taxonID	type	description	language	source
03BD2655FFA3D216FF7CFD24A75FE5E7.taxon	materials_examined	Type locality. Morocco, Middle Atlas Mts., Tazzeka National Park, Bab-Bou-Idir env., 34 ° 04 ′ N 4 ° 07 ′ W. Type material. Holotype: Ƌ (micropterous), ‘ MOROCCO, 28. v. - 3. vi. / 1999. Tazzeka N. P., Bab- / Bou-Idir env., 34 ° 04 ′ N, / 4 ° 07 ′ W, P. Průdek leg. [p, white label] // EX COLLECTIO / Z. JINDRA, PRAGUE [p, white label] // Ƌ [p, white label] // HOLOTYPUS / HEBRUS / ATLAS sp. nov. / det. P. KMENT, Z. JINDRA / & G. M. BERCHI 2016 ’ (ZJPC, to be deposited in NMPC). Holotype card-mounted, complete. Paratypes: 1 Ƌ (micropterous), ‘ MOROCCO, 28. v. – 3. vi. / 1999. Tazzeka N. P., Bab- / Bou-Idir env., 34 ° 04 ′ N, / 4 ° 07 ′ W, P. Průdek leg. [p, white label] // EX COLLECTIO / Z. JINDRA, PRAGUE [p, white label] // Ƌ [p, white label] // (ZJPC); 1 ♀ (micropterous), ‘ MOROCCO, 28. v. – 3. vi. / 1999. Tazzeka N. P., Bab- / Bou-Idir env., 34 ° 04 ′ N, / 4 ° 07 ′ W, P. Průdek leg. [p, white label] // EX COLLECTIO / Z. JINDRA, PRAGUE [p, white label] // COLLECTIO / NATIONAL MUSEUM / Praha, Czech Republic [p, white label] // ♀ [p, white label] ’ (NMPC); 1 Ƌ (micropterous), ‘ MAROCCO, 10 October 1993 / Ifrane env., cca 1600 m / 60 km S of Fes / P. Bílek lgt. [p, white label] // EX COLLECTIO / Z. JINDRA, PRAGUE [p, white label] // COLLECTIO / NATIONAL MUSEUM / Praha, Czech Republic [p, white label] // Ƌ [p, white label] ’ (NMPC); 1 ♀ (micropterous), ‘ MAROCCO, 10 October 1993 / Ifrane env., cca 1600 m / 60 km S of Fes / P. Bílek lgt. [p, white label] // EX COLLECTIO / Z. JINDRA, PRAGUE [p, white label] // ♀ [p, white label] ’ (ZJPC). All the paratypes bear the following labels: ‘ PARATYPUS / HEBRUS / ATLAS sp. nov. / det. P. KMENT, Z. JINDRA / & G. M. BERCHI 2016 ’.	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFA3D216FF7CFD24A75FE5E7.taxon	description	Description. Known only in micropterous form. Colouration (Figs. 1 – 3). Head and thorax reddish (with metanotal elevation somewhat darker) in males (Figs. 1 – 2), blackish brown (with head and pronotum posteriorly with reddish tinge) in females (Fig. 3). Remaining body parts with same colouration in both sexes. Eyes dark reddish. Bucculae yellowish. Legs and antennal segments I and II yellowish with apices somewhat infuscated; antennal segments III – IVb brownish. Corium blackish brown, concolorous with disc of pronotum; clavus with small white spot basally, rest of corium and entire membrane dark brown. Abdomen black, only apically somewhat reddish. Vestiture. Pubescence on head, pronotum, sides of thorax, mesoscutellum and metanotal elevation (‘ scutellum’) very short, in optical microscope indistinct (see Figs. 4 – 9). Mandibular plates (Fig. 7), corium, clavus posteriorly (Fig. 10), abdomen dorsally and ventrally (Figs. 4, 10), apical thirds of femora (Fig. 11), tibiae (Fig. 11), tarsi (Fig. 12), and antennal segments (Figs. 13 – 14) with longer pale hairs appearing silver or gold. Structure. Body elongate (Figs. 1 – 4), 2.63 – 2.70 times longer than wide across humeral angles, but body widest across gibbose metapleura (see Measurements). Vertex with median sulcus shallow, hardly visible in optical microscope (Figs. 4, 6). Bucculae low, ventrally straight, posteriorly conically produced (Fig. 7); head laterally with prominent oblique ridge running from base of antenniferous tubercle to postero-ventral margin of eye (Fig. 7). Labium not observed on card-mounted specimens. Antennal segment I stoutest, cylindrical, slightly curved (Fig. 13); segment II clavate (Fig. 13); segments III, IVa and IVb terete (Figs. 1 3 – 14). Antennal segments from longest to shortest (Figs. 13 – 14): I> III> IVb> II> IVa; segment I 1.3 3 – 1.50 × longer than II as well as maximum width of compound eye; segment IV divided by membranous junction into IVa and IVb (Figs. 15 – 16). Pronotum (Figs. 1 – 4, 8) transversely hexagonal, anterior collar not delimited posteriorly, middle transverse constriction well developed, especially laterally. Metanotal elevation shortly trapezoid, about 2.9 × wider than long, posteriorly widely obtusely arcuate, without any incision (Figs. 8 – 9); disc of metanotal elevation with low median carina accompanied by two shallow submedian depressions (Figs. 8 – 9), not turned upwards posteriorly (Fig. 5). Metapleuron strongly gibbose, therefore metathorax slightly wider than pronotum across humeral angles (Figs. 1, 3, 8). Hemelytra (Fig. 10) reaching posterior margin of mediotergite II (Figs. 1 – 4). Metafemur of male strongly curved (Figs. 1 – 2, 11), only indistinctly curved in female (Fig. 3), metatibiae of both sexes straight, without long hairs (Figs. 1 – 3, 11). Mediotergites I and II with two thin, submedian, posteriad slightly diverging longitudinal keels (Figs. 3 – 4). Male genitalia: Pygophore elliptic in dorsal view (Figs. 17 – 18); proctiger large, posteriorly oval (Figs. 19 – 20), distinctly protruding posteriad (Figs. 17 – 18); paramere with group of long setae apically (Fig. 21); aedeagus as in Fig. 22. Measurements (mm). Micropterous form. Male / female (n = 2; holotype, paratype / 3 (for antennal segments III – IVb n = 2); median (minimum – maximum )). Body length: 1.62, 1.74 / 1.82 (1.76 – 1.92). Pronotum: anterior width: 0.38, 0.40 / 0.40 (0.40 – 0.42), humeral width: 0.61, 0.65 / 0.69 (0.67 – 0.71). Metathorax: maximum width across metapleura: 0.63, 0.71 / 0.71 (0.71 – 0.75). Lengths of antennal segments: I: 0.24, 0.24 / 0.24 (0.24 – 0.26), II: 0.16, 0.18 / 0.18 (no span), III: 0.24, 0.24 / 0.26 – 0.28, IVa: 0.14, 0.14 / 0.12 – 0.14, IVb: 0.20, 0.20 / 0.20 – 0.22). Length of metafemur: 0.52, 0.48 / 0.52 (0.50 – 0.53), metatibia: 0.61, 0.65 / 0.67 (0.61 – 0.67), metatarsus: 0.22, 0.26 / 0.24 (0.24 – 0.26). Variation. There is remarkable sexual dimorphism in colouration (head and thorax reddish in males, blackish brown in females) and shape of the metafemora (curved strongly in males, only indistinctly in females). Concerning individual variability in colouration, we noticed only the different shade of colouration of the metanotal plate in the three males examined, varying from reddish (Fig. 2) to dark reddish brown (Fig. 1).	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFA3D216FF7CFD24A75FE5E7.taxon	diagnosis	Differential diagnosis. Hebrus atlas sp. nov. most resembles the three Palaearctic species of Hebrus with known short-winged (H. fulvinervis, H. ruficeps) or wingless form (H. franzi), but it differs especially in the following unique combination of characters: Antennal segment IV subdivided by a membranous joint (fully divided in H. fulvinervis and H. ruficeps, undivided in H. franzi according to Cianferoni et al. 2016), head laterally with narrow ridge above bucculae (unique character), sexual dichroism in colouration of head and thorax, body widest across gibbose metapleura (body widest across humeral angles of pronotum in H. fulvinervis and H. ruficeps, across abdomen in H. franzi), short-winged form micropterous with hemelytra reaching posterior margin of mediotergite II (reaching posterior margin of mediotergite I in H. ruficeps and mediotergite IV in H. fulvinervis), and metafemur of male strongly curved (straight in H. fulvinervis and H. ruficeps, slightly curved in H. franzi). Hebrus iheriri Poisson, 1953, the only North African species of the subgenus Hebrusella, is known only as macropterous females but it should differ in the antennal segment IV fully divided and colouration of entire body yellowish (Poisson 1953). Despite the lack of knowledge of other important characters potentially distinguishing H. atlas and H. iheriri, we consider the possibility that H. atlas is the micropterous form of the latter as improbable. Three micropterous Afrotropical species of Hebrus are known, all with shortened metanotal elevations (a character apparently related to reduction of wings): H. aristomides Linnavuori, 1981 (Nigeria) and H. gembuanus Linnavuori, 1981 (Nigeria), both originally assigned to the subgenus Hebrus s. str. (Linnavuori 1981) and H. leleupi Cobben, 1982 (Democratic Republic of the Congo) described without subgeneric placement but regarded closely related to H. gembuanus (Cobben 1982 a). Hebrus aristomides differs among others in shape of the bucculae, rudiments of the hemelytra shorter than the metanotal elevation, metanotal elevation 3.2 × wider than long, shape of the parameres, and colouration of the head, pronotum and scutellum blackish with bluish metallic lustre (Linnavuori 1981).	