identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03BB87AA0B426373FF0827C5FAA0F866.text	03BB87AA0B426373FF0827C5FAA0F866.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euconnus Thomson	<div><p>Genus Euconnus Thomson</p><p>Euconnus Thomson, 1859: 61 .</p><p>Type species: Pselaphus hirticollis Illiger, 1798 (des. orig.).</p><p>Remarks. The subgeneric system of Euconnus was a subject of a long series of publications (Jałoszyński 2012, 2013b, 2015a, b, c, 2016a, b, c, d, 2017a, b, c, d, e, f, 2018a, b, 2019a, 2020a, b, 2021a, b, 2022b, Jałoszyński &amp; Newton 2017). Many taxonomic issues have been solved, but the current subgeneric division of Euconnus is not phylogeny-based and certainly will require profound changes.</p><p>Euconnus is characterized by the following set of features: eyes (if present) situated closer to mandibular bases and antennal fossae than to occipital constriction; lateral submental sutures lacking; mesoventral intercoxal process present, carinate; notosternal sutures complete; hypomeral ridges at least partly developed or at least inner regions of hypomera delimited from outer regions by abrupt change in microsculpture; each elytron with two asetose basal pits (reduced in some species); metaventral intercoxal process widely or narrowly (but clearly) separating metacoxae; and aedeagus with free rod-like parameres. Euconnus can be identified using a preliminary key (and comments related to the yet unclear diagnosis) to world Stenichnini genera published recently (Jałoszyński 2024).</p></div>	https://treatment.plazi.org/id/03BB87AA0B426373FF0827C5FAA0F866	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jałoszyński, Paweł	Jałoszyński, Paweł (2024): Subgenera Cladoconnus Reitter and Tetramelus Motschulsky of Euconnus Thomson discovered in China (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 5514 (3): 232-256, DOI: 10.11646/zootaxa.5514.3.2, URL: http://dx.doi.org/10.11646/zootaxa.5514.3.2
03BB87AA0B416370FF0822B6FCE5FD71.text	03BB87AA0B416370FF0822B6FCE5FD71.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euconnus (Cladoconnus) Reitter	<div><p>Subgenus Cladoconnus Reitter</p><p>Cladoconnus Reitter, 1909: 226 (as subgenus of Euconnus). Type species: Scydmaenus motschulskii Sturm, 1838 (des. by Franz in Newton &amp; Franz 1998).</p><p>Remarks. Morphological characters were illustrated and the diagnosis emended in Jałoszyński (2018b); species of the Balkans, Turkey and Caucasus were revised in Hlaváč &amp; Stevanović (2013), Eastern Palaearctic species were treated by Hoshina (2004), Hoshina &amp; Park (2020), Jałoszyński (2019b), and Kurbatov (1988), and recently the North American species were studied by Caterino (2022). One extinct species is known from Eocene Rovno amber (Jałoszyński &amp; Perkovsky 2021).</p><p>Distribution of extant species. Asia: Japan (Honshu, Shikoku), Russian Far East (Primorie), South Korea, Turkey; Europe: Albania, Armenia, Austria, Belgium, Bosnia &amp; Herzegovina, Croatia, Czech Rep., Denmark, Estonia, France (incl. Corsica), Germany, Great Britain, Georgia, Greece (incl. Rhodos), Hungary, Italy (incl. Elba), Latvia, Lithuania, Macedonia, Montenegro, Netherlands, Poland, Portugal, Romania, Russia, Serbia, Slovakia, Slovenia, Spain, Sweden, Switzerland, Ukraine, ‘Yugoslavia’; North America: USA (Alabama, Georgia, North Carolina, South Carolina, Tennessee).</p><p>Composition. 51 species (incl. 1 extinct) and 3 subspecies.</p></div>	https://treatment.plazi.org/id/03BB87AA0B416370FF0822B6FCE5FD71	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jałoszyński, Paweł	Jałoszyński, Paweł (2024): Subgenera Cladoconnus Reitter and Tetramelus Motschulsky of Euconnus Thomson discovered in China (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 5514 (3): 232-256, DOI: 10.11646/zootaxa.5514.3.2, URL: http://dx.doi.org/10.11646/zootaxa.5514.3.2
03BB87AA0B416371FF082118FC25FC16.text	03BB87AA0B416371FF082118FC25FC16.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euconnus (Cladoconnus) pingpong Jałoszyński 2024	<div><p>Euconnus (Cladoconnus) pingpong sp. nov.</p><p>(Figs 1, 3, 5–8)</p><p>Material studied. Holotype: ♂ (CHINA: Gansu): "CHINA: S-Gansu [CH2-09] / W-Qinling <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.166664&amp;materialsCitation.latitude=34.053886" title="Search Plazi for locations around (long 104.166664/lat 34.053886)">Shan</a>, 101 km NW Longnan, / 34°03’14’’N, 104°10’00’’E, 2200 m, / SW-slope with shrubs, litter sifted / 1.VIII.2012, leg. M. Schülke " [white, printed], " EUCONNUS / ( CLADOCONNUS) / pingpong m. / P. Jałoszyński, 2024 / HOLOTYPUS" [red, printed] (MNHB).</p><p>Diagnosis. Ventral plate of median lobe shaped like table tennis paddle, with slender distal ‘handle’ and broad proximal part; endophallus with three asymmetrical sclerites, of which one is extremely long and projecting from ostium, while two others are extremely short and situated in proximal region of endophallus, in ventral view left long sclerite is nearly straight except hook-like curved apex, left short sclerite is simple, situated at base of long sclerite and directed mesally and slightly distally, and right short sclerite is bifurcate Y-shaped with both apices directed distally and slightly laterally and with its proximal stalk touching base of long left sclerite.</p><p>Description. Body of male (Fig. 1) elongate but moderately slender, moderately convex, with long appendages, BL 1.68 mm; cuticle glossy, pigmentation dark brown, head and pronotum indistinctly darker than elytra, vestiture of setae light brown.</p><p>Head in dorsal view round, broadest across eyes, HL 0.30 mm, HW 0.33 mm; tempora in dorsal view nearly twice as long as eyes, but in lateral view each temple subequal to longest diameter of eye; vertex and frons confluent, weakly and evenly convex, vertex slightly bulging posterodorsally; supraantennal tubercles small but distinctly elevated. Eyes moderately large, coarsely faceted, distinctly but not strongly projecting laterally from head silhouette, in lateral view oval. Punctures on vertex and frons inconspicuous, small and sparse; setae short and sparse, suberect, thick bristles present on vertex and tempora. Antennae (Fig. 3) slender, about as long as half BL, AnL 0.90 mm; antennomeres 1–2 each elongate, 3 indistinctly elongate, 4–7 each about as long as broad, 8 modified, at base broader than 7, elongate, with its mesal margin straight and with strongly projecting mesal proximal corner, mesal distal corner not projecting, antennomere 9 modified, as long as 8, with its mesal margin indistinctly concave and its mesal distal corner distinctly projecting, 10 distinctly shorter than 9, about as long as broad, 11 longer than 10 but slightly shorter than 9–10 combined, about twice as long as broad.</p><p>Pronotum broadest in front of middle, PL 0.40 mm, PW 0.36 mm; anterior margin arcuate, lateral margins rounded in anterior half and sinuate in posterior half, posterior corners slightly obtuse-angled, blunt. Pronotal base with narrow indistinct median longitudinal wrinkle flanked by pair of large pits, each posterior corner with short and fine sublateral carina and pit between carina and lateral margin. Punctures on pronotal disc inconspicuous; setae moderately dense and long, suberect and mixed with thick bristles.</p><p>Elytra oval, broadest slightly in front of middle, EL 0.98 mm, EW 0.73 mm, EI 1.34; elongate adsutural area in anterior third slightly impressed; basal impressions deep, humeral calli prominent, elytral apices separately rounded. Punctures on elytral disc inconspicuous, superficial; setae longer than those on pronotum, moderately dense and suberect to erect. Hind wings long, functional.</p><p>Legs long and slender, unmodified.</p><p>Aedeagus (Figs 5–8) stout, drop-shaped, AeL 0.38 mm; median lobe broadest near middle, rapidly narrowed in distal half, with dorsal apical plate elongate subtriangular with nearly straight lateral margins in distal region; ventral apical plate shaped like table tennis paddle, with broad proximal part and slender abruptly delimited distal ‘handle’ broadly rounded at apex; endophallus with three asymmetrical sclerites, of which one is extremely long and projecting from ostium, while two others are extremely short and situated in proximal region of endophallus, in ventral view left long sclerite is nearly straight except hook-like curved apex, left short sclerite is simple, situated at base of long sclerite and directed mesally and slightly distally, and right short sclerite is bifurcate Y-shaped with both apices directed distally and slightly laterally and with its proximal stalk touching base of long left sclerite. Setae on dorsal wall short and visible only in lateral view (Fig. 8); parameres broad and short, slightly broadened in subapical region, each with two long apical setae.