P. abbreviaticornis the female has a strongly reduced carina and the male has it complete and well developed and similar differences could be observed in the other species too.]</p><p>1. T1 short and wide, postcarinal area shorter than wide at apex (Fig. 7A); transverse carina of T1 strongly bulging and followed in lateral view by a depression. S2 shallowly depressed basally, almost smoothly passing into S1 (Fig. 7B).................................................................................................. P. henrica (Cameron)</p><p>- T1 elongate, postcarinal area longer than wide at apex (Figs 1H, 2E, 3F, 4F, 5E, 6F, 6H, 8F, 9F); transverse carina of T1 weakly bulging and not followed in lateral view by a depression. S2 basally convex, clearly separated from S1 (Figs 1B, 2B, 3B, 4B, 5B, 5F, 6B, 8B, 9B)................................................................................... 2</p><p>2. T1 more elongate, 1.3–1.55× as long as apically wide, and with smaller and sparser punctures, interspaces wide and flattened (Figs 2E, 6F, 6H). Apical margin of T2 at least weakly reflexed, without a hyaline lamella following the pigmented part (Figs 2E, 6F, 6H).......................................................................................... 3</p><p>- T1 less elongate, 1.0–1.15× as long as apically wide, and with larger and denser punctures, interspaces narrow and mostly reduced to ridges (Figs 1H, 3F, 4F, 5E, 8F, 9F). Apical area of T2 variably depressed and placed below level of the rest of the tergite, preapical margin variably swollen and followed by a variably developed but always evident hyaline lamella (Figs 1H, 3F, 4F, 5E, 8F, 9F).................................................................................... 4</p><p>3. Clypeus distinctly emarginate at apex, apical teeth subtriangular (Fig. 2C). Anterior margin of pronotum entirely rounded, pronotal carina entirely absent on the dorsal part; pronotal humeri sharply spiniform (Fig. 2D). T1 shorter, 1.3× as long as apically wide, and more strongly convex, apical margin distinctly reflexed (Fig. 2E). T2 with a depressed and densely punctate preapical area, then strongly reflexed; apical margin irregular, weakly but evidently crenate (Fig. 2E). Pale markings bright yellow and more extensive on head and tegula; legs mostly bright ferruginous (Figs 2A–B)....... P. accliva (Gusenleitner)</p><p>- Clypeus subtruncate at apex, apical teeth blunt and slightly pointing out (Figs 6D, 6G). Anterior margin of pronotum not entirely rounded, with traces of pronotal carina in the middle of the dorsal part; pronotal humeri blunt or sharp, but not spiniform (Fig. 6E). T1 longer, 1.4–1.55× as long as apically wide, and less convex, apical margin nearly flattened (Figs 6F, 6H). T2 not particularly depressed or densely punctate in the preapical area, and weakly reflexed; apical margin regular and not crenate (Figs 6F, 6H). Pale markings ivory and less extensive on head and absent on tegula; legs mostly blackish brown (Figs 6A–B)................................................................................... P. gujaratica (Nurse)</p><p>4. Clypeus densely and strongly punctate (Fig. 9D). Occipital carina strongly developed and reflexed, strongly angled in lower third (Fig. 9B). Pronotal carina complete (Fig. 9E). T2 and S2 with strongly bulging preapical margin, followed by a long and reflexed yellowish-hyaline lamella (Fig. 9F); S2 distinctly longer than T2 (Fig. 9B).................. P. lamellata sp. nov.</p><p>- Clypeus very sparsely and finely punctate (Figs 1D–E, 3D, 4D, 5C, 8D). Occipital carina weakly reflexed, nearly straight on whole gena (Figs 1B, 3B, 4B, 5B, 5F, 8B). Pronotal carina largely absent dorsally, reduced to lateral teeth and sometimes to very short median portion (Figs 1G, 3E, 4E, 5D, 8E). T2 and S2 at most shallowly bulging preapically, with a variably long smooth pigmented part and a very short apical lamella (Figs 1H, 3F, 4F, 5E, 8F); S2 as long as T2 or very slightly longer (Figs 1B, 3B, 4B, 5B, 5F, 8B)................................................................................ 