identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03B8222866745502FF75FA20FBE5F8E5.text	03B8222866745502FF75FA20FBE5F8E5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudohomonyx continentalis Yang & Pathomwattananurak 2023	<div><p>Pseudohomonyx continentalis Yang &amp; Pathomwattananurak, new species</p><p>(Figs. 1A–1E)</p><p>Type material (2♁♁, 1♀). Holotype: Malaysia: ♁ (CAU), “ Genting Highlands, Pahang State, Malaysia, IV.2022, local collector leg.” Paratypes: Thailand: 1♁ (THNHM), “S Thailand: Surat Thani Prov., Ban Na Sarn Dist., Plai Nam Subdist., 15–28 April 1986, J. Nabhitabhata leg. // THNHM-I-10375”; 1♀ (THNHM), “larva collected in 2023, Nopphitam Dist., Nakhon Sri Thammarat Prov., Chayaphol Photaworn leg., adult emerged 19.V.2023.”</p><p>Description (holotype male). General (Figs. 1A–1B): Body dark brown, surface mildly shiny; ventral surface covered with dense, long reddish-brown setae. Head (Fig. 1C): Surface mostly dark brown. Clypeus with dense transverse rugopunctures throughout; clypeal apex almost flat; lateral and apical margins of clypeus bordered. Frons with transverse rugopunctures throughout. Transverse carina separating frons and clypeus indicated by an obtuse tubercle at each side. Mandible at outer margin rounded near apex, almost straight at middle. Pronotum (Fig. 1C): Surface dark brown; sparsely micropunctate throughout, only with a few slightly larger punctures sparsely distributed near lateral margin. impression absent. Side roundly curved, widest near posterior one-fourth; all margins emarginate except narrowly interrupted in lateral margin; anterior margin without any tubercle at middle. Anterior and posterior angles rounded. Scutellum: Surface dark brown; sparsely punctate near base. Elytra: Surface mostly dark brown, except dark reddish sutural ridge; with sparse micropunctures and dense, round, large punctures in striae. Sutural striae narrow, deeply sulcate; other striae wider, even intervals slightly wider than their adjacent odd intervals. Intervals micropunctate, with dense, larger punctures on fourth and sixth intervals posteriorly. Apex densely punctured. Humeral and apical calli obtuse. Epipleuron of elytron moderately dilated horizontally from base to posterior one-fifth, widest behind middle. Metasternum: Surface with dense setae; punctures small and dense close to sides; median region with sparser and smaller, with a longitudinal depression. Abdomen: Pygidium with dark brown surface, slightly convex, with very dense punctures throughout. Abdominal ventrites with dark brown surface, with reddish anterior margin; with sparse micropunctures, larger punctures near base and longitudinal midline, and a row of punctures with long setae at sides. Legs: All legs dark brown, slightly more reddish than other parts of the body. Protibia tridentate, protarsi very strongly thickened, inner protibial claw very strongly enlarged and elongated. Parameres (Figs. 1D–1E): In caudal view (Fig.1D), slender and symmetric; without any setae; outer sides with a pair of short, acute, outward and strongly backward processes at posterior one-sixth; inner sides simply curved; apex rounded; in lateral view (Fig. 1E), apex slightly bent downward.</p><p>Paratype (male). Characters stable, except punctures on fourth and sixth intervals sparser, lesser in number (Fig. 2A).</p><p>Paratype (female). Characters mainly stable, except color much more reddish (it died two days after emerging before full melanization, thus the color is likely not its natural color); transverse carina separating frons and clypeus absent; punctures on fourth and sixth intervals sparser, lesser in number; protarsi not thickened, inner protibial claw not enlarged and elongated (Fig. 2B–2C).