identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03BADF229332FFF8FF22368DFD5E5CCF.text	03BADF229332FFF8FF22368DFD5E5CCF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heteroscelis Latreille 1829	<div><p>Heteroscelis Latreille, 1829</p><p>Heteroscelis Latreille, 1829: 194 (not Latreille, 1828: 574, nor Latreille, 1829: 18); Laporte 1833: 86; Stål 1870: 31; Distant 1880: 33; Distant 1900: 55; Schouteden 1907: 18; Kormilev 1951: 40; Pirán 1954: 19; Pirán 1961: 84; Thomas 1992: 45; Grazia &amp; Schwertner 2011: 713; Grazia et al. 2015: 686; Cazorla 2021: 10; Rider &amp; Swanson 2021: 7; Castro-Huertas et al. 2022: 9.</p><p>Phyllocheirus Spinola, 1837: 306; Kirkaldy 1909: 30; Pirán 1961: 87 (syn.).</p><p>Phyllochirus Amyot &amp; Serville, 1843: 80 (emendation); Stål 1862: 93; Stål 1870: 31 (syn.); Kirkaldy 1909: 30.</p><p>Agerrus Stål, 1859: 434; Stål 1862: 93 (syn.); Schouteden 1907: 19; Kirkaldy 1909: 30.</p><p>Bodetria Walker, 1867: 119; Walker 1868: 528; Stål 1870: 31 (syn.); Distant 1900: 63; Schouteden 1907: 19; Kirkaldy 1909: 30.</p><p>Heterosceloides Schouteden, 1907: 19; Kirkaldy 1909: 30 (syn.); McDonald 1966.</p><p>Type species: Heteroscelis servillii Laporte, 1833, by subsequent monotypy (ICZN 1999: Article 69.3)</p><p>Diagnosis. Head elongated, with lateral post-ocular projections; proportion of labiomeres: I&lt;II&gt;III&lt;IV; pronotal depression anterior to the cicatrices forming a collar; humeral angles laterally projected, bifid, with the anterior portion longer than the posterior; pro-, meso- and metasternum covered by small setae; prosternum flat, mesosternum with a low longitudinal carina dilated anteriorly and posteriorly, metasternum slightly elevated with a shallow longitudinal groove.</p><p>Redescription</p><p>BODY: Elongated oval, with variable color patterns.</p><p>HEAD: Rectangular, longer than wide, longitudinally convex, slightly punctured; mandibular plates longer than clypeus, with the anterior margin rounded; bucculae low, rounded at apex and evanescent posteriorly; eyes separated from the anterolateral angles of pronotum by at least half the eye diameter; post-ocular lateral triangular projection contiguous to the eyes; antennomere I shorter and wider than II, which is thinner than the others; antennomeres III, IV and V similar in length and more setose than the others.</p><p>THORAX: Pronotum trapezoidal, uniformly and deeply punctured, twice as wide as long; scutellum uniform and shallowly punctured; frenal constriction width more than half the basal width, postfrenal lobe with parallel margins, broadly rounded at apex; coria with posterior margin sinuous; mesopleural evaporatoria occupying the entire posterior margin and the anterolateral mesopleural angle; metapleural evaporatoria surrounding the disc-shaped peritreme, occupying more than half of the metapleural width; ostiole guttiform in ventral view, laterally oriented; peritremal furrow extending more than half the length of the peritreme; dorsal and ventral foretibial expansions present, the dorsal expansion wider than the ventral.</p><p>ABDOMEN: Uniformly punctured; mid tubercle on urosternite III; posterior angles of urosternites III to VII ending in minute spine; male bearing ventrolateral glandular setose patches, extending from near the anterior margin of urosternite V to the anterior margin of urosternite VII.</p><p>Male</p><p>GENITALIA: Pygophore bowl-shaped, with setae on the entire surface, ventral rim slightly sinuous; inferior layer excavated; posterolateral angles rounded; segment X trapezoidal, with rounded apex, ventrally directed, dorsally sclerotized, setose on apex; pseudoclaspers concave, sculptured, each placed between the paramere and the lateral portion of the dorsal rim; parameres sculptured, head triangular, spatulate, occupying in dorsal view most of the space between the segment X and the posterolateral angles. Phallus: phallotheca divided in a basal theca and thecal shield; ejaculatory reservoir contained inside basal theca; basal foramen elliptical, reinforced by basal plates; thecal shield longer and wider than basal theca; vesica oval in dorsal and lateral view; ductus seminis with uniform diameter, running between filaments of vesica, ending on a secondary gonopore, ventrally directed; conjuntiva with: two ventral lobes with sclerotized apex, and one dorsal lobe; all lobes posteriorly directed.</p><p>Female</p><p>GENITALIA: Valvifers VIII longer than wide, subtriangular, posterior margins sinuous, sutural margins straight, juxtaposed; laterotergites VIII subtriangular; exposed portion of valvifer IX trapezoidal, wider than long, juxtaposed to the laterotergites IX; exposed portion of laterotergites IX digitiform, attaining the mediotergite VIII; segment X trapezoidal with posterior margin straight; thickening of vaginal intima elongated; pars intermedialis small, narrower than the median duct of vesicular area; capsula seminalis longer and wider than pars intermedialis.</p><p>Distribution: United States of America (Texas), Mexico, Guatemala, Honduras, El Salvador, Costa Rica, Panama, Colombia, Ecuador (new record), Trinidad and Tobago, Venezuela, Guyana, Surinam, French Guyana, Brazil, Peru, Bolivia, Paraguay and Argentina (Fig. 1).</p></div>	https://treatment.plazi.org/id/03BADF229332FFF8FF22368DFD5E5CCF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sampaio, Vinicius Gomes;Roell, Talita;Campos, Luiz Alexandre	Sampaio, Vinicius Gomes, Roell, Talita, Campos, Luiz Alexandre (2023): Revision of Heteroscelis Latreille, 1829 (Heteroptera: Pentatomidae: Asopinae) with the description of a new species from Colombia. Zootaxa 5278 (3): 401-438, DOI: 10.11646/zootaxa.5278.3.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.3.1
