taxonID	type	description	language	source
03B8222866745502FF75FA20FBE5F8E5.taxon	description	(Figs. 1 A – 1 E)	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B8222866745502FF75FA20FBE5F8E5.taxon	materials_examined	Type material (2 ♁♁, 1 ♀). Holotype: Malaysia: ♁ (CAU), “ Genting Highlands, Pahang State, Malaysia, IV. 2022, local collector leg. ” Paratypes: Thailand: 1 ♁ (THNHM), “ S Thailand: Surat Thani Prov., Ban Na Sarn Dist., Plai Nam Subdist., 15 – 28 April 1986, J. Nabhitabhata leg. // THNHM-I- 10375 ”; 1 ♀ (THNHM), “ larva collected in 2023, Nopphitam Dist., Nakhon Sri Thammarat Prov., Chayaphol Photaworn leg., adult emerged 19. V. 2023. ”	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B8222866745502FF75FA20FBE5F8E5.taxon	description	Description (holotype male). General (Figs. 1 A – 1 B): Body dark brown, surface mildly shiny; ventral surface covered with dense, long reddish-brown setae. Head (Fig. 1 C): Surface mostly dark brown. Clypeus with dense transverse rugopunctures throughout; clypeal apex almost flat; lateral and apical margins of clypeus bordered. Frons with transverse rugopunctures throughout. Transverse carina separating frons and clypeus indicated by an obtuse tubercle at each side. Mandible at outer margin rounded near apex, almost straight at middle. Pronotum (Fig. 1 C): Surface dark brown; sparsely micropunctate throughout, only with a few slightly larger punctures sparsely distributed near lateral margin. impression absent. Side roundly curved, widest near posterior one-fourth; all margins emarginate except narrowly interrupted in lateral margin; anterior margin without any tubercle at middle. Anterior and posterior angles rounded. Scutellum: Surface dark brown; sparsely punctate near base. Elytra: Surface mostly dark brown, except dark reddish sutural ridge; with sparse micropunctures and dense, round, large punctures in striae. Sutural striae narrow, deeply sulcate; other striae wider, even intervals slightly wider than their adjacent odd intervals. Intervals micropunctate, with dense, larger punctures on fourth and sixth intervals posteriorly. Apex densely punctured. Humeral and apical calli obtuse. Epipleuron of elytron moderately dilated horizontally from base to posterior one-fifth, widest behind middle. Metasternum: Surface with dense setae; punctures small and dense close to sides; median region with sparser and smaller, with a longitudinal depression. Abdomen: Pygidium with dark brown surface, slightly convex, with very dense punctures throughout. Abdominal ventrites with dark brown surface, with reddish anterior margin; with sparse micropunctures, larger punctures near base and longitudinal midline, and a row of punctures with long setae at sides. Legs: All legs dark brown, slightly more reddish than other parts of the body. Protibia tridentate, protarsi very strongly thickened, inner protibial claw very strongly enlarged and elongated. Parameres (Figs. 1 D – 1 E): In caudal view (Fig. 1 D), slender and symmetric; without any setae; outer sides with a pair of short, acute, outward and strongly backward processes at posterior one-sixth; inner sides simply curved; apex rounded; in lateral view (Fig. 1 E), apex slightly bent downward. Paratype (male). Characters stable, except punctures on fourth and sixth intervals sparser, lesser in number (Fig. 2 A). Paratype (female). Characters mainly stable, except color much more reddish (it died two days after emerging before full melanization, thus the color is likely not its natural color); transverse carina separating frons and clypeus absent; punctures on fourth and sixth intervals sparser, lesser in number; protarsi not thickened, inner protibial claw not enlarged and elongated (Fig. 2 B – 2 C). Measurements. Body length: male 19.5 – 20.0 mm (holotype 20.0 mm), female 20.7 mm; body width: male 10.3 – 11.0 mm (holotype 10.3 mm), female 11.6 mm.	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B8222866745502FF75FA20FBE5F8E5.taxon	diagnosis	Diagnosis. This species is relatively unique in its appearances and can be easily distinguished from all other species of the genus by absent of the frontal impression and frontal tubercle on the pronotum in both sexes, the combination of the following characters can also distinguish this species from all other species: male with very strongly thickened protarsi and very strongly enlarged and elongated inner protibial claw; fourth and sixth intervals with punctures posteriorly; and the shape of the parameres.	