taxonID	type	description	language	source
03BADF229332FFF8FF22368DFD5E5CCF.taxon	type_taxon	Type species: Heteroscelis servillii Laporte, 1833, by subsequent monotypy (ICZN 1999: Article 69.3)	en	Sampaio, Vinicius Gomes, Roell, Talita, Campos, Luiz Alexandre (2023): Revision of Heteroscelis Latreille, 1829 (Heteroptera: Pentatomidae: Asopinae) with the description of a new species from Colombia. Zootaxa 5278 (3): 401-438, DOI: 10.11646/zootaxa.5278.3.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.3.1
03BADF229332FFF8FF22368DFD5E5CCF.taxon	diagnosis	Diagnosis. Head elongated, with lateral post-ocular projections; proportion of labiomeres: I <II> III <IV; pronotal depression anterior to the cicatrices forming a collar; humeral angles laterally projected, bifid, with the anterior portion longer than the posterior; pro-, meso- and metasternum covered by small setae; prosternum flat, mesosternum with a low longitudinal carina dilated anteriorly and posteriorly, metasternum slightly elevated with a shallow longitudinal groove.	en	Sampaio, Vinicius Gomes, Roell, Talita, Campos, Luiz Alexandre (2023): Revision of Heteroscelis Latreille, 1829 (Heteroptera: Pentatomidae: Asopinae) with the description of a new species from Colombia. Zootaxa 5278 (3): 401-438, DOI: 10.11646/zootaxa.5278.3.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.3.1
03BADF229332FFF8FF22368DFD5E5CCF.taxon	description	Redescription BODY: Elongated oval, with variable color patterns. HEAD: Rectangular, longer than wide, longitudinally convex, slightly punctured; mandibular plates longer than clypeus, with the anterior margin rounded; bucculae low, rounded at apex and evanescent posteriorly; eyes separated from the anterolateral angles of pronotum by at least half the eye diameter; post-ocular lateral triangular projection contiguous to the eyes; antennomere I shorter and wider than II, which is thinner than the others; antennomeres III, IV and V similar in length and more setose than the others. THORAX: Pronotum trapezoidal, uniformly and deeply punctured, twice as wide as long; scutellum uniform and shallowly punctured; frenal constriction width more than half the basal width, postfrenal lobe with parallel margins, broadly rounded at apex; coria with posterior margin sinuous; mesopleural evaporatoria occupying the entire posterior margin and the anterolateral mesopleural angle; metapleural evaporatoria surrounding the disc-shaped peritreme, occupying more than half of the metapleural width; ostiole guttiform in ventral view, laterally oriented; peritremal furrow extending more than half the length of the peritreme; dorsal and ventral foretibial expansions present, the dorsal expansion wider than the ventral. ABDOMEN: Uniformly punctured; mid tubercle on urosternite III; posterior angles of urosternites III to VII ending in minute spine; male bearing ventrolateral glandular setose patches, extending from near the anterior margin of urosternite V to the anterior margin of urosternite VII. Male GENITALIA: Pygophore bowl-shaped, with setae on the entire surface, ventral rim slightly sinuous; inferior layer excavated; posterolateral angles rounded; segment X trapezoidal, with rounded apex, ventrally directed, dorsally sclerotized, setose on apex; pseudoclaspers concave, sculptured, each placed between the paramere and the lateral portion of the dorsal rim; parameres sculptured, head triangular, spatulate, occupying in dorsal view most of the space between the segment X and the posterolateral angles. Phallus: phallotheca divided in a basal theca and thecal shield; ejaculatory reservoir contained inside basal theca; basal foramen elliptical, reinforced by basal plates; thecal shield longer and wider than basal theca; vesica oval in dorsal and lateral view; ductus seminis with uniform diameter, running between filaments of vesica, ending on a secondary gonopore, ventrally directed; conjuntiva with: two ventral lobes with sclerotized apex, and one dorsal lobe; all lobes posteriorly directed. Female GENITALIA: Valvifers VIII longer than wide, subtriangular, posterior margins sinuous, sutural margins straight, juxtaposed; laterotergites VIII subtriangular; exposed portion of valvifer IX trapezoidal, wider than long, juxtaposed to the laterotergites IX; exposed portion of laterotergites IX digitiform, attaining the mediotergite VIII; segment X trapezoidal with posterior margin straight; thickening of vaginal intima elongated; pars intermedialis small, narrower than the median duct of vesicular area; capsula seminalis longer and wider than pars intermedialis.	en	Sampaio, Vinicius Gomes, Roell, Talita, Campos, Luiz Alexandre (2023): Revision of Heteroscelis Latreille, 1829 (Heteroptera: Pentatomidae: Asopinae) with the description of a new species from Colombia. Zootaxa 5278 (3): 401-438, DOI: 10.11646/zootaxa.5278.3.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.3.1