03BADF229335FFFFFF2232D7FA385D78.text	03BADF229335FFFFFF2232D7FA385D78.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heteroscelis Latreille 1829	<div><p>Key to the Species of Heteroscelis Latreille</p><p>1 Dorsal foretibial expansion wider than the femur............................................................ 2</p><p>- Dorsal foretibial expansion narrower than the femur......................................................... 4</p><p>2 Mandibular plates as long as or slightly surpassing clypeus; urosternite III tubercle concolor, or completely yellow or orange, from apex to the posterior margin of the urosternite.......................................................... 3</p><p>- Mandibular plates distinctly surpassing clypeus; urosternite III tubercle yellow or orange from apex to the middle of the urosternite........................................................................... H. servillii Laporte</p><p>3 Corium with a circular yellow spot near the posterior margin; urosternites VI and VII with yellow spots; urosternite III tubercle yellow or orange................................................................... H. bimaculata (Walker)</p><p>- Body entirely metallic-blue with a triangular red or orange spot on scutellum......................... H. lepida (Stål)</p><p>4 Pronotum and scutellum with a yellow midline............................................. H. robustus Thomas</p><p>- Pronotum and scutellum red to orange................................ H. vandembergi Sampaio &amp; Campos, sp.nov.</p></div>	https://treatment.plazi.org/id/03BADF229335FFFFFF2232D7FA385D78	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sampaio, Vinicius Gomes;Roell, Talita;Campos, Luiz Alexandre	Sampaio, Vinicius Gomes, Roell, Talita, Campos, Luiz Alexandre (2023): Revision of Heteroscelis Latreille, 1829 (Heteroptera: Pentatomidae: Asopinae) with the description of a new species from Colombia. Zootaxa 5278 (3): 401-438, DOI: 10.11646/zootaxa.5278.3.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.3.1
03BADF229335FFF4FF223095FCFB5DD5.text	03BADF229335FFF4FF223095FCFB5DD5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heteroscelis servillii Laporte 1833	<div><p>Heteroscelis servillii Laporte, 1833</p><p>(Figs. 1–14)</p><p>Heteroscelis servillii Laporte, 1833: 86; Stål 1870: 31; Kirkaldy 1909: 30 (syn.).</p><p>Phyllocheirus servillei: Spinola 1837: 306</p><p>Phyllochirus servillei: Amyot &amp; Serville 1843: 80 (emen); Stål 1862: 93; Stål 1870: 31 (syn.); Kirkaldy 1909: 30.</p><p>Agerrus remipes Stål, 1859: 434; Stål 1862: 93 (syn.); Schouteden 1907: 19; Kirkaldy 1909: 30.</p><p>Bodetria brenthoides Walker, 1867: 119; Stål 1870: 31 (syn.); Distant 1900: 63; Schouteden 1907: 19; Kirkaldy 1909: 30.</p><p>Bodetria chrysoclora Walker, 1868: 528; Stål 1870: 31 (syn.); Distant 1900: 63; Schouteden 1907: 19; Kirkaldy 1909: 30.</p><p>Bodetria indecora Walker, 1868: 528; Stål 1870: 31 (syn.); Distant 1900: 63; Schouteden 1907: 19; Kirkaldy 1909: 30.</p><p>Heteroscelis brenthoides: Schouteden 1907: 19; Kirkaldy 1909: 30 (syn.).</p><p>Heteroscelis servillei: Distant 1900: 63; Schouteden 1907: 19; Pirán 1961: 87; Thomas 1992: 46; Grazia &amp; Schwertner 2011: 713; Grazia et al. 2015: 686; Silva et al. 2018: 404; Lupoli 2019: 80; Cazorla 2021: 10; Castro-Huertas et al. 2022: 9; Roell et al. 2023: 28, 29, 47.</p><p>Phyllocheirus brenthoides: Kirkaldy 1909: 30 .</p><p>Phyllocheirus servillii: Kirkaldy 1909: 30; Pirán, 1961: 87 (syn.).</p><p>Heteroscelis zischkai Kormilev, 1951: 40; Pirán 1961: 88; Thomas 1992: 46 (syn.).</p><p>Heteroscelis dureti Pirán, 1954: 19; Thomas 1992: 46 (syn.).</p><p>Heteroscelis bergi Pirán, 1954: 20; Thomas 1992: 46 (syn.).</p><p>Heteroscelis carcavalloi Pirán, 1961: 96; Thomas 1992: 46 (syn.).</p><p>Heteroscelis amazonica Pirán, 1961: 98; Thomas 1992: 46 (syn.).</p><p>Types examined</p><p>SURINAM. Agerrus remipes Stål, 1859; female syntype; labels: “Surinam.”, “Stål”, “ Agerrus remipes Stål. Typ. ”, “111”, “TYPUS”, “NHRS—GULI 000096153”; (NHRS). Photo examined (Fig. 3).</p><p>BRAZIL. Heteroscelis amazonica Pirán, 1961; female holotype; labels: “Ig do Passarinho Manaus. Am. 24.II.55. Costa Leite e N. Cerqueira—cols. 07. 9/59”, “ Heteroscelis amazonica Pirán 1960 ”, “ HOLOTYPUS ”; (MNRJ) (Fig. 4).</p><p>BRAZIL. Heteroscelis carcavalloi Pirán, 1961; female alotype; labels: “Sítio Vieiralves Manaus—Am. 21.Xll.65. Elias e Rappa—cols. 02. 9/59”, “ Heteroscelis carcavalloi Pirán 1960 ”, “ ALLOTYPUS ”; (MNRJ).