</p><p>Female. Unknown.</p><p>Distribution. Central China: SE Gansu.</p><p>Etymology. The name pingpong (noun in apposition) refers to the ventral plate of the aedeagus shaped like a table tennis paddle.</p><p>Remarks. Eastern Palaearctic species of Cladoconnus are externally difficult or impossible to identify, but the aedeagus of each strongly differs from those of all remaining species. The aedeagus of E. pingpong is the only one with a ping pong paddle-shaped ventral plate. This species has the aedeagus strongly differing from those in the Eastern Palaearctic E. ussuriensis species group (Jałoszyński 2019b).</p></div>	https://treatment.plazi.org/id/03BB87AA0B416371FF082118FC25FC16	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jałoszyński, Paweł	Jałoszyński, Paweł (2024): Subgenera Cladoconnus Reitter and Tetramelus Motschulsky of Euconnus Thomson discovered in China (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 5514 (3): 232-256, DOI: 10.11646/zootaxa.5514.3.2, URL: http://dx.doi.org/10.11646/zootaxa.5514.3.2
03BB87AA0B406377FF0821F2FDCFFDF6.text	03BB87AA0B406377FF0821F2FDCFFDF6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euconnus (Cladoconnus) sagittarius Jałoszyński 2024	<div><p>Euconnus (Cladoconnus) sagittarius sp. nov.</p><p>(Figs 2, 4, 9–12)</p><p>Material studied. Holotype: ♂ (CHINA: Gansu): "CHINA: S-Gansu [CH12-07] / Mts. 36 km SE <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.25278&amp;materialsCitation.latitude=33.22222" title="Search Plazi for locations around (long 105.25278/lat 33.22222)">Longnan</a>, / 33°13’20’’N, 105°15’10’’E, 2170 m, / N-slope with shrubs and scattered / coniferous trees, litter &amp; mushrooms / sifted, 31.VII.2012, leg. M. Schülke " [white, printed], " EUCONNUS / ( CLADOCONNUS) / sagittarius m. / P. Jałoszyński, 2024 / HOLOTYPUS" [red, printed] (MNHB) . Paratypes: 2 ♀♀, same data as for holotype (cPJ, MNHB) .</p><p>Diagnosis. Ventral plate of median lobe broad and arrowhead-shaped with proximal and distal regions subequal in width; endophallus with three asymmetrical sclerites, of which two are extremely long and one short and situated in proximal region of endophallus, in ventral view left long sclerite weakly curved except strongly curved apex, median long sclerite bifurcate, in distal half divided into two slender and pointed arms parallel to long axis of aedeagus, and right short sclerite bifurcate L-shaped with distal arm directed distally and laterally, and proximal arm transverse in relation to long axis of aedeagus and directed laterally.</p><p>Description. Body of male (Fig. 2) elongate but moderately slender, moderately convex, with long appendages, BL 1.68 mm; cuticle glossy, pigmentation dark brown, head and pronotum slightly darker than elytra, vestiture of setae light brown.</p><p>Head in dorsal view round, broadest across eyes, HL 0.30 mm, HW 0.33 mm; tempora in dorsal view nearly twice as long as eyes, but in lateral view each temple subequal to longest diameter of eye; vertex and frons confluent, weakly and evenly convex, vertex slightly bulging posterodorsally; supraantennal tubercles small but distinctly elevated. Eyes moderately large, coarsely faceted, distinctly but not strongly projecting laterally from head silhouette, in lateral view oval. Punctures on vertex and frons inconspicuous, small and sparse; setae short and sparse, suberect, thick bristles present on vertex and tempora. Antennae (Fig. 4) slender, indistinctly longer than half BL, AnL 0.98 mm; antennomeres 1–2 each elongate, 3 indistinctly elongate, 4–7 each about as long as broad, 8 modified, at base broader than 7, elongate, with its mesal margin concave and with strongly projecting mesal proximal corner, mesal distal corner weakly projecting, antennomere 9 modified, indistinctly longer than 8, with its mesal margin concave and its mesal distal corner distinctly projecting, 10 distinctly shorter than 9, about as long as broad, 11 longer than 10 but much shorter than 9–10 combined, about twice as long as broad.</p><p>Pronotum broadest in front of middle, PL 0.40 mm, PW 0.36 mm; anterior margin arcuate, lateral margins rounded in anterior half and sinuate in posterior half, posterior corners slightly obtuse-angled, blunt. Pronotal base with barely marked median longitudinal wrinkle separating pair of large pits, each posterior corner with short and fine sublateral carina and pit between carina and lateral margin. Punctures on pronotal disc inconspicuous; setae moderately dense and long, suberect and mixed with thick bristles.</p><p>Elytra oval, broadest slightly in front of middle, EL 0.98 mm, EW 0.73 mm, EI 1.34; elongate adsutural area in anterior third flattened, but not impressed; basal impressions deep, humeral calli prominent, elytral apices separately rounded. Punctures on elytral disc inconspicuous, superficial; setae longer than those on pronotum, moderately dense and suberect to erect. Hind wings long, functional.</p><p>Legs long and slender, unmodified.</p><p>Aedeagus (Figs 9–12) stout, drop-shaped, AeL 0.38 mm; median lobe broadest near middle, rapidly narrowed in distal half, with dorsal apical plate elongate subtriangular with convex lateral margins; ventral apical plate arrowhead-shaped with remarkably large distal region; endophallus with three asymmetrical sclerites, of which two are extremely long and one short and situated in proximal region of endophallus, in ventral view left long sclerite weakly curved except strongly curved apex, median long sclerite bifurcate, in distal half divided into two slender and pointed arms parallel to long axis of aedeagus, and right short sclerite bifurcate L-shaped with distal arm directed distally and laterally, and proximal arm transverse in relation to long axis of aedeagus and directed laterally. Setae on dorsal wall short and visible only in lateral view (Fig. 12); parameres broad and short, each with two long apical setae.</p><p>Female. Similar to male but slightly larger, antennae shorter in relation to BL and with unmodified antennomeres 8 and 9, and with distinctly smaller eyes, in dorsal view tempora almost 3 times as long as eye. BL 1.73–1.75 mm; HL 0.33–0.35 mm, HW 0.33–0.35 mm, AnL 0.85 mm; PL 0.43 mm, PW 0.36–0.38 mm; EL 0.98 mm, EW 0.70– 0.73 mm, EI 1.34–1.39.</p><p>Distribution. Central China: SE Gansu.</p><p>Etymology. The name sagittarius (noun in apposition) refers to the ventral plate of the aedeagus being shaped like an arrowhead.</p><p>Remarks. Euconnus sagittarius has the ventral plate of the median lobe arrowhead-shaped, and this structure is similar to those known in the members of the E. ussuriensis species group: E. ussuriensis Kurbatov, 1988 (Russian Far East), E. nakahamai Hoshina &amp; Miyata, 2018 (in Hoshina et al. 2018) (Shikoku), and E. rudimentalis Jałoszyński, 2019b (South Korea). However, in these species the ventral plate of the aedeagus has the arrowhead apex less than half as wide as its base, while in E. sagittarius the distal region is almost as wide as the proximal portion. Moreover, the distal arrowhead region in E. sagittarius is much longer than in the remaining species, and the shapes and arrangement of endophallic sclerites is different. Interestingly, males of E. pingpong and E. sagittarius have almost all measurements identical, with the exception of AnL, which is slightly greater for the latter species. They differ externally in antennal modifications, E. sagittarius having more concave mesal margins of antennomeres 8 and 9, and their aedeagi are remarkably different.</p></div>	https://treatment.plazi.org/id/03BB87AA0B406377FF0821F2FDCFFDF6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jałoszyński, Paweł	Jałoszyński, Paweł (2024): Subgenera Cladoconnus Reitter and Tetramelus Motschulsky of Euconnus Thomson discovered in China (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 5514 (3): 232-256, DOI: 10.11646/zootaxa.5514.3.2, URL: http://dx.doi.org/10.11646/zootaxa.5514.3.2