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFA3D216FF7CFD24A75FE5E7.taxon	etymology	Etymology. Atlas (genitive Atlantis, masculine) the giant of ancient Greek mythology bearing the Cope of Heaven, as well as the northwest African Atlas Mountains named after him, the homeland of the new species. The species name is noun in apposition. Habitat. Unknown.	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFA3D216FF7CFD24A75FE5E7.taxon	distribution	Distribution (Fig. 65). Africa: Morocco: Al-Atlas al-Mutawassit [= Middle Atlas] Mts. (this paper). Subgeneric placement. Despite the strong superficial resemblance of H. atlas to H. fulvinervis and H. ruficeps, which are currently classified in the subgenus Hebrusella due to the fully divided antennal segment IV, H. atlas has antennal segment IV subdivided by membranous joint and should be placed in the subgenus Hebrus s. str. as defined by Poisson (1944). This may suggest relationships of the new species to some of the Afrotropical species of Hebrus s. str, such as H. gembuanus or H. leleupi. Unfortunatelly, the subgeneric classification of Hebrus is problematic and requires revision (see Summary).	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFA8D216FF7CFB12A0F4E665.taxon	description	Naeogeus eckerleini Jordan, 1954: 601. Holotype: ♂, France, Digne, Champ du Bés (ZMHB).	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFA8D216FF7CFB12A0F4E665.taxon	distribution	Distribution (Fig. 65). Europe: France: Alpes-de-Haute-Provence (Jordan 1954, Poisson 1957 a), Italy: Calabria (Bertrand 1958, Servadei 1967).	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFA8D216FF7CFB12A0F4E665.taxon	discussion	Comments. Jordan (1954) described the species based on the holotype (‘ Type’) deposited in his private collection, and an unspecified number of paratypes (‘ Cotypen’) deposited in Eckerlein’s private collection and in SDEI (1 Ƌ 1 ♀; actually 2 ♀ — D. Rédei, pers. comm.). Andersen (1995) indicated the depository of the holotype as? ZSMC, but according to the collection list (Anonymus 2016) no type of H. eckerleini is deposited in ZSMC. The Jordan Collection is deposited in ZMHB, including the male holotype of H. eckerleini (J. Deckert, pers. comm.). This taxon has not been recorded since the 1950 s and Tamanini (1979) expressed his doubts about the validity of the species, which may be a synonym of H. montanus or H. pusillus pusillus. Theoretically, Hebrus sericeus A. Costa, 1864, described from environs of Benevento in southern Italy and considered a synonym of H. pusillus pusillus, may be the senior synonym of this taxon. This taxon is in urgent need of revision.	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFA8D217FF7CF893A6B2E1F7.taxon	distribution	Distribution. Asia: Tajikistan (Kanyukova 1997).	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFA9D217FF7CFCBAA1ECE7B5.taxon	description	Hebrus jeanneli (f.) djaneti Poisson, 1953: 69. Syntypes: 1 ♂ 5 ♀, Algeria: Djanet (USNM).	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFA9D217FF7CFCBAA1ECE7B5.taxon	materials_examined	Material examined. ALGERIA: Hoggar Mts., near Tamanrasset, 7. – 8. v. 1975, 2 ♀, R. Linnavuori lgt. & det. (NMPC). MOROCCO: Haut Atlas Mts., Qued Reraïa, 33 km south of Marakech, 31. vii. 1959, 1 ♀, Eckerlein lgt., L. Hoberlandt 1965 det. as H. dupuisi, P. Kment revid. (NMPC); Tamegroute env., 540 m a. s. l., 14. iv. 2012, 1 Ƌ, P. Krásenský lgt. (ZJPC).	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFA9D217FF7CFCBAA1ECE7B5.taxon	distribution	Distribution (Fig. 66). North Africa: Algeria (south-east): Tassili Mts. (Poisson 1953), Hoggar Mts. (this paper); Morocco (new record). New species for Morocco.	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFA9D217FF7CFCBAA1ECE7B5.taxon	discussion	Comments. In the original description, the taxon was treated twice as a form (Hebrus jeanneli f. djaneti — pp. 69, 70) and three times as a trinomen which is fitting subspecific status (Hebrus jeanneli djaneti — pp. 71, 81). According to the Article 45.6.4 (ICZN 1999), the name djaneti must be regarded as available in subspecific rank. This subspecies has not been recorded since its description. According to the original description, the species is very similar to H. montanus, but the hind tibia of the male, as depicted by Poisson (1953: Fig. 3 D) and also in the only examined male, bears no long hairs. The three examined females (Figs. 23 – 24) fits the original description well. The male from Morocco: Asni (Fig. 25), identified as H. montanus by Kment & Kanyukova (2010), has similar colouration but it bears a row of erect setae in the middle of the upper surface of the hind tibia. No depository of the types was mentioned in the original description (Poisson 1953) nor in the Palaearctic catalogue by Andersen (1995). The collection at the USNM in Washington includes four card-mounted and two slide-mounted specimens: ‘ I found two slides of H. jeanneli djaneti, one with the genitalia only and labelled “ Hebrus jeanneli djaneti nov., [?] Tassili, F. Bernard. ”, the second one has the genitalia, hemelytra, and legs and is labelled “ Hebrus jeanneli djaneti nov., [?], Tassili, F. Bernard. ” There are four carded H. jeanneli djaneti, two on separate cards (a male and female) and pins and two on separate cards (apparently both females) but one pin, all are from Djanet. The separate male and female have a label saying “ scutellum different ”. ’ (T. J. Henry, pers. comm).	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFA9D217FF7CFEE2A48AE24F.taxon	distribution	Distribution (Fig. 66). North Africa: Egypt (Gadalla & Saleh Ahmad 2005, Saleh Ahmad & Gadalla 2005). Asia: Sinai (Gadalla & Saleh Ahmad 2005, Saleh Ahmad & Gadalla 2005). Afrotropical Region: Ethiopia (Cobben 1982 b), Kenya (Poisson 1944), Niger (Linnavuori 1981), Tanzania: Zanzibar (Poisson 1944).	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFA9D217FF7CFEE2A48AE24F.taxon	discussion	Comments. The head in lateral view was figured by Poisson (1957 b: 353). No depository of the types was mentioned in the original description (Poisson 1953) nor in the Palaearctic catalogue by Andersen (1995). The collection at the USNM in Washington includes five card-mounted specimens, one from Tiwi (male), two from Taveta (apparent females), one from Voï (female), and one from Ethiopia, Hynes, Lac Awasa (apparent male); three of the specimens are certainly syntypes of H. jeanneli jeanneli (T. J. Henry, pers. comm).	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFAAD214FF7CF9CEA59AE655.taxon	distribution	Distribution (Fig. 67). Asia: Afghanistan (Kment & Kanyukova 2010), Tajikistan (Kanyukova 1997), Uzbekistan (Kanyukova 1997).	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFAAD215FF7CF883A0E5E0B2.taxon	distribution	Distribution. Caucasus (without details) (Horváth 1929). Andersen (1995) interpreted the distribution of H. liliimacula as ‘ Transcaucasia’ and Kanyukova (1997) expected its distribution in Armenia or Azerbaijan. However, examination of rich material of Hebrus from the Caucasus, Turkey and Iran did not led to rediscovery of the species (Kanyukova 1997, 2006; Kment & Kanyukova 2010; Fent et al. 2011).	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFAAD215FF7CF883A0E5E0B2.taxon	discussion	Comment. This species with characteristic ‘ lilly-shaped’ whitish pattern on the membrane was described by Horváth (1929) based on male and female syntypes from the Caucasus collected by H. Leder. The species has not been collected since. Kanyukova (1997) redescribed and keyed the species based on a single female originating from the same series as the syntypes. Proper description of the male is therefore missing. Checking for a male syntype within the type series deposited in HNHM might shed new light on this problem.	