03BADF229332FFF8FF22368DFD5E5CCF.taxon	distribution	Distribution: United States of America (Texas), Mexico, Guatemala, Honduras, El Salvador, Costa Rica, Panama, Colombia, Ecuador (new record), Trinidad and Tobago, Venezuela, Guyana, Surinam, French Guyana, Brazil, Peru, Bolivia, Paraguay and Argentina (Fig. 1).	en	Sampaio, Vinicius Gomes, Roell, Talita, Campos, Luiz Alexandre (2023): Revision of Heteroscelis Latreille, 1829 (Heteroptera: Pentatomidae: Asopinae) with the description of a new species from Colombia. Zootaxa 5278 (3): 401-438, DOI: 10.11646/zootaxa.5278.3.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.3.1
03BADF229335FFF4FF223095FCFB5DD5.taxon	description	(Figs. 1 – 14)	en	Sampaio, Vinicius Gomes, Roell, Talita, Campos, Luiz Alexandre (2023): Revision of Heteroscelis Latreille, 1829 (Heteroptera: Pentatomidae: Asopinae) with the description of a new species from Colombia. Zootaxa 5278 (3): 401-438, DOI: 10.11646/zootaxa.5278.3.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.3.1
03BADF229335FFF4FF223095FCFB5DD5.taxon	materials_examined	Types examined SURINAM. Agerrus remipes Stål, 1859; female syntype; labels: “ Surinam. ”, “ Stål ”, “ Agerrus remipes Stål. Typ. ”, “ 111 ”, “ TYPUS ”, “ NHRS — GULI 000096153 ”; (NHRS). Photo examined (Fig. 3). BRAZIL. Heteroscelis amazonica Pirán, 1961; female holotype; labels: “ Ig do Passarinho Manaus. Am. 24. II. 55. Costa Leite e N. Cerqueira — cols. 07. 9 / 59 ”, “ Heteroscelis amazonica Pirán 1960 ”, “ HOLOTYPUS ”; (MNRJ) (Fig. 4). BRAZIL. Heteroscelis carcavalloi Pirán, 1961; female alotype; labels: “ Sítio Vieiralves Manaus — Am. 21. Xll. 65. Elias e Rappa — cols. 02. 9 / 59 ”, “ Heteroscelis carcavalloi Pirán 1960 ”, “ ALLOTYPUS ”; (MNRJ). BRAZIL. Bodetria brenthoides Walker, 1867; female holotype; labels: “ Amazon St. Paul / 60 32 ” “ Type ”, “ Holotype ”, “ 1. Bodetria brenthoides ”; “ NHMUK 010592390 ”; (NHMUK). Photo examined (Fig. 5). BRAZIL. Bodetria chrysoclora Walker, 1868; male holotype; labels: “ Braz / 62 57 ” “ Type ”, “ Holotype ”, “ Bodetria chrysoclora ”; “ NHMUK 010592391 ”; (NHMUK). Photo examined (Fig. 6). BRAZIL. Bodetria indecora Walker, 1868; male holotype; labels: “ Braz / 62 57 ” “ Type ”, “ Holotype ”, “ Bodetria indecora ”; “ NHMUK 010592393 ”; (NHMUK). Photo examined (Fig. 7). BOLIVIA. Heteroscelis carcavalloi Pirán, 1961; male holotype; labels: “ HOLOTYPUS ”, “ Yungas del Palmar. IV – 954. A. Martinez. ”, “ Heteroscelis carcavalloi Pirán, 1960. ”; (MACN). Photo examined (Fig. 8). BOLIVIA. Heteroscelis zischkai Kormilev, 1951; female alotype; labels: “ BOLIVIA, Region Chaparé, Dep. Cochabamba (400 mtr), 30. VIII. 1951. Dirings. ”. Heteroscelis zischkai Kormilev. Museo Argentino de Ciencias Naturales. ”. “ ALLOTYPE ”; (MACN) (Fig. 9). PARAGUAY. Heteroscelis dureti Pirán, 1954; female holotype; labels: “ Heteroscelis dureti Det. Pirán. 952 ”, “ HOLOTYPUS ”, “ Paraguay, Villarrica, Schade col. 15. IV. 46 ”; (MACN). Photo examined (Fig. 10). ARGENTINA. Heteroscelis bergi Pirán, 1954; male holotype; labels: “ Missiones — Argentina. Dep. Concep. — Sta. Maria. M. J. Viana ”, “ 53.484 ”, “ HOLOTYPUS ”, “ Heteroscelis bergi Det. Pirán. 952 ”; (MACN). Photo examined (Fig. 11). Other material examined. VENEZUELA, Guayana: (Conuco, Rio Moroco), 3 males, 18 / 19. I. 1972 G. & E. Scherer (NMPC); Zulia: (Kasmera, 9.94907, - 72.75292), 1 female, 20. IX. 1961 F. Fernandez Y. & C. J. Rosales (UFRG); Cx. Panamexicana El Vigia Coloneito: (Km 8.32611, - 72.08742), 1 female, 6. I. 1955. F. Fernandez Y. & C. J. Rosales (UFRG); Aragua: (El Limón, 10.30589, - 67.63212), 1 male, 2. III. 1951 F. Fernandes (UFRG); Aragua: (Maracay, 10.23535, - 67.59113), 1 female, 2. V. 1953 N. Angeles (UFRG). TRINIDAD AND TOBAGO: 2 males, XII. 1953 G. u. Helga Frey. (NMPC). BRAZIL, 1 female, 11. IV. 1962 Cerq. (UFRG); Pará: (Belém Mocambo, - 1.45583, - 48.50444), 1 female, 6. II. 1979 P Tadeu (UFRG); (Peixe-boi, - 1.10168, - 47.27259), 1 male, 12. IV. 1977 P Waldir. (UFRG); (Serra Norte Manganês - 6.01748, - 50.30386), 1 female, 22. X. 1984 T. Pimentel; Amazonas: (Manaus, CEPLAC, - 2.8905, - 59.9576), 1 female, 15. VII. 1977 (UFRG); (Serra dos Porcos, 0.416667, - 69.366667), 1 male, 25. II. 1977 Franklim (INPA); (Manaus, CEPLAC, Am 010 Km 31, - 2.8905, - 59.9576), 1 male, 7. V. 1988 Albuquerque (UFRG); (Manaus, - 3.04361, - 60.01282), 1 male, 5. V. 1976 Paraluppi (UFRG); (Manaus, Br 174 Km 41, Agricultura Geológica - 2.6770, - 60.04668), 1 male, 15. VI. 1981 C. A. Fonseca (INPA); (Itacoatiara, - 3.1373, - 58.4418), 1 female, Dirings (MCNZ); Amapá: (Serra do Navio, 1.65803, - 52.28195), 1 female, 13. VII. 1961 AP. J & B. Bechyné (UFRG); Rio Grande do Norte: (Natal, - 5.8101, - 35.22674), 1 male, I. 1952 M. Alvarenga (UFRG); Pernambuco: (Paudalho, Acerolândia, - 7.850, - 35.250. 160 m), 1 male, II. 2016 Grossi & Parizotto (CERPE); (Paudalho, Acerolândia, - 7.850, - 35.250. 