</p><p>Measurements. Body length: male 19.5–20.0 mm (holotype 20.0 mm), female 20.7 mm; body width: male 10.3–11.0 mm (holotype 10.3 mm), female 11.6 mm.</p><p>Diagnosis. This species is relatively unique in its appearances and can be easily distinguished from all other species of the genus by absent of the frontal impression and frontal tubercle on the pronotum in both sexes, the combination of the following characters can also distinguish this species from all other species: male with very strongly thickened protarsi and very strongly enlarged and elongated inner protibial claw; fourth and sixth intervals with punctures posteriorly; and the shape of the parameres.</p><p>Etymology. This species is named for its limited distribution on the continent, which is currently the only species known exclusively from mainland Southeast Asia.</p><p>Distribution. Malaysia: Malay Peninsula; Thailand.</p><p>Remarks. This species belongs to a group who lacks setation on the parameres (the small white dots on the photographs of the parameres (Figs. 1D–1E) are small pieces of dust instead of setae). Other species of this group are: Pseudohomonyx belitungensis Miyake &amp; Yamaya, 1997, P. borneensis Arrow, 1908, P. javanus (Burmeister, 1847), P. loksadoensis Jákl &amp; Zídek, 2017 and P. minor Jákl &amp; Zídek, 2017 (Jákl &amp; Zídek 2017).</p><p>Pseudohomonyx continentalis Yang &amp; Pathomwattananurak, new species, alongside with P. javanus (more information given in the according section in the catalogue), represents the first records of this genus from mainland Southeast Asia. It is also the only species endemic to the mainland.</p><p>It should be noted that the female paratype was assembled from pieces after it died before full melanization. Fortunately, characters can still be clearly visible and compared with the holotype.</p></div>	https://treatment.plazi.org/id/03B8222866745502FF75FA20FBE5F8E5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yang, Qiao-Zhi;Geshev, Georgi;Pathomwattananurak, Wuttipon	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B8222866715504FF75FF4CFCC6FB1E.text	03B8222866715504FF75FF4CFCC6FB1E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudohomonyx apoensis Miyake & Yamaya 1997	<div><p>Pseudohomonyx apoensis Miyake &amp; Yamaya, 1997</p><p>(Figs. 6A–6C)</p><p>Pseudohomonyx apoensis Miyake &amp; Yamaya, 1997: 15, figs. 17 (head and apical part of pronotum), 20h (habitus).</p><p>Type material examined. None.</p><p>Additional material examined (1♁). The Philippines: 1♁ (CQZY), Agco Hot Spring, Mt. Apo, Kidapawan City, Cotabato Prov., Mindanao Island, III.2017, Noel Mohagan leg.</p><p>Description (male). General (Figs. 3A–3B): Body black, with slight reddish color, surface mildly shiny; ventral surface covered with dense, long reddish-brown setae. Head (Fig. 3C): Surface mostly black, slight reddish near apex, mandible reddish. Clypeus with dense small punctures throughout, sometimes rugopunctate; clypeal apex almost flat, with a very mild, uplifted, triangular protuberance at middle; lateral and apical margins of clypeus bordered. Frons more sparsely punctate than clypeus, punctures mostly larger, slightly rugopunctate near center. Transverse carina separating frons and clypeus indicated by an obtuse tubercle and a distinct, deep, longitudinal depression behind the tubercle at each side. Mandible simply rounded at outer margin. Pronotum (Fig. 2C): Surface black; sparsely micropunctate throughout, only with a few slightly larger punctures sparsely distributed near lateral margin. Anterior region with a small, deep, round frontal impression, reaching one-fifth the length of pronotum, and a faint line posterior to impression reaching mid-length. Side roundly curved, widest near posterior one-third; all margins emarginate except widely interrupted in lateral margin; anterior margin with a distinct, large frontal tubercle at middle. Anterior and posterior angle rounded. Scutellum: Surface black; impunctate. Elytra: Surface mostly black, except by dark reddish suture; with sparse micropunctures and dense, round, small punctures in striae. Sutural striae narrow, deeply sulcate; other striae wider, equally wide. Intervals micropunctate, with sparse, larger punctures in sixth intervals posteriorly. Apex moderately punctured. Humeral and apical calli obtuse. Epipleuron of elytron strongly dilated horizontally from base to posterior one-fifth, widest behind middle. Metasternum: Surface with dense setae; punctures large and dense close to sides; median region impunctate, with a longitudinal depression. Abdomen: Pygidium with black surface, slightly convex, with dense punctures throughout except impunctate near apex, punctures denser at sides; punctures mainly small. Abdominal ventrites with black surface, with reddish anterior margin; with sparse micropunctures, larger punctures near base and a row of punctures with long setae at sides. Legs: All legs slightly reddish. Protibia tridentate, protarsi strongly thickened, inner protibial claw moderately enlarged and elongated. Parameres (Figs. 3D–3E): In caudal view (Fig. 3D), slender and nearly symmetric; apex with many setae-bearing punctures, setae very short; outer sides with a pair of short, acute, outward and backward processes at posterior one-seventh; inner sides sinuate; apex almost flat; in lateral view (Fig. 3E), apex bent downward.</p><p>Measurements. Body length: 18.8 mm; body width: 10.7 mm.</p><p>Remarks. The identification of this specimen is based on its characters highly matching the holotype (female) description and the locality. It is most closely related to P. negros Miyake &amp; Yamaya, 1997 due to the strong similarity between characters on the parameres (apex of parameres with setae) and their appearances (frontal impression small and round, frontal tubercle distinct, elytral intervals not densely punctate before middle or impunctate, which are often densely punctate in other species whose parameres bears setae at apex). The setae on the parameres of this species is much shorter than all other Philippine species.</p></div>	https://treatment.plazi.org/id/03B8222866715504FF75FF4CFCC6FB1E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yang, Qiao-Zhi;Geshev, Georgi;Pathomwattananurak, Wuttipon	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B8222866705504FF75F96DFD38F81C.text	03B8222866705504FF75F96DFD38F81C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudohomonyx apoensis Miyake & Yamaya 1997	<div><p>Pseudohomonyx apoensis Miyake &amp; Yamaya, 1997</p><p>Pseudohomonyx apoensis Miyake &amp; Yamaya, 1997: 15, figs. 17 (head and apical part of pronotum), 20h (habitus).</p><p>Type locality: “ Mt. Apo, Mindanao Island, Philippines ” .</p><p>Type depository: Holotype: ♀ (NMSM).</p><p>Distribution: The Philippines: Mindanao Island.