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFABD212FF7CFDA1A59DE006.taxon	materials_examined	Material examined. ALBANIA: Shkopet pr. Lezha, Mat river, 2. vii. 2012, 1 ♀, L. Blažej lgt. (MMBC). BULGARIA: Blagoevgrad Province: Strumyani, 8. vi. 2013, 1 ♀, E. Ezer lgt., P. Kment det. (NMPC). FRANCE: Corse: Navaccia River, 27. vii. 1955, 1 Ƌ 1 ♀, Eckerlein lgt., P. Kment det. (NMPC). ISRAEL: Huleh, 26. viii. 1980, 1 ♀, D. Furth lgt., P. Kment det. (NMPC). ROMANIA: Mehedinți county: Cerneți [stream], 44.64 ° N 22.701944 ° E, 53 m a. s. l., 26. vii. 2012, 1 Ƌ, G. M. Berchi lgt., P. Kment det. (GBCR). TURKEY (Asian Part): Adana Province: Taurus Mts., Pozanti, 24. v. 1960, 1 Ƌ, Eckerlein lgt., L. Hoberlandt 1965 det. as H. eckerleini, P. Kment revid. (NMPC).	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFABD212FF7CFDA1A59DE006.taxon	distribution	Distribution (Fig. 68). Europe: Albania (Josifov 1970), Bosnia Herzegovina (Kment & Jindra 2006), Bulgaria (Josifov 1958, 1960, 1963; Kment & Jindra 2006), France: Vendée, Corse (Poisson 1944, 1957 a; Kment & Kanyukova 2010), Greece (Kment & Kanyukova 2010), Macedonia (Wagner 1962, Protić 1998), Romania (Horváth 1897, this paper), Russia: South European Territory (Kanyukova 1997, 2006; Golub et al. 2014), Serbia (Kment & Jindra 2006). North Africa:? Egypt (Stichel 1956, Andersen 1995, Gadalla & Saleh Ahmed 2005, Salleh Ahmed & Gadalla 2005), Morocco (Poisson 1944, Kment & Kanyukova 2010),? Tunisia (Slimani et al. 2016). Asia: Armenia (Kanyukova 1997, 2006), Azerbaijan (Kanyukova 1997, 2006; Kment & Jindra 2006; Kment & Kanyukova 2010), Georgia (Kanyukova 1997, 2006), Iran: Fars, Gilan, Golestan, Khorasan, Tehran (Kanyukova 1997, 2006; Kment & Kanyukova 2010; Ghahari et al. 2013), Israel (Nieser 1995, this paper), Jordan (Kment & Kanyukova 2010), Lebanon (Kment & Kanyukova 2010),? Syria (Stichel 1956, Andersen 1995), Turkey (Asian part) (Kment & Jindra 2006, Fent et al. 2011). A record of H. montanus from Slovakia referring to Kment et al. (2003) was cited in error by Aukema et al. (2013 a); the record refers to H. ruficeps. A record from Hormozgan Province of Iran (Kment & Kanyukova 2010) was based on a misidentified specimen of H. pusillus arabicus (see below). Slimani et al. (2016) recorded H. montanus from Tunisia; however, the specimens on the photograph they provided are very dark, probably greasy, and it is difficult to determine to which species they really belong. Previous records from Egypt and Syria require further confirmation (see Fent et al. 2011). The occurrence of H. montanus in Israel and Romania is confirmed herein.	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFABD212FF7CFDA1A59DE006.taxon	discussion	Comments. Kolenati (1857) listed the type locality of H. montanus as follows: ‘ Habitat in superficie aquae lacus montani sic dicti Sullü-ghöll (lacu hirudinum), ubi Pinacobdella Kolenatii Diesing frequens, in monte Koepesdagh provinciae Transcaucasicae Karabagh prope pagum Tschai-Kent, insidiator ovulis hujusce Pinacobdellae. ’ [= Habitat on water surface of mountain lake called as Sullü-ghöll (lake of leeches; in Turkish sülük = leech, göl = lake), where Pinacobdella kolenatii Diesing is frequent, in Kopetdag Mt., in Karabagh Province of Transcaucasia, near Tschai-Kent village, being here predator of Pinacobdella eggs]. So far, nobody has succeeded in precisely identifying the type locality, other than its location in Mountain Karabagh Republic, an Armenian exclave in Azerbaijan. The names of the lake and mountain seem to disappear nowdays, but there are two villages of the name Chaykend or Çaykənd (which means ‘ built near river) ’ with the following coordinates: 39 ° 47 ′ 30.8 ″ N 46 ° 35 ′ 58.6 ″ E and 40 ° 07 ′ 47.1 ″ N 46 ° 03 ′ 38.1 ″ E; the first is inside Mountain Karabagh Republic, the second is under the control of Azerbaijan (M. Kalashian, pers. comm.). We follow here the concept of the species as redescribed and illustrated by Kanyukova (1997). Some potential problems concerning taxonomy and identification of the species are discussed under H. eckerleini, H. jeanneli djaneti and H. pusillus pusillus.	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFACD212FF7CFE6EA08AE3FD.taxon	materials_examined	Material examined. UZBEKISTAN: Zeravshan river valley, reservation near Samarkand, 12. v. 1990, 3 Ƌ 1 ♀, Z. Jindra lgt., P. Kment det. (ZJPC, NMPC).	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFACD212FF7CFE6EA08AE3FD.taxon	distribution	Distribution (Fig. 69). Asia: Afghanistan (Kment & Kanyukova 2010), Iran (Kment & Kanyukova 2010), Tajikistan (Kanyukova 2007), Turkmenistan (Kanyukova 2007), Uzbekistan (Kanyukova 2007).	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFACD212FF7CFCEBA542E5EB.taxon	distribution	Distribution (Fig. 70). Europe: Russia: South of the European Territory (Kanyukova 1997, 2006; Prokin et al. 2009), Ukraine: Crimea (Kanyukova 1997, 2006). Asia: Afghanistan (Kment & Kanyukova 2010), Armenia (Kanyukova 1997, 2006), Azerbaijan (Kanyukova 1997, 2006), Georgia (Kanyukova 1997, 2006), Iran: Golestan, Khorasan, Tehran, West Azerbaijan (Kanyukova 1997, 2006; Kment & Kanyukova 2010; Ghahari et al. 2013), Kazakhstan: Asian part (Kanyukova 1997, 2006), Tajikistan (Kanyukova 1997, 2006), Turkmenistan (Kanyukova 1997, 2006), Turkey (Asian part) (Kment & Jindra 2006, Fent et al. 2011), Uzbekistan (Kanyukova 1997, 2006).	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFACD211FF7CFB17A131E3E3.taxon	materials_examined	Material examined. Hebrus pusillus pusillus: BULGARIA: Blagoevgrad Province: Sandanski, 18. vii. 1956, 1 Ƌ 1 ♀, L. Hoberlandt lgt., P. Kment det. (NMPC). Pleven Province: Kajlăka [= Kayl’ka] hotel 2 km E of Pleven, 100 – 150 m a. s. l., Locality No. 52 / 87, 24. vii. 1978, 1 ♀, P. Lauterer lgt., P. Kment det. (MMBC). CROATIA: Privlaka, Bosut river, 45.21327 ° N 18.85828 ° E, 74 m a. s. l., 16. viii. 2015, 1 Ƌ, G. M. Berchi lgt. & det. (GBCR). CZECH REPUBLIC: Bohemia: Heřmanice (5156), flooded basalt quarry, 26. iv. 2002, 2 ♀, M. Fikáček lgt., P. Kment det. (NMPC); Mařeničky, Rašeliniště Mařeničky NR (5254; 50 ° 47 ′ 41.2 ″ N 14 ° 40 ′ 27.8 ″ E), peatbog, 1. x. 2004, 1 Ƌ 2 ♀, P. Kment lgt. & det. (NMPC); Staré Splavy, Swamp NNM (5460; 50 ° 34 ′ 48.3 ″ N 14 ° 40 ′ 02.8 ″ E), peatbog, 265 m a. s. l., 25. viii. 2009, 1 spec., P. Kment lgt. & det. (JSBC); Jabkenice (5656), 5. i. 1975, 1 ♀, P. Bílek lgt., Z. Jindra det. (ZJPC); Neudau [= Nejda] (5743), 24. x. 1937, 1 Ƌ, P. Kment det. (NMPC); Otovice u Karlových Varů (5743), 1 Ƌ 1 ♀, P. Kment det. (NMPC); Čelákovice (5854), 2. v. 1971, 1 Ƌ, Martiš lgt., P. Kment det. (NMPC); Horní Ředice (5961 b; 50 ° 05.508 ′ N 15 ° 57.064 ′ E), NR Žernov, marsh at Mordýř pond, 20. iv. 2007, 13 Ƌ 11 ♀, M. Boukal lgt., P. Kment det. (NMPC); Lnáře (6548), 14. vi. 1944, 1 Ƌ 2 ♀, Mařan and Z. Tesař lgt., L. Hoberlandt det. (NMPC); Lnáře (6548), Malý Pálenec [pond?], ix. 1942, 3 Ƌ 3 ♀, Mařan lgt., L. Hoberlandt det. (NMPC); Horusice, Ruda NNR (6854, 49 ° 09 ′ 03.9 ″ N 14 ° 41 ′ 27.7 ″ E), peatbog, extracted from wet Sphagnum, 416 m a. s. l., 13. ix. 2010, 2 ♀, P. Kment lgt. & det. (NMPC); Ponědrážka, Rašeliniště Hovízna NR (6854; 49 ° 08 ′ 34.6 ″ N 14 ° 41 ′ 40.2 ″ E), peatbog with pools, 16. ix. 2010, 1 Ƌ, P. Kment lgt. & det. (NMPC); Hluboká nad Vltavou (6952), 20. vi. 1974, 1 Ƌ, P. Bílek lgt., Z. Jindra det. (ZJPC); Dolní / Horní Slověnice u Českých Budějovic (6954), 3. viii. 1971, 1 Ƌ 1 ♀, Z. Jindra lgt. & det. (ZJPC); Stráž nad Nežárkou (6955), vii. 1957, 1 Ƌ, J. Dostál lgt., P. Kment det. (MMBC). Moravia: Děhylov (6175), Štěpán NR, pond, 22. iii. 2003, 1 ♀, M. Fikáček lgt., P. Kment det. (NMPC); Křižánky, Kyšperský rybník pond (6362; 49 ° 41 ′ 10.1 ″ N 16 ° 03 ′ 32.8 ″ E), waterlogged shore, 21. v. 2000, 1 Ƌ 3 ♀, P. Kment lgt. & det. (NMPC); Rudice, Rudická jezírka (6666, 49 ° 20 ′ 09.7 ″ N 16 ° 43 ′ 24.4 ″ E), flooded pits after iron ore mining, 3. iv. 1999, 1 Ƌ 1 ♀, 1. v. 1999, 5 Ƌ 13 ♀, 6. vi. 1999, 1 ♀, P. Kment lgt. & det. (NMPC, MHBC); Velká Bíteš, Dolní Klečanský rybník pond (6763; 49 ° 16 ′ 39 ″ N 16 ° 14 ′ 28 ″ E), waterlogged grassy shore, 1. v. 2000, 6 Ƌ 11 ♀, P. Kment lgt. & det. (NMPC, MUBC); Milotice, Písečný rybník pond (7068, 48 ° 57 ′ 46 ″ N 17 ° 09 ′ 22 ″ E), 26. vii. 2000, 1 ♀, M. Horsák lgt., P. Kment