taxonID	type	description	language	source
03BC8436FFD1C82DFF5AFC8FE4BBFAB8.taxon	distribution	Distribution. China, Guangxi Zhuang Autonomous Region, Lo Ch’eng. Previously known only from the holotype male, which is lost (Wu 1938 a, Zwick 1984).	en	Mo, Raorao, Wang, Guoquan, Li, Weihai, Murányi, Dávid (2020): Review of the Oriental genus Neoperlops Banks, 1939 (Plecoptera: Perlidae). Zootaxa 4763 (3): 405-418, DOI: 10.11646/zootaxa.4763.3.5
03BC8436FFD1C82DFF5AFC8FE4BBFAB8.taxon	discussion	Remarks. Zwick (1984) noted that only on the basis of the original description, this species is indistinguishable from N. gressitti. Despite that this is the only species hitherto known from Guangxi, below we report our gressitti group specimen as N. gressitti. Wu (1938, figs. 308 – 309) originally illustrated an undivided sensilla basiconica patch on tergum 9, probably a species specific character of Neoperlops. Our specimen has a divided sensilla patch, similar to the redescription of N. gressitti (Zwick 1984: fig. 2 d). However, topotypes need to be examined from Luocheng City (Lo Ch’eng) to clarify this the status of this species.	en	Mo, Raorao, Wang, Guoquan, Li, Weihai, Murányi, Dávid (2020): Review of the Oriental genus Neoperlops Banks, 1939 (Plecoptera: Perlidae). Zootaxa 4763 (3): 405-418, DOI: 10.11646/zootaxa.4763.3.5
03BC8436FFD1C82DFF5AFDBAE47AFCE0.taxon	discussion	This morphological species group is characterized by the elongated hemitergal apex of the male, and distinctly punctuated chorion of the eggs. Three species are recognized:	en	Mo, Raorao, Wang, Guoquan, Li, Weihai, Murányi, Dávid (2020): Review of the Oriental genus Neoperlops Banks, 1939 (Plecoptera: Perlidae). Zootaxa 4763 (3): 405-418, DOI: 10.11646/zootaxa.4763.3.5
03BC8436FFD1C828FF5AFA67E270FEA1.taxon	description	(Figs. 1 – 2)	en	Mo, Raorao, Wang, Guoquan, Li, Weihai, Murányi, Dávid (2020): Review of the Oriental genus Neoperlops Banks, 1939 (Plecoptera: Perlidae). Zootaxa 4763 (3): 405-418, DOI: 10.11646/zootaxa.4763.3.5
03BC8436FFD1C828FF5AFA67E270FEA1.taxon	materials_examined	Material examined. 1 male (HIST): CHINA, Guangxi Zhuang Autonomous Region, Nanning City, Wuming County, Liangjiang Town, Neichao village, 23 ° 29.664 ' N, 108 ° 21.622 ' E, 230 m, 2014. V. 10, G. Q. Wang.	en	Mo, Raorao, Wang, Guoquan, Li, Weihai, Murányi, Dávid (2020): Review of the Oriental genus Neoperlops Banks, 1939 (Plecoptera: Perlidae). Zootaxa 4763 (3): 405-418, DOI: 10.11646/zootaxa.4763.3.5
03BC8436FFD1C828FF5AFA67E270FEA1.taxon	distribution	Distribution. China, described from Guangdong and Jiangxi Provinces, on the basis of two males and two females (Banks 1939), recently reported from Fujian Province (Zhang et al. 2019) and herein from Guangxi Zhuang Autonomous Region.	en	Mo, Raorao, Wang, Guoquan, Li, Weihai, Murányi, Dávid (2020): Review of the Oriental genus Neoperlops Banks, 1939 (Plecoptera: Perlidae). Zootaxa 4763 (3): 405-418, DOI: 10.11646/zootaxa.4763.3.5
03BC8436FFD1C828FF5AFA67E270FEA1.taxon	discussion	Remarks. Zwick (1984) presented a redescription, including the everted aedeagus of N. gressitti based on type material. The head ocellar pattern and the main aedeagal armatures of the present Guangxi male generally agree with the types, in addition to the identical external terminalia. Slight differences in the aedeagus of the Guangxi male include a larger trapezoidal dorsomedial process in lateral view; in dorsal aspect the process is bilobed (Figs. 2 a, c – f). The basal half of aedeagus in the type seems twisted (Zwick 1984: fig. 2 g). These differences may be due to the preserved condition of the aedeagus. Herein, we present color illustrations of external features and everted aedeagal structures to assist further identification of the species (Figs. 1 – 2). The specimens recently listed from Fujian (Zhang et al. 2019) may refer to N. huanghuye, described from the same area (Wuyishan). Further molecular studies are needed to confirm identity of the three, apparently closely related species of the gressitti group.	