03BB87AA0B466374FF08253BFB7DFC16.text	03BB87AA0B466374FF08253BFB7DFC16.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euconnus (Tetramelus) Motschulsky	<div><p>Subgenus Tetramelus Motschulsky</p><p>Tetramelus Motschulsky, 1869: 257. Type species: Scydmaenus oblongus Sturm, 1838 (des. by Franz in Newton &amp; Franz (1998): 147).</p><p>Remarks. Morphological structures were illustrated and an emended diagnosis was given in Jałoszyński (2015c); further notes on character variability can be found in Jałoszyński (2017a, 2021a).</p><p>Tetramelus, as currently defined, can be identified using the following set of co-occurring characters: hypostomal ridges running posteromesally; head ventrally lacking deep narrow longitudinal groove connecting gular plate and submentum; pronotum lacking longitudinal hypomeral grooves; pronotum broadest in front of middle; antenna in male unmodified, gradually thickened or with variously distinct tetramerous club; and tarsomere 4 in all legs lacking long distoventral lobe (for key to subgenera see Jałoszyński (2022b)).</p><p>Distribution. Africa: Algeria, Cameroon, Comoros, Kenya, Lesotho, Madagascar, RSA, Swaziland, Tanzania; Asia: Asian Turkey, Nagorno-Karabakh, Nepal, “Syria”; Australia: Australia (incl. Franklin Is., Lord Howe Is., Reevesby Is., Tasmania); Caribbean: Jamaica; Europe: Albania, Andorra, Armenia, Austria, Belgium, Bosnia &amp; Herzegovina, Bulgaria, Croatia, Czech Rep., France, Georgia, Germany, Greece, Hungary, Italy, Kosovo, Lithuania, Macedonia, Montenegro, Netherlands, Poland, Portugal, Romania, European Russia, Slovakia, Slovenia, Spain, Switzerland, European Turkey, Ukraine; North America: Canada, USA; Pacific: New Caledonia, New Zealand; South America: Bolivia, Brazil, Peru.</p><p>Composition. At the time of submission of the present study there were 233 species and 2 subspecies. However, in a parallel study on Malagasy species some Stenichnini are transferred into Tetramelus (Jałoszyński, in press), which results in 17 more species included in this subgenus.</p><p>All new Chinese species described below are microphthalmous, wingless and have reduced wing-related exoskeletal structures, including lack of humeral calli and the elytra strongly narrowing anteriorly; males have a large and deep median impression on the metaventrite, and all species except one have modified trochanters. All species are externally very similar, they differ in body size, pigmentation, modifications of male trochanters, and genital structures. The Chinese Tetramelus species are morphologically so close to their Western Palaearctic congeners that they can be included in the same Euconnus oblongus group (Jałoszyński 2021a).</p></div>	https://treatment.plazi.org/id/03BB87AA0B466374FF08253BFB7DFC16	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jałoszyński, Paweł	Jałoszyński, Paweł (2024): Subgenera Cladoconnus Reitter and Tetramelus Motschulsky of Euconnus Thomson discovered in China (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 5514 (3): 232-256, DOI: 10.11646/zootaxa.5514.3.2, URL: http://dx.doi.org/10.11646/zootaxa.5514.3.2
03BB87AA0B45637BFF0821E9FEFBFED6.text	03BB87AA0B45637BFF0821E9FEFBFED6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euconnus (Tetramelus) punctator Jałoszyński 2024	<div><p>Euconnus (Tetramelus) punctator sp. nov.</p><p>(Figs 13, 21, 31–38)</p><p>Material studied. Holotype: ♂ (CHINA: Sichuan): “CHINA: N-Sichuan [CH12-24] / pass 35 km NNW <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.43222&amp;materialsCitation.latitude=32.925556" title="Search Plazi for locations around (long 103.43222/lat 32.925556)">Songpan</a> / 32°55’32’’N, 103°25’56’’E, 3600 m, / moist N-slope with Salix and other / shrubs, litter, grass roots &amp; moss / sifted, 11.VIII.2012, leg. M. Schülke ” [white, printed], “ EUCONNUS / ( TETRAMELUS) / punctator m. / P. Jałoszyński, 2024 / HOLOTYPUS” [red, printed] (MNHB) . Paratypes: 8 ♀♀, same data as for holotype (cPJ, MNHB) .</p><p>Diagnosis. Body rufous brown; male metatrochanter distally produced into long and slender subtriangular spine, ventral margin of trochanter evenly concave in distal half; aedeagus in ventral view about twice as long as wide, its ventral plate wider than long, with sinuate lateral margins and rounded apex, dorsal plate elongate subtriangular with broadly rounded apex, endophallus in ventral view with one elongate and curved sclerite surrounded by membranous folds densely covered with cuticular spines.</p><p>Description. Body of male (Fig. 13) elongate and slender, moderately convex, with long appendages, BL 2.04 mm; cuticle glossy, pigmentation uniformly rufous brown, vestiture of setae lighter than cuticle.</p><p>Head in dorsal view round, broadest across eyes, HL 0.34 mm, HW 0.31 mm; in lateral view each temple 3 times as long as eye; vertex weakly convex and only slightly bulging posterodorsally; frons flattened; supraantennal tubercles small and indistinctly elevated. Eyes small and finely faceted, hardly projecting laterally from head silhouette, in lateral view oval. Punctures on vertex and frons inconspicuous, small and sparse; setae short and sparse, suberect, sparse and short thick bristles present on vertex and tempora.Antennae slender, indistinctly shorter than half BL, AnL 0.85 mm; antennomeres 1–5 each elongate (5 weakly so), 6–8 each about as long as broad, 9 and 10 each slightly transverse, 11 distinctly shorter than 9–10 combined, about 1.6 times as long as broad.</p><p>Pronotum broadest in front of middle, PL 0.50 mm, PW 0.40 mm; anterior margin arcuate, lateral margins rounded in anterior half and nearly straight in posterior third, posterior corners slightly obtuse-angled, blunt; posterior margin weakly rounded. Pronotal base with single outer pair of small pits. Punctures on pronotal disc inconspicuous; setae moderately dense and short, suberect and on sides mixed with thick bristles.</p><p>Elytra oval, broadest distinctly in front of middle, EL 1.20 mm, EW 0.75 mm, EI 1.60; basal impressions and humeral calli lacking, elytral apices rounded together. Punctures on elytral disc inconspicuous, superficial; setae similar to those on pronotum, moderately dense and weakly suberect.</p><p>Legs long and slender, metatrochanter (Fig. 21) modified, with long distal spine, ventral margin of trochanter nearly evenly concave in distal half.</p><p>Metaventrite with deep oval median impression occupying most of its surface, extending on almost entire length.</p><p>Aedeagus (Figs 31–38) stout, AeL 0.50 mm; median lobe in ventral view broadest near distal third, and rapidly narrowed here, ventral plate broader than long, with strongly sinuate lateral margins and narrow rounded apex, dorsal plate elongate subtriangular with lateral margins rounded in apical half, apex broadly rounded; endophallus in ventral view with one elongate and curved sclerite surrounded by membranous folds densely covered with cuticular spines. In lateral view (Fig. 37) short setae can be seen on dorsal wall in subapical region; parameres slender, each with 3 long subapical setae.</p><p>Female. Similar to male (with similarly small eyes), lacking metaventral impression and with unmodified metatrochanters. BL 1.85–1.98 mm; HL 0.33–0.35 mm, HW 0.33–0.34 mm, AnL 0.78–0.80 mm; PL 0.45–0.48 mm, PW 0.38–0.40 mm; EL 1.08–1.15 mm, EW 0.69–0.73 mm, EI 1.54–1.64.</p><p>Distribution. Central China: NC Sichuan.</p><p>Etymology. The name punctator (noun in apposition) alludes to the pointed metatrochanteral spine.</p><p>Remarks. The modification of the metatrochanter in this species is most similar to those of T. magnirufus sp. nov. and E. mediorufus sp. nov. described below. In these three species the ventral margin of the metatrochanter is more or less evenly concave in the distal region, and not abruptly bent or nearly straight, as in the remaining Chinese species. Trochanteral modifications are not known in species occurring outside China. Euconnus punctator differs from E. magnirufus in a distinctly smaller body. Aedeagi of these species have a similar shape, but the dorsal plate in E. punctator is elongate with broadly rounded apex, whereas that of E. magnirufus is wider than long and has strongly sinuate lateral margins, so that the apical region is projecting as a narrow subtriangular distomedian lobe. Euconnus mediorufus has a body length similar to that of E. punctator, but its aedeagus has the dorsal plate remarkably narrower than that in E. punctator . These three species share a similar endophallic structure and may be closely related.</p></div>	https://treatment.plazi.org/id/03BB87AA0B45637BFF0821E9FEFBFED6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jałoszyński, Paweł	Jałoszyński, Paweł (2024): Subgenera Cladoconnus Reitter and Tetramelus Motschulsky of Euconnus Thomson discovered in China (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 5514 (3): 232-256, DOI: 10.11646/zootaxa.5514.3.2, URL: http://dx.doi.org/10.11646/zootaxa.5514.3.2