det. (NMPC); 16. iv. 2009, 2 ♀, M. Straka lgt., P. Kment det. (MSHC); Lednice, Lednické rybníky NNR, Mlýnský rybník pond, camp Apollo env. (7266), 12. vi. 1982, 1 ♀, Z. Jindra lgt. & det. (ZJPC); Lednice, Lednické rybníky NNR, northern shore of Mlýnský rybník pond (7266), walking on wet mud, 4. vii. 1998, 1 ♀, P. Kment & M. Horsák lgt., P. Kment det. (NMPC); Sedlec + Hlohovec + Valtice, Lednické rybníky NNR, Nesyt pond (7266), 17. vii. 1997, 1 ♀, P. Kment lgt. & det. (NMPC); Valtice, Úvaly, Úvalský rybník pond (7266, 48 ° 44 ′ 47 ″ N 16 ° 42 ′ 46 ″ E), 29. iv. 2009, 1 ♀, M. Straka lgt., P. Kment det. (MSHC); Lanžhot, Soutok game preserve, Dědova pískovna (7367; 48 ° 38 ′ 02.0 ″ N 16 ° 57 ′ 33.6 ″ E), vegetated pool in abandoned sandpit, 14. v. 2016, 1 Ƌ, P. Kment lgt. & det. (NMPC). FINLAND: Lojo, 10. vi. 1916, 1 ♀, H. Lindberg lgt., P. Kment det. (NMPC). GREECE: 3 km NW of Igoumenítsa, salt soil, 29. v. 1997, 1 ♀, P. Bulirsch lgt. (ZJPC). LITHUANIA: Borussia, Kurische Nehrung [= Prussia, Kuršių nerija Peninsula], 1 ♀, L. Hoberlandt det. (NMPC). NETHERLANDS: Holland, Winterswijk, 1894, 1 ♀, Fokker lgt., L. Hoberlandt det. (NMPC). ROMANIA: Caraș-Severin county: Caransebeș, litter on pond banks (45.4188 ° N 22.1875 ° E), 217 m a. s. l., 24. xii. 2014, 2 Ƌ 4 ♀ (macropterous), G. M. Berchi lgt. & det. (GBCR). Mehedinți county: Izimșa [stream], 44.175277 ° N 22.960277 ° E, 55 m a. s. l., 27. vii. 2012, 1 ♀, G. M. Berchi lgt. & det (GBCR). Timiș county: Dumbrăvița, litter on pond banks (45.80638 ° N 21.26416 ° E), 96 m a. s. l., 19. xii. 2014, 1 ♀ (macropterous), I. A. Rădac leg., G. M. Berchi det. (GBCR); Jimbolia, litter on pond banks (45.8099 ° N 20.7177 ° E), 75 m a. s. l., 26. xii. 2014, 1 ♀, I. A. Rădac leg., G. M. Berchi det. (GBCR). Vâlcea county: Marcea, stream, 44.918055 ° N 24.238055 ° E, 186 m a. s. l., 27. vii. 2012, 1 Ƌ, G. M. Berchi lgt. & det (GBCR). RUSSIA: North European Territory: Leningrad Province, Karelian Isthmus, 4 km S of Dibuny railway station, river bank, 13. vi. 2004, 1 ♀, P. Kment lgt. & det. (NMPC). SLOVAKIA: Gánovce (6987), Gánovké travertíny NM, on mud, 18. v. 2002, 3 Ƌ 16 ♀, M. Mantič lgt., P. Kment det. (MMHC); Kúty (7267), 31. v. 1986, 1 ♀, Z. Jindra lgt. & det. (ZJPC); Veľké Leváre, Abrod NNR (7467), 17. v. 1982, 1 Ƌ 2 ♀, O. Majzlan lgt., P. Kment det., (MMBC); Račištorf [= Bratislava, Rača] (7768), 23. iii. 1941, 1 Ƌ, L. Korbel lgt., P. Kment det. (MMBC); Tvrdošovce (7974), small pond, in water, 20. iv. 1999, 1 ♀, M. Mantič lgt., P. Kment det. (MMHC); Veľký Meder, Čalovo (8072), 15. vii. 1953, 1 ♀, P. Lauterer lgt., J. L. Stehlík 1962 det. (MMBC). Hebrus dupuisi: FRANCE: Pyren. orient., Font Estramer, 26.7. – 11.8.1956, 1 Ƌ 1 ♀, Weber lgt., L. Hoberlandt 1985 det. (NMPC). SPAIN: Pyren. or., Ripoli, 29. – 30. vii. 1956, 3 ♀, L. Hoberlandt 1965 det. (NMPC).	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFACD211FF7CFB17A131E3E3.taxon	distribution	Distribution (Fig. 71). Europe: Albania (Josifov 1970), Austria (Rabitsch 2005), Belgium (Stoffelen et al. 2013), Bosnia Herzegovina (Benedek 1970, Protić 1998), Bulgaria (Josifov 1960, 1963, 1964), Byelorussia (Lukashuk 1997), Croatia (Horváth 1897, Benedek 1970, Protić 1998, Kment & Beran 2011), Czech Republic (Hoberlandt 1977), Denmark (Andersen & Gaun 1974), Estonia (Lukashuk 1997, Coulianos 2005), Finland (Rintala & Rinne 2010), France (Wagner 1954, as H. dupuisi; Poisson 1957 a; Jacquemin 2006), Germany (Hoffmann & Melber 2003, Wachmann et al. 2006, Strauss & Niedringhaus 2014), Great Britain (Savage 1989), Greece (Lindberg 1922, Royer 1924, Linnavuori 1954, Josifov 1959, Zimmermann 1982, this paper), Hungary (Horváth 1897, Benedek 1970, Boda et al. 2015), Italy (Servadei 1967, Tamanini 1979), Latvia (Lukashuk 1997), Lithuania (new record), Luxembourg (Reichling & Gerend 1994), Macedonia (Protić 1998), Moldavia (Derzhansky 1984, 1997), Montenegro (Protić 1998), the Netherlands (Aukema et al. 2002), Norway (Coulianos 1998, Coulianos & Ossiannilsson 1976, Roth & Coulianos 2014), Poland (Jaczewski & Wróblewski 1976, Wróblewski 1980), Portugal (Seabra 1926, 1941; Baena & Vázquez 1986; Baena 1996; Kment 2006), Romania (Horváth 1897, Benedek 1970), Russia: North, Central, and South European Terriotory (Kanyukova 1997, 2006), Serbia (Horváth 1897, Benedek 1970, Protić 1998), Slovakia (Benedek 1970, Hoberlandt 1977, Reduciendo Klementová et al. 2015), Slovenia (Protić 1998, Gogala 2003), Spain (Ribes & Sauleda 1979, as H. dupuisi; Nieser & Montes 1984; Baena & Vázquez 1986; Baena 1996; Pagola-Carte & Ribes 2007), Sweden (Coulianos & Ossiannilsson 1976), Switzerland (Dethier & Matthey 1977), Ukraine (incl. Crimea) (Putshkov & Putshkov 1996, Kanyukova 1997, Grandova 2014),? Turkey (European part) (see Fent et al. 2011). Macaronesia: Canary Islands: La Gomera, Gran Canaria, Tenerife (Poisson 1954 a, b, as H. pusillus canariensis; Baena 1996; Aukema et al. 2013 b). North Africa:? Algeria (Lindberg 1922),? Egypt (Lindberg 1922, Linnavuori 1964, Gadalla & Saleh Ahmed 2005, Saleh Ahmed & Gadalla 2005),? Libya (Mancini 1936),? Morocco (Gheit 1995),? Tunisia (Lindberg 1922). Asia:? Iran: Gilan, Kordestan (Ghahari et al. 2013),? Israel (Lindberg 1922; Linnavuori 1960, 1986),? Lebanon (Lindberg 1922, Nieser & Moubayed 1985), Russia: West Siberia, East Siberia, Far East (Kanyukova 1997, 2006; Vinokurov 2005; Vinokurov et al. 2010),? Sinai (Gadalla & Saleh Ahmed 2005, Saleh Ahmed & Gadalla 2005),? Syria (Stichel 1956, Andersen 1995),? Turkey (Asian part) (Linnavuori 1986, but see Fent et al. 2011). Confirmed occurrence in Greece, new species for Lithuania. All previous records of H. pusillus from Turkey, Transcaucasia, Iran and central Asia revised recently are misidentifications of other species (see Kanyukova 1997, Kment & Jindra 2006). The occurrence of the species in Morocco, Algeria, Libya, Egypt, Sinai, Israel, Syria, Turkey and Iran requires confirmation (cf. Andersen 1995, Kanyukova 1997, Fent et al. 2011, Aukema et al. 2013 a). The record of H. pusillus from Kazakhstan by Esenbekova et al. (2015) might be based on mistaking the names H. pilipes as H. pusillus, resulting in the omission of H. pilipes in that paper.	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFACD211FF7CFB17A131E3E3.taxon	discussion	Comments. The subspecies status of H. pusillus canariensis was questioned by Nieser & Montes (1984) and Baena & Báez (1990). Andersen (1995) mentioned in his catalogue that H. dupuisi is probably a synonym of H. pusillus (Fallén, 1807). Finally, Baena (1996) synonymized both taxa with H. pusillus pusillus, using material of the latter species from Europe, Iberian Peninsula and North Africa. Currently, there are 3 species of Hebrus s. str. recorded from North Africa, but no confirmed record of H. pusillus. This suggests that Baena might have been working with mixture of species. The examined specimens of H. dupuisi from East Pyrenees Mts. have pale yellowish legs and colouration and golden pilosity similar to that of H. montanus, but the structure of the metanotal elevation and erect hairs on the metatibia of the single examined male resembles that of H. pusillus. The status of H. dupuisi, H. pusillus canariensis and H. pusillus pombasi require further confirmation.	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFACD211FF7CFB17A131E3E3.taxon	description	Linnavuori (1986) provided illustrations of the pronotum and metanotal elevations and commented on colouration of populations from Israel and Turkey ascribed by him as H. pusillus pusillus. Kment & Jindra (2006), Kment & Kanyukova (2010) and Fent et al. (2011) recorded only H. montanus from that area instead of H. pusillus pusillus. However, material from the Near East is scarce and identification of isolated females is tentative. The reexamination of voucher specimens of Linnavuori (1960, 1986) is needed. Andersen (1995) gave the type depository for Hebrus letzneri Scholtz, 1847 as UWCP with a question mark. However, the majority of types in Wrocław were destroyed during World War II (J. Gorczyca, pers. comm.).	