en	Mo, Raorao, Wang, Guoquan, Li, Weihai, Murányi, Dávid (2020): Review of the Oriental genus Neoperlops Banks, 1939 (Plecoptera: Perlidae). Zootaxa 4763 (3): 405-418, DOI: 10.11646/zootaxa.4763.3.5
03BC8436FFD4C828FF5AFE53E6B6FCB6.taxon	distribution	Distribution. China, Fujian Province, Mt. Wuyishan. Known from five males and six females, all collected at a single locality (Chen 2020).	en	Mo, Raorao, Wang, Guoquan, Li, Weihai, Murányi, Dávid (2020): Review of the Oriental genus Neoperlops Banks, 1939 (Plecoptera: Perlidae). Zootaxa 4763 (3): 405-418, DOI: 10.11646/zootaxa.4763.3.5
03BC8436FFD4C828FF5AFE53E6B6FCB6.taxon	discussion	Remarks. The species was recently described and placed in Phanoperla. However, it is a species of Neoperlops: besides obvious similarity of the male and female terminalia, aedeagus and egg to those of N. gressitti, fig. 2 C in Chen (2020) clearly shows the elongated tentorial callosities, characteristic for Neoperlops and Tetropina (Zwick 1984, Sivec et al. 1988). The species can be distinguished from N. gressitti based on the slightly different head pattern, and denser sensilla basiconica of tergum 9. As mentioned above, identity and distinction should be confirmed by molecular studies.	en	Mo, Raorao, Wang, Guoquan, Li, Weihai, Murányi, Dávid (2020): Review of the Oriental genus Neoperlops Banks, 1939 (Plecoptera: Perlidae). Zootaxa 4763 (3): 405-418, DOI: 10.11646/zootaxa.4763.3.5
03BC8436FFD4C828FF5AFBB8E575F921.taxon	distribution	Distribution. China, described from Guangdong Province, Yim Na San, based on four females (Banks 1939). Stark & Sivec (2008) reported several specimens from Vietnam, including males, females and larvae (Lào Cai, Ngh ệ An and Vĭnh Phúc provinces).	en	Mo, Raorao, Wang, Guoquan, Li, Weihai, Murányi, Dávid (2020): Review of the Oriental genus Neoperlops Banks, 1939 (Plecoptera: Perlidae). Zootaxa 4763 (3): 405-418, DOI: 10.11646/zootaxa.4763.3.5
03BC8436FFD4C828FF5AFBB8E575F921.taxon	discussion	Remarks. The female types of N. obscuripennis are distinctive by having elongated oval eggs, contrary to nearly globular eggs of the congeners (Zwick 1984). The putative male and larva from Vietnam were associated with this species based on the female and eggs similar to the redescription of the types (Stark & Sivec 2008). However, the illustrated Vietnamese eggs are nearly globular (Stark & Sivec 2008, fig. 7), so conspecificity can be questioned. In addition, Stark & Sivec (2008) mention one male specimen that lacked spines on the apex of male hemiterga, a character that is characteristic for N. vietnamellus and the proposed new species. Molecular studies of topotypes are needed to clear identity of the species.	en	Mo, Raorao, Wang, Guoquan, Li, Weihai, Murányi, Dávid (2020): Review of the Oriental genus Neoperlops Banks, 1939 (Plecoptera: Perlidae). Zootaxa 4763 (3): 405-418, DOI: 10.11646/zootaxa.4763.3.5
03BC8436FFD4C828FF5AFC60E534FBCE.taxon	discussion	This morphological species group is characterized by the short hemitergal apex of the male, and smooth chorion of the eggs. Contains three species:	en	Mo, Raorao, Wang, Guoquan, Li, Weihai, Murányi, Dávid (2020): Review of the Oriental genus Neoperlops Banks, 1939 (Plecoptera: Perlidae). Zootaxa 4763 (3): 405-418, DOI: 10.11646/zootaxa.4763.3.5
