taxonID	type	description	language	source
03BA87FBFFD1FFDB9CD3F8CEFB01EFA2.taxon	description	(Fig. 2 A)	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD1FFDB9CD3F8CEFB01EFA2.taxon	materials_examined	Material examined. 1 male (5.6 x 5.0 mm) (DABFUK), Minicoy, 29 August 2013; 1 male (7.1 x 6.4 mm) (DAB- FUK), Kavaratti, 7 November 2013; 1 ovigerous female (7.3 x 6.1 mm) (DABFUK), Kilton, 22 October 2014. Comparative material: Guam — 1 ovig. female (7.2 x 6.5 mm) (ZRC 2000.0751), Gun Beach, fore reef, 3 – 5 m depth, at night, coll. L. Kirkendale, 5 July 1999, det. C. McLay, Apr. 2000; Philippines — 1 male (9.2 x 8.3 mm) (ZRC 2011.0891), Panglao I., Gak-ang, stn. M 18, 0 – 1 m depth, coll. PANGLAO 2004 Expedition, 10 & 12 June 2004, det. C. McLay, ca. 2011; Vanuatu — 1 male (6.3 x 5.9 mm) (ZRC 2011.0909), stn. DB 80, Bruat Channel, 18 m depth, sand & corals on submarine hill, coll. SANTO 2006 Expedition, 2 October 2006; 1 female (7.0 x 5.7 mm) (ZRC 2011.0902), stn. VM 36, south of Aés I., intertidal, coll. SANTO 2006 Expedition, 25 Sep. 2006; 1 male (8.5 x 8.0 mm) (ZRC 2011.0904), stn. VM 58, Palikulo Peninsula, intertidal, coll. SANTO 2006 Expedition, 7 October 2006; all det. C. McLay, ca. 2011.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD1FFDB9CD3F8CEFB01EFA2.taxon	discussion	Remarks. Cryptodromia hirsuta was described by Borradaile (1903) from the Maldives and has since been synonymized under C. fallax (Latreille, in Milbert, 1812), described from Réunion Island (McLay 1993, 2003). Examination of the type material of C. fallax and representative specimens in the ZRC, suggests that it is probably a species complex. The neotype of C. fallax is deposited in the MNHN (Cleva et al. 2007: 240, fig. 7 C), and the shape of the carapace and the dentition of the anterolateral margins differ with the present material from Lakshadweep (Fig. 2 A). The carapace of the neotype is more globose and the first two anterolateral teeth are distinctly separated. In the Lakshadweep material, the carapace is relatively flatter and more trapezoidal in outline, with the first two anterolateral teeth fused. Likewise, Borradaile’s figure of C. hirsuta (Borradaile, 1903: pl. 33 fig. 3 A) shows the first two anterolateral teeth separate from each other. In all three cases, the median rostral tooth is not much narrower than the adjacent lateral teeth. The Lakshadweep specimens, however, are smaller than the types of C. fallax (CW: 13.3 mm) or C. hirsuta (CW: 8.0 mm), which could explain the minor differences in morphology. We therefore, follow McLay’s (1993) synonymization of C. hirsuta Borradaile under C. fallax (Latreille) for the time being. More substantial differences distinguish the Lakshadweep specimens from C. fallax collected from the western Pacific (e. g. Guam and Vanuatu): the median rostral tooth is only slightly narrower than the two lateral teeth (versus median tooth much narrower than the laterals in the Pacific material), the supraorbital teeth are relatively blunt and positioned closer together (versus more acute and spaced further apart in the Pacific material), and the infraorbital tooth is subtruncate (versus more acutely triangular in the Pacific material). It is very likely that the Pacific material is not conspecific with C. fallax; this will require a more thorough examination of the available material and synonyms, which are beyond the scope of the present work. This species has been recorded from Lakshadweep by Dev Roy & Nandi (2005). Among the present specimens, the smaller male (5.6 x 5.0 mm) from Minicoy was collected carrying a colonial ascidian.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD2FFDB9CD3F980FD01EDD7.taxon	materials_examined	Material examined. 1 male (56.4 x 34.2 mm) (DABFUK), Agatti, 6 January 2014; 1 male (52.2 x 31.7 mm) (DAB- FUK), Kalpeni, 30 October 2013; 1 female (50.5 x 32.8 mm) (DABFUK), Minicoy, 29 August 2013.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD2FFDB9CD3F980FD01EDD7.taxon	discussion	Remarks. Previously recorded from Lakshadweep by Alcock (1894, 1896); Thomas (1969); Suvarna Devi et al. (2014); and Kumaralingam et al. (2017).	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD2FFDA9CD3F8DCFAD4EBB2.taxon	materials_examined	Material examined. 5 males (47.4 x 35.0 mm, 48.6 x 33.0 mm, 49.4 x 33.6 mm, 53.8 x 38.7 mm, 60.4 x 44.6 mm) (DABFUK), Agatti, 7 January 2014; 1 female (40.9 x 27.0 mm), 2 males (36.2 x 24.2 mm, 39.8 x 27.6 mm) (DAB- FUK), Agatti, 20 May 2014.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD2FFDA9CD3F8DCFAD4EBB2.taxon	discussion	Remarks. Previously recorded from Lakshadweep by Rao et al. (1989) and Suvarna Devi et al. (2014).	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD3FFDA9CD3FDF0FC1FE847.taxon	materials_examined	Material examined. 