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFAFD211FF7CFD09A7F8E4C7.taxon	materials_examined	Material examined. IRAN: Hormozgan: Isin (Loc. No. 198), 11. – 15. iv. 1973, 1 ♀, Exp. Nat. Mus. Praha, E. Kanyukova 2006 det. as H. montanus, P. Kment det. (NMPC). OMAN: Nakhl, Wadi Mistal, 24. iii. 1997, 3 Ƌ 1 ♀, E. Heiss lgt., P. Kment det. (EHIA, NMPC).	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFAFD211FF7CFD09A7F8E4C7.taxon	distribution	Distribution (Fig. 71). Asia: Iran: Hormozgan (Kment & Kanyukova 2010, as H. montanus), Iraq (Linnavuori 1986, 1994), Oman (new record), Saudi Arabia (Linnavuori 1986), United Arab Emirates (Linnavuori et al. 2011), Yemen (Brown 1951, as H. pusillus var. rufescens; Linnavuori 1971; Linnavuori & van Harten 1997, 2002). Afrotropical Region: Sudan (Port Sudan on Red Sea Coast) (Linnavuori 1986). New species for Iran and Oman.	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFAFD211FF7CFD09A7F8E4C7.taxon	discussion	Comments. Linnavuori (1971) illustrated only parameres of the new subspecies based on Yemeni specimens. Linnavuori (1986) provided illustrations of metanotal elevations, hemelytron and paramere of specimens from Iraq and Saudi Arabia, and compared the subspecies to Near East populations ascribed by him as H. pusillus pusillus. However, using the key by Kanyukova (1997, 2006), H. pusillus arabicus with pale yellowish legs, would run to H. montanus, although the pilosity of the male metatibia is more similar to that of H. pusillus pusillus. The relationship and subspecific status of H. pusillus arabicus and the nominotypical H. pusillus pusillus is worthy of re-examination; H. pusillus arabicus possibly deserves specific rank.	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFAFD211FF7CFA2AA5F6E6B1.taxon	distribution	Distribution (Fig. 71). Macaronesia: Cape Verde Islands: Brava, Santo Antão (Lindberg 1959, Báez et al. 2005). Afrotropical Region: Ivory Coast (Poisson 1968).	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFAFD211FF7CFA2AA5F6E6B1.taxon	discussion	Comments. The species has not been recorded since its description, except a single record from the Ivory Coast by Poisson (1968). However, the latter record requires confirmation. The subspecific status of this taxon and its relationship with H. pusillus pusillus requires re-examination. In the MZHF are 9 specimens from the type locality: ‘ holotype nr. 11635, 3 spec. on the same pin, no locality label, determination handwritten label by Poisson, no year; allotype nr. 11636, 1 spec., Santo Antão: Pombas, 23. – 25.12.1953, Lindberg, + Poisson determination label in 1956; paratype nr. 11637, 2 spec. on the same pin, Santo Antão: Pombas, 23. – 25.12.1953, Lindberg, + Poisson determination label in 1956; paratype (red label no number), 3 spec. on two pins, Santo Antão: Pombas, 23. – 25.12.1953, Lindberg, + Poisson determination label in 1956. ’ (L. Huldén, pers. comm.).	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFB0D20CFF7CFF71A65FE51F.taxon	materials_examined	Material examined. CYPRUS: Kyrenis, Ammos, 7. vii. 1939, 1 ♀, H. Lindberg lgt. & det., P. Kment revid. (NMPC). Descriptive notes. The female examined (Figs. 26, 29 – 39) fits the redescription by Linnavuori (1994), although the scutellum seems less bifid apically (see Figs. 36 – 37) and the membrane of the hemelytra is uniformly milky-brown and of the same colour as the posterior portion of the clavus (Fig. 26). The clavus has sparse pilosity and corium relatively dense pilosity (Fig. 38). The membrane reaches the posterior 2 / 3 of mediotergite VII, leaving the apex of the dorsum free (Fig. 26).	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFB0D20CFF7CFF71A65FE51F.taxon	description	Measurements (mm): Body length: 1.66. Head width (across eyes): 0.4. Pronotum: median length: 0.38, anterior width: 0.38, humeral width: 0.69. Lengths of antennal segments: I: 0.18, II: 0.14, III: 0.20, IVa: 0.08, IVb: 0.24. Length of metafemur: 0.51, metatibia: 0.63, metatarsus: 0.22.	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFB0D20CFF7CFF71A65FE51F.taxon	distribution	Distribution (Fig. 67). Asia: Cyprus (new record), Israel (Horváth 1896, as Syria; Linnavuori 1961, 1994), Jordan (Linnavuori 1961),? Syria (Stichel 1956, Andersen 1995). New species for Cyprus. The type locality in the original description was given as ‘ Syria: Kaiffa (Reitter) ’ (Horváth 1896). Linnavuori (1961) listed four more specimens, one from Jordan and three from Judea, all from J. Sahlberg’s collection. Linnavuori (1994) interpreted the type locality as ‘ Israel, Haifa’ and speculated that ‘ since all new finds from Israel belong to H. pusillus pusillus (Fallén, 1807), H. syriacus has possibly become extinct. ’ Distribution of H. syriacus in Syria requires confirmation as it seems to be based on the above mentioned error.	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFB0D20CFF7CFF71A65FE51F.taxon	discussion	Comments. Horváth (1896) described the species based on the macropterous female, but did not specify the number of specimens, although there was probably only one. Andersen (1995) listed the type material as syntypes from Syria. Linnavuori (1994) redescribed the species based on a single female, presenting it as a ‘ type’, and explicitely stated: ‘ The holotype is the only known specimen’. This statement fulfilled the requirements of Article 74.6 of the ICZN (1999) for fixation of the lectotype by inference of the holotype. Horváth (1896) compared the new species to H. ruficeps: ‘ Similar to H. ruficeps, easily distinguished by body smaller and narrower, colouration of body pale, pronotum anteriorly not declivous, and posteriorly less dilated. Length 1.5 mm. ’ Linnavuori (1994) redescribed the species based on the ‘ holotype’ and provided illustrations of the head in lateral view and thorax and hemelytron in dorsal view, and provided the following differential diagnosis: ‘ H. syriacus is readily distinguished from H. pusillus and the related species by the smaller and narrower body, much narrower and more parallel-sided pronotum and, especially, the apically bifid scutellum’. Neither Horváth (1896) nor Linnavuori (1994) described apical antennal segments. Jordan (1954) included H. syriacus in a comparative table of seven European Hebrus species, and provided lengths of the antennal segments (mm): I — 0.18, II — 0.16, III — 0.20, IVa — 0.10, IVb — 0.24. However, the material Jordan (1954) used for comparison was not listed in that paper and remains unknown. Linnavuori (1994) did not assign H. syriacus to any of the subgenera; however, the metanotal elevation apically truncated with a wide V-shaped incision (Linnavuori 1994: Fig. 4 b), antennal segments I and II subequal in length (Stichel 1955, Linnavuori 1994), antennal segment IVb more than twice as long as IVa (Jordan 1954), antennal segment I little longer than eye diameter (Horváth 1896, Stichel 1955), and its small size (1.50 – 1.66 mm) suggest placement of H. syriacus in Hebrusella. (Body length of Euro-Mediterranean Hebrusella is 1.30 – 1.80 (exceptionally 1.93) mm, in Hebrus s. str.> 1.70 (usually 1.80) mm — see Poisson 1944, 1953; Jordan 1954; Wagner 1954, 1957; Kanyukova 1997, 2006 )). On the other hand, examination of the antenna by scanning electron microscope revealed that segment IV is subdivided only by a membranous joint (Figs. 34 – 35), which supports its placement in Hebrus s. str. sensu Poisson (1944). However, the subgeneric classification of Hebrus is problematic and requires revision (see Summary).	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFB2D20CFF7CFB4AA6E2E6F5.taxon	materials_examined	Material examined. MOROCCO: Massa River, 30 km NE Tiznit, 27. and 29. vii. 1959, 162 spec. (Ƌ ♀), Eckerlein lgt., P. Kment det. (NMPC); Tamegroute env., 540 m a. s. l., 14. iv. 2012, 1 Ƌ, P. Krásenský lgt. (ZJPC).	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFB2D20CFF7CFB4AA6E2E6F5.taxon	distribution	Distribution (Fig. 72). North Africa: Algeria (south-east): Tassili Mts. (Poisson 1953), Morocco (new record). Afrotropical Region: Burkina Faso, Cameroon, Chad, Nigeria (all Linnavuori 1981). New species for Morocco.	