03BB87AA0B4A637FFF08253EFAD0FE3A.text	03BB87AA0B4A637FFF08253EFAD0FE3A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euconnus (Tetramelus) aurilobatus Jałoszyński 2024	<div><p>Euconnus (Tetramelus) aurilobatus sp. nov.</p><p>(Figs 14, 22, 39–68)</p><p>Material studied. Holotype: ♂ (CHINA: Gansu): “CHINA: Gansu Prov. [CH 11-27] / Lenglong Ling Mts., 60 km NNW / Honggu, Jin Sha Gorge, mixed forest / (Picea, Populus, Betula), 36°51’56.9’’N, / 102°38’55.3’’E, litter, moss, mushrooms / sifted, 12.VII.2011, leg. M. Schülke” [white, printed], “ EUCONNUS / ( TETRAMELUS) /</p><p>aurilobatus m. / P. Jałoszyński, 2024 / HOLOTYPUS ” [red, printed] (MNHB). Paratypes (22 exx.): 1 ♂, same data as for holotype; 3 ♂♂, “ CHINA: Gansu Prov. [CH11-27a] / Lenglong Ling Mts., 60 km NNW / Honggu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.64869&amp;materialsCitation.latitude=36.865803" title="Search Plazi for locations around (long 102.64869/lat 36.865803)">Jin Sha Gorge</a>, mixed forest / (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.64869&amp;materialsCitation.latitude=36.865803" title="Search Plazi for locations around (long 102.64869/lat 36.865803)">Picea</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.64869&amp;materialsCitation.latitude=36.865803" title="Search Plazi for locations around (long 102.64869/lat 36.865803)">Populus</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.64869&amp;materialsCitation.latitude=36.865803" title="Search Plazi for locations around (long 102.64869/lat 36.865803)">Betula</a>), 36°51’56.9’’N, / 102°38’55.3’’E, litter, moss, mushrooms / sifted, 14/ 15.VII.2011, leg. M. Schülke ” [white, printed] ; 5 ♂♂, “ CHINA: Qinghai Prov. [CH11-08] / Daban Shan, 60 km NW Honggu / 36°49’10.7’’N, 102°31’’22.8’’E, 2366- / 2400 m, mixed forest (Betula, Populus, / Picea) dead wood, litter sifted / 25. VI.2011, leg. M. Schülke ” [white, printed] ; 2 ♂♂, “ CHINA: Gansu Prov. [CH11-13d] / Lenglong Ling Mts., 80 km NNW / <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.66589&amp;materialsCitation.latitude=37.063972" title="Search Plazi for locations around (long 102.66589/lat 37.063972)">Honggu</a>, 37°03’50.3’’N, 102°39’57.2’’E, / 3392 m, alpine pasture with / Rhododendron, litter, grass roots sifted / 1.VII.2011, leg. M. Schülke ” [white, printed] ; 3 ♂♂, 2 ♀♀, “ CHINA: Qinghai Prov. [CH11-08a] / Daban Shan, 60 km NW Honggu / 36°49’10.7’’N, 102°31’’22.8’’E, 2366- / 2400 m, mixed forest (Betula, Populus, / Picea) dead wood, litter sifted / 11.VII.2011, leg. M. Schülke ” [white, printed] ; 2 ♂♂, 1 ♀, “ CHINA: Qinghai Prov. [CH11-09] / Daban Shan, 62 km NNW Honggu / creek valley, Picea, Populus, Betula / forest, 36°51’15-28’’N, 102°36’’24’’- / 37’07’’E, 2236-2350 m, litter, dead wood / &amp; moss sifted, 26. VI.2011, leg. M. Schülke ” [white, printed] ; 1 ♂, “ CHINA: Gansu Prov. [CH11-13] / Lenglong Ling Mts., 80 km NNW / <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.66589&amp;materialsCitation.latitude=37.063972" title="Search Plazi for locations around (long 102.66589/lat 37.063972)">Honggu</a>, 37°03’50.3’’N, 102°39’57.2’’E, / 3392 m, alpine pasture with / Rhododendron, litter, grass roots sifted / 30. VI.2011, leg. M. Schülke ” [white, printed] ; 1 ♂, “ CHINA: Qinghai Prov. [CH11-09e] / Daban Shan, 62 km NNW Honggu / creek valley, Picea, Populus, Betula / forest, 36°51’15-28’’N, 102°36’’24’’- / 37’07’’E, 2236-2350 m, creek valley, / litter, dead wood / &amp; moss sifted, / 10.VII.2011, leg. M. Schülke ” [white, printed] ; 1 ♂, “ CHINA: Gansu Prov. [CH11-12] / Lenglong Ling Mts., pass road 61.5 km / N <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.75855&amp;materialsCitation.latitude=36.88639" title="Search Plazi for locations around (long 102.75855/lat 36.88639)">Honggu</a>, 2875-2900 m, 36°53’11.0’’N, / 102°45’30.8’’E, dry grass steppe and / field edges, under stones, running on / trail, 29. VI.2011, leg. M. Schülke ” [white, printed], (cPJ, MNHB) .</p><p>Diagnosis. Body nearly black; male metatrochanter distally produced into short and stout subtriangular spine, ventral margin of trochanter bent distally at an obtuse angle; aedeagus in ventral view distinctly more than twice as long as wide, its ventral plate wider than long, with distomedian subtriangular projection and pair of large subtriangular or rounded lateral lobes, dorsal plate elongate subtriangular with broadened subapical region and narrow truncate apex, endophallus in ventral view with two strongly elongate sub-parallel sclerites narrowing distally.</p><p>Description. Body of male (Fig. 14) elongate and slender, moderately convex, with long appendages, BL 1.68–1.90 mm; cuticle glossy, pigmentation nearly black with distinctly lighter, dark brown appendages, vestiture of setae lighter than cuticle.</p><p>Head in dorsal view round, broadest across eyes, HL 0.28–0.31 mm, HW 0.29–0.31 mm; in lateral view each temple 2.2–2.5 times as long as eye; vertex weakly convex and only slightly bulging posterodorsally; frons flattened; supraantennal tubercles small and indistinctly elevated. Eyes small and finely faceted, hardly projecting laterally from head silhouette, in lateral view oval. Punctures on vertex and frons inconspicuous, small and sparse; setae short and sparse, suberect, sparse and short thick bristles present on vertex and tempora. Antennae slender, indistinctly shorter than half BL, AnL 0.78–0.85 mm; antennomeres 1–2 strongly, 3–6 slightly elongate, 7–8 each about as long as broad, 9 and 10 each weakly transverse, 11 much shorter than 9–10 combined, about 1.6 times as long as broad.</p><p>Pronotum broadest in front of middle, PL 0.40–0.45 mm, PW 0.35–0.40 mm; anterior margin arcuate, lateral margins rounded in anterior half and nearly straight in posterior third, posterior corners slightly obtuse-angled, blunt; posterior margin weakly rounded. Pronotal base with single outer pair of faint pits. Punctures on pronotal disc inconspicuous; setae moderately dense and short, suberect and on sides mixed with thick bristles.</p><p>Elytra oval, broadest distinctly anterior to middle, EL 1.00– 1.13 mm, EW 0.65–0.70 mm, EI 1.50–1.69; basal impressions and humeral calli lacking, elytral apices separately rounded. Punctures on elytral disc inconspicuous, superficial; setae similar to those on pronotum, moderately dense and weakly suberect.</p><p>Legs long and slender, metatrochanter (Fig. 22) modified, with short and stout subtriangular distal spine, ventral margin of trochanter bent at an obtuse angle in distal region.</p><p>Metaventrite with deep oval median impression occupying most of its surface, extending on almost entire length.</p><p>Aedeagus (Figs 39–58) elongate, AeL 0.50 mm; median lobe in ventral view showing some variability, from nearly as long as broad to distinctly elongate, rapidly narrowing distally, ventral plate broader than long, with lateral pair of large subtriangular or rounded lobes and variously long distomedian projection; endophallus in ventral view (Figs 59–68) with two strongly elongate and approximately parallel sclerites narrowing distally, variable in shape. Dorsal wall lacking setae; parameres slender, each with 3 long subapical setae.</p><p>Female. Similar to male but with slightly smaller eyes (in lateral view temple nearly as long as 3 diameters of eye), lacking metaventral impression and with unmodified metatrochanters. BL 1.73–1.78 mm; HL 0.30 mm, HW 0.30 mm, AnL 0.73–0.80 mm; PL 0.43 mm, PW 0.38 mm; EL 1.00– 1.05 mm, EW 0.63–0.65 mm, EI 1.54–1.62.</p><p>Distribution. Central China: Central Gansu / easternmost Qinghai border area.</p><p>Etymology. The adjective aurilobatus alludes to the lateral earlike lobes of the aedeagus (after the Latin word auris, an ear).</p><p>Remarks. This species is most easily identifiable by its remarkable aedeagus, strongly differing from all Chinese congeners in the presence of large lateral lobes of the ventral plate. Variously shaped lateral lobes of the ventral plate are characteristic of Tetramelus of the reitteri group distributed in Caucasus and Transcaucasia (Vít &amp; Hlaváč 1997, 1998), but none of these was reported to have a modified metatrochanter.