160 m), 1 female, XI. 2017 Grossi & Parizotto (CERPE); Mato Grosso do Sul: (Bonito, Praia da Figueira - 21.2972, - 56.5005), 1 female, 01. III. 2009 Grossi & Parizotto (DZUP); Rio Grande do Sul: (Cerro Azul - 27.7403, - 51.5199), 1 female, I. 1931 Pe. Buck leg. (MCNZ); ECUADOR, Amazonia: 1 female, II. 1998 lgt. Bezdek (NMPC); Napo: (Tiputini Biodiversity Station, 216 m, - 0.6379, - 76.1495), 1 male, 06. II. 1999 T. L. Erwin, et al. (USNM); (Tiputini Biodiversity Station, 216 m, - 0.6379, - 76.1495), 1 female, 22. X. 1998 T. L. Erwin, et al. (USNM); PERU, Pucalipa: (Rio Ucayali, - 8.4084, - 74.6058), 1 female, Dirings (MCNZ); BOLIVIA, Cochabamba (Chapare, 400 m - 16.80922, - 65.71926), 1 female, 07. IX. 1951 Dirings (MACN). Records retrieved from iNaturalist TRINIDAD AND TOBAGO, San Juan-Laventille: (10.605137, - 61.42763), 14. II. 2021. BRAZIL, Amazonas: (Manaus, - 3.095784, - 59.989879), 13. VIII. 2020. BRAZIL, Espírito Santo: (S „ o Gabriel de Palha, - 18.906837, - 40.478105), II. 2014. BRAZIL, S „ o Paulo: (Campinas, - 22.819691, - 47.064677), 16. II. 2023. BRAZIL, Rio de Janeiro: (Cabo Frio, - 22.886901, - 42.026649), 8. I. 2017.	en	Sampaio, Vinicius Gomes, Roell, Talita, Campos, Luiz Alexandre (2023): Revision of Heteroscelis Latreille, 1829 (Heteroptera: Pentatomidae: Asopinae) with the description of a new species from Colombia. Zootaxa 5278 (3): 401-438, DOI: 10.11646/zootaxa.5278.3.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.3.1
03BADF229335FFF4FF223095FCFB5DD5.taxon	diagnosis	Diagnosis. Mandibular plates distinctly longer than clypeus, converging apically; head length anterior to the eyes more than 2 / 3 of the total length of the head (Fig. 2, D); urosternite III tubercle yellow or orange at apex, and surpassing the posterior margin of metasternum (Fig. 2, B).	en	Sampaio, Vinicius Gomes, Roell, Talita, Campos, Luiz Alexandre (2023): Revision of Heteroscelis Latreille, 1829 (Heteroptera: Pentatomidae: Asopinae) with the description of a new species from Colombia. Zootaxa 5278 (3): 401-438, DOI: 10.11646/zootaxa.5278.3.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.3.1
03BADF229335FFF4FF223095FCFB5DD5.taxon	description	Redescription BODY: General color ranging from brown and dark red to iridescent hues of green, yellow, blue and purple, with or without yellow spots on the pronotum, scutellum and coria (more commonly observed in non-iridescent specimens). HEAD: Labium almost reaching the metasternum (Fig. 2, B); proportion of antennomeres: I <II <III ≅ IV ≅ V. THORAX: Coria longer than scutellum, not reaching the posterior margin of connexival segment VI, uniformly punctured; membrane dark and surpassing posteriorly the abdomen; metapleural evaporatoria surpassing posteriorly the anterior limit of coxae; peritreme reaching or surpassing half the pleural width; apex of peritreme rounded, posterior margin a little more arched than the anterior, both convex (Fig. 2, E); profemora with rudimentary ante-apical spine; foretibial length on average 3.5 x larger than the width of the tibial dorsal expansion (Fig. 2, F). ABDOMEN: Posterior angles of urosternites III to VII ending in minute spine, more developed in VII (Fig. 2, B). Male: MEASUREMENTS (n = 13): Head length 1.76 ± 0.28 (1.30 – 1.84); width 1.29 ± 0.07 (1.20 – 1.50); pronotum length 2.00 ± 0.26 (1.50 – 2.40); width 4.21 ± 0.26 (3.80 – 4.50); scutellum length 2.38 ± 0.16 (2.08 – 2.60); width 2.30 ± 0.13 (2.08 – 2.56); length of antennomeres: I (n = 11) 0.39 ± 0.03 (0.30 – 0.40), II (n = 11) 0.71 ± 0.09 (0.60 – 0.90), III (n = 11) 0.95 ± 0.10 (0.80 – 1.04), IV (n = 11) 1.03 ± 0.14 (0.72 – 1.20); V (n = 8) 1.09 ± 0.20 (0.72 – 1.40); length of labiomeres: I (n = 8) 0.67 ± 0.08 (0.60 – 0.80), II (n = 8) 0.87 ± 0.07 (0.80 – 1.00), III (n = 8) 0.45 ± 0.07 (0.40 – 0.60), IV (n = 8) 0.68 ± 0.09 (0.60 – 0.90); width of foretibal expansion (n = 10) 0.59 ± 0.07 (0.48 – 0.72); width of abdomen (n = 11) 3.21 ± 0.18 (2.96 – 3.50); total length 7.04 ± 0.65 (6.10 – 8.30). GENITALIA: Pygophore (Fig. 12): setae denser on posterolateral angles and on medial portion of dorsal rim; dorsal rim sinuous, medially concave; ventral rim slightly sinuous, medially concave; segment X with surface corrugated and with a membranous longitudinal line; head of parameres lateroventrally directed in posterior view. Phallus (Fig. 13): conjunctival dorsal lobe with one pair of long projections and one single forked central projection, ventral lobes with two projections each. Female: MEASUREMENTS (n = 15): Head length (n = 11) 2.06 ± 0.38 (1.40 – 2.60); width (n = 11) 1.47 ± 0.20 (1.20 – 1.90); pronotum length (n = 13) 2.45 ± 0.26 (1.92 – 2.80); width (n = 13) 4.83 ± 0.42 (3.92 – 5.60); scutellum length 2.88 ± 0.32 (2.24 – 3.40); width 2.56 ± 0.40 (1.84 – 3.00); length of antennomeres: I (n = 12) 0.42 ± 0.06 (0.30 – 0.50), II (n = 12) 0.79 ± 0.15 (0.56 – 1.10), III (n = 12) 1.04 ± 0.21 (0.60 – 1.30), IV (n = 8) 1.22 ± 0.39 (0.80 – 2.00); V (n = 5) 1.03 ± 0.18 (0.96 – 1.30); length of labiomeres: I (n = 9) 0.77 ± 0.11 (0.60 – 0.90), II (n = 9) 1.02 ± 0.16 (0.70 – 1.30), III (n = 9) 0.50 ± 0.10 (0.40 – 0.70), IV (n = 9) 0.85 ± 0.14 (0.60 – 1.00); width of foretibal expansion (n = 11) 0.72 ± 0.09 (0.56 – 0.90); width of abdomen 3.70 ± 0.33 (3.04 – 4.30); total length (n = 8) 8.28 ± 1.05 (6.88 – 9.50). GENITALIA (Fig. 14): Valvifers VIII posterior margin bisinuous, slightly projected over the basal angles of laterotergites IX; laterotergites VIII wider than long, with spiracles equidistant from the basal angle and the lateral margin; valvulae IX with 1 + 1 bean-shaped secondary thickenings; vesicular area half the length of the proximal part of the ductus receptaculi; capsula seminalis oval.	en	Sampaio, Vinicius Gomes, Roell, Talita, Campos, Luiz Alexandre (2023): Revision of Heteroscelis Latreille, 1829 (Heteroptera: Pentatomidae: Asopinae) with the description of a new species from Colombia. Zootaxa 5278 (3): 401-438, DOI: 10.11646/zootaxa.5278.3.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.3.1