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFB2D20CFF7CFB4AA6E2E6F5.taxon	discussion	Comment. The parameres of the species were illustrated by Linnavuori (1981). The species differs from H. montanus and H. pusillus by the nearly completely brown clavus and male metatibia without long erect setae. No depository of the types was mentioned in the original description (Poisson 1 953). Andersen (1995) indicated the type depository as MNHN in Paris, but a subsequent search for the types was unsuccesfull (E. Guilbert, pers. comm.). The collection at the USNM in Washington includes four card-mounted and two slidemounted specimens: ‘ I found two slides of H. vaillanti, one with only the antennae and the labelled “ H. vaillanti n. sp., something [can’t read], Tassili, Antennae, female symbol. ”, the second slide has the male genitalia under one cover slip and the hemelytra and three legs under the other and the labels “ Hebrus vaillanti n. sp., male symbol; and + 2 hemelytra, female symbol [I presume the hemelytra and legs belong to the female], Tassili, F. Bernard, st- 14 et 20. ” There are also four carded specimens, two on each of two pins. One male and one female are on one, and two females are on the other. There is no locality data, but one has the number 14 and the other 20 (and one has Tassili, Bernard) ’ (T. J. Henry, pers. comm).	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFB3D20DFF7CFF7EA563E355.taxon	diagnosis	Diagnosis. Poisson (1944, 1957 a) diagnosed the new sugbenus Hebrusella based on a single character: ‘ Antennal segment IV completely divided into two segments (IVa and IVb) by true articulation’ (see Poisson 1944: Fig. 2 C; Poisson 1957 a: Fig. 129 B; Andersen 1982: Fig. 114 c; Kanyukova 2006: Fig. 372), whereas the subgenus Hebrus s. str. has ‘ segment IV incompletely divided by membranous ring, not fully articulated’ (see Poisson 1944: Figs. 1 D, 3 D, 5 D; Poisson 1957 a: Figs. 130 B, 134 C; Andersen 1982: Fig. 114 b and Pl. C; Kanyukova 2006: Fig. 373). This character is difficult to observe and thus far, no other reliable character has been added to the diagnosis of the subgenus.	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFB3D209FF7CFD83A7CDE6C3.taxon	description	Hebrus ruficeps (misidentification): Lindberg (1922): 16 (faunistics).	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFB3D209FF7CFD83A7CDE6C3.taxon	materials_examined	Type material examined. Syntypes: 1 ♀ (macropterous), ‘ Malomviz / Pável [p, white label] // fulvinervis [hw, underlined with red] / det. Horváth [p, white label] // TYPUS [p, red label] ’ (HNHM); 1 Ƌ 1 ♀ (brachypterous), ‘ Nagyszeben / Coll. Fuss [p, white label] // pusillus [hw] Coll. Fuss [p, white label] // fulvinervis [hw, underlined with red] / det. Horváth [p, white label] // Ƌ or ♀ [hw; respectively] / TYPUS [p, red label] ’ (HNHM). Additional material examined. BULGARIA: Blagoevgrad Province: Gara Sandanski, Struma, 20. vii. 1956, 3 Ƌ 3 ♀, L. Hoberlandt lgt., P. Kment det. (NMPC); Gotse Delchev (→ Ognianovo), Mesta river banks, 500 m a. s. l., Locality No. 13 B / 72, 16. viii. 1972, 1 Ƌ (brachypterous), A. Merta lgt. (MMBC).	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFB3D209FF7CFD83A7CDE6C3.taxon	description	Redescription. Macropterous and brachypterous form. Colouration (Figs. 40 – 43). Head yellowish to reddish brown, in some specimens somewhat darker anteriorly, sometimes narrowly infuscate around ocelli. Eyes red to dark brown. Antennal segments brown, basal two thirds of segment I and basal third of segment II paler, yellowish brown. Bucculae and labium yellowish brown. Pronotum and mesoscutellum yellowish to reddish brown, in some specimens pronotum anterolaterally dark brown; metanotal elevation (‘ scutellum’) darker than pronotum, brown to dark brown. Propleura and meso- and metacoxal cavities yellowish to reddish brown, remaining portions of meso- and metathorax ventrally brown. Legs yellowish to pale brown. Clavus anteriorly milky white, in its posterior one to two thirds gradually becoming brown. Veins of corium yellowish to reddish brown, concolorous with disc of pronotum, only anteriorly infuscated with dark brown; corium between veins white. Membrane of brachypterous form completely brown (Fig. 41), in macropterous form brown with conspicuous oval whitish central spot, and two indistinct paler spots anterolaterally (Figs. 40, 42 – 43). Abdomen dark brown, pygophore slightly paler. Vestiture. Pubescence on head, pronotum, sides of thorax, mesoscutellum and metanotal elevation (‘ scutellum’) very short, in optical microscope indistinct (see Figs. 46 – 53). Clavus completely bare (Figs. 50 – 53). Vein of corium with distinct, short and fine pale setae (Figs. 46, 50). Thorax and abdomen ventrally with dense silver pubescence. Antennae and legs with long pale hairs (Figs. 48 – 49, 55 – 56). Structure. Body elongate (Figs. 40 – 43, 46 – 47), about 2.3 – 2.6 × longer than wide across humeral angles. Vertex with median sulcus not visible (Figs. 48 – 49). Bucculae low, ventrally straight, posteriorly conically produced (Fig. 55). Labium reaching anterior margin of metacoxae. Antennal segment I stoutest, cylindrical, slightly curved (Figs. 42 – 43, 48 – 49, 54 – 55); segment II clavate (Figs. 42 – 43, 54); segments III and IVa terete (Figs. 4 2 – 43, 54); segment IVb narrowly fusiform (Figs. 42 – 43, 54). Antennal segments from longest to shortest (Fig. 54): III> IVb ≥ I ≥ II ≥ IVa; segment I usually slightly longer than II (1.14 ×) and as long as compound eye (Fig. 55), rarely as long as II; segment IV fully divided into IVa and IVb (Figs. 54, 56). Pronotum (Figs. 46 – 47, 50 – 51) transverselly hexagonal, anterior collar narrow, middle transverse constriction deeper and humeral angles more arcuately produced in macropterous form (pronotum humeral width 0.69 – 0.79 mm; Figs. 46, 50) than in brachypterous form (pronotum humeral width 0.59 – 0.67; Figs. 47, 51). Metanotal elevation trapezoid; in macropterous form longer (2.8 × wider than long), posteriorly with wide and deep U-shaped incision (Fig. 52), in brachypterous form shorter (3.1 × wider than long), posteriorly widely truncate with only shallow U-shaped incision (Fig. 53); disc of metanotal elevation anteriorly depressed with more or less well developed median carina, posteriorly flat (Figs. 52 – 53), not turned upwards (Fig. 54). Metapleuron only slightly gibbose, metathorax therefore not wider than pronotum. Hemelytra of macropterous form reaching (Figs. 42 – 43, 46) or nearly reaching (Fig. 40) apex of abdomen. Hemelytra of brachypterous form reaching posterior margin of mediotergite IV (about midlength of abdomen), veins of corium reaching only midlength of mediotergite III (Figs. 41, 47). Metafemora and metatibiae of both sexes straight, without long hairs (Figs. 42 – 43, 46, 54). Mediotergites I and II with two thin, submedian, posteriad slightly diverging longitudinal keels. Male genitalia: Pygophore obovate in dorsal view (Figs. 57 – 59); proctiger short, posteriorly oval (Fig. 62), only slightly protruding from pygophore outline posteriorly (Figs. 57 – 59); paramere hook-shaped (Figs. 60 – 61); aedeagus as in Figs. 63 – 64. Measurements (mm). Brachypterous form. Male / female (n = 2 / 1). Body length: 1.51 – 1.74 / 1.66. Pronotum: anterior width: 0.34 – 0.36 / 0.36, humeral width: 0.59 – 0.67 / 0.65. Lengths of antennal segments: I: 0.14 / 0.14, II: 0.12 – 0.14 / 0.14, III: – / 0.18, IVa: – / 0.12, IVb: – / 0.14. Length of metafemur: 0.44 / 0.5, metatibia: 0.52 / 0.55, metatarsus: 0.18 / –. Macropterous form. Male / female (n = 3 / 4; median (minimum – maximum )). Body length: 1.66 (1.60 – 1.68) / 1.80 (1.78 – 1.82). Pronotum: anterior width: 0.36 (0.34 – 0.36) / 0.38 (0.36 – 0.38), humeral width: 0.71 (0.69 – 0.71) / 0.77 (0.75 – 0.79). Lengths of antennal segments: I: 0.14 (0.14 – 0.16) / 0.14 (0.14 – 0.16), II: 0.12 (0.12 – 0.14) / 0.13 (0.12 – 0.14), III: 0.20 (0.20 – 0.20) / 0.20 (0.20 – 0.22), IVa: 0.12 (0.12 – 0.12) / 0.12 (0.10 – 0.14), IVb: 0.16 (0.16 – 0.16) / 0.16 (0.16 – 0.16). Length of metafemur: 0.50 (0.46 – 0.50) / 0.48 (0.48 – 0.50), metatibia: 0.55 (0.53 – 0.57) / 0.55 (0.55 – 0.59), metatarsus: 0.16 (0.16 – 0.16) / 0.16 (0.16 – 0.20).	