</p><p>BRAZIL. Bodetria brenthoides Walker, 1867; female holotype; labels: “Amazon St. Paul / 60 32” “Type”, “ Holotype ”, “1. Bodetria brenthoides ”; “NHMUK 010592390”; (NHMUK). Photo examined (Fig. 5).</p><p>BRAZIL. Bodetria chrysoclora Walker, 1868; male holotype; labels: “Braz / 62 57” “Type”, “ Holotype ”, “ Bodetria chrysoclora ”; “NHMUK 010592391”; (NHMUK). Photo examined (Fig. 6).</p><p>BRAZIL. Bodetria indecora Walker, 1868; male holotype; labels: “Braz / 62 57” “Type”, “ Holotype ”, “ Bodetria indecora ”; “NHMUK 010592393”; (NHMUK). Photo examined (Fig. 7).</p><p>BOLIVIA. Heteroscelis carcavalloi Pirán, 1961; male holotype; labels: “ HOLOTYPUS ”, “Yungas del Palmar. IV–954. A. Martinez.”, “ Heteroscelis carcavalloi Pirán, 1960 .”; (MACN). Photo examined (Fig. 8).</p><p>BOLIVIA. Heteroscelis zischkai Kormilev, 1951; female alotype; labels: “ BOLIVIA, Region Chaparé, Dep. Cochabamba (400 mtr), 30. VIII. 1951. Dirings.”. Heteroscelis zischkai Kormilev. Museo Argentino de Ciencias Naturales. ”. “ ALLOTYPE ”; (MACN) (Fig. 9).</p><p>PARAGUAY. Heteroscelis dureti Pirán, 1954; female holotype; labels: “ Heteroscelis dureti Det. Pirán. 952”, “ HOLOTYPUS ”, “ Paraguay, Villarrica, Schade col. 15.IV.46”; (MACN). Photo examined (Fig. 10).</p><p>ARGENTINA. Heteroscelis bergi Pirán, 1954; male holotype; labels: “Missiones—Argentina. Dep. Concep.— Sta. Maria. M.J. Viana”, “53.484”, “ HOLOTYPUS ”, “ Heteroscelis bergi Det. Pirán. 952”; (MACN). Photo examined (Fig. 11).</p><p>Other material examined. VENEZUELA, Guayana: (Conuco, Rio Moroco), 3 males, 18/ 19.I.1972 G. &amp; E. Scherer (NMPC); Zulia: (Kasmera, 9.94907, -72.75292), 1 female, 20.IX.1961 F. Fernandez Y. &amp; C. J. Rosales (UFRG); Cx. Panamexicana El Vigia Coloneito: (Km 8.32611, -72.08742), 1 female, 6.I.1955. F. Fernandez Y. &amp; C. J. Rosales (UFRG); Aragua: (El Limón, 10.30589, -67.63212), 1 male, 2.III.1951 F. Fernandes (UFRG); Aragua: (Maracay, 10.23535, -67.59113), 1 female, 2.V.1953 N. Angeles (UFRG). TRINIDAD AND TOBAGO: 2 males, XII.1953 G. u. Helga Frey. (NMPC). BRAZIL, 1 female, 11.IV.1962 Cerq. (UFRG); Pará: (Belém Mocambo, - 1.45583, -48.50444), 1 female, 6.II.1979 P Tadeu (UFRG); (Peixe-boi, -1.10168, -47.27259), 1 male, 12.IV.1977 P Waldir. (UFRG); (Serra Norte Manganês -6.01748, -50.30386), 1 female, 22.X.1984 T. Pimentel; Amazonas: (Manaus, CEPLAC, -2.8905, -59.9576), 1 female, 15.VII.1977 (UFRG); (Serra dos Porcos, 0.416667, -69.366667), 1 male, 25.II.1977 Franklim (INPA); (Manaus, CEPLAC, Am 010 Km 31, -2.8905, -59.9576), 1 male, 7.V.1988 Albuquerque (UFRG); (Manaus, -3.04361, -60.01282), 1 male, 5.V.1976 Paraluppi (UFRG); (Manaus, Br 174 Km 41, Agricultura Geológica -2.6770, -60.04668), 1 male, 15.VI.1981 C.A. Fonseca (INPA); (Itacoatiara, -3.1373, - 58.4418), 1 female, Dirings (MCNZ); Amapá: (Serra do Navio, 1.65803, -52.28195), 1 female, 13.VII.1961 AP. J &amp; B. Bechyné (UFRG); Rio Grande do Norte: (Natal, -5.8101, -35.22674), 1 male, I.1952 M. Alvarenga (UFRG); Pernambuco: (Paudalho, Acerolândia, -7.850, -35.250. 160m), 1 male, II.2016 Grossi &amp; Parizotto (CERPE); (Paudalho, Acerolândia, -7.850, -35.250. 160m), 1 female, XI.2017 Grossi &amp; Parizotto (CERPE); Mato Grosso do Sul: (Bonito, Praia da Figueira -21.2972, -56.5005), 1 female, 01.III.2009 Grossi &amp; Parizotto (DZUP); Rio Grande do Sul: (Cerro Azul -27.7403, -51.5199), 1 female, I.1931 Pe. Buck leg. (MCNZ); ECUADOR, Amazonia: 1 female, II.1998 lgt. Bezdek (NMPC); Napo: (Tiputini Biodiversity Station, 216m, -0.6379, -76.1495), 1 male, 06.II.1999 T.L. Erwin, et al. (USNM); (Tiputini Biodiversity Station, 216m, -0.6379, -76.1495), 1 female, 22.X.1998 T.L. Erwin, et al. (USNM); PERU, Pucalipa: (Rio Ucayali, -8.4084, -74.6058), 1 female, Dirings (MCNZ); BOLIVIA, Cochabamba (Chapare, 400m -16.80922, -65.71926), 1 female, 07.IX.1951 Dirings (MACN).</p><p>Records retrieved from iNaturalist</p><p>TRINIDAD AND TOBAGO, San Juan-Laventille: (10.605137, -61.42763), 14.II.2021.</p><p>BRAZIL, Amazonas: (Manaus, -3.095784, -59.989879), 13.VIII.2020.</p><p>BRAZIL, Espírito Santo: (S„o Gabriel de Palha, -18.906837, -40.478105), II.2014.</p><p>BRAZIL, S„o Paulo: (Campinas, -22.819691, -47.064677), 16.II.2023.</p><p>BRAZIL, Rio de Janeiro: (Cabo Frio, -22.886901, -42.026649), 8.I.2017.</p><p>Diagnosis. Mandibular plates distinctly longer than clypeus, converging apically; head length anterior to the eyes more than 2/3 of the total length of the head (Fig. 2, D); urosternite III tubercle yellow or orange at apex, and surpassing the posterior margin of metasternum (Fig. 2, B).</p><p>Redescription</p><p>BODY: General color ranging from brown and dark red to iridescent hues of green, yellow, blue and purple, with or without yellow spots on the pronotum, scutellum and coria (more commonly observed in non-iridescent specimens).</p><p>HEAD: Labium almost reaching the metasternum (Fig. 2, B); proportion of antennomeres: I&lt;II&lt;III ≅ IV ≅ V.