</p></div>	https://treatment.plazi.org/id/03B8222866705504FF75F96DFD38F81C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yang, Qiao-Zhi;Geshev, Georgi;Pathomwattananurak, Wuttipon	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B8222866705504FF75FAE1FAE6F9DA.text	03B8222866705504FF75FAE1FAE6F9DA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudohomonyx Arrow 1908	<div><p>Genus Pseudohomonyx Arrow, 1908</p><p>Pseudohomonyx Arrow, 1908: 326 (original description of the genus); Arrow 1937: 32 (checklist); Endrődi 1969: 436 (revision of the genus, with redescription and identification key); Endrődi 1985: 315 (revision of the genus, with redescription and identification key); Krajčík 2005: 55 (checklist); Krajčík 2012: 228 (checklist).</p><p>Type species: Pseudohomonyx borneensis Arrow, 1908, by original designation.</p><p>Remarks. A distribution map of the genus based on all available information from literature (Endrődi 1969; Miyake &amp; Yamaya 1997, 1999, 2000; Jákl &amp; Zídek 2017) and the current paper is provided here (Fig. 4).</p></div>	https://treatment.plazi.org/id/03B8222866705504FF75FAE1FAE6F9DA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yang, Qiao-Zhi;Geshev, Georgi;Pathomwattananurak, Wuttipon	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B8222866735507FF75FA77FD4FF926.text	03B8222866735507FF75FA77FD4FF926.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudohomonyx belitungensis Miyake & Yamaya 1997	<div><p>Pseudohomonyx belitungensis Miyake &amp; Yamaya, 1997</p><p>Pseudohomonyx belitungensis Miyake &amp; Yamaya, 1997: 10, figs. 7a–b (parameres), 20k (habitus); Jákl &amp; Zídek 2017: 27.</p><p>Type locality: “ Belitung Island, east of Sumatra, Indonesia ”.</p><p>Type depository: Holotype: ♁ (NMSM); paratype: 1♁ (NMSM).</p><p>Distribution: Indonesia: Belitung Island.</p></div>	https://treatment.plazi.org/id/03B8222866735507FF75FA77FD4FF926	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yang, Qiao-Zhi;Geshev, Georgi;Pathomwattananurak, Wuttipon	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B8222866735506FF75F930FC90FF6B.text	03B8222866735506FF75F930FC90FF6B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudohomonyx borneensis Arrow 1908	<div><p>Pseudohomonyx borneensis Arrow, 1908</p><p>Pseudohomonyx borneensis Arrow, 1908: 327; Arrow 1937: 32; Endrődi 1969: 438, figs. 32–33 (parameres); Endrődi 1985: 315, figs. 1216–1217 (parameres); Miyake &amp; Yamaya 1997: 11, figs. 10 (head and apical part of pronotum), 12a–b (parameres), 20i (habitus); Krajčík 2005: 55; Krajčík 2012: 228; Jákl &amp; Zídek 2017: 24, figs. 10 (habitus), 11 (head and pronotum), 12–13 (parameres).</p><p>Type locality: “Borneo, Bowring”</p><p>Type depository: Neotype: ♁ (BMNH).</p><p>Distribution: Malaysia: Borneo; Indonesia: Kalimantan.</p></div>	https://treatment.plazi.org/id/03B8222866735506FF75F930FC90FF6B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yang, Qiao-Zhi;Geshev, Georgi;Pathomwattananurak, Wuttipon	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B8222866725506FF75FEFDFCC2FE6F.text	03B8222866725506FF75FEFDFCC2FE6F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudohomonyx continentalis Yang & Pathomwattananurak 2023	<div><p>Pseudohomonyx continentalis Yang &amp; Pathomwattananurak, new species</p><p>Type locality: “Genting Highlands, Pahang State, Malaysia”</p><p>Type depository: Holotype: ♁ (CAU); paratypes: 1♁, 1♀ (THNHM).</p><p>Distribution: Malaysia: Malay Peninsula; Thailand.</p></div>	https://treatment.plazi.org/id/03B8222866725506FF75FEFDFCC2FE6F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yang, Qiao-Zhi;Geshev, Georgi;Pathomwattananurak, Wuttipon	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B8222866725506FF75FDF9FBD2FC63.text	03B8222866725506FF75FDF9FBD2FC63.