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFB3D209FF7CFD83A7CDE6C3.taxon	diagnosis	Systematic placement and differential diagnosis. The complete subdivision of antennal segment IV into two (IVa and IVb — see Fig. 56) as well as other characters relating it to H. ruficeps, justify the transfer of Hebrus fulvinervis to the subgenus Hebrusella as defined by Poisson (1944).	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFB3D209FF7CFD83A7CDE6C3.taxon	description	Habitat. According to the locality database of the MMBC, the specimen from Gotse Delchev was collected on the left bank of Mesta River, most probably in a muddy habitat surrounded by Salix spp. and Alnus glutinosa (sunny weather, temperature 25 – 38 ° C, 10: 30 – 15: 00) (I. Malenovský, pers. comm.).	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFB3D209FF7CFD83A7CDE6C3.taxon	distribution	Distribution (Fig. 65). Europe: Bosnia Herzegovina: Ilidža (= Ilidze) [43 ° 49 ′ 51 ″ N 18 ° 17 ′ 56 ″ E] (Lindberg 1922, as H. ruficeps; Benedek 1970), Sarajevo [43 ° 51 ′ 27 ″ N 18 ° 24 ′ 45 ″ E], Jablanica (= Jablanitza) [43 ° 39 ′ 27 ″ N 17 ° 45 ′ 38 ″ E] (Horváth 1929); Bulgaria: Gotse Delchev [41 ° 34 ′ 21 ″ N 23 ° 43 ′ 43 ″ E], Sandanski [41 ° 33 ′ 34 ″ N 23 ° 16 ′ 23 ″ E] (this paper); Romania: Sibiu county: Sibiu (= Cibinium, Nagyszeben) [45 ° 47 ′ 54 ″ N 24 ° 07 ′ 32 ″ E], Hunedoara county: Râu de Mori (= Malomvíz) [45 ° 29 ′ 48 ″ N 22 ° 51 ′ 15 ″ E], Suceava county: Crucea (= Cruce) [47 ° 21 ′ 03 ″ N 25 ° 36 ′ 29 ″ E] (Horváth 1929). New species for Bulgaria.	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFBBD202FF7CF8F2A667E0EB.taxon	distribution	Distribution (Fig. 65). North Africa: Algeria (southeast): Tassili Mts. (Poisson 1953).	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFBBD202FF7CF8F2A667E0EB.taxon	discussion	Comments. The species has not been found since its original description and the male is unknown. No type depository was mentioned in the original description by Poisson (1953). Andersen (1995) indicated the type depository as MNHN in Paris, but a subsequent search for the types was unsuccesfull (E. Guilbert, pers. comm.). All four syntypes (3 card-mounted, one slide-mounted) are deposited in the USNM in Washington: ‘ I found one slide, an apparent syntype, of H. iheriri with the head, hemelytra and wings and labelled Hebrus iheriri Tassili n. sp., two letter [can’t read them] 76, E. Bernard. We also have three carded specimens, all apparent syntypes, one of which is labelled “ Type ”. There is not locality data, but two have the number 52 and the third has a 76. ’ (T. J. Henry, pers. comm).	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFBCD200FF7CFE19A5B4E2D9.taxon	materials_examined	Material examined. BULGARIA: Sofia Province: Bajana [= Bayana], 17. vii. 1956, 1 ♀ (micropterous), L. Hoberlandt lgt., P. Kment det. (NMPC). CZECH REPUBLIC: Bohemia: Filipovka (5056), peatbog, 26. iv. 2002, 1 ♀ (micropterous), M. Fikáček lgt., P. Kment det. (NMPC); Česká Kamenice, Noldenteich NM (5152; 50 ° 48 ′ 43.3 ″ N 14 ° 26 ′ 15.0 ″ E), peatbog, 2. viii. 2004, 3 Ƌ 3 ♀ (micropterous), 23. viii. 2004, 2 ♀ (micropterous), 3. x. 2004, 2 Ƌ 1 ♀ (micropterous), P. Kment lgt. & det. (NMPC); Heřmanice (5156), flooded basalt quarry, 26. iv. 2002, 2 ♀ (micropterous), M. Fikáček lgt., P. Kment det. (NMPC); Mařeničky, Rašeliniště Mařeničky NM (5254; 50 ° 47 ′ 41.2 ″ N 14 ° 40 ′ 27.8 ″ E), peatbog, 29. vi. 2004, 2 ♀ (micropterous), 1. x. 2004, 1 Ƌ 3 ♀ (micropterous), P. Kment lgt. & det. (NMPC); Provodín, sand-pit NW of the village (5353), 8. v. 2002, 1 ♀ (micropterous), M. Fikáček & H. Poláček lgt., P. Kment det. (NMPC); Karlsbad, Zinnbach [= Lužní potok near Karlovy Vary (56 – 5739)], 9. ix. 1937, 1 ♀ (micropterous), W. Reinhard lgt., P. Kment det. (NMPC); Lomnice (5742), experimental reservoirs on recultivated spoil tips 2 km NE of the village, 20. iv. 2002, 1 ♀ (micropterous), M. Fikáček lgt., P. Kment det. (NMPC); Karlsbad, b. Flugplatz, Waldrand bei Espenthor [= Karlovy Vary, near airport, forest margin near Olšová Vrata] (5743), 1. xi. 1938, 1 Ƌ (micropterous), L. Hoberlandt det. (NMPC); Neudau [= Nejda] (5743), 24. x. 1937, 1 Ƌ 2 ♀ (micropterous), P. Kment det. (NMPC); Otovice u Karlových Varů (5743), 5 ♀ (micropterous), P. Kment det. (NMPC); Račice — Předonín (5552), 22. iv. 1976, 2 ♀ (micropterous), I. Kovář & J. Ježek lgt., P. Kment det. (NMPC); Rožďalovice (5656 – 57), 19. v. 1930, 1 Ƌ (micropterous), Dr. Klička lgt., L. Hoberlandt det. (NMPC); Kateřina, Soos NNR (5840), 27. v. 1987, 3 ♀ (micropterous), J. Vilímová lgt., P. Kment det. (NMPC); Čelákovice (5854), ix. 1946, 1 ♀ (micropterous), Mat lgt., J. L. Stehlík det. (MMBC); Sedlčánky, oxbow-lakes on left bank of Labe river (5854; 50 ° 10 ′ 05 ″ N 14 ° 47 ′ 53.1 ″ E), 27. v. 2003, 1 ♀ (micropterous), P. Kment lgt. & det. (NMPC); Přerov [nad Labem] (5854 – 55), 5. v. 1930, 1 ♀ (micropterous), Dr. Klička lgt., P. Kment det. (NMPC); Prameny, Mokřady pod Vlčkem NR (5942), vegetated pool in spring fen, 750 – 760 m a. s. l., 13. ix. 2011, 1 Ƌ 1 ♀ (micropterous), J. Škodová lgt., P. Kment det. (NMPC); [Praha], Krč (5952), 1 Ƌ 1 ♀ (micropterous), Zeman lgt., L. Hoberlandt det. (NMPC); Horní Ředice (5961 b; 50 ° 05.508 ′ N 15 ° 57.064 ′ E), NR Žernov, marsh at Mordýř pond, 20. iv. 2007, 1 ♀ (micropterous), M. Boukal lgt., P. Kment det. (NMPC); Benešov (6254), 31. vii. 1973, 1 ♀ (micropterous), Z. Jindra lgt., P. Kment det. (VURV); Lnáře (6548), Malý Pálenec [pond?], iv. 1942, Mařan lgt., L. Hoberlandt det. (NMPC); Jindřichův Hradec (6855 – 56), 18. vii. 1885, 1 ♀ (micropterous), coll. L. Duda, L. Hoberlandt det. (NMPC); Horusice, NPR Ruda (6854; 49 ° 09 ′ 03.9 ″ N 14 ° 41 ′ 27.7 ″ E), peatbog, 416 m a. s. l., 18. vii. 1970, 2 Ƌ 1 ♀ (micropterous), 18. vii. 1971, 1 ♀ (micropterous), L. Škapec lgt., P. Kment det.; 29. iv. 1975, 7 Ƌ 4 ♀ (micropterous), P. Štys lgt., J. Vilímová det.; 18. vii. 1976, 1 Ƌ 1 ♀ (micropterous), L. Škapec lgt., P. Kment det. (all NMPC); 13. ix. 2010, 2 Ƌ 5 ♀ (micropterous), extracted from wet Sphagnum, P. Kment lgt. & det. (NMPC); viii. 2011, 1 Ƌ 1 ♀ (micropterous), J. Vilímová lgt., P. Kment det. (NMPC); 29. viii. 2014, 5 Ƌ 3 ♀ (micropterous), extracted from wet Sphagnum, J. Vilímová lgt., P. Kment det. (NMPC); Ponědrážka env., W bank of Švarcenberk pond (6854; 49 ° 08.72 ′ N 14 ° 42.03 ″ E), 27. vii. 2014, 1 Ƌ (micropterous), J. Vilímová lgt., P. Kment det. (NMPC); Lomnice nad Lužnicí (6954), 16. vi. 1992, 1 ♀ (micropterous), J. Vilímová lgt., P. Kment det. (NMPC); Stožec, Mrtvý Luh (7048; 48 ° 51 ′ 34 ″ N 13 ° 53 ′ 35 ″ E), peatbog, 740 m a. s. l., extracted from wet Sphagnum, 18. viii. 2015, 1 ♀ (micropterous), P. Kment lgt. & det. (NMPC); České Budějovice, Čtyři Dvory, small peatbog near U hada street (7052; 48 ° 58 ′ 39.1 ″ N 14 ° 25 ′ 33.3 ″ E), 11. vii. 2000, 1 ♀ (micropterous), P. Kment lgt. & det. (NMPC); Nová Pec, Pěkná env. (7149; 48 ° 50 ′ 49 ″ N 13 ° 55 ′ 27 ″ E), Vltavský luh, oxbow lake of Vltava river, 18. viii. 2015, 1 ♀ (micropterous), P. Kment lgt. & det. (NMPC). Moravia: Rychlebské hory Mts., Javorník env., Uhelná (5668), 15. vii. 1981, 1 Ƌ 2 ♀ (micropterous), Z. Jindra lgt. & det. (ZJPC); Rejvíz, Rejvíz NNR (5769; 50 ° 13 ′ 15.5 ″ N 17 ° 17 ′ 58.5 ″ E), alluvial peaty meadow, 736 m a. s. l., in wet Sphagnum, 15. viii. 2011, 3 Ƌ 3 ♀ (micropterous), P. Kment lgt. & det. (NMPC); Ostrava, Martinov, Štěpán pond (6175), shore, sifting, 22. ix. 1998, 1 Ƌ (micropterous), 12. ii. 2000, 1 Ƌ (micropterous), M. Mantič lgt., P. Kment det. (MMHC); Křižánky, Kyšperský rybník pond (6362; 49 ° 41 ′ 10.1 ″ N 16 ° 03 ′ 32.8 ″ E), waterlogged shore, 24. vi. 2000, 1 Ƌ (micropterous), P. Kment lgt. & det. (NMPC); Rudice, Rudická jezírka (6666, 49 ° 20 ′ 09.7 ″ N 16 ° 43 ′ 24.4 ″ E), flooded pits after iron ore mining, 1. v. 1999, 2 Ƌ 1 ♀ (micropterous), 6. vi. 1999, 2 Ƌ 2 ♀ (micropterous), 6. vii. 1999, 2 ♀ (micropterous), P. Kment lgt. & det. (NMPC, MHBC, MUBC); Velká Bíteš, Dolní Klečanský rybník pond (6763; 49 ° 16 ′ 39 ″ N 16 ° 14 ′ 28 ″ E), waterlogged grassy shore, 1. v. 2000, 3 Ƌ 10 ♀ (micropterous), P. Kment lgt. & det. (NMPC, MUBC); Střelice (6865), ix. 1943, 1 ♀ (micropterous), Rauscher lgt., L. Hoberlandt det. (MMBC); Pouzdřany (7065), 7. vi. 1967, 1 ♀ (macropterous), Volák lgt., P. Kment det. (SMOC); Šafov env., Janský rybník pond (7160), litter on banks (sifting) + in water (high water