</p><p>THORAX: Coria longer than scutellum, not reaching the posterior margin of connexival segment VI, uniformly punctured; membrane dark and surpassing posteriorly the abdomen; metapleural evaporatoria surpassing posteriorly the anterior limit of coxae; peritreme reaching or surpassing half the pleural width; apex of peritreme rounded, posterior margin a little more arched than the anterior, both convex (Fig. 2, E); profemora with rudimentary ante-apical spine; foretibial length on average 3.5x larger than the width of the tibial dorsal expansion (Fig. 2, F).</p><p>ABDOMEN: Posterior angles of urosternites III to VII ending in minute spine, more developed in VII (Fig. 2, B).</p><p>Male:</p><p>MEASUREMENTS (n=13): Head length 1.76 ± 0.28 (1.30–1.84); width 1.29 ± 0.07 (1.20–1.50); pronotum length 2.00 ± 0.26 (1.50–2.40); width 4.21 ± 0.26 (3.80–4.50); scutellum length 2.38 ± 0.16 (2.08–2.60); width 2.30 ± 0.13 (2.08–2.56); length of antennomeres: I (n=11) 0.39 ± 0.03 (0.30–0.40), II (n=11) 0.71 ± 0.09 (0.60 –0.90), III (n=11) 0.95 ± 0.10 (0.80–1.04), IV (n=11) 1.03 ± 0.14 (0.72–1.20); V (n=8) 1.09 ± 0.20 (0.72–1.40); length of labiomeres: I (n=8) 0.67 ± 0.08 (0.60–0.80), II (n=8) 0.87 ± 0.07 (0.80–1.00), III (n=8) 0.45 ± 0.07 (0.40–0.60), IV (n=8) 0.68 ± 0.09 (0.60–0.90); width of foretibal expansion (n=10) 0.59 ± 0.07 (0.48–0.72); width of abdomen (n=11) 3.21 ± 0.18 (2.96–3.50); total length 7.04 ± 0.65 (6.10–8.30).</p><p>GENITALIA: Pygophore (Fig. 12): setae denser on posterolateral angles and on medial portion of dorsal rim; dorsal rim sinuous, medially concave; ventral rim slightly sinuous, medially concave; segment X with surface corrugated and with a membranous longitudinal line; head of parameres lateroventrally directed in posterior view. Phallus (Fig. 13): conjunctival dorsal lobe with one pair of long projections and one single forked central projection, ventral lobes with two projections each.</p><p>Female:</p><p>MEASUREMENTS (n=15): Head length (n=11) 2.06 ± 0.38 (1.40–2.60); width (n=11) 1.47 ± 0.20 (1.20– 1.90); pronotum length (n=13) 2.45 ± 0.26 (1.92–2.80); width (n=13) 4.83 ± 0.42 (3.92–5.60); scutellum length 2.88 ± 0.32 (2.24–3.40); width 2.56 ± 0.40 (1.84–3.00); length of antennomeres: I (n=12) 0.42 ± 0.06 (0.30–0.50), II (n=12) 0.79 ± 0.15 (0.56 –1.10), III (n=12) 1.04 ± 0.21 (0.60–1.30), IV (n=8) 1.22 ± 0.39 (0.80–2.00); V (n=5) 1.03 ± 0.18 (0.96–1.30); length of labiomeres: I (n=9) 0.77 ± 0.11 (0.60–0.90), II (n=9) 1.02 ± 0.16 (0.70–1.30), III (n=9) 0.50 ± 0.10 (0.40–0.70), IV (n=9) 0.85 ± 0.14 (0.60–1.00); width of foretibal expansion (n=11) 0.72 ± 0.09 (0.56–0.90); width of abdomen 3.70 ± 0.33 (3.04–4.30); total length (n=8) 8.28 ± 1.05 (6.88–9.50).</p><p>GENITALIA (Fig. 14): Valvifers VIII posterior margin bisinuous, slightly projected over the basal angles of laterotergites IX; laterotergites VIII wider than long, with spiracles equidistant from the basal angle and the lateral margin; valvulae IX with 1+1 bean-shaped secondary thickenings; vesicular area half the length of the proximal part of the ductus receptaculi; capsula seminalis oval.</p><p>Distribution: Colombia, Ecuador (new record), Trinidad and Tobago, Venezuela, Guyana, Surinam, French Guyana, Brazil, Peru, Bolivia, Paraguay and Argentina (Fig. 1).</p></div>	https://treatment.plazi.org/id/03BADF229335FFF4FF223095FCFB5DD5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sampaio, Vinicius Gomes;Roell, Talita;Campos, Luiz Alexandre	Sampaio, Vinicius Gomes, Roell, Talita, Campos, Luiz Alexandre (2023): Revision of Heteroscelis Latreille, 1829 (Heteroptera: Pentatomidae: Asopinae) with the description of a new species from Colombia. Zootaxa 5278 (3): 401-438, DOI: 10.11646/zootaxa.5278.3.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.3.1
03BADF229321FFEFFF2233CFFC7C59CE.text	03BADF229321FFEFFF2233CFFC7C59CE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heteroscelis lepida (Stal 1862)	<div><p>Heteroscelis lepida (Stål, 1862)</p><p>(Figs. 1, 15–19)</p><p>Phyllochirus lepidus Stål, 1862: 93; Stål 1870: 31 (syn.).</p><p>Bodetria scutellaris Walker, 1868: 529; Stål 1870: 31 (syn.); Distant 1880: 33, 1900: 63; Thomas 1992: 48.</p><p>Heteroscelis lepida: Stål 1870: 31; Distant 1880: 33, 1900: 63; Pirán 1961: 88; Thomas 1992: 48; Brailovsky &amp; Mayorga 1994: 34; Arismendi &amp; Thomas 2003: 224; Cambra et al. 2018: 5; Lupoli 2019: 80; Cazorla 2021: 10; Rider &amp; Swanson 2021: 7; Castro-Huertas et al. 2022: 9; Roell et al. 2023: 77.</p><p>Heterosceloides lepida: Schouteden 1907: 20; Kirkaldy 1909: 30; Pirán 1961: 88 (syn.).</p><p>Types examined</p><p>MEXICO. Phyllochirus lepidus Stål, 1862; female holotype; “ México. Coll. Signoret”, “ lepidus set. Stål”, “ lepidus ”, “ Heterosceloides lepidus Stål ”; (NHMW). Photo examined (Fig. 16).</p><p>MEXICO. Bodetria scutellaris Walker, 1868; male holotype; “ Holotype ”, “Type”, “Vera cruz / 5466”, “ Bodetria scutellaris ”; “NHMUK 010592395”; (NHMUK). Photo examined (Fig. 17).</p><p>Other material examined. PANAMA. Panama: (Canal Zone, Madden Forest, 9.05, -79.37), 1 male, 8.I.1971 H. Stockwell (STRI), Smithsonian Tropical Research institute, photo examined. VENEZUELA, Falcon: (Las dos Bocas, 11.30129, -69.41226), 1 female, 29.I.1967 R. Cassares (UFRG). COLOMBIA, Antioquia: (Santa Fé de Antioquia, Hacienda Cotové - UNAL, 600 m. 6.5316, -75.8260), 1 female, 6.X.2000 A. Botero, et al. (UNAB). 