</p><p>The available sample of specimens (all collected in mountains between Xining in Qinghai and Wuwei in Gansu) shows a great variability of the body length and proportions of body parts, most notably the elytral index, as well as some variability in the shape of the median lobe and the endophallic sclerites. The variability of external structures is not uncommon within West Palaearctic populations of wingless Tetramelus species (Jałoszyński, unpublished obs.). The aedeagus of each available specimen was studied, photographed and compared with all others in order to verify a hypothesis of more than one species being present in the sample. Examples are shown in Figs 39–68. Differences can be seen in shapes and proportions of various regions of the median lobe (lateral lobes, distomedian process of ventral plate, distal and proximal regions of median lobe), and, most notably, in the shapes of two endophallic sclerites. In the latter case, the sclerites are in all specimens asymmetrical, one being longer and more slender than the other, but their shapes vary and their placement in some specimens is inverted (compare Fig. 41 vs. 45, and 59 vs. 64). All males have the same shape of modified metatrochanters. In my opinion, differences found in the aedeagi represent intraspecific variability, and I treat all observed variants as belonging to a single species.</p></div>	https://treatment.plazi.org/id/03BB87AA0B4A637FFF08253EFAD0FE3A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jałoszyński, Paweł	Jałoszyński, Paweł (2024): Subgenera Cladoconnus Reitter and Tetramelus Motschulsky of Euconnus Thomson discovered in China (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 5514 (3): 232-256, DOI: 10.11646/zootaxa.5514.3.2, URL: http://dx.doi.org/10.11646/zootaxa.5514.3.2
03BB87AA0B4E637CFF0824C6FC4FF976.text	03BB87AA0B4E637CFF0824C6FC4FF976.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euconnus (Tetramelus) magnirufus Jałoszyński 2024	<div><p>Euconnus (Tetramelus) magnirufus sp. nov.</p><p>(Figs 15, 23, 69–72)</p><p>Material studied. Holotype: ♂ (CHINA: Yunnan): “CHINA: N-Yunnan [C03-07] / Zhongdian Co., 55 km N Zhong- / dian, 28°19.8’ N, / 99°45.7’E, / 3800m, primary mixed forest, / Rhodod., dead wood, mushrooms, / moss, 18.VIII.2003, M. Schülke ” [white, printed], “ EUCONNUS / ( TETRAMELUS) / magnirufus m. / P. Jałoszyński, 2024 / HOLOTYPUS” [red, printed] (MNHB) . Paratypes (19 exx.): 12 ♂♂, 4 ♀♀, same data as for holotype; 1 ♂, 2 ♀♀,</p><p>“ CHINA (N-Yunnan) <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.761665&amp;materialsCitation.latitude=28.33" title="Search Plazi for locations around (long 99.761665/lat 28.33)">Zhongdian</a> / Co., 55 km N Zhongdian, / 28°19.8’N / 99°45.7’E, 3800m / (primary mixed forest, / Rhodod., dead wood, leaf / litter, mushrooms, moss) / 18.VIII.2003, Wrase [07]” [white, printed] (cPJ, MNHB) .</p><p>Diagnosis. Body rufous brown; male metatrochanter distally produced into moderately long subtriangular spine, ventral margin of trochanter bent at an obtuse angle; aedeagus in ventral view about twice as long as wide, its ventral plate much wider than long, with sinuate lateral margins converging distally to rounded apex, dorsal plate distinctly wider than long, with strongly sinuate lateral margins converging distally to subtriangular and rounded apex, endophallus in ventral view with one slender and nearly L-shaped sclerite narrowing distally, surrounded by membranous folds densely covered with cuticular denticles.</p><p>Description. Body of male (Fig. 15) elongate and slender, moderately convex, with long appendages, BL 2.35–2.43 mm; cuticle glossy, pigmentation uniformly rufous brown, vestiture of setae lighter than cuticle.</p><p>Head in dorsal view round, broadest across eyes, HL 0.38–0.40 mm, HW 0.35–0.38 mm; in lateral view each temple 5 times as long as eye; vertex weakly convex and only slightly bulging posterodorsally; frons flattened; supraantennal tubercles small and indistinctly elevated. Eyes very small but not vestigial, finely faceted, hardly projecting laterally from head silhouette, in lateral view oval. Punctures on vertex and frons inconspicuous, small and sparse; setae short and sparse, suberect, sparse and short thick bristles present on vertex and tempora. Antennae slender, indistinctly shorter than half BL, AnL 0.93–1.03 mm; antennomeres 1–6 elongate (6 weakly so), 7–8 each about as long as broad, 9 and 10 each weakly transverse, 11 slightly shorter than 9–10 combined, about 1.7 times as long as broad.</p><p>Pronotum broadest in anterior to middle, PL 0.50–0.55 mm, PW 0.45–0.48 mm; anterior margin arcuate, lateral margins rounded in anterior half and nearly straight in posterior third, posterior corners slightly obtuse-angled, blunt; posterior margin weakly rounded. Pronotal base with single outer pair of small and shallow but distinct pits. Punctures on pronotal disc inconspicuous; setae moderately dense and short, suberect and on sides mixed with thick bristles.</p><p>Elytra oval, broadest slightly anterior to middle, EL 1.38–1.50 mm, EW 0.85–0.90 mm, EI 1.57–1.74; basal impressions and humeral calli lacking, elytral apices rounded together. Punctures on elytral disc inconspicuous, superficial; setae similar to those on pronotum, moderately dense and weakly suberect.</p><p>Legs long and slender, metatrochanter (Fig. 23) modified, distally produced into moderately long subtriangular spine, ventral margin of trochanter bent at an obtuse angle.</p><p>Metaventrite with deep oval median impression occupying most of its surface, extending on almost entire length.</p><p>Aedeagus (Figs 69–72) elongate, AeL 0.63 mm; median lobe in ventral view broadest in subapical region, rapidly narrowing distally, ventral plate much wider than long, with sinuate lateral margins converging distally to rounded apex, dorsal plate distinctly wider than long, with strongly sinuate lateral margins converging distally to subtriangular and rounded apex, endophallus in ventral view with one slender and nearly L-shaped sclerite narrowing distally, surrounded by membranous folds densely covered with cuticular denticles. Dorsal wall lacking setae; parameres slender, each with 3 long subapical setae.</p><p>Female. Similar to male but with indistinctly smaller and more circular eyes, lacking metaventral impression and with unmodified metatrochanters. BL 2.28–2.43 mm; HL 0.38–0.40 mm, HW 0.38 mm, AnL 0.93–1.00 mm; PL 0.53–0.55 mm, PW 0.45–0.48 mm; EL 1.38–1.50 mm, EW 0.88–0.95 mm, EI 1.47–1.67.</p><p>Distribution. WS China: Yunnan, Diqing Tibetan Autonomous Prefecture.</p><p>Etymology. The adjective magnirufus alludes to the large body size and brown pigmentation with a faint reddish hue.</p><p>Remarks. Euconnus magnirufus is the largest Chinese Tetramelus known so far. It has the endophallic structures similar to those in E. punctator and E. mediorufus sp. nov. described below. However, shapes of the ventral and dorsal plates of the median lobe in each of these species are clearly different.</p></div>	https://treatment.plazi.org/id/03BB87AA0B4E637CFF0824C6FC4FF976	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jałoszyński, Paweł	Jałoszyński, Paweł (2024): Subgenera Cladoconnus Reitter and Tetramelus Motschulsky of Euconnus Thomson discovered in China (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 5514 (3): 232-256, DOI: 10.11646/zootaxa.5514.3.2, URL: http://dx.doi.org/10.11646/zootaxa.5514.3.2
03BB87AA0B4D6362FF082512FB55FAEE.text	03BB87AA0B4D6362FF082512FB55FAEE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euconnus (Tetramelus) tetraspinosus Jałoszyński 2024	<div><p>Euconnus (Tetramelus) tetraspinosus sp. nov.</p><p>(Figs 16, 24–25, 73–76)</p><p>Material studied. Holotype: ♂ (CHINA: Gansu): “CHINA: Gansu Prov. [CH11-12] / <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.75855&amp;materialsCitation.latitude=36.88639" title="Search Plazi for locations around (long 102.75855/lat 36.88639)">Lenglong Ling Mts.</a>, pass road 61.5 km / N Honggu, 2875-2900 m, 36°53’11.0’’N, / 102°45’30.8’’E, dry grass steppe and / field edges,</p><p>under stones, running on / trail, 29. VI .2011, leg. M. Schülke ” [white, printed], “ EUCONNUS / ( TETRAMELUS) / tetraspinosus m. / P. Jałoszyński, 2024 / HOLOTYPUS ” [red, printed] (MNHB).</p><p>Diagnosis. Body dark brown; male with peculiar modifications of pro- and metatrochanters: protrochanter with conspicuously long and straight distal process broadening distally and truncate at apex, metatrochanter with long and strongly curved distal process broadening distally and truncate at apex; aedeagus in ventral view nearly four times as long as wide, its ventral plate elongate, with lateral margins weakly converging distally to broadly rounded apex, dorsal plate strongly elongate, with weakly sinuate lateral margins strongly converging distally to narrow rounded apex, endophallus in ventral view with three elongate sclerites, of which lateral ones are conspicuously long, and median sclerite is much shorter, all sclerites surrounded by membranous folds densely covered with cuticular denticles.