5</p><p>5. Propodeal shelf in the middle longer than metanotum (Fig. 4E). T2 with coarse and dense punctures in the basal third, becoming abruptly much finer and sparser posteriorly, except for a preapical series of coarse and dense punctures; smooth apical part of T2 long, about 4× as long as the apical lamella (Fig. 4F)................................... P. camarinensis sp. nov.</p><p>- Propodeal shelf in the middle as long as metanotum or shorter (Figs 1G, 3E, 5D, 8E). T2 with coarse punctures on whole surface, becoming very slightly finer on disc; smooth apical part of T2 short, 1.5–2.5× as long as the apical lamella (Figs 1H, 3F, 5E, 8F).......................................................................................... 6</p><p>6. Anterior vertical face of T1 with many dense and deep punctures. Smooth apical part of T2 preceded by a serie of elongate pits, distinctly larger than the punctures preceding them; margin produced in long and pointed costulations above apical lamella (Fig. 8F)............................................................................... P. incola sp. nov.</p><p>- Anterior vertical face of T1 with few sparse punctures, mostly impunctate. Smooth apical part of T2 not preceded by elongate pits; margin produced in short rounded digitations above apical lamella (Figs 1H, 3F, 5E)............................ 7</p><p>7. S 2 in lateral view straight for most of the surface, then strongly and abruptly curved in the basal fourth (Fig. 3B)....................................................................................... P. bhamensis Giordani Soika</p><p>- S 2 in lateral view gently and evenly curved in the basal half or basal third (Figs 1B, 5B, 5F).......................... 8</p><p>8. Propodeal shelf strongly developed and about at same level as scutellum, as long as metanotum and with a deep and narrow median longitudinal furrow (Fig. 1G). Punctures on T2 becoming abruptly coarser and denser in front of apical smooth part; interspaces on T2 with sparse micropunctures and pubescence, appearing markedly shiny (Fig. 1H)............................................................................................. P. abbreviaticornis Giordani Soika</p><p>- Propodeal shelf little developed and placed well below level of scutellum, shorter than metanotum and widely and shallowly depressed (Fig. 5D). Punctures on T2 becoming progressively and weakly coarser and denser in front of apical smooth part; interspaces on T2 with dense micropunctures and pubescence, with a weak silky shine, almost matte (Fig. 5E)....................................................................................... P. flavolimbata Giordani Soika</p></div>	https://treatment.plazi.org/id/03BF8784E3686869FF4FFC7826FFFC63	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Selis, Marco;Nguyen, Lien Thi Phuong	Selis, Marco, Nguyen, Lien Thi Phuong (2024): The Oriental species of the genus Pseudonortonia Giordani Soika (Hymenoptera Vespidae: Eumeninae). Zootaxa 5477 (2): 171-194, DOI: 10.11646/zootaxa.5477.2.4, URL: http://dx.doi.org/10.11646/zootaxa.5477.2.4
03BF8784E36B6869FF4FFB9021F2F8C1.text	03BF8784E36B6869FF4FFB9021F2F8C1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Symmorphus (Symmorphus) jucundus (Gusenleitner 2008) Selis & Nguyen 2024	<div><p>Symmorphus (Symmorphus) jucundus (Gusenleitner, 2008) comb. nov.</p><p>(Fig. 11)</p><p>Pseudonortonia jucundus Gusenleitner 2008: 1495, ♀, ♂ —“ Thailand NW, Chiang Dao, 8 km W Chiang Dao ” (holotype female OLML).</p><p>Material examined. HOLOTYPE OF PSEUDONORTONIA JUCUNDUS: ♀, labeled: “THAILAND WN / Chiang Dao, 8kmW / Chiang Dao, Waldbach / 10.-12.3.07, lg. Risch // Pseudonortonia / jucundus nov. spec. ♀ / J.Gusenleitner,det.2008 / Holotypus // HOLOTYPUS [red label] // Biologiezentrum Linz / Austria (LI) / ex. Coll. J. Gusenleitner / Eingang 2023” (OLML). THAILAND: same data as holotype, 1♂ (OLML, paratype of Pseudonortonia jucunda).</p><p>Distribution. Thailand (Gusenleitner 2008).</p><p>Notes. Gusenleitner (2008) compared this taxon to Pseudonortonia bhamensis and differentiated it by the sparser sculpture of the mesosoma, the obliquely sloping propodeum and the coarse preapical punctures of T2. Examination of the type series however revealed that Pseudonortonia jucunda (not jucundus, since Pseudonortonia is feminine in gender) is in fact a Symmorphus Wesmael. The characters that support this new placement are those already listed by Cumming (1989) as diagnostic of Symmorphus: T1 longitudinally furrowed behind transverse carina, female with cephalic foveae widely separated (Fig. 11E), and male with simple antennal apex (Fig. 11G).