identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
B6C835D2254F643DCE4E65D17CC823AB.text	B6C835D2254F643DCE4E65D17CC823AB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amastigogonus Broelemann 1913	<div><p>Amastigogonus Broelemann, 1913</p><p>Amastigogonus Brölemann 1913: 152. Verhoeff 1924: 75, 84; 1932: 1732, 1737; 1944: 36, 41. Jeekel 1971: 107; 1981: 39; 2009: 35. Hoffman 1972: 204; 1980: 91. Mauriès 1987: 198. Mauriès et al. 2001: 585. Korsós and Johns 2009: 3. Edward and Harvey 2010: 5. Korsós and Read 2012: 44.</p><p>Euethogonus Chamberlin 1920: 166. Hoffman 1972: 204 (synonymised with Amastigogonus); 1980: 91.</p><p>Type species.</p><p>Amastigogonus tasmanianus Brölemann, 1913, by original designation.</p><p>Other assigned species.</p><p>Amastigogonus danpicola sp. n., Amastigogonus elephas sp. n., Amastigogonus fossuliger Verhoeff, 1944, Amastigogonus hardyi (Chamberlin, 1920), Amastigogonus hellyeri sp. n., Amastigogonus michaelsae sp. n., Amastigogonus orientalis sp. n., Amastigogonus peninsulensis sp. n., Amastigogonus verreauxii (Gervais, 1847).</p><p>Diagnosis.</p><p>Like Victoriocambala Verhoeff, 1944 in having the coxite process on the anterior gonopod close to the telopodite and nearly as long, forming a chamber resembling a bird’s beak in which the pseudoflagellum is protected. Differences between Amastigogonus / Victoriocambala, as noted by Jeekel (2009: 35), are leg 1 with free/fused tibia and tarsus, posterior gonopod without/with reduced telopodite.</p><p>Description.</p><p>Living animals usually with black or dark grey rings with annular pale band at rear of each metazonite, often with a greenish tinge (live Amastigogonus fossuliger more consistently green, see species description); head, collum and telson often faintly reddish brown; legs pale. With long storage in alcohol and staining by defensive secretion, animals dull grey with faintly reddish legs.</p><p>Observed midbody diameter of larger males 2.5-4.2 mm, 55-71 podous rings. Head smooth, slightly convex, vertigial sulcus reaching to level of dorsalmost ocellar row. Ocellar area of larger males lenticular with ca 25-50 ocelli in 4-6 somewhat irregular horizontal rows. Antennae short, barely reaching past posterior edge of collum when manipulated dorsally; relative antennomere lengths (2=3)&gt;6&gt;(4=5); antennomere 6 widest; 4 apical cones; socket ca 1 socket diameter from lateral margin of head capsule. Gnathochilarium with lateral edges of mentum slightly convex, mentum about as wide as combined lingual plates; mentum-promentum junction slightly concave anteriorly; a prominent pit with small seta anteriorly on each gnathochilarial stipes. Collum convex, laterally narrowing with rounded corner, margins straight. Prozonites only slightly narrower than metazonites; suture weakly defined; fine longitudinal striae on lower portion of metazonite, anterior end of each stria (Fig. 3C; s) bent obliquely upwards towards suture (see Remarks, below); prozonites and metazonites with surface otherwise smooth, free of setae. Ozopores (Fig. 3C; o) small, round, beginning ring 6 at a little over 1/2 ring height, ring 6 ozopore distinctly lower than ring 7 ozopore; each ozopore at ca 1/3 the distance from suture to posterior metazonite margin, and usually absent from apodous rings. Limbus lamellar, undivided. Preanal ring smooth, epiproct broadly rounded, extending slightly over anal valves; hypoproct with slightly convex margin.</p><p>Legpair 1 separate on coxosternite, each leg 1 with 5 podomeres without setae, anteroposteriorly somewhat flattened; relative podomere lengths typically femur&gt;tibia&gt;(prefemur=postfemur)&gt;tarsus, relative widths typically prefemur&gt; femur&gt;tibia&gt;postfemur&gt;tarsus (see Remarks, below); no claw. Small brushes of setae on legpair 1 coxosternite anterior to and between legs, and laterally on coxosternite corners. Leg 2 leg-like with large claw and reduced prefemur; penis arising basally on posterior coxal surface, barrel-shaped with a few long setae in apical crown. Leg 7 (and sometimes other legs near gonopod aperture; see Remarks, below) with elongated coxa (Fig. 2). Midbody legs short, ca 2/3 ring diameter when extended; relative podomere lengths prefemur&gt;(femur=tarsus)&gt;(postfemur=tibia). Most prefemora distally with conical prefemoral pad (Fig. 5A, B; pa); pads first appear on ring 8 legs and diminish in size posteriorly; pads small or absent on last 2-4 legpairs (see Remarks, below).</p><p>Posterior margin of gonopod aperture raised and thickened on either side, adjoining tips of retracted anterior gonopods. Anterior gonopods (Figs 3A, 3B, 3D, 3E, 6, 8) parallel, closely appressed. Anterior gonopod coxite (Figs 3A, B, D, E; cx) massive, rounded laterally, extending thin, finger-shaped process (Figs 3A, 3B, 3D, 3E, 6, 8; cxp) from anteromedial surface, process slightly concave laterally, thickened medially to ca 2/3 process height (coxite process more complex in Amastigogonus danpicola sp. n., see species description). Telopodite (Figs 3A, 3B, 3D, 3E, 6, 8; te) arising from wide, shallow recess on coxite, paralleling coxite process and slightly longer; thin and slightly concave medially; somewhat thickened distally from near posterior margin to midline near apex, the thickening usually with row of more or less evenly spaced, prominent setae on posterior side and sometimes a separate row or group of setae on anterior side. Posterior surface of telopodite produced basally as rounded flange. Pseudoflagellum (Figs 3C, 6, 8; ps) thinly lamellar, usually arising at ca 1/2 telopodite height, usually supported by setae on distomedial surface of telopodite. Prostatic groove (Figs 3C, 6, 8; pg) running anterodistally from posterobasal corner of telopodite under rounded flange to pseudoflagellum, following anterior side of pseudoflagellum and terminating at pseudoflagellum tip. Posterior gonopod (Fig. 3F) ca 1/3 height of anterior gonopod, subcylindrical with apical recess posterolaterally, a crown of short setae around recess and an arm-like cylindrical process with rounded apex arising at ca 1/2 gonopod height on posterolateral surface and directed distally. In situ, posterior gonopod grips base of anterior gonopod telopodite between arm-like process and body of posterior gonopod, the posterior gonopod apex pressed against prostatic groove: "The rudimentary posterior gonopods (Fig. 3H) appear to be attached like forceps to the wall at the very base of the caudomedian ridges of the anterior gonopods (Fig. 3E)" ( Mauriès, Golovatch and Hoffman 2001: 585).</p><p>Female slightly larger in diameter than male with same ring count; leg 1 normally leg-like, claw-bearing; no prefemoral pads on any legs.</p><p>Remarks.</p><p>Amastigogonus species are closely similar in size, general appearance and habits, and males can only be positively identified by close inspection, and usually dissection, of the anterior gonopods. There is also some variation in non-gonopodal male structures, as noted here and in the species descriptions. The most reliable of these differences are in cardo shape and modifications of near-aperture legs:</p><p>Cardo. In Amastigogonus danpicola sp. n. the cardo extends further ventrally in its posterior half than in its anterior half, i.e. the cardo is deeper posteriorly (Fig. 4A). In other Amastigogonus species the ventral edge of the cardo is either evenly convex or is deeper anteriorly (Fig. 4B).</p><p>Near-aperture legs. In all Amastigogonus species, leg 7 has an elongated coxa (Fig. 2A, B; arrow). Less elongated coxae are also found on legs 10 and 11 in Amastigogonus hellyeri sp. n. and on legs 6, 10 and 11 in Amastigogonus danpicola sp. n. (Fig. 2C).