03BADF229335FFF4FF223095FCFB5DD5.taxon	distribution	Distribution: Colombia, Ecuador (new record), Trinidad and Tobago, Venezuela, Guyana, Surinam, French Guyana, Brazil, Peru, Bolivia, Paraguay and Argentina (Fig. 1).	en	Sampaio, Vinicius Gomes, Roell, Talita, Campos, Luiz Alexandre (2023): Revision of Heteroscelis Latreille, 1829 (Heteroptera: Pentatomidae: Asopinae) with the description of a new species from Colombia. Zootaxa 5278 (3): 401-438, DOI: 10.11646/zootaxa.5278.3.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.3.1
03BADF229321FFEFFF2233CFFC7C59CE.taxon	description	(Figs. 1, 15 – 19)	en	Sampaio, Vinicius Gomes, Roell, Talita, Campos, Luiz Alexandre (2023): Revision of Heteroscelis Latreille, 1829 (Heteroptera: Pentatomidae: Asopinae) with the description of a new species from Colombia. Zootaxa 5278 (3): 401-438, DOI: 10.11646/zootaxa.5278.3.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.3.1
03BADF229321FFEFFF2233CFFC7C59CE.taxon	materials_examined	Types examined MEXICO. Phyllochirus lepidus Stål, 1862; female holotype; “ México. Coll. Signoret ”, “ lepidus set. Stål ”, “ lepidus ”, “ Heterosceloides lepidus Stål ”; (NHMW). Photo examined (Fig. 16). MEXICO. Bodetria scutellaris Walker, 1868; male holotype; “ Holotype ”, “ Type ”, “ Vera cruz / 5466 ”, “ Bodetria scutellaris ”; “ NHMUK 010592395 ”; (NHMUK). Photo examined (Fig. 17). Other material examined. PANAMA. Panama: (Canal Zone, Madden Forest, 9.05, - 79.37), 1 male, 8. I. 1971 H. Stockwell (STRI), Smithsonian Tropical Research institute, photo examined. VENEZUELA, Falcon: (Las dos Bocas, 11.30129, - 69.41226), 1 female, 29. I. 1967 R. Cassares (UFRG). COLOMBIA, Antioquia: (Santa Fé de Antioquia, Hacienda Cotové - UNAL, 600 m. 6.5316, - 75.8260), 1 female, 6. X. 2000 A. Botero, et al. (UNAB). 1 female, no data (UFRG). Records retrieved from iNaturalist MEXICO, Veracruz: (Puente Nacional, 19.34434, - 96.464912), 31. X. 2016. MEXICO, Veracruz: (Puente Nacional, 19.343728, - 96.464263), 25. IX. 2017. COSTA RICA, Puntarenas: (Puntarenas, 10.275075, - 84.812452) 25. V. 2017. COLOMBIA, Sucre: (Galeras, 9.147859, - 74.978428), 18. II. 2017. VENEZUELA, Falcon: (Miranda, 11.41002, - 69.679581), 5. XII. 2022.	en	Sampaio, Vinicius Gomes, Roell, Talita, Campos, Luiz Alexandre (2023): Revision of Heteroscelis Latreille, 1829 (Heteroptera: Pentatomidae: Asopinae) with the description of a new species from Colombia. Zootaxa 5278 (3): 401-438, DOI: 10.11646/zootaxa.5278.3.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.3.1
03BADF229321FFEFFF2233CFFC7C59CE.taxon	diagnosis	Diagnosis. Body iridescent-blue, with a large triangular red or orange spot on scutellum; vesicular area of female genitalia subequal in length to the proximal part of the ductus receptaculi.	en	Sampaio, Vinicius Gomes, Roell, Talita, Campos, Luiz Alexandre (2023): Revision of Heteroscelis Latreille, 1829 (Heteroptera: Pentatomidae: Asopinae) with the description of a new species from Colombia. Zootaxa 5278 (3): 401-438, DOI: 10.11646/zootaxa.5278.3.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.3.1
03BADF229321FFEFFF2233CFFC7C59CE.taxon	description	Redescription HEAD: Mandibular plates equal to or slightly surpassing clypeus, anteriorly rounded, separated at apex; head length anterior to the eyes equal to 2 / 3 of the total length of the head (Fig. 15, D); labium reaching the metasternum; proportion of labiomeres: I <II> III <IV; proportion of antennomeres: I <II <III <IV ≅ V. THORAX: Scutellum with a large triangular red or orange spot, covering the entire frenal lobe and the central portion of the postfrenal lobe; coria longer than the scutellum, not reaching the posterior margin of the connexival segment VI, uniformly punctured; membranes dark, surpassing posteriorly the abdomen; metapleural evaporatoria not surpassing posteriorly the anterior limit of coxae; peritreme disc-shaped, surpassing half the pleural width; apex of peritreme rounded, posterior margin arched, convex, anterior margin almost straight (Fig. 15, E); foretibial length on average 3.5 x larger than the width of the dorsal tibial expansion (Fig. 15, F). ABDOMEN: Uniformly punctured; urosternite III tubercle not surpassing the posterior margin of metasternum; posterior angles of urosternites III to VII ending in minute