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B8222866745502FF75FA20FBE5F8E5.taxon	etymology	Etymology. This species is named for its limited distribution on the continent, which is currently the only species known exclusively from mainland Southeast Asia.	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B8222866745502FF75FA20FBE5F8E5.taxon	distribution	Distribution. Malaysia: Malay Peninsula; Thailand.	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B8222866745502FF75FA20FBE5F8E5.taxon	discussion	Remarks. This species belongs to a group who lacks setation on the parameres (the small white dots on the photographs of the parameres (Figs. 1 D – 1 E) are small pieces of dust instead of setae). Other species of this group are: Pseudohomonyx belitungensis Miyake & Yamaya, 1997, P. borneensis Arrow, 1908, P. javanus (Burmeister, 1847), P. loksadoensis Jákl & Zídek, 2017 and P. minor Jákl & Zídek, 2017 (Jákl & Zídek 2017). Pseudohomonyx continentalis Yang & Pathomwattananurak, new species, alongside with P. javanus (more information given in the according section in the catalogue), represents the first records of this genus from mainland Southeast Asia. It is also the only species endemic to the mainland. It should be noted that the female paratype was assembled from pieces after it died before full melanization. Fortunately, characters can still be clearly visible and compared with the holotype.	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B8222866715504FF75FF4CFCC6FB1E.taxon	materials_examined	Type material examined. None. Additional material examined (1 ♁). The Philippines: 1 ♁ (CQZY), Agco Hot Spring, Mt. Apo, Kidapawan City, Cotabato Prov., Mindanao Island, III. 2017, Noel Mohagan leg.	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B8222866715504FF75FF4CFCC6FB1E.taxon	description	Description (male). General (Figs. 3 A – 3 B): Body black, with slight reddish color, surface mildly shiny; ventral surface covered with dense, long reddish-brown setae. Head (Fig. 3 C): Surface mostly black, slight reddish near apex, mandible reddish. Clypeus with dense small punctures throughout, sometimes rugopunctate; clypeal apex almost flat, with a very mild, uplifted, triangular protuberance at middle; lateral and apical margins of clypeus bordered. Frons more sparsely punctate than clypeus, punctures mostly larger, slightly rugopunctate near center. Transverse carina separating frons and clypeus indicated by an obtuse tubercle and a distinct, deep, longitudinal depression behind the tubercle at each side. Mandible simply rounded at outer margin. Pronotum (Fig. 2 C): Surface black; sparsely micropunctate throughout, only with a few slightly larger punctures sparsely distributed near lateral margin. Anterior region with a small, deep, round frontal impression, reaching one-fifth the length of pronotum, and a faint line posterior to impression reaching mid-length. Side roundly curved, widest near posterior one-third; all margins emarginate except widely interrupted in lateral margin; anterior margin with a distinct, large frontal tubercle at middle. Anterior and posterior angle rounded. Scutellum: Surface black; impunctate. Elytra: Surface mostly black, except by dark reddish suture; with sparse micropunctures and dense, round, small punctures in striae. Sutural striae narrow, deeply sulcate; other striae wider, equally wide. Intervals micropunctate, with sparse, larger punctures in sixth intervals posteriorly. Apex moderately punctured. Humeral and apical calli obtuse. Epipleuron of elytron strongly dilated horizontally from base to posterior one-fifth, widest behind middle. Metasternum: Surface with dense setae; punctures large and dense close to sides; median region impunctate, with a longitudinal depression. Abdomen: Pygidium with black surface, slightly convex, with dense punctures throughout except impunctate near apex, punctures denser at sides; punctures mainly small. Abdominal ventrites with black surface, with reddish anterior margin; with sparse micropunctures, larger punctures near base and a row of punctures with long setae at sides. Legs: All legs slightly reddish. Protibia tridentate, protarsi strongly thickened, inner protibial claw moderately enlarged and elongated. Parameres (Figs. 3 D – 3 E): In caudal view (Fig. 3 D), slender and nearly symmetric; apex with many setae-bearing punctures, setae very short; outer sides with a pair of short, acute, outward and backward processes at posterior one-seventh; inner sides sinuate; apex almost flat; in lateral view (Fig. 3 E), apex bent downward. Measurements. Body length: 18.8 mm; body width: 10.7 mm.	