1 male (46.6 x 32.1 mm) (DABFUK), Minicoy, 28 September 2014; 1 male (35.6 x 24.6 mm) (DABFUK), Bitra, 13 May 2015.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD3FFDA9CD3FDF0FC1FE847.taxon	discussion	Remarks. Previously recorded from Lakshadweep by Alcock (1894) and Rao et al. (1989). Specimens were also observed in Kavaratti (R. Ravinesh pers. obs.) but not collected.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD3FFDA9CD3FB97FBB6EFDF.taxon	materials_examined	Material examined. 1 male (13.7 x 9.7 mm) (DABFUK), Kadamat, 5 May 2014; 1 female (46.8 x 32.7 mm) (DAB- FUK), Chetlat, 4 November 2015.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD3FFDA9CD3FB97FBB6EFDF.taxon	discussion	Remarks. Previously recorded from Lakshadweep by Alcock (1894, 1898), Thomas (1969) and Suvarna Devi et al. (2014), this species has a wide range in the Indo-West Pacific (Serène 1984).	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD3FFDA9CD3FA20FD18EDBF.taxon	materials_examined	Material examined. 2 males (19.2 x 12.7 mm, 20.7 x 14.8 mm) (DABFUK), Kilton, 17 May 2015; 1 male (22.0 x 15.0 mm) (DABFUK), Kavaratti, 25 May 2015; 1 male (25.2 x 17.1 mm) (DABFUK), Amini, 29 May 2014; 1 male (18.9 x 12.9 mm) (DABFUK), Bitra, 10 May 2015.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD3FFDA9CD3FA20FD18EDBF.taxon	discussion	Remarks. Previously recorded from Lakshadweep by Alcock (1898) and Borradaile (1902 b). This is a common species in the Indo-West Pacific (Koh & Ng 2008) and was observed on the reefs from the 10 islands in the group, although not all the specimens were collected.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD3FFDA9CD3FF5BFE81EAA5.taxon	materials_examined	Material examined. 1 male (34.5 x 24.3 mm) (DABFUK), Minicoy, 29 August 2013; 1 female (31.0 x 22.7 mm) (DABFUK), Kavaratti, 7 November 2013.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD3FFDA9CD3FF5BFE81EAA5.taxon	discussion	Remarks. New record from Lakshadweep. This species was also observed from Agatti but material was not collected.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD3FFDD9CD3F960FBA3EBFB.taxon	materials_examined	Material examined. 2 males (49.6 x 38.3 mm, 65.3 x 49.6 mm) (DABFUK), Kilton, 20 May 2015; 1 male (28.6 x 21.6 mm) (DABFUK), Minicoy, 18 January 2015; 1 female (53.6 x 41.6 mm) (DABFUK), Agatti, 20 May 2014; 1 female (47.7 x 36.4 mm) (DABFUK), Chetlat, 4 November 2015; 1 female (29.5 x 22.6 mm) (DABFUK), Amini, 25 May 2014; 1 female (20.9 x 13.3 mm) (DABFUK), Androth, 29 January 2015.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD3FFDD9CD3F960FBA3EBFB.taxon	discussion	Remarks. The taxonomy of E. sebana and allied species was clarified by Koh & Ng (2008); it has been previously recorded from Lakshadweep by Alcock (1894, 1898) and Suvarna Devi et al. (2014). This species was also seen in Kalpeni, Kavaratti, Kadamat and Bitra, but specimens were not collected.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD4FFDD9CD3FE5BFDB0EF78.taxon	description	(Fig. 3 A – F)	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD4FFDD9CD3FE5BFDB0EF78.taxon	materials_examined	Material examined. Lectotype (here designated): male (5.0 x 4.4 mm) (CMZ I 64238), on reef, Hulule, Male Atoll, Maldives, coll. J. S. Gardner, June 1900. Paralectotypes: 1 ovigerous female (7.3 x 6.2 mm), same data as lectotype; 1 female (6.0 x 5.1 mm) (CMZ I 64236), on reef, Minicoy, Lakshadweep, coll. J. S. Gardner, June 1900. Others: 1 female (6.0 x 5.3 mm) (DABFUK), Kavaratti, 15 October 2014.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD4FFDD9CD3FE5BFDB0EF78.taxon	discussion	Remarks. The species was first described based on material collected from Minicoy (Lakshadweep) and Male Atoll (Maldives) by Borradaile (1903 c), with Gordon (1940) redescribing the species and figuring its gonopods for the first time. The species was subsequently listed by Dev Roy & Nandi (2005). Nevertheless, E. gracilis is not well known, so the types are figured here. Borradaile (1903 c) and Gordon (1940) did not designate a holotype for E. gracilis so all the three specimens are syntypes. The male from Male Atoll is here selected as the lectotype of the species. Gordon (1940) compared E. gracilis with the similar E. sindensis Alcock, 1900, and E. mathoei (Desmarest, 1823) from the Indian Ocean and showed they were distinct in several carapace, ambulatory leg and G 1 characters (see also Ng & Chuang 1996). The carapace shape varies slightly, with the lectotype male proportionately longer and the anterolateral margins slightly more concave (Fig. 3 A) than in other specimens; the female carapace may also be proportionally wider (Fig. 3 B, C).	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD4FFDD9CD3FA25FAE0EDDC.taxon	materials_examined	Material examined. 1 male (11.5 x 15.3 mm) (DABFUK), Androth, 10 November 2013; 1 ovigerous female (8.3 x 10.8 mm) (DABFUK), Bitra, 8 May 2015.