</p><p>Description. Body of male (Fig. 16) elongate and slender, moderately convex, with long appendages, BL 1.95 mm; cuticle glossy, pigmentation dark brown with indistinctly lighter appendages, vestiture of setae lighter than cuticle.</p><p>Head in dorsal view round, broadest across eyes, HL 0.33 mm, HW 0.35 mm; in lateral view each temple 3 times as long as eye; vertex weakly convex and only slightly bulging posterodorsally; frons flattened; supraantennal tubercles small and indistinctly elevated. Eyes small, finely faceted, hardly projecting laterally from head silhouette, in lateral view oval. Punctures on vertex and frons inconspicuous, small and sparse; setae short and sparse, suberect, sparse and short thick bristles present on vertex and tempora. Antennae slender, indistinctly shorter than half BL, AnL 0.88 mm; antennomeres 1–6 elongate (6 weakly so), 7–8 each about as long as broad, 9 and 10 each weakly transverse, 11 slightly shorter than 9–10 combined, about 1.7 times as long as broad.</p><p>Pronotum broadest anterior to middle, PL 0.50 mm, PW 0.40 mm; anterior margin arcuate, lateral margins rounded in anterior half and nearly straight in posterior third, posterior corners slightly obtuse-angled, blunt; posterior margin weakly rounded. Pronotal base with single outer pair of small and shallow but distinct pits. Punctures on pronotal disc inconspicuous; setae moderately dense and short, suberect and on sides mixed with thick bristles.</p><p>Elytra oval, broadest slightly anterior to middle, EL 1.13 mm, EW 0.73 mm, EI 1.55; basal impressions and humeral calli lacking, elytral apices separately rounded. Punctures on elytral disc inconspicuous, superficial; setae similar to those on pronotum, moderately dense and weakly suberect.</p><p>Legs long and slender, protrochanter (Fig. 24) with conspicuously long and straight distal process broadening distally and truncate at apex, ventral margin of protrochanter bent at a nearly right angle, metatrochanter (Fig. 25) with long and strongly curved distal process broadening distally and truncate at apex, ventral margin of metatrochanter proximally straight and distally strongly curved.</p><p>Metaventrite with deep oval median impression occupying most of its surface, extending on almost entire length.</p><p>Aedeagus (Figs 73–76) conspicuously elongate, AeL 0.85 mm; median lobe in ventral view broadest near middle, in distal third rapidly narrowing distally, ventral plate elongate, with lateral margins weakly converging distally to broadly rounded apex, dorsal plate strongly elongate, with weakly sinuate lateral margins strongly converging distally to narrow rounded apex, endophallus in ventral view with three elongate sclerites, of which lateral ones are conspicuously long, and median sclerite is much shorter, all sclerites are surrounded by membranous folds densely covered with cuticular denticles. Dorsal wall lacking setae; parameres slender, each with 3 long subapical setae.</p><p>Female. Not known.</p><p>Distribution. Central China: Central Gansu.</p><p>Etymology. The adjective tetraspinosus refers to the spines present on the pro- and metatrochanters.</p><p>Remarks. This species is the easiest to identify of all Chinese Tetramelus . No other species of world Euconnus has so uniquely modified pro- and metatrochanters in males, and among the Chinese species E. tetraspinosus has a remarkably elongate aedeagus, much more slender than any other Tetramelus found in this country.</p></div>	https://treatment.plazi.org/id/03BB87AA0B4D6362FF082512FB55FAEE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jałoszyński, Paweł	Jałoszyński, Paweł (2024): Subgenera Cladoconnus Reitter and Tetramelus Motschulsky of Euconnus Thomson discovered in China (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 5514 (3): 232-256, DOI: 10.11646/zootaxa.5514.3.2, URL: http://dx.doi.org/10.11646/zootaxa.5514.3.2
03BB87AA0B536363FF0827EAFE2FFA9A.text	03BB87AA0B536363FF0827EAFE2FFA9A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euconnus (Tetramelus) gansuminor Jałoszyński 2024	<div><p>Euconnus (Tetramelus) gansuminor sp. nov.</p><p>(Figs 17, 26, 77–80)</p><p>Material studied. Holotype: ♂ (CHINA: Gansu): “CHINA: Gansu Prov. [CH11-12 a] / <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.75855&amp;materialsCitation.latitude=36.88639" title="Search Plazi for locations around (long 102.75855/lat 36.88639)">Lenglong Ling Mts.</a>, pass road 61.5 km / N Honggu, 2875-2900 m, 36°53’11.0’’N, / 102°45’30.8’’E, dry grass steppe and / field edges, under stones, running on / trail, 30.VI.2011, leg. M. Schülke ” [white, printed], “ EUCONNUS / ( TETRAMELUS) / tetraspinosus m. / P. Jałoszyński, 2024 / HOLOTYPUS” [red, printed] (MNHB).</p><p>Diagnosis. Body dark brown; male metatrochanter distally produced into short and stout subtriangular spine, ventral margin of trochanter distally bent at an obtuse angle; aedeagus in ventral view about twice as long as wide, its ventral plate conspicuously small, elongate and subtriangular with rounded apex, dorsal plate elongate, with sinuate lateral margins strongly converging distally to narrow truncate apex, endophallus in ventral view with two short and curved elongate sclerites of unequal lengths surrounded by membranous folds densely covered with cuticular denticles.</p><p>Description. Body of male (Fig. 17) elongate and slender, moderately convex, with long appendages, BL 1.40 mm; cuticle glossy, pigmentation uniformly dark brown, vestiture of setae lighter than cuticle.</p><p>Head in dorsal view round, broadest across eyes, HL 0.23 mm, HW 0.23 mm; in lateral view each temple slightly more than 3 times as long as eye; vertex weakly convex and only slightly bulging posterodorsally; frons flattened; supraantennal tubercles small and indistinctly elevated. Eyes small, finely faceted, hardly projecting laterally from head silhouette, in lateral view oval. Punctures on vertex and frons inconspicuous, small and sparse; setae short and sparse, suberect, sparse and short thick bristles present on vertex and tempora. Antennae slender, almost as long as half BL, AnL 0.68 mm; antennomeres 1–2 elongate, 3–8 each about as long as broad, 9 and 10 each weakly transverse, 11 slightly shorter than 9–10 combined, about 1.7 times as long as broad.</p><p>Pronotum broadest anterior to middle, PL 0.35 mm, PW 0.28 mm; anterior margin arcuate, lateral margins rounded in anterior half and nearly straight in posterior third, posterior corners slightly obtuse-angled, blunt; posterior margin weakly rounded. Pronotal base with single outer pair of faint pits. Punctures on pronotal disc inconspicuous; setae moderately dense and short, suberect and on sides mixed with thick bristles.</p><p>Elytra oval, broadest near middle, EL 0.83 mm, EW 0.53 mm, EI 1.57; basal impressions and humeral calli lacking, elytral apices rounded together. Punctures on elytral disc inconspicuous, superficial; setae similar to those on pronotum, moderately dense and weakly suberect.</p><p>Legs long and slender, metatrochanter (Fig. 26) distally produced into short and stout subtriangular spine, ventral margin of trochanter distally bent at an obtuse angle.</p><p>Metaventrite with deep oval median impression occupying most of its surface, extending on almost entire length.</p><p>Aedeagus (Figs 77–80) stout, AeL 0.38 mm; median lobe in ventral view broadest distally to middle, capsular part distally truncate, ventral plate conspicuously small, elongate and subtriangular with rounded apex, dorsal plate elongate, with sinuate lateral margins strongly converging distally to narrow truncate apex, endophallus in ventral view with two short and curved elongate sclerites of unequal lengths surrounded by membranous folds densely covered with cuticular denticles. Dorsal wall with short setae; parameres slender, each with 3 long subapical setae.</p><p>Female. Not known.</p><p>Distribution. Central China: Central Gansu.</p><p>Etymology. The adjective gansuminor alludes to the locality where this species has been collected (Gansu province) and to the very small body size.</p><p>Remarks. This species is the smallest of all Chinese Tetramelus . The metatrochanter in E. gansuminor is nearly identical to those in E. aurilobatus and E. microrufus . Aedeagi of these three species differ remarkably, with only that of E. aurilobatus having large distolateral lobes. Aedeagi of E. gansuminor and E. microrufus show clearly different general shapes, different shapes of both ventral and dorsal plates, and a different arrangement of endophallic structures.</p></div>	https://treatment.plazi.org/id/03BB87AA0B536363FF0827EAFE2FFA9A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jałoszyński, Paweł	Jałoszyński, Paweł (2024): Subgenera Cladoconnus Reitter and Tetramelus Motschulsky of Euconnus Thomson discovered in China (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 5514 (3): 232-256, DOI: 10.11646/zootaxa.5514.3.2, URL: http://dx.doi.org/10.11646/zootaxa.5514.3.2