</p><p>Other differences between species do not seem to be large enough or consistent enough to be useful for taxonomic purposes:</p><p>Legpair 1. The relative lengths and widths of the leg 1 podomeres vary a little between species (Fig. 4 C–G), between individuals, and sometimes between right and left legs.</p><p>Prefemoral pads. There are differences between species in pad length, as shown in Fig. 5A and 5B (pa), but these differences are masked by the anteroposterior size gradient on single individuals as well as by variability from individual to individual.</p><p>Metazonal striae. The height reached by the horizontal striae on the metazonites diminishes slightly from anterior to posterior. At ca 2/3 body length, the topmost horizontal stria lies at ca 1/2 or ca 3/4 of the height to the ozopore (Fig. 5C; st), depending on species, but with considerable variation between rings and between individuals.</p></div>	https://treatment.plazi.org/id/B6C835D2254F643DCE4E65D17CC823AB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Mesibov, Robert	Mesibov, Robert (2017): Iulomorphid millipedes (Diplopoda, Spirostreptida, Iulomorphidae) of Tasmania, Australia. ZooKeys 652: 1-36, DOI: http://dx.doi.org/10.3897/zookeys.652.12035, URL: http://dx.doi.org/10.3897/zookeys.652.12035
FD0A76D1C223E554498C39F0734B3D50.text	FD0A76D1C223E554498C39F0734B3D50.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amastigogonus danpicola	<div><p>Amastigogonus danpicola sp. n. Figs 2C, 4A, 4C, 6A</p><p>Holotype.</p><p>Male, Apsley River, Tas, -41.7992 148.1508 ± 250 m [label "EP 955 717" (= 55G 595500 5371700, AGD66)], 300 m a.s.l., 5 July 1988, R. Mesibov, QVM 23:54377 (specimen in 3 pieces).</p><p>Paratypes.</p><p>1 male, locality and collector as for holotype but -41.7972 148.1544 ± 250 m [label "EP 958 719" (= 55G 595800 5371900, AGD66)], 320 m a.s.l., 6 July 1988, R. Mesibov, QVM 23:54371; 1 male, same locality and collector but -41.7964 48.1592 ± 250 m [label "EP 962 720" (= 55G 596200 5372000, AGD66)], 350 m a.s.l., 18 July 1988, R. Mesibov, QVM 23:54373.</p><p>Other material.</p><p>22 males, 4 probable females from 16 unique localities; details in Suppl. material 1.</p><p>Diagnosis.</p><p>Coxite process of the anterior gonopod divided by narrow fossae apically and anterobasally; legs 6, 7, 10 and 11 with elongated coxae.</p><p>Description.</p><p>Mature males observed with (48+4) rings, 2.6 mm midbody diameter to (71+1) rings, 3.2 mm. Cardo deeper posteriorly than anteriorly (Fig. 4A; ce). Legs 6, 7, 10 and 11 with elongated coxae, swollen distomedially (Fig. 2C); leg 7 coxa longest and most swollen. Prefemoral pad ca 3/4 femur length. Striae on posterior metazonites reaching ca 1/2 ozopore height.</p><p>Coxite process on anterior gonopod (Fig. 6A; cxp) with basal portion divided anteriorly by fossa (Fig. 6A; fo) into lateral and medial longitudinal flanges, the medial flange with deep, V-shaped notch at ca 1/2 coxite height on anterior margin; process divided apically by shallow fossa (Fig. 6A; fo) separating distolateral and distomedial margins; process not reaching level of telopodite apex. Telopodite (Fig. 6A; te) with row of minute setae on posterior side of medial thickening, behind pseudoflagellum (Fig. 6A; ps). Pseudoflagellum ca 1/2 telopodite width at base, truncate at ca 1/2 pseudoflagellum height, extending anterodistally from anterior corner of truncate basal portion as thin, flexible ribbon, the latter usually curving laterally over telopodite apex in preserved specimens.</p><p>Distribution.</p><p>Eucalypt forest over ca 1000 km2 on the East Coast of Tasmania, mainly in the Apsley, Douglas, St Pauls and Swan River catchments (Fig. 7C), from near sea level to at least 600 m. Possibly parapatric with Amastigogonus elephas sp. n. in the upper St Pauls River catchment and with Amastigogonus michaelsae sp. n. near Swansea. Sympatric with Amastigogonus fossuliger and with Amastigogonus orientalis sp. n.</p><p>Name.</p><p>Abbreviation in lower case “danp” for Douglas-Apsley National Park plus Latin cola, inhabitant; noun in apposition. This species is abundant in the Park, which also contains the type locality.</p><p>Remarks.</p><p>Amastigogonus danpicola sp. n. is the most apomorphic species within the group included here in Amastigogonus . I place it in this genus because the structure and position of the pseudoflagellum and its supporting setae conform to the general pattern seen in the other Amastigogonus species.</p></div>	https://treatment.plazi.org/id/FD0A76D1C223E554498C39F0734B3D50	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Mesibov, Robert	Mesibov, Robert (2017): Iulomorphid millipedes (Diplopoda, Spirostreptida, Iulomorphidae) of Tasmania, Australia. ZooKeys 652: 1-36, DOI: http://dx.doi.org/10.3897/zookeys.652.12035, URL: http://dx.doi.org/10.3897/zookeys.652.12035
747D260FDBEC16707B062ABB52811B8E.text	747D260FDBEC16707B062ABB52811B8E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amastigogonus elephas	<div><p>Amastigogonus elephas sp. n. Fig. 6B</p><p>Holotype .</p><p>Male, Mt Elephant, Tas, -41.6338 148.2421 ± 25 m, 420 m a.s.l., 13 May 2016, R. Mesibov, QVM 23:54519.</p><p>Paratypes.</p><p>3 males, 3 females, details as for holotype, QVM 23:54520; 1 male, same locality but -41.6244 148.2425 ± 250 m [label "FP 034 910" (= 55G 603400 5391000, AGD66)], 19 January 2002, R. Mesibov and T. Moule, QVM 23:54369.</p><p>Other material.</p><p>6 males from 5 unique localities; details in Suppl. material 1.</p><p>Diagnosis.</p><p>Like Amastigogonus verreauxii in having a telopodite with a subquadrate extension of the posterobasal margin and a posteriorly curving pseudoflagellum; distinguished from Amastigogonus verreauxii in the pseudoflagellum having a small, tooth-like, distally directed extension basal to the posterobasally directed tip.</p><p>Description.</p><p>Mature males observed with (47+3) rings, 3.2 mm midbody diameter to (69+0) rings, 3.6 mm. Cardo not deeper posteriorly. Leg 7 (only) with elongated coxa. Prefemoral pad ca 1/2 femur length. Striae on posterior metazonites reaching ca 3/4 of ozopore height.</p><p>Coxite process on anterior gonopod (Fig. 6B) with small portion of posterodistal margin extended as rounded tab (Fig. 6B; t), sharply folded over laterally; process nearly reaching level of telopodite apex. Telopodite with posterobasal surface produced posteriorly as subquadrate flange (Fig. 6B; fl) and with single row of prominent setae on posterior side of medial thickening. Pseudoflagellum with small rounded swelling adjoining base posteromedially; arising at 1/3-1/2 telopodite height, ca 2/3 width of telopodite at base, slightly curving posteriorly, slightly expanded posteriorly at 1/3-1/2 pseudoflagellum height, apically with small, rounded, distally directed, tooth-like extension, the prostatic groove opening on posterobasally directed pseudoflagellum tip.</p><p>Distribution.</p><p>Eucalypt forest at the eastern end of the Fingal Valley on the Tasmanian East Coast (Fig. 7C), to at least 700 m elevation. The small range of this species is home to other locally endemic invertebrates, including the millipede Tasmaniosoma nicolaus Mesibov, 2015 and the onychophoran Tasmanipatus anophthalmus Ruhberg, Mesibov, Briscoe &amp; Tait, 1991. Amastigogonus elephas sp. n. may be parapatric with Amastigogonus danpicola sp. n. in the upper St Pauls River catchment.</p><p>Name.</p><p>Latin elephas, elephant; noun in apposition. For the type locality, Mt Elephant.</p><p>Remarks.</p><p>The anterior gonopod of Amastigogonus elephas sp. n. (Fig. 6B) is similar to that of Amastigogonus verreauxii (Fig. 8D). The wide disjunction in the species ranges (more than 100 km) and the remarkable consistency of form in the anterior gonopod of Amastigogonus verreauxii (across a linear range extent of ca 400 km) suggests that the two lineages have long been separated.</p></div>	https://treatment.plazi.org/id/747D260FDBEC16707B062ABB52811B8E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Mesibov, Robert	Mesibov, Robert (2017): Iulomorphid millipedes (Diplopoda, Spirostreptida, Iulomorphidae) of Tasmania, Australia. ZooKeys 652: 1-36, DOI: http://dx.doi.org/10.3897/zookeys.652.12035, URL: http://dx.doi.org/10.3897/zookeys.652.12035