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD4FFDD9CD3FA25FAE0EDDC.taxon	discussion	Remarks. The long, basally narrow rostrum, distinctly areolate dorsal carapace surface and non-cristate meri of the ambulatory legs distinguishes this widely distributed species from the closely related M. orientalis (Sakai, 1969) (see Griffin & Tranter 1986). This species is common in Agatti, Minicoy, Androth, Bitra, Kilton, Kalpeni and Kadamat but material was not collected. Menaethius monoceros has previously been reported from Lakshadweep (Thomas 1969; Bakus 1994; Venkataraman et al. 2004; Suvarna Devi et al. 2014; Kumaralingam et al. 2017).	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD4FFDF9CD3F8EBFEAEEBD6.taxon	materials_examined	Material examined. 1 female (13.2 x 14.1 mm) (DABFUK), Amini, 28 May 2014.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD4FFDF9CD3F8EBFEAEEBD6.taxon	discussion	Remarks. A new record for Lakshadweep. This so-called ‘ decorator crab’ is a widespread species in the Indo- West Pacific.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD6FFDF9CD3FE2AFC15E9C5.taxon	materials_examined	Material examined. 2 males (11.0 x 10.9 mm, 26.4 x 24.2 mm) (DABFUK), Kavaratti, 30 December 2014; 1 female (30.2 x 29.4 mm) (DABFUK), Amini, 26 May 2014; 1 ovigerous female (27.2 x 24.6 mm) (DABFUK), Kavaratti, 25 May 2014	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD6FFDF9CD3FE2AFC15E9C5.taxon	discussion	Remarks. The species has been recorded from Tamil Nadu (Alcock 1895; Jeyabaskaran et al. 2000), the Andaman and Nicobar Islands (Alcock 1895; Dev Roy 2008), and from Lakshadweep (Suvarna Devi et al. 2014). This species was observed at Kadamat but specimens were not collected.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD6FFDF9CD3FA0FFEDAED56.taxon	materials_examined	Material examined. 1 ovigerous female (18.0 x 12.9 mm) (DABFUK), Amini, 26 May 2014.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD6FFDF9CD3FA0FFEDAED56.taxon	discussion	Remarks. Pilumnus longicornis is widespread in the Indo-West Pacific (Takeda & Miyake 1968), typically found at the edges of reefs. It has been previously recorded from Lakshadweep by Rao et al. (1989) and Venkataraman et al. (2004).	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD6FFDF9CD3FD6AFC1FE899.taxon	materials_examined	Material examined. 1 male (22.8 x 25.9 mm) (DABFUK), Amini, 3 June 2015. Remarks. A new record for Lakshadweep. Griffin (1966) treated the taxonomy of this species at length. It is widely distributed in the Indo-West Pacific (Griffin & Tranter 1986).	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD6FFDF9CD3FC38FA0AEFB0.taxon	materials_examined	Material examined. 1 male (4.7 x 7.6 mm) (DABFUK), Kavaratti, 11 October 2014.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD6FFDF9CD3FC38FA0AEFB0.taxon	discussion	Remarks. The present specimen is a subadult and was probably freshly moulted before it was collected. It does not appear to belong to any of the 14 recognised species of Pseudomicippe Heller, 1861 (Griffin & Tranter 1986; Ng et al. 2008). It has a different carapace shape from P. maldivensis Griffin & Tranter, 1986, also described from the same area. Intact adult specimens, especially males, will need to be obtained before this species can be described.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD6FFDE9CD3F8F1FC5FEBD6.taxon	description	(Fig. 4 A)	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD6FFDE9CD3F8F1FC5FEBD6.taxon	materials_examined	Material examined. 1 female (14.8 x 9.7 mm) (DABFUK), Bitra, 9 May 2015.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD6FFDE9CD3F8F1FC5FEBD6.taxon	discussion	Remarks. A new record for Lakshadweep. Previously recorded from Tamil Nadu (Alcock 1894, 1899 b) as well as Andaman and Nicobar Islands (Alcock 1899 b; Dev Roy & Nandi 2012)	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD7FFDE9CD3FB73FF65EE9E.taxon	materials_examined	Material examined. 1 male, (57.1 x 37.0 mm) (DABFUK), Agatti, 10 January 2014. Remarks. Previously recorded from Lakshadweep (Thomas 1969; Rao et al. 1989; Venkataraman et al. 2004).	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD7FFDE9CD3FC37FAFDEFC6.taxon	description	(Fig. 2 B)	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD7FFDE9CD3FC37FAFDEFC6.taxon	materials_examined	Material examined. 1 female (35.9 x 27.0 mm) (DABFUK), Agatti, 20 May 2014	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD7FFDE9CD3FC37FAFDEFC6.taxon	discussion	Remarks. The single specimen of this widespread species presents an interesting new record for India (viz. Apel & Spiridinov 1998). Galil et al. (2018) recently reappraised the taxonomy of Thalamita giardi Nobili, 1905, an Indian Ocean species long synonymized under G. paucidentatus, and showed that it to be a valid species. The shape of the frontal margin of the present specimen, however, indicates that the present specimen is G. paucidentatus.