03BB87AA0B526360FF08277EFD66FB36.text	03BB87AA0B526360FF08277EFD66FB36.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euconnus (Tetramelus) microrufus Jałoszyński 2024	<div><p>Euconnus (Tetramelus) microrufus sp. nov.</p><p>(Figs 18, 27, 81–83)</p><p>Material studied. Holotype: ♂ (CHINA: Yunnan): “CHINA: N-Yunnan [C03-12] / Zhongdian Co., pass 28 km ESE Zhong- / dian, devastated primary forest with / young Abies, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.97&amp;materialsCitation.latitude=27.731667" title="Search Plazi for locations around (long 99.97/lat 27.731667)">Larix</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.97&amp;materialsCitation.latitude=27.731667" title="Search Plazi for locations around (long 99.97/lat 27.731667)">Betula</a>, Rhodod., / 27°43.9’ N, / 99°58.2’E, / 3700-3750m, / 22.VIII.2003, leg. M. Schülke ” [white, printed], “ EUCONNUS / ( TETRAMELUS) / microrufus m. / P. Jałoszyński, 2024 / HOLOTYPUS” [red, printed] (MNHB).</p><p>Diagnosis. Body rufous brown; male metatrochanter distally produced into short and stout subtriangular spine, ventral margin of trochanter distally weakly bent at an obtuse angle; aedeagus in ventral view nearly 3 times as long as wide, its ventral plate about as long as broad, with lateral margins sinuate and strongly converging distally to narrow rounded apex, dorsal plate elongate, with nearly straight lateral margins strongly converging distally to broad truncate apex, endophallus in ventral view with three short elongate sclerites of unequal lengths surrounded by membranous folds densely covered with cuticular denticles.</p><p>Description. Body of male (Fig. 18) elongate and slender, moderately convex, with long appendages, BL 1.80 mm; cuticle glossy, pigmentation uniformly rufous brown, vestiture of setae lighter than cuticle.</p><p>Head in dorsal view round, broadest at eyes, HL 0.28 mm, HW 0.29 mm; in lateral view each temple slightly more than 3 times as long as eye; vertex weakly convex and only slightly bulging posterodorsally; frons flattened; supraantennal tubercles small and indistinctly elevated. Eyes small, finely faceted, hardly projecting laterally from head silhouette, in lateral view oval. Punctures on vertex and frons inconspicuous, small and sparse; setae short and sparse, suberect, sparse and short thick bristles present on vertex and tempora.Antennae slender, indistinctly shorter than half BL, AnL 0.80 mm; antennomeres 1–5 elongate, 6–8 each about as long as broad, 9 and 10 each weakly transverse, 11 slightly shorter than 9–10 combined, about 1.6 times as long as broad.</p><p>Pronotum broadest in front of middle, PL 0.45 mm, PW 0.38 mm; anterior margin arcuate, lateral margins rounded in anterior half and nearly straight in posterior third, posterior corners slightly obtuse-angled, blunt; posterior margin weakly rounded. Pronotal base with one outer pair of faint pits. Punctures on pronotal disc inconspicuous; setae moderately dense and short, suberect and on sides mixed with thick bristles.</p><p>Elytra oval, broadest distinctly in front of middle, EL 1.08 mm, EW 0.65 mm, EI 1.65; basal impressions and humeral calli lacking, elytral apices rounded together. Punctures on elytral disc inconspicuous, superficial; setae similar to those on pronotum, moderately dense and weakly suberect.</p><p>Legs long and slender, metatrochanter (Fig. 27) distally produced into short and stout subtriangular spine, ventral margin of trochanter distally weakly bent at an obtuse angle.</p><p>Metaventrite with deep oval median impression occupying most of its surface, extending on almost entire length.</p><p>Aedeagus (Figs 81–84) slender, AeL 0.55 mm; median lobe in ventral view broadest near middle, distally rapidly narrowing, ventral plate about as long as broad, with lateral margins sinuate and strongly converging distally to narrow rounded apex, dorsal plate elongate, with nearly straight lateral margins strongly converging distally to broad truncate apex, endophallus in ventral view with three short elongate sclerites of unequal lengths surrounded by membranous folds densely covered with cuticular denticles. Dorsal wall with short setae; parameres slender, each with 3 long subapical setae.</p><p>Female. Not known.</p><p>Distribution. WS China: Yunnan, Diqing Tibetan Autonomous Prefecture.</p><p>Etymology. The adjective microrufus refers to the small body size and to reddish-brown pigmentation.</p><p>Remarks. As mentioned above, the metatrochanter in E. microrufus is nearly identical to those in E. gansuminor and E. aurilobatus, but the aedeagi of these three species strongly differ. Of these three species, E. gansuminor has the aedeagus most similar to that of E. microrufus . However, all important structures, i.e., the general shape of the median lobe, shapes and proportions of the ventral and dorsal plate, and shape and arrangement of endophallic structures, are clearly different in these two species.</p></div>	https://treatment.plazi.org/id/03BB87AA0B526360FF08277EFD66FB36	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jałoszyński, Paweł	Jałoszyński, Paweł (2024): Subgenera Cladoconnus Reitter and Tetramelus Motschulsky of Euconnus Thomson discovered in China (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 5514 (3): 232-256, DOI: 10.11646/zootaxa.5514.3.2, URL: http://dx.doi.org/10.11646/zootaxa.5514.3.2
03BB87AA0B516366FF0826D2FD2DFB86.text	03BB87AA0B516366FF0826D2FD2DFB86.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euconnus (Tetramelus) sinespinosus Jałoszyński 2024	<div><p>Euconnus (Tetramelus) sinespinosus sp. nov.</p><p>(Figs 19, 28, 85–88)</p><p>Material studied. Holotype: ♂ (CHINA: Sichuan): “CHINA: N-Sichuan [CH12-24] / pass 35 km NNW <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.43222&amp;materialsCitation.latitude=32.925556" title="Search Plazi for locations around (long 103.43222/lat 32.925556)">Songpan</a> / 32°55’32’’N, 103°25’56’’E, 3600 m, / moist N-slope with Salix and other / shrubs, litter, grass roots &amp; moss / sifted, 11.VIII.2012, leg. M. Schülke ” [white, printed], “ EUCONNUS / ( TETRAMELUS) / sinespinosus m. / P. Jałoszyński, 2024 / HOLOTYPUS” [red, printed] (MNHB).</p><p>Diagnosis. Body nearly black with distinctly lighter proximal region of antennae and infuscate clubs; male trochanters unmodified; aedeagus in ventral view slightly more than twice as long as wide, its ventral plate much wider than long, with lateral margins weakly concave and strongly converging distally to narrow rounded apex, dorsal plate elongate, with broad subtriangular proximal half and rapidly narrowed long distal portion slightly broadened subapically and with narrow rounded apex, endophallus in ventral view with three long sclerites of unequal lengths, of which two left sclerites are parallel and long, and one right sclerite is much shorter and divergent in relation to left pair, between sclerites endophallic folds covered densely with cuticular denticles.</p><p>Description. Body of male (Fig. 19) elongate and slender, moderately convex, with long appendages, BL 1.53 mm; cuticle glossy, pigmentation nearly black with slightly lighter legs and distinctly lighter proximal antennomeres and infuscate clubs, vestiture of setae lighter than cuticle.</p><p>Head in dorsal view round, broadest across eyes, HL 0.25 mm, HW 0.25 mm; in lateral view each temple about 1.5 times as long as eye; vertex weakly convex and only slightly bulging posterodorsally; frons flattened; supraantennal tubercles small and indistinctly elevated. Eyes small, finely faceted, hardly projecting laterally from head silhouette, in lateral view oval. Punctures on vertex and frons inconspicuous, small and sparse; setae short and sparse, suberect, sparse and short thick bristles present on vertex and tempora.Antennae slender, indistinctly shorter than half BL, AnL 0.65 mm; antennomeres 1–3 elongate, 4–8 each about as long as broad, 9 and 10 each weakly transverse, 11 slightly shorter than 9–10 combined, about 1.6 times as long as broad.