07EFBAA26E03F7DD2E7FBE3719CE74D1.text	07EFBAA26E03F7DD2E7FBE3719CE74D1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amastigogonus fossuliger Verhoeff 1944	<div><p>Amastigogonus fossuliger Verhoeff, 1944 Figs 1, 2A, B, 3C, 4B, 5B, 6C</p><p>Amastigogonus fossuliger Verhoeff 1944: 43, figs 6-8. Jeekel 1981: 43. Korsós and Read 2012: 45.</p><p>Syntypes.</p><p>At least 1 male and 1 female, Lake Leake, Tasmania, date and collector unknown (see Remarks, below), specimens not located.</p><p>Other material.</p><p>130 males and 12 females from 66 unique localities; details in Suppl. material 1.</p><p>Diagnosis.</p><p>Coxite process of anterior gonopod with posterodistal margin not extended; pseudoflagellum with dense field of short setae on telopodite behind pseudoflagellum tip; pseudoflagellum with distinct shoulder, the prostatic groove making an S-bend before entering the abruptly tapered tip of the pseudoflagellum.</p><p>Description.</p><p>Live males and females with more or less greenish-grey rings (Fig. 1A, C). Mature males observed with (39+4) rings, 2.1 mm midbody diameter to (63+1) rings, 3.6 mm. Cardo not deeper posteriorly (Fig. 4B). Leg 7 (only) with elongated coxa (Fig. 2A, C). Prefemoral pad ca 3/4 or more femur length (Fig. 5B). Striae on posterior metazonites reaching ca 2/3-3/4 of ozopore height.</p><p>Coxite process on anterior gonopod (Fig. 6C) with posterodistal margin not extended and folded over laterally. Telopodite with single row of prominent setae on posterior side of medial thickening to near telopodite apex, the thickening then widening and bearing dense brush of numerous minute setae. Pseudoflagellum ca 1/2 width of telopodite at base, tapering abruptly at ca 1/2 pseudoflagellum height to narrow, sharply pointed tip with rounded shoulder on anterior side of tapered section (Fig. 3C; ps), sometimes with small, tooth-like, anterodistal extension on shoulder; prostatic groove (Fig. 3C; pg) making S-bend from anterior side of pseudoflagellum into tapered tip.</p><p>Distribution.</p><p>Widespread in the eastern half of Tasmania (Fig. 7B) in dry and wet eucalypt forest from near sea level to at least 1050 m, extending across the Central Plateau to the Cradle Mountain area. Parapatric with Amastigogonus hellyeri sp. n. along the Mersey Break, a well-documented faunal divide for millipedes in north central Tasmania (Mesibov 1999). Co-occurs with Amastigogonus tasmanianus in northeast Tasmania. Overlaps with Amastigogonus verreauxii on the Central Plateau and possibly in southern Tasmania, and to a small extent with Amastigogonus danpicola sp. n. southeast of the Fingal Valley. The far southern record on the distribution map (Fig. 7B) is for two Amastigogonus fossuliger males found on the verandah of a house at Francistown, and may represent an accidental translocation rather than a natural occurrence.</p><p>Remarks .</p><p>At least one male and one female of this species from the Lake Leake area were probably sent to Verhoeff by George Edward Nicholls, a Western Australian biologist who collected in Tasmania in 1928, 1929 and 1939 (Nicholls 1943). Verhoeff (1936: 14) had previously thanked Nicholls for providing specimens of an unrelated millipede species from Lake Leake.</p><p>I have trouble understanding the differences in the two anterior gonopods illustrated by Verhoeff (1944), both presumably from Lake Leake specimens and possibly from the same male. Verhoeff’s fig. 6 shows a right gonopod tip in posterior view, fig. 7 a left gonopod tip in medial view. The thread-like pseudoflagellum in fig. 6 has a tooth-like extension on the shoulder and an intact posterobasal margin, while the shorter, tapered pseudoflagellum in fig. 7 has no tooth-like extension and a notched posterobasal margin. The remarkable thinness and fragility of the Amastigogonus fossuliger pseudoflagellum may be the explanation for Verhoeff’s difficulties in seeing and drawing these features. My Fig. 6C is based on a male from near the type locality, while the image in Fig. 3C is of a male from Ansons Bay, ca 100 km to the north. The only significant difference is the absence of a tooth-like extension in the latter.</p><p>Amastigogonus fossuliger is more consistently and more obviously green than other Amastigogonus species, but the green colour varies in intensity from individual to individual.</p></div>	https://treatment.plazi.org/id/07EFBAA26E03F7DD2E7FBE3719CE74D1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Mesibov, Robert	Mesibov, Robert (2017): Iulomorphid millipedes (Diplopoda, Spirostreptida, Iulomorphidae) of Tasmania, Australia. ZooKeys 652: 1-36, DOI: http://dx.doi.org/10.3897/zookeys.652.12035, URL: http://dx.doi.org/10.3897/zookeys.652.12035
A6E76A6F229B4014CE94870A09D1F29D.text	A6E76A6F229B4014CE94870A09D1F29D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amastigogonus hardyi (Chamberlin 1920) Chamberlin 1920	<div><p>Amastigogonus hardyi (Chamberlin, 1920) Figs 3A, B, D, E, 5C, 6D</p><p>Euethogonus hardyi Chamberlin 1920: 166. Jeekel 1971: 109.</p><p>Amastigogonus hardyi Hoffman 1972: 204 (new combination), figs 6-8. Jeekel 1981: 43. Korsós and Read 2012: 45.</p><p>Amastigogonus nichollsii Verhoeff 1944: 43, figs 1-5. Hoffman 1972: 204 (synonymised with Amastigogonus hardyi).</p><p>Holotype of Euethogonus hardyi .</p><p>Male, Tasmania, date unknown, G.H. Hardy, MCZ 4817. Illustrated by Hoffman (1972); not examined.</p><p>Paratypes of Euethogonus hardyi .</p><p>At least 1 female, details as for holotype, MCZ 4818. Not examined.</p><p>Lectotype of Amastigogonus nichollsii</p><p>(here designated). Male (slide mount of gonopods), Mt Nelson, Tasmania, date and collector unknown, ZMB 12642a (= Verhoeff collection slide 3777). Illustrated by Verhoeff (1944); material examined as image.</p><p>Paralectotypes of Amastigogonus nichollsii .</p><p>1 female, same collection details, ZMB 12642; not examined. Verhoeff (1944: 44) describes a second male specimen, but this has not been located; this male is another paralectotype.</p><p>Other material.</p><p>3 males and 3 females from 2 unique localities; details in Suppl. material 1.</p><p>Diagnosis.</p><p>Like Amastigogonus fossuliger in having the pseudoflagellum with a narrow, tapering, pointed tip; distinguished by the prostatic groove following a straight course on the pseudoflagellum rather than having an S-bend, and by the lack of a dense field of short setae on the telopodite behind the pseudoflagellum tip.</p><p>Description.</p><p>Three mature males examined: (52+3) rings, 2.8 mm midbody diameter, (61+1) rings, 3.1 mm and (65+1) rings, 3.2 mm. Cardo not deeper posteriorly. Leg 7 (only) with elongated coxa. Prefemoral pad ca 1/2 femur length. Striae on posterior metazonites reaching ca 1/2 of ozopore height (Fig. 5C).</p><p>Coxite process on anterior gonopod (Fig. 6D) with small portion of posterodistal margin extended as rounded tab and folded over laterally. Telopodite with single row of prominent setae on posterior side of medial thickening and short row on anterior side near apex. Pseudoflagellum ca 1/2 telopodite width at base, slightly extended posteriorly at base, then gradually tapering to sharp, posterodistally directed point.</p><p>Distribution.</p><p>Currently known only from eucalypt forest at two sites in the Hobart metropolitan area (Fig. 7B) in Tasmania: Mt Nelson (including 1973 collections at this type locality for Amastigogonus nichollsii) and Mt Rumney.