spine (Fig. 15, B). Male GENITALIA (Fig. 18): Pygophore with dorsal rim concave; ventral rim slightly sinuous, medially concave; segment X with surface corrugated; head of parameres laterodorsally directed in posterior view. Remarks: The only description of the internal male genitalia was made by McDonald (1966). Based on his description and illustrations (McDonald 1966: 121, figs. 300 – 302) we interpret the phallus having the conjunctival dorsal lobe (conjunctival appendages sic McDonald) with one pair of projections, and the ventral lobes with three projections each. Female MEASUREMENTS (n = 3). Head length 1.24 ± 0.22 (1.00 – 1.44); width 1.34 ± 0.07 (1.26 – 1.40); pronotum length 1.75 ± 0.21 (1.57 – 2.00); width 3.88 ± 0.24 (3.70 – 4.16); scutellum length 2.25 ± 0.27 (2.01 – 2.56); width 2.18 ± 0.11 (2.10 – 2.32); length of antennomeres: I 0.33 ± 0.04 (0.30 – 0.37), II 0.62 ± 0.15 (0.50 – 0.80), III 0.81 ± 0.19 (0.69 – 1.04), IV 1.09 ± 0.17 (0.90 – 1.20); V 1.00 ± 0.11 (0.88 – 1.12); length of labiomeres: I 0.53 ± 0.14 (0.37 – 0.64), II 0.71 ± 0.08 (0.63 – 0.80), III 0.40 ± 0.09 (0.30 – 0.48), IV 0.61 ± 0.11 (0.50 – 0.72); width of foretibal expansion 0.65 ± 0.26 (0.50 – 0.96); width of abdomen 3.41 ± 0.36 (3.20 – 3.84); total length 6.31 ± 0.64 (5.67 – 6.96). GENITALIA (Fig. 19): Valvifers VIII posterior margin shallowly concave over the plates of sternite IX, truncate over the base of laterotergites VIII; laterotergites VIII wider than long, with spiracles close to the lateral margin; segment X with the posterior margin straight; valvulae IX with 1 + 1 bean-shaped secondary thickenings with the proximal margin laterally elongated; capsula seminalis oval.	en	Sampaio, Vinicius Gomes, Roell, Talita, Campos, Luiz Alexandre (2023): Revision of Heteroscelis Latreille, 1829 (Heteroptera: Pentatomidae: Asopinae) with the description of a new species from Colombia. Zootaxa 5278 (3): 401-438, DOI: 10.11646/zootaxa.5278.3.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.3.1
03BADF229321FFEFFF2233CFFC7C59CE.taxon	discussion	Distribution: United States of America (Texas), Mexico, Guatemala, Honduras, El Salvador, Costa Rica (new record), Panama, Colombia, Ecuador, Venezuela, French Guyana. (Fig. 1).	en	Sampaio, Vinicius Gomes, Roell, Talita, Campos, Luiz Alexandre (2023): Revision of Heteroscelis Latreille, 1829 (Heteroptera: Pentatomidae: Asopinae) with the description of a new species from Colombia. Zootaxa 5278 (3): 401-438, DOI: 10.11646/zootaxa.5278.3.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.3.1
03BADF229324FFE3FF22368DFADD5CD2.taxon	description	(Figs. 1, 20 – 24)	en	Sampaio, Vinicius Gomes, Roell, Talita, Campos, Luiz Alexandre (2023): Revision of Heteroscelis Latreille, 1829 (Heteroptera: Pentatomidae: Asopinae) with the description of a new species from Colombia. Zootaxa 5278 (3): 401-438, DOI: 10.11646/zootaxa.5278.3.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.3.1
03BADF229324FFE3FF22368DFADD5CD2.taxon	materials_examined	Types examined BRAZIL. Platynopus bimaculatus Walker, 1867; male holotype; labels: “ PETROPOLIS Feb. 1857. J. Gray ” “ 6. Platynopus bimaculatus ”, “ Type ”, “ Holotype ”; “ NHMUK 010592394 ”; (NHMUK). Photo examined (Fig. 21). ARGENTINA. Heteroscelis doello-juradoi Pirán, 1961; male holotype; labels: “ HOLOTYPUS ”, “ Heteroscelis doello-juradoi Pirán 1959 ”, “ Provincia Jujuy ”; (MACN). Photo examined (Fig. 22). BOLIVIA. Heteroscelis denieri Pirán, 1961; male holotype; “ HOLOTYPUS ”, “ Heteroscelis Latr.? ”, “ Chirca 7. IV. 31 ”, “ Heteroscelis denieri Pirán 1959 ”, “ 1420 ”; (MLPA). Photo examined (Fig. 23 A, B). BOLIVIA. Heteroscelis durionei Pirán, 1961; female holotype; labels: “ HOLOTYPUS ”, “ Coroico ”, “ Heteroscelis durionei Pirán 1959 ”, “ 1421 ”; (MLPA). Photo examined (Fig. 23 C, D). Other material examined. BRAZIL, Rio de Janeiro: (Itatiaia, 700 m, - 22.41099, - 44.57621), 1 female, 1. IV. 1925 J. F. Zikán (CEIOC); ECUADOR, Napo: (Tiputini Biodiversity Station, 216 m, - 0.6379, - 76.1495), 1 female, 6. II. 1999 T. L. Erwin, et al. (USNM); PERU, Cuzco: (Pilcopata, - 14.13262, - 69.27723), 1 female (UFRG).	en	Sampaio, Vinicius Gomes, Roell, Talita, Campos, Luiz Alexandre (2023): Revision of Heteroscelis Latreille, 1829 (Heteroptera: Pentatomidae: Asopinae) with the description of a new species from Colombia. Zootaxa 5278 (3): 401-438, DOI: 10.11646/zootaxa.5278.3.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.3.1
03BADF229324FFE3FF22368DFADD5CD2.taxon	diagnosis	Diagnosis. Coria with a circular yellow spot near to the posterior margin (Fig. 20, A); urosternite III tubercle yellow or orange, from apex to the posterior margin of the urosternite; two pairs of yellow abdominal spots, the first pair lateral in urosternites VI and VII, and the second pair mesial in urosternite VII; vesicular area of female genitalia twice longer than the proximal part of the ductus receptaculi.	en	Sampaio, Vinicius Gomes, Roell, Talita, Campos, Luiz Alexandre (2023): Revision of Heteroscelis Latreille, 1829 (Heteroptera: Pentatomidae: Asopinae) with the description of a new species from Colombia. Zootaxa 5278 (3): 401-438, DOI: 10.11646/zootaxa.5278.3.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.3.1