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B8222866715504FF75FF4CFCC6FB1E.taxon	discussion	Remarks. The identification of this specimen is based on its characters highly matching the holotype (female) description and the locality. It is most closely related to P. negros Miyake & Yamaya, 1997 due to the strong similarity between characters on the parameres (apex of parameres with setae) and their appearances (frontal impression small and round, frontal tubercle distinct, elytral intervals not densely punctate before middle or impunctate, which are often densely punctate in other species whose parameres bears setae at apex). The setae on the parameres of this species is much shorter than all other Philippine species.	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B8222866705504FF75F96DFD38F81C.taxon	materials_examined	Type locality: “ Mt. Apo, Mindanao Island, Philippines ”. Type depository: Holotype: ♀ (NMSM).	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B8222866705504FF75F96DFD38F81C.taxon	distribution	Distribution: The Philippines: Mindanao Island.	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B8222866705504FF75FAE1FAE6F9DA.taxon	type_taxon	Type species: Pseudohomonyx borneensis Arrow, 1908, by original designation.	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B8222866705504FF75FAE1FAE6F9DA.taxon	discussion	Remarks. A distribution map of the genus based on all available information from literature (Endrődi 1969; Miyake & Yamaya 1997, 1999, 2000; Jákl & Zídek 2017) and the current paper is provided here (Fig. 4).	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B8222866735507FF75FA77FD4FF926.taxon	materials_examined	Type locality: “ Belitung Island, east of Sumatra, Indonesia ”. Type depository: Holotype: ♁ (NMSM); paratype: 1 ♁ (NMSM).	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B8222866735507FF75FA77FD4FF926.taxon	distribution	Distribution: Indonesia: Belitung Island.	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B8222866735506FF75F930FC90FF6B.taxon	materials_examined	Type locality: “ Borneo, Bowring ” Type depository: Neotype: ♁ (BMNH).	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B8222866735506FF75F930FC90FF6B.taxon	distribution	Distribution: Malaysia: Borneo; Indonesia: Kalimantan.	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B8222866725506FF75FEFDFCC2FE6F.taxon	materials_examined	Type locality: “ Genting Highlands, Pahang State, Malaysia ” Type depository: Holotype: ♁ (CAU); paratypes: 1 ♁, 1 ♀ (THNHM).	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B8222866725506FF75FEFDFCC2FE6F.taxon	distribution	Distribution: Malaysia: Malay Peninsula; Thailand.	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B8222866725506FF75FDF9FBD2FC63.taxon	materials_examined	Type locality: “ Kina Balu, N. - Borneo ” Type depository: Lectotype: ♁ (BMNH).	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B8222866725506FF75FDF9FBD2FC63.taxon	distribution	Distribution: Malaysia: Borneo; Indonesia: Kalimantan.	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B8222866725506FF75FDF9FBD2FC63.taxon	discussion	Remarks. As mentioned by Endrődi (1969), a “ P. craceus ♁ - Type ” was found in Prell’s collection (now in MFNB) which is presented in the section of Pseudohomonyx crassus Arrow, 1911. However, it has been identified by Endrődi as P. borneensis Arrow, 1908, with dubious locality of “ Java ”; and without any publication from Prell regarding this specimen, it was excluded from any type series (Endrődi 1969).	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B8222866725506FF75FBF5FDEFFA85.taxon	materials_examined	Type locality: “ Keningau, Sabah, North Borneo ”. Type depository: Holotype: ♁ (NMSM); paratypes: 4 ♁♁, 5 ♀♀ (NMSM).	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B8222866725506FF75FBF5FDEFFA85.taxon	distribution	Distribution: Malaysia: Borneo.	