</p><p>Remarks.</p><p>The types of Euethogonus hardyi were collected by the entomologist George H. H. Hardy, probably during Hardy’s tenure as Assistant Curator of the Tasmanian Museum in Hobart, 1913-1917 (Marks 1991: 216). Hoffman (1972) examined and illustrated the Euethogonus hardyi holotype and assigned the species to Amastigogonus . The locality given for the types is simply “Tasmania” (Hoffman 1972: 204). Chamberlin (1920: 166-167) did not specify the number or gender of the Euethogonus hardyi paratypes.</p><p>Two males and one female of Amastigogonus nichollsii were presumably sent to Verhoeff by G.E. Nicholls, collector of the Amastigogonus fossuliger types. Hoffman (1972) synonymised Amastigogonus nichollsii with Amastigogonus hardyi after comparing Verhoeff’s drawings of slide-mounted nichollsii gonopods with the unmounted gonopods of the hardyi holotype. After examining males from the nichollsii type locality, Mt Nelson (TMAG J5897 and J5926), I agree with Hoffman that nichollsii is a junior synonym of hardyi .</p></div>	https://treatment.plazi.org/id/A6E76A6F229B4014CE94870A09D1F29D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Mesibov, Robert	Mesibov, Robert (2017): Iulomorphid millipedes (Diplopoda, Spirostreptida, Iulomorphidae) of Tasmania, Australia. ZooKeys 652: 1-36, DOI: http://dx.doi.org/10.3897/zookeys.652.12035, URL: http://dx.doi.org/10.3897/zookeys.652.12035
6F2053EE20523E53F5C757BE75F7D303.text	6F2053EE20523E53F5C757BE75F7D303.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amastigogonus hellyeri	<div><p>Amastigogonus hellyeri sp. n. Figs 4D, 6E</p><p>Holotype.</p><p>Male, Keddies Creek area, Tas, -41.1704 146.0545 ± 25 m, 60 m a.s.l., 7 May 2016, R. Mesibov, QVM 23:54471.</p><p>Paratypes.</p><p>1 male, details as for holotype, QVM 23:54472; 2 males, same locality and collector but -41.1672 146.0569 ± 250 m, 50-120 m a.s.l., 11 May 2016, tree trunks along Dial Road at night, QVM 23:54515.</p><p>Other material.</p><p>142 males and 10 females from 26 unique localities; details in Suppl. material 1.</p><p>Diagnosis.</p><p>Like Amastigogonus orientalis sp. n. in having a broad pseudoflagellum abruptly reduced apically; distinguished by having a smoothly curving rather than a subquadrate extension of the posterobasal telopodite margin, with a notch anteriorly at the base of the reduced pseudoflagellum tip and the tip relatively long and curving laterally or medially.</p><p>Description.</p><p>Mature males observed with (40+4) rings, 2.5 mm midbody diameter to (63+1) rings, 3.9 mm. Cardo not deeper posteriorly. Leg 7 with elongated coxa, legs 10 and 11 with less elongated coxae. Prefemoral pad ca 1/2 femur length. Striae on posterior metazonites reaching ca 3/4 of ozopore height.</p><p>Coxite process on anterior gonopod (Fig. 6E) with posterodistal margin extended as rounded tab and sharply folded over laterally. Telopodite with single row of prominent setae on posterior side of medial thickening. Pseudoflagellum ca 2/3 telopodite width at base, posterior margin sometimes sinuous in transverse plane; abruptly narrowing apically with a shallow notch dividing the apex into small, rounded, anterior tooth and short, thinly laminar, acutely pointed extension, the latter usually folded over laterally and carrying the prostatic groove to its pointed tip.</p><p>Distribution.</p><p>Eucalypt forest and cool temperate rainforest in northwest Tasmania (Fig. 7B), from near sea level to at least 760 m. Meets Amastigogonus fossuliger parapatrically along the Mersey Break (Mesibov 1999) and overlaps to a small extent in far northwest Tasmania with Amastigogonus verreauxii .</p><p>Name.</p><p>For Henry Hellyer (1790-1832), explorer of northwest Tasmania; noun in the genitive case.</p><p>Remarks.</p><p>The tip of the pseudoflagellum in preserved males is often bent laterally into the space between the pseudoflagellum and the rest of the telopodite, as shown in Fig. 6E. In other specimens it may be bent medially towards the coxite tip, or extend past the tip of the telopodite. This species otherwise varies little across its range.</p></div>	https://treatment.plazi.org/id/6F2053EE20523E53F5C757BE75F7D303	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Mesibov, Robert	Mesibov, Robert (2017): Iulomorphid millipedes (Diplopoda, Spirostreptida, Iulomorphidae) of Tasmania, Australia. ZooKeys 652: 1-36, DOI: http://dx.doi.org/10.3897/zookeys.652.12035, URL: http://dx.doi.org/10.3897/zookeys.652.12035
234E5716110BE335B8878CAD6184C8BB.text	234E5716110BE335B8878CAD6184C8BB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amastigogonus michaelsae	<div><p>Amastigogonus michaelsae sp. n. Figs 4E, 6F</p><p>Holotype .</p><p>Male, Douglas Creek, Tas, -42.5139 147.7767 ± 100 m [label "EN 637 927" (= 55G 563700 5292700, AGD66)], 210 m a.s.l., 24 April 1991, R. Mesibov, QVM 23:54374; dissected, with head and anterior rings in genitalia vial.</p><p>Paratypes.</p><p>2 males, Montgomery Road, Tas, -42.6863 147.7111 ± 50 m, 330 m a.s.l., 21 June 2016, R. Mesibov, QVM 23:54548.</p><p>Other material.</p><p>11 males and 1 probable female from 8 unique localities; details in Suppl. material 1.</p><p>Diagnosis .</p><p>Like Amastigogonus peninsulensis sp. n. in having a relatively broad, gently tapering pseudoflagellum; distinguished by the pseudoflagellum tip directed distally rather than posteriorly and with a prominent, posterodistally directed tooth basally on the posterior margin.</p><p>Description.</p><p>Mature males observed with (55+1) rings, 3.3 mm midbody diameter to (66+1) rings, 3.4 mm. Cardo not deeper posteriorly. Leg 7 (only) with elongated coxa. Prefemoral pad ca 3/4 femur length. Striae on posterior metazonites reaching 3/4 ozopore height.</p><p>Coxite process on anterior gonopod (Fig. 6F) with posterodistal margin not extended and folded over. Telopodite without pronounced medial thickening, but with usual row of setae from near posterior margin to midline at telopodite apex, continued basally as group of sparse setae (not shown in Fig. 6F) behind tip of pseudoflagellum. Pseudoflagellum ca 1/3 telopodite width at base, curving slightly posteriorly and tapering gradually to rounded, posteriorly directed apex, and with short, sharp, posterodistally directed tooth at ca 1/4 pseudoflagellum height on posterior margin.</p><p>Distribution.</p><p>Eucalypt forest in southeast Tasmania from Coles Bay south to the Nugent area, including Maria Island (Fig. 7C), from sea level to at least 590 m. Possibly parapatric with Amastigogonus danpicola sp. n. near Swansea, parapatric or overlapping with Amastigogonus orientalis sp. n. west of Triabunna.</p><p>Name.</p><p>For the ecologist Karyl Michaels, who trapped specimens in the previously little-sampled dry forests of southeast Tasmania; noun in the genitive case.</p><p>Remarks.</p><p>Most of the non-type males are partial or fragmented specimens.</p></div>	https://treatment.plazi.org/id/234E5716110BE335B8878CAD6184C8BB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Mesibov, Robert	Mesibov, Robert (2017): Iulomorphid millipedes (Diplopoda, Spirostreptida, Iulomorphidae) of Tasmania, Australia. ZooKeys 652: 1-36, DOI: http://dx.doi.org/10.3897/zookeys.652.12035, URL: http://dx.doi.org/10.3897/zookeys.652.12035