1 female, no data (UFRG).</p><p>Records retrieved from iNaturalist</p><p>MEXICO, Veracruz: (Puente Nacional, 19.34434, -96.464912), 31.X.2016.</p><p>MEXICO, Veracruz: (Puente Nacional, 19.343728, -96.464263), 25.IX.2017.</p><p>COSTA RICA, Puntarenas: (Puntarenas, 10.275075, -84.812452) 25.V.2017.</p><p>COLOMBIA, Sucre: (Galeras, 9.147859, -74.978428), 18.II.2017.</p><p>VENEZUELA, Falcon: (Miranda, 11.41002, -69.679581), 5.XII.2022.</p><p>Diagnosis. Body iridescent-blue, with a large triangular red or orange spot on scutellum; vesicular area of female genitalia subequal in length to the proximal part of the ductus receptaculi.</p><p>Redescription</p><p>HEAD: Mandibular plates equal to or slightly surpassing clypeus, anteriorly rounded, separated at apex; head length anterior to the eyes equal to 2/3 of the total length of the head (Fig. 15, D); labium reaching the metasternum; proportion of labiomeres: I&lt;II&gt;III&lt;IV; proportion of antennomeres: I&lt;II&lt;III&lt;IV ≅ V.</p><p>THORAX: Scutellum with a large triangular red or orange spot, covering the entire frenal lobe and the central portion of the postfrenal lobe; coria longer than the scutellum, not reaching the posterior margin of the connexival segment VI, uniformly punctured; membranes dark, surpassing posteriorly the abdomen; metapleural evaporatoria not surpassing posteriorly the anterior limit of coxae; peritreme disc-shaped, surpassing half the pleural width; apex of peritreme rounded, posterior margin arched, convex, anterior margin almost straight (Fig. 15, E); foretibial length on average 3.5x larger than the width of the dorsal tibial expansion (Fig. 15, F).</p><p>ABDOMEN: Uniformly punctured; urosternite III tubercle not surpassing the posterior margin of metasternum; posterior angles of urosternites III to VII ending in minute spine (Fig. 15, B).</p><p>Male</p><p>GENITALIA (Fig. 18): Pygophore with dorsal rim concave; ventral rim slightly sinuous, medially concave; segment X with surface corrugated; head of parameres laterodorsally directed in posterior view.</p><p>Remarks: The only description of the internal male genitalia was made by McDonald (1966). Based on his description and illustrations (McDonald 1966: 121, figs. 300–302) we interpret the phallus having the conjunctival dorsal lobe (conjunctival appendages sic McDonald) with one pair of projections, and the ventral lobes with three projections each.</p><p>Female</p><p>MEASUREMENTS (n=3). Head length 1.24 ± 0.22 (1.00–1.44); width 1.34 ± 0.07 (1.26 –1.40); pronotum length 1.75 ± 0.21 (1.57–2.00); width 3.88 ± 0.24 (3.70–4.16); scutellum length 2.25 ± 0.27 (2.01–2.56); width 2.18 ± 0.11 (2.10–2.32); length of antennomeres: I 0.33 ± 0.04 (0.30–0.37), II 0.62 ± 0.15 (0.50–0.80), III 0.81 ± 0.19 (0.69–1.04), IV 1.09 ± 0.17 (0.90–1.20); V 1.00 ± 0.11 (0.88–1.12); length of labiomeres: I 0.53 ± 0.14 (0.37–0.64), II 0.71 ± 0.08 (0.63–0.80), III 0.40 ± 0.09 (0.30–0.48), IV 0.61 ± 0.11 (0.50–0.72); width of foretibal expansion 0.65 ± 0.26 (0.50–0.96); width of abdomen 3.41 ± 0.36 (3.20–3.84); total length 6.31 ± 0.64 (5.67–6.96).</p><p>GENITALIA (Fig. 19): Valvifers VIII posterior margin shallowly concave over the plates of sternite IX, truncate over the base of laterotergites VIII; laterotergites VIII wider than long, with spiracles close to the lateral margin; segment X with the posterior margin straight; valvulae IX with 1+1 bean-shaped secondary thickenings with the proximal margin laterally elongated; capsula seminalis oval.</p><p>Distribution: United States of America (Texas), Mexico, Guatemala, Honduras, El Salvador, Costa Rica (new record), Panama, Colombia, Ecuador, Venezuela, French Guyana. (Fig. 1).</p></div>	https://treatment.plazi.org/id/03BADF229321FFEFFF2233CFFC7C59CE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sampaio, Vinicius Gomes;Roell, Talita;Campos, Luiz Alexandre	Sampaio, Vinicius Gomes, Roell, Talita, Campos, Luiz Alexandre (2023): Revision of Heteroscelis Latreille, 1829 (Heteroptera: Pentatomidae: Asopinae) with the description of a new species from Colombia. Zootaxa 5278 (3): 401-438, DOI: 10.11646/zootaxa.5278.3.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.3.1