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B8222866725509FF75FA50FF20FEFF.taxon	materials_examined	Type locality: “ Java occident. [western Java], Sukabumi, 2000 ” Type depository: Neotype of Pseudohomonyx javanus javanus (Burmeister, 1847): ♁ (HNHM); Lectotype of Pseudohomonyx javanus sumatrensis (Fairmaire, 1896): ♁ (MNHN); Paratype of Pseudohomonyx javanus sumatrensis (Fairmaire, 1896): 1 [sex unknown] (HNHM) (information regarding HNHM type materials are referenced from a datasheet of Dynastinae collection in HNHM provided by the late Ottó Merkl, private communication, 2020). Material examined (5 ♁♁). Thailand: 1 ♁ (CWP), 5 ° 53 ’ 07.05 ” N, 101 ° 01 ’ 17.06 ” E, Betong, Yala Prov., 24. V. 2022, Suradet Sae-Liang Leg. Malaysia: 4 ♁♁ (EUMJ), Univ. Malaya Field Studies Centre, Ulu Gombak, Daerah Gombak, Selangor State, alt. 250 m, 12 – 18. III. 2009, Y. Senda leg.	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B8222866725509FF75FA50FF20FEFF.taxon	distribution	Distribution: Indonesia: Sumatra Island, Nias Island, Java Island; Malaysia (new record); Thailand (new record).	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B82228667D5509FF75FE49FDA6FDC0.taxon	materials_examined	Type locality: “ Indonesia, SE kalimantan, Kandangan District, Loksado village env. ” Type depository: Holotype: ♁ (CSJ); paratypes: 3 ♀♀ (CSJ).	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B82228667D5509FF75FE49FDA6FDC0.taxon	distribution	Distribution: Indonesia: Kalimantan.	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B82228667D5509FF75FD0BFD12FC63.taxon	materials_examined	Type locality: “ Mt. Banahaw, Luzon Is., Philippines ”. Type depository: Holotype: ♁ (NMSM); paratypes: 2 ♀♀ (NMSM).	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B82228667D5509FF75FD0BFD12FC63.taxon	distribution	Distribution: The Philippines: Luzon Island.	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B82228667D5509FF75FBF5FD08FA86.taxon	materials_examined	Type locality: “ Mt. Halcon, Mindro [Mindoro] Is., Philippines ”. Type depository: Holotype: ♁ (NMSM).	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B82228667D5509FF75FBF5FD08FA86.taxon	distribution	Distribution: The Philippines: Mindoro Island.	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B82228667D5509FF75FA50FD40F97F.taxon	materials_examined	Type locality: “ Indonesia, Sumatra Island, West Sumatra Province, Mt. Sanggul, Landai village env., 1250 m m alt. ” Type depository: Holotype: ♁ (CSJ); paratypes: 9 – 14 ♁♁, 16 – 22 ♀♀ [as mentioned in Jákl & Zídek (2017)] (CSJ).	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B82228667D5509FF75FA50FD40F97F.taxon	distribution	Distribution: Indonesia: Sumatra Island.	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B82228667D5508FF75F8C9FD1BFF0F.taxon	materials_examined	Type locality: “ Mt. Canlaon, Mambcal, Negros Island, Philippines ”. Type depository: Holotype: ♁ (NMSM), paratypes: 9 ♁♁, 18 ♀♀ (NMSM).	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B82228667D5508FF75F8C9FD1BFF0F.taxon	distribution	Distribution: The Philippines: Negros Island.	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B82228667C5508FF75FED9FD0DFDB2.taxon	materials_examined	Type locality: “ Palawan (Süd · Palawan) [southern Palawan] ” Type depository: Holotype: ♁ (MFNB).	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B82228667C5508FF75FED9FD0DFDB2.taxon	distribution	Distribution: The Philippines: Palawan Island.	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B82228667C5508FF75FD84FD12FCF3.taxon	materials_examined	Type locality: “ Samar Island, Philippines ”. Type depository: Holotype: ♁ (NMSM).	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B82228667C5508FF75FD84FD12FCF3.taxon	distribution	Distribution: The Philippines: Samar Island.	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B82228667C5508FF75FC45FDEFFB11.taxon	materials_examined	Type locality: “ Mt. Tangga Saran, 400 – 900 m alt., S. W. Engkili, Sarawak, Borneo Is. ” Type depository: Holotype: ♁ (NMSM).	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
03B82228667C5508FF75FC45FDEFFB11.taxon	distribution	Distribution: Malaysia: Borneo.	en	Yang, Qiao-Zhi, Geshev, Georgi, Pathomwattananurak, Wuttipon (2023): A new species of the genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with first description of P. apoensis Miyake & Yamaya, 1997 male and a catalogue of the genus. Zootaxa 5361 (1): 114-124, DOI: 10.11646/zootaxa.5361.1.6, URL: https://www.mapress.com/zt/article/download/zootaxa.5361.1.6/52164