03BADF229324FFE3FF22368DFADD5CD2.taxon	description	Redescription BODY: General color ranging from brown to iridescent purple; some specimens with yellow spots, callous or not, on the head, pronotum and scutellum. HEAD: Mandibular plates equal to or slightly surpassing clypeus, anteriorly rounded, separated at apex; head length anterior to the eyes equal to 2 / 3 of the total length of the head (Fig. 20, D); labium almost reaching the metasternum; proportion of labiomeres: I <II> III <IV; proportion of antennomeres: I <II <III ≅ IV ≅ V. THORAX: Coria longer than scutellum, not reaching the posterior margin of the connexival segment VI, uniformly punctured; membrane dark and surpassing posteriorly the abdomen; metapleural evaporatoria surpassing posteriorly the anterior limit of coxae; peritreme disc-shaped, surpassing half the pleural width; apex of peritreme rounded, posterior margin strongly arched, convex, anterior margin almost straight (Fig. 20, E); profemora with rudimentary ante-apical spine; foretibial length on average 3.5 x larger than the width of the tibial dorsal expansion (Fig. 20, F). ABDOMEN: Posterior angles of urosternites III to VII ending in minute spine (Fig. 20, B). Male GENITALIA (Fig. 21, C; Fig. 22, D): Pygophore with ventral rim slightly sinuous, shallowly concave medially. Female MEASUREMENTS (n = 4). Head length 1.04 ± 0.23 (0.75 – 1.32); width 1.37 ± 0.13 (1.25 – 1.52); pronotum length 1.70 ± 0.36 (1.38 – 2.16); width 4.21 ± 0.53 (3.78 – 4.88); scutellum length 2.46 ± 0.39 (2.20 – 3.04); width 2.35 ± 0.39 (2.01 – 2.88); length of antennomeres: I (n = 3) 0.31 ± 0.06 (0.25 – 0.37), II (n = 3) 0.67 ± 0.09 (0.56 – 0.75), III (n = 3) 0.79 ± 0.13 (0.69 – 0.94), IV (n = 2) 0.88 ± 0.17 (0.75 – 1.00); V (n = 1) 0.94; length of labiomeres: I 0.70 ± 0.11 (0.63 – 0.88), II 0.82 ± 0.15 (0.69 – 1.04), III 0.52 ± 0.09 (0.44 – 0.64), IV 0.67 ± 0.08 (0.63 – 0.80); width of foretibal expansion 0.53 ± 0.06 (0.50 – 0.56); width of abdomen 3.56 ± 0.45 (3.21 – 4.16); total length 6.26 ± 0.95 (5.41 – 7.60). GENITALIA (Fig. 24): Valvifers VIII posterior margin bisinuous, uniformly convex over the base of laterotergites IX; laterotergites VIII wider than long, with spiracles close to the lateral margin. Segment X with the posterior margin straight; valvulae IX with a median hourglass-shaped secondary thickening; capsula seminalis oval.	en	Sampaio, Vinicius Gomes, Roell, Talita, Campos, Luiz Alexandre (2023): Revision of Heteroscelis Latreille, 1829 (Heteroptera: Pentatomidae: Asopinae) with the description of a new species from Colombia. Zootaxa 5278 (3): 401-438, DOI: 10.11646/zootaxa.5278.3.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.3.1
03BADF229324FFE3FF22368DFADD5CD2.taxon	distribution	Distribution: Panama, Ecuador (new record), French Guyana, Brazil, Peru, Bolivia and Argentina (Fig. 1).	en	Sampaio, Vinicius Gomes, Roell, Talita, Campos, Luiz Alexandre (2023): Revision of Heteroscelis Latreille, 1829 (Heteroptera: Pentatomidae: Asopinae) with the description of a new species from Colombia. Zootaxa 5278 (3): 401-438, DOI: 10.11646/zootaxa.5278.3.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.3.1
03BADF229328FFE6FF2233E4FC6459BA.taxon	description	(Figs. 1, 25 – 29)	en	Sampaio, Vinicius Gomes, Roell, Talita, Campos, Luiz Alexandre (2023): Revision of Heteroscelis Latreille, 1829 (Heteroptera: Pentatomidae: Asopinae) with the description of a new species from Colombia. Zootaxa 5278 (3): 401-438, DOI: 10.11646/zootaxa.5278.3.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.3.1
03BADF229328FFE6FF2233E4FC6459BA.taxon	materials_examined	Other material examined. BRAZIL, Amazonas: (Manaus, Conj. Manca. - 3.04361, - 60.01282), 1 female, 10. V. 1988 P. E. S. Silva (INPA); Roraima: (Boa Vista, 3.18861, - 60.61212), 1 female, 31. V. 1974 (UFRG); Pará: (Peixe-Boi, - 1.10168, - 47.27259), 1 female, 12. IV. 1977 P Waldir (UFRG); (Marituba, - 1.37399, - 48.3142), 1 female, 07. VI. 1961 J & B. Bechyné (UFRG); Mato Grosso: (Chapada dos Guimar „ es, - 15.02015, - 55.53735), 1 female, 20. I. 1961 J & B. Bechyné (UFRG); São Paulo: 3 females, J. Mráz (NMPC); Rio Grande do Sul: (Salvador do Sul, - 29.48006, - 51.51407), 1 male, 16. X. 1994 (MCTP); COLOMBIA, (Meta, San Martín, Reserva Natural El Caduceo, Km 4.5, Via San Francisco, 3.671389, - 73.659444), 1 male, 30. IX – 4. X. 2013 M Arango (MPUJ _ ENT).	en	Sampaio, Vinicius Gomes, Roell, Talita, Campos, Luiz Alexandre (2023): Revision of Heteroscelis Latreille, 1829 (Heteroptera: Pentatomidae: Asopinae) with the description of a new species from Colombia. Zootaxa 5278 (3): 401-438, DOI: 10.11646/zootaxa.5278.3.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.3.1