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD7FFDE9CD3FE83FC37EAAE.taxon	materials_examined	Material examined. 1 male (10.1 x 8.8 mm) (DABFUK), Kavaratti, 19 May 2015. Remarks. This species, which is always associated with sea cucumbers, has previously been recorded from Lakshadweep (Alcock 1899 b; Thomas 1969; Dev Roy & Nandi 2005).	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD7FFDE9CD3FA4BFB6BEDBE.taxon	materials_examined	Material examined. 1 female (15.0 x 10.7 mm) (DABFUK), Kavaratti, 18 January 2015. This species was also observed from Minicoy (R. Ravinesh) but specimens not collected.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD7FFDE9CD3FA4BFB6BEDBE.taxon	discussion	Remarks. Thalamita admete was also observed from Minicoy but specimens were not collected. This is a widespread species in the Indo-West Pacific (Wee & Ng 1995; Apel & Spiridinov 1998); and it has previously been recorded from Lakshadweep by Thomas (1969), Rao et al. (1989), and Venkataraman et al. (2004).	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD7FFDE9CD3F96BFB03EC96.taxon	materials_examined	Material examined. 1 female (37.2 x 23.7 mm) (DABFUK), Agatti 20 May 2014. Remarks. Thalamita prymna was previously recorded from Lakshadweep by Borradaile (1902 a). The taxonomy of this species was discussed by Wee & Ng (1995) and Apel & Spiridonov (1998).	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD7FFDE9CD3FDCFFCC7E816.taxon	description	(Fig. 2 C)	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD7FFDE9CD3FDCFFCC7E816.taxon	materials_examined	Material examined. 1 male (24.7 x 11.6 mm), 1 ovig. female (26.1 x 11.8 mm) (DABFUK), Agatti, 8 January 2014. Remarks. This distinctive species is recorded for the first time from India. It has otherwise been reported from various parts of the Indian Ocean (Apel & Spiridonov 1998).	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD8FFD19CD3FA4AFB7EEC06.taxon	materials_examined	Material examined. 3 males (21.1 x 13.2 mm, 25.2 x 15.0 mm, 25.8 x 15.8 mm) (DABFUK), Kalpeni, 4 January 2013; 1 male (25.5 x 15.7 mm), 1 female (15.2 x 8.9 mm) (DABFUK), Kavaratti, 31 October 2013; 2 males (25.0 x 15.5 mm, 26.5 x 16.2 mm) (DABFUK), Minicoy, 2 December 2013; 2 females (17.6 x 10.8 mm, 22.8 x 14.1 mm) (DABFUK), Agatti, 0 8 January 2014; 2 females (17.1 x 11.0 mm, 18.3 x 11.7 mm) (DABFUK), Kilton, 22 October 2014; 1 male (26.8 x 16.5 mm) (DABFUK), Bitra, 0 8 May 2015.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD8FFD19CD3FA4AFB7EEC06.taxon	discussion	Remarks. Actaeodes tomentosus was previously recorded from Lakshadweep by Alcock (1898), Dev Roy (2008), and Kumaralingam et al. (2017). It has a wide range in the Indo-West Pacific (Serène 1984).	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD8FFD19CD3FEAEFE4AE91A.taxon	materials_examined	Material examined. 1 ovigerous female (5.7 x 4.1 mm) (DABFUK), Minicoy, 16 September 2014.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD8FFD19CD3FEAEFE4AE91A.taxon	discussion	Remarks. Cherusius was better known as Jonesius Sankarankutty, 1962, but Low & Ng (2012) showed that the name was preoccupied by the cestode Jonesius Yamaguti, 1959. As such, they proposed a replacement name, Cherusius. Cherusius triunguiculatus has been previously reported from Lakshadweep by Rao et al. (1989) and Venkataraman et al. (2004).	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD8FFD19CD3F8B2FEFFECDE.taxon	materials_examined	Material examined. 1 male (13.7 x 10.0 mm), 1 female (18.1 x 13.6 mm) (DABFUK), Amini, 26. May 2014. Remarks. A new record for Lakshadweep. This species is widely distributed in the Indo-West Pacific (Guinot 1976; Serène 1984).	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD8FFD19CD3FD32FB53E885.taxon	materials_examined	Material examined. 1 female (15.9 x 12.1 mm) (DABFUK), Kavaratti, 15 October 2014.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD8FFD19CD3FD32FB53E885.taxon	discussion	Remarks. Trapezia bidentata is better known throughout its range as Trapezia ferruginea Latreille, 1828 (cf. Serène 1984) but Castro et al. (2004) showed that T. bidentata (Forskål, 1775) was actually its senior synonym. It has previously been recorded from Lakshadweep by Rao et al. (1989) and Sankarankutty (1961).	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD8FFD19CD3FC36FB46EFA2.taxon	description	(Fig. 4 B)	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD8FFD19CD3FC36FB46EFA2.taxon	materials_examined	Material examined. 1 male (12.6 x 10.8 mm) (DABFUK), Bitra, 13 May 2015.