03BC8436FFD4C820FF5AF8D3E5ADFD81.taxon	description	(Figs. 3 – 8)	en	Mo, Raorao, Wang, Guoquan, Li, Weihai, Murányi, Dávid (2020): Review of the Oriental genus Neoperlops Banks, 1939 (Plecoptera: Perlidae). Zootaxa 4763 (3): 405-418, DOI: 10.11646/zootaxa.4763.3.5
03BC8436FFD4C820FF5AF8D3E5ADFD81.taxon	description	Adult habitus. General color brown. Head brown with a large dark brown trapezoidal marking, covering ocellar area and extending forward to pale M-line; tentorial callosities slender with curved tips, included in the dark brown patch (Figs. 3 a, 5 a). Biocellate, distance between ocelli about twice as wide as diameter of the ocellus, tentorial cal- losities set close to ocelli. Palpi brown; antennae dark brown, scape paler. Pronotum brown and elongate oval, with a pair of pale brown areas at lateral surfaces; lateral and posterior margins dark brown; rugosities large but obscure except two darker and distinct rugosities present on mediolateral surfaces; slightly narrower than head with eyes (Figs. 3 a, 5 a). Meso- and metathorax ventrally pale, dorsally with brown to dark brown pattern; metathorax with anteromedial brush, being much sparser on the female. Wings membrane brownish with darker veins. Legs pale brown but distal one fourth of femora to tarsi dark brown (Fig. 4 a). Abdomen pale, sterna 5 – 7 with medial brush; cerci brown, relatively long. Male (Figs. 3 – 4). Forewing length 16.0 – 17.5 mm; hindwing length 15.0 – 16.0 mm. Terga 7 – 9 bear prominent mesal patches of sensilla basiconica, among them, the patch of sensilla basiconica of tergum 9 is interrupted by a narrow bare band (Figs. 3 b – d). Hemitergal anterior processes short and flat, spatula shaped in lateral aspect, lacking strong setae; inner basal surface of hemiterga inflated, cushion-like and covered with minute sensilla basiconica (Figs. 3 b – d). Aedeagus (Figs. 4 b – e) mostly membranous but weak sclerotization occurs in basal half; slightly constricted medially and strongly curved ventrally, shape like a “ hair dryer; ” apical half distinctly enlarged. Basal portion with dorsal field of small scales, a large patch of fine triangular spines covers most of the enlarged apical half but is interrupted by the triangular membranous lobe on the dorsum; apex membranous, bare. Female. Forewing length 27.0 – 28.5 mm; hindwing length 22.0 – 23.0 mm. abdominal sterna generally pale, but sterna 5 – 7 medially darker around poorly developed brush. Sternum 8 with small, brown and triangular subgenital plate, forming a distinct bump in lateral view (Figs. 5 c – d). Sternum 9 with mediolateral pair of brown spots, sternum 10 unmodified, paraproct pale and blunt. Vagina simple, membranous. Egg (Fig. 5 b). Length 0.32 – 0.40 mm, width 0.28 – 0.36 mm (N = 8). Outline oval, nearly globular, chorion dark brown. Opercular line is at one fourth to opercular end. Chorion smooth, micropyles indistinct. Collar short but distinctly flanged, collar rim sinuous. Anchor consists of tentacle-like fibers. Larva. Body length of female larvae 21.0 – 22.5 mm, male exuviae 14.0 – 16.5 mm, female exuviae 21.0 – 23.0 mm. General color brown, with dark brown pattern on head and thorax, and reddish terminal terga and cerci (Fig. 8 a); ventral surface generally pale (Figs. 6 e – f). Head mostly dark brown but with pale occiput, tentorial callosities, a transverse patch anterior to M-line, and smaller anteromedial patch on