</p><p>Pronotum broadest anterior to middle, PL 0.38 mm, PW 0.33 mm; anterior margin arcuate, lateral margins rounded in anterior half and nearly straight in posterior third, posterior corners slightly obtuse-angled, blunt; posterior margin weakly rounded. Pronotal base with single outer pair of small but distinct pits. Punctures on pronotal disc inconspicuous; setae moderately dense and short, suberect and on sides mixed with thick bristles.</p><p>Elytra oval, broadest slightly in front of middle, EL 0.90 mm, EW 0.58 mm, EI 1.57; basal impressions and humeral calli lacking, elytral apices rounded together. Punctures on elytral disc inconspicuous, superficial; setae similar to those on pronotum, moderately dense and weakly suberect.</p><p>Legs long and slender, metatrochanter (Fig. 28) unmodified.</p><p>Metaventrite with deep oval median impression occupying most of its surface, extending on almost entire length.</p><p>Aedeagus (Figs 85–88) stout with conspicuously slender apical region, AeL 0.50 mm; median lobe in ventral view broadest distally to middle, distally rapidly narrowing, ventral plate much wider than long, with lateral margins weakly concave and strongly converging distally to narrow rounded apex, dorsal plate elongate, with broad subtriangular proximal half and rapidly narrowed long distal portion slightly broadened subapically and with narrow rounded apex, endophallus in ventral view with three long sclerites of unequal lengths, of which two left sclerites are parallel and long, and one right sclerite is much shorter and divergent in relation to left pair, between sclerites endophallic folds covered densely with cuticular denticles. Dorsal wall lacking setae; parameres slender, each with 3 long subapical setae.</p><p>Female. Not known.</p><p>Distribution. Central China, NC Sichuan.</p><p>Etymology. The adjective sinespinosus is derived from a Latin phrase sine spinae, “without spines,” and refers to the unmodified metatrochanters.</p><p>Remarks. Euconnus sinespinosus is the only known Chinese Tetramelus with all trochanters in male unmodified. Its aedeagus is most similar to that of E. mediorufus, but shapes of the ventral and distal plates, as well as arrangement of endophallic structures are clearly different.</p></div>	https://treatment.plazi.org/id/03BB87AA0B516366FF0826D2FD2DFB86	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jałoszyński, Paweł	Jałoszyński, Paweł (2024): Subgenera Cladoconnus Reitter and Tetramelus Motschulsky of Euconnus Thomson discovered in China (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 5514 (3): 232-256, DOI: 10.11646/zootaxa.5514.3.2, URL: http://dx.doi.org/10.11646/zootaxa.5514.3.2
03BB87AA0B576364FF082642FB1DFE46.text	03BB87AA0B576364FF082642FB1DFE46.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euconnus (Tetramelus) mediorufus Jałoszyński 2024	<div><p>Euconnus (Tetramelus) mediorufus sp. nov.</p><p>(Figs 20, 29–30, 89–96)</p><p>Material studied. Holotype: ♂ (CHINA: Yunnan): “CHINA: N-Yunnan [C2005-05] / Diqing Tibet. Aut. Pref., / Zhongdian Co., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.64167&amp;materialsCitation.latitude=27.618334" title="Search Plazi for locations around (long 99.64167/lat 27.618334)">Xue Shan</a> near / lake 23 km S Zhongdian, 3895 m ”, “ 27°37.1’ N, 99°38.5 E, devast. / mixed forest, meadows, lake / border, leaf litter, dead wood, / sifted, 5.VI.2005, / leg. M. Schülke ” [white, printed], “ EUCONNUS / ( TETRAMELUS) / mediorufus m. / P. Jałoszyński, 2024 / HOLOTYPUS” [red, printed] (MNHB) . Paratype: ♂, “CHINA, Yunnan, / Haba Shan, N27°21’57’’ / E100°06’55’’, / 29.vi.2012, 3362m, / sift36 V. Grebennikov” [white, printed] (cPJ) .</p><p>Diagnosis. Body rufous brown; male metatrochanter distally produced into long subtriangular spine, ventral margin of trochanter evenly concave in distal half; aedeagus in ventral view slightly less than twice as long as wide, its ventral plate much wider than long, with lateral margins strongly sinuate and strongly converging distally to narrow rounded apex, dorsal plate elongate, with sinuate lateral margins strongly converging to narrowly rounded apex, endophallus in ventral view with one short elongate sclerite surrounded by endophallic folds covered densely with cuticular denticles.</p><p>Description. Body of male (Fig. 20) elongate and slender, moderately convex, with long appendages, BL 1.98–2.13 mm; cuticle glossy, pigmentation rufous brown, vestiture of setae lighter than cuticle.</p><p>Head in dorsal view round, broadest across eyes, HL 0.33–0.35 mm, HW 0.33–0.35 mm; in lateral view each temple about 3.5 times as long as eye; vertex weakly convex and only slightly bulging posterodorsally; frons flattened; supraantennal tubercles small and indistinctly elevated. Eyes small, finely faceted, hardly projecting laterally from head silhouette, in lateral view oval. Punctures on vertex and frons inconspicuous, small and sparse; setae short and sparse, suberect, sparse and short thick bristles present on vertex and tempora. Antennae slender, indistinctly shorter than half BL, AnL 0.88–0.90 mm; antennomeres 1–5 elongate (3–5 indistinctly so), 6–8 each about as long as broad, 9 and 10 each weakly transverse, 11 slightly shorter than 9–10 combined, about 1.5 times as long as broad.</p><p>Pronotum broadest anterior to middle, PL 0.48–0.53 mm, PW 0.43–0.45 mm; anterior margin arcuate, lateral margins rounded in anterior half and nearly straight in posterior third, posterior corners slightly obtuse-angled, blunt; posterior margin weakly rounded. Pronotal base with single outer pair of small but distinct pits. Punctures on pronotal disc inconspicuous; setae moderately dense and short, suberect and on sides mixed with thick bristles.</p><p>Elytra oval, broadest slightly in front of middle, EL 1.18–1.25 mm, EW 0.80 mm, EI 1.47–1.56; basal impressions and humeral calli lacking, elytral apices rounded together. Punctures on elytral disc inconspicuous, superficial; setae similar to those on pronotum, moderately dense and weakly suberect.</p><p>Legs long and slender, metatrochanter (Figs 29, 30) distally produced into long subtriangular spine, ventral margin of trochanter evenly concave in distal half.</p><p>Metaventrite with deep oval median impression occupying most of its surface, extending on almost entire length.</p><p>Aedeagus (Figs 89–96) stout with slender apical region, AeL 0.55 mm; median lobe in ventral view broadest distally to middle or near middle, distally rapidly narrowing, ventral plate much wider than long, with lateral margins strongly sinuate and strongly converging distally to narrow rounded apex, dorsal plate elongate, with sinuate lateral margins strongly converging to narrowly rounded apex, endophallus in ventral view with one short elongate sclerite surrounded by endophallic folds covered densely with cuticular denticles. Dorsal wall with short setae; parameres slender, each with 3 long subapical setae.</p><p>Female. Not known.</p><p>Distribution. WS China: Yunnan, Diqing Tibetan Autonomous Prefecture.</p><p>Etymology. The adjective mediorufus refers to the medium body size and to the reddish-brown pigmentation.</p><p>Remarks. Males of Euconnus mediorufus have the metatrochanters modified similarly to those in E. punctator and E. magnirufus, with a relatively long subtriangular distal spine and the ventral margin arcuate, not bent. The three species share also a similar endophallic structure, with membranous folds densely covered with numerous cuticular denticles. The dorsal plate in E. mediorufus is conspicuously elongate and narrow, whereas in E. punctator it is elongate but broad, and in E. magnirufus the plate is nearly as long as broad.</p><p>The two studied males differ slightly in metatrochanters (Figs 29 and 30) and in aedeagi (Figs 89–92, and 93–96). However, these differences are too small to justify establishing two separate species.</p></div>	https://treatment.plazi.org/id/03BB87AA0B576364FF082642FB1DFE46	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jałoszyński, Paweł	Jałoszyński, Paweł (2024): Subgenera Cladoconnus Reitter and Tetramelus Motschulsky of Euconnus Thomson discovered in China (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 5514 (3): 232-256, DOI: 10.11646/zootaxa.5514.3.2, URL: http://dx.doi.org/10.11646/zootaxa.5514.3.2