03BADF229328FFE6FF2233E4FC6459BA.taxon	diagnosis	Diagnosis. Body robust compared to the other species, dark-brown to black; unpunctured, yellowish longitudinal line extending from the anterior margin of the pronotum to the posterior margin of the scutellum; radial vein of coria usually yellowish; foretibial length on average ten times larger than the width of the dorsal tibial expansion (Fig. 25, F); capsula seminalis digitiform.	en	Sampaio, Vinicius Gomes, Roell, Talita, Campos, Luiz Alexandre (2023): Revision of Heteroscelis Latreille, 1829 (Heteroptera: Pentatomidae: Asopinae) with the description of a new species from Colombia. Zootaxa 5278 (3): 401-438, DOI: 10.11646/zootaxa.5278.3.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.3.1
03BADF229328FFE6FF2233E4FC6459BA.taxon	description	Redescription HEAD: Mandibular plates equal to or slightly surpassing clypeus, rounded at apex, never touching each other; head length anterior to the eyes smaller than 2 / 3 of the total length of the head (Fig. 25, D); labium reaching the metasternum; proportion of labiomeres: I <II> III ≅ IV; proportion of antennomeres: I <II <III ≅ IV ≅ V. THORAX: Coria longer than the scutellum, not reaching the posterior margin of the connexival segment VI; metapleural evaporatoria not surpassing posteriorly the anterior limit of coxae; peritreme disc-shaped, surpassing half the pleural width; apex of peritreme rounded, posterior margin arched, convex, anterior margin almost straight (Fig. 25, E). ABDOMEN: Uniformly punctured; urosternite III tubercle not surpassing the posterior margin of metasternum; posterior angles of urosternites III to VII ending in minute spine, more developed in VII (Fig. 25, B). Male: MEASUREMENTS (n = 2). Head length 1.24 ± 0.05 (1.20 – 1.28); width 1.48 ± 0.05 (1.44 – 1.52); pronotum length 1.88 ± 0.16 (1.76 – 2.00); width 4.12 ± 0.16 (4.00 – 4.24); scutellum length 2.60 ± 0.05 (2.56 – 2.64); width 2.60 ± 0.05 (2.24 – 2.40); length of antennomeres: I (n = 1) 0.32, II 0.64 ± 0.11 (0.56 – 0.72), III 0.84 ± 0.05 (0.80 – 0.88), IV 0.96 ± 0.11 (0.88 – 1.40); V (n = 1) 1.12; length of labiomeres: I 0.68 ± 0.05 (0.64 – 0.72), II 0.72 ± 0.00 (0.72 – 0.72), III 0.40 ± 0.00 (0.40 – 0.40), IV 0.48 ± 0.00 (0.48 – 0.48); width of foretibal expansion 0.20 ± 0.05 (0.16 – 0.24); width of abdomen 3.40 ± 0.28 (3.20 – 3.60); total length (n = 1) 7.20. GENITALIA: Pygophore (Fig. 27): setae denser on medial portion of dorsal rim; dorsal rim sinuous, medially concave; ventral rim sinuous, shallowly concave medially; segment X setose on the entire surface; heads of parameres lateroventrally directed in posterior view. Phallus (Fig. 28): conjunctival dorsal lobe with one pair of long projections, ventral lobes with two projections each. Female: MEASUREMENTS (n = 8). Head length 1.12 ± 0.09 (1.00 – 1.28); width 1.40 ± 0.05 (1.44 – 1.53); pronotum length 1.96 ± 0.21 (1.76 – 2.20); width 4.42 ± 0.33 (4.16 – 5.00); scutellum length 2.99 ± 0.26 (2.72 – 3.50); width 2.66 ± 0.25 (2.40 – 3.10); length of antennomeres: I (n = 6) 0.38 ± 0.03 (0.32 – 0.44), II (n = 4) 0.71 ± 0.10 (0.60 – 0.80), III (n = 4) 0.98 ± 0.09 (0.88 – 1.10), IV (n = 1) 1.12; V (n = 1) 1.04; length of labiomeres: I (n = 7) 0.76 ± 0.07 (0.70 – 0.90), II (n = 7) 0.90 ± 0.11 (0.80 – 1.10), III (n = 7) 0.40 ± 0.05 (0.32 – 0.50), IV (n = 7) 0.50 ± 0.04 (0.48 – 0.60); width of foretibal expansion (n = 5) 0.25 ± 0.06 (0.16 – 0.30); width of abdomen 4.06 ± 0.25 (3.68 – 4.50); total length (n = 6) 7.67 ± 0.88 (6.56 – 8.90). GENITALIA (Fig. 29): Valvifers VIII posterior margin bisinuous, truncate over the laterotergites VIII, and slightly projected over the basal angles of laterotergites IX, setae on sutural margins; laterotergites VIII slightly longer than wide, with spiracles close to the lateral margin; segment X with the posterior margin straight; valvulae IX with 1 + 1 secondary thickenings, these bean-shaped with the anterior margin elongated with acute apices; vesicular area half the length of the proximal part of the ductus receptaculi.	en	Sampaio, Vinicius Gomes, Roell, Talita, Campos, Luiz Alexandre (2023): Revision of Heteroscelis Latreille, 1829 (Heteroptera: Pentatomidae: Asopinae) with the description of a new species from Colombia. Zootaxa 5278 (3): 401-438, DOI: 10.11646/zootaxa.5278.3.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.3.1
03BADF229328FFE6FF2233E4FC6459BA.taxon	distribution	Distribution: Paraguay, Colombia (new record) and Brazil (Fig. 1).	en	Sampaio, Vinicius Gomes, Roell, Talita, Campos, Luiz Alexandre (2023): Revision of Heteroscelis Latreille, 1829 (Heteroptera: Pentatomidae: Asopinae) with the description of a new species from Colombia. Zootaxa 5278 (3): 401-438, DOI: 10.11646/zootaxa.5278.3.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.3.1
03BADF22932FFFDAFF2233B5FD9E5A92.taxon	description	(Figs. 1, 30 – 32)	en	Sampaio, Vinicius Gomes, Roell, Talita, Campos, Luiz Alexandre (2023): Revision of Heteroscelis Latreille, 1829 (Heteroptera: Pentatomidae: Asopinae) with the description of a new species from Colombia. Zootaxa 5278 (3): 401-438, DOI: 10.11646/zootaxa.5278.3.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.3.1