frons; occipital area not elongated, eyes normal sized. Biocellate, distance between ocelli about twice as wide as diameter of one ocellus, tentorial callosities longitudinally elongated and set close to ocelli. Occipital ridge distinct, row of occipital setae complete, setae rather long around eyes; remainder of the head covered by fine, black clothing hairs (Fig. 6 a). Antenna brown, palpi pale brown (Fig. 6 c). Postmentum undivided, glossa and paraglossa usual for the family (Fig. 6 e). Lacinia bidentate, apical tooth longer than subapical tooth, marginal fringe consists of 6 – 8 strong setae and several thin hairs; galea as long as apical tooth of lacinia, bears only apical setae (Fig. 7 f). Mandible wide, having two molar and two or three incisors, molar brush dense and relatively long (Fig. 7 e). Pronotum with rounded corners, less than twice wider than long, marginal row of setae complete, moderately long; the disc is brown to pale brown but submarginal edges dark brown, rugosities numerous but not distinct (Fig. 6 b). Meso- and metanotum similar in coloration, all thoracic segments covered by fine, black clothing hairs. Mesosternum with long furcasternal pit, furcasternal arms also long (Fig. 6 e). Thoracic gills moderately long and with dense branches (Fig. 6 e). Proventricular teeth denser on the basal section of folds (Fig. 7 b). Legs long, dorsal row of swimming hairs dense, long and distinct on femora, similar on tibiae; surface setation consists of black clothing hairs and regularly placed, short brown setae (Figs. 6 c, 7 a, 8 b). Abdominal terga lack distinct pattern, but gradually darkened towards terminalia, with a reddish hint (Fig. 8 b). All terga bear dense apical row of setae with equal length, about of one fourth of segment length; most of the surface is covered with fine, black clothing hairs, and short setae similar to those on femora; medial silky hairs incomplete, scarce (Figs. 6 d, 7 a, 7 c). Clothing hairs pale on sterna, stout brown setae present on lateral areas; the apical row medially widely interrupted on sterna 2 – 5, complete on sterna 6 – 10 in case of males while on female sternum 8 it is narrowly interrupted by the shallow genital notch (Fig. 6 f). Paraproct simple, bears a long and dense anal gill (Figs. 6 f, 7 c – d). Cercus as long as the body, reddish, lacks swimming hairs; apical row consists of short but erect, brown setae, intercalary hairs long and pale, fit tight (Figs. 7 c – d).	en	Mo, Raorao, Wang, Guoquan, Li, Weihai, Murányi, Dávid (2020): Review of the Oriental genus Neoperlops Banks, 1939 (Plecoptera: Perlidae). Zootaxa 4763 (3): 405-418, DOI: 10.11646/zootaxa.4763.3.5
03BC8436FFD4C820FF5AF8D3E5ADFD81.taxon	materials_examined	Type material. Holotype male (HIST): CHINA, Guangxi Zhuang Autonomous Region, Shangsi County, Shiwanda Mts, Wanggui Village, Pinglong River at the village, 21 ° 52.411 ' N, 107 ° 49.996 ' E, 260 m, 2015. iii. 28, J. Kontschán, W. H. Li, D. Murányi, G. Q. Wang. Paratypes: same data as holotype: 3 males 1 female (HIST), 2 males, 1 female with its exuviae, 3 male and 3 female exuviae (HNHM); Guangxi Zhuang Autonomous Region, Shangsi County, Shiwandashan National Forest Park, Minan River, 21 ° 51.869 ' N, 107 ° 50.712 ' E, 300 m, 2015. iii. 28, J. Kontschán, W. H. Li, D. Murányi, G. Q. Wang: 1 female larva (HIST), 1 female larva (HNHM); Guangxi Zhuang Autonomous Region, Shangsi County, Shiwandashan National Forest Park, Pearl River below tourist route bridge, 21 ° 54.122 ' N, 107 ° 54.292 ' E, 315 m, 2015. iii. 29, J. Kontschán, J. N. Li, S. Li, W. H. Li, D. Murányi, G. Q. Wang: 1 female larva, 1 male exuviae (HNHM).	en	Mo, Raorao, Wang, Guoquan, Li, Weihai, Murányi, Dávid (2020): Review of the Oriental genus Neoperlops Banks, 1939 (Plecoptera: Perlidae). Zootaxa 4763 (3): 405-418, DOI: 10.11646/zootaxa.4763.3.5