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD8FFD19CD3FC36FB46EFA2.taxon	discussion	Remarks. Trapezia lutea is superficially similar to Trapezia cymodoce (Herbst, 1801) and many old records of the latter species will need to be checked (see Castro 1997). Both species have a wide Indo-West Pacific distribution and can occur together in the same reef. Trapezia lutea is recorded for the first time from India.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD9FFD09CD3FDA3FBDEE9F3.taxon	materials_examined	Material examined. 1 female (12.3 x 8.6 mm) (DABFUK), Minicoy, 13 August 2013, 1 female (15.3 x 10.1 mm) (DABFUK), Kavaratti, 4 January 2014.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD9FFD09CD3FDA3FBDEE9F3.taxon	discussion	Remarks. Cyclodius granulosus has previously been listed from Tamil Nadu by Jeyabaskaran et al. (2000) and Dev Roy & Nandi (2005); the present specimen is the first record from Lakshadweep.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD9FFD09CD3FCA7FC17E8CA.taxon	materials_examined	Material examined. 2 males (23.4 x 15.7 mm, 17.1 x 12.2 mm) (DABFUK), Bitra, 11 May 2015. Remarks. A common species on Indo-West Pacific coral reefs (Serène 1984). Previously recorded from Lakshadweep (Rao et al. 1989; Bakus 1994; Venkataraman et al. 2004).	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD9FFD09CD3FA4BFE78ED77.taxon	materials_examined	Material examined. 1 female (79.6 x 54.4 mm) (DABFUK), Kavaratti, 8 January 2013. Remarks. Previously recorded by Suvarna Devi et al. (2014) from Kavaratti. Specimens were also observed from Minicoy but not collected.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD9FFD39CD3F882FD63EBFA.taxon	description	(Fig. 2 D)	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD9FFD39CD3F882FD63EBFA.taxon	materials_examined	Material examined. 1 juvenile male (10.3 x 6.8 mm) (DABFUK), Kavaratti, 7 January 2013.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD9FFD39CD3F882FD63EBFA.taxon	discussion	Remarks. A new record for Lakshadweep. The present specimen of E. rugosus is a juvenile, but it already has the rugosities on the areolae of the dorsal surface of the carapace and the prominent outwardly directed projection on the carpus of the cheliped, a character typical for the species (cf. Guinot-Dumortier 1960; Serene 1984). The only other Indian record of E. rugosus is that of Dev Roy & Nandi (2012) from the Nicobar and Andaman Islands. Given the similarity of the present species with E. exsculptus (Herbst, 1790), previous records of E. exsculptus by Rao et al. (1989) and Bakus (1994) will need to be verified.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD9FFD09CD3FC7FFB73EE33.taxon	materials_examined	Material examined. 1 female (12.1 x 8.4 mm) (DABFUK), Kavaratti, 7 January 2013; 1 male (23.3 x 15.2 mm), (DABFUK) 19 August 2013; 2 females (13.8 x 9.5 mm, 20.1 x 13.2 mm) (DABFUK), Androth, 10 November 2013; 1 ovigerous female (13.5 x 9.8 mm) (DABFUK), Agatti, 9 January 2014; 2 females (10.2 x 7.2 mm, 15.6 x 10.7 mm) (DABFUK), Bitra, 6 May 2015; 1 female (14.0 x 10.5 mm) (DABFUK), Kilton, 18 May 2015; 2 males (13.9 x 9.7 mm, 18.7 x 12.8 mm) (DABFUK), Chetlat, 7 November 2015.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD9FFD09CD3FC7FFB73EE33.taxon	discussion	Remarks. Previously recorded from Lakshadweep by Dev Roy & Nandi (2005) and Suvarna Devi et al. (2014). The species was also observed from Kalpeni, Amini and Minicoy but specimens were not collected.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD9FFD09CD3FF1FFAA9EA8A.taxon	description	(Fig. 4 C)	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD9FFD09CD3FF1FFAA9EA8A.taxon	materials_examined	Material examined. 1 female (17.2 x 10.9 mm) (DABFUK), Minicoy, 3 December 2013.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFD9FFD09CD3FF1FFAA9EA8A.taxon	discussion	Remarks. The present specimen represents the first record of Pseudoliomera granosimana from Lakshadweep. A specimen was also seen in Agatti but not collected. This species has been recorded from many localities in the Indo-West Pacific (Serène 1984), including the Andaman and Nicobar Islands (Dev Roy & Nandi 2012).	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDAFFD39CD3FE12FE38E83A.taxon	description	(Fig. 3 D)	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDAFFD39CD3FE12FE38E83A.taxon	materials_examined	Material examined. 1 female (20.9 x 12.1 mm) (DABFUK), Minicoy, 26 September 2014.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDAFFD39CD3FE12FE38E83A.taxon	discussion	Remarks. The sole specimen from Minicoy is a female, so its identification is tentative until a male specimen can be collected for confirmation. Nevertheless, it is identified as L. laevis on the basis of the absence of a groove separating 2 F + 1 M from 2 M, the relatively shallow and poorly defined furrow separating 3 M from 2 M, and the poorly defined posterior boundaries of 2 L + 3 L and 5 L + 6 L (viz. Serène & Nguyen 1960; Serne 1984). This species is also quite similar to L. tristis, which has previously been recorded from Lakshadweep (Thomas 1969; Dev Roy 2008). The taxonomy of these two species will need to be reviewed. The present specimen of L. laevis represents a new record for Lakshadweep, the species having been previously found in the Andaman and Nicobar Islands by Dev Roy & Nandi (2005).	