03BADF229324FFE3FF22368DFADD5CD2.text	03BADF229324FFE3FF22368DFADD5CD2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heteroscelis bimaculata (Walker 1867)	<div><p>Heteroscelis bimaculata (Walker, 1867)</p><p>(Figs. 1, 20–24)</p><p>Platynopus bimaculatus Walker, 1867: 124; Distant 1900: 55 (syn.); Schouteden 1907: 20.</p><p>Heterosceloides bimaculata: Schouteden 1907: 20; Kirkaldy 1909: 30; Pirán 1961: 87 (syn.).</p><p>Heteroscelis bimaculata: Distant 1900: 55; Pirán 1961: 87; Thomas 1992: 49; Cambra et al. 2018: 5; Lupoli 2019: 79; Roell et al. 2023: 26.</p><p>Heteroscelis doello-juradoi Pirán, 1961: 92 new syn. (removed from the synonymy with H. servillii).</p><p>Heteroscelis denieri Pirán, 1961: 94; Thomas 1992: 49 (syn.).</p><p>Heteroscelis durionei Pirán, 1961: 95; Thomas 1992: 49 (syn.).</p><p>Types examined</p><p>BRAZIL. Platynopus bimaculatus Walker, 1867; male holotype; labels: “PETROPOLIS Feb. 1857. J. Gray” “6. Platynopus bimaculatus ”, “Type”, “ Holotype ”; “NHMUK 010592394”; (NHMUK). Photo examined (Fig. 21).</p><p>ARGENTINA. Heteroscelis doello-juradoi Pirán, 1961; male holotype; labels: “ HOLOTYPUS ”, “ Heteroscelis doello-juradoi Pirán 1959 ”, “Provincia Jujuy ”; (MACN). Photo examined (Fig. 22).</p><p>BOLIVIA. Heteroscelis denieri Pirán, 1961; male holotype; “ HOLOTYPUS ”, “ Heteroscelis Latr. ?”, “Chirca 7.IV.31”, “ Heteroscelis denieri Pirán 1959 ”, “1420”; (MLPA). Photo examined (Fig. 23 A, B).</p><p>BOLIVIA. Heteroscelis durionei Pirán, 1961; female holotype; labels:“ HOLOTYPUS ”, “Coroico”, “ Heteroscelis durionei Pirán 1959 ”, “1421”; (MLPA). Photo examined (Fig. 23 C, D).</p><p>Other material examined. BRAZIL, Rio de Janeiro: (Itatiaia, 700m, -22.41099, -44.57621), 1 female, 1.IV.1925 J.F. Zikán (CEIOC); ECUADOR, Napo: (Tiputini Biodiversity Station, 216m, -0.6379, -76.1495), 1 female, 6.II.1999 T.L. Erwin, et al. (USNM); PERU, Cuzco: (Pilcopata, -14.13262, -69.27723), 1 female (UFRG).</p><p>Diagnosis. Coria with a circular yellow spot near to the posterior margin (Fig. 20, A); urosternite III tubercle yellow or orange, from apex to the posterior margin of the urosternite; two pairs of yellow abdominal spots, the first pair lateral in urosternites VI and VII, and the second pair mesial in urosternite VII; vesicular area of female genitalia twice longer than the proximal part of the ductus receptaculi.</p><p>Redescription</p><p>BODY: General color ranging from brown to iridescent purple; some specimens with yellow spots, callous or not, on the head, pronotum and scutellum.</p><p>HEAD: Mandibular plates equal to or slightly surpassing clypeus, anteriorly rounded, separated at apex; head length anterior to the eyes equal to 2/3 of the total length of the head (Fig. 20, D); labium almost reaching the metasternum; proportion of labiomeres: I&lt;II&gt;III&lt;IV; proportion of antennomeres: I&lt;II&lt;III ≅ IV ≅ V.</p><p>THORAX: Coria longer than scutellum, not reaching the posterior margin of the connexival segment VI, uniformly punctured; membrane dark and surpassing posteriorly the abdomen; metapleural evaporatoria surpassing posteriorly the anterior limit of coxae; peritreme disc-shaped, surpassing half the pleural width; apex of peritreme rounded, posterior margin strongly arched, convex, anterior margin almost straight (Fig. 20, E); profemora with rudimentary ante-apical spine; foretibial length on average 3.5x larger than the width of the tibial dorsal expansion (Fig. 20, F).</p><p>ABDOMEN: Posterior angles of urosternites III to VII ending in minute spine (Fig. 20, B).</p><p>Male</p><p>GENITALIA (Fig. 21, C; Fig. 22, D): Pygophore with ventral rim slightly sinuous, shallowly concave medially.</p><p>Female</p><p>MEASUREMENTS (n=4). Head length 1.04 ± 0.23 (0.75–1.32); width 1.37 ± 0.13 (1.25–1.52); pronotum length 1.70 ± 0.36 (1.38–2.16); width 4.21 ± 0.53 (3.78–4.88); scutellum length 2.46 ± 0.39 (2.20–3.04); width 2.35 ± 0.39 (2.01–2.88); length of antennomeres: I (n=3) 0.31 ± 0.06 (0.25–0.37), II (n=3) 0.67 ± 0.09 (0.56–0.75), III (n=3) 0.79 ± 0.13 (0.69–0.94), IV (n=2) 0.88 ± 0.17 (0.75–1.00); V (n=1) 0.94; length of labiomeres: I 0.70 ± 0.11 (0.63–0.88), II 0.82 ± 0.15 (0.69–1.04), III 0.52 ± 0.09 (0.44–0.64), IV 0.67 ± 0.08 (0.63–0.80); width of foretibal expansion 0.53 ± 0.06 (0.50–0.56); width of abdomen 3.56 ± 0.45 (3.21–4.16); total length 6.26 ± 0.95 (5.41–7.60).</p><p>GENITALIA (Fig. 24):Valvifers VIII posterior margin bisinuous, uniformly convex over the base of laterotergites IX; laterotergites VIII wider than long, with spiracles close to the lateral margin. Segment X with the posterior margin straight; valvulae IX with a median hourglass-shaped secondary thickening; capsula seminalis oval.</p><p>Distribution: Panama, Ecuador (new record), French Guyana, Brazil, Peru, Bolivia and Argentina (Fig. 1).</p></div>	https://treatment.plazi.org/id/03BADF229324FFE3FF22368DFADD5CD2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sampaio, Vinicius Gomes;Roell, Talita;Campos, Luiz Alexandre	Sampaio, Vinicius Gomes, Roell, Talita, Campos, Luiz Alexandre (2023): Revision of Heteroscelis Latreille, 1829 (Heteroptera: Pentatomidae: Asopinae) with the description of a new species from Colombia. Zootaxa 5278 (3): 401-438, DOI: 10.11646/zootaxa.5278.3.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.3.1