03BC8436FFD4C820FF5AF8D3E5ADFD81.taxon	etymology	Etymology. The specific name is composed of the Latin words “ tri ” (three) and “ angulus ” (angle) and refers to the shape of the triangular lobe in dorsum of aedeagus.	en	Mo, Raorao, Wang, Guoquan, Li, Weihai, Murányi, Dávid (2020): Review of the Oriental genus Neoperlops Banks, 1939 (Plecoptera: Perlidae). Zootaxa 4763 (3): 405-418, DOI: 10.11646/zootaxa.4763.3.5
03BC8436FFD4C820FF5AF8D3E5ADFD81.taxon	distribution	Distribution and ecology. China, Guangxi Zhuang Autonomous Region, Shiwandashan Mountains. Larvae and exuviae were found at three nearby small stony rivers. Adults were emerging only at the Pinglong River (Fig. 8 a). No adults were attracted to light. The species was found at the Pinglong River with several other Perlidae, a few Nemouridae and Leuctridae species; exuviae of a Chloroperlidae species was found.	en	Mo, Raorao, Wang, Guoquan, Li, Weihai, Murányi, Dávid (2020): Review of the Oriental genus Neoperlops Banks, 1939 (Plecoptera: Perlidae). Zootaxa 4763 (3): 405-418, DOI: 10.11646/zootaxa.4763.3.5
03BC8436FFD4C820FF5AF8D3E5ADFD81.taxon	discussion	Remarks. The female of the new species is similar to the types of N. obscuripennis in coloration but differs with nearly globular eggs differing from elongated oval eggs of the types. As mentioned above, Vietnamese specimens reported by Stark & Sivec (2008) as N. obscuripennis, clearly belong to two species. The illustrated Vietnamese adults differ from the new species by small dark pattern on head, undivided sensilla basiconica patch on male tergum 9, presence of spines on the apex of male hemiterga, much more scales on the basal half of the aedeagus, and nearly erect shape of the aedeagus (Stark & Sivec 2008, figs. 2 – 4). Neoperlops vietnamellus, the third species of the obscuripennis group, differs from the new species by partly divided sensilla basiconica patch on tergum 9, fingerlike apex of the male hemiterga, and more dense surface structures of the aedeagus (Cao & Bae 2008, figs. 4 – 7). The larva can be distinguished from N. vietnamellus and the Vietnamese specimens illustrated by Stark & Sivec (2008) based on different head pattern, in addition, from N. vietnamellus by complete marginal row of setae on pronotum, and scarce medial silky hairs on the abdomen.	en	Mo, Raorao, Wang, Guoquan, Li, Weihai, Murányi, Dávid (2020): Review of the Oriental genus Neoperlops Banks, 1939 (Plecoptera: Perlidae). Zootaxa 4763 (3): 405-418, DOI: 10.11646/zootaxa.4763.3.5
03BC8436FFDCC820FF5AFD73E594FC06.taxon	distribution	Distribution. Vietnam, described from Hà Tĭnh, Lào Cai, Phú Th ọ and Vĭnh Phúc provinces on the basis of several specimens including males, females and larvae (Cao & Bae 2008).	en	Mo, Raorao, Wang, Guoquan, Li, Weihai, Murányi, Dávid (2020): Review of the Oriental genus Neoperlops Banks, 1939 (Plecoptera: Perlidae). Zootaxa 4763 (3): 405-418, DOI: 10.11646/zootaxa.4763.3.5
03BC8436FFDCC820FF5AFD73E594FC06.taxon	discussion	Remarks. Affinities of the species were discussed above. Some specimens reported as N. obscuripennis by Stark & Sivec (2008) may refer to this species.	en	Mo, Raorao, Wang, Guoquan, Li, Weihai, Murányi, Dávid (2020): Review of the Oriental genus Neoperlops Banks, 1939 (Plecoptera: Perlidae). Zootaxa 4763 (3): 405-418, DOI: 10.11646/zootaxa.4763.3.5