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudohomonyx crassus Arrow 1911	<div><p>Pseudohomonyx crassus Arrow, 1911</p><p>Pseudohomonyx crassus Arrow, 1911: 159; Arrow 1937: 32; Endrődi 1969: 441, figs. 36–37 (parameres); Endrődi 1985: 316, figs. 1220–1221 (parameres); Miyake &amp; Yamaya 1997: 10, figs. 8a–b (parameres); Krajčík 2005: 55; Krajčík 2012: 228; Jákl &amp; Zídek 2017: 22, figs. 6 (habitus), 7 (head and pronotum), 8–9 (parameres).</p><p>Type locality: “ Kina Balu, N. -Borneo”</p><p>Type depository: Lectotype: ♁ (BMNH).</p><p>Distribution: Malaysia: Borneo; Indonesia: Kalimantan.</p><p>Remarks. As mentioned by Endrődi (1969), a “ P. craceus ♁-Type” was found in Prell’s collection (now in MFNB) which is presented in the section of Pseudohomonyx crassus Arrow, 1911 . However, it has been identified by Endrődi as P. borneensis Arrow, 1908, with dubious locality of “ Java ”; and without any publication from Prell regarding this specimen, it was excluded from any type series (Endrődi 1969).</p></div>	https://treatment.plazi.org/id/03B8222866725506FF75FDF9FBD2FC63	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yang, Qiao-Zhi;Geshev, Georgi;Pathomwattananurak, Wuttipon	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B8222866725506FF75FBF5FDEFFA85.text	03B8222866725506FF75FBF5FDEFFA85.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudohomonyx extradentatus Miyake & Yamaya 1997	<div><p>Pseudohomonyx extradentatus Miyake &amp; Yamaya, 1997</p><p>Pseudohomonyx extradentatus Miyake &amp; Yamaya, 1997: 12, figs. 9 (head and apical part of pronotum), 11a–b (parameres), 20I (habitus); Jákl &amp; Zídek 2017: 27.</p><p>Type locality: “Keningau, Sabah, North Borneo”.</p><p>Type depository: Holotype: ♁ (NMSM); paratypes: 4♁♁, 5♀♀ (NMSM).</p><p>Distribution: Malaysia: Borneo.</p></div>	https://treatment.plazi.org/id/03B8222866725506FF75FBF5FDEFFA85	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yang, Qiao-Zhi;Geshev, Georgi;Pathomwattananurak, Wuttipon	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B8222866725509FF75FA50FF20FEFF.text	03B8222866725509FF75FA50FF20FEFF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudohomonyx javanus (Burmeister 1847)	<div><p>Pseudohomonyx javanus (Burmeister, 1847)</p><p>Heteronychus javanus Burmeister, 1847: 98 .</p><p>Pseudohomonyx javanus: Arrow, 1908: 326; Arrow 1937: 32; Endrődi 1969: 442, figs. 38–39 (parameres); Endrődi 1985: 316, figs. 1222–1223 (parameres); Krajčík 2005: 55; Krajčík 2012: 228; Jákl &amp; Zídek 2017: 25, figs. 23, 27 (habitus), 24, 28 (head and pronotum), 25–26, 29–30 (parameres).</p><p>Pseudohomonyx javanus javanus: Endrődi 1985: 317.</p><p>Pseudohomonyx morator sensu Arrow, 1908 (nec Fabricius, 1798).</p><p>Heteronychus sumatrensis Fairmaire, 1896: 225 .</p><p>Pseudohomonyx sumatrensis: Arrow, 1908: 326; Arrow 1937: 32.</p><p>Pseudohomonyx javanus sumatrensis: Endrődi 1969: 442, Endrődi 1985: 317; Krajčík 2005: 55; Krajčík 2012: 228; Jákl &amp; Zídek 2017: 25 (synonymized with Pseudohomonyx javanus).</p><p>Type locality: “ Java occident. [western Java], Sukabumi, 2000”</p><p>Type depository: Neotype of Pseudohomonyx javanus javanus (Burmeister, 1847): ♁ (HNHM); Lectotype of Pseudohomonyx javanus sumatrensis (Fairmaire, 1896): ♁ (MNHN); Paratype of Pseudohomonyx javanus sumatrensis (Fairmaire,1896):1[sex unknown](HNHM) (information regarding HNHM type materials are referenced from a datasheet of Dynastinae collection in HNHM provided by the late Ottó Merkl, private communication, 2020).