</p><p>The strongly elongate T1 observed in this species is an uncommon feature in the genus Symmorphus, being currently known in two Central American species only: S. centralis Carpenter &amp; Grandinete and S. chiriquiensis (Gusenleitner) (Grandinete et al. 2016). This character immediately allows the recognition of S. jucundus from all other known Old World species of the genus.</p></div>	https://treatment.plazi.org/id/03BF8784E36B6869FF4FFB9021F2F8C1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Selis, Marco;Nguyen, Lien Thi Phuong	Selis, Marco, Nguyen, Lien Thi Phuong (2024): The Oriental species of the genus Pseudonortonia Giordani Soika (Hymenoptera Vespidae: Eumeninae). Zootaxa 5477 (2): 171-194, DOI: 10.11646/zootaxa.5477.2.4, URL: http://dx.doi.org/10.11646/zootaxa.5477.2.4
03BF8784E36A686FFF4FFA7B2293F980.text	03BF8784E36A686FFF4FFA7B2293F980.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tachyancistrocerus komarowi (Morawitz 1885)	<div><p>Tachyancistrocerus komarowi (Morawitz, 1885)</p><p>(Fig. 12)</p><p>Ancistrocerus komarowi Morawitz 1885: 175, ♀ —[Turkmenistan] “ In territorio transcaspico. Asschabad ” (ZIN).</p><p>Pseudonortonia bushirensis Giordani Soika 1943: 1, fig. 1 (1-4), ♂ —“ Persia: Bushire ” (MSNVE).</p><p>Pseudonortonia pretiosissima Giordani Soika 1943: 4, fig. 2 (1–3), ♀ —“ N. Assam: Dschaipur ” (MSNVE). Syn. nov.</p><p>Tachyancistrocerus komarowi derufata Gusenleitner in Bytinksi-Salz &amp; Gusenleitner 1971: 239, 271, 294, ♀, ♂ —“ Israel … En Gedi ” (holotype female ZMB).</p><p>Material examined. HOLOTYPE OF PSEUDONORTONIA PRETIOSISSIMA: ♀, labeled: “Dschaipur / 30.X.-2.XI.29 / Dr. Enslin // ♀ // HOLOTYPUS / Pseudonortonia / pretiosissima / n. sp. / A.Giordani Soika [red label]” (MSNVE). IRAN: Khuzestan, Haft Tapeh, 300 km N of Abadan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=34.9&amp;materialsCitation.latitude=30.966667" title="Search Plazi for locations around (long 34.9/lat 30.966667)">Dez river</a>, 29.VI-1.VII.1965, leg. A. Giordani Soika, 1♀ (MSVI). ISRAEL: 2 km SW of Jerucham, 30°58’N 34°54’E, 470 m, 18.IV.2018, leg. A. Kudrna jr., 1♂ (MSVI). PAKISTAN: Karachi, leg. W.D. Cumming, 1♀ (MSVI).</p><p>Distribution. Afghanistan, India (Rajasthan), Iran, Israel, Kazakhstan, Oman, Pakistan, Turkey, Turkmenistan, Uzbekistan (Blüthgen 1955, 1961; Bytinksi-Salz &amp; Gusenleitner 1971; Fateryga et al. 2023; Giordani Soika 1943, 1970; Gusenleitner 1973, 1988, 2006b; Kostylev 1940; Morawitz 1885, 1895; Schmid-Egger et al. 2021).</p><p>Notes. Giordani Soika (1943) described Pseudonortonia bushirensis and Pseudonortonia pretiosissima in the same paper, comparing them with the African species of the genus, in particular with P. aegyptiaca (de Saussure) . Later, Blüthgen (1955) synonymized P. bushirensis under Tachyancistrocerus komarowi and stated that the type of P. pretiosissima was probably in Zürich, in the von Schulthess collection. The holotype of P. pretiosissima is found in MSNVE in the Giordani Soika collection, as reported in the original description, and its examination revealed that this species is also actually a synonym of Tachyancistrocerus komarowi . The locality reported on the label is “Dschaipur”, which Giordani Soika (1943) reports to be Jaipur in northern Assam, but which Blüthgen (1955) reports to be in Rajputana based on information obtained from the collector of the specimen, Eduard Enslin. Given the known distribution of the species, the collection locality is undoubtedly that reported by Blüthgen and Enslin, today in the state of Rajasthan.</p></div>	https://treatment.plazi.org/id/03BF8784E36A686FFF4FFA7B2293F980	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Selis, Marco;Nguyen, Lien Thi Phuong	Selis, Marco, Nguyen, Lien Thi Phuong (2024): The Oriental species of the genus Pseudonortonia Giordani Soika (Hymenoptera Vespidae: Eumeninae). Zootaxa 5477 (2): 171-194, DOI: 10.11646/zootaxa.5477.2.4, URL: http://dx.doi.org/10.11646/zootaxa.5477.2.4