level), 10. iv. 2007, 5 ♀ (micropterous), M. Fikáček lgt., P. Kment det. (NMPC); Hrabětice, Travní dvůr SAC, Za drahou pool (7164; 48 ° 48 ′ 55.9 ″ N 16 ° 27 ′ 8.8 ″ E), 15. v. 2011, 1 Ƌ (micropterous), J. Sychra lgt., P. Kment det. (NMPC); Lednice, Nejdek (7166), Nejdecké louky meadows, Bažina u Azontu reed marsh, 18. ix. 1998, 2 ♀ (micropterous), P. Kment & M. Horsák lgt., P. Kment det. (NMPC); Podivín, pools by road to Lednice (7167; 48 ° 49 ′ 24 ″ N 16 ° 50 ′ 41 ″ E), 26. iv. 2003, 2 Ƌ 1 ♀ (micropterous), M. Horsák lgt., P. Kment det. (NMPC); Valtice, Alach IV (7266), small silting pond in forest, 9. v. 1998, 1 ♀ (macropterous), P. Kment lgt. & det. (NMPC). POLAND: Lesser Poland Voivodeship: Chochołów, sepage on south margin of village, 14. vi. 2002, 1 ♀ (micropterous), M. Horsák lgt., P. Kment det. (NMPC). ROMANIA: Caraș-Severin county: Caransebeș, litter on pond banks (45.4188 ° N 22.1875 ° E), 217 m a. s. l., 24. xii. 2014, 1 ♀ (micropterous), G. M. Berchi lgt. & det. (GBCR). Cluj county: Turda, litter on pond banks (46.47388 ° N 23.77694 ° E), 317 m a. s. l., 22. xii. 2014, 1 ♀ (macropterous) 2 Ƌ (micropterous), G. M. Berchi lgt. & det (GBCR). Timiș county: Dumbrăvița, litter on pond banks (45.80638 ° N 21.26416 ° E), 96 m a. s. l., 19. xii. 2014, 1 Ƌ 1 ♀ (micropterous), I. A. Rădac leg., G. M. Berchi det. (GBCR); Jimbolia, litter on pond banks (45.8099 ° N 20.7177 ° E), 75 m a. s. l., 26. xii. 2014, 2 ♀ (micropterous), I. A. Rădac leg., G. M. Berchi det. (GBCR); Satchinez, collected by pitfall trap in farmland (45.9622 ° N 21.0466 ° E), 92 m a. s. l., 10. vii. 2012, 1 ♀ (micropterous), I. A. Rădac leg., G. M. Berchi det. (GBCR). RUSSIA: North European Territory: Leningrad Province: Karelian Isthmus, St. Petersburg reg., Svetloe ozero Lake (ca. 60 ° 18 ′ N 29 ° 50 ′ E), surrounding peat bog, 21. vii. 2002, 2 ♀ (micropterous), P. Kment lgt. & det. (NMPC); Karelian Isthmus, 4 km S of Dibuny railway station, river bank, 13. vi. 2004, 1 ♀ (macropterous) 9 Ƌ 5 ♀ (micropterous), P. Kment lgt. & det. (NMPC). SLOVAKIA: Svätý Jur, Šúr NNR (7768), 4. vi. 1941, 1 Ƌ 3 ♀ (micropterous), L. Korbel lgt., P. Kment det. (MMBC); Palárikovo (7974), salty clay-pit, in water, 1. v. 1999, 1 ♀ (micropterous), M. Mantič lgt., P. Kment det. (MMHC); Tvrdošovce (7974), meadow behind railway, canal, 1. v. 1999, 1 ♀ (micropterous), M. Mantič lgt., P. Kment det. (MMHC).	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFBCD200FF7CFE19A5B4E2D9.taxon	distribution	Distribution (Fig. 73). Europe: Albania (Josifov 1970), Austria (Benedek 1970, Rabitsch 2005), Belgium (Stoffelen et al. 2013), Bulgaria (Josifov 1964), Byelorussia (Lukashuk 1997, Kanyukova 2006), Croatia (Protić 1998, Kment & Beran 2011), Czech Republic (Hoberlandt 1977), Denmark (Andersen & Gaun 1974), Estonia (Coulianos 2005), Finland (Rintala & Rinne 2010), France (Poisson 1957 a), Germany (Hoffmann & Melber 2003, Wachmann et al. 2006, Strauss & Niedringhaus 2014), Great Britain (Savage 1989), Hungary (Horváth 1897, Benedek 1970, Boda et al. 2015), Ireland (Savage 1989), Italy (Servadei 1967, Tamanini 1979, Cianferoni & Mazza 2012), Latvia (Lukashuk 1997, Spuņģis 2008), Luxembourg (Reichling & Gerend 1994), Macedonia (Protić 1998), Moldavia (Derzhansky 1984, 1997; Kanyukova 2006), the Netherlands (Aukema et al. 2002), Norway (Coulianos 1998, Coulianos et al. 2008), Poland (Jaczewski & Wróblewski 1976, Wróblewski 1980), Romania (Horváth 1897, Benedek 1970), Russia: North, Central and South European Territory of Russia (Kanyukova 1997, 2006), Serbia (Horváth 1897, Benedek 1970, Protić 1998), Slovakia (Horváth 1897, Benedek 1970, Kment et al. 2003, Reduciendo Klementová et al. 2015), Slovenia (Gogala 2003), Sweden (Coulianos & Ossiannilsson 1976), Switzerland (Dethier & Matthey 1977, Matthey & Fiora 1979), Ukraine (Putshkov & Putshkov 1996; Kanyukova 1997, 2006; Grandova 2013). Asia: Iran: Mazandaran (Ghahari et al. 2013), Kazakhstan: Asian part (Andersen 1995; Kanyukova 1997, 2006; Esenbekova et al. 2015), Russia: East Siberia, Far East (Kanyukova 1997, 2006; Vinokurov et al. 2010), Turkey: Asian part (Fent et al. 2011), Uzbekistan (Kanyukova 1997, 2006). The species was listed from Bosnia Herzegovina by Andersen (1995), probably based on Lindberg’s (1922) record from the locality Ilidze, which probably is a misidentification of H. fulvinervis recorded from the same locality by Benedek (1970).	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFBED200FF7CFB78A11BE71C.taxon	description	Hebrus acutiscutata [sic] Brown, 1951: 239. Holotype: ♀, Saudi Arabia: Al Birza near Jeddah (BMNH).	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFBED200FF7CFB78A11BE71C.taxon	materials_examined	Material examined. ISRAEL: E. Gedi [= Ein Gedi], 13 / 3. A, 25. x. 1980, 1 Ƌ, D. Furth lgt., P. Kment det. (NMPC). YEMEN: Al Hudaydah Province, Jabal Bura valley forest NP (stream valley), 14 ° 52.4 – 5 ′ N 43 ° 24.6 – 25.2 ′ E, 240 – 350 m a. s. l., at light, 4. xi. 2010, 2 Ƌ 1 ♀, J. Hájek lgt., P. Kment det. (NMPC).	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFBED200FF7CFB78A11BE71C.taxon	distribution	Distribution (Fig. 72). Asia: Israel (this paper), Saudia Arabia: Hijaz (Brown 1951, Linnavuori 1980), Yemen (Linnavuori & van Harten 1997, 2002). Afrotropical Region: Sudan (Poisson 1964, as H. soudani; Linnavuori 1971, 1981). New species for Israel.	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFBED200FF7CFB78A11BE71C.taxon	discussion	Comment. Linnavuori (1981) listed this species in the subgenus Hebrus s. str. and illustrated the scutellum.	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFBED201FF7CF901A0E0E0EA.taxon	distribution	Distribution (Fig. 65). Italy: Emilia-Romagna (Cianferoni et al. 2016); Tuscany (Wagner 1957; Servadei 1958, 1967, 1969; Cianferoni et al. 2016).	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFBED201FF7CF901A0E0E0EA.taxon	discussion	Comment. Tamanini (1979) expressed his doubts about the validity of this species. However, the identity of this exceptional apterous species was confirmed recently based on newly collected material (Cianferoni 2015, Cianferoni et al. 2016). Subgeneric placement. Cianferoni et al. (2016) mentioned that the antenna of H. franzi is tetramerous, without distinct membranous subdivision of antennal segment IV. This would exclude H. franzi from the subgenera Hebrusella as well Hebrus s. str. as defined by Poisson (1944). They commented that subgeneric division needs further study on a world-wide scale and phylogenetic context, and did not consider the subgeneric assignment of H. franzi. We prefer to list this taxon as Hebrus incertae sedis without subgeneric placement.	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFBFD201FF7CFD9FA55DE4C4.taxon	materials_examined	Material examined. CANARY ISLANDS: Gran Canaria: Maspalomas, 282 - 120, sand dunes, water, 3. viii. 2003, 3 ♀, M. Mantič lgt. (2 ♀ NMPC, 1 ♀ MMHC).	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFBFD201FF7CFD9FA55DE4C4.taxon	distribution	Distribution. North Africa: Canary Islands (Poisson 1954 a, b; Aukema et al. 2013 b). Nearctic and Neotropical Region: From southern Canada to Argentina (except Chile), West Indies and Netherland Antilles (Drake 1917, Blatchley 1926, Drake & Chapman 1958, Drake & Cobben 1960, Polhemus & Polhemus 1988). Hawaiian Islands (White 1877). The distribution in Cape Verde Islands (Drake & Chapman 1958) is an apparent error for Canary Islands, which were not listed by those authors.	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
03BD2655FFBFD201FF7CFD9FA55DE4C4.taxon	discussion	Comment. The synonym Merragata lindbergi was described according to 2 male and 4 female syntypes deposited in MZHF. However, in the collection of NMPC there is deposited also one female ‘ paratype’ of M. lindbergi (see Kment & Kolínová 2013). Most probably there are the same problems with redundant types as in H. pusillus canariensis (see Comment on p. 15) but we did not receive the detailed information from L. Huldén to be provided here.	en	Kment, Petr, Jindra, Zdeněk, Berchi, Gavril Marius (2016): Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera). Zootaxa 4147 (3): 201-239, DOI: 10.11646/zootaxa.4147.3.1