59C6E74646C2DC5F6DBAF25F700D3BB7.text	59C6E74646C2DC5F6DBAF25F700D3BB7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amastigogonus orientalis	<div><p>Amastigogonus orientalis sp. n. Fig. 8A</p><p>Holotype.</p><p>Male, Maclaines Creek, Tas, -42.4628 147.8564 ± 100 m [label "EN 703 983" (= 55G 570300 5298300, AGD66)], 260 m a.s.l., 26 April 1991, R. Mesibov, QVM 23:54523 (in 2 pieces).</p><p>Paratypes.</p><p>2 males, details as for holotype, QVM 23:54401.</p><p>Other material.</p><p>14 males and 3 probable females from 14 unique localities; details in Suppl. material 1.</p><p>Diagnosis.</p><p>Like Amastigogonus hellyeri sp. n. in having a broad pseudoflagellum abruptly narrowed apically; distinguished by having a subquadrate extension of the posterobasal telopodite margin, with the prostatic groove opening on a very short tooth-like extension of the reduced pseudoflagellum tip, rather than on a relatively long, flexible extension.</p><p>Description.</p><p>Mature males observed with (43+3) rings, 2.6 mm midbody diameter to (67+0) rings, 4.2 mm. Cardo not deeper posteriorly. Leg 7 (only) with elongated coxa. Prefemoral pad ca 1/2 femur length. Striae on posterior metazonites reaching 3/4 ozopore height.</p><p>Coxite process on anterior gonopod (Fig. 8A) with posterodistal margin substantially extended as rounded tab and folded over laterally to ca 1/2 process width. Telopodite with posterobasal surface produced posteriorly as large rounded flange (Fig. 8A; fl) and with single row of prominent setae on posterior side of medial thickening . Pseudoflagellum ca 1/2 width of telopodite at base, expanded slightly posteriorly, tapering gradually before expanding distally in wedge shape, often curving laterally, with very short pointed extension at middle of distal margin; prostatic groove making S-bend from anterior side of pseudoflagellum into tip, terminating in pointed extension. Posterior margin of pseudoflagellum sometimes sinuous in transverse plane.</p><p>Distribution.</p><p>Eucalypt forest over ca 100 km linear extent in the Eastern Tiers of Tasmania (Fig. 7A) from ca 150 to at least 600 m elevation; the disjunction in the distribution map is likely to be a sampling artefact. Possibly parapatric with Amastigogonus elephas sp. n. near Gray; overlapping to a small extent with Amastigogonus danpicola sp. n. in the Douglas-Apsley National Park and with Amastigogonus fossuliger northwest of Triabunna; parapatric or overlapping with Amastigogonus michaelsae sp. n. in the southern Eastern Tiers.</p><p>Name.</p><p>Latin orientalis, eastern; adjective. This species is restricted to the East Coast region of Tasmania.</p></div>	https://treatment.plazi.org/id/59C6E74646C2DC5F6DBAF25F700D3BB7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Mesibov, Robert	Mesibov, Robert (2017): Iulomorphid millipedes (Diplopoda, Spirostreptida, Iulomorphidae) of Tasmania, Australia. ZooKeys 652: 1-36, DOI: http://dx.doi.org/10.3897/zookeys.652.12035, URL: http://dx.doi.org/10.3897/zookeys.652.12035
E0EE2CD020F941F28D3A8A9BCFF5ED5C.text	E0EE2CD020F941F28D3A8A9BCFF5ED5C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amastigogonus peninsulensis	<div><p>Amastigogonus peninsulensis sp. n. Fig. 8B</p><p>Holotype .</p><p>Male, Coal Mine Hill, Tas, -42.9852 147.7113 ± 25 m, 40 m a.s.l., 20 June 2016, R. Mesibov, QVM 23:54544.</p><p>Paratypes.</p><p>13 males and 5 females, details as for holotype, QVM 23:54545.</p><p>Other material.</p><p>3 males and 1 female from 3 unique localities; details in Suppl. material 1.</p><p>Diagnosis.</p><p>Like Amastigogonus michaelsae sp. n. in having a relatively broad, gently tapering pseudoflagellum, but with the tip directed distally and without a prominent tooth on the posterior margin.</p><p>Description.</p><p>Mature males observed with (40+3) rings, 1.8 mm midbody diameter to (60+1) rings, 2.5 mm. Cardo not deeper posteriorly. Leg 7 (only) with elongated coxa. Prefemoral pad ca 3/4 femur length. Striae on posterior metazonites reaching 1/2 ozopore height.</p><p>Coxite process on anterior gonopod (Fig. 8B) with very small portion of posterodistal margin slightly extended as rounded tab, bent laterally rather than folded over. Telopodite with single row of prominent setae on posterior side of medial thickening and shorter row of similarly prominent setae on anterior side. Pseudoflagellum ca 1/2 width of telopodite at base, posterior margin a little expanded at 1/3-1/2 pseudoflagellum height, apex acuminate and directed posteriorly.</p><p>Distribution.</p><p>Eucalypt forest on Forestier and Tasman Peninsulas (Fig. 7A) in Tasmania, from near sea level to at least 260 m.</p><p>Name.</p><p>For the Tasman Peninsula, type locality of this species; adjective.</p></div>	https://treatment.plazi.org/id/E0EE2CD020F941F28D3A8A9BCFF5ED5C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Mesibov, Robert	Mesibov, Robert (2017): Iulomorphid millipedes (Diplopoda, Spirostreptida, Iulomorphidae) of Tasmania, Australia. ZooKeys 652: 1-36, DOI: http://dx.doi.org/10.3897/zookeys.652.12035, URL: http://dx.doi.org/10.3897/zookeys.652.12035
B320447A060D7F1AEDE6FE54F4A5504A.text	B320447A060D7F1AEDE6FE54F4A5504A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amastigogonus tasmanianus Broelemann 1913	<div><p>Amastigogonus tasmanianus Broelemann, 1913 Figs 3F, 4F, 5A, 8C</p><p>Amastigogonus tasmanianus Brölemann 1913: 153, figs 32-37. Attems 1914: 293. Chamberlin 1920: 167. Verhoeff 1944: 43. Jeekel 1971: 107; 1981: 40. Korsós and Read 2012: 45.</p><p>Lectotype</p><p>(here designated). Male, Tasmania, date and collector unknown, AM KS.125304 (ex KS.37403). Intact specimen in 12 mm glass vial in 80% alcohol with original label " Amastigogonus / tasmanianus m. / (H.W.B.)".</p><p>Paralectotypes.</p><p>Collection details as for lectotype, AM KS.37403. 1 female and parts of 2 dissected males: last 23 podous rings + apodous rings + telson, midbody 31 rings, head capsule + collum, and ring 7 (gonopods missing) and following 2 podous rings. Also 8 small filter-paper envelopes containing cleared body parts illustrated by Brölemann (1913). In 25 mm glass vial with original label " Amastigogonus / tasmanianus m. / (H.W.B.)".</p><p>Other material.</p><p>92 males, 1 probable female and 1 possible juvenile from 8 unique localities; details in Suppl. material 1.</p><p>Diagnosis.</p><p>Distinguished from all other Amastigogonus species by having a long, gradually tapering, narrowly ribbon-like pseudoflagellum.</p><p>Description.</p><p>Mature males observed with (46+2) rings, midbody diameter 2.5 mm to (67+1) rings, 3.4 mm. Cardo not deeper posteriorly. Leg 7 (only) with elongated coxa. Prefemoral pad 1/3-1/2 femur length (Fig. 5A). Striae on posterior metazonites reaching ca 1/2 of ozopore height.</p><p>Coxite process on anterior gonopod (Fig. 8C) with posterodistal margin not extended and folded over. Telopodite with single row of prominent setae on posterior and anterior sides of medial thickening. Pseudoflagellum arising at 1/3-1/2 telopodite height, ca 1/2 width of telopodite at base, ribbon-like, curving sinuously first posterodistally, then anterodistally, tapering abruptly to sharp point (ribbon-like apex of pseudoflagellum shown edge-on in Fig. 8C).</p><p>Distribution.</p><p>Eucalypt forest and cool temperate rainforest at scattered locations in northeast Tasmania (Fig. 7A), from ca 100 m elevation to at least 1000 m. Co-occurs with Amastigogonus fossuliger .</p><p>Remarks.</p><p>The types were in excellent condition when examined in 2016. Because it is not possible to decide which of the two dissected male syntypes (or both) was illustrated by Brölemann, I am unable to follow Recommendation 74B (Preference for illustrated specimen) of the International Code of Zoological Nomenclature in choosing a lectotype, and instead have selected the intact male syntype. The lectotype is the third of the three males listed by Brölemann (1913: 154): "length 44 m/m; diameter 2.80 m/m; 56 segments; three segments apodous; 99 pair of legs".</p><p>I suspect that the type locality is the Launceston area, and the QVM male illustrated in Figs 3F, 4F and 5A is from Mowbray in Launceston.</p></div>	https://treatment.plazi.org/id/B320447A060D7F1AEDE6FE54F4A5504A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Mesibov, Robert	Mesibov, Robert (2017): Iulomorphid millipedes (Diplopoda, Spirostreptida, Iulomorphidae) of Tasmania, Australia. ZooKeys 652: 1-36, DOI: http://dx.doi.org/10.3897/zookeys.652.12035, URL: http://dx.doi.org/10.3897/zookeys.652.12035