</p><p>Material examined (5♁♁). Thailand: 1♁ (CWP), 5°53’07.05”N, 101°01’17.06”E, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.02141&amp;materialsCitation.latitude=5.8852916" title="Search Plazi for locations around (long 101.02141/lat 5.8852916)">Betong</a>, Yala Prov., 24.V.2022, Suradet Sae-Liang Leg . Malaysia: 4♁♁ (EUMJ), Univ. Malaya Field Studies Centre, Ulu Gombak, Daerah Gombak, Selangor State, alt. 250 m, 12–18.III.2009, Y. Senda leg.</p><p>Distribution: Indonesia: Sumatra Island, Nias Island, Java Island; Malaysia (new record); Thailand (new record).</p></div>	https://treatment.plazi.org/id/03B8222866725509FF75FA50FF20FEFF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yang, Qiao-Zhi;Geshev, Georgi;Pathomwattananurak, Wuttipon	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B82228667D5509FF75FE49FDA6FDC0.text	03B82228667D5509FF75FE49FDA6FDC0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudohomonyx loksadoensis Jakl & Zidek 2017	<div><p>Pseudohomonyx loksadoensis Jákl &amp; Zídek, 2017</p><p>Pseudohomonyx loksadoensis Jákl &amp; Zídek, 2017: 20 figs. 1–2, 19–20 (habitus), 3 (head and pronotum), 4–5 (parameres).</p><p>Type locality: “Indonesia, SE kalimantan, Kandangan District, Loksado village env.”</p><p>Type depository: Holotype: ♁ (CSJ); paratypes: 3♀♀ (CSJ).</p><p>Distribution: Indonesia: Kalimantan.</p></div>	https://treatment.plazi.org/id/03B82228667D5509FF75FE49FDA6FDC0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yang, Qiao-Zhi;Geshev, Georgi;Pathomwattananurak, Wuttipon	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B82228667D5509FF75FD0BFD12FC63.text	03B82228667D5509FF75FD0BFD12FC63.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudohomonyx luzonicus Miyake & Yamaya 2000	<div><p>Pseudohomonyx luzonicus Miyake &amp; Yamaya, 2000</p><p>Pseudohomonyx luzonicus Miyake &amp; Yamaya, 2000: 7, figs. 1 (head and apical part of pronotum), 2a–b (parameres), 5–6 (habitus); Krajčík 2005: 55; Krajčík 2012: 228.</p><p>Type locality: “Mt. Banahaw, Luzon Is., Philippines”.</p><p>Type depository: Holotype: ♁ (NMSM); paratypes: 2♀♀ (NMSM).</p><p>Distribution: The Philippines: Luzon Island.</p></div>	https://treatment.plazi.org/id/03B82228667D5509FF75FD0BFD12FC63	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yang, Qiao-Zhi;Geshev, Georgi;Pathomwattananurak, Wuttipon	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B82228667D5509FF75FBF5FD08FA86.text	03B82228667D5509FF75FBF5FD08FA86.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudohomonyx mindroensis Miyake & Yamaya 2000	<div><p>Pseudohomonyx mindroensis Miyake &amp; Yamaya, 2000</p><p>Pseudohomonyx mindroensis Miyake &amp; Yamaya, 2000: 9, figs. 3 (head and apical part of pronotum), 4a–b (parameres), 7 (habitus); Krajčík 2005: 55; Krajčík 2012: 228.</p><p>Type locality: “Mt. Halcon, Mindro [Mindoro] Is., Philippines”.</p><p>Type depository: Holotype: ♁ (NMSM).</p><p>Distribution: The Philippines: Mindoro Island.</p></div>	https://treatment.plazi.org/id/03B82228667D5509FF75FBF5FD08FA86	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yang, Qiao-Zhi;Geshev, Georgi;Pathomwattananurak, Wuttipon	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B82228667D5509FF75FA50FD40F97F.text	03B82228667D5509FF75FA50FD40F97F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudohomonyx minor Jakl & Zidek 2017	<div><p>Pseudohomonyx minor Jákl &amp; Zídek, 2017</p><p>Pseudohomonyx minor Jákl &amp; Zídek, 2017: 24 figs. 14–15, 21–22 (habitus), 16 (head and pronotum), 17–18 (parameres).