03BADF229328FFE6FF2233E4FC6459BA.text	03BADF229328FFE6FF2233E4FC6459BA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heteroscelis robustus Thomas 1992	<div><p>Heteroscelis robustus Thomas, 1992</p><p>(Figs. 1, 25–29)</p><p>Heteroscelis robustus Thomas, 1992: 49; Grazia &amp; Schwertner 2011: 713; Silva et al. 2018: 404.</p><p>Types examined. BRAZIL. Heteroscelis robustus Thomas, 1992; male holotype; labels: “ HOLOTYPE. Heteroscelis robustus Thom. ”, “Braz. Mato Grosso — Barra do Tapirape. 26–XII–62. B. Malkin ”. “ California Academy of Sciences ”. “Type n°: 16773” (CAS). Photo examined (Fig. 26) .</p><p>Other material examined. BRAZIL, Amazonas: (Manaus, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-60.01282&amp;materialsCitation.latitude=-3.04361" title="Search Plazi for locations around (long -60.01282/lat -3.04361)">Conj. Manca.</a> -3.04361, -60.01282), 1 female, 10.V.1988 P.E.S. Silva (INPA); Roraima: (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-60.61212&amp;materialsCitation.latitude=3.18861" title="Search Plazi for locations around (long -60.61212/lat 3.18861)">Boa Vista</a>, 3.18861, -60.61212), 1 female, 31.V.1974 (UFRG); Pará: (Peixe-Boi, -1.10168, -47.27259), 1 female, 12.IV.1977 P Waldir (UFRG); (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.3142&amp;materialsCitation.latitude=-1.37399" title="Search Plazi for locations around (long -48.3142/lat -1.37399)">Marituba</a>, -1.37399, -48.3142), 1 female, 07.VI.1961 J &amp; B. Bechyné (UFRG); Mato Grosso: (Chapada dos <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-55.53735&amp;materialsCitation.latitude=-15.02015" title="Search Plazi for locations around (long -55.53735/lat -15.02015)">Guimar</a> „es, -15.02015, -55.53735), 1 female, 20.I.1961 J &amp; B. Bechyné (UFRG); São Paulo: 3 females, J. Mráz (NMPC); Rio Grande do Sul: (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-51.51407&amp;materialsCitation.latitude=-29.48006" title="Search Plazi for locations around (long -51.51407/lat -29.48006)">Salvador do Sul</a>, -29.48006, -51.51407), 1 male, 16.X.1994 (MCTP); COLOMBIA, (Meta, San Martín, Reserva Natural El Caduceo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.65945&amp;materialsCitation.latitude=3.671389" title="Search Plazi for locations around (long -73.65945/lat 3.671389)">Km</a> 4.5, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.65945&amp;materialsCitation.latitude=3.671389" title="Search Plazi for locations around (long -73.65945/lat 3.671389)">Via San Francisco</a>, 3.671389, -73.659444), 1 male, 30.IX–4.X.2013 M Arango (MPUJ _ENT).</p><p>Diagnosis. Body robust compared to the other species, dark-brown to black; unpunctured, yellowish longitudinal line extending from the anterior margin of the pronotum to the posterior margin of the scutellum; radial vein of coria usually yellowish; foretibial length on average ten times larger than the width of the dorsal tibial expansion (Fig. 25, F); capsula seminalis digitiform.</p><p>Redescription</p><p>HEAD: Mandibular plates equal to or slightly surpassing clypeus, rounded at apex, never touching each other; head length anterior to the eyes smaller than 2/3 of the total length of the head (Fig. 25, D); labium reaching the metasternum; proportion of labiomeres: I&lt;II&gt;III ≅ IV; proportion of antennomeres: I&lt;II&lt;III ≅ IV ≅ V.</p><p>THORAX: Coria longer than the scutellum, not reaching the posterior margin of the connexival segment VI; metapleural evaporatoria not surpassing posteriorly the anterior limit of coxae; peritreme disc-shaped, surpassing half the pleural width; apex of peritreme rounded, posterior margin arched, convex, anterior margin almost straight (Fig. 25, E).</p><p>ABDOMEN: Uniformly punctured; urosternite III tubercle not surpassing the posterior margin of metasternum; posterior angles of urosternites III to VII ending in minute spine, more developed in VII (Fig. 25, B).</p><p>Male:</p><p>MEASUREMENTS (n=2). Head length 1.24 ± 0.05 (1.20–1.28); width 1.48 ± 0.05 (1.44–1.52); pronotum length 1.88 ± 0.16 (1.76–2.00); width 4.12 ± 0.16 (4.00–4.24); scutellum length 2.60 ± 0.05 (2.56–2.64); width 2.60 ± 0.05 (2.24–2.40); length of antennomeres: I (n=1) 0.32, II 0.64 ± 0.11 (0.56–0.72), III 0.84 ± 0.05 (0.80–0.88), IV 0.96 ± 0.11 (0.88–1.40); V (n=1) 1.12; length of labiomeres: I 0.68 ± 0.05 (0.64–0.72), II 0.72 ± 0.00 (0.72–0.72), III 0.40 ± 0.00 (0.40–0.40), IV 0.48 ± 0.00 (0.48–0.48); width of foretibal expansion 0.20 ± 0.05 (0.16–0.24); width of abdomen 3.40 ± 0.28 (3.20–3.60); total length (n=1) 7.20.</p><p>GENITALIA: Pygophore (Fig. 27): setae denser on medial portion of dorsal rim; dorsal rim sinuous, medially concave; ventral rim sinuous, shallowly concave medially; segment X setose on the entire surface; heads of parameres lateroventrally directed in posterior view. Phallus (Fig. 28): conjunctival dorsal lobe with one pair of long projections, ventral lobes with two projections each.</p><p>Female:</p><p>MEASUREMENTS (n=8). Head length 1.12 ± 0.09 (1.00–1.28); width 1.40 ± 0.05 (1.44 –1.53); pronotum length 1.96 ± 0.21 (1.76–2.20); width 4.42 ± 0.33 (4.16–5.00); scutellum length 2.99 ± 0.26 (2.72–3.50); width 2.66 ±0.25 (2.40–3.10); length of antennomeres: I (n=6) 0.38 ± 0.03 (0.32–0.44), II (n=4) 0.71 ± 0.10 (0.60–0.80), III (n=4) 0.98 ± 0.09 (0.88–1.10), IV (n=1) 1.12; V (n=1) 1.04; length of labiomeres: I (n=7) 0.76 ± 0.07 (0.70–0.90), II (n=7) 0.90 ± 0.11 (0.80–1.10), III (n=7) 0.40 ± 0.05 (0.32–0.50), IV (n=7) 0.50 ± 0.04 (0.48–0.60); width of foretibal expansion (n=5) 0.25 ± 0.06 (0.16 –0.30); width of abdomen 4.06 ± 0.25 (3.68–4.50); total length (n=6) 7.67 ± 0.88 (6.56 –8.90).