106E9E764652D4EF393A81C603A91B4A.text	106E9E764652D4EF393A81C603A91B4A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amastigogonus verreauxii (Gervais 1847) Gervais 1847	<div><p>Amastigogonus verreauxii (Gervais, 1847) Figs 4G, 8D</p><p>Iulus verreauxii Gervais 1847: 175.</p><p>Julus Verreauxii Preudhomme de Borre 1884: 62.</p><p>" Julus " verreauxii Jeekel 1981: 43.</p><p>Amastigogonus verreauxii Mauriès et al. 2001: 585 (new combination), fig. 3. Korsós and Read 2012: 45.</p><p>Holotype.</p><p>Male, "De la Nouvelle-Hollande, sur le penchant du mont Wellington, par M. Jules Verreaux" (Gervais 1847: 175), MNHN GA031. Described and illustrated by Mauriès et al. (2001), specimen not re-examined.</p><p>Other material.</p><p>360 males, 100 probable females and 34 possible juveniles from 111 unique localities; details in Suppl. material 1.</p><p>Diagnosis.</p><p>Like Amastigogonus elephas sp. n. in having a telopodite with a subquadrate extension of the posterobasal margin and a posteriorly curving pseudoflagellum; distinguished from Amastigogonus elephas sp. n. in the pseudoflagellum lacking a small, tooth-like, distally directed extension on the tip.</p><p>Description.</p><p>Mature males observed with (38+4) rings, midbody diameter 2.2 mm to (55+1) rings, 3.2 mm in single 1-month pitfall sample, QVM 23:54197. Cardo not deeper posteriorly. Leg 7 (only) with elongated coxa. Prefemoral pad ca 3/4 femur length. Striae on posterior metazonites reaching ca 3/4 of ozopore height.</p><p>Coxite process on anterior gonopod (Fig. 8D) with small portion of posterodistal margin extended as rounded tab, sharply folded over laterally. Telopodite with posterobasal surface produced posteriorly as subquadrate flange (Fig. 8D; fl) and with single row of prominent setae on posterior side of medial thickening, on anterior side of thickening continued to telopodite apex as short row of more closely spaced setae (not shown in Fig. 8D). Pseudoflagellum arising at ca 2/3 telopodite height, ca 1/2 width of telopodite at base, curving posterodistally, the apex narrowing to truncate, posterobasally directed tip.</p><p>Distribution.</p><p>Widespread in forested and some non-forested habitats in western and southern Tasmania and on the Central Plateau (Fig. 7A) from sea level to at least 1260 m; also found on Hunter Island in the Hunter Group in western Bass Strait. Overlaps (with some parapatry?) with Amastigogonus hellyeri sp. n. in northwest Tasmania, and with Amastigogonus fossuliger on the Central Plateau and possibly in far southern Tasmania (see distribution notes for Amastigogonus fossuliger).</p><p>Remarks.</p><p>Gervais (1847) described Iulus Verreauxii from material in the Muséum national d’Histoire naturelle in Paris. A presumed holotype was still in the Muséum more than 150 years later and was redescribed and illustrated by Mauriès, Golovatch and Hoffman (2001), who assigned the species to Amastigogonus . The type material had probably been collected on Mt Wellington by the naturalist Jules Pierre Verreaux ca 1843 during his residency in Tasmania (Maiden 1910: 153).</p><p>Mauriès, Golovatch and Hoffman (2001: 585) refer to a "very faint axial line" on the promentum of the Amastigogonus verreauxii holotype. The line appears to be an artefact of long preservation, as I have not observed it in any Amastigogonus verreauxii specimens.</p><p>Amastigogonus verreauxii varies remarkably little in size or gonopod details over its large range.</p></div>	https://treatment.plazi.org/id/106E9E764652D4EF393A81C603A91B4A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Mesibov, Robert	Mesibov, Robert (2017): Iulomorphid millipedes (Diplopoda, Spirostreptida, Iulomorphidae) of Tasmania, Australia. ZooKeys 652: 1-36, DOI: http://dx.doi.org/10.3897/zookeys.652.12035, URL: http://dx.doi.org/10.3897/zookeys.652.12035
5AE1D28C91E5138DD669B9D1D56A2745.text	5AE1D28C91E5138DD669B9D1D56A2745.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atelomastix Attems 1911	<div><p>Atelomastix Attems, 1911</p><p>Atelomastix Attems 1911: 183 (in genus key), 192 (first description); 1926: 206; 1928: 312. Verhoeff 1913: 59; 1924: 74, 83; 1932: 1728, 1732, 1735, 1741; 1944: 33. Jeekel 1971: 107 (type species designated); 1985: 106; 2009: 31. Hoffman 1980: 91. Mauriès 1987: 196, 198. Korsós and Johns 2009: 3. Edward and Harvey 2010: 6. Korsós and Read 2012: 44.</p><p>Type species.</p><p>Atelomastix albanyensis Attems, 1911, by subsequent designation.</p><p>Other assigned species.</p><p>Atelomastix albanyensis Attems, 1911, Atelomastix anancita Edward &amp; Harvey, 2010, Atelomastix attemsi Edward &amp; Harvey, 2010, Atelomastix bamfordi Edward &amp; Harvey, 2010, Atelomastix bonhami sp. n., Atelomastix brennani Edward &amp; Harvey, 2010, Atelomastix culleni Edward &amp; Harvey, 2010, Atelomastix danksi Edward &amp; Harvey, 2010, Atelomastix dendritica Edward &amp; Harvey, 2010, Atelomastix ellenae Edward &amp; Harvey, 2010, Atelomastix flavognatha Edward &amp; Harvey, 2010, Atelomastix francesae Edward &amp; Harvey, 2010, Atelomastix gibsoni Edward &amp; Harvey, 2010, Atelomastix grandis Edward &amp; Harvey, 2010, Atelomastix julianneae Edward &amp; Harvey, 2010, Atelomastix lengae Edward &amp; Harvey, 2010, Atelomastix longbottomi Edward &amp; Harvey, 2010, Atelomastix mainae Edward &amp; Harvey, 2010, Atelomastix melindae Edward &amp; Harvey, 2010, Atelomastix montana Edward &amp; Harvey, 2010, Atelomastix nigrescens Attems, 1911, Atelomastix poustiei Edward &amp; Harvey, 2010, Atelomastix priona Edward &amp; Harvey, 2010, Atelomastix psittacina Edward &amp; Harvey, 2010, Atelomastix rubricephala Edward &amp; Harvey, 2010, Atelomastix sarahae Edward &amp; Harvey, 2010, Atelomastix smithi sp. n., Atelomastix solitaria Jeekel, 2009, Atelomastix tigrina Edward &amp; Harvey, 2010, Atelomastix tumula Edward &amp; Harvey, 2010.</p></div>	https://treatment.plazi.org/id/5AE1D28C91E5138DD669B9D1D56A2745	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Mesibov, Robert	Mesibov, Robert (2017): Iulomorphid millipedes (Diplopoda, Spirostreptida, Iulomorphidae) of Tasmania, Australia. ZooKeys 652: 1-36, DOI: http://dx.doi.org/10.3897/zookeys.652.12035, URL: http://dx.doi.org/10.3897/zookeys.652.12035
02AC9988230EA7AE272B666759A2990B.text	02AC9988230EA7AE272B666759A2990B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atelomastix bonhami	<div><p>Atelomastix bonhami sp. n. Figs 9, 10A, C, D</p><p>Holotype.</p><p>Male, White Spur, Tas, -42.7764 146.0369 ± 100 m [label "DN 211 634" (= 55G 421100 5263400, AGD66)], 320 m a.s.l., 2 February 1994, R. Mesibov, QVM 23:54460.</p><p>Paratypes.</p><p>1 male, 1 female, details as for holotype, QVM 23:54176.</p><p>Other material.</p><p>7 males, 5 females and 5 juveniles from 7 unique localities in Tasmania; details in Suppl. material 1.</p><p>Diagnosis.</p><p>Most similar to Atelomastix gibsoni Edward &amp; Harvey, 2010 from the Ravensthorpe Ranges in Western Australia; both species have a wedge-shaped sclerite “b” and a non-bifurcate sclerite “c” . Differences between Atelomastix bonhami / Atelomastix gibsoni: rounded tab present under sclerite “a” near pseudoflagellum/no tab; anterior corner of distal margin of sclerite “b” higher than posterior corner/anterior corner lower than posterior corner; sclerite “c” taller than sclerite “b” / sclerite “c” shorter than sclerite “b” .</p><p>Description .</p><p>Atelomastix as a genus has been well characterised by Edward and Harvey (2010), who described or redescribed 27 species from Western Australia. Atelomastix bonhami sp. n. fits the genus description and only key details are noted here.