03BADF22932FFFDAFF2233B5FD9E5A92.taxon	materials_examined	Type Material Holotype: male, here designated. COLOMBIA. Santander. Charalá. Virolin. Mar 1981. 18 - III. Santander. Charalá. Virolin. Heteroscelis lepida Stål, 1862 — ICN. Det: F. Rua Alvear, 2006. ICN 055894. (6.1080, - 73.2004). Deposited in: ICN (Fig. 30). Paratype: male, here designated. COLOMBIA. Boyacá. Villa de Leyva. SFF Iguaque. 05 ° 38 ’ N 73 ° 29 ’ W WGS 84. 2450 m. Captura manual. 19. VIII. 1998. J. Castro. IAvH-E-VillaDeLeyva. Deposited in: IAvH-E (Fig. 31).	en	Sampaio, Vinicius Gomes, Roell, Talita, Campos, Luiz Alexandre (2023): Revision of Heteroscelis Latreille, 1829 (Heteroptera: Pentatomidae: Asopinae) with the description of a new species from Colombia. Zootaxa 5278 (3): 401-438, DOI: 10.11646/zootaxa.5278.3.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.3.1
03BADF22932FFFDAFF2233B5FD9E5A92.taxon	etymology	Etymology. Named after the father of the first author, Vandemberg de Freitas Sampaio (in memoriam), who made a great effort to provide a good education for his children, but unfortunately could not reap the fruits he sowed.	en	Sampaio, Vinicius Gomes, Roell, Talita, Campos, Luiz Alexandre (2023): Revision of Heteroscelis Latreille, 1829 (Heteroptera: Pentatomidae: Asopinae) with the description of a new species from Colombia. Zootaxa 5278 (3): 401-438, DOI: 10.11646/zootaxa.5278.3.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.3.1
03BADF22932FFFDAFF2233B5FD9E5A92.taxon	diagnosis	Diagnosis. Head, thorax and abdomen orange to red, coria iridescent-blue, radial vein usually reddish; foretibial length on average 9.5 x larger than the width of the dorsal expansion (Fig. 30, F).	en	Sampaio, Vinicius Gomes, Roell, Talita, Campos, Luiz Alexandre (2023): Revision of Heteroscelis Latreille, 1829 (Heteroptera: Pentatomidae: Asopinae) with the description of a new species from Colombia. Zootaxa 5278 (3): 401-438, DOI: 10.11646/zootaxa.5278.3.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.3.1
03BADF22932FFFDAFF2233B5FD9E5A92.taxon	description	Description HEAD: Mandibular plates slightly surpassing clypeus, anteriorly rounded, separated at apex; head length anterior to the eyes shorter than 2 / 3 of the total length of the head (Fig. 30, D); proportion of labiomeres: I <II> III <IV; proportion of antennomeres: I <II <III, IV and V missing. THORAX: Coria iridescent-blue, uniformly punctured, longer than the scutellum, not reaching the posterior margin of connexival segment VI; radial vein usually reddish; membrane dark, surpassing posteriorly the abdomen; metapleural evaporatoria not surpassing posteriorly the anterior limit of coxae; peritreme disc-shaped, surpassing half the pleural width; apex of peritreme rounded, posterior margin arched, convex, anterior margin almost straight (Fig. 30, E). ABDOMEN: Uniformly punctured; urosternite III tubercle not surpassing the posterior margin of metasternum; posterior angles of urosternites III to VII ending in minute spine (Fig. 30, B). Male: MEASUREMENTS (n = 1). Head length 1.52; width 1.36; pronotum length 1.84; width 4.08; scutellum length 2.48; width 3.04; length of antennomeres: I 0.32, II 0.64, III 1.04, IV –, V –; length of labiomeres: I 0.60, II 0.90, III 0.40, IV 0.50; width of foretibal expansion 0.24; width of abdomen 3.60; total length 6.72. GENITALIA: Pygophore (Fig. 32): setae denser on posterolateral angles; dorsal rim concave; ventral rim sinuous, medially concave; segment X with surface corrugated; head of parameres laterodorsally directed in posterior view.	en	Sampaio, Vinicius Gomes, Roell, Talita, Campos, Luiz Alexandre (2023): Revision of Heteroscelis Latreille, 1829 (Heteroptera: Pentatomidae: Asopinae) with the description of a new species from Colombia. Zootaxa 5278 (3): 401-438, DOI: 10.11646/zootaxa.5278.3.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.3.1
03BADF22932FFFDAFF2233B5FD9E5A92.taxon	discussion	Remarks: This species is quite similar to H. lepida, however, without considering the difference in color, H. vandembergi sp. nov. also differs from H. lepida in the proportion of the area anterior to the eyes, and in the proportion of the dorsal foretibial expansions in relation to the femur, a trait that does not vary within the other species of the genus. Female unknown.	en	Sampaio, Vinicius Gomes, Roell, Talita, Campos, Luiz Alexandre (2023): Revision of Heteroscelis Latreille, 1829 (Heteroptera: Pentatomidae: Asopinae) with the description of a new species from Colombia. Zootaxa 5278 (3): 401-438, DOI: 10.11646/zootaxa.5278.3.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.3.1
03BADF22932FFFDAFF2233B5FD9E5A92.taxon	distribution	Distribution: Colombia (Fig. 1).	en	Sampaio, Vinicius Gomes, Roell, Talita, Campos, Luiz Alexandre (2023): Revision of Heteroscelis Latreille, 1829 (Heteroptera: Pentatomidae: Asopinae) with the description of a new species from Colombia. Zootaxa 5278 (3): 401-438, DOI: 10.11646/zootaxa.5278.3.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.3.1