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDBFFD29CD3FCBEFE16EF2D.taxon	description	(Figs. 2 E, 4 E)	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDBFFD29CD3FCBEFE16EF2D.taxon	materials_examined	Material examined. 1 male (43.5 x 27.2 mm), 1 female (39.5 x 24.8 mm) (DABFUK), Kavaratti, coll. 7 January 2013.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDBFFD29CD3FCBEFE16EF2D.taxon	discussion	Remarks. Previously recorded by Suvarna Devi et al. (2014) from Lakshadweep. Serène (1984) reports it from Mauritius (type locality) and Chagos Archipelago, and there are also records from Sumatra, Japan and Marshall Islands (Serène 1984).	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDBFFD29CD3FBDEFA1CEE05.taxon	description	(Fig. 2 F)	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDBFFD29CD3FBDEFA1CEE05.taxon	materials_examined	Material examined. 1 male (14.0 x 9.3 mm), 1 female (12.9 x 8.9 mm) (DABFUK), Minicoy, 7 August 2014. Remarks. A new record for Lakshadweep. This species is widespread in the Indo-West Pacific (Serène 1984).	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDBFFD29CD3FAB6FBCFEDFD.taxon	materials_examined	Material examined. 1 male (13.3 x 9.4 mm), Kalpeni, 4 January 2013; 1 female (26.7 x 17.0 mm) (DABFUK), Kadamat, 8 June 2013; 2 females (18.8 x 12.6 mm, 22.8 x 15.8 mm), 1 ovigerous female (28.5 x 18.1 mm) (DABFUK), Kadamat, 8 June 2013; 1 male (13.5 x 9.1 mm) (DABFUK), Kadamat, 22 June 2013; 2 males (20.4 x 13.8 mm, 22.7 x 14.8 mm) (DABFUK), Kalpeni, 25 October 2013; 2 males (18.2 x 12.5 mm, 13.5 x 9.6 mm) (DABFUK), Kavaratti, 2 November 2013; 2 males (17.3 x 11.8 mm, 18.6 x 12.5 mm) (DABFUK), Androth, 9 November 2013; 2 females (19.5 x 13.2 mm, 23.3 x 15.1 mm) (DABFUK), Amini, 28 May 2014; 2 males (22.0 x 14.4 mm, 29.1 x 18.3 mm) (DABFUK), Androth, 2 February 2015; 2 females (27.3 x 17.7 mm, 17.2 x 11.4 mm) (DABFUK), Bitra, 8 May 2015; 2 males (39.3 x 24.3 mm, 39.9 x 24.7 mm) (DABFUK), Kilton, 16 May 2015; 1 female (15.5 x 10.2 mm) (DABFUK), Chetlat, 6 November 2015.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDBFFD29CD3FAB6FBCFEDFD.taxon	discussion	Remarks. Previously recorded from Lakshadweep by Alcock (1898) and Rao et al. (1989). This common species was observed on the reefs of the 10 islands in the group, often in large numbers.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDBFFD29CD3FF1FFB94EA66.taxon	materials_examined	Material examined. 3 males (9.1 x 4.9 mm, 10.7 x 5.9 mm, 12.0 x 6.4 mm), 2 females (9.6 x 5.3 mm, 9.8 x 5.3 mm) (DABFUK), Minicoy, 26 September 2014.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDBFFD29CD3FF1FFB94EA66.taxon	discussion	Remarks. This is a distinctive and widespread species in the Indo-West Pacific (Serène 1984). Previously recorded from Lakshadweep Islands (Rao et al. 1989; Venkataraman et al. 2004).	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDBFFD29CD3FD87FCBAE9AA.taxon	materials_examined	Material examined. 1 female (8.9 x 6.8 mm) (DABFUK), Minicoy, 20 January 2015. Remarks. This widespread Indo-West Pacific species (Guinot 1976; Serène 1984) has previously been recorded from Lakshadweep (Borradaile 1902; Thomas 1969).	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDBFFD29CD3F8AEFB56ECF9.taxon	description	(Fig. 2 G)	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDBFFD29CD3F8AEFB56ECF9.taxon	materials_examined	Material examined. 1 male (34.7 x 22.1 mm) (DABFUK), Kavaratti, 10 October 2014. Remarks. A new record from Lakshadweep. This species has a wide distribution and has been reported from the Red Sea and the Indian Ocean to the western Pacific and all the way to the Hawaiian Islands.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDCFFD59CD3F9E9FA01EDF8.taxon	materials_examined	Material examined. 1 female (35.1 x 42.9 mm) (DABFUK), Minicoy, 17 August 2013.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDCFFD59CD3F9E9FA01EDF8.taxon	discussion	Remarks. Previously recorded from Lakshadweep (Thomas 1969). This is the most widely distributed gecarcinid species in the Indo-West Pacific (Türkay 1974), and is the only one known from the western Indian Ocean.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDCFFD59CD3FCDFFB80E8FC.taxon	materials_examined	Material examined. 1 male (15.4 x 10.2 mm) (DABFUK), Kavaratti, 13 October 2014. Remarks. A new record for Lakshadweep. Previously recorded from Tamil Nadu (Henderson 1893; Alcock 1894); Andaman and Nicobar Islands (Alcock 1898; Dev Roy & Nandi 2012).	