</p><p>Colour in alcohol variable, dark blueish grey to dark brown anteriorly on metazonites, pale posteriorly; some specimens largely brown. Ocelli 30-40 in 4-5 horizontal rows. Mature males with (37+5) rings, 2.0 mm midbody diameter to (60+1) rings, 2.3 mm. Longitudinal striae on metazonites meeting suture almost at right angle. Female substantially more robust than male with similar ring number, e.g. male with (51+1) rings, midbody diameter 2.2 mm vs female with (52+1) rings, midbody diameter 2.8 mm, both in QVM 23:54140.</p><p>Male gonopod aperture with sides slightly raised (Fig. 9A). Anterior gonopods nearly touching along midline. Sclerite “a” (Fig. 10A) curving posteriorly with pseudoflagellum (Fig. 10A; ps) arising at ca 3/4 gonopod height, extending posteriorly and slightly distally and gradually tapering to rounded apex; a thin, rounded tab (Fig. 10A; t) arising just distal to pseudoflagellum origin along sclerite midline. Sclerite “b” arising medially at ca 1/3 gonopod height, ca 3x as wide distally as at origin, thickly lamellar with distal half slightly bent medially, the distal margin with rounded anterior corner, sloping posterobasally with small, rounded notch at posterior corner; a double row of small, short setae near distal margin. Sclerite “c” more or less cylindrical basally, the apex curving slightly anteriorly and spatulate, the concave surface facing anterolaterally; a few long setae on anterior surface of sclerite at level of sclerite “b” distal margin.</p><p>Posterior gonopods (Fig. 10C) appressed basally along midline, 1/3-1/2 anterior gonopod height. Each posterior gonopod basally conical, tapering and with anterolateral extension; extension slightly flattened with a few long, mediodistally directed setae near base of extension on posteromedial surface and a few short setae at ca 1/2 gonopod height on medial surface.</p><p>Leg 1 (Fig. 10D) with prefemur wider than long, slightly tapering, with triangular tab (Figs 9B, 10D; at) on anteromedial surface of prefemur; relative lengths of podomeres femur&gt;prefemur&gt;tarsus&gt;postfemur&gt;tibia; tarsus distally excavate on anteromedial surface; prefemur, femur and postfemur with a few small setae laterally. Legpair 1 clearly separated on coxosternite, the latter with a field of long setae anterolateral to each prefemur.</p><p>Posterolateral margin of preanal ring meets epiproct margin at obtuse angle, making anal valves appear more prominent in lateral view than in other Tasmanian Iulomorphidae (Fig. 9C).</p><p>Distribution.</p><p>Known from wet forest, rainforest and scrub over ca 1500 km2 in southwest Tasmania at elevations ca 300-1100 m (Fig. 7D).</p><p>Name.</p><p>For Kevin Bonham, Tasmanian malacologist and diligent millipede hunter, who collected this species at two remote sites in 2016; noun in the genitive case.</p><p>Remarks.</p><p>The posterior gonopod of Atelomastix bonhami sp. n. is similar to that of Atelomastix nigrescens as illustrated in figs 13, 14 and 118 of Edward and Harvey (2010), and leg 1 of Atelomastix bonhami sp. n. closely resembles leg 1 of Atelomastix solitaria as illustrated in fig. 2 of Jeekel (2009).</p><p>Jeekel (2009: figs 3, 4) does not show the Atelomastix solitaria anterior gonopod in lateral view, so the shapes of the sclerites are unclear. From Jeekel’s description of Atelomastix solitaria and his fig. 4, it appears that sclerite “c” is sharply bent with an acuminate apex, unlike the smoothly curving, apically rounded sclerite “c” of Atelomastix bonhami sp. n. and the following species, and is shorter than “b”, rather than longer as in Atelomastix bonhami sp. n. and the following species.</p></div>	https://treatment.plazi.org/id/02AC9988230EA7AE272B666759A2990B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Mesibov, Robert	Mesibov, Robert (2017): Iulomorphid millipedes (Diplopoda, Spirostreptida, Iulomorphidae) of Tasmania, Australia. ZooKeys 652: 1-36, DOI: http://dx.doi.org/10.3897/zookeys.652.12035, URL: http://dx.doi.org/10.3897/zookeys.652.12035
581B713C108982A963C5272DE136EBD8.text	581B713C108982A963C5272DE136EBD8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atelomastix smithi	<div><p>Atelomastix smithi sp. n. Fig. 10B</p><p>Holotype .</p><p>Male, Ooze Lake, Tas, -43.5003 146.7019 ± 100 m [label "DM 758 834" (= 55G 475800 5183400, AGD66)], 870 m a.s.l., 16 February 1988, S.J. Smith, QVM 23:54179; dissected.</p><p>Paratypes.</p><p>2 males and 1 juvenile, details as for holotype, QVM 23:54484; 1 male, Promontory Lake, Tas, -43.1667 146.3653 ± 1 km [label "DN 483 203" (= 55G 448300 5220300, AGD66)], 25 February 2004, P. Sugden, QVM 23:24959.</p><p>Other material.</p><p>None.</p><p>Diagnosis.</p><p>Readily distinguished from the otherwise similar Atelomastix bonhami sp. n. by the pseudoflagellum curving distally; this difference can be seen in undissected males.</p><p>Description.</p><p>As for Atelomastix bonhami sp. n., but the two known males with 40-50 ocelli in 5-6 horizontal rows; holotype with (54+0) rings, 2.2 mm midbody diameter, paratype with (47+1) rings, 2.1 mm. Females (QVM 23:54484) more robust than males: (42+1) and (47+0) rings, both 2.5 mm in midbody diameter.</p><p>Anterior gonopod (Fig. 10B) with sclerite “a” bending posteriorly near bluntly rounded, thickened apex; pseudoflagellum (Fig. 10B; ps) arising at ca 3/4 gonopod height, curving smoothly distally, bending medially and terminating just beneath sclerite “a” apex; a longitudinally divided, thickened tab (Fig. 10B; t) arising just distal to pseudoflagellum origin along sclerite “a” midline. Sclerite “b” arising medially at ca 1/3 gonopod height, ca 2x as wide distally as at origin, thickly lamellar, the distal margin with rounded corners, sloping anterobasally; a double row of small, short setae near distal margin. Sclerite “c” more or less cylindrical, slightly tapered, apex thickened with shallow notch on anterior surface; a few long setae on anterior surface of sclerite at level of sclerite “b” apex.</p><p>Distribution.</p><p>Known from two localities ca 45 km apart at ca 900 m elevation, south and west of the known range of Atelomastix bonhami sp. n. in southwest Tasmania (Fig. 7D). Atelomastix smithi sp. n. is likely to be more widely distributed in this little-sampled wilderness area.</p><p>Name.</p><p>For Steven J. Smith, first collector of this species and formerly a senior zoologist with the Tasmanian Parks and Wildlife Service; noun in the genitive case.</p></div>	https://treatment.plazi.org/id/581B713C108982A963C5272DE136EBD8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Mesibov, Robert	Mesibov, Robert (2017): Iulomorphid millipedes (Diplopoda, Spirostreptida, Iulomorphidae) of Tasmania, Australia. ZooKeys 652: 1-36, DOI: http://dx.doi.org/10.3897/zookeys.652.12035, URL: http://dx.doi.org/10.3897/zookeys.652.12035
CB57DED4FF69A75115719E7170680F55.text	CB57DED4FF69A75115719E7170680F55.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Equestrigonus	<div><p>Equestrigonus gen. n.</p><p>Diagnosis.</p><p>Like Amastigogonus and Victoriocambala Verhoeff, 1944 in having greatly reduced posterior gonopods with a single lateral process, and relatively simple, two-branched anterior gonopods with the prostatic groove entering a pseudoflagellum on the telopodite. Distinguished from the other two genera by the form of the anterior gonopods (Figs 11A, 13A): coxite process a bluntly pointed rod, longer than telopodite; telopodite slender with a distal, spreading crown of setae. Distinguished from all other Tasmanian Iulomorphidae by the posteroventral extension of the cardo (Fig. 12B; ce) in males.</p><p>Type species.</p><p>Equestrigonus tasmaniensis sp. n., by present designation.</p><p>Name.</p><p>From Latin equestris, genitive singular of equester, pertaining to horse-riding, plus -gonus, a suffix commonly used in millipede names, referring to the gonopods; masculine gender. The tips of the anterior gonopod of the type species (Fig. 11A) resemble the ends of prick and rowel spurs.</p></div>	https://treatment.plazi.org/id/CB57DED4FF69A75115719E7170680F55	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Mesibov, Robert	Mesibov, Robert (2017): Iulomorphid millipedes (Diplopoda, Spirostreptida, Iulomorphidae) of Tasmania, Australia. ZooKeys 652: 1-36, DOI: http://dx.doi.org/10.3897/zookeys.652.12035, URL: http://dx.doi.org/10.3897/zookeys.652.12035