</p><p>Type locality: “Indonesia, Sumatra Island, West Sumatra Province, Mt. Sanggul, Landai village env., 1250m m alt.”</p><p>Type depository: Holotype: ♁ (CSJ); paratypes: 9–14♁♁, 16– 22♀♀ [as mentioned in Jákl &amp; Zídek (2017)] (CSJ).</p><p>Distribution: Indonesia: Sumatra Island.</p></div>	https://treatment.plazi.org/id/03B82228667D5509FF75FA50FD40F97F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yang, Qiao-Zhi;Geshev, Georgi;Pathomwattananurak, Wuttipon	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B82228667D5508FF75F8C9FD1BFF0F.text	03B82228667D5508FF75F8C9FD1BFF0F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudohomonyx negros Miyake & Yamaya 1997	<div><p>Pseudohomonyx negros Miyake &amp; Yamaya, 1997</p><p>Pseudohomonyx negros Miyake &amp; Yamaya, 1997: 14, figs. 15a–b (parameres), 16 (head and apical part of pronotum), 20j (habitus).</p><p>Type locality: “Mt. Canlaon, Mambcal, Negros Island, Philippines”.</p><p>Type depository: Holotype: ♁ (NMSM), paratypes: 9♁♁, 18♀♀ (NMSM).</p><p>Distribution: The Philippines: Negros Island.</p></div>	https://treatment.plazi.org/id/03B82228667D5508FF75F8C9FD1BFF0F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yang, Qiao-Zhi;Geshev, Georgi;Pathomwattananurak, Wuttipon	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B82228667C5508FF75FED9FD0DFDB2.text	03B82228667C5508FF75FED9FD0DFDB2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudohomonyx palavanus Prell 1912	<div><p>Pseudohomonyx palavanus Prell, 1912</p><p>Pseudohomonyx palavanus Prell, 1912: 186; Arrow 1937: 32; Endrődi 1969: 440, figs. 34–35 (parameres); Endrődi 1985: 316, figs. 1218–1219 (parameres); Krajčík 2005: 55; Krajčík 2012: 228.</p><p>Type locality: “Palawan (Süd·Palawan) [southern Palawan]”</p><p>Type depository: Holotype: ♁ (MFNB).</p><p>Distribution: The Philippines: Palawan Island.</p></div>	https://treatment.plazi.org/id/03B82228667C5508FF75FED9FD0DFDB2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yang, Qiao-Zhi;Geshev, Georgi;Pathomwattananurak, Wuttipon	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B82228667C5508FF75FD84FD12FCF3.text	03B82228667C5508FF75FD84FD12FCF3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudohomonyx samarensis Miyake & Yamaya 1997	<div><p>Pseudohomonyx samarensis Miyake &amp; Yamaya, 1997</p><p>Pseudohomonyx samarensis Miyake &amp; Yamaya, 1997: 13, figs. 13 (head and apical part of pronotum), 14a–b (parameres).</p><p>Type locality: “Samar Island, Philippines”.</p><p>Type depository: Holotype: ♁ (NMSM).</p><p>Distribution: The Philippines: Samar Island.</p></div>	https://treatment.plazi.org/id/03B82228667C5508FF75FD84FD12FCF3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yang, Qiao-Zhi;Geshev, Georgi;Pathomwattananurak, Wuttipon	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B82228667C5508FF75FC45FDEFFB11.text	03B82228667C5508FF75FC45FDEFFB11.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudohomonyx similis Miyake & Yamaya 1999	<div><p>Pseudohomonyx similis Miyake &amp; Yamaya, 1999</p><p>Pseudohomonyx similis Miyake &amp; Yamaya, 1999: 101, figs. 1a–b (parameres), 8 (habitus); Krajčík 2005: 55; Krajčík 2012: 228; Jákl &amp; Zídek 2017: 27.</p><p>Type locality: “Mt. Tangga Saran, 400–900m alt., S. W. Engkili, Sarawak, Borneo Is.”</p><p>Type depository: Holotype: ♁ (NMSM).</p><p>Distribution: Malaysia: Borneo.</p></div>	https://treatment.plazi.org/id/03B82228667C5508FF75FC45FDEFFB11	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yang, Qiao-Zhi;Geshev, Georgi;Pathomwattananurak, Wuttipon	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