</p><p>GENITALIA (Fig. 29): Valvifers VIII posterior margin bisinuous, truncate over the laterotergites VIII, and slightly projected over the basal angles of laterotergites IX, setae on sutural margins; laterotergites VIII slightly longer than wide, with spiracles close to the lateral margin; segment X with the posterior margin straight; valvulae IX with 1+1 secondary thickenings, these bean-shaped with the anterior margin elongated with acute apices; vesicular area half the length of the proximal part of the ductus receptaculi.</p><p>Distribution: Paraguay, Colombia (new record) and Brazil (Fig. 1).</p></div>	https://treatment.plazi.org/id/03BADF229328FFE6FF2233E4FC6459BA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sampaio, Vinicius Gomes;Roell, Talita;Campos, Luiz Alexandre	Sampaio, Vinicius Gomes, Roell, Talita, Campos, Luiz Alexandre (2023): Revision of Heteroscelis Latreille, 1829 (Heteroptera: Pentatomidae: Asopinae) with the description of a new species from Colombia. Zootaxa 5278 (3): 401-438, DOI: 10.11646/zootaxa.5278.3.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.3.1
03BADF22932FFFDAFF2233B5FD9E5A92.text	03BADF22932FFFDAFF2233B5FD9E5A92.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heteroscelis vandembergi Sampaio & Roell & Campos 2023	<div><p>Heteroscelis vandembergi Sampaio &amp; Campos, sp. nov.</p><p>(Figs. 1, 30–32)</p><p>Type Material</p><p>Holotype: male, here designated. COLOMBIA. Santander. Charalá. Virolin. Mar 1981. 18-III. Santander. Charalá. Virolin. Heteroscelis lepida Stål, 1862 —ICN. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.2004&amp;materialsCitation.latitude=6.108" title="Search Plazi for locations around (long -73.2004/lat 6.108)">Det</a>: F. Rua Alvear, 2006. ICN 055894. (6.1080, -73.2004). Deposited in: ICN (Fig. 30).</p><p>Paratype: male, here designated. COLOMBIA. Boyacá. Villa de Leyva. SFF <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.48333&amp;materialsCitation.latitude=5.633333" title="Search Plazi for locations around (long -73.48333/lat 5.633333)">Iguaque</a>. 05°38’N 73°29’W WGS 84. 2450 m. Captura manual. 19.VIII.1998. J. Castro. IAvH-E-VillaDeLeyva. Deposited in: IAvH-E (Fig. 31).</p><p>Etymology. Named after the father of the first author, Vandemberg de Freitas Sampaio (in memoriam), who made a great effort to provide a good education for his children, but unfortunately could not reap the fruits he sowed.</p><p>Diagnosis. Head, thorax and abdomen orange to red, coria iridescent-blue, radial vein usually reddish; foretibial length on average 9.5x larger than the width of the dorsal expansion (Fig. 30, F).</p><p>Description</p><p>HEAD: Mandibular plates slightly surpassing clypeus, anteriorly rounded, separated at apex; head length anterior to the eyes shorter than 2/3 of the total length of the head (Fig. 30, D); proportion of labiomeres: I&lt;II&gt;III&lt;IV; proportion of antennomeres: I&lt;II&lt;III, IV and V missing.</p><p>THORAX: Coria iridescent-blue, uniformly punctured, longer than the scutellum, not reaching the posterior margin of connexival segment VI; radial vein usually reddish; membrane dark, surpassing posteriorly the abdomen; metapleural evaporatoria not surpassing posteriorly the anterior limit of coxae; peritreme disc-shaped, surpassing half the pleural width; apex of peritreme rounded, posterior margin arched, convex, anterior margin almost straight (Fig. 30, E).</p><p>ABDOMEN: Uniformly punctured; urosternite III tubercle not surpassing the posterior margin of metasternum; posterior angles of urosternites III to VII ending in minute spine (Fig. 30, B).</p><p>Male:</p><p>MEASUREMENTS (n=1). Head length 1.52; width 1.36; pronotum length 1.84; width 4.08; scutellum length 2.48; width 3.04; length of antennomeres: I 0.32, II 0.64, III 1.04, IV–, V–; length of labiomeres: I 0.60, II 0.90, III 0.40, IV 0.50; width of foretibal expansion 0.24; width of abdomen 3.60; total length 6.72.</p><p>GENITALIA: Pygophore (Fig.32): setae denser on posterolateral angles; dorsal rim concave; ventral rim sinuous, medially concave; segment X with surface corrugated; head of parameres laterodorsally directed in posterior view.</p><p>Remarks: This species is quite similar to H. lepida, however, without considering the difference in color, H. vandembergi sp.nov. also differs from H. lepida in the proportion of the area anterior to the eyes, and in the proportion of the dorsal foretibial expansions in relation to the femur, a trait that does not vary within the other species of the genus.</p><p>Female unknown.</p><p>Distribution: Colombia (Fig. 1).</p></div>	https://treatment.plazi.org/id/03BADF22932FFFDAFF2233B5FD9E5A92	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sampaio, Vinicius Gomes;Roell, Talita;Campos, Luiz Alexandre	Sampaio, Vinicius Gomes, Roell, Talita, Campos, Luiz Alexandre (2023): Revision of Heteroscelis Latreille, 1829 (Heteroptera: Pentatomidae: Asopinae) with the description of a new species from Colombia. Zootaxa 5278 (3): 401-438, DOI: 10.11646/zootaxa.5278.3.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.3.1