09821AF776CF8C4CBDDEF44E5AB359FC.text	09821AF776CF8C4CBDDEF44E5AB359FC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Equestrigonus tasmaniensis	<div><p>Equestrigonus tasmaniensis sp. n. Figs 11, 12, 13</p><p>Holotype.</p><p>Male, Sideling Range, Tas, -41.2358 147.4131 ± 100 m [label "EQ 345 348" (= 55G 534500 5434800, AGD66)], 550 m a.s.l., 7 July 1993, T. Kingston, QVM 23:54173 (ex 23:11638).</p><p>Paratypes.</p><p>3 males, Sideling Range, Tas, -41.2303 147.4117 ± 100 m [label "EQ 344 354" (= 55G 534400 5435400, AGD66)], 550 m a.s.l., 15 June 1992, T. Kingston and R. D’Orazio, QVM 23:12719; 6 males, same details but 13 September 1993, T. Kingston et al., QVM 23:54057; 6 males, same locality but -41.2411 147.4106 ± 100 m [label "EQ 343 342" (= 55G 534300 5434200, AGD66)], 540 m a.s.l., 14 September 1993, T. Kingston et al., QVM 23:54094; 2 males, same locality but -41.2394 147.4083 ± 100 m [label "EQ 341 344" (= 55G 534100 5434400, AGD66)], 500 m a.s.l., 14 September 1993, T. Kingston et al., QVM 23:54095; 1 male, details as for holotype, 23:11637; 1 male, details as for holotype, QVM 23:11638.</p><p>Other material.</p><p>1013 males, 82 probable females and 12 possible juveniles from 68 unique localities in Tasmania; details in Suppl. material 1.</p><p>Description.</p><p>Living animals (Fig. 12A) have dark grey rings with annular pale band at rear of each metazonite; head, collum and last 1-2 rings before telson tinged with light brown; legs pale. With long storage in alcohol, ring colour fades to dark blue-grey, annular band darkens to light grey; often with brownish tinge on anterior rings; head and legs pale golden yellow (Fig. 11B).</p><p>Mature males (Fig. 11B) with (34+4) rings, 2.0 mm midbody diameter to (50+1) rings, 3.0 mm. Head (Fig. 12B) smooth, frons flattened, vertigial sulcus reaching to level of dorsalmost ocellar row. Posteroventral margin of cardo thickened and extended ventrally as large, rounded lobe (Fig. 12B; ce). Ocellar area narrow-triangular, triangle apex medial; ca 27 ocelli in 4 more or less regular horizontal rows, dorsal&gt;ventral typically 9+8+6+4. Antennae short, barely reaching past posterior edge of collum when manipulated dorsally; relative antennomere lengths (2=3)&gt;6&gt;(4=5); antennomere 6 widest; 4 apical cones; socket ca 1 socket diameter from lateral margin of head capsule. Gnathochilarium (Fig. 12E) with lateral edges of mentum (Fig. 12E, me) slightly convex, mentum wider than combined lingual plates (Fig. 12E, lp); mentum-promentum (Fig. 12E, pm) junction nearly straight or slightly concave anteriorly; a prominent pit (Fig. 12E, sp) with small seta anteriorly on each gnathochilarial stipes (Fig. 12E, gs). Collum convex, laterally narrowing with rounded corner, margins straight. Ventral margin of ring 2 swollen posteriorly. Prozonites only slightly narrower than metazonites; prozonites with weakly defined annular striae anteriorly; suture weakly defined; fine longitudinal striae on lower half of metazonite, anterior end of each stria bent obliquely upwards towards suture; prozonites and metazonites with surface otherwise smooth, free of setae. Ozopores small, beginning ring 6, opening just above 1/2 ring height at ca 1/3 the distance between suture and posterior metazonite margin. Limbus lamellar, undivided. Preanal ring smooth, epiproct broadly rounded, extend ing slightly over anal valves; hypoproct with margin slightly convex dorsally. Midbody legs short, ca 2/3 ring diameter when extended; relative podomere lengths (prefemur=femur)&gt;tarsus&gt;(postfemur=tibia), claw ca 1/2 tarsus length. Prefemur distally with ventral flattening and conical prefemoral pad ca 1/2 femur length on midbody legs; pads first appear on ring 5 legs, diminish in posterior 1/3 of body and are greatly reduced or absent on last few legpairs.</p><p>Legpair 1 (Fig. 12C, D) separate on coxosternite, each leg 1 composed of broad basal and leg-like distal portion. Basal portion here assumed to be prefemur; widest at base, tapering medially, extending anterolaterally at base as large lobe (Fig. 12D; all) with a few distal setae; prefemur marked with several quasi-annular chitinous ridges and anteriorly bearing small field of coarse setae (Fig. 12D; sf), with large subquadrate tab (Fig. 12D; at) extending anteriorly. Distal portion with 3 well-demarcated podomeres, here assumed to be femur, postfemur and fused tibia+tarsus; femur widest, the tibia+tarsus narrowest and longest and with faint annular subdivision; all 3 podomeres sparsely and shortly setose; no claw.</p><p>Leg 2 (Fig. 12C, F) incrassate with large claw, prefemur reduced, coxa expanded anteriorly and distally with thick, tab-like apex (Figs 12C, F; ct); penis (Fig. 12F; pe) arising basally on posterior coxal surface, barrel-shaped with a few long setae in distal, marginal crown.</p><p>Coxae not elongated on near-aperture legs. Rear portion of gonopod aperture flat, not raised behind gonopods.</p><p>Anterior gonopods (Figs 11A, 13A) with coxite process (Figs 11A, 13A; cx) tapering to blunt spine extending further distally than telopodite (Figs 11A, 13A; te), and with short, spine-like process (Fig. 13A; pr) arising on anterior surface at level of pseudoflagellum on telopodite and directed distally. Telopodite tapering strongly, apex spatulate with spreading, marginal crown of ca 15 long, well-spaced setae; pseudoflagellum (Figs 11A, 13A; ps) branching off at ca 3/4 telopodite height on anterior surface, tapering to blunt tip and curving medially to terminate proximal to telopodite apex.</p><p>Posterior gonopods (Fig. 13B) ca 1/2 length of anterior gonopods, tapering from base, apex truncate distolaterally with apical crown of long setae; lateral process arising at ca 1/2 gonopod height, terminating at same level as body of gonopod, tip slightly expanded and flattened with convex distal margin.</p><p>Mature females (specimens in QVM 23:54050 and 23:54467) a little larger than males with same ring number; cardo not extended ventrally as in male.</p><p>Distribution.</p><p>Wet eucalypt forest and cool temperate rainforest in northern Tasmania (Fig. 7D), from 50 m to at least 900 m elevation; not yet collected south of 41°30'S latitude. Wanders on the forest floor and climbs tree trunks at night; shelters during the day in and under rotting logs and in leaf litter.</p><p>Name.</p><p>For the occurrence of this species in Tasmania; adjective.</p><p>Remarks.</p><p>The distinctive tips of the anterior gonopods invariably protrude from the gonopod aperture (Fig. 11A), allowing males of Equestrigonus tasmaniensis gen. n., sp. n. to be recognised without dissection. Western Tasmanian males are generally a little larger in diameter than eastern males.</p></div>	https://treatment.plazi.org/id/09821AF776CF8C4CBDDEF44E5AB359FC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Mesibov, Robert	Mesibov, Robert (2017): Iulomorphid millipedes (Diplopoda, Spirostreptida, Iulomorphidae) of Tasmania, Australia. ZooKeys 652: 1-36, DOI: http://dx.doi.org/10.3897/zookeys.652.12035, URL: http://dx.doi.org/10.3897/zookeys.652.12035