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDCFFD59CD3FF1FFC85EA66.taxon	materials_examined	Material examined. 1 male (16.7 x 11.1 mm) (DABFUK), Kavaratti, 10 October 2014; 1 male (10.0 x 7.1 mm) (DABFUK), Kalpeni, 4 October 2014; 1 male (8.8 x 6.0 mm) (DABFUK), Minicoy, 24 September 2014.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDCFFD59CD3FF1FFC85EA66.taxon	discussion	Remarks. Paraxanthias notatus was recorded from Lakshadweep by Dev Roy & Nandi (2005) but is otherwise found throughout the Indo-West Pacific (Serène 1984).	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDCFFD59CD3FE13FC98E9CF.taxon	materials_examined	Material examined. 1 male (9.3 x 6.0 mm) (DABFUK), Minicoy, 30 December 2012; 2 males (8.8 x 5.5 mm, 16.0 x 10.3 mm) (DABFUK), Kavaratti, 8 January 2013; 1 male (7.3 x 4.3 mm) (DABFUK), Kalpeni, 28 October 2013; 2 females (13.7 x 8.9 mm, 14.9 x 9.4 mm) (DABFUK), Androth, 9 November 2013; 1 female (11.7 x 7.5 mm), 1 ovigerous female (11.9 x 7.7 mm) (DABFUK), Agatti, 8 January 2014; 2 males (9.2 x 5.8 mm, 11.1 7.2 mm) (DABFUK), Kilton, 20 October 2014.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDCFFD59CD3FE13FC98E9CF.taxon	discussion	Remarks. Previously recorded from Lakshadweep by Rao et al. (1989). This species was also seen at Chetlet, Bitra, Amini and Kadamat but material was not collected.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDCFFD59CD3FBA1FA17EE61.taxon	materials_examined	Material examined. 2 males (45.9 x 30.5 mm, 67.6 x 45.7 mm) (DABFUK), Kavaratti, 8 January 2013; 2 males (30.8 x 20.9 mm, 43.5 x 29.8 mm) (DABFUK), Kadamat, 20 June 2013; 1 male (31.3 x 21.7 mm), 1 female (66.6 x 44.3 mm) (DABFUK), Minicoy, 25 September 2014; 2 females (41.9 x 20.4 mm, 51.2 x 39.8 mm) (DABFUK), Kilton, 19 October 2014; 2 females (52.6 x 33.5 mm, 65.8 x 44.3 mm) (DABFUK), Kalpeni, 2 October 2014; 1 juvenile male (13.8 x 9.4 mm) (DABFUK), Kavaratti, 17 November 2015.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDCFFD59CD3FBA1FA17EE61.taxon	discussion	Remarks. Previously recorded from Lakshadweep (Alcock 1898; Thomas 1969; Suvarna Devi et al. 2014; Kumaralingam et al. 2017). This is a common poisonous species widespread in the Indo-West Pacific (Serène 1984).	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDCFFD49CD3F804FC4DEA8A.taxon	materials_examined	Material examined. One male (69.6 x 54.5 mm) (DABFUK), Kilton, 20 October 2014.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDCFFD49CD3F804FC4DEA8A.taxon	discussion	Remarks. This species was observed in all 10 islands, viz. Agatti, Kavaratti, Minicoy, Amini, Chetlat, Androth, Bitra, Kilton, Kalpeni and Kadamat. It can, however, be confused with the similar G. tenuicrustatus (Herbst, 1783), but easily separated by a number of frontal margin, cheliped and male pleonal differences: the frontal margin is smooth and entire (serrated in G. tenuicrustatus); the chelae have a purple tinge, and the inner angle of the cheliped carpus has an acutely triangular tooth (chelae dark brown, and inner angle of cheliped carpus has a broader tooth with a subtruncate base in G. tenuicrustatus); the sixth pleonal somite is shorter than the fifth (sixth pleonal somite longer than the fifth in G. tenuicrustatus) (viz. Banerjee 1960; Ng 1998).	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDDFFD49CD3FE3FFADEE93E.taxon	materials_examined	Material examined. 1 male (7.3 x 5.5 mm) (DABFUK), Kavaratti, 10 October 2014. Remarks. Previously recorded from Lakshadweep by Borradaile (1903 a) and Dev Roy & Nandi (2005).	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDDFFD49CD3F91CFC48ECF9.taxon	description	(Fig. 2 H)	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDDFFD49CD3F91CFC48ECF9.taxon	materials_examined	Material examined. 1 megalopa (6.9 x 5.1 mm) (DABFUK), Kavaratti, 11 October 2014.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDDFFD49CD3F91CFC48ECF9.taxon	discussion	Remarks. The distinctive characters identifying the megalopa as a species of Percnon are the carapace outline, the three long rostral spines on the anterior of the carapace, and the short spines on the ventral (flexor) margin of the ambulatory dactyli. It resembles the megalopa of the Atlantic species, Percnon gibbesi (H. Milne Edwards, 1853) (cf. Paula & Hartnoll 1989: 31, fig. 11), but the carapace of the present specimen is much narrower and the legs are relatively longer. It is difficult to ascertain which of the two preceding, sympatric species of Percnon it belongs to, as megalopal differences between the species have not been documented.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDDFFD49CD3FB24FDB1EE71.taxon	materials_examined	Material examined. 2 males (8.6 x 7.0 mm, 14.1 x 12.8 mm) (DABFUK), Minicoy, 16 August 2013; 1 female (10.9 x 10.8 mm) (DABFUK), Kalpeni, 20 October 2015.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDDFFD49CD3FB24FDB1EE71.taxon	discussion	Remarks. Kumaralingam et al. (2017) first recorded this species from Lakshadweep, which has a wide range in the Indian Ocean (Crosnier 1965).	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDDFFD49CD3FA18FD68ED6D.taxon	materials_examined	Material examined. 1 female (12.9 x 12.6 mm) (DABFUK), Kalpeni, 4 October 2014.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDDFFD49CD3FA18FD68ED6D.taxon	discussion	Remarks. Previously recorded from Lakshadweep (Alcock 1900 b; Thomas 1969; Dev Roy 2008; Kumaralingam et al. 2017). This is a widespread species in the Indo-West Pacific (Crosnier 1965). Specimens were also observed on Kavaratti and Minicoy but not collected.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDDFFD49CD3FD5FFD35EF12.taxon	materials_examined	Material examined. 1 male (36.6 x 35.4 mm) (DABFUK), Kavaratti, 13 October 2014; 1 female (21.2 x 18.7 mm) (DABFUK), Kilton, 20 October 2014; 2 females (20.5 x 19.2 mm, 13.02 x 12.98 mm) (DABFUK), Androth, 2 February 2015.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDDFFD49CD3FD5FFD35EF12.taxon	discussion	Remarks. This species was also seen in the islands of Amini, Bitra, Chetlat, Kadamat, Kalpeni and Minicoy, but specimens were not collected. Schubart & Ng (2000) showed that the better known name, Plagusia tuberculata Lamarck, 1818, is a junior synonym of P. squamosa (Herbst, 1790). The species has a wide Indo-West Pacific distribution (Crosnier 1965; Schubart & Ng 2000).	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDEFFD79CD3FF65FAB3EAD5.taxon	materials_examined	Material examined. 1 male (9.2 x 8.4 mm) (DABFUK), Kavaratti, 10 October 2014.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDEFFD79CD3FF65FAB3EAD5.taxon	discussion	Remarks. This is a common supralittoral species widely distributed in the Indo-West Pacific. The species was also seen at Kalpeni but specimens were not collected. Previously recorded from Lakshadweep (Alcock 1900 b; Dev Roy 2008; Kumaralingam et al. 2017). The species is better known as Metasesarma rousseauxi H. Milne Edwards, 1853, but Ng & Schubart (2003) showed that Sesarma obesa Dana, 1851, is a senior subjective synonym.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDFFFD69CD3FDEBFA13EFA3.taxon	materials_examined	Material examined. 2 males (40.0 x 34.9 mm, 64.6 x 52.7 mm) (DABFUK), Kadamat, 28 June 2013; 1 female (67.7 x 59.2 mm) (DABFUK), Minicoy, 20 August 2013.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDFFFD69CD3FDEBFA13EFA3.taxon	discussion	Remarks. Ocypode ceratophthalmus (Pallas, 1772) as currently understood is a species complex, currently under study by P. J. F. Davie, H. - T. Shih and P. K. L. Ng, with three genetically distinct forms present in the Indian Ocean. Two are the commonly recognised white-and-maroon morphs and are morphologically close, but can be separated by details of the stridulatory apparatus on its chela. One white-maroon morph represents Ocypode ceratophthalmus sensu stricto, which occurs on islands as well as continental landmasses as well as in most of Southeast Asia and East Asia. The second white-maroon morph is known mostly from isolated islands and appears allied to O. pallidula Hombron & Jacquinot, 1846. Whether it is a new species or there is older name now listed under its extensive synonymy will need to be established. The third morph is the most distinct, with a prominent green to greenishyellow coloration in life and a diagnostic stridulatory structure on its chela. This green morph is almost certainly a new species and is known only from islands in the Indian Ocean (see also Orchard 2012; Ma et al. 2018). All the material observed in the islands were of the green morph. The species was also observed on eight islands: viz; Agatti, Kavaratti, Amini, Chetlat, Androth, Bitra, Kilton and Kalpeni but specimens were not collected.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDFFFD69CD3FF1EFF4FEA8B.taxon	materials_examined	Material examined. 1 male (9.2 x 8.2 mm) (DABFUK), Kavaratti, 4 January 2014.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
03BA87FBFFDFFFD69CD3FF1EFF4FEA8B.taxon	discussion	Remarks. A new record from Lakshadweep. The identity of the Indian material referred to this species (see Trivedi et al. 2018) requires confirmation. There are slight differences in the chela and gonopods of material from the Indian Ocean and the east Pacific (where P. pictum s. str. occurs), suggesting there may be more than one species.	en	Devi, S. Suvarna, Mendoza, Jose C. E., Ravinesh, R., Idress Babu, K. K., Kumar, A. Biju, Ng, Peter K. L. (2019): On a collection of brachyuran crabs from Lakshadweep, Indian Ocean (Crustacea: Decapoda: Brachyura). Zootaxa 4613 